311 lines
34 KiB
XML
311 lines
34 KiB
XML
<document id="ED4B451C33D8355D24BD178981AC0AEC" ID-DOI="10.1186/s13358-022-00259-x" ID-ISSN="1664-2384" ID-PMC="PMC9681704" ID-PubMed="36439694" ID-Zenodo-Dep="12000082" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1718653166208" checkinUser="felipe" docAuthor="Leu, Marc, Bucher, Hugo, Vennemann, Torsten, Bagherpour, Borhan, Ji, Cheng, Brosse, Morgane & Goudemand, Nicolas" docDate="2022" docId="038B1D1D644AFF95FCA1ECDBD165FC07" docLanguage="en" docName="SwissJPalaeontol.141.19.pdf" docOrigin="Swiss Journal of Palaeontology (19) 141 (1)" docSource="http://dx.doi.org/10.1186/s13358-022-00259-x" docStyle="DocumentStyle:DBDC680DC95E08776BDF6354C30A315B.1:SwissJPalaeontol.2022-.journal_article" docStyleId="DBDC680DC95E08776BDF6354C30A315B" docStyleName="SwissJPalaeontol.2022-.journal_article" docStyleVersion="1" docTitle="Novispathodus praebrevissimus Leu & Bucher & Vennemann & Bagherpour & Ji & Brosse & Goudemand 2022, n. sp." docType="treatment" docVersion="11" lastPageNumber="37" masterDocId="FFB265656469FFB1FF8CED10D031FFE4" masterDocTitle="A Unitary Association-based conodont biozonation of the Smithian-Spathian boundary (Early Triassic) and associated biotic crisis from South China" masterLastPageNumber="61" masterPageNumber="1" pageNumber="36" updateTime="1722285166916" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="6186806A49F444C2918F99E5EDE9DE05">A Unitary Association-based conodont biozonation of the Smithian-Spathian boundary (Early Triassic) and associated biotic crisis from South China</mods:title>
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<mods:namePart id="B31C4223399A4013DD2766FD9D8ACC8F">Leu, Marc</mods:namePart>
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<mods:affiliation id="3EEE85AD35D73786FB06A3CD690C76EA">Paleontological Institute and Museum, University of Zurich, Karl-Schmid-Strasse 4, 8006 Zurich, Switzerland</mods:affiliation>
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<mods:namePart id="FE0649242758A2241770855C393D4519">Bucher, Hugo</mods:namePart>
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<mods:affiliation id="B025632041CF1637A26BF982912D09D7">Paleontological Institute and Museum, University of Zurich, Karl-Schmid-Strasse 4, 8006 Zurich, Switzerland</mods:affiliation>
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<mods:namePart id="9E7DA4F1FDC52EE325BA2E5D9D7A648D">Vennemann, Torsten</mods:namePart>
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<mods:namePart id="1B804073D34EAC94368E28C1D94A92D5">Bagherpour, Borhan</mods:namePart>
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<mods:namePart id="64FEBB1DAEC06FEC0A411384C2530DA4">Ji, Cheng</mods:namePart>
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<mods:namePart id="B3C66023DACA723874864F05CFFB07D2">Brosse, Morgane</mods:namePart>
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<mods:namePart id="46F7BE64D9148E6615DEA3B4F4168CB3">Goudemand, Nicolas</mods:namePart>
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<mods:title id="8051635133E14F03AB078199A4A9F3B0">Swiss Journal of Palaeontology</mods:title>
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<mods:date id="A0700DAC7A1BDCAC5C0D9F7931063188">2022</mods:date>
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<treatment id="038B1D1D644AFF95FCA1ECDBD165FC07" ID-DOI="http://doi.org/10.5281/zenodo.13128029" ID-Zenodo-Dep="13128029" LSID="urn:lsid:zoobank.org:act:244F954B-2E0B-49DA-A316-F854DA0DE2F4" httpUri="http://treatment.plazi.org/id/038B1D1D644AFF95FCA1ECDBD165FC07" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
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<subSubSection id="C338FF80644AFF92FCA1ECDBD4F3FE07" box="[813,1218,459,483]" pageId="35" pageNumber="36" type="nomenclature">
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<paragraph id="8B9DAC0B644AFF92FCA1ECDBD4F3FE07" blockId="35.[813,1303,459,515]" box="[813,1218,459,483]" pageId="35" pageNumber="36">
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<heading id="D0D51B67644AFF92FCA1ECDBD4F3FE07" box="[813,1218,459,483]" fontSize="9" level="2" pageId="35" pageNumber="36" reason="8">
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<taxonomicName id="4C22D788644AFF92FCA1ECDBD4B3FE07" authority="Leu & Bucher & Vennemann & Bagherpour & Ji & Brosse & Goudemand, 2022" authorityName="Leu & Bucher & Vennemann & Bagherpour & Ji & Brosse & Goudemand" authorityYear="2022" box="[813,1154,459,483]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ozarkodinida" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="praebrevissimus" status="sp. nov.">
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<emphasis id="B9567019644AFF92FCA1ECDBD4B3FE07" box="[813,1154,459,483]" italics="true" pageId="35" pageNumber="36">Novispathodus praebrevissimus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A265CD62644AFF92FB05ECDCD4F3FE07" box="[1161,1218,460,483]" pageId="35" pageNumber="36" rank="species">n. sp.</taxonomicNameLabel>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C338FF80644AFF95FCA1ECFBD165FC07" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="description">
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<paragraph id="8B9DAC0B644AFF92FCA1ECFBD526FDE7" blockId="35.[813,1303,459,515]" box="[813,1303,491,515]" pageId="35" pageNumber="36">
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<figureCitation id="1319B08E644AFF92FCA1ECFBD3A2FDE7" box="[813,915,491,515]" captionStart="Fig" captionStartId="26.[169,200,1663,1683]" captionTargetBox="[152,1438,238,1791]" captionTargetId="figure-11@27.[232,1358,254,1754]" captionTargetPageId="27" captionText="Fig. 17 Novispathodinae from Qiakong,Laren, Shanggang,and Lilong. Magnification is × 80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. A–C, J, T, AA Novispathodus ex gr. pingdingshanensis (Zhao & Orchard);A, QIA138,PIMUZ 39226; B QIA138, PIMUZ 39227; C LAR202, PIMUZ 39228; J QIA136, PIMUZ 39229;T LIL504, PIMUZ 39230; AA LIL508,PIMUZ 39231. D–F, M–P, R, S, U, X, AB, AD Novispathodus pingdingshanensis (Zhao & Orchard); D QIA135, PIMUZ 39259; E LIL508,PIMUZ 39260; F LIL506, PIMUZ 39261; M LIL506, PIMUZ 39262; N LIL506,PIMUZ 39263; O LIL507, PIMUZ 39264; P LIL506, PIMUZ 39265; R LIL508,PIMUZ 39266; S LAR204,PIMUZ 39267; U LIL507, PIMUZ 39268; X LIL508, PIMUZ 39269; AB LIL508, PIMUZ 39270; AD LIL508, PIMUZ 39271. G, I, Q Novispathodus cf.?gryphus n. sp.; G QIA136, PIMUZ 39204; I QIA136, PIMUZ 39205; Q QIA135, PIMUZ 39206. H, K, L Novispathodus gryphus n. sp.; H QIA135, PIMUZ 39243; K LIL506, PIMUZ 39244; L LIL506,PIMUZ 39245. V Novispathodus praebrevissimus n. sp.; LIL507, PIMUZ 39273. W, Y, Z, AE Novispathodus ex gr. abruptus (Orchard); W LIL505, PIMUZ 39208; Y SHA346,PIMUZ 39209; Z LIL506, PIMUZ 39210; AE SHA346, PIMUZ 39211. AC sp. indet.; LIL504, PIMUZ 39295. AF Novispathodus pingdingshanensis (P1 cluster) (Zhao & Orchard); QIA136, PIMUZ 39272" figureDoi="http://doi.org/10.5281/zenodo.13127510" httpUri="https://zenodo.org/record/13127510/files/figure.png" pageId="35" pageNumber="36">Figs. 17V</figureCitation>
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;
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<figureCitation id="1319B08E644AFF92FC10ECFBD46DFDE7" box="[924,1116,491,515]" captionStart="Fig" captionStartId="28.[169,200,1717,1737]" captionTargetBox="[152,1438,238,1801]" captionTargetId="figure-11@29.[228,1362,254,1763]" captionTargetPageId="29" captionText="Fig. 18 Novispathodinae from Laren and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise.A, B, D, F, L–N, Q, S, T Novispathodus praebrevissimus n. sp.; A LIL506,PIMUZ 39274; B LAR207, PIMUZ 39275; D LIL508, PIMUZ 39276; F LIL507, PIMUZ 39277;L LIL507, PIMUZ 39278; M LIL509, PIMUZ 39279; N LIL507,PIMUZ 39280; Q LIL507, PIMUZ 39281; S LIL507, PIMUZ 39282; T LIL507, PIMUZ 39283. C, O, R Novispathodus?praebrevissimus n. sp.; C LAR204,PIMUZ 39284; O LIL507, PIMUZ 39285; R LIL507, PIMUZ 39286. E, I Novispathodus ex gr. pingdingshanensis (Zhao & Orchard); E LIL507,PIMUZ 39232, I LIL507, PIMUZ 39233. G Triassospathodus aff. symmetricus (Orchard);LIL507, PIMUZ 39309. H Novispathodus ex gr. abruptus (Orchard); LIL507, PIMUZ 39212. J, K Novispathodus praebrevissimus (juvenile) n. sp.; J LIL507, PIMUZ 39287; K LIL507,PIMUZ 39288. P Novispathodus robustispinus (Zhao & Orchard); LIL507,PIMUZ 39290" figureDoi="http://doi.org/10.5281/zenodo.13127512" httpUri="https://zenodo.org/record/13127512/files/figure.png" pageId="35" pageNumber="36">18A–D, F, J–N, O</figureCitation>
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?, P?, Q, S, T; 19R
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||
</paragraph>
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<paragraph id="8B9DAC0B644AFF92FCA1EF3BD470FD87" blockId="35.[813,1437,555,899]" pageId="35" pageNumber="36">
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1984
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<taxonomicName id="4C22D788644AFF92FCE1EF3BD400FDA7" box="[877,1073,555,579]" family="Gondolellidae" genus="Neospathodus" kingdom="Animalia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="undetermined">
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<emphasis id="B9567019644AFF92FCE1EF3BD437FDA7" box="[877,1030,555,579]" italics="true" pageId="35" pageNumber="36">Neospathodus</emphasis>
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sp.
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</taxonomicName>
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<taxonomicName id="4C22D788644AFF92FBB0EF3BD579FDA7" authority="Bender" authorityName="Bender" box="[1084,1352,555,579]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="triangularis">
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aff.
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<emphasis id="B9567019644AFF92FBE5EF3BD4DAFDA7" box="[1129,1259,555,579]" italics="true" pageId="35" pageNumber="36">triangularis</emphasis>
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Bender
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</taxonomicName>
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; Hatleberg & Clark, pl. 3, fig. 16
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</paragraph>
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<paragraph id="8B9DAC0B644AFF92FCA1EF7BD4E0FD47" blockId="35.[813,1437,555,899]" pageId="35" pageNumber="36">
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2014
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<taxonomicName id="4C22D788644AFF92FCE1EF7BD484FD67" authority="(Buryi)" baseAuthorityName="Buryi" baseAuthorityYear="1979" box="[877,1205,619,643]" family="Gondolellidae" genus="Icriospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="zaksi">
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<emphasis id="B9567019644AFF92FCE1EF7BD43CFD67" box="[877,1037,619,643]" italics="true" pageId="35" pageNumber="36">Icriospathodus</emphasis>
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?
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<emphasis id="B9567019644AFF92FBAFEF7BD46BFD67" box="[1059,1114,619,643]" italics="true" pageId="35" pageNumber="36">zaksi</emphasis>
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(Buryi)
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</taxonomicName>
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; Maekawa & Igo in Shigeta et al., fig. 192, nr. 10–13 (only).
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</paragraph>
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<paragraph id="8B9DAC0B644AFF92FCA1EFBBD3F8FD07" blockId="35.[813,1437,555,899]" pageId="35" pageNumber="36">
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2015
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<taxonomicName id="4C22D788644AFF92FCE7EFBBD53CFD27" authority="(Orchard)" baseAuthorityName="Orchard" box="[875,1293,683,707]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="brevissimus">
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<emphasis id="B9567019644AFF92FCE7EFBBD4A5FD27" box="[875,1172,683,707]" italics="true" pageId="35" pageNumber="36">Novispathodus brevissimus</emphasis>
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(Orchard)
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</taxonomicName>
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; Chen et al., p. 111, fig. 7.8.
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</paragraph>
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<paragraph id="8B9DAC0B644AFF92FCA1EFFBD342FCA7" blockId="35.[813,1437,555,899]" pageId="35" pageNumber="36">
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2019
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<taxonomicName id="4C22D788644AFF92FC0DEFFBD3A5FCC7" authority="(Zhao & Orchard)" baseAuthorityName="Zhao & Orchard" baseAuthorityYear="2007" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="pingdingshanensis">
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<emphasis id="B9567019644AFF92FC0DEFFBD536FCE7" box="[897,1287,747,771]" italics="true" pageId="35" pageNumber="36">Novispathodus pingdingshanensis</emphasis>
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(Zhao & Orchard)
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</taxonomicName>
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; Chen et al., fig. 3 nr. 5, 9 (only) fig. 5, nr. 6 (only).
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</paragraph>
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<paragraph id="8B9DAC0B644AFF92FCA1EE5BD532FC67" blockId="35.[813,1437,555,899]" pageId="35" pageNumber="36">
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2019
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||
<taxonomicName id="4C22D788644AFF92FC0DEE5BD3A5FC67" authority="(Zhao & Orchard)" baseAuthorityName="Zhao & Orchard" baseAuthorityYear="2007" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="pingdingshanensis">
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<emphasis id="B9567019644AFF92FC0DEE5BD536FC87" box="[897,1287,843,867]" italics="true" pageId="35" pageNumber="36">Novispathodus pingdingshanensis</emphasis>
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(Zhao & Orchard)
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</taxonomicName>
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||
; Liu et al., p. 13, pl. 3, fig. 7 (only).
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</paragraph>
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<paragraph id="8B9DAC0B644AFF92FCA1EEBBD4B1FC07" blockId="35.[813,1437,939,1251]" pageId="35" pageNumber="36">
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<emphasis id="B9567019644AFF92FCA1EEBBD3AFFC27" box="[813,926,939,963]" italics="true" pageId="35" pageNumber="36">Etymology</emphasis>
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: According to its presumed relationship as a predecessor of
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<taxonomicName id="4C22D788644AFF92FC5DEEDCD44BFC07" box="[977,1146,971,995]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="brevissimus">
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<emphasis id="B9567019644AFF92FC5DEEDCD3C1FC07" box="[977,1008,972,995]" italics="true" pageId="35" pageNumber="36">Nv</emphasis>
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||
.
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<emphasis id="B9567019644AFF92FC70EEDBD44BFC07" box="[1020,1146,971,995]" italics="true" pageId="35" pageNumber="36">brevissimus</emphasis>
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</taxonomicName>
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.
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</paragraph>
|
||
<paragraph id="8B9DAC0B644AFF92FCA1EEFBD4DBFBE7" blockId="35.[813,1437,939,1251]" box="[813,1258,1003,1027]" pageId="35" pageNumber="36">
|
||
<emphasis id="B9567019644AFF92FCA1EEFBD3A5FBE7" box="[813,916,1003,1027]" italics="true" pageId="35" pageNumber="36">
|
||
<typeStatus id="549912A9644AFF92FCA1EEFBD3BFFBE7" box="[813,910,1003,1027]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
specimen illustrated in fig. 24R
|
||
</paragraph>
|
||
<paragraph id="8B9DAC0B644AFF92FCA1E91CD4DEFBC7" blockId="35.[813,1437,939,1251]" box="[813,1263,1035,1059]" pageId="35" pageNumber="36">
|
||
<emphasis id="B9567019644AFF92FCA1E91CD3A5FBC7" box="[813,916,1036,1059]" italics="true" pageId="35" pageNumber="36">
|
||
<typeStatus id="549912A9644AFF92FCA1E91CD3BEFBC7" box="[813,911,1036,1059]" pageId="35" pageNumber="36" type="paratype">Paratype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
specimen illustrated in fig. 24Q
|
||
</paragraph>
|
||
<paragraph id="8B9DAC0B644AFF92FCA1E93CD3D3FB87" blockId="35.[813,1437,939,1251]" pageId="35" pageNumber="36">
|
||
<emphasis id="B9567019644AFF92FCA1E93CD3F4FBA7" box="[813,965,1067,1091]" italics="true" pageId="35" pageNumber="36">Type locality:</emphasis>
|
||
Lilong cliff, Luolou formation,
|
||
<collectingRegion id="49E662E9644AFF92FAB3E93BD3BEFB87" country="China" name="Guangxi" pageId="35" pageNumber="36">Guangxi Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F335EC9B644AFF92FC15E95BD3EFFB87" box="[921,990,1099,1123]" name="China" pageId="35" pageNumber="36">China</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9DAC0B644AFF92FCA1E97CD4ABFB07" blockId="35.[813,1437,939,1251]" pageId="35" pageNumber="36">
|
||
<emphasis id="B9567019644AFF92FCA1E97CD394FB67" box="[813,933,1131,1155]" italics="true" pageId="35" pageNumber="36">Type level</emphasis>
|
||
: Luolou Formation, within latest Smithian black shales. Present in UAZ
|
||
<subScript id="17A6AE4E644AFF92FB3CE987D488FB4C" attach="left" box="[1200,1209,1175,1192]" fontSize="7" pageId="35" pageNumber="36">5</subScript>
|
||
which corresponds to the peak of the positive δ
|
||
<superScript id="7C570143644AFF92FBF1E9B6D4BEFB53" attach="both" box="[1149,1167,1190,1207]" fontSize="7" pageId="35" pageNumber="36">13</superScript>
|
||
<subScript id="17A6AE4E644AFF92FB03E9BBD4F3FB2C" attach="left" box="[1167,1218,1195,1224]" fontSize="7" pageId="35" pageNumber="36">Ccarb</subScript>
|
||
excursion and the
|
||
<taxonomicName id="4C22D788644AFF92FCA1E9DBD39FFB07" authorityName="Spath" authorityYear="1930" box="[813,942,1227,1251]" class="Cephalopoda" family="Xenoceltitidae" genus="Xenoceltites" kingdom="Animalia" pageId="35" pageNumber="36" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B9567019644AFF92FCA1E9DBD39FFB07" box="[813,942,1227,1251]" italics="true" pageId="35" pageNumber="36">Xenoceltites</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C22D788644AFF92FC3BE9DBD46BFB07" authorityName="Spath" authorityYear="1930" box="[951,1114,1227,1251]" class="Cephalopoda" family="Xenoceltitidae" genus="Glyptophiceras" kingdom="Animalia" pageId="35" pageNumber="36" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B9567019644AFF92FC3BE9DBD46BFB07" box="[951,1114,1227,1251]" italics="true" pageId="35" pageNumber="36">Glyptophiceras</emphasis>
|
||
</taxonomicName>
|
||
beds.
|
||
</paragraph>
|
||
<paragraph id="8B9DAC0B644AFF92FCA1E81BD460FAC7" blockId="35.[813,1437,1291,1923]" box="[813,1105,1291,1315]" pageId="35" pageNumber="36">
|
||
<emphasis id="B9567019644AFF92FCA1E81BD422FAC7" box="[813,1043,1291,1315]" italics="true" pageId="35" pageNumber="36">Number of specimens</emphasis>
|
||
.> 40.
|
||
</paragraph>
|
||
<paragraph id="8B9DAC0B644AFF92FCCEE83CD4FBFA47" blockId="35.[813,1437,1291,1923]" pageId="35" pageNumber="36">
|
||
<emphasis id="B9567019644AFF92FCCEE83CD39DFAA7" box="[834,940,1324,1347]" italics="true" pageId="35" pageNumber="36">Diagnosis</emphasis>
|
||
. Short robust segminate P
|
||
<subScript id="17A6AE4E644AFF92FB6DE827D4DBFAAC" attach="left" box="[1249,1258,1335,1352]" fontSize="7" pageId="35" pageNumber="36">1</subScript>
|
||
element. Large rounded-to-sub-rounded basal cavity. Posterior margin often slightly concave. Small, largely fused denticles. Groove from basal pit to anterior end.
|
||
</paragraph>
|
||
<paragraph id="8B9DAC0B644AFF92FCCEE8BCD52FF8E7" blockId="35.[813,1437,1291,1923]" pageId="35" pageNumber="36">
|
||
<emphasis id="B9567019644AFF92FCCEE8BCD38EFA27" box="[834,959,1452,1475]" italics="true" pageId="35" pageNumber="36">Description</emphasis>
|
||
. Te segminate P
|
||
<subScript id="17A6AE4E644AFF92FB03E8A7D4A9FA2C" attach="left" box="[1167,1176,1463,1480]" fontSize="7" pageId="35" pageNumber="36">1</subScript>
|
||
element shows a very large rounded to sub-rounded basal cavity in the posterior half of the element. A very deep basal groove is present from the basal pit to the anterior end of the lower margin. Te carina is relatively high with numerous small, fused denticles. Te cusp is undistinguishable. Te posterior lateral margin (posteriormost edge between basal cavity and last denticle) is slightly to strongly curved. Sometimes, the height of the posteriormost denticles is gradually but rapidly decreasing. In lateral view, the denticles tend to be radiating or recurved.
|
||
</paragraph>
|
||
<paragraph id="8B9DAC0B644AFF95FCCEEA1BD336FD67" blockId="35.[813,1437,1291,1923]" lastBlockId="36.[151,776,235,995]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
|
||
<emphasis id="B9567019644AFF92FCCEEA1BD391F8C7" box="[834,928,1803,1827]" italics="true" pageId="35" pageNumber="36">Remarks</emphasis>
|
||
. Tis species is thought to be intermediate between
|
||
<taxonomicName id="4C22D788644AFF92FC1DEA3CD4B0F8A7" baseAuthorityName="Zhao & Orchard" baseAuthorityYear="2007" box="[913,1153,1835,1859]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="pingdingshanensis">
|
||
<emphasis id="B9567019644AFF92FC1DEA3CD4B0F8A7" box="[913,1153,1835,1859]" italics="true" pageId="35" pageNumber="36">Nv. pingdingshanensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C22D788644AFF92FB37EA3CD554F8A7" box="[1211,1381,1835,1859]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="brevissimus">
|
||
<emphasis id="B9567019644AFF92FB37EA3CD554F8A7" box="[1211,1381,1835,1859]" italics="true" pageId="35" pageNumber="36">Nv. brevissimus</emphasis>
|
||
</taxonomicName>
|
||
. Te P
|
||
<subScript id="17A6AE4E644AFF92FCB7EA47D375F88C" attach="left" box="[827,836,1879,1896]" fontSize="7" pageId="35" pageNumber="36">1</subScript>
|
||
element of this species is very similar to that of
|
||
<taxonomicName id="4C22D788644AFF92FAF4EA5CD3C0F867" baseAuthorityName="Zhao & Orchard" baseAuthorityYear="2007" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="pingdingshanensis">
|
||
<emphasis id="B9567019644AFF92FAF4EA5CD5A6F887" box="[1400,1431,1868,1891]" italics="true" pageId="35" pageNumber="36">Nv</emphasis>
|
||
.
|
||
<emphasis id="B9567019644AFF92FCA1EA7BD3C0F867" box="[813,1009,1899,1923]" italics="true" pageId="35" pageNumber="36">pingdingshanensis</emphasis>
|
||
</taxonomicName>
|
||
, but the basal cavity is much larger and rounded. In comparison to
|
||
<taxonomicName id="4C22D788644DFF95FD95EDFCD2FDFEE7" box="[537,716,235,259]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="brevissimus">
|
||
<emphasis id="B9567019644DFF95FD95EDFCD209FEE7" box="[537,568,236,259]" italics="true" pageId="36" pageNumber="37">Nv</emphasis>
|
||
.
|
||
<emphasis id="B9567019644DFF95FDC0EDFBD2FDFEE7" box="[588,716,235,259]" italics="true" pageId="36" pageNumber="37">brevissimus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9567019644DFF95FD40EDFBD2E1FEE7" box="[716,720,235,259]" italics="true" pageId="36" pageNumber="37">,</emphasis>
|
||
this species is usually smaller and has a relatively bigger and more rounded basal cavity. Furthermore, the carina is lower, with less numerous denticles. Te new species is distinguished from the similar
|
||
<taxonomicName id="4C22D788644DFF95FE64EC7BD1BEFE47" authority="(Maekawa et al., 2018)" baseAuthorityName="Maekawa" baseAuthorityYear="2018" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="shirokawai">
|
||
<emphasis id="B9567019644DFF95FE64EC7BD336FE67" box="[488,775,363,387]" italics="true" pageId="36" pageNumber="37">Novispathodus shirokawai</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB3D1FA644DFF95FF12EC9BD1B7FE47" author="Maekawa" box="[158,390,395,419]" etAl="et al." firstAuthor="Maekawa" pageId="36" pageNumber="37" pagination="1 - 62" refId="ref44774" refString="Maekawa, T., Komatsu, T., & Koike, T. (2018). Early Triassic conodonts from the Tahogawa Member of the Taho Formation, Ehime Prefecture, Southwest Japan. Paleontological Research, 22 (s 1), 1 - 62. https: // doi. org / 10. 2517 / 2018 PR 001" type="journal article" year="2018">Maekawa et al., 2018</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
by reclined and less pointy denticles, a larger, rounded basal cavity and a less triangular shape in lateral view. Compared to
|
||
<taxonomicName id="4C22D788644DFF95FDA4ECDCD2B3FE07" baseAuthorityName="Buryi" baseAuthorityYear="1979" box="[552,642,459,483]" family="Gondolellidae" genus="Icriospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="zaksi">
|
||
<emphasis id="B9567019644DFF95FDA4ECDCD2B3FE07" box="[552,642,459,483]" italics="true" pageId="36" pageNumber="37">Ic. zaksi</emphasis>
|
||
</taxonomicName>
|
||
, the P
|
||
<subScript id="17A6AE4E644DFF95FD40ECC7D2E4FE0C" attach="left" box="[716,725,471,488]" fontSize="7" pageId="36" pageNumber="37">1</subScript>
|
||
element of this species is less robust and has a relatively higher carina.
|
||
<taxonomicName id="4C22D788644DFF95FEBAEF1CD2A3FDC7" authority="(Zhao & Orchard)" baseAuthorityName="Zhao & Orchard" box="[310,658,523,547]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="expansus">
|
||
<emphasis id="B9567019644DFF95FEBAEF1CD1F2FDC7" box="[310,451,524,547]" italics="true" pageId="36" pageNumber="37">Ns. expansus</emphasis>
|
||
(Zhao & Orchard)
|
||
</taxonomicName>
|
||
has a very similar morphology to
|
||
<taxonomicName id="4C22D788644DFF95FE16EF3CD244FDA7" authority="Leu & Bucher & Vennemann & Bagherpour & Ji & Brosse & Goudemand, 2022" authorityName="Leu & Bucher & Vennemann & Bagherpour & Ji & Brosse & Goudemand" authorityYear="2022" box="[410,629,555,579]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="praebrevissimus" status="n. sp.">
|
||
<emphasis id="B9567019644DFF95FE16EF3CD244FDA7" box="[410,629,555,579]" italics="true" pageId="36" pageNumber="37">Nv. praebrevissimus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A265CD62644DFF95FDF2EF3CD28DFDA7" box="[638,700,556,579]" pageId="36" pageNumber="37" rank="species">n. sp.</taxonomicNameLabel>
|
||
, but it has a distinct conspicuous thickening in the middle part of its unit. Some specimens of
|
||
<taxonomicName id="4C22D788644DFF95FE60EF7CD2F4FD67" authority="Leu & Bucher & Vennemann & Bagherpour & Ji & Brosse & Goudemand, 2022" authorityName="Leu & Bucher & Vennemann & Bagherpour & Ji & Brosse & Goudemand" authorityYear="2022" box="[492,709,619,643]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="praebrevissimus" status="n. sp.">
|
||
<emphasis id="B9567019644DFF95FE60EF7CD2F4FD67" box="[492,709,619,643]" italics="true" pageId="36" pageNumber="37">Nv. praebrevissimus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A265CD62644DFF95FD41EF7CD336FD67" box="[717,775,620,643]" pageId="36" pageNumber="37" rank="species">n. sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8B9DAC0B644DFF95FF1BEF9BD11BFD07" blockId="36.[151,776,235,995]" pageId="36" pageNumber="37">
|
||
(
|
||
<figureCitation id="1319B08E644DFF95FF13EF9BD16CFD47" box="[159,349,651,675]" captionStart="Fig" captionStartId="28.[169,200,1717,1737]" captionTargetBox="[152,1438,238,1801]" captionTargetId="figure-11@29.[228,1362,254,1763]" captionTargetPageId="29" captionText="Fig. 18 Novispathodinae from Laren and Lilong. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise.A, B, D, F, L–N, Q, S, T Novispathodus praebrevissimus n. sp.; A LIL506,PIMUZ 39274; B LAR207, PIMUZ 39275; D LIL508, PIMUZ 39276; F LIL507, PIMUZ 39277;L LIL507, PIMUZ 39278; M LIL509, PIMUZ 39279; N LIL507,PIMUZ 39280; Q LIL507, PIMUZ 39281; S LIL507, PIMUZ 39282; T LIL507, PIMUZ 39283. C, O, R Novispathodus?praebrevissimus n. sp.; C LAR204,PIMUZ 39284; O LIL507, PIMUZ 39285; R LIL507, PIMUZ 39286. E, I Novispathodus ex gr. pingdingshanensis (Zhao & Orchard); E LIL507,PIMUZ 39232, I LIL507, PIMUZ 39233. G Triassospathodus aff. symmetricus (Orchard);LIL507, PIMUZ 39309. H Novispathodus ex gr. abruptus (Orchard); LIL507, PIMUZ 39212. J, K Novispathodus praebrevissimus (juvenile) n. sp.; J LIL507, PIMUZ 39287; K LIL507,PIMUZ 39288. P Novispathodus robustispinus (Zhao & Orchard); LIL507,PIMUZ 39290" figureDoi="http://doi.org/10.5281/zenodo.13127512" httpUri="https://zenodo.org/record/13127512/files/figure.png" pageId="36" pageNumber="37">Fig. 18L, O and P</figureCitation>
|
||
) show a bulbous thickening in the posterior part of the carina but never as pronounced as in
|
||
<taxonomicName id="4C22D788644DFF95FF1BEFDCD115FD07" box="[151,292,716,739]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="expansus">
|
||
<emphasis id="B9567019644DFF95FF1BEFDCD115FD07" box="[151,292,716,739]" italics="true" pageId="36" pageNumber="37">Ns. expansus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9DAC0B644DFF95FF21EFFCD165FC07" blockId="36.[151,776,235,995]" pageId="36" pageNumber="37">
|
||
<emphasis id="B9567019644DFF95FF21EFFCD119FCE7" box="[173,296,748,771]" italics="true" pageId="36" pageNumber="37">Occurrence</emphasis>
|
||
. Spitsbergen, Botneheia Formation (
|
||
<bibRefCitation id="EFB3D1FA644DFF95FD4DEFFBD144FCC7" author="Hatleberg & Clark" firstAuthor="Hatleberg" pageId="36" pageNumber="37" pagination="101 - 125" refId="ref41527" refString="Hatleberg, E. W., & Clark, D. L. (1984). Lower Triassic conodonts and biofacies interpretations: Nepal and Svalbard. Geologica Et Palaeontologica, 18 (12), 101 - 125." type="journal article" year="1984">Hatleberg & Clark, 1984</bibRefCitation>
|
||
). South
|
||
<collectingCountry id="F335EC9B644DFF95FE50EE1BD211FCC7" box="[476,544,779,803]" name="China" pageId="36" pageNumber="37">China</collectingCountry>
|
||
, Nanpanjiang basin: Jiarong, southern
|
||
<collectingRegion id="49E662E9644DFF95FEE9EE3BD1F5FCA7" box="[357,452,811,835]" country="China" name="Guizhou" pageId="36" pageNumber="37">Guizhou</collectingRegion>
|
||
(
|
||
<bibRefCitation id="EFB3D1FA644DFF95FE54EE3BD2AEFCA7" author="Chen" box="[472,671,811,835]" etAl="et al." firstAuthor="Chen" pageId="36" pageNumber="37" pagination="104 - 121" refId="ref39362" refString="Chen, Y., Jiang, H., Lai, X., Yan, C., Richoz, S., Liu, X., & Wang, L. (2015). Early Triassic conodonts of Jiarong, Nanpanjiang Basin, southern Guizhou Province, South China. Journal of Asian Earth Sciences, 105, 104 - 121. https: // doi. org / 10.1016 / j. jseaes. 2015.03. 014" type="journal article" year="2015">Chen et al., 2015</bibRefCitation>
|
||
). Luolou Formation, within latest Smithian black shales. Present in UAZ
|
||
<subScript id="17A6AE4E644DFF95FF60EE67D0C4FC6C" attach="left" box="[236,245,887,904]" fontSize="7" pageId="36" pageNumber="37">5</subScript>
|
||
which corresponds to the peak of the positive δ
|
||
<superScript id="7C570143644DFF95FF29EE96D086FC73" attach="both" box="[165,183,902,919]" fontSize="7" pageId="36" pageNumber="37">13</superScript>
|
||
<subScript id="17A6AE4E644DFF95FF3BEE9BD0DBFC4C" attach="left" box="[183,234,907,936]" fontSize="7" pageId="36" pageNumber="37">Ccarb</subScript>
|
||
excursion and the
|
||
<taxonomicName id="4C22D788644DFF95FE55EE9BD26BFC47" authorityName="Spath" authorityYear="1930" box="[473,602,907,931]" class="Cephalopoda" family="Xenoceltitidae" genus="Xenoceltites" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B9567019644DFF95FE55EE9BD26BFC47" box="[473,602,907,931]" italics="true" pageId="36" pageNumber="37">Xenoceltites</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C22D788644DFF95FDE8EE9BD336FC47" authorityName="Spath" authorityYear="1930" box="[612,775,907,931]" class="Cephalopoda" family="Xenoceltitidae" genus="Glyptophiceras" kingdom="Animalia" pageId="36" pageNumber="37" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B9567019644DFF95FDE8EE9BD336FC47" box="[612,775,907,931]" italics="true" pageId="36" pageNumber="37">Glyptophiceras</emphasis>
|
||
</taxonomicName>
|
||
beds,
|
||
<collectingRegion id="49E662E9644DFF95FF52EEBBD10CFC27" box="[222,317,939,963]" country="China" name="Guangxi" pageId="36" pageNumber="37">Guangxi</collectingRegion>
|
||
, South
|
||
<collectingCountry id="F335EC9B644DFF95FE11EEBBD1EEFC27" box="[413,479,939,963]" name="China" pageId="36" pageNumber="37">China</collectingCountry>
|
||
(this study).
|
||
<collectingCountry id="F335EC9B644DFF95FD0FEEBBD2F9FC27" box="[643,712,939,963]" name="Oman" pageId="36" pageNumber="37">Oman</collectingCountry>
|
||
(Leu et al., submitted).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |