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<document id="56E98BBD2FCA131D89885C9D1A369FA0" ID-CLB-Dataset="20416" ID-DOI="10.5852/ejt.2022.798.1671" ID-GBIF-Dataset="dc185c1a-2bc2-48f1-bbdd-e0ae42f852f1" ID-ISSN="2118-9773" ID-Zenodo-Dep="6323040" ID-ZooBank="F136E044-62C8-4FB3-8160-7DAE663D9600" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646225490206" checkinUser="felipe" docAuthor="Houben, Albrecht M., Monnens, Marlies, Proesmans, Willem &amp; Artois, Tom J." docDate="2022" docId="038A87DAA765FF870412FD52FE800819" docLanguage="en" docName="ejt-798_70-102.pdf" docOrigin="European Journal of Taxonomy 798" docSource="http://dx.doi.org/10.5852/ejt.2022.798.1671" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Protoplanella simplex Reisinger 1924" docType="treatment" docVersion="12" lastPageNumber="99" masterDocId="FFB3FFA2A779FF9A0626FF8AFFA50F38" masterDocTitle="Limnoterrestrial ‘ Typhloplanidae’ (Rhabdocoela, Platyhelminthes), with the description of four new species and a new genus" masterLastPageNumber="102" masterPageNumber="70" pageNumber="98" updateTime="1699316596528" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="7E3EB7854BA9D9C53E52C73212CC8878" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1664F1BA34A639B6923B11852D077C99">
<mods:title id="664F631E96373391971FD581BA72228C">Limnoterrestrial ‘ Typhloplanidae’ (Rhabdocoela, Platyhelminthes), with the description of four new species and a new genus</mods:title>
</mods:titleInfo>
<mods:name id="BBCDAFFEA238C4A787A22403AEB720EE" type="personal">
<mods:role id="BAECEB8DE6D590D241EA639B44DB7491">
<mods:roleTerm id="0DDC163E41E29CF02B80A7148FC7A614">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B93EE7F3EA73B774C081303B6A14A8F9">Houben, Albrecht M.</mods:namePart>
<mods:nameIdentifier id="1732282B11C7AFB9728B28A00547A585" type="LSID">47A2BBE9-0820-4E83-B8BD-B41A66C214ED</mods:nameIdentifier>
<mods:affiliation id="C19C3F7AECD627F4E5ED574F6CFA0DFC">Biodiversity &amp; Toxicology, Agoralaan Gebouw D, B- 3590 Diepenbeek, Belgium.</mods:affiliation>
<mods:nameIdentifier id="826DFEE71019A1000E1ABFAFAE9826F1" type="email">albrecht.houben@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="B6DBD319E75F5A63246FA315FAA399C6" type="personal">
<mods:role id="70ECBA53BBAFC438F6B053B000BDBF9F">
<mods:roleTerm id="08B85DAF29AA8BE994E5A4D2FBD21250">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BD319F2D22DCBFE2F87289D9602F980F">Monnens, Marlies</mods:namePart>
<mods:nameIdentifier id="7A73C76794E721ADD88EA219E1652DFC" type="LSID">782F71E0-EF84-48DA-BE72-8E205CB78EAC</mods:nameIdentifier>
<mods:affiliation id="D9E03F0C998D8112827E059B922F4CC2">Biodiversity &amp; Toxicology, Agoralaan Gebouw D, B- 3590 Diepenbeek, Belgium.</mods:affiliation>
<mods:nameIdentifier id="F680697CA2F4E63B3630A3A234D241DA" type="email">marlies.monnens@uhasselt.be</mods:nameIdentifier>
</mods:name>
<mods:name id="BE73C7206FAA63EF3A23E9246EA1DF41" type="personal">
<mods:role id="006AFF4AB246A65F93A932C3759F52A3">
<mods:roleTerm id="927FD2D440B6B6251304C2F9FB6A2C4F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="95CD41B54F89F2692CDD926B8CE39D6D">Proesmans, Willem</mods:namePart>
<mods:nameIdentifier id="08AD5F40B5C1F847ED1855A75A4F9C57" type="LSID">41B81434-06EE-4BD0-8DCE-4DE73F2A6B30</mods:nameIdentifier>
<mods:affiliation id="8F7D69E2761108F1290C80C05FC9ED0A">Agroécologie, INRAE, 17 Rue Sully, F- 21065 Dijon, France.</mods:affiliation>
<mods:nameIdentifier id="9A4CBC6AD289832BF994E683325E0308" type="email">willem.proesmans@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="D3D0184A0005300D6F8DCE464A195127" type="personal">
<mods:role id="148C995F26F5763967B3007E7DD7E328">
<mods:roleTerm id="EB045D18BF5101EF5CE957A389BF0C12">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="285D88237DC113A51A66D812E5968B9D">Artois, Tom J.</mods:namePart>
<mods:nameIdentifier id="C6D7DEE5F85D720C395A2EF0E462CC3F" type="LSID">2EDDE35C-A2F0-4CA2-84AA-2A7893C40AC4</mods:nameIdentifier>
<mods:affiliation id="EDF19670E17EEFAF67EBBD3E130AC2A1">Biodiversity &amp; Toxicology, Agoralaan Gebouw D, B- 3590 Diepenbeek, Belgium.</mods:affiliation>
<mods:nameIdentifier id="CE345CBC378B8FA091B701A30FF0867A" type="email">tom.artois@uhasselt.be</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="8CE0D84A10C797C3243B4899A1A6F436">text</mods:typeOfResource>
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<mods:titleInfo id="79F4016ABACA969A7770307598DE18C6">
<mods:title id="305E769CE5DDC0B3CE990B52B3978627">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="FB976A781D05FA4D40FBF31C979FC8FA">
<mods:date id="DA412006819CD50FF9484F42146BEE99">2022</mods:date>
<mods:detail id="E5E51F1DDB52C15808CDC31D1A5A33C4" type="pubDate">
<mods:number id="D23AB39FB3061BE99EBB6C5A4DE6806E">2022-03-01</mods:number>
</mods:detail>
<mods:detail id="BF60D1EB2B34D392BF5C62AD8480EB9C" type="volume">
<mods:number id="EF9DB3E1EF495AC051DB67AB879D0FD7">798</mods:number>
</mods:detail>
<mods:extent id="BF4ED310AB676C1FCD8BDB1CD691BFF7" unit="page">
<mods:start id="941C1DE05E7F24BE49B99534C43B1B5E">70</mods:start>
<mods:end id="56B366E680B5194C357F3D6439A817E5">102</mods:end>
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<mods:url id="34F51954910BB769C54ED8BDC98C82F5">http://dx.doi.org/10.5852/ejt.2022.798.1671</mods:url>
</mods:location>
<mods:classification id="1542DBC6648A7D11E4F9E1E3FD52800C">journal article</mods:classification>
<mods:identifier id="F925EB04F5DC0B8E9C98945A4648753B" type="CLB-Dataset">20416</mods:identifier>
<mods:identifier id="A0FD748CA48658F472F60128CDC10E9B" type="DOI">10.5852/ejt.2022.798.1671</mods:identifier>
<mods:identifier id="00E17567FEBCE35C85FF592796406398" type="GBIF-Dataset">dc185c1a-2bc2-48f1-bbdd-e0ae42f852f1</mods:identifier>
<mods:identifier id="1C0FE01F554423D3BE18F1AF577799DA" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="05EB62985D47DEACE58AB8A541D5B52D" type="Zenodo-Dep">6323040</mods:identifier>
<mods:identifier id="5417BE3EEC563C42E58B95FBE91803F8" type="ZooBank">F136E044-62C8-4FB3-8160-7DAE663D9600</mods:identifier>
</mods:mods>
<treatment id="038A87DAA765FF870412FD52FE800819" ID-DOI="http://doi.org/10.5281/zenodo.7450121" ID-GBIF-Taxon="193858138" ID-Zenodo-Dep="7450121" LSID="urn:lsid:plazi:treatment:038A87DAA765FF870412FD52FE800819" httpUri="http://treatment.plazi.org/id/038A87DAA765FF870412FD52FE800819" lastPageId="29" lastPageNumber="99" pageId="28" pageNumber="98">
<subSubSection id="C3396547A765FF860412FD52FBA50DCB" box="[564,1024,728,755]" pageId="28" pageNumber="98" type="nomenclature">
<paragraph id="8B9C36CCA765FF860412FD52FBA50DCB" blockId="28.[564,1024,728,790]" box="[564,1024,728,755]" pageId="28" pageNumber="98">
<heading id="D0D481A0A765FF860412FD52FBA50DCB" box="[564,1024,728,755]" centered="true" fontSize="11" level="2" pageId="28" pageNumber="98" reason="2">
<taxonomicName id="4C234D4FA765FF860412FD52FBA50DCB" ID-CoL="4N6ML" authority="Reisinger, 1924" authorityName="Reisinger" authorityYear="1924" box="[564,1024,728,755]" family="Typhloplanidae" genus="Protoplanella" higherTaxonomySource="CoL" kingdom="Animalia" order="Rhabdocoela" pageId="28" pageNumber="98" phylum="Platyhelminthes" rank="species" species="simplex">
<emphasis id="B957EADEA765FF860412FD52FCE50DCB" bold="true" box="[564,832,728,755]" italics="true" pageId="28" pageNumber="98">Protoplanella simplex</emphasis>
<bibRefCitation id="EFB24B3DA765FF860561FD53FBA50DCB" author="Reisinger E." box="[839,1024,729,755]" pageId="28" pageNumber="98" pagination="128 - 143" refId="ref15836" refString="Reisinger E. 1924. Die terricolen Rhabdocoelen Steiermarks. Zoologischer Anzeiger 59: 33 - 48, 72 - 86, 128 - 143." type="journal article" year="1924">Reisinger, 1924</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3396547A765FF8604D7FD76FCE70C2E" box="[753,834,764,790]" pageId="28" pageNumber="98" type="description">
<paragraph id="8B9C36CCA765FF8604D7FD76FCE70C2E" blockId="28.[564,1024,728,790]" box="[753,834,764,790]" pageId="28" pageNumber="98">
<figureCitation id="13182A49A765FF8604D7FD76FCE70C2E" box="[753,834,764,790]" captionStart="Fig" captionStartId="30.[189,232,1928,1954]" captionTargetBox="[224,1353,955,1890]" captionTargetId="figure-156@30.[189,1398,910,1890]" captionTargetPageId="30" captionText="Fig. 11. Protoplanella simplex Reisinger, 1924. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 825)." figureDoi="http://doi.org/10.5281/zenodo.6323067" httpUri="https://zenodo.org/record/6323067/files/figure.png" pageId="28" pageNumber="98">Fig. 11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3396547A765FF86069BFCB7FB190A28" pageId="28" pageNumber="98" type="materials_examined">
<paragraph id="8B9C36CCA765FF86069BFCB7FE080C6F" blockId="28.[189,429,829,855]" box="[189,429,829,855]" pageId="28" pageNumber="98">
<heading id="D0D481A0A765FF86069BFCB7FE080C6F" bold="true" box="[189,429,829,855]" fontSize="11" level="3" pageId="28" pageNumber="98" reason="3">
<emphasis id="B957EADEA765FF86069BFCB7FE080C6F" bold="true" box="[189,429,829,855]" pageId="28" pageNumber="98">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B9C36CCA765FF8606CCFCE6FAA00CF5" blockId="28.[188,1399,876,973]" pageId="28" pageNumber="98">
<materialsCitation id="3B4B3C91A765FF8606CCFCE6FAA00CF5" ID-GBIF-Occurrence="3695141318" collectedFrom="mosses growing on a wall" collectingDate="2011-07-24" collectionCode="HU" collectorName="A. M. Houben &amp; W. Proesmans" country="Germany" latitude="49.823334" location="Kordel" longLatPrecision="18" longitude="6.6349998" pageId="28" pageNumber="98" specimenCode="825" specimenCount="1" typeStatus="neotype">
<emphasis id="B957EADEA765FF8606CCFCE6FEF70CBE" bold="true" box="[234,338,876,902]" pageId="28" pageNumber="98">
<typeStatus id="5498886EA765FF8606CCFCE6FEF70CBE" box="[234,338,876,902]" pageId="28" pageNumber="98" type="neotype">Neotype</typeStatus>
</emphasis>
<collectingCountry id="F334765CA765FF86069BFC05FEF50C91" box="[189,336,911,937]" name="Germany" pageId="28" pageNumber="98">GERMANY</collectingCountry>
•
<specimenCount id="9D25FD45A765FF860741FC05FE1E0C92" box="[359,443,911,938]" pageId="28" pageNumber="98" type="generic">1 spec.</specimenCount>
, live observations and sagittal sections;
<location id="8EFC6017A765FF8605AAFC05FC440C91" LSID="urn:lsid:plazi:treatment:038A87DAA765FF870412FD52FE800819:8EFC6017A765FF8605AAFC05FC440C91" box="[908,993,911,937]" country="Germany" latitude="49.823334" longLatPrecision="18" longitude="6.6349998" name="Kordel" pageId="28" pageNumber="98">Kordel</location>
;
<geoCoordinate id="EE17500BA765FF8605CAFC05FB240C92" box="[1004,1153,909,938]" degrees="49" direction="north" minutes="49" orientation="latitude" pageId="28" pageNumber="98" precision="15" seconds="24" value="49.823334">49°49′24″ N</geoCoordinate>
,
<geoCoordinate id="EE17500BA765FF8602AAFC05FABB0C92" box="[1164,1310,909,938]" degrees="06" direction="east" minutes="38" orientation="longitude" pageId="28" pageNumber="98" precision="15" seconds="06" value="6.6349998">06°38′06″ E</geoCoordinate>
;
<date id="FF9D100CA765FF86030FFC05FF5C0CF4" pageId="28" pageNumber="98" value="2011-07-24">
<collectingDate id="EFD9E9E4A765FF86030FFC05FF5C0CF4" pageId="28" pageNumber="98" value="2011-07-24">24 Jul. 2011</collectingDate>
</date>
;
<collectorName id="26D6531AA765FF860722FC38FE030CF4" box="[260,422,946,972]" pageId="28" pageNumber="98">A.M. Houben</collectorName>
and
<collectorName id="26D6531AA765FF8607F8FC39FD200CF5" box="[478,645,947,973]" pageId="28" pageNumber="98">W. Proesmans</collectorName>
leg.;
<collectedFrom id="383F7531A765FF8604E1FC39FC5E0CF4" box="[711,1019,946,973]" pageId="28" pageNumber="98">mosses growing on a wall</collectedFrom>
; neotype no.
<specimenCode id="DB859EB7A765FF8602B1FC38FB630CF5" box="[1175,1222,946,973]" pageId="28" pageNumber="98">825</specimenCode>
;
<collectionCode id="ED32AE09A765FF8602F5FC39FAA40CF5" box="[1235,1281,947,973]" pageId="28" pageNumber="98">HU</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B9C36CCA765FF86069BFC79FEDE0B35" blockId="28.[189,379,1011,1037]" box="[189,379,1011,1037]" pageId="28" pageNumber="98">
<heading id="D0D481A0A765FF86069BFC79FEDE0B35" bold="true" box="[189,379,1011,1037]" fontSize="11" level="3" pageId="28" pageNumber="98" reason="3">
<emphasis id="B957EADEA765FF86069BFC79FEDE0B35" bold="true" box="[189,379,1011,1037]" pageId="28" pageNumber="98">Other material</emphasis>
</heading>
</paragraph>
<paragraph id="8B9C36CCA765FF86069BFBA8FB190A28" blockId="28.[188,1400,1058,1296]" pageId="28" pageNumber="98">
<materialsCitation id="3B4B3C91A765FF86069BFBA8FB5B0B05" ID-GBIF-Occurrence="3695141313" box="[189,1278,1058,1085]" collectedFrom="mosses growing on a wall" collectingDate="2011-07-24" collectionCode="HU" collectorName="A. M. Houben &amp; W. Proesmans" country="Germany" latitude="49.823334" location="GERMANY" longLatPrecision="18" longitude="6.6349998" pageId="28" pageNumber="98" specimenCount="1">
<collectingCountry id="F334765CA765FF86069BFBA8FEF50B04" box="[189,336,1058,1084]" name="Germany" pageId="28" pageNumber="98">GERMANY</collectingCountry>
•
<specimenCount id="9D25FD45A765FF86074DFBA8FE1A0B05" box="[363,447,1058,1085]" pageId="28" pageNumber="98" type="generic">1 spec.</specimenCount>
, serially sectioned; same collection data as for neotype; XIV.3.11;
<collectionCode id="ED32AE09A765FF8602F2FBA9FB5B0B05" box="[1236,1278,1059,1085]" pageId="28" pageNumber="98">HU</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B4B3C91A765FF86033FFBA8FC540BBB" ID-GBIF-Occurrence="3695141326" collectedFrom="forest litter" collectingDate="2011-07-13" collectionCode="HU" collectorName="A. M. Houben &amp; W. Proesmans" country="Germany" latitude="7.559167" location="Oberau" longLatPrecision="21" longitude="11.115833" pageId="28" pageNumber="98" specimenCount="2" stateProvince="Bavaria">
<specimenCount id="9D25FD45A765FF86033FFBA8FAD70B05" box="[1305,1394,1058,1085]" pageId="28" pageNumber="98">2 specs</specimenCount>
, live observations, one of which serially sectioned;
<collectingRegion id="49E7F82EA765FF860533FBCFFCD60B67" box="[789,883,1093,1119]" country="Germany" name="Bayern" pageId="28" pageNumber="98">Bavaria</collectingRegion>
,
<location id="8EFC6017A765FF8605A6FBCFFC7F0B67" LSID="urn:lsid:plazi:treatment:038A87DAA765FF870412FD52FE800819:8EFC6017A765FF8605A6FBCFFC7F0B67" box="[896,986,1093,1119]" country="Germany" latitude="7.559167" longLatPrecision="21" longitude="11.115833" name="Oberau" pageId="28" pageNumber="98" stateProvince="Bavaria">Oberau</location>
;
<geoCoordinate id="EE17500BA765FF8605C0FBCCFBD90B58" box="[998,1148,1092,1120]" degrees="07" direction="north" minutes="33" orientation="latitude" pageId="28" pageNumber="98" precision="15" seconds="33" value="7.559167">07°33′33″ N</geoCoordinate>
,
<geoCoordinate id="EE17500BA765FF8602AEFBCFFABF0B58" box="[1160,1306,1092,1120]" degrees="11" direction="east" minutes="06" orientation="longitude" pageId="28" pageNumber="98" precision="15" seconds="57" value="11.115833">11°06′57″ E</geoCoordinate>
;
<date id="FF9D100CA765FF86030EFBCCFF5E0BBB" pageId="28" pageNumber="98" value="2011-07-13">
<collectingDate id="EFD9E9E4A765FF86030EFBCCFF5E0BBB" pageId="28" pageNumber="98" value="2011-07-13">13 Jul. 2011</collectingDate>
</date>
;
<collectorName id="26D6531AA765FF860722FBE3FE000BBB" box="[260,421,1129,1155]" pageId="28" pageNumber="98">A.M. Houben</collectorName>
and
<collectorName id="26D6531AA765FF8607FCFBE3FD250BBB" box="[474,640,1129,1155]" pageId="28" pageNumber="98">W. Proesmans</collectorName>
leg.;
<collectedFrom id="383F7531A765FF860499FBE3FCE60BBB" box="[703,835,1129,1155]" pageId="28" pageNumber="98">forest litter</collectedFrom>
; XIV.3.12;
<collectionCode id="ED32AE09A765FF8605E1FBE3FC540BBB" box="[967,1009,1129,1155]" pageId="28" pageNumber="98">HU</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B4B3C91A765FF860221FBE2FC730BF2" ID-GBIF-Occurrence="3695141309" collectedFrom="mosses at the forest edge" collectingDate="2011-07-27" collectionCode="HU" collectorName="A. M. Houben &amp; W. Proesmans" country="Germany" latitude="50.934166" location="Lanaken" longLatPrecision="17" longitude="5.6600003" pageId="28" pageNumber="98" specimenCount="5">
<specimenCount id="9D25FD45A765FF860221FBE2FBFB0BBB" box="[1031,1118,1128,1155]" pageId="28" pageNumber="98">5 specs</specimenCount>
, live observations, three of which serially sectioned;
<location id="8EFC6017A765FF860429FB06FDDD0B9E" LSID="urn:lsid:plazi:treatment:038A87DAA765FF870412FD52FE800819:8EFC6017A765FF860429FB06FDDD0B9E" box="[527,632,1164,1190]" country="Germany" latitude="50.934166" longLatPrecision="17" longitude="5.6600003" name="Lanaken" pageId="28" pageNumber="98">Lanaken</location>
;
<geoCoordinate id="EE17500BA765FF8604A0FB06FCBA0B9E" box="[646,799,1162,1191]" degrees="50" direction="north" minutes="56" orientation="latitude" pageId="28" pageNumber="98" precision="15" seconds="03" value="50.934166">50°56′03″ N</geoCoordinate>
;
<geoCoordinate id="EE17500BA765FF860508FB06FC660B9E" box="[814,963,1162,1191]" degrees="05" direction="east" minutes="39" orientation="longitude" pageId="28" pageNumber="98" precision="15" seconds="36" value="5.6600003">05°39′36″ E</geoCoordinate>
;
<date id="FF9D100CA765FF8605F7FB06FBCD0B9E" box="[977,1128,1164,1190]" pageId="28" pageNumber="98" value="2011-07-27">
<collectingDate id="EFD9E9E4A765FF8605F7FB06FBCD0B9E" box="[977,1128,1164,1190]" pageId="28" pageNumber="98" value="2011-07-27">27 Jul. 2011</collectingDate>
</date>
;
<collectorName id="26D6531AA765FF860253FB06FABC0B9E" box="[1141,1305,1164,1190]" pageId="28" pageNumber="98">A.M. Houben</collectorName>
and
<collectorName id="26D6531AA765FF860370FB06FE9B0BF2" pageId="28" pageNumber="98">W. Proesmans</collectorName>
leg.;
<collectedFrom id="383F7531A765FF8607A7FB3AFD0E0BF1" box="[385,683,1199,1226]" pageId="28" pageNumber="98">mosses at the forest edge</collectedFrom>
; XIV.3.13–XIV.3.15;
<collectionCode id="ED32AE09A765FF86058AFB3AFC730BF2" box="[940,982,1200,1226]" pageId="28" pageNumber="98">HU</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B4B3C91A765FF8605D6FB25FB1D0A28" ID-GBIF-Occurrence="3695141311" collectedFrom="moist forest soil" collectingDate="2011-07-27" collectionCode="HU" collectorName="A. M. Houben &amp; W. Proesmans" country="Germany" county="Lanaken" latitude="50.93389" location="Hoge Kempen'" longLatPrecision="17" longitude="5.660556" municipality="National Park" pageId="28" pageNumber="98" specimenCount="4">
<specimenCount id="9D25FD45A765FF8605D6FB25FBEC0BF2" box="[1008,1097,1199,1226]" pageId="28" pageNumber="98">4 specs</specimenCount>
, live observations, two of which serially sectioned;
<collectingCounty id="62FD4E40A765FF8607CBFB58FDF00BD4" box="[493,597,1234,1260]" pageId="28" pageNumber="98">Lanaken</collectingCounty>
,
<collectingMunicipality id="6BF8ACB6A765FF860444FB58FCA30BD4" box="[610,774,1234,1260]" pageId="28" pageNumber="98">National Park</collectingMunicipality>
‘
<location id="8EFC6017A765FF860535FB58FC770BD4" LSID="urn:lsid:plazi:treatment:038A87DAA765FF870412FD52FE800819:8EFC6017A765FF860535FB58FC770BD4" box="[787,978,1234,1260]" country="Germany" county="Lanaken" latitude="50.93389" longLatPrecision="17" longitude="5.660556" municipality="National Park" name="Hoge Kempen'" pageId="28" pageNumber="98">Hoge Kempen’</location>
;
<geoCoordinate id="EE17500BA765FF8605F9FB58FBD30BD5" box="[991,1142,1233,1261]" degrees="50" direction="north" minutes="56" orientation="latitude" pageId="28" pageNumber="98" precision="15" seconds="02" value="50.93389">50°56′02″ N</geoCoordinate>
;
<geoCoordinate id="EE17500BA765FF8602A2FB58FABC0BD5" box="[1156,1305,1233,1261]" degrees="05" direction="east" minutes="39" orientation="longitude" pageId="28" pageNumber="98" precision="15" seconds="38" value="5.660556">05°39′38″ E</geoCoordinate>
;
<date id="FF9D100CA765FF860301FB58FF590A28" pageId="28" pageNumber="98" value="2011-07-27">
<collectingDate id="EFD9E9E4A765FF860301FB58FF590A28" pageId="28" pageNumber="98" value="2011-07-27">27 Jul. 2011</collectingDate>
</date>
;
<collectorName id="26D6531AA765FF860720FB7CFE0D0A37" box="[262,424,1269,1296]" pageId="28" pageNumber="98">A.M. Houben</collectorName>
and
<collectorName id="26D6531AA765FF8607C7FB7CFD220A28" box="[481,647,1270,1296]" pageId="28" pageNumber="98">W. Proesmans</collectorName>
leg.;
<collectedFrom id="383F7531A765FF8604EFFB7FFC2F0A37" box="[713,906,1269,1295]" pageId="28" pageNumber="98">moist forest soil</collectedFrom>
; XIV.3.16–XIV.3.17;
<collectionCode id="ED32AE09A765FF8602ACFB7CFB1D0A28" box="[1162,1208,1270,1296]" pageId="28" pageNumber="98">HU</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3396547A765FF87069BFABDFB690E88" lastPageId="29" lastPageNumber="99" pageId="28" pageNumber="98" type="description">
<paragraph id="8B9C36CCA765FF86069BFABDFEEA0A69" blockId="28.[189,335,1335,1361]" box="[189,335,1335,1361]" pageId="28" pageNumber="98">
<heading id="D0D481A0A765FF86069BFABDFEEA0A69" bold="true" box="[189,335,1335,1361]" fontSize="11" level="3" pageId="28" pageNumber="98" reason="3">
<emphasis id="B957EADEA765FF86069BFABDFEEA0A69" bold="true" box="[189,335,1335,1361]" pageId="28" pageNumber="98">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B9C36CCA765FF86069BFAEFFBC70934" blockId="28.[188,1399,1381,1549]" pageId="28" pageNumber="98">
The studied specimens are about
<quantity id="4CDB9B29A765FF86047BFAECFD190AB8" box="[605,700,1382,1408]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="28" pageNumber="98" unit="mm" value="0.8">0.8 mm</quantity>
long. The anterior and posterior body ends are rounded (
<figureCitation id="13182A49A765FF8606E0FA03FE940A9B" box="[198,305,1417,1443]" captionStart="Fig" captionStartId="30.[189,232,1928,1954]" captionTargetBox="[224,1353,955,1890]" captionTargetId="figure-156@30.[189,1398,910,1890]" captionTargetPageId="30" captionText="Fig. 11. Protoplanella simplex Reisinger, 1924. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 825)." figureDoi="http://doi.org/10.5281/zenodo.6323067" httpUri="https://zenodo.org/record/6323067/files/figure.png" pageId="28" pageNumber="98">Fig. 11A</figureCitation>
). Rhabdite glands are present and arranged into two groups behind the brain (
<figureCitation id="13182A49A765FF860289FA03FABF0A9B" box="[1199,1306,1417,1443]" captionStart="Fig" captionStartId="30.[189,232,1928,1954]" captionTargetBox="[224,1353,955,1890]" captionTargetId="figure-156@30.[189,1398,910,1890]" captionTargetPageId="30" captionText="Fig. 11. Protoplanella simplex Reisinger, 1924. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 825)." figureDoi="http://doi.org/10.5281/zenodo.6323067" httpUri="https://zenodo.org/record/6323067/files/figure.png" pageId="28" pageNumber="98">Fig. 11A</figureCitation>
: ar); the rhabdites themselves extend forward in two anastomosing tracts. Dermal rhabdites were not observed. The paired protonephridiopores (
<figureCitation id="13182A49A765FF860476FA45FD640AD1" box="[592,705,1487,1513]" captionStart="Fig" captionStartId="30.[189,232,1928,1954]" captionTargetBox="[224,1353,955,1890]" captionTargetId="figure-156@30.[189,1398,910,1890]" captionTargetPageId="30" captionText="Fig. 11. Protoplanella simplex Reisinger, 1924. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 825)." figureDoi="http://doi.org/10.5281/zenodo.6323067" httpUri="https://zenodo.org/record/6323067/files/figure.png" pageId="28" pageNumber="98">Fig. 11A</figureCitation>
: pp) open laterally and somewhat caudal to the rosulate pharynx (
<figureCitation id="13182A49A765FF86070BFA78FE1E0934" box="[301,443,1522,1548]" captionStart="Fig" captionStartId="30.[189,232,1928,1954]" captionTargetBox="[224,1353,955,1890]" captionTargetId="figure-156@30.[189,1398,910,1890]" captionTargetPageId="30" captionText="Fig. 11. Protoplanella simplex Reisinger, 1924. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 825)." figureDoi="http://doi.org/10.5281/zenodo.6323067" httpUri="https://zenodo.org/record/6323067/files/figure.png" pageId="28" pageNumber="98">Fig. 11A, C</figureCitation>
: ph), which is situated just behind the middle of the body.
</paragraph>
<paragraph id="8B9C36CCA765FF86069BF9BEFF5A09AC" blockId="28.[189,1399,1588,1684]" pageId="28" pageNumber="98">
The gonopore (
<figureCitation id="13182A49A765FF86075FF9BEFDA80976" box="[377,525,1588,1614]" captionStart="Fig" captionStartId="30.[189,232,1928,1954]" captionTargetBox="[224,1353,955,1890]" captionTargetId="figure-156@30.[189,1398,910,1890]" captionTargetPageId="30" captionText="Fig. 11. Protoplanella simplex Reisinger, 1924. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 825)." figureDoi="http://doi.org/10.5281/zenodo.6323067" httpUri="https://zenodo.org/record/6323067/files/figure.png" pageId="28" pageNumber="98">Fig. 11A–C</figureCitation>
: gp) is located at ±80% of the body and connected to a genital atrium (
<figureCitation id="13182A49A765FF8606E0F9DDFEF30949" box="[198,342,1623,1649]" captionStart="Fig" captionStartId="30.[189,232,1928,1954]" captionTargetBox="[224,1353,955,1890]" captionTargetId="figure-156@30.[189,1398,910,1890]" captionTargetPageId="30" captionText="Fig. 11. Protoplanella simplex Reisinger, 1924. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 825)." figureDoi="http://doi.org/10.5281/zenodo.6323067" httpUri="https://zenodo.org/record/6323067/files/figure.png" pageId="28" pageNumber="98">Fig. 11A–C</figureCitation>
: ga). The genital atrium is surrounded by an inner circular and outer longitudinal muscle layer.
</paragraph>
<paragraph id="8B9C36CCA765FF86069BF931FDE708F4" blockId="28.[189,1401,1723,1996]" pageId="28" pageNumber="98">
The small, round testes (
<figureCitation id="13182A49A765FF8607D0F931FD2909EE" box="[502,652,1723,1750]" captionStart="Fig" captionStartId="30.[189,232,1928,1954]" captionTargetBox="[224,1353,955,1890]" captionTargetId="figure-156@30.[189,1398,910,1890]" captionTargetPageId="30" captionText="Fig. 11. Protoplanella simplex Reisinger, 1924. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 825)." figureDoi="http://doi.org/10.5281/zenodo.6323067" httpUri="https://zenodo.org/record/6323067/files/figure.png" pageId="28" pageNumber="98">Fig. 11A–B</figureCitation>
: t) lie posterior to the pharynx and ventral to the vitellaria (
<figureCitation id="13182A49A765FF8606E0F955FEFD09C1" box="[198,344,1759,1785]" captionStart="Fig" captionStartId="30.[189,232,1928,1954]" captionTargetBox="[224,1353,955,1890]" captionTargetId="figure-156@30.[189,1398,910,1890]" captionTargetPageId="30" captionText="Fig. 11. Protoplanella simplex Reisinger, 1924. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 825)." figureDoi="http://doi.org/10.5281/zenodo.6323067" httpUri="https://zenodo.org/record/6323067/files/figure.png" pageId="28" pageNumber="98">Fig. 11A–C</figureCitation>
: vi). Both vasa deferentia (
<figureCitation id="13182A49A765FF860482F955FC9109C1" box="[676,820,1759,1785]" captionStart="Fig" captionStartId="30.[189,232,1928,1954]" captionTargetBox="[224,1353,955,1890]" captionTargetId="figure-156@30.[189,1398,910,1890]" captionTargetPageId="30" captionText="Fig. 11. Protoplanella simplex Reisinger, 1924. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 825)." figureDoi="http://doi.org/10.5281/zenodo.6323067" httpUri="https://zenodo.org/record/6323067/files/figure.png" pageId="28" pageNumber="98">Fig. 11B–C</figureCitation>
: vde) enter the copulatory organ (
<figureCitation id="13182A49A765FF8602F5F955FAE709C1" box="[1235,1346,1759,1785]" captionStart="Fig" captionStartId="30.[189,232,1928,1954]" captionTargetBox="[224,1353,955,1890]" captionTargetId="figure-156@30.[189,1398,910,1890]" captionTargetPageId="30" captionText="Fig. 11. Protoplanella simplex Reisinger, 1924. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 825)." figureDoi="http://doi.org/10.5281/zenodo.6323067" httpUri="https://zenodo.org/record/6323067/files/figure.png" pageId="28" pageNumber="98">Fig. 11A</figureCitation>
: co) separately from the lateral side. This 22 µm long, oval-shaped copulatory organ is surrounded by two layers of spiral muscles and contains an intracapsular seminal vesicle (
<figureCitation id="13182A49A765FF860239F8AFFB0A0807" box="[1055,1199,1829,1855]" captionStart="Fig" captionStartId="30.[189,232,1928,1954]" captionTargetBox="[224,1353,955,1890]" captionTargetId="figure-156@30.[189,1398,910,1890]" captionTargetPageId="30" captionText="Fig. 11. Protoplanella simplex Reisinger, 1924. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 825)." figureDoi="http://doi.org/10.5281/zenodo.6323067" httpUri="https://zenodo.org/record/6323067/files/figure.png" pageId="28" pageNumber="98">Fig. 11B–C</figureCitation>
: vs) with a low, nucleated epithelium and an ejaculatory duct (
<figureCitation id="13182A49A765FF8604C0F8C2FCD0085A" box="[742,885,1864,1890]" captionStart="Fig" captionStartId="30.[189,232,1928,1954]" captionTargetBox="[224,1353,955,1890]" captionTargetId="figure-156@30.[189,1398,910,1890]" captionTargetPageId="30" captionText="Fig. 11. Protoplanella simplex Reisinger, 1924. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 825)." figureDoi="http://doi.org/10.5281/zenodo.6323067" httpUri="https://zenodo.org/record/6323067/files/figure.png" pageId="28" pageNumber="98">Fig. 11B–C</figureCitation>
: de), which is surrounded by two layers of spiral muscles. Extracapsular, eosinophilic prostate glands (
<figureCitation id="13182A49A765FF8605B3F8E1FB8C08BE" box="[917,1065,1899,1926]" captionStart="Fig" captionStartId="30.[189,232,1928,1954]" captionTargetBox="[224,1353,955,1890]" captionTargetId="figure-156@30.[189,1398,910,1890]" captionTargetPageId="30" captionText="Fig. 11. Protoplanella simplex Reisinger, 1924. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 825)." figureDoi="http://doi.org/10.5281/zenodo.6323067" httpUri="https://zenodo.org/record/6323067/files/figure.png" pageId="28" pageNumber="98">Fig. 11A–C</figureCitation>
: gg) open laterally into the copulatory organ. A male duct (
<figureCitation id="13182A49A765FF860408F805FD3C0891" box="[558,665,1935,1961]" captionStart="Fig" captionStartId="30.[189,232,1928,1954]" captionTargetBox="[224,1353,955,1890]" captionTargetId="figure-156@30.[189,1398,910,1890]" captionTargetPageId="30" captionText="Fig. 11. Protoplanella simplex Reisinger, 1924. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 825)." figureDoi="http://doi.org/10.5281/zenodo.6323067" httpUri="https://zenodo.org/record/6323067/files/figure.png" pageId="28" pageNumber="98">Fig. 11C</figureCitation>
: md) with the same musculature as the genital atrium connects the copulatory organ to the latter.
</paragraph>
<paragraph id="8B9C36CCA764FF87069BFE83FB690E88" blockId="29.[188,1401,265,433]" pageId="29" pageNumber="99">
The paired vitellaria extend from 25% of the body to the posterior end where they fuse via a broad anastomosis. A single ovary (
<figureCitation id="13182A49A764FF87043BFEA6FD0B0E7E" box="[541,686,300,326]" captionStart="Fig" captionStartId="30.[189,232,1928,1954]" captionTargetBox="[224,1353,955,1890]" captionTargetId="figure-156@30.[189,1398,910,1890]" captionTargetPageId="30" captionText="Fig. 11. Protoplanella simplex Reisinger, 1924. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 825)." figureDoi="http://doi.org/10.5281/zenodo.6323067" httpUri="https://zenodo.org/record/6323067/files/figure.png" pageId="29" pageNumber="99">Fig. 11A–C</figureCitation>
: ov) is closely associated with one of the vitellaria forming an ovovitellarium. The female duct (
<figureCitation id="13182A49A764FF87044AFEC5FD720E51" box="[620,727,335,361]" captionStart="Fig" captionStartId="30.[189,232,1928,1954]" captionTargetBox="[224,1353,955,1890]" captionTargetId="figure-156@30.[189,1398,910,1890]" captionTargetPageId="30" captionText="Fig. 11. Protoplanella simplex Reisinger, 1924. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 825)." figureDoi="http://doi.org/10.5281/zenodo.6323067" httpUri="https://zenodo.org/record/6323067/files/figure.png" pageId="29" pageNumber="99">Fig. 11C</figureCitation>
: fd) is surrounded by the same musculature as the genital atrium and receives a club-shaped seminal receptacle (
<figureCitation id="13182A49A764FF870512FEF8FC670EB5" box="[820,962,370,397]" captionStart="Fig" captionStartId="30.[189,232,1928,1954]" captionTargetBox="[224,1353,955,1890]" captionTargetId="figure-156@30.[189,1398,910,1890]" captionTargetPageId="30" captionText="Fig. 11. Protoplanella simplex Reisinger, 1924. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 825)." figureDoi="http://doi.org/10.5281/zenodo.6323067" httpUri="https://zenodo.org/record/6323067/files/figure.png" pageId="29" pageNumber="99">Fig. 11A–C</figureCitation>
: rs) and ovovitelloduct at its proximal end. Fine-grained, eosinophilic glands (
<figureCitation id="13182A49A764FF8704ABFE1CFD5D0E88" box="[653,760,406,432]" captionStart="Fig" captionStartId="30.[189,232,1928,1954]" captionTargetBox="[224,1353,955,1890]" captionTargetId="figure-156@30.[189,1398,910,1890]" captionTargetPageId="30" captionText="Fig. 11. Protoplanella simplex Reisinger, 1924. A. Internal organisation of a live specimen. B. Organisation of the genital system of a live specimen. C. Reconstruction of reproductive organs based on sagittal sections of the neotype (neotype no. 825)." figureDoi="http://doi.org/10.5281/zenodo.6323067" httpUri="https://zenodo.org/record/6323067/files/figure.png" pageId="29" pageNumber="99">Fig. 11C</figureCitation>
: fg) surround and enter the female duct.
</paragraph>
</subSubSection>
<subSubSection id="C3396547A764FF87069BFE5DFD060DC1" pageId="29" pageNumber="99" type="discussion">
<paragraph id="8B9C36CCA764FF87069BFE5DFEE70EC9" blockId="29.[189,322,471,497]" box="[189,322,471,497]" pageId="29" pageNumber="99">
<heading id="D0D481A0A764FF87069BFE5DFEE70EC9" bold="true" box="[189,322,471,497]" fontSize="11" level="3" pageId="29" pageNumber="99" reason="3">
<emphasis id="B957EADEA764FF87069BFE5DFEE70EC9" bold="true" box="[189,322,471,497]" pageId="29" pageNumber="99">Discussion</emphasis>
</heading>
</paragraph>
<paragraph id="8B9C36CCA764FF87069BFD8CFC9A0D18" blockId="29.[189,831,518,544]" box="[189,831,518,544]" pageId="29" pageNumber="99">
See the general discussion on the genus
<taxonomicName id="4C234D4FA764FF8704B3FD8CFC9C0D18" authorityName="Reisinger" authorityYear="1924" box="[661,825,518,544]" family="Typhloplanidae" genus="Protoplanella" higherTaxonomySource="CoL" kingdom="Animalia" order="Rhabdocoela" pageId="29" pageNumber="99" phylum="Platyhelminthes" rank="genus">
<emphasis id="B957EADEA764FF8704B3FD8CFC9C0D18" box="[661,825,518,544]" italics="true" pageId="29" pageNumber="99">Protoplanella</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9C36CCA764FF87069BFDCDFE950D59" blockId="29.[189,304,583,609]" box="[189,304,583,609]" pageId="29" pageNumber="99">
<heading id="D0D481A0A764FF87069BFDCDFE950D59" bold="true" box="[189,304,583,609]" fontSize="11" level="3" pageId="29" pageNumber="99" reason="3">
<emphasis id="B957EADEA764FF87069BFDCDFE950D59" bold="true" box="[189,304,583,609]" pageId="29" pageNumber="99">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B9C36CCA764FF87069BFDFCFD060DC1" blockId="29.[189,1399,630,762]" pageId="29" pageNumber="99">
Intensive surveys at the type locality yielded no specimens. The specimens from Oberau,
<collectingRegion id="49E7F82EA764FF870332FDFCFAD70DA8" box="[1300,1394,630,656]" country="Germany" name="Bayern" pageId="29" pageNumber="99">Bavaria</collectingRegion>
,
<collectingCountry id="F334765CA764FF87069BFD13FE8E0D8B" box="[189,299,665,691]" name="Germany" pageId="29" pageNumber="99">Germany</collectingCountry>
, which is relatively close to the original type locality in Graz,
<collectingCountry id="F334765CA764FF870201FD13FB250D8B" box="[1063,1152,665,691]" name="Austria" pageId="29" pageNumber="99">Austria</collectingCountry>
, are of poor quality. Therefore, a specimen from Kordel,
<collectingCountry id="F334765CA764FF87044DFD36FD7D0DEE" box="[619,728,700,726]" name="Germany" pageId="29" pageNumber="99">Germany</collectingCountry>
was designated
<typeStatus id="5498886EA764FF8705BCFD37FC5F0DEF" box="[922,1018,701,727]" pageId="29" pageNumber="99" type="neotype">neotype</typeStatus>
, since all diagnostic features are exactly as described by
<bibRefCitation id="EFB24B3DA764FF8707F3FD55FD3A0DC1" author="Reisinger E." box="[469,671,735,761]" pageId="29" pageNumber="99" pagination="128 - 143" refId="ref15836" refString="Reisinger E. 1924. Die terricolen Rhabdocoelen Steiermarks. Zoologischer Anzeiger 59: 33 - 48, 72 - 86, 128 - 143." type="journal article" year="1924">Reisinger (1924)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3396547A764FF87069BFCAAFEAC0C88" pageId="29" pageNumber="99" type="distribution">
<paragraph id="8B9C36CCA764FF87069BFCAAFD990C02" blockId="29.[189,572,800,827]" box="[189,572,800,827]" pageId="29" pageNumber="99">
<heading id="D0D481A0A764FF87069BFCAAFD990C02" bold="true" box="[189,572,800,827]" fontSize="11" level="3" pageId="29" pageNumber="99" reason="3">
<emphasis id="B957EADEA764FF87069BFCAAFD990C02" bold="true" box="[189,572,800,827]" pageId="29" pageNumber="99">Previously known distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B9C36CCA764FF87069BFCDAFEAC0C88" blockId="29.[189,1399,847,944]" pageId="29" pageNumber="99">
In the vicinity of Graz,
<collectingCountry id="F334765CA764FF8707F3FCC5FD8E0C51" box="[469,555,847,873]" name="Austria" pageId="29" pageNumber="99">Austria</collectingCountry>
in forest humus (
<bibRefCitation id="EFB24B3DA764FF8704D8FCC5FC130C51" author="Reisinger E." box="[766,950,847,873]" pageId="29" pageNumber="99" pagination="128 - 143" refId="ref15836" refString="Reisinger E. 1924. Die terricolen Rhabdocoelen Steiermarks. Zoologischer Anzeiger 59: 33 - 48, 72 - 86, 128 - 143." type="journal article" year="1924">Reisinger 1924</bibRefCitation>
,
<bibRefCitation id="EFB24B3DA764FF8705E5FCC5FBA40C52" author="Reisinger E." box="[963,1025,847,874]" pageId="29" pageNumber="99" pagination="106 - 123" refId="ref15892" refString="Reisinger E. 1954. Edaphische Kleinturbellarien als bodenkundliche Leitformen. Mitteilungen des Naturwissenschaftlichen Vereines fur Karnten 64: 106 - 123." type="journal article" year="1954">1954</bibRefCitation>
;
<bibRefCitation id="EFB24B3DA764FF87022BFCC5FAE20C52" author="An der Lan H. &amp; Franz H." box="[1037,1351,847,874]" pageId="29" pageNumber="99" pagination="129 - 134" refId="ref15191" refString="An der Lan H. &amp; Franz H. 1954. Turbellaria. In: Franz H. (ed.) Die Nordost-Alpen im Spiegel ihrer Landtierwelt: 129 - 134. Universitatsverlag Wagner, Innsbruck." type="book chapter" year="1954">An der Lan &amp; Franz 1954</bibRefCitation>
;
<bibRefCitation id="EFB24B3DA764FF870375FCC5FEFA0CB5" author="An der Lan H." pageId="29" pageNumber="99" pagination="227 - 234" refId="ref15165" refString="An der Lan H. 1963. Zur Verbreitung edaphischer Kleinturbellarien in Osterreich. Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck 53: 227 - 234." type="journal article" year="1963">An der Lan 1963</bibRefCitation>
), on the
<collectingCountry id="F334765CA764FF8707E0FCF9FDC10CB4" box="[454,612,882,909]" name="Faroe Islands" pageId="29" pageNumber="99">Faroe Islands</collectingCountry>
(
<bibRefCitation id="EFB24B3DA764FF870455FCF8FC970CB5" author="Steinbock O." box="[627,818,882,909]" pageId="29" pageNumber="99" pagination="1 - 32" refId="ref16111" refString="Steinbock O. 1931. Freshwater Turbellaria. In: Jensen S., Lundbeck W. &amp; Mortensen Th. (eds) Zoology of the Faroes: 1 - 32. Copenhagen." type="book chapter" year="1931">Steinböck 1931</bibRefCitation>
); near Poznań,
<collectingCountry id="F334765CA764FF8705C1FCF8FB9E0CB4" box="[999,1083,882,908]" name="Poland" pageId="29" pageNumber="99">Poland</collectingCountry>
, in moss and litter (
<bibRefCitation id="EFB24B3DA764FF870302FCF8FF590C88" author="Kolasa J." pageId="29" pageNumber="99" pagination="7 - 31" refId="ref15535" refString="Kolasa J. 1974. Badania nad wirkami (Turbellaria) zrodel ze szczegolnym uwzglednieniem okolic Poznania (The investigations on spring Turbellaria with a special regard to surroundings of Poznan). Badania Fizjograficzne nad polska zachodnia 26: 7 - 31." type="journal article" year="1974">Kolasa 1974</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3396547A764FF87069BFC5DFE800819" pageId="29" pageNumber="99" type="discussion">
<paragraph id="8B9C36CCA764FF87069BFC5DFD250CCA" blockId="29.[189,640,983,1010]" box="[189,640,983,1010]" pageId="29" pageNumber="99">
<emphasis id="B957EADEA764FF87069BFC5DFD250CCA" bold="true" box="[189,640,983,1010]" pageId="29" pageNumber="99">
General discussion on
<taxonomicName id="4C234D4FA764FF8707FCFC5DFD250CCA" authorityName="Reisinger" authorityYear="1924" box="[474,640,983,1010]" family="Typhloplanidae" genus="Protoplanella" higherTaxonomySource="CoL" kingdom="Animalia" order="Rhabdocoela" pageId="29" pageNumber="99" phylum="Platyhelminthes" rank="genus">
<emphasis id="B957EADEA764FF8707FCFC5DFD250CCA" bold="true" box="[474,640,983,1010]" italics="true" pageId="29" pageNumber="99">Protoplanella</emphasis>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8B9C36CCA764FF87069BFB8CFE350A9B" blockId="29.[188,1399,1030,1443]" pageId="29" pageNumber="99">
As mentioned by
<bibRefCitation id="EFB24B3DA764FF8707AFFB8CFD680B18" author="Van Steenkiste N. &amp; Tessens B. &amp; Krznaric K. &amp; Artois T." box="[393,717,1030,1056]" pageId="29" pageNumber="99" pagination="1 - 29" refId="ref16304" refString="Van Steenkiste N., Tessens B., Krznaric K. &amp; Artois T. 2011. Dalytyphloplanida (Platyhelminthes: Rhabdocoela) from Andalusia, Spain, with the description of four new species. Zootaxa 2791: 1 - 29. https: // doi. org / 10.11646 / zootaxa. 2791.1.1" type="journal article" year="2011">
Van Steenkiste
<emphasis id="B957EADEA764FF87041DFB8CFDD30B18" box="[571,630,1030,1056]" italics="true" pageId="29" pageNumber="99">et al.</emphasis>
(2011)
</bibRefCitation>
, the identification of
<taxonomicName id="4C234D4FA764FF8705E4FB8CFB630B18" authorityName="Reisinger" authorityYear="1924" box="[962,1222,1030,1056]" family="Typhloplanidae" genus="Protoplanella" higherTaxonomySource="CoL" kingdom="Animalia" order="Rhabdocoela" pageId="29" pageNumber="99" phylum="Platyhelminthes" rank="species" species="simplex">
<emphasis id="B957EADEA764FF8705E4FB8CFB630B18" box="[962,1222,1030,1056]" italics="true" pageId="29" pageNumber="99">Protoplanella simplex</emphasis>
</taxonomicName>
is challenging. New material found during several sampling trips showed a need for
<typeStatus id="5498886EA764FF870239FBA3FBF70B7B" box="[1055,1106,1065,1091]" pageId="29" pageNumber="99">type</typeStatus>
material to unravel the morphological differences between all known descriptions of
<taxonomicName id="4C234D4FA764FF870580FBC7FB850B5E" authorityName="Reisinger" authorityYear="1924" box="[934,1056,1100,1126]" family="Typhloplanidae" genus="Protoplanella" higherTaxonomySource="CoL" kingdom="Animalia" order="Rhabdocoela" pageId="29" pageNumber="99" phylum="Platyhelminthes" rank="species" species="simplex">
<emphasis id="B957EADEA764FF870580FBC7FB850B5E" box="[934,1056,1100,1126]" italics="true" pageId="29" pageNumber="99">P. simplex</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFB24B3DA764FF870214FBC6FB5F0B5E" author="Reisinger E." box="[1074,1274,1100,1126]" pageId="29" pageNumber="99" pagination="128 - 143" refId="ref15836" refString="Reisinger E. 1924. Die terricolen Rhabdocoelen Steiermarks. Zoologischer Anzeiger 59: 33 - 48, 72 - 86, 128 - 143." type="journal article" year="1924">Reisinger (1924)</bibRefCitation>
originally described animals with round testes; vasa deferentia that open separately from the lateral side of the copulatory organ; a vitellarium and ovovitellarium connected to each other over a broad anastomosis; and a seminal receptacle that opens into the proximal part of the female duct.
<bibRefCitation id="EFB24B3DA764FF87027DFB3CFAA50BE9" author="Luther A." box="[1115,1280,1206,1233]" pageId="29" pageNumber="99" pagination="1 - 163" refId="ref15760" refString="Luther A. 1963. Die Turbellarien Ostfennoskandiens IV. Neorhabdocoela 2. Typhloplanoida: Typhloplanidae, Solenopharyngidae und Carcharodopharyngidae. Fauna Fennica 16: 1 - 163." type="journal article" year="1963">Luther (1963)</bibRefCitation>
described a specimen with elongated testes; a bursa that could be a seminal receptacle near the male copulatory organ; and, judging from his illustration, most probably paired ovovitellaria. Also,
<bibRefCitation id="EFB24B3DA764FF8702AEFB77FEAF0A01" author="Van Steenkiste N. &amp; Tessens B. &amp; Krznaric K. &amp; Artois T." pageId="29" pageNumber="99" pagination="1 - 29" refId="ref16304" refString="Van Steenkiste N., Tessens B., Krznaric K. &amp; Artois T. 2011. Dalytyphloplanida (Platyhelminthes: Rhabdocoela) from Andalusia, Spain, with the description of four new species. Zootaxa 2791: 1 - 29. https: // doi. org / 10.11646 / zootaxa. 2791.1.1" type="journal article" year="2011">
Van Steenkiste
<emphasis id="B957EADEA764FF87031DFB77FAD30A2E" box="[1339,1398,1276,1302]" italics="true" pageId="29" pageNumber="99">et al.</emphasis>
(2011)
</bibRefCitation>
describe a sack-like protrusion at the genital atrium (which was not mentioned by
<bibRefCitation id="EFB24B3DA764FF870320FA95FF590A65" author="Reisinger E." pageId="29" pageNumber="99" pagination="128 - 143" refId="ref15836" refString="Reisinger E. 1924. Die terricolen Rhabdocoelen Steiermarks. Zoologischer Anzeiger 59: 33 - 48, 72 - 86, 128 - 143." type="journal article" year="1924">Reisinger 1924</bibRefCitation>
); a female bursa (which was called a seminal receptacle by
<bibRefCitation id="EFB24B3DA764FF8705E4FAC8FBD30A65" author="Reisinger E." box="[962,1142,1346,1373]" pageId="29" pageNumber="99" pagination="128 - 143" refId="ref15836" refString="Reisinger E. 1924. Die terricolen Rhabdocoelen Steiermarks. Zoologischer Anzeiger 59: 33 - 48, 72 - 86, 128 - 143." type="journal article" year="1924">Reisinger 1924</bibRefCitation>
) containing remnants of sperm; and their illustration shows the vasa deferentia uniting while entering the copulatory organ at the proximal side.
</paragraph>
<paragraph id="8B9C36CCA764FF87069BFA40FE800819" blockId="29.[189,1400,1482,1825]" pageId="29" pageNumber="99">
The descriptions of
<bibRefCitation id="EFB24B3DA764FF870794FA40FDFF0ADD" author="Luther A." box="[434,602,1482,1509]" pageId="29" pageNumber="99" pagination="1 - 163" refId="ref15760" refString="Luther A. 1963. Die Turbellarien Ostfennoskandiens IV. Neorhabdocoela 2. Typhloplanoida: Typhloplanidae, Solenopharyngidae und Carcharodopharyngidae. Fauna Fennica 16: 1 - 163." type="journal article" year="1963">Luther (1963)</bibRefCitation>
and
<bibRefCitation id="EFB24B3DA764FF8704BFFA41FC480ADC" author="Van Steenkiste N. &amp; Tessens B. &amp; Krznaric K. &amp; Artois T." box="[665,1005,1482,1509]" pageId="29" pageNumber="99" pagination="1 - 29" refId="ref16304" refString="Van Steenkiste N., Tessens B., Krznaric K. &amp; Artois T. 2011. Dalytyphloplanida (Platyhelminthes: Rhabdocoela) from Andalusia, Spain, with the description of four new species. Zootaxa 2791: 1 - 29. https: // doi. org / 10.11646 / zootaxa. 2791.1.1" type="journal article" year="2011">
Van Steenkiste
<emphasis id="B957EADEA764FF870573FA41FC300ADC" box="[853,917,1482,1508]" italics="true" pageId="29" pageNumber="99">et al.</emphasis>
(2011)
</bibRefCitation>
differ substantially from that of
<bibRefCitation id="EFB24B3DA764FF87069BFA67FE200930" author="Reisinger E." box="[189,389,1517,1544]" pageId="29" pageNumber="99" pagination="128 - 143" refId="ref15836" refString="Reisinger E. 1924. Die terricolen Rhabdocoelen Steiermarks. Zoologischer Anzeiger 59: 33 - 48, 72 - 86, 128 - 143." type="journal article" year="1924">Reisinger (1924)</bibRefCitation>
and, therefore, we consider them not to refer to representatives of
<taxonomicName id="4C234D4FA764FF870288FA64FA83093F" authorityName="Reisinger" authorityYear="1924" box="[1198,1318,1517,1543]" family="Typhloplanidae" genus="Protoplanella" higherTaxonomySource="CoL" kingdom="Animalia" order="Rhabdocoela" pageId="29" pageNumber="99" phylum="Platyhelminthes" rank="species" species="simplex">
<emphasis id="B957EADEA764FF870288FA64FA83093F" box="[1198,1318,1517,1543]" italics="true" pageId="29" pageNumber="99">P. simplex</emphasis>
</taxonomicName>
. Here, we consider these descriptions as referring to a new species:
<taxonomicName id="4C234D4FA764FF8705A4F99BFEA30976" authority="Houben, Proesmans &amp; Artois" authorityName="Houben, Proesmans &amp; Artois" authorityYear="2022" family="Typhloplanidae" genus="Protoplanella" higherTaxonomySource="CoL" kingdom="Animalia" order="Rhabdocoela" pageId="29" pageNumber="99" phylum="Platyhelminthes" rank="species" species="leiae" status="sp. nov.">
<emphasis id="B957EADEA764FF8705A4F99BFBCD0913" box="[898,1128,1553,1579]" italics="true" pageId="29" pageNumber="99">Protoplanella leiae</emphasis>
Houben, Proesmans &amp; Artois
</taxonomicName>
<taxonomicNameLabel id="A26457A5A764FF870729F9BEFEC90976" box="[271,364,1588,1614]" pageId="29" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
Moreover, the description of
<bibRefCitation id="EFB24B3DA764FF8704F2F9BEFCDF0977" author="Luther A." box="[724,890,1588,1615]" pageId="29" pageNumber="99" pagination="1 - 163" refId="ref15760" refString="Luther A. 1963. Die Turbellarien Ostfennoskandiens IV. Neorhabdocoela 2. Typhloplanoida: Typhloplanidae, Solenopharyngidae und Carcharodopharyngidae. Fauna Fennica 16: 1 - 163." type="journal article" year="1963">Luther (1963)</bibRefCitation>
indicates some, albeit smaller, differences with that of
<bibRefCitation id="EFB24B3DA764FF870770F9D2FD0D0949" author="Van Steenkiste N. &amp; Tessens B. &amp; Krznaric K. &amp; Artois T." box="[342,680,1623,1650]" pageId="29" pageNumber="99" pagination="1 - 29" refId="ref16304" refString="Van Steenkiste N., Tessens B., Krznaric K. &amp; Artois T. 2011. Dalytyphloplanida (Platyhelminthes: Rhabdocoela) from Andalusia, Spain, with the description of four new species. Zootaxa 2791: 1 - 29. https: // doi. org / 10.11646 / zootaxa. 2791.1.1" type="journal article" year="2011">
Van Steenkiste
<emphasis id="B957EADEA764FF870428F9D2FDE90949" box="[526,588,1623,1649]" italics="true" pageId="29" pageNumber="99">et al.</emphasis>
(2011)
</bibRefCitation>
. The illustration made by
<bibRefCitation id="EFB24B3DA764FF8705C1F9DDFB2B094A" author="Luther A." box="[999,1166,1623,1650]" pageId="29" pageNumber="99" pagination="1 - 163" refId="ref15760" refString="Luther A. 1963. Die Turbellarien Ostfennoskandiens IV. Neorhabdocoela 2. Typhloplanoida: Typhloplanidae, Solenopharyngidae und Carcharodopharyngidae. Fauna Fennica 16: 1 - 163." type="journal article" year="1963">Luther (1963)</bibRefCitation>
is not detailed and only made from live specimens. Since Luther’s drawings are in general more detailed, we assume this illustration was based on sub-ideal circumstances, which yielded a quickly made sketch. For now, we consider the specimens described by
<bibRefCitation id="EFB24B3DA764FF87045EF94BFCB809E4" author="Luther A." box="[632,797,1729,1756]" pageId="29" pageNumber="99" pagination="1 - 163" refId="ref15760" refString="Luther A. 1963. Die Turbellarien Ostfennoskandiens IV. Neorhabdocoela 2. Typhloplanoida: Typhloplanidae, Solenopharyngidae und Carcharodopharyngidae. Fauna Fennica 16: 1 - 163." type="journal article" year="1963">Luther (1963)</bibRefCitation>
and
<bibRefCitation id="EFB24B3DA764FF870571F94BFB3B09E3" author="Van Steenkiste N. &amp; Tessens B. &amp; Krznaric K. &amp; Artois T." box="[855,1182,1729,1755]" pageId="29" pageNumber="99" pagination="1 - 29" refId="ref16304" refString="Van Steenkiste N., Tessens B., Krznaric K. &amp; Artois T. 2011. Dalytyphloplanida (Platyhelminthes: Rhabdocoela) from Andalusia, Spain, with the description of four new species. Zootaxa 2791: 1 - 29. https: // doi. org / 10.11646 / zootaxa. 2791.1.1" type="journal article" year="2011">
Van Steenkiste
<emphasis id="B957EADEA764FF87022BF94BFBEF09E3" box="[1037,1098,1729,1755]" italics="true" pageId="29" pageNumber="99">et al.</emphasis>
(2011)
</bibRefCitation>
to be members of the same species since differences are small and, in the case of Luther, not based on thoroughly studied material.
</paragraph>
</subSubSection>
</treatment>
</document>
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