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<document id="D997160DD2BA2A32212B632C129D208E" ID-DOI="10.18590/euscorpius.2003.vol2003.iss11.1" ID-ISSN="1536-9307" ID-Zenodo-Dep="13237351" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="existingObjects" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1721353212353" checkinUser="felipe" docAuthor="Soleglad, Michael E. &amp; Fet, Victor" docDate="2003" docId="038A87D5D738F53DFC9F5F8DFAD452DD" docLanguage="en" docName="Euscorpius.11.1-175.pdf" docOrigin="Euscorpius 2003 (11)" docSource="http://dx.doi.org/10.18590/euscorpius.2003.vol2003.iss11.1" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Liochelinae Fet &amp; Bechly 2001" docType="treatment" docVersion="5" lastPageNumber="113" masterDocId="FFB3FFADD74CF549FFA15E77FFB0560F" masterDocTitle="High-level systematics and phylogeny of the extant scorpions (Scorpiones: Orthosterni)" masterLastPageNumber="175" masterPageNumber="1" pageNumber="113" updateTime="1722954143772" updateUser="ExternalLinkService">
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<mods:title id="223EB469199AD71189E0308A27A39FAE">High-level systematics and phylogeny of the extant scorpions (Scorpiones: Orthosterni)</mods:title>
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<mods:namePart id="641BED3D9AE3CBF0F6BBA3E7DC27CA14">Soleglad, Michael E.</mods:namePart>
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<mods:namePart id="3FC555D3EED616330FC3A4433D15E6C7">Fet, Victor</mods:namePart>
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<mods:title id="A3984F87FC2E528DBB60A6E90DCB1399">Euscorpius</mods:title>
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<mods:date id="4255BB74D3463889060E1B117BD6149D">2003</mods:date>
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<mods:number id="78155568255D51D95A51C2982D50C873">2003-12-26</mods:number>
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<mods:number id="B19741F90F2BB91F6A35797C1250BD71">2003</mods:number>
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<treatment id="038A87D5D738F53DFC9F5F8DFAD452DD" ID-DOI="http://doi.org/10.5281/zenodo.12785281" ID-Zenodo-Dep="12785281" LSID="urn:lsid:plazi:treatment:038A87D5D738F53DFC9F5F8DFAD452DD" httpUri="http://treatment.plazi.org/id/038A87D5D738F53DFC9F5F8DFAD452DD" lastPageNumber="113" pageId="116" pageNumber="113">
<subSubSection id="C3396548D738F53DFC9F5F8DFADB541E" box="[830,1387,506,529]" pageId="116" pageNumber="113" type="nomenclature">
<paragraph id="8B9C36C3D738F53DFC9F5F8DFADB541E" blockId="116.[830,1440,506,1234]" box="[830,1387,506,529]" pageId="116" pageNumber="113">
<heading id="D0D481AFD738F53DFC9F5F8DFADB541E" box="[830,1387,506,529]" fontSize="10" level="2" pageId="116" pageNumber="113" reason="5">
<emphasis id="B957EAD1D738F53DFC9F5F8DFADB541E" bold="true" box="[830,1387,506,529]" pageId="116" pageNumber="113">
Subfamily
<taxonomicName id="4C234D40D738F53DFC1B5F8DFAAD541E" authority="Fet &amp; Bechly, 2001" authorityName="Fet &amp; Bechly" authorityYear="2001" box="[954,1309,506,529]" class="Arachnida" family="Hormuridae" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="113" phylum="Arthropoda" rank="subFamily" subFamily="Liochelinae">Liochelinae Fet &amp; Bechly, 2001</taxonomicName>
(1879)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3396548D738F53DFCCF5C6FFBE455F4" pageId="116" pageNumber="113" type="type_taxon">
<paragraph id="8B9C36C3D738F53DFCCF5C6FFB285462" blockId="116.[830,1440,506,1234]" pageId="116" pageNumber="113">
<typeStatus id="54988861D738F53DFCCF5C6FFC135420" box="[878,931,536,559]" pageId="116" pageNumber="113">Type</typeStatus>
Genus.
<taxonomicName id="4C234D40D738F53DFBA05C6FFA905420" ID-CoL="5F2D" authority="Sundevall, 1833" authorityName="Sundevall" authorityYear="1833" box="[1025,1312,536,559]" class="Arachnida" family="Hormuridae" genus="Liocheles" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="B957EAD1D738F53DFBA05C6FFBD75420" box="[1025,1127,536,559]" italics="true" pageId="116" pageNumber="113">Liocheles</emphasis>
Sundevall, 1833
</taxonomicName>
;
<typeStatus id="54988861D738F53DFA8D5C6FFAEA5420" box="[1324,1370,536,559]" pageId="116" pageNumber="113">type</typeStatus>
genus
<taxonomicName id="4C234D40D738F53DFC9F5C40FBCE5441" ID-CoL="8WGQ3" authority="C. L. Koch, 1837" authorityName="C. L. Koch" authorityYear="1837" box="[830,1150,567,590]" class="Liliopsida" family="Poaceae" genus="Ischnurus" kingdom="Plantae" order="Poales" pageId="116" pageNumber="113" phylum="Tracheophyta" rank="genus">
<emphasis id="B957EAD1D738F53DFC9F5C40FC175441" box="[830,935,567,590]" italics="true" pageId="116" pageNumber="113">Ischnurus</emphasis>
C.L. Koch, 1837
</taxonomicName>
(=
<taxonomicName id="4C234D40D738F53DFB195C40FCC75462" ID-CoL="5F2D" authority="Sundevall, 1833" authorityName="Sundevall" authorityYear="1833" class="Arachnida" family="Hormuridae" genus="Liocheles" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="B957EAD1D738F53DFB195C40FAAE5441" box="[1208,1310,567,590]" italics="true" pageId="116" pageNumber="113">Liocheles</emphasis>
Sundevall, 1833
</taxonomicName>
); see Fet &amp; Bechly (2001).
</paragraph>
<paragraph id="8B9C36C3D738F53DFCCF5C03FC565484" blockId="116.[830,1440,506,1234]" box="[878,998,628,651]" pageId="116" pageNumber="113">
<emphasis id="B957EAD1D738F53DFCCF5C03FC6F5484" bold="true" box="[878,991,628,651]" italics="true" pageId="116" pageNumber="113">Synonyms</emphasis>
.
</paragraph>
<paragraph id="8B9C36C3D738F53DFC3F5CE4FAA854C6" blockId="116.[830,1440,506,1234]" pageId="116" pageNumber="113">
Hormurini Laurie, 1896;
<typeStatus id="54988861D738F53DFB155CE4FB5254A5" box="[1204,1250,659,682]" pageId="116" pageNumber="113">type</typeStatus>
genus
<taxonomicName id="4C234D40D738F53DFA935CE4FC6054C6" ID-CoL="62V9C" authority="Thorell, 1876" authorityName="Thorell" authorityYear="1876" class="Arachnida" family="Hormuridae" genus="Hormurus" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="B957EAD1D738F53DFA935CE4FA1054A5" box="[1330,1440,659,682]" italics="true" pageId="116" pageNumber="113">Hormurus</emphasis>
<bibRefCitation id="EFB24B32D738F53DFC9F5CC5FC6054C6" author="THORELL" box="[830,976,690,713]" pageId="116" pageNumber="113" pagination="1 - 15" refId="ref98099" refString="THORELL, T. 1876 a. On the classification of scorpions. Annals and Magazine of Natural History, 4 (17): 1 - 15." type="journal article" year="1876">Thorell, 1876</bibRefCitation>
</taxonomicName>
(=
<taxonomicName id="4C234D40D738F53DFC4F5CC5FABC54C6" ID-CoL="5F2D" authority="Sundevall, 1833" authorityName="Sundevall" authorityYear="1833" box="[1006,1292,690,713]" class="Arachnida" family="Hormuridae" genus="Liocheles" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="B957EAD1D738F53DFC4F5CC5FBE454C6" box="[1006,1108,690,713]" italics="true" pageId="116" pageNumber="113">Liocheles</emphasis>
Sundevall, 1833
</taxonomicName>
).
</paragraph>
<paragraph id="8B9C36C3D738F53DFC3F5CA7FBC05509" blockId="116.[830,1440,506,1234]" pageId="116" pageNumber="113">
Opisthacanthinae Kraepelin, 1905;
<typeStatus id="54988861D738F53DFA895CA7FAE654E8" box="[1320,1366,720,743]" pageId="116" pageNumber="113">type</typeStatus>
genus
<taxonomicName id="4C234D40D738F53DFC9F5C98FBC05509" ID-CoL="694T" authority="Peters, 1861" authorityName="Peters" authorityYear="1861" box="[830,1136,751,774]" class="Arachnida" family="Hormuridae" genus="Opisthacanthus" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="B957EAD1D738F53DFC9F5C98FC555509" box="[830,997,751,774]" italics="true" pageId="116" pageNumber="113">Opisthacanthus</emphasis>
<bibRefCitation id="EFB24B32D738F53DFC4C5C98FBC05509" author="PETERS" box="[1005,1136,751,774]" pageId="116" pageNumber="113" pagination="507 - 516" refId="ref95496" refString="PETERS, W. 1861. (Ueber eine neue Eintheilung der Skorpione und ueber die von ihm in Mossambique gesammelten Arten von Skorpionen). Monatsberichte der Koniglichen Preussischen Akademie der Wissenschaften zu Berlin, 1861: 507 - 516." type="journal article" year="1861">Peters, 1861</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8B9C36C3D738F53DFC3F5D79FBC0554C" blockId="116.[830,1440,506,1234]" pageId="116" pageNumber="113">
<taxonomicName id="4C234D40D738F53DFC3F5D79FB9B552A" authorityName="Soleglad &amp; Fet" authorityYear="2003" box="[926,1067,782,805]" class="Arachnida" family="Scorpionidae" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="129" phylum="Arthropoda" rank="subFamily" status="subfam. nov." subFamily="Hadogeninae">Hadogeninae</taxonomicName>
Lourenço, 1999;
<typeStatus id="54988861D738F53DFABD5D79FAFA552A" box="[1308,1354,782,805]" pageId="116" pageNumber="113">type</typeStatus>
genus
<taxonomicName id="4C234D40D738F53DFC9F5D5BFBDD554C" ID-CoL="4SBM" authority="Kraepelin, 1894" authorityName="Kraepelin" authorityYear="1894" box="[830,1133,812,835]" class="Arachnida" family="Hormuridae" genus="Hadogenes" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="B957EAD1D738F53DFC9F5D5BFC06554C" box="[830,950,812,835]" italics="true" pageId="116" pageNumber="113">Hadogenes</emphasis>
Kraepelin, 1894
</taxonomicName>
.
</paragraph>
<paragraph id="8B9C36C3D738F53DFCCF5D3CFBE4556D" blockId="116.[830,1440,506,1234]" box="[878,1108,843,866]" pageId="116" pageNumber="113">
<emphasis id="B957EAD1D738F53DFCCF5D3CFBE4556D" bold="true" box="[878,1108,843,866]" italics="true" pageId="116" pageNumber="113">Non-available name.</emphasis>
</paragraph>
<paragraph id="8B9C36C3D738F53DFC3F5D1DFAFB558E" blockId="116.[830,1440,506,1234]" box="[926,1355,874,897]" pageId="116" pageNumber="113">
<taxonomicName id="4C234D40D738F53DFC3F5D1DFBAA558E" box="[926,1050,874,897]" higherTaxonomySource="Manual Input" pageId="116" pageNumber="113" rank="subFamily" subFamily="Ischnurinae">Ischnurinae</taxonomicName>
<bibRefCitation id="EFB24B32D738F53DFB835D1DFB19558E" author="SIMON" box="[1058,1193,874,897]" pageId="116" pageNumber="113" pagination="79 - 115" refId="ref96627" refString="SIMON, E. 1879. 3 e Ordre. - Scorpiones. Pp. 79 - 115 in: Les Arachnides de France. VII. Contenant les Ordres des Chernetes, Scorpiones et Opiliones. Paris: Roret." type="book chapter" year="1879">Simon, 1879</bibRefCitation>
(ICZN, 2003).
</paragraph>
<paragraph id="8B9C36C3D738F53DFCCF5DFFFBE455F4" blockId="116.[830,1440,506,1234]" pageId="116" pageNumber="113">
<emphasis id="B957EAD1D738F53DFCCF5DFFFBB05590" bold="true" box="[878,1024,904,927]" italics="true" pageId="116" pageNumber="113">Composition.</emphasis>
This subfamily includes nine genera, predominantly from the Old World (only some species of
<taxonomicName id="4C234D40D738F53DFCC15DB1FBB755D2" ID-CoL="694T" authorityName="Peters" authorityYear="1861" box="[864,1031,966,989]" class="Arachnida" family="Hormuridae" genus="Opisthacanthus" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="B957EAD1D738F53DFCC15DB1FBB755D2" box="[864,1031,966,989]" italics="true" pageId="116" pageNumber="113">Opisthacanthus</emphasis>
</taxonomicName>
are found in the New World) (Fet, 2000c; Lourenço, 2001b).
</paragraph>
</subSubSection>
<subSubSection id="C3396548D738F53DFCCF5A74FC7B529A" pageId="116" pageNumber="113" type="distribution">
<paragraph id="8B9C36C3D738F53DFCCF5A74FAA75236" blockId="116.[830,1440,506,1234]" pageId="116" pageNumber="113">
<emphasis id="B957EAD1D738F53DFCCF5A74FC4A5215" bold="true" box="[878,1018,1027,1050]" italics="true" pageId="116" pageNumber="113">Distribution.</emphasis>
Asia, Africa,
<collectingCountry id="F3347653D738F53DFB635A74FA975215" box="[1218,1319,1027,1050]" name="Australia" pageId="116" pageNumber="113">Australia</collectingCountry>
, Oceania, North America, Central and South America.
</paragraph>
<paragraph id="8B9C36C3D738F53DFCCF5A37FC7B529A" blockId="116.[830,1440,506,1234]" pageId="116" pageNumber="113">
<emphasis id="B957EAD1D738F53DFCCF5A37FBF85258" bold="true" box="[878,1096,1088,1111]" italics="true" pageId="116" pageNumber="113">Taxonomic history</emphasis>
.
<bibRefCitation id="EFB24B32D738F53DFBC15A37FAA25258" author="PRENDINI" box="[1120,1298,1088,1111]" pageId="116" pageNumber="113" pagination="1 - 78" refId="ref95824" refString="PRENDINI, L. 2000. Phylogeny and classification of the superfamily Scorpionoidea Latreille 1802 (Chelicerata, Scorpiones): An exemplar approach. Cladistics, 16: 1 - 78. (March 2000)." type="journal article" year="2000">Prendini (2000)</bibRefCitation>
provided a detailed phylogenetic analysis of the liocheline taxa (as
<taxonomicName id="4C234D40D738F53DFC9F5A09FC70529A" authorityName="Simon" authorityYear="1879" box="[830,960,1150,1173]" class="Arachnida" family="Ischnuridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="113" phylum="Arthropoda" rank="family">Ischnuridae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3396548D738F53DFCCF5AEBFAD452DD" pageId="116" pageNumber="113" type="diagnosis">
<paragraph id="8B9C36C3D738F53DFCCF5AEBFAD452DD" blockId="116.[830,1440,506,1234]" pageId="116" pageNumber="113">
<emphasis id="B957EAD1D738F53DFCCF5AEBFC6C52BC" bold="true" box="[878,988,1180,1203]" italics="true" pageId="116" pageNumber="113">Diagnosis</emphasis>
. See
<bibRefCitation id="EFB24B32D738F53DFB835AEBFB7F52BC" author="PRENDINI" box="[1058,1231,1180,1203]" pageId="116" pageNumber="113" pagination="1 - 78" refId="ref95824" refString="PRENDINI, L. 2000. Phylogeny and classification of the superfamily Scorpionoidea Latreille 1802 (Chelicerata, Scorpiones): An exemplar approach. Cladistics, 16: 1 - 78. (March 2000)." type="journal article" year="2000">Prendini (2000)</bibRefCitation>
for details on the diagnosis of this subfamily (as family
<taxonomicName id="4C234D40D738F53DFB765ACCFAE952DD" authorityName="Simon" authorityYear="1879" box="[1239,1369,1211,1234]" class="Arachnida" family="Ischnuridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="116" pageNumber="113" phylum="Arthropoda" rank="family">Ischnuridae</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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