527 lines
83 KiB
XML
527 lines
83 KiB
XML
<document id="E184B0F6942DC8E025F1230CBF77CF85" ID-CLB-Dataset="39268" ID-DOI="10.11646/zootaxa.4058.1.5" ID-GBIF-Dataset="1735f2e0-7ff1-4e3d-9cea-8cfcfe47da56" ID-ISSN="1175-5326" ID-Zenodo-Dep="241131" ID-ZooBank="605E2A8F-6CC5-4037-90C7-BA3AE70F8ED6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461358083118" checkinUser="plazi" docAuthor="Komai, Tomoyuki" docDate="2015" docId="03908813FFB1FFCE24E9FD9DB0898E8E" docLanguage="en" docName="zt04058p111.pdf" docOrigin="Zootaxa 4058 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Alpheus ikedosoma Komai, 2015, n. sp." docType="treatment" docVersion="9" lastPageNumber="109" masterDocId="FFA9F06BFFB0FFC6247EFFF9B5148F5E" masterDocTitle="A new species of the snapping shrimp genus Alpheus (Crustacea: Decapoda: Caridea: Alpheidae) from Japan, associated with the innkeeper worm Ikedosoma elegans (Annelida: Echiura: Echiuridae)" masterLastPageNumber="111" masterPageNumber="101" pageNumber="102" updateTime="1698661618968" updateUser="ExternalLinkService">
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<mods:mods id="3C00A0E50A23BAC98CE8F0EAAF9DADA2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="860CF71F2AD6745309374FAFC42EBC2A">
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<mods:title id="F2F54E58CB83CFC85554B5DE60ECD783">A new species of the snapping shrimp genus Alpheus (Crustacea: Decapoda: Caridea: Alpheidae) from Japan, associated with the innkeeper worm Ikedosoma elegans (Annelida: Echiura: Echiuridae)</mods:title>
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</mods:titleInfo>
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<mods:name id="654747B2307C7CF7ADEADCB54768F070" type="personal">
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<mods:role id="C208E6CE9817B3AEA8F14696E0693EDA">
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<mods:roleTerm id="AAE62A02474DDF93E2CDF8131553E639">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="413D0B52A4F9E7D517E77869DC32231B">Komai, Tomoyuki</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="271A490CD5F3D167C8944467922DB9A9">text</mods:typeOfResource>
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<mods:titleInfo id="0BE9DCFADB09FBE950F2C153B992392A">
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<mods:title id="66D34C438A503BA1E01DD0EC90145BE0">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="E170B284D2AD6305EFECCBFFBD14443C">
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<mods:date id="FDEBEDAFB47509EDDEAFA217C317EB20">2015</mods:date>
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<mods:detail id="561F11395B209EE5745088CABF2AB1FD" type="volume">
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<mods:number id="404311F446CDB394C17B20E48D5B581F">4058</mods:number>
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</mods:detail>
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<mods:detail id="CFE501A7857E523B3C6D470B67713880" type="issue">
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<mods:number id="9DA56570C577C14582606B27A504523A">1</mods:number>
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</mods:detail>
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<mods:extent id="E351B46DFF83060A3BF6B00A4D480DE1" unit="page">
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<mods:start id="C557963BD512AC4A6217031B1C2FEA5F">101</mods:start>
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<mods:end id="15A90B2F836FD3FD881DD20E75B43C21">111</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="778C42737A854B80DF4B02DF8368F1A7">journal article</mods:classification>
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<mods:identifier id="8EF084673AF2883B5EE2589961A4E0C6" type="CLB-Dataset">39268</mods:identifier>
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<mods:identifier id="D14441DF1C58A448DB5E7DA9D2F27837" type="DOI">10.11646/zootaxa.4058.1.5</mods:identifier>
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<mods:identifier id="A0E70B29023523BCDE9682C26983D13A" type="GBIF-Dataset">1735f2e0-7ff1-4e3d-9cea-8cfcfe47da56</mods:identifier>
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<mods:identifier id="97A8EBFDD2A92889C1E6BF8E77E30565" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="47FD6BB8B7F6E82EE28DA8B6631D5CF4" type="Zenodo-Dep">241131</mods:identifier>
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<mods:identifier id="47D33B78D2AAAAAAC7E05CF822166220" type="ZooBank">605E2A8F-6CC5-4037-90C7-BA3AE70F8ED6</mods:identifier>
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</mods:mods>
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<treatment id="03908813FFB1FFCE24E9FD9DB0898E8E" ID-DOI="http://doi.org/10.5281/zenodo.6100386" ID-GBIF-Taxon="119647826" ID-Zenodo-Dep="6100386" LSID="urn:lsid:plazi:treatment:03908813FFB1FFCE24E9FD9DB0898E8E" httpUri="http://treatment.plazi.org/id/03908813FFB1FFCE24E9FD9DB0898E8E" lastPageId="8" lastPageNumber="109" pageId="1" pageNumber="102">
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<subSubSection id="C3236A8EFFB1FFC724E9FD9DB4148D9D" pageId="1" pageNumber="102" type="nomenclature">
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<paragraph id="8B863905FFB1FFC724E9FD9DB4D88D20" blockId="1.[151,712,612,707]" box="[151,460,612,638]" pageId="1" pageNumber="102">
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<heading id="D0CE8E69FFB1FFC724E9FD9DB4D88D20" bold="true" box="[151,460,612,638]" fontSize="11" level="1" pageId="1" pageNumber="102" reason="1">
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<emphasis id="B94DE517FFB1FFC724E9FD9DB4D88D20" bold="true" box="[151,460,612,638]" pageId="1" pageNumber="102">
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<taxonomicName id="4C394286FFB1FFC724E9FD9DB4968D20" ID-CoL="85LJX" box="[151,386,612,638]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="102" phylum="Arthropoda" rank="species" species="ikedosoma" status="sp. nov.">
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<emphasis id="B94DE517FFB1FFC724E9FD9DB4968D20" bold="true" box="[151,386,612,638]" italics="true" pageId="1" pageNumber="102">Alpheus ikedosoma</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27E586CFFB1FFC725F7FD9DB4D88D20" box="[393,460,612,638]" pageId="1" pageNumber="102" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B863905FFB1FFC724E9FD7EB4148D9D" blockId="1.[151,712,612,707]" pageId="1" pageNumber="102">
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[New Japanese name: Yume-yumushi-teppou-ebi]
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<figureCitation id="13022580FFB1FFC724E9FD53B4148D9D" box="[151,256,682,707]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1517,1539]" captionStartId-1="4.[151,250,1818,1840]" captionStartId-2="5.[151,250,1837,1859]" captionStartId-3="6.[151,250,1907,1929]" captionStartId-4="7.[151,250,878,900]" captionTargetBox-0="[227,1359,325,1496]" captionTargetBox-1="[163,1421,212,1779]" captionTargetBox-2="[241,1337,208,1802]" captionTargetBox-3="[200,1354,198,1853]" captionTargetBox-4="[154,1431,201,847]" captionTargetId-0="figure@2.[226,1360,325,1496]" captionTargetId-1="figure@4.[151,1436,193,1797]" captionTargetId-2="figure@5.[225,1348,193,1812]" captionTargetId-3="figure@6.[195,1367,193,1868]" captionTargetId-4="figure@7.[151,1436,193,857]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURE 1. Alpheus ikedosoma n. sp. A, holotype, male (cl 8.8 mm), CBM-ZC 13128; B, allotype, female (cl 11.1 mm), spouse of male holotype, CBM-ZC 13129." captionText-1="FIGURE 2. Alpheus ikedosoma n. sp., holotype, male (cl 8.8 mm), CBM-ZC 13128. A, carapace and cephalic appendages, lateral view; B, anterior part of carapace and left antennular and antennal peduncle, lateral view; C, close up of rostrum and left eye, lateral view; D, anterior part of carapace, and antennular and antennal peduncles, dorsal view (marginal setae on left antennal scale omitted); E, posterior margin of carapace, lateral view; F, telson, dorsal view (setae omitted); G, left antennal scale, dorsal view (setae omitted); H, appendices internal and masculina on left second pleopod, dorsal view; I, right uropod, dorsal view (perpendicular) (marginal setae on rami omitted). Scale bars: 2 mm for A; 1 mm for B, D – G, I; 0.5 mm for C, H." captionText-2="FIGURE 3. Alpheus ikedosoma n. sp., holotype, male (cl 8.8 mm), CBM-ZC 13128, left thoracic appendages. A, third maxilliped, lateral view; B, same, coxa, exopod and antepenultimate segment of endopod, ventral view (setae omitted); C, same, penultimate segment (= carpus), ventral view; D, second pereopod, lateral view; E, third pereopod, lateral view; F, same, ischium, ventral view; G, same, propodus and dactylus, lateral view; H, same, distal part of propodus and dactylus, flexor view; I, fourth pereopod, lateral view; J, same, ischium, ventral view; K, fifth pereopod, lateral view. Scale bars: 1 mm for A – F, I – K; 0.5 mm for G, H." captionText-3="FIGURE 4. Alpheus ikedosoma n. sp., holotype, male (cl 8.8 mm), CBM-ZC 13128. A, left major cheliped (first pereopod), ischium to carpus, lateral view; B, same, mesial view; C, same pereopod, chela, dorsal (outer) view; D, same, mesial (upper) view; E, same, lateral (lower) view; F, right minor cheliped, lateral view; G, same, mesial view; H, same pereopod, chela, upper view; I, same, lower view. Scale bars: 2 mm." captionText-4="FIGURE 5. Alpheus ikedosoma n. sp., paratype, ovigerous female (cl 12.2 mm), CBM-ZC 13135. A, left major cheliped, ischium and merus, mesial view; B, same pereopod, chela, lateral view; C, right minor cheliped, mesial view; D, same pereopod, chela, mesial view. Scale bars: 2 mm." httpUri-0="https://zenodo.org/record/241132/files/figure.png" httpUri-1="https://zenodo.org/record/241133/files/figure.png" httpUri-2="https://zenodo.org/record/241134/files/figure.png" httpUri-3="https://zenodo.org/record/241135/files/figure.png" httpUri-4="https://zenodo.org/record/241136/files/figure.png" pageId="1" pageNumber="102">Figs. 1–5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3236A8EFFB1FFC724E9FD0BB1068BE5" pageId="1" pageNumber="102" type="materials_examined">
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<paragraph id="8B863905FFB1FFC724E9FD0BB1048C0D" blockId="1.[151,1437,754,2003]" pageId="1" pageNumber="102">
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<emphasis id="B94DE517FFB1FFC724E9FD0BB46D8C55" bold="true" box="[151,377,754,779]" pageId="1" pageNumber="102">Material examined</emphasis>
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. All the specimens examined were extracted from burrows of echiuran worm near water edge at low tide by using yabby pump.
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<typeStatus id="548287A7FFB1FFC72665FCEFB79C8C71" box="[539,648,790,815]" pageId="1" pageNumber="102" type="holotype">Holotype</typeStatus>
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: Hojyo Beach, Tateyama, Boso Peninsula, central
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<collectingCountry id="F32E7995FFB1FFC720BAFCEEB0138C70" box="[1220,1287,791,814]" name="Japan" pageId="1" pageNumber="102">Japan</collectingCountry>
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,
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<geoCoordinate id="EE0D5FC2FFB1FFC7216DFCEFB0838C71" box="[1299,1431,790,815]" direction="north" orientation="latitude" pageId="1" pageNumber="102" precision="9" value="35.001835">35°00.11’N</geoCoordinate>
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,
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<geoCoordinate id="EE0D5FC2FFB1FFC724E9FCC3B43C8C0D" box="[151,296,826,851]" direction="east" orientation="longitude" pageId="1" pageNumber="102" precision="9" value="139.855">139°51.30’E</geoCoordinate>
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, coll. T. Komai,
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<date id="FF871FC5FFB1FFC7259DFCC2B7628C0D" box="[483,630,826,851]" pageId="1" pageNumber="102" value="2015-07-30">30 July 2015</date>
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, male (cl
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<quantity id="4CC194E0FFB1FFC7269DFCC2B6298C0D" box="[739,829,827,851]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="1" pageNumber="102" unit="mm" value="8.8">8.8 mm</quantity>
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), CBM-ZC 13128.
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</paragraph>
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<paragraph id="8B863905FFB1FFC724B9FCA7B16A8C29" blockId="1.[151,1437,754,2003]" box="[199,1150,862,887]" pageId="1" pageNumber="102">
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Allotype: spouse of male
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<typeStatus id="548287A7FFB1FFC72598FCA7B75F8C29" box="[486,587,862,887]" pageId="1" pageNumber="102" type="holotype">holotype</typeStatus>
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, ovigerous female (cl
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<quantity id="4CC194E0FFB1FFC7273AFCA6B6BF8C29" box="[836,939,863,887]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="1" pageNumber="102" unit="mm" value="11.1">11.1 mm</quantity>
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), CBM-ZC 13129.
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</paragraph>
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<paragraph id="8B863905FFB1FFC724B9FC7AB1068BE5" blockId="1.[151,1437,754,2003]" pageId="1" pageNumber="102">
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<typeStatus id="548287A7FFB1FFC724B9FC7AB42D8CC5" box="[199,313,899,923]" pageId="1" pageNumber="102" type="paratype">Paratypes</typeStatus>
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: Tatara Beach, Minami-Boso, Boso Peninsula,
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<geoCoordinate id="EE0D5FC2FFB1FFC72737FC7BB6DB8CC5" box="[841,975,898,923]" direction="north" orientation="latitude" pageId="1" pageNumber="102" precision="9" value="35.036167">35°02.17’N</geoCoordinate>
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,
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<geoCoordinate id="EE0D5FC2FFB1FFC727A5FC7AB1798CC5" box="[987,1133,899,923]" direction="east" orientation="longitude" pageId="1" pageNumber="102" precision="9" value="139.8215">139°49.29’E</geoCoordinate>
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, coll. T. Komai,
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<date id="FF871FC5FFB1FFC72156FC7AB5C68CE1" pageId="1" pageNumber="102" value="2013-08-22">22 August 2013</date>
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,
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<specimenCount id="9D3FF28CFFB1FFC724A3FC5EB43C8CE1" box="[221,296,934,959]" pageId="1" pageNumber="102" type="male">1 male</specimenCount>
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(cl 8.0 mm), 1 ovigerous female (cl
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<quantity id="4CC194E0FFB1FFC726BAFC5FB60A8CE1" box="[708,798,934,959]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="1" pageNumber="102" unit="mm" value="9.6">9.6 mm</quantity>
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), CBM-ZC 13130; same data as
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<typeStatus id="548287A7FFB1FFC720F3FC5FB1FB8CE1" box="[1165,1263,934,959]" pageId="1" pageNumber="102" type="holotype">holotype</typeStatus>
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and allotype,
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<specimenCount id="9D3FF28CFFB1FFC721F0FC5EB5D98CBD" pageId="1" pageNumber="102" type="male">1 male</specimenCount>
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(cl
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<quantity id="4CC194E0FFB1FFC72489FC32B4458CBD" box="[247,337,971,995]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="1" pageNumber="102" unit="mm" value="8.4">8.4 mm</quantity>
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),
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<specimenCount id="9D3FF28CFFB1FFC7251DFC32B4D08CBD" box="[355,452,970,995]" pageId="1" pageNumber="102" type="female">1 female</specimenCount>
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(cl
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<quantity id="4CC194E0FFB1FFC72590FC32B7418CBD" box="[494,597,971,995]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="1" pageNumber="102" unit="mm" value="11.2">11.2 mm</quantity>
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), CBM-ZC 13131; same locality as
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<typeStatus id="548287A7FFB1FFC7279BFC33B1538CBD" box="[997,1095,970,995]" pageId="1" pageNumber="102" type="holotype">holotype</typeStatus>
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and allotype,
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<date id="FF871FC5FFB1FFC7209AFC32B08C8CBD" box="[1252,1432,970,995]" pageId="1" pageNumber="102" value="2015-08-31">31 August 2015</date>
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, coll. M. Tanaka,
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<specimenCount id="9D3FF28CFFB1FFC72523FC16B4BF8B59" box="[349,427,1006,1031]" pageId="1" pageNumber="102" type="male">1 male</specimenCount>
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(cl
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<quantity id="4CC194E0FFB1FFC725A3FC17B72E8B59" box="[477,570,1006,1031]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="1" pageNumber="102" unit="mm" value="9.6">9.6 mm</quantity>
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), 1 ovigerous female (cl
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<quantity id="4CC194E0FFB1FFC72721FC16B6A88B59" box="[863,956,1007,1031]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="1" pageNumber="102" unit="mm" value="9.7">9.7 mm</quantity>
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), CBM-ZC 13132; Hondo flat, Amakusa, Kumamoto Prefecture, Ariake Sea, Kyushu,
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<geoCoordinate id="EE0D5FC2FFB1FFC726ECFBEAB60C8B75" box="[658,792,1043,1067]" direction="north" orientation="latitude" pageId="1" pageNumber="102" precision="9" value="32.4515">32°27.09’N</geoCoordinate>
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,
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<geoCoordinate id="EE0D5FC2FFB1FFC7275BFBEAB6A28B75" box="[805,950,1043,1067]" direction="east" orientation="longitude" pageId="1" pageNumber="102" precision="9" value="130.20317">130°12.19’E</geoCoordinate>
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,
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<date id="FF871FC5FFB1FFC727BCFBEAB1488B75" box="[962,1116,1043,1067]" pageId="1" pageNumber="102" value="2013-05-11">11 May 2013</date>
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,
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<specimenCount id="9D3FF28CFFB1FFC72016FBEAB1D48B75" box="[1128,1216,1042,1067]" pageId="1" pageNumber="102" type="male">3 males</specimenCount>
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(cl
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<quantity id="4CC194E0FFB1FFC72090FBEBB06E8B75" box="[1262,1402,1042,1067]" metricMagnitude="-3" metricUnit="m" metricValue="7.35" metricValueMax="8.2" metricValueMin="6.5" pageId="1" pageNumber="102" unit="mm" value="7.35" valueMax="8.2" valueMin="6.5">6.5–8.2 mm</quantity>
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),
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<specimenCount id="9D3FF28CFFB1FFC721F0FBEBB5FA8B11" pageId="1" pageNumber="102" type="female">6 females</specimenCount>
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(cl
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<quantity id="4CC194E0FFB1FFC7255EFBCFB4A88B11" box="[288,444,1078,1103]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" metricValueMax="10.5" metricValueMin="6.9" pageId="1" pageNumber="102" unit="mm" value="8.7" valueMax="10.5" valueMin="6.9">6.9–10.5 mm</quantity>
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), 1 ovigerous female (cl
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<quantity id="4CC194E0FFB1FFC7269FFBCEB62A8B11" box="[737,830,1079,1103]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="1" pageNumber="102" unit="mm" value="9.4">9.4 mm</quantity>
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), CBM-ZC13133; same data,
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<specimenCount id="9D3FF28CFFB1FFC720E5FBCEB1FD8B11" box="[1179,1257,1078,1103]" pageId="1" pageNumber="102" type="male">1 male</specimenCount>
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(cl 8.0 mm), 1 ovigerous female (cl
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<quantity id="4CC194E0FFB1FFC725F0FBA2B4FF8B2D" box="[398,491,1115,1139]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="1" pageNumber="102" unit="mm" value="9.8">9.8 mm</quantity>
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),
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<collectionCode id="ED28A1C0FFB1FFC7267FFBA2B7428B2C" box="[513,598,1115,1138]" pageId="1" pageNumber="102">KMNH</collectionCode>
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IvR 500845; same locality,
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<date id="FF871FC5FFB1FFC727DCFBA2B1558B2D" box="[930,1089,1115,1139]" pageId="1" pageNumber="102" value="2013-05-12">12 May 2013</date>
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,
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<specimenCount id="9D3FF28CFFB1FFC7202EFBA2B1AB8B2D" box="[1104,1215,1114,1139]" pageId="1" pageNumber="102" type="female">2 females</specimenCount>
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||
(cl 9.7,
|
||
<quantity id="4CC194E0FFB1FFC7215BFBA2B0848B2D" box="[1317,1424,1115,1139]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="1" pageNumber="102" unit="mm" value="10.7">10.7 mm</quantity>
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), CBM-ZC 13134; Okoshiki Beach, Uto, Kumamoto Prefecture, Ariake Sea,
|
||
<geoCoordinate id="EE0D5FC2FFB1FFC72075FB87B1868BC9" box="[1035,1170,1150,1175]" direction="north" orientation="latitude" pageId="1" pageNumber="102" precision="9" value="32.65917">32°39.55’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0D5FC2FFB1FFC720DFFB86B0268BC9" box="[1185,1330,1151,1175]" direction="east" orientation="longitude" pageId="1" pageNumber="102" precision="9" value="130.5355">130°32.13’E</geoCoordinate>
|
||
,
|
||
<date id="FF871FC5FFB1FFC7213CFB86B5C68BE5" pageId="1" pageNumber="102" value="2013-05-13">13 May 2013</date>
|
||
, coll. T. Komai, 2 ovigerous females (cl 11.7,
|
||
<quantity id="4CC194E0FFB1FFC726A9FB5AB62B8BE5" box="[727,831,1187,1211]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="1" pageNumber="102" unit="mm" value="12.2">12.2 mm</quantity>
|
||
), CBM-ZC 13135.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3236A8EFFB1FFC124B9FB3FB4758B87" lastPageId="7" lastPageNumber="108" pageId="1" pageNumber="102" type="description">
|
||
<paragraph id="8B863905FFB1FFC724B9FB3FB1EE8B81" blockId="1.[151,1437,754,2003]" box="[199,1274,1222,1247]" pageId="1" pageNumber="102">
|
||
<emphasis id="B94DE517FFB1FFC724B9FB3FB4458B81" bold="true" box="[199,337,1222,1247]" pageId="1" pageNumber="102">Description</emphasis>
|
||
.
|
||
<emphasis id="B94DE517FFB1FFC72521FB3FB4B78B80" box="[351,419,1222,1246]" italics="true" pageId="1" pageNumber="102">Males</emphasis>
|
||
. Body (
|
||
<figureCitation id="13022580FFB1FFC72580FB3FB7288B81" box="[510,572,1222,1247]" captionStart="FIGURE 1" captionStartId="2.[151,250,1517,1539]" captionTargetBox="[227,1359,325,1496]" captionTargetId="figure@2.[226,1360,325,1496]" captionTargetPageId="2" captionText="FIGURE 1. Alpheus ikedosoma n. sp. A, holotype, male (cl 8.8 mm), CBM-ZC 13128; B, allotype, female (cl 11.1 mm), spouse of male holotype, CBM-ZC 13129." httpUri="https://zenodo.org/record/241132/files/figure.png" pageId="1" pageNumber="102">Fig. 1</figureCitation>
|
||
A) relatively slender; integument rather soft, surface glabrous.
|
||
</paragraph>
|
||
<paragraph id="8B863905FFB1FFC724B9FB12B7748AE9" blockId="1.[151,1437,754,2003]" pageId="1" pageNumber="102">
|
||
Rostrum (
|
||
<figureCitation id="13022580FFB1FFC72542FB13B4908A5D" box="[316,388,1258,1283]" captionStart="FIGURE 2" captionStartId="4.[151,250,1818,1840]" captionTargetBox="[163,1421,212,1779]" captionTargetId="figure@4.[151,1436,193,1797]" captionTargetPageId="4" captionText="FIGURE 2. Alpheus ikedosoma n. sp., holotype, male (cl 8.8 mm), CBM-ZC 13128. A, carapace and cephalic appendages, lateral view; B, anterior part of carapace and left antennular and antennal peduncle, lateral view; C, close up of rostrum and left eye, lateral view; D, anterior part of carapace, and antennular and antennal peduncles, dorsal view (marginal setae on left antennal scale omitted); E, posterior margin of carapace, lateral view; F, telson, dorsal view (setae omitted); G, left antennal scale, dorsal view (setae omitted); H, appendices internal and masculina on left second pleopod, dorsal view; I, right uropod, dorsal view (perpendicular) (marginal setae on rami omitted). Scale bars: 2 mm for A; 1 mm for B, D – G, I; 0.5 mm for C, H." httpUri="https://zenodo.org/record/241133/files/figure.png" pageId="1" pageNumber="102">Fig. 2</figureCitation>
|
||
A–D) very short, falling far short of midlength of first segment of antennular peduncle, triangular in dorsal view, slightly to strongly curved downward, distally acute or subacute; dorsal surface strongly sloping, non-carinate. Carapace (
|
||
<figureCitation id="13022580FFB1FFC7266DFACBB74D8A15" box="[531,601,1330,1355]" captionStart="FIGURE 2" captionStartId="4.[151,250,1818,1840]" captionTargetBox="[163,1421,212,1779]" captionTargetId="figure@4.[151,1436,193,1797]" captionTargetPageId="4" captionText="FIGURE 2. Alpheus ikedosoma n. sp., holotype, male (cl 8.8 mm), CBM-ZC 13128. A, carapace and cephalic appendages, lateral view; B, anterior part of carapace and left antennular and antennal peduncle, lateral view; C, close up of rostrum and left eye, lateral view; D, anterior part of carapace, and antennular and antennal peduncles, dorsal view (marginal setae on left antennal scale omitted); E, posterior margin of carapace, lateral view; F, telson, dorsal view (setae omitted); G, left antennal scale, dorsal view (setae omitted); H, appendices internal and masculina on left second pleopod, dorsal view; I, right uropod, dorsal view (perpendicular) (marginal setae on rami omitted). Scale bars: 2 mm for A; 1 mm for B, D – G, I; 0.5 mm for C, H." httpUri="https://zenodo.org/record/241133/files/figure.png" pageId="1" pageNumber="102">Fig. 2</figureCitation>
|
||
A, B, D) smooth, glabrous, moderately compressed laterally; postrostral carina and adrostral grooves absent; dorsum nearly straight in lateral view; orbital hoods not inflated, anteriorly unarmed; anterior margin between rostrum and orbital hoods nearly straight; pterygostomial angle (Fig.) broadly rounded; cardiac notch minute (
|
||
<figureCitation id="13022580FFB1FFC72582FA67B72B8AE9" box="[508,575,1438,1463]" captionStart="FIGURE 2" captionStartId="4.[151,250,1818,1840]" captionTargetBox="[163,1421,212,1779]" captionTargetId="figure@4.[151,1436,193,1797]" captionTargetPageId="4" captionText="FIGURE 2. Alpheus ikedosoma n. sp., holotype, male (cl 8.8 mm), CBM-ZC 13128. A, carapace and cephalic appendages, lateral view; B, anterior part of carapace and left antennular and antennal peduncle, lateral view; C, close up of rostrum and left eye, lateral view; D, anterior part of carapace, and antennular and antennal peduncles, dorsal view (marginal setae on left antennal scale omitted); E, posterior margin of carapace, lateral view; F, telson, dorsal view (setae omitted); G, left antennal scale, dorsal view (setae omitted); H, appendices internal and masculina on left second pleopod, dorsal view; I, right uropod, dorsal view (perpendicular) (marginal setae on rami omitted). Scale bars: 2 mm for A; 1 mm for B, D – G, I; 0.5 mm for C, H." httpUri="https://zenodo.org/record/241133/files/figure.png" pageId="1" pageNumber="102">Fig. 2</figureCitation>
|
||
E).
|
||
</paragraph>
|
||
<paragraph id="8B863905FFB1FFC724B9FA3BB6C589D1" blockId="1.[151,1437,754,2003]" pageId="1" pageNumber="102">
|
||
Pleon (
|
||
<figureCitation id="13022580FFB1FFC72562FA3BB4748A85" box="[284,352,1474,1499]" captionStart="FIGURE 1" captionStartId="2.[151,250,1517,1539]" captionTargetBox="[227,1359,325,1496]" captionTargetId="figure@2.[226,1360,325,1496]" captionTargetPageId="2" captionText="FIGURE 1. Alpheus ikedosoma n. sp. A, holotype, male (cl 8.8 mm), CBM-ZC 13128; B, allotype, female (cl 11.1 mm), spouse of male holotype, CBM-ZC 13129." httpUri="https://zenodo.org/record/241132/files/figure.png" pageId="1" pageNumber="102">Fig. 1</figureCitation>
|
||
A) without distinctive features; pleura broadly rounded marginally; sixth pleomere without articulated flap, preanal area not protruding as tooth. Telson (
|
||
<figureCitation id="13022580FFB1FFC7272EFA1FB6808AA1" box="[848,916,1510,1535]" captionStart="FIGURE 2" captionStartId="4.[151,250,1818,1840]" captionTargetBox="[163,1421,212,1779]" captionTargetId="figure@4.[151,1436,193,1797]" captionTargetPageId="4" captionText="FIGURE 2. Alpheus ikedosoma n. sp., holotype, male (cl 8.8 mm), CBM-ZC 13128. A, carapace and cephalic appendages, lateral view; B, anterior part of carapace and left antennular and antennal peduncle, lateral view; C, close up of rostrum and left eye, lateral view; D, anterior part of carapace, and antennular and antennal peduncles, dorsal view (marginal setae on left antennal scale omitted); E, posterior margin of carapace, lateral view; F, telson, dorsal view (setae omitted); G, left antennal scale, dorsal view (setae omitted); H, appendices internal and masculina on left second pleopod, dorsal view; I, right uropod, dorsal view (perpendicular) (marginal setae on rami omitted). Scale bars: 2 mm for A; 1 mm for B, D – G, I; 0.5 mm for C, H." httpUri="https://zenodo.org/record/241133/files/figure.png" pageId="1" pageNumber="102">Fig. 2</figureCitation>
|
||
F) relatively broad, subtrapezoidal, narrowing posteriorly; lateral margins without concavity; dorsal surface with 2 pairs of minute dorsolateral spines, situated at some distance from lateral margins, first pair located at 0.6 of telson length, second pair at 0.8; posterior margin broadly rounded, with 2 pairs of unequal spines at posterolateral angles (mesial spine much longer than lateral spine), and with row of long plumose setae between posterolateral spines.
|
||
</paragraph>
|
||
<paragraph id="8B863905FFB1FFC724B9F962B5C489A5" blockId="1.[151,1437,754,2003]" pageId="1" pageNumber="102">
|
||
Eyes (
|
||
<figureCitation id="13022580FFB1FFC72571F963B44089ED" box="[271,340,1690,1715]" captionStart="FIGURE 2" captionStartId="4.[151,250,1818,1840]" captionTargetBox="[163,1421,212,1779]" captionTargetId="figure@4.[151,1436,193,1797]" captionTargetPageId="4" captionText="FIGURE 2. Alpheus ikedosoma n. sp., holotype, male (cl 8.8 mm), CBM-ZC 13128. A, carapace and cephalic appendages, lateral view; B, anterior part of carapace and left antennular and antennal peduncle, lateral view; C, close up of rostrum and left eye, lateral view; D, anterior part of carapace, and antennular and antennal peduncles, dorsal view (marginal setae on left antennal scale omitted); E, posterior margin of carapace, lateral view; F, telson, dorsal view (setae omitted); G, left antennal scale, dorsal view (setae omitted); H, appendices internal and masculina on left second pleopod, dorsal view; I, right uropod, dorsal view (perpendicular) (marginal setae on rami omitted). Scale bars: 2 mm for A; 1 mm for B, D – G, I; 0.5 mm for C, H." httpUri="https://zenodo.org/record/241133/files/figure.png" pageId="1" pageNumber="102">Fig. 2</figureCitation>
|
||
A–D) concealed in dorsal view, partially visible in lateral view, anteromesial margin produced, with minute sharp tubercle; cornea small, darkly pigmented. Ocellar beak not conspicuously protruding between eyes.
|
||
</paragraph>
|
||
<paragraph id="8B863905FFB1FFC724B9F8FFB75D88F1" blockId="1.[151,1437,754,2003]" pageId="1" pageNumber="102">
|
||
Antennular peduncle (
|
||
<figureCitation id="13022580FFB1FFC725BBF8FFB71D8841" box="[453,521,1798,1823]" captionStart="FIGURE 2" captionStartId="4.[151,250,1818,1840]" captionTargetBox="[163,1421,212,1779]" captionTargetId="figure@4.[151,1436,193,1797]" captionTargetPageId="4" captionText="FIGURE 2. Alpheus ikedosoma n. sp., holotype, male (cl 8.8 mm), CBM-ZC 13128. A, carapace and cephalic appendages, lateral view; B, anterior part of carapace and left antennular and antennal peduncle, lateral view; C, close up of rostrum and left eye, lateral view; D, anterior part of carapace, and antennular and antennal peduncles, dorsal view (marginal setae on left antennal scale omitted); E, posterior margin of carapace, lateral view; F, telson, dorsal view (setae omitted); G, left antennal scale, dorsal view (setae omitted); H, appendices internal and masculina on left second pleopod, dorsal view; I, right uropod, dorsal view (perpendicular) (marginal setae on rami omitted). Scale bars: 2 mm for A; 1 mm for B, D – G, I; 0.5 mm for C, H." httpUri="https://zenodo.org/record/241133/files/figure.png" pageId="1" pageNumber="102">Fig. 2</figureCitation>
|
||
A, B, D) moderately stout, overreaching distal margin of scaphocerite (antennal scale). First segment with ventromesial carina forming large crest-like lamina; stylocerite somewhat inflated, terminating in blunt or subacute tip, slightly falling short of or reaching distal margin of first peduncular segment. Second segment about 2.0 times as long as broad. Lateral flagellum with numerous tufts of aesthetascs on about 10 articles; secondary ramus rudimentary.
|
||
</paragraph>
|
||
<paragraph id="8B863905FFB1FFC724B9F842B089888D" blockId="1.[151,1437,754,2003]" box="[199,1437,1978,2003]" pageId="1" pageNumber="102">
|
||
Antenna (
|
||
<figureCitation id="13022580FFB1FFC72547F843B494888D" box="[313,384,1978,2003]" captionStart="FIGURE 2" captionStartId="4.[151,250,1818,1840]" captionTargetBox="[163,1421,212,1779]" captionTargetId="figure@4.[151,1436,193,1797]" captionTargetPageId="4" captionText="FIGURE 2. Alpheus ikedosoma n. sp., holotype, male (cl 8.8 mm), CBM-ZC 13128. A, carapace and cephalic appendages, lateral view; B, anterior part of carapace and left antennular and antennal peduncle, lateral view; C, close up of rostrum and left eye, lateral view; D, anterior part of carapace, and antennular and antennal peduncles, dorsal view (marginal setae on left antennal scale omitted); E, posterior margin of carapace, lateral view; F, telson, dorsal view (setae omitted); G, left antennal scale, dorsal view (setae omitted); H, appendices internal and masculina on left second pleopod, dorsal view; I, right uropod, dorsal view (perpendicular) (marginal setae on rami omitted). Scale bars: 2 mm for A; 1 mm for B, D – G, I; 0.5 mm for C, H." httpUri="https://zenodo.org/record/241133/files/figure.png" pageId="1" pageNumber="102">Fig. 2</figureCitation>
|
||
A, B, D) with basicerite unarmed at ventrolateral distal angle or dorsolateral distal angle.
|
||
</paragraph>
|
||
<paragraph id="8B863905FFB2FFC424E9FF6EB6E88E43" blockId="2.[151,1436,151,285]" pageId="2" pageNumber="103">
|
||
Carpocerite stout, overreaching lamella of scaphocerite. Scaphocerite (
|
||
<figureCitation id="13022580FFB2FFC427B7FF6EB11B8FEE" box="[969,1039,151,176]" captionStart="FIGURE 2" captionStartId="4.[151,250,1818,1840]" captionTargetBox="[163,1421,212,1779]" captionTargetId="figure@4.[151,1436,193,1797]" captionTargetPageId="4" captionText="FIGURE 2. Alpheus ikedosoma n. sp., holotype, male (cl 8.8 mm), CBM-ZC 13128. A, carapace and cephalic appendages, lateral view; B, anterior part of carapace and left antennular and antennal peduncle, lateral view; C, close up of rostrum and left eye, lateral view; D, anterior part of carapace, and antennular and antennal peduncles, dorsal view (marginal setae on left antennal scale omitted); E, posterior margin of carapace, lateral view; F, telson, dorsal view (setae omitted); G, left antennal scale, dorsal view (setae omitted); H, appendices internal and masculina on left second pleopod, dorsal view; I, right uropod, dorsal view (perpendicular) (marginal setae on rami omitted). Scale bars: 2 mm for A; 1 mm for B, D – G, I; 0.5 mm for C, H." httpUri="https://zenodo.org/record/241133/files/figure.png" pageId="2" pageNumber="103">Fig. 2</figureCitation>
|
||
G) short, 0.2–0.3 times as long as carapace and about 2.6 times as long as wide, greatest width at proximal 0.2 of length; lateral margin faintly sinuous, terminating in small distolateral tooth slightly falling short of distal margin of lamella; cleft between lamella and distolateral tooth slit-like, deep; lamella broadly rounded distally.
|
||
</paragraph>
|
||
<caption id="DF46698DFFB2FFC424E9FA14B75C897C" httpUri="https://zenodo.org/record/241132/files/figure.png" pageId="2" pageNumber="103" targetBox="[227,1359,325,1496]" targetPageId="2">
|
||
<paragraph id="8B863905FFB2FFC424E9FA14B75C897C" blockId="2.[151,1436,1517,1570]" pageId="2" pageNumber="103">
|
||
<emphasis id="B94DE517FFB2FFC424E9FA14B404895D" bold="true" box="[151,272,1517,1539]" pageId="2" pageNumber="103">FIGURE 1</emphasis>
|
||
.
|
||
<taxonomicName id="4C394286FFB2FFC42560FA14B4F1895D" box="[286,485,1517,1539]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="ikedosoma" status="sp. nov.">
|
||
<emphasis id="B94DE517FFB2FFC42560FA14B4F1895D" box="[286,485,1517,1539]" italics="true" pageId="2" pageNumber="103">Alpheus ikedosoma</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27E586CFFB2FFC42591FA17B73E895D" box="[495,554,1518,1539]" pageId="2" pageNumber="103" rank="species">
|
||
<emphasis id="B94DE517FFB2FFC42591FA17B730895D" bold="true" box="[495,548,1518,1539]" pageId="2" pageNumber="103">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
A, holotype, male (cl 8.8 mm), CBM-ZC 13128; B, allotype, female (cl 11.1 mm), spouse of male holotype, CBM-ZC 13129.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B863905FFB2FFC424B9F9B0B7B68867" blockId="2.[151,1437,1609,2030]" pageId="2" pageNumber="103">
|
||
Mouthparts typical for
|
||
<taxonomicName id="4C394286FFB2FFC425AEF9B3B73E893C" box="[464,554,1610,1634]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="103" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94DE517FFB2FFC425AEF9B3B73E893C" box="[464,554,1610,1634]" italics="true" pageId="2" pageNumber="103">Alpheus</emphasis>
|
||
</taxonomicName>
|
||
(only external observation made, not figured). Third maxilliped (
|
||
<figureCitation id="13022580FFB2FFC42167F9B0B048893C" box="[1305,1372,1609,1634]" captionStart="FIGURE 3" captionStartId="5.[151,250,1837,1859]" captionTargetBox="[241,1337,208,1802]" captionTargetId="figure@5.[225,1348,193,1812]" captionTargetPageId="5" captionText="FIGURE 3. Alpheus ikedosoma n. sp., holotype, male (cl 8.8 mm), CBM-ZC 13128, left thoracic appendages. A, third maxilliped, lateral view; B, same, coxa, exopod and antepenultimate segment of endopod, ventral view (setae omitted); C, same, penultimate segment (= carpus), ventral view; D, second pereopod, lateral view; E, third pereopod, lateral view; F, same, ischium, ventral view; G, same, propodus and dactylus, lateral view; H, same, distal part of propodus and dactylus, flexor view; I, fourth pereopod, lateral view; J, same, ischium, ventral view; K, fifth pereopod, lateral view. Scale bars: 1 mm for A – F, I – K; 0.5 mm for G, H." httpUri="https://zenodo.org/record/241134/files/figure.png" pageId="2" pageNumber="103">Fig. 3</figureCitation>
|
||
A–C) moderately stout, slightly overreaching distal end of antennal carpocerite. Lateral plate on coxa extending to base of exopod, with subacute point. Antepenultimate segment strongly flattened dorsoventrally in proximal half, with sharply carinate lateral and mesial margins; ventral surface with obtuse ridge adjacent to mesial margin. Penultimate segment (= carpus) slightly flattened dorsoventrally, about 2.6 times as long as wide in dorsal or ventral view, with 3 long spiniform setae on distal half of mesial surface. Ultimate segment tapering, trigonal in cross section, tip blunt with tufts of long setae.
|
||
</paragraph>
|
||
<paragraph id="8B863905FFB2FFC524B9F8BCB7C38E3B" blockId="2.[151,1437,1609,2030]" lastBlockId="3.[151,1437,151,2013]" lastPageId="3" lastPageNumber="104" pageId="2" pageNumber="103">
|
||
Major cheliped (
|
||
<figureCitation id="13022580FFB2FFC425FDF8BCB4C58800" box="[387,465,1861,1886]" captionStart="FIGURE 1" captionStartId="2.[151,250,1517,1539]" captionTargetBox="[227,1359,325,1496]" captionTargetId="figure@2.[226,1360,325,1496]" captionTargetPageId="2" captionText="FIGURE 1. Alpheus ikedosoma n. sp. A, holotype, male (cl 8.8 mm), CBM-ZC 13128; B, allotype, female (cl 11.1 mm), spouse of male holotype, CBM-ZC 13129." httpUri="https://zenodo.org/record/241132/files/figure.png" pageId="2" pageNumber="103">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="13022580FFB2FFC4259EF8BFB4FA8800" box="[480,494,1862,1886]" captionStart="FIGURE 4" captionStartId="6.[151,250,1907,1929]" captionTargetBox="[200,1354,198,1853]" captionTargetId="figure@6.[195,1367,193,1868]" captionTargetPageId="6" captionText="FIGURE 4. Alpheus ikedosoma n. sp., holotype, male (cl 8.8 mm), CBM-ZC 13128. A, left major cheliped (first pereopod), ischium to carpus, lateral view; B, same, mesial view; C, same pereopod, chela, dorsal (outer) view; D, same, mesial (upper) view; E, same, lateral (lower) view; F, right minor cheliped, lateral view; G, same, mesial view; H, same pereopod, chela, upper view; I, same, lower view. Scale bars: 2 mm." httpUri="https://zenodo.org/record/241135/files/figure.png" pageId="2" pageNumber="103">4</figureCitation>
|
||
A–E) much larger than minor cheliped. Ischium short, unarmed. Merus moderately stout, broadened distally, trigonal in cross-section; dorsal margin nearly straight, dorsodistal margin not produced; ventrolateral margin faintly sinuous, unarmed; ventromesial margin straight or faintly sinuous, armed usually with small subdistal tooth and 1 minute movable spinule (entirely unarmed in
|
||
<typeStatus id="548287A7FFB2FFC427B6F848B13B8894" box="[968,1071,1969,1994]" pageId="2" pageNumber="103" type="holotype">holotype</typeStatus>
|
||
). Carpus very short, cup-shaped. Chela strongly compressed, oval in cross section, slightly elongate, about 3.0–3.2 times as long as wide, 1.1–1.4 times as long as carapace; lateral margin nearly straight and mesial margin slightly sinuous, greatest width at proximal 0.4 of palm. Palm about 2.2 times as long as wide, without any grooves or depressions on surfaces; surfaces smooth, glabrous, ventral surface slightly inflated. Pollex less than half-length of palm, terminating in upturned, acute tip; dorsal side of socket forming convexity bearing row of prominent setae. Dactylus slightly arcuate in longitudinal plane, not double-headed, tip acuminate, crossed with pollex; plunger much reduced, obliquely truncate, defined only by proximal angle.
|
||
</paragraph>
|
||
<paragraph id="8B863905FFB3FFC524B9FE96B0308DDB" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="104">
|
||
Minor cheliped (
|
||
<figureCitation id="13022580FFB3FFC525FAFE96B4C68ED6" box="[388,466,367,392]" captionStart="FIGURE 1" captionStartId="2.[151,250,1517,1539]" captionTargetBox="[227,1359,325,1496]" captionTargetId="figure@2.[226,1360,325,1496]" captionTargetPageId="2" captionText="FIGURE 1. Alpheus ikedosoma n. sp. A, holotype, male (cl 8.8 mm), CBM-ZC 13128; B, allotype, female (cl 11.1 mm), spouse of male holotype, CBM-ZC 13129." httpUri="https://zenodo.org/record/241132/files/figure.png" pageId="3" pageNumber="104">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="13022580FFB3FFC5259FFE89B4FB8ED6" box="[481,495,368,392]" captionStart="FIGURE 4" captionStartId="6.[151,250,1907,1929]" captionTargetBox="[200,1354,198,1853]" captionTargetId="figure@6.[195,1367,193,1868]" captionTargetPageId="6" captionText="FIGURE 4. Alpheus ikedosoma n. sp., holotype, male (cl 8.8 mm), CBM-ZC 13128. A, left major cheliped (first pereopod), ischium to carpus, lateral view; B, same, mesial view; C, same pereopod, chela, dorsal (outer) view; D, same, mesial (upper) view; E, same, lateral (lower) view; F, right minor cheliped, lateral view; G, same, mesial view; H, same pereopod, chela, upper view; I, same, lower view. Scale bars: 2 mm." httpUri="https://zenodo.org/record/241135/files/figure.png" pageId="3" pageNumber="104">4</figureCitation>
|
||
F–I) not balaeniceps form, reaching level of midlength of left major chela. Ischium similar to that of major cheliped. Merus with dorsal margin faintly sinuous, dorsodistal margin not produced; ventrolateral margin faintly convex; ventromesial margin also faintly convex, small subterminal tooth present or absent, no movable spinules. Carpus short, cup-shaped, similar to that of major cheliped. Chela slender, subcylindrical, about 4.7 times as long as wide, without any depressions or grooves; mesial margin faintly sinuous. Palm about 2.0 times as long as wide, surfaces almost glabrous, mesial face slightly inflated proximally; lateral margin slightly convex. Fingers slender, about 1.5 times as long as palm, with strongly curved, crossing tips; ventral side forming longitudinal excavation; occlusal margins each bordered with thin, chitinous blade.
|
||
</paragraph>
|
||
<paragraph id="8B863905FFB3FFC524B9FD76B62A8DAE" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="104">
|
||
Second pereopod (
|
||
<figureCitation id="13022580FFB3FFC525E9FD76B4CC8DF6" box="[407,472,655,680]" captionStart="FIGURE 3" captionStartId="5.[151,250,1837,1859]" captionTargetBox="[241,1337,208,1802]" captionTargetId="figure@5.[225,1348,193,1812]" captionTargetPageId="5" captionText="FIGURE 3. Alpheus ikedosoma n. sp., holotype, male (cl 8.8 mm), CBM-ZC 13128, left thoracic appendages. A, third maxilliped, lateral view; B, same, coxa, exopod and antepenultimate segment of endopod, ventral view (setae omitted); C, same, penultimate segment (= carpus), ventral view; D, second pereopod, lateral view; E, third pereopod, lateral view; F, same, ischium, ventral view; G, same, propodus and dactylus, lateral view; H, same, distal part of propodus and dactylus, flexor view; I, fourth pereopod, lateral view; J, same, ischium, ventral view; K, fifth pereopod, lateral view. Scale bars: 1 mm for A – F, I – K; 0.5 mm for G, H." httpUri="https://zenodo.org/record/241134/files/figure.png" pageId="3" pageNumber="104">Fig. 3</figureCitation>
|
||
D) slender, overreaching distal end of antennal carpocerite by length of chela. Ischium subequal in length to merus; carpus divided into five articles, with approximate ratio 1: 0.4: 0.2: 0.2: 0.4; chela about 0.25 times as long as carpus, fingers longer than palm.
|
||
</paragraph>
|
||
<paragraph id="8B863905FFB3FFC524B9FD05B7318C96" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="104">
|
||
Third pereopod (
|
||
<figureCitation id="13022580FFB3FFC525F8FD05B4DC8C4A" box="[390,456,764,789]" captionStart="FIGURE 3" captionStartId="5.[151,250,1837,1859]" captionTargetBox="[241,1337,208,1802]" captionTargetId="figure@5.[225,1348,193,1812]" captionTargetPageId="5" captionText="FIGURE 3. Alpheus ikedosoma n. sp., holotype, male (cl 8.8 mm), CBM-ZC 13128, left thoracic appendages. A, third maxilliped, lateral view; B, same, coxa, exopod and antepenultimate segment of endopod, ventral view (setae omitted); C, same, penultimate segment (= carpus), ventral view; D, second pereopod, lateral view; E, third pereopod, lateral view; F, same, ischium, ventral view; G, same, propodus and dactylus, lateral view; H, same, distal part of propodus and dactylus, flexor view; I, fourth pereopod, lateral view; J, same, ischium, ventral view; K, fifth pereopod, lateral view. Scale bars: 1 mm for A – F, I – K; 0.5 mm for G, H." httpUri="https://zenodo.org/record/241134/files/figure.png" pageId="3" pageNumber="104">Fig. 3</figureCitation>
|
||
E–H) moderately slender, overreaching distal end of antennal carpocerite by length of dactylus. Ischium with 1 small movable spine on lateral surface ventrally. Merus unarmed, about 7.0 times as long as wide. Carpus unarmed. Propodus 1.1–1.2 times as long as carpus, armed with 2 widely spaced, slender movable spines in proximal half of ventral (flexor) margin and with pair of unequal movable spines at ventrodistal margin. Dactylus subspatulate, terminating in acute tip, about half as long as propodus; extensor surface with only few short setae, flexor surface glabrous.
|
||
</paragraph>
|
||
<paragraph id="8B863905FFB3FFC524B9FC2DB40F8B4E" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="104">
|
||
Fourth pereopod (
|
||
<figureCitation id="13022580FFB3FFC525ECFC2DB4C18CB2" box="[402,469,980,1005]" captionStart="FIGURE 3" captionStartId="5.[151,250,1837,1859]" captionTargetBox="[241,1337,208,1802]" captionTargetId="figure@5.[225,1348,193,1812]" captionTargetPageId="5" captionText="FIGURE 3. Alpheus ikedosoma n. sp., holotype, male (cl 8.8 mm), CBM-ZC 13128, left thoracic appendages. A, third maxilliped, lateral view; B, same, coxa, exopod and antepenultimate segment of endopod, ventral view (setae omitted); C, same, penultimate segment (= carpus), ventral view; D, second pereopod, lateral view; E, third pereopod, lateral view; F, same, ischium, ventral view; G, same, propodus and dactylus, lateral view; H, same, distal part of propodus and dactylus, flexor view; I, fourth pereopod, lateral view; J, same, ischium, ventral view; K, fifth pereopod, lateral view. Scale bars: 1 mm for A – F, I – K; 0.5 mm for G, H." httpUri="https://zenodo.org/record/241134/files/figure.png" pageId="3" pageNumber="104">
|
||
Fig.
|
||
<date id="FF871FC5FFB3FFC525B9FC2DB4C18CB2" box="[455,469,980,1004]" pageId="3" pageNumber="104">3</date>
|
||
</figureCitation>
|
||
I, J) similar to third pereopod, but slightly shorter, not reaching distal end of antennal carpocerite.
|
||
</paragraph>
|
||
<paragraph id="8B863905FFB3FFC524B9FBE5B00B8BFE" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="104">Fifth pereopod (Fig. K) shorter and slenderer than third and fourth pereopods. Ischium unarmed or armed with 1 movable spine on lateral face ventrally. Propodus with grooming brush consisting of several transverse rows of stiff setulose setae on distal 0.6 length, one small spiniform seta present at ventrodistal angle (usually concealed by grooming setae, not visible in lateral view); dactylus narrower than those of third and fourth pereopods.</paragraph>
|
||
<paragraph id="8B863905FFB3FFC524B9FB55B7C38BB6" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="104">
|
||
Male second pleopod with appendix masculina (
|
||
<figureCitation id="13022580FFB3FFC52686FB55B6298B9A" box="[760,829,1196,1221]" captionStart="FIGURE 2" captionStartId="4.[151,250,1818,1840]" captionTargetBox="[163,1421,212,1779]" captionTargetId="figure@4.[151,1436,193,1797]" captionTargetPageId="4" captionText="FIGURE 2. Alpheus ikedosoma n. sp., holotype, male (cl 8.8 mm), CBM-ZC 13128. A, carapace and cephalic appendages, lateral view; B, anterior part of carapace and left antennular and antennal peduncle, lateral view; C, close up of rostrum and left eye, lateral view; D, anterior part of carapace, and antennular and antennal peduncles, dorsal view (marginal setae on left antennal scale omitted); E, posterior margin of carapace, lateral view; F, telson, dorsal view (setae omitted); G, left antennal scale, dorsal view (setae omitted); H, appendices internal and masculina on left second pleopod, dorsal view; I, right uropod, dorsal view (perpendicular) (marginal setae on rami omitted). Scale bars: 2 mm for A; 1 mm for B, D – G, I; 0.5 mm for C, H." httpUri="https://zenodo.org/record/241133/files/figure.png" pageId="3" pageNumber="104">Fig. 2</figureCitation>
|
||
H) subequal in length to appendix interna, furnished with numerous stiff setae on dorsal surface to apex.
|
||
</paragraph>
|
||
<paragraph id="8B863905FFB3FFC524B9FB0DB7DF8A26" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="104">
|
||
Uropod (
|
||
<figureCitation id="13022580FFB3FFC52551FB0DB4638A52" box="[303,375,1268,1293]" captionStart="FIGURE 2" captionStartId="4.[151,250,1818,1840]" captionTargetBox="[163,1421,212,1779]" captionTargetId="figure@4.[151,1436,193,1797]" captionTargetPageId="4" captionText="FIGURE 2. Alpheus ikedosoma n. sp., holotype, male (cl 8.8 mm), CBM-ZC 13128. A, carapace and cephalic appendages, lateral view; B, anterior part of carapace and left antennular and antennal peduncle, lateral view; C, close up of rostrum and left eye, lateral view; D, anterior part of carapace, and antennular and antennal peduncles, dorsal view (marginal setae on left antennal scale omitted); E, posterior margin of carapace, lateral view; F, telson, dorsal view (setae omitted); G, left antennal scale, dorsal view (setae omitted); H, appendices internal and masculina on left second pleopod, dorsal view; I, right uropod, dorsal view (perpendicular) (marginal setae on rami omitted). Scale bars: 2 mm for A; 1 mm for B, D – G, I; 0.5 mm for C, H." httpUri="https://zenodo.org/record/241133/files/figure.png" pageId="3" pageNumber="104">
|
||
Fig.
|
||
<date id="FF871FC5FFB3FFC52516FB0DB4638A52" box="[360,375,1268,1292]" pageId="3" pageNumber="104">2</date>
|
||
</figureCitation>
|
||
I) with protopod armed with 2 large acute distal teeth. Exopod with slightly convex lateral margin, terminating in small posterolateral tooth, with adjacent 2 movable spinules; diaeresis faintly trilobate; distal margin with row of minute spinules dorsal to main setal fringe. Endopod with scattered distal margin with row of minute spinules dorsal to main setal fringe.
|
||
</paragraph>
|
||
<paragraph id="8B863905FFB3FFC524B9FA7DB79B8AC2" blockId="3.[151,1437,151,2013]" box="[199,655,1412,1437]" pageId="3" pageNumber="104">
|
||
Gill/exopod formula typical for
|
||
<taxonomicName id="4C394286FFB3FFC52650FA7DB79C8AC2" box="[558,648,1412,1436]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="104" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94DE517FFB3FFC52650FA7DB79C8AC2" box="[558,648,1412,1436]" italics="true" pageId="3" pageNumber="104">Alpheus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B863905FFB3FFC524B9FA51B5C2890E" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="104">
|
||
<emphasis id="B94DE517FFB3FFC524B9FA51B4328A9E" box="[199,294,1448,1472]" italics="true" pageId="3" pageNumber="104">Females</emphasis>
|
||
. Non-spawning females generally similar to males. In spawning molt, rostrum acute to blunt; dorsum of carapace becoming weakly convex along with development of ovary. Pleon fairly widened (
|
||
<figureCitation id="13022580FFB3FFC5209BFA35B0338ABA" box="[1253,1319,1484,1509]" captionStart="FIGURE 1" captionStartId="2.[151,250,1517,1539]" captionTargetBox="[227,1359,325,1496]" captionTargetId="figure@2.[226,1360,325,1496]" captionTargetPageId="2" captionText="FIGURE 1. Alpheus ikedosoma n. sp. A, holotype, male (cl 8.8 mm), CBM-ZC 13128; B, allotype, female (cl 11.1 mm), spouse of male holotype, CBM-ZC 13129." httpUri="https://zenodo.org/record/241132/files/figure.png" pageId="3" pageNumber="104">Fig. 1</figureCitation>
|
||
B). Major cheliped (
|
||
<figureCitation id="13022580FFB3FFC5257BFA16B4528956" box="[261,326,1519,1544]" captionStart="FIGURE 5" captionStartId="7.[151,250,878,900]" captionTargetBox="[154,1431,201,847]" captionTargetId="figure@7.[151,1436,193,857]" captionTargetPageId="7" captionText="FIGURE 5. Alpheus ikedosoma n. sp., paratype, ovigerous female (cl 12.2 mm), CBM-ZC 13135. A, left major cheliped, ischium and merus, mesial view; B, same pereopod, chela, lateral view; C, right minor cheliped, mesial view; D, same pereopod, chela, mesial view. Scale bars: 2 mm." httpUri="https://zenodo.org/record/241136/files/figure.png" pageId="3" pageNumber="104">Fig. 5</figureCitation>
|
||
A, B) comparatively small, with chela 0.8–1.0 times as long as carapace; tip of fingers blunt. Minor chela (
|
||
<figureCitation id="13022580FFB3FFC5249FF9EDB4378973" box="[225,291,1556,1581]" captionStart="FIGURE 5" captionStartId="7.[151,250,878,900]" captionTargetBox="[154,1431,201,847]" captionTargetId="figure@7.[151,1436,193,857]" captionTargetPageId="7" captionText="FIGURE 5. Alpheus ikedosoma n. sp., paratype, ovigerous female (cl 12.2 mm), CBM-ZC 13135. A, left major cheliped, ischium and merus, mesial view; B, same pereopod, chela, lateral view; C, right minor cheliped, mesial view; D, same pereopod, chela, mesial view. Scale bars: 2 mm." httpUri="https://zenodo.org/record/241136/files/figure.png" pageId="3" pageNumber="104">Fig. 5</figureCitation>
|
||
C, D) stouter than in males, with palm about 1.7 times as long as wide; fingers 1.3–1.4 times as long as palm.
|
||
</paragraph>
|
||
<paragraph id="8B863905FFB3FFC524B9F9A5B1CC892A" blockId="3.[151,1437,151,2013]" box="[199,1240,1628,1653]" pageId="3" pageNumber="104">
|
||
<emphasis id="B94DE517FFB3FFC524B9F9A5B5E3892B" bold="true" box="[199,247,1628,1653]" pageId="3" pageNumber="104">Size</emphasis>
|
||
. Largest male cl
|
||
<quantity id="4CC194E0FFB3FFC525C6F9A5B705892A" box="[440,529,1628,1652]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="3" pageNumber="104" unit="mm" value="8.8">8.8 mm</quantity>
|
||
, largest female cl
|
||
<quantity id="4CC194E0FFB3FFC526A5F9A5B655892A" box="[731,833,1628,1652]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="3" pageNumber="104" unit="mm" value="12.2">12.2 mm</quantity>
|
||
, ovigerous females cl
|
||
<quantity id="4CC194E0FFB3FFC52042F9A5B1C0892A" box="[1084,1236,1628,1653]" metricMagnitude="-2" metricUnit="m" metricValue="1.0899999999999999" metricValueMax="1.22" metricValueMin="0.96" pageId="3" pageNumber="104" unit="mm" value="10.899999999999999" valueMax="12.2" valueMin="9.6">9.6–12.2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B863905FFB3FFC524B9F986B1F189E3" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="104">
|
||
<emphasis id="B94DE517FFB3FFC524B9F986B4B589C6" bold="true" box="[199,417,1663,1688]" pageId="3" pageNumber="104">Colouration in life</emphasis>
|
||
. Males: Entirely semitranslucent, with tinge of pink on posterior part of each pleomere and distal part of major cheliped; eyes dark gray; yellowish hepatopancrea visible through integument.
|
||
</paragraph>
|
||
<paragraph id="8B863905FFB3FFC524B9F93EB4A0885B" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="104">Females: More pinkish than in males, particularly in anterior part of carapace, pleon and chelipeds; ovary orange; eggs also orange.</paragraph>
|
||
<paragraph id="8B863905FFB3FFC524B9F8F6B42E882E" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="104">
|
||
<emphasis id="B94DE517FFB3FFC524B9F8F6B4218876" bold="true" box="[199,309,1807,1832]" pageId="3" pageNumber="104">Variation</emphasis>
|
||
. As is apparent from the above description, sexual dimorphism influences the stoutness of the body, the shape of the dorsum of the carapace and the relative length of the chelipeds. Females seem to attain larger size than males do.
|
||
</paragraph>
|
||
<paragraph id="8B863905FFB3FFC124B9F885B49B8B14" blockId="3.[151,1437,151,2013]" lastBlockId="7.[151,1436,1001,2034]" lastPageId="7" lastPageNumber="108" pageId="3" pageNumber="104">
|
||
<emphasis id="B94DE517FFB3FFC524B9F885B4D388CB" bold="true" box="[199,455,1916,1941]" pageId="3" pageNumber="104">Symbiotic association</emphasis>
|
||
. All the specimens of shrimp examined were extracted from burrows of echiuran worm constructed on easily accessible intertidal sand beaches or sand flats, with the aid of a bait suction pump. Specimens of the host worm, collected at Boso Peninsula, were identified with
|
||
<emphasis id="B94DE517FFB3FFC52076F83DB1C88882" box="[1032,1244,1988,2012]" italics="true" pageId="3" pageNumber="104">
|
||
<taxonomicName id="4C394286FFB3FFC52076F83DB1958882" box="[1032,1153,1988,2012]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="109" phylum="Arthropoda" rank="species" species="ikedosoma" status="sp. nov.">Ikedosoma</taxonomicName>
|
||
elegans
|
||
</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA844F4FFB3FFC52095F83DB0658882" author="Ikeda" box="[1259,1393,1988,2013]" pageId="3" pageNumber="104" refString="Ikeda, I. (1904) The Gephyrea of Japan. Journal of College of Science, Imperial University of Tokyo, 20, 1 - 87." type="journal article" year="1904">Ikeda, 1904</bibRefCitation>
|
||
) by Dr. Masaatsu Tanaka (Faculty of Science, Toho University) (
|
||
<figureCitation id="13022580FFB7FFC12735FC10B69B8B5C" box="[843,911,1001,1026]" captionStart="FIGURE 6" captionStartId="8.[151,250,1642,1664]" captionTargetBox="[366,1220,522,1620]" captionTargetId="figure@8.[365,1222,499,1621]" captionTargetPageId="8" captionText="FIGURE 6. Host innkeeper worm of Alpheus ikedosoma n. sp. A, Ikedosoma elegans (Ikeda, 1904), CBM-ZW, Hojo Beach, Tateyama, Chiba Prefecture, Japan, 30 July 2015, scale circle: 33 mm; B, unidentified specimen, not preserved, Hondo flat, Amakusa, Kumamto Prefecture, 11 May 2013." httpUri="https://zenodo.org/record/241137/files/figure.png" pageId="7" pageNumber="108">Fig. 6</figureCitation>
|
||
A). Five heterosexual pairs of the present new species were collected from two nearby locations in Boso Peninsula (Tatara and Hojo beaches). One pair was found to inhabit one burrow.
|
||
</paragraph>
|
||
<caption id="DF46698DFFB4FFC224E9F8E3B0928897" httpUri="https://zenodo.org/record/241133/files/figure.png" pageId="4" pageNumber="105" targetBox="[163,1421,212,1779]" targetPageId="4">
|
||
<paragraph id="8B863905FFB4FFC224E9F8E3B0928897" blockId="4.[151,1436,1818,1993]" pageId="4" pageNumber="105">
|
||
<emphasis id="B94DE517FFB4FFC224E9F8E3B41A8871" bold="true" box="[151,270,1818,1840]" pageId="4" pageNumber="105">FIGURE 2</emphasis>
|
||
.
|
||
<taxonomicName id="4C394286FFB4FFC22562F8E3B4F6886E" box="[284,482,1818,1840]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="ikedosoma" status="sp. nov.">
|
||
<emphasis id="B94DE517FFB4FFC22562F8E3B4F6886E" box="[284,482,1818,1840]" italics="true" pageId="4" pageNumber="105">Alpheus ikedosoma</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27E586CFFB4FFC22595F8E3B731886E" box="[491,549,1818,1840]" pageId="4" pageNumber="105" rank="species">
|
||
<emphasis id="B94DE517FFB4FFC22595F8E3B70B8871" bold="true" box="[491,543,1818,1839]" pageId="4" pageNumber="105">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, holotype, male (cl 8.8 mm), CBM-ZC 13128. A, carapace and cephalic appendages, lateral view; B, anterior part of carapace and left antennular and antennal peduncle, lateral view; C, close up of rostrum and left eye, lateral view; D, anterior part of carapace, and antennular and antennal peduncles, dorsal view (marginal setae on left antennal scale omitted); E, posterior margin of carapace, lateral view; F, telson, dorsal view (setae omitted); G, left antennal scale, dorsal view (setae omitted); H, appendices internal and masculina on left second pleopod, dorsal view; I, right uropod, dorsal view (perpendicular) (marginal setae on rami omitted). Scale bars: 2 mm for A; 1 mm for B, D–G, I; 0.5 mm for C, H.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF46698DFFB5FFC324E9F8D4B4578882" httpUri="https://zenodo.org/record/241134/files/figure.png" pageId="5" pageNumber="106" targetBox="[241,1337,208,1802]" targetPageId="5">
|
||
<paragraph id="8B863905FFB5FFC324E9F8D4B4578882" blockId="5.[151,1436,1836,2012]" pageId="5" pageNumber="106">
|
||
<emphasis id="B94DE517FFB5FFC324E9F8D4B405881C" bold="true" box="[151,273,1837,1859]" pageId="5" pageNumber="106">FIGURE 3</emphasis>
|
||
.
|
||
<taxonomicName id="4C394286FFB5FFC3255CF8D5B4FF881C" box="[290,491,1836,1858]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="ikedosoma" status="sp. nov.">
|
||
<emphasis id="B94DE517FFB5FFC3255CF8D5B4FF881C" box="[290,491,1836,1858]" italics="true" pageId="5" pageNumber="106">Alpheus ikedosoma</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27E586CFFB5FFC32588F8D4B727881C" box="[502,563,1837,1858]" pageId="5" pageNumber="106" rank="species">
|
||
<emphasis id="B94DE517FFB5FFC32588F8D4B739881C" bold="true" box="[502,557,1837,1858]" pageId="5" pageNumber="106">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, holotype, male (cl 8.8 mm), CBM-ZC 13128, left thoracic appendages. A, third maxilliped, lateral view; B, same, coxa, exopod and antepenultimate segment of endopod, ventral view (setae omitted); C, same, penultimate segment (= carpus), ventral view; D, second pereopod, lateral view; E, third pereopod, lateral view; F, same, ischium, ventral view; G, same, propodus and dactylus, lateral view; H, same, distal part of propodus and dactylus, flexor view; I, fourth pereopod, lateral view; J, same, ischium, ventral view; K, fifth pereopod, lateral view. Scale bars: 1 mm for A–F, I–K; 0.5 mm for G, H.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF46698DFFB6FFC024E9F88AB74988BB" httpUri="https://zenodo.org/record/241135/files/figure.png" pageId="6" pageNumber="107" targetBox="[200,1354,198,1853]" targetPageId="6">
|
||
<paragraph id="8B863905FFB6FFC024E9F88AB74988BB" blockId="6.[151,1436,1907,2021]" pageId="6" pageNumber="107">
|
||
<emphasis id="B94DE517FFB6FFC024E9F88AB41A88D6" bold="true" box="[151,270,1907,1929]" pageId="6" pageNumber="107">FIGURE 4</emphasis>
|
||
.
|
||
<taxonomicName id="4C394286FFB6FFC02565F88AB4F488D7" box="[283,480,1907,1929]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="107" phylum="Arthropoda" rank="species" species="ikedosoma" status="sp. nov.">
|
||
<emphasis id="B94DE517FFB6FFC02565F88AB4F488D7" box="[283,480,1907,1929]" italics="true" pageId="6" pageNumber="107">Alpheus ikedosoma</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27E586CFFB6FFC02596F88AB73688D7" box="[488,546,1907,1929]" pageId="6" pageNumber="107" rank="species">
|
||
<emphasis id="B94DE517FFB6FFC02596F88AB70888D6" bold="true" box="[488,540,1907,1928]" pageId="6" pageNumber="107">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, holotype, male (cl 8.8 mm), CBM-ZC 13128. A, left major cheliped (first pereopod), ischium to carpus, lateral view; B, same, mesial view; C, same pereopod, chela, dorsal (outer) view; D, same, mesial (upper) view; E, same, lateral (lower) view; F, right minor cheliped, lateral view; G, same, mesial view; H, same pereopod, chela, upper view; I, same, lower view. Scale bars: 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF46698DFFB7FFC124E9FC97B76A8C9C" httpUri="https://zenodo.org/record/241136/files/figure.png" pageId="7" pageNumber="108" targetBox="[154,1431,201,847]" targetPageId="7">
|
||
<paragraph id="8B863905FFB7FFC124E9FC97B76A8C9C" blockId="7.[151,1436,878,962]" pageId="7" pageNumber="108">
|
||
<emphasis id="B94DE517FFB7FFC124E9FC97B4048CDA" bold="true" box="[151,272,878,900]" pageId="7" pageNumber="108">FIGURE 5</emphasis>
|
||
.
|
||
<taxonomicName id="4C394286FFB7FFC12561FC97B4F28CDA" box="[287,486,878,900]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="108" phylum="Arthropoda" rank="species" species="ikedosoma" status="sp. nov.">
|
||
<emphasis id="B94DE517FFB7FFC12561FC97B4F28CDA" box="[287,486,878,900]" italics="true" pageId="7" pageNumber="108">Alpheus ikedosoma</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27E586CFFB7FFC12591FC96B73E8CDA" box="[495,554,879,900]" pageId="7" pageNumber="108" rank="species">
|
||
<emphasis id="B94DE517FFB7FFC12591FC96B7318CDA" bold="true" box="[495,549,879,900]" pageId="7" pageNumber="108">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, paratype, ovigerous female (cl 12.2 mm), CBM-ZC 13135. A, left major cheliped, ischium and merus, mesial view; B, same pereopod, chela, lateral view; C, right minor cheliped, mesial view; D, same pereopod, chela, mesial view. Scale bars: 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B863905FFB7FFC124B9FBACB4758B87" blockId="7.[151,1436,1001,2034]" pageId="7" pageNumber="108">
|
||
Specimens collected at Hondo flat, Amakusa, and Okoshiki, Uto, both in Kumamoto Prefecture, were not always paired, but this may be due to sampling condition. The host worms collected at these two localities in Kumamoto Prefecture were not preserved, but the specimen photographed at Hondo flat is very similar to
|
||
<emphasis id="B94DE517FFB7FFC121F2FB67B5F88B87" italics="true" pageId="7" pageNumber="108">I. elegans</emphasis>
|
||
(
|
||
<figureCitation id="13022580FFB7FFC12482FB38B42A8B84" box="[252,318,1217,1242]" captionStart="FIGURE 6" captionStartId="8.[151,250,1642,1664]" captionTargetBox="[366,1220,522,1620]" captionTargetId="figure@8.[365,1222,499,1621]" captionTargetPageId="8" captionText="FIGURE 6. Host innkeeper worm of Alpheus ikedosoma n. sp. A, Ikedosoma elegans (Ikeda, 1904), CBM-ZW, Hojo Beach, Tateyama, Chiba Prefecture, Japan, 30 July 2015, scale circle: 33 mm; B, unidentified specimen, not preserved, Hondo flat, Amakusa, Kumamto Prefecture, 11 May 2013." httpUri="https://zenodo.org/record/241137/files/figure.png" pageId="7" pageNumber="108">Fig. 6</figureCitation>
|
||
B).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3236A8EFFB7FFC124B9FB1CB02E8AD3" pageId="7" pageNumber="108" type="distribution">
|
||
<paragraph id="8B863905FFB7FFC124B9FB1CB02E8AD3" blockId="7.[151,1436,1001,2034]" pageId="7" pageNumber="108">
|
||
<emphasis id="B94DE517FFB7FFC124B9FB1CB44D8BA0" bold="true" box="[199,345,1253,1278]" pageId="7" pageNumber="108">Distribution</emphasis>
|
||
. Presently known only from Japanese main islands: Boso Peninsula (Honshu, Chiba Prefecture), and Amakusa and Uto, Ariake Sea (Kyushu, Kumamoto Prefecture); intertidal. The host innkeeper worm
|
||
<emphasis id="B94DE517FFB7FFC124E9FAD4B47B8A1B" box="[151,367,1325,1349]" italics="true" pageId="7" pageNumber="108">
|
||
<taxonomicName id="4C394286FFB7FFC124E9FAD4B4048A1B" box="[151,272,1325,1349]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="109" phylum="Arthropoda" rank="species" species="ikedosoma" status="sp. nov.">Ikedosoma</taxonomicName>
|
||
elegans
|
||
</emphasis>
|
||
has been recorded only in
|
||
<collectingCountry id="F32E7995FFB7FFC126D2FAD7B7FB8A1B" box="[684,751,1326,1349]" name="Japan" pageId="7" pageNumber="108">Japan</collectingCountry>
|
||
, viz., Misaki, Sagami Bay (Kanagawa Prefecture), Hamana Lake (Shizuoka Prefecture), and Takasu, Seto Inland Sea (Okayama Prefecture) (
|
||
<bibRefCitation id="EFA844F4FFB7FFC12055FAA8B0168A34" author="Tanaka" box="[1067,1282,1361,1386]" pageId="7" pageNumber="108" refString="Tanaka, M., Kon, T. & Nishikawa, T. (2014) Unraveling a 70 - year-old taxonomic puzzle: redefining the genus Ikedosoma (Annelida: Echiura) on the basis of morphological and molecular analyses. Zoological Science, 31, 849 - 861. http: // dx. doi. org / 10.2108 / zs 140155" type="journal article" year="2014">
|
||
Tanaka
|
||
<emphasis id="B94DE517FFB7FFC120FBFAABB1A28A37" box="[1157,1206,1361,1385]" italics="true" pageId="7" pageNumber="108">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
), and it could be expected that the present new species possibly occurs in those localities with records of the host worm.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3236A8EFFB7FFCE24B9FA60B4628EF3" lastPageId="8" lastPageNumber="109" pageId="7" pageNumber="108" type="discussion">
|
||
<paragraph id="8B863905FFB7FFC124B9FA60B4C98844" blockId="7.[151,1436,1001,2034]" pageId="7" pageNumber="108">
|
||
<emphasis id="B94DE517FFB7FFC124B9FA60B4208AEC" bold="true" box="[199,308,1433,1458]" pageId="7" pageNumber="108">Remarks</emphasis>
|
||
. The present new species is tentatively placed in the
|
||
<taxonomicName id="4C394286FFB7FFC127EBFA60B16C8AEF" box="[917,1144,1433,1457]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="108" phylum="Arthropoda" rank="species" species="brevirostris">
|
||
<emphasis id="B94DE517FFB7FFC127EBFA60B16C8AEF" box="[917,1144,1433,1457]" italics="true" pageId="7" pageNumber="108">Alpheus brevirostris</emphasis>
|
||
</taxonomicName>
|
||
species group because of the compressed major chela without conspicuous sculpture or armature and the subspatulate dactyli of the third and fourth pereopods (
|
||
<bibRefCitation id="EFA844F4FFB7FFC1251DFA1BB7788AA4" author="Banner" box="[355,620,1506,1530]" pageId="7" pageNumber="108" refString="Banner, D. M. & Banner, A. H. (1982) The alpheid shrimp of Australia. Part III: The remaining alpheids, principally the genus Alpheus and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner & Banner 1982</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA844F4FFB7FFC12608FA18B7EA8AA4" author="Chace" box="[630,766,1505,1530]" pageId="7" pageNumber="108" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, Part 5: Family Alpheidae. Smithsonian Contributions to Zoology, 466, 1 - 99." type="journal article" year="1988">Chace 1988</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA844F4FFB7FFC12777FA1BB69B8AA4" author="Bruce" box="[777,911,1506,1530]" pageId="7" pageNumber="108" refString="Bruce, A. J. (1994) Alpheus fenneri sp. nov. and A. williamsi sp. nov., two new Indo-West Pacific alpheid shrimps of the brevirostris species group. The Beagle, Records of the Museums and Art Galleries of the Northern Territory, 11, 15 - 28." type="journal article" year="1994">Bruce 1994</bibRefCitation>
|
||
). The species group is currently represented by 44 species (
|
||
<bibRefCitation id="EFA844F4FFB7FFC12565F9FFB4B78940" author="Bruce" box="[283,419,1542,1566]" pageId="7" pageNumber="108" refString="Bruce, A. J. (1994) Alpheus fenneri sp. nov. and A. williamsi sp. nov., two new Indo-West Pacific alpheid shrimps of the brevirostris species group. The Beagle, Records of the Museums and Art Galleries of the Northern Territory, 11, 15 - 28." type="journal article" year="1994">Bruce 1994</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA844F4FFB7FFC125D1F9FCB7D68940" author="Hayashi" box="[431,706,1541,1566]" pageId="7" pageNumber="108" refString="Hayashi, K. & Nagata, M. (2000) A new species of Alpheus (Decapoda, Caridea, Alpheidae) from Kagoshima Bay, Japan. Crustaceana, 73, 1109 - 1120. http: // dx. doi. org / 10.1163 / 156854000505119" type="journal article" year="2000">Hayashi & Nagata 2000</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFA844F4FFB7FFC126B0F9FFB61E8940" author="Hayashi" box="[718,778,1542,1566]" pageId="7" pageNumber="108" refString="Hayashi, K. & Nagata, M. (2002) Identity of Alpheus digitalis De Haan, 1844 and description of a new closely related species from the northwest Pacific (Decapoda: Caridea: Alpheidae). Crustacean Research, 31, 73 - 90." type="journal article" year="2002">2002</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA844F4FFB7FFC1276BF9FCB1578940" author="Anker" box="[789,1091,1541,1566]" pageId="7" pageNumber="108" refString="Anker, A & Dworschak, P. C. (2004) A new species of Alpheus from the tropical eastern Atlantic (Crustacea: Decapoda: Alpheidae). Annalen des Naturhistorischen Museums in Wien, 105 B, 47 - 58." type="journal article" year="2004">Anker & Dworschak 2004</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA844F4FFB7FFC12031F9FCB00D8940" author="Anker" box="[1103,1305,1541,1566]" pageId="7" pageNumber="108" refString="Anker, A., Hurt, C. & Knowlton, N. (2007) Three transisthmian snapping shrimps (Crustacea: Decapoda: Alpheidae: Alpheus) associated with innkeeper worms (Echiura: Thalassematidae). Zootaxa, 1626, 1 - 23." type="journal article" year="2007">
|
||
Anker
|
||
<emphasis id="B94DE517FFB7FFC120E3F9FFB1DA8943" box="[1181,1230,1541,1565]" italics="true" pageId="7" pageNumber="108">et al</emphasis>
|
||
. 2007
|
||
</bibRefCitation>
|
||
), several of them burrowing soft sediments or associated with gobiid fishes (
|
||
<bibRefCitation id="EFA844F4FFB7FFC127F7F9D3B188891C" author="Banner" box="[905,1180,1578,1602]" pageId="7" pageNumber="108" refString="Banner, D. M. & Banner, A. H. (1982) The alpheid shrimp of Australia. Part III: The remaining alpheids, principally the genus Alpheus and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner & Banner 1982</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA844F4FFB7FFC120D2F9D0B05E891C" author="Karplus" box="[1196,1354,1577,1602]" pageId="7" pageNumber="108" refString="Karplus, I. (1987) The associations between gobiid fishes and burrowing alpheid shrimp. Ocenography and Marine Biology: Annual Review, 25, 507 - 562." type="journal article" year="1987">Karplus 1987</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA844F4FFB7FFC12127F9D3B5C78938" author="Bruce" pageId="7" pageNumber="108" refString="Bruce, A. J. (1994) Alpheus fenneri sp. nov. and A. williamsi sp. nov., two new Indo-West Pacific alpheid shrimps of the brevirostris species group. The Beagle, Records of the Museums and Art Galleries of the Northern Territory, 11, 15 - 28." type="journal article" year="1994">Bruce 1994</bibRefCitation>
|
||
). In spite of the group assignment,
|
||
<taxonomicName id="4C394286FFB7FFC12612F9B7B610893B" box="[620,772,1613,1637]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="108" phylum="Arthropoda" rank="species" species="ikedosoma" status="sp. nov.">
|
||
<emphasis id="B94DE517FFB7FFC12612F9B7B610893B" box="[620,772,1613,1637]" italics="true" pageId="7" pageNumber="108">A. ikedosoma</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27E586CFFB7FFC12771F9B4B6448938" box="[783,848,1613,1638]" pageId="7" pageNumber="108" rank="species">
|
||
<emphasis id="B94DE517FFB7FFC12771F9B4B6448938" bold="true" box="[783,848,1613,1638]" pageId="7" pageNumber="108">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
does not exhibit close similarities to any of the 44 species in this informal species group. It is characterized by a suit of characters, including the very short, often down-curved rostrum without a dorsal ridge, the absence of adrostral grooves on the frontal region of the carapace, the strongly reduced dorsolateral spines on the telson, the lack of a ventrolateral distal tooth on the antennal basicerite, and the almost glabrous major chela without setal rows on margins. The real relationship of the present new species remains unclear.
|
||
</paragraph>
|
||
<paragraph id="8B863905FFB7FFC124B9F8DFB4B888F4" blockId="7.[151,1436,1001,2034]" pageId="7" pageNumber="108">
|
||
In the general shape of the major cheliped,
|
||
<taxonomicName id="4C394286FFB7FFC126CCF8DFB65E8863" box="[690,842,1829,1853]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="108" phylum="Arthropoda" rank="species" species="ikedosoma" status="sp. nov.">
|
||
<emphasis id="B94DE517FFB7FFC126CCF8DFB65E8863" box="[690,842,1829,1853]" italics="true" pageId="7" pageNumber="108">A. ikedosoma</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27E586CFFB7FFC1272CF8DCB6868860" box="[850,914,1829,1854]" pageId="7" pageNumber="108" rank="species">
|
||
<emphasis id="B94DE517FFB7FFC1272CF8DCB6868860" bold="true" box="[850,914,1829,1854]" pageId="7" pageNumber="108">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is superficially similar to
|
||
<taxonomicName id="4C394286FFB7FFC120C2F8DFB0888860" authority="Dana, 1852" authorityName="Dana" authorityYear="1852" box="[1212,1436,1829,1854]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="108" phylum="Arthropoda" rank="species" species="mitis">
|
||
<emphasis id="B94DE517FFB7FFC120C2F8DFB0068863" box="[1212,1298,1830,1853]" italics="true" pageId="7" pageNumber="108">A. mitis</emphasis>
|
||
<bibRefCitation id="EFA844F4FFB7FFC12164F8DFB0888860" author="Dana" box="[1306,1436,1829,1854]" pageId="7" pageNumber="108" refString="Dana, J. D. (1852) Crustacea, Part 1. United States Exploring Expedition during the years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U. S. N., 13, 1 - 685. Atlas 1 - 27, pls. 1 - 96 [1855]. C. Sherman, Philadelphia." type="journal article" year="1852">Dana, 1852</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C394286FFB7FFC124B9F8B3B716883C" authority="Miers, 1881" authorityName="Miers" authorityYear="1881" box="[199,514,1865,1890]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="108" phylum="Arthropoda" rank="species" species="paracrinitus">
|
||
<emphasis id="B94DE517FFB7FFC124B9F8B3B464883F" box="[199,368,1866,1889]" italics="true" pageId="7" pageNumber="108">A. paracrinitus</emphasis>
|
||
<bibRefCitation id="EFA844F4FFB7FFC12506F8B0B716883C" author="Miers" box="[376,514,1865,1890]" pageId="7" pageNumber="108" refString="Miers, E. J. (1881) On a collection of Crustacea made by Baron Hermann-Maltzan at Goree island, Senegambia. Annals and Magazine of Natural History, (5) 9, 192 - 195." type="journal article" year="1881">Miers, 1881</bibRefCitation>
|
||
</taxonomicName>
|
||
, currently referred to the
|
||
<taxonomicName id="4C394286FFB7FFC1275DF8B3B138883C" authority="Dana, 1852" authorityName="Dana" authorityYear="1852" box="[803,1068,1865,1890]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="108" phylum="Arthropoda" rank="species" species="diadema">
|
||
<emphasis id="B94DE517FFB7FFC1275DF8B3B6B7883F" box="[803,931,1865,1889]" italics="true" pageId="7" pageNumber="108">A. diadema</emphasis>
|
||
<bibRefCitation id="EFA844F4FFB7FFC127D4F8B3B138883C" author="Dana" box="[938,1068,1865,1890]" pageId="7" pageNumber="108" refString="Dana, J. D. (1852) Crustacea, Part 1. United States Exploring Expedition during the years 1838, 1839, 1840, 1841, 1842, under the Command of Charles Wilkes, U. S. N., 13, 1 - 685. Atlas 1 - 27, pls. 1 - 96 [1855]. C. Sherman, Philadelphia." type="journal article" year="1852">Dana, 1852</bibRefCitation>
|
||
</taxonomicName>
|
||
species group, but the latter two species have a less compressed major chela and subconical, simple dactyli of the third and fourth pereopods (cf.
|
||
<bibRefCitation id="EFA844F4FFB7FFC124E9F86BB48B88F4" author="Banner" box="[151,415,1938,1962]" pageId="7" pageNumber="108" refString="Banner, D. M. & Banner, A. H. (1982) The alpheid shrimp of Australia. Part III: The remaining alpheids, principally the genus Alpheus and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner & Banner 1982</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B863905FFB7FFCE24B9F84FB4628EF3" blockId="7.[151,1436,1001,2034]" lastBlockId="8.[151,1437,151,464]" lastPageId="8" lastPageNumber="109" pageId="7" pageNumber="108">
|
||
As mentioned above, association with echiuran burrows has been reported for the five species of
|
||
<taxonomicName id="4C394286FFB7FFC12175F84CB0718893" box="[1291,1381,1973,1997]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="108" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94DE517FFB7FFC12175F84CB0718893" box="[1291,1381,1973,1997]" italics="true" pageId="7" pageNumber="108">Alpheus</emphasis>
|
||
</taxonomicName>
|
||
, and the present new species represents the sixth example of the association. Of the five species, four are morphologically similar for one another within the
|
||
<taxonomicName id="4C394286FFB8FFCE268BFF61B68E8FEE" box="[757,922,152,176]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="109" phylum="Arthropoda" rank="species" species="brevirostris">
|
||
<emphasis id="B94DE517FFB8FFCE268BFF61B68E8FEE" box="[757,922,152,176]" italics="true" pageId="8" pageNumber="109">A. brevirostris</emphasis>
|
||
</taxonomicName>
|
||
group (constituting the
|
||
<taxonomicName id="4C394286FFB8FFCE20C6FF61B0548FEE" box="[1208,1344,152,176]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="109" phylum="Arthropoda" rank="species" species="barbatus">
|
||
<emphasis id="B94DE517FFB8FFCE20C6FF61B0548FEE" box="[1208,1344,152,176]" italics="true" pageId="8" pageNumber="109">A. barbatus</emphasis>
|
||
</taxonomicName>
|
||
species complex; see
|
||
<bibRefCitation id="EFA844F4FFB8FFCE254FFF45B4ED8F8A" author="Anker" box="[305,505,188,213]" pageId="8" pageNumber="109" refString="Anker, A., Hurt, C. & Knowlton, N. (2007) Three transisthmian snapping shrimps (Crustacea: Decapoda: Alpheidae: Alpheus) associated with innkeeper worms (Echiura: Thalassematidae). Zootaxa, 1626, 1 - 23." type="journal article" year="2007">
|
||
Anker
|
||
<emphasis id="B94DE517FFB8FFCE2503FF44B4BB8F8A" box="[381,431,188,212]" italics="true" pageId="8" pageNumber="109">et al</emphasis>
|
||
. 2007
|
||
</bibRefCitation>
|
||
). Nevertheless,
|
||
<taxonomicName id="4C394286FFB8FFCE26D4FF45B6548F8A" box="[682,832,188,212]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="109" phylum="Arthropoda" rank="species" species="ikedosoma" status="sp. nov.">
|
||
<emphasis id="B94DE517FFB8FFCE26D4FF45B6548F8A" box="[682,832,188,212]" italics="true" pageId="8" pageNumber="109">A. ikedosoma</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27E586CFFB8FFCE2736FF45B6938F8B" box="[840,903,188,213]" pageId="8" pageNumber="109" rank="species">
|
||
<emphasis id="B94DE517FFB8FFCE2736FF45B6938F8B" bold="true" box="[840,903,188,213]" pageId="8" pageNumber="109">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is not closely related to the
|
||
<taxonomicName id="4C394286FFB8FFCE20BFFF45B0508F8A" box="[1217,1348,188,212]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="109" phylum="Arthropoda" rank="species" species="barbatus">
|
||
<emphasis id="B94DE517FFB8FFCE20BFFF45B0508F8A" box="[1217,1348,188,212]" italics="true" pageId="8" pageNumber="109">A. barbatus</emphasis>
|
||
</taxonomicName>
|
||
species complex, because it is primarily distinguished from the latter species complex by the lack of a transverse groove or cleft on the palm of the major cheliped proximal to the dactylar articulation (versus a distinct groove or cleft is present) and the subspatulate dactyli of the third and fourth pereopods (versus the dactyli are subconical in the shape) (
|
||
<bibRefCitation id="EFA844F4FFB8FFCE2491FEB5B4AF8E3A" author="Anker" box="[239,443,332,357]" pageId="8" pageNumber="109" refString="Anker, A., Hurt, C. & Knowlton, N. (2007) Three transisthmian snapping shrimps (Crustacea: Decapoda: Alpheidae: Alpheus) associated with innkeeper worms (Echiura: Thalassematidae). Zootaxa, 1626, 1 - 23." type="journal article" year="2007">
|
||
Anker
|
||
<emphasis id="B94DE517FFB8FFCE2540FEB4B4648E3A" box="[318,368,332,356]" italics="true" pageId="8" pageNumber="109">et al</emphasis>
|
||
. 2007
|
||
</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C394286FFB8FFCE25B1FEB5B7DE8E3A" box="[463,714,332,356]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="109" phylum="Arthropoda" rank="species" species="echiurophilus">
|
||
<emphasis id="B94DE517FFB8FFCE25B1FEB5B7DE8E3A" box="[463,714,332,356]" italics="true" pageId="8" pageNumber="109">Alpheus echiurophilus</emphasis>
|
||
</taxonomicName>
|
||
, another representative of the echiuran associate, belongs to the separate species group,
|
||
<taxonomicName id="4C394286FFB8FFCE25D7FE89B75C8ED6" box="[425,584,368,392]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="109" phylum="Arthropoda" rank="species" species="leviusculus">
|
||
<emphasis id="B94DE517FFB8FFCE25D7FE89B75C8ED6" box="[425,584,368,392]" italics="true" pageId="8" pageNumber="109">A. leviusculus</emphasis>
|
||
</taxonomicName>
|
||
species group, suggesting a remote phylogenetic relationship to the other echiuran associates.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3236A8EFFB8FFCE24B9FE4EB0898E8E" box="[199,1437,439,464]" pageId="8" pageNumber="109" type="etymology">
|
||
<paragraph id="8B863905FFB8FFCE24B9FE4EB0898E8E" blockId="8.[151,1437,151,464]" box="[199,1437,439,464]" pageId="8" pageNumber="109">
|
||
<emphasis id="B94DE517FFB8FFCE24B9FE4EB4528E8E" bold="true" box="[199,326,439,464]" pageId="8" pageNumber="109">Etymology</emphasis>
|
||
. The new species is named after the host animal,
|
||
<emphasis id="B94DE517FFB8FFCE270EFE41B1518E8E" box="[880,1093,440,464]" italics="true" pageId="8" pageNumber="109">
|
||
<taxonomicName id="4C394286FFB8FFCE270EFE41B6FD8E8E" box="[880,1001,440,464]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="109" phylum="Arthropoda" rank="species" species="ikedosoma" status="sp. nov.">Ikedosoma</taxonomicName>
|
||
elegans
|
||
</emphasis>
|
||
. Used as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |