340 lines
29 KiB
XML
340 lines
29 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.6210813" ID-GBIF-Dataset="7485373c-0219-4ed7-bb5c-ac664acf6775" ID-GBIF-Taxon="114923704" ID-Zenodo-Dep="6210813" LSID="urn:lsid:zoobank.org:act:E8ACB7E2-9473-4521-B654-F8A923F10AF0" checkinTime="1427905504904" checkinUser="donat" docAuthor="Alberto Ballerio & Munetoshi Maruyama" docDate="2010" docId="038951095C1A3D0466B6FF63ABB6FD92" docLanguage="en" docName="ZK_article_2140.pdf" docOrigin="Zookeys 34" docTitle="Pterorthochaetes tsurui Ballerio & Maruyama, 2010, sp. n." docType="treatment" docUuid="E8ACB7E2-9473-4521-B654-F8A923F10AF0" docUuidSource="ZooBank" docVersion="9" lastPageId="24" lastPageNumber="101" masterDocId="FFB029715C083D1C663AFFC4A802FFD6" masterDocTitle="The Ceratocanthinae of Ulu Gombak: high species richness at a single site, with descriptions of three new species and an annotated checklist of the Ceratocanthinae of Western Malaysia and Singapore (Coleoptera, Scarabaeoidea, Hybosoridae)" masterLastPageNumber="104" masterPageNumber="77" pageId="18" pageNumber="95" updateTime="1645477292090" updateUser="ExternalLinkService">
|
|
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo>
|
|
<mods:title>The Ceratocanthinae of Ulu Gombak: high species richness at a single site, with descriptions of three new species and an annotated checklist of the Ceratocanthinae of Western Malaysia and Singapore (Coleoptera, Scarabaeoidea, Hybosoridae)</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Alberto Ballerio</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Munetoshi Maruyama</mods:namePart>
|
|
</mods:name>
|
|
<mods:typeOfResource>text</mods:typeOfResource>
|
|
<mods:relatedItem type="host">
|
|
<mods:titleInfo>
|
|
<mods:title>Zookeys</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part>
|
|
<mods:date>2010</mods:date>
|
|
<mods:detail type="volume">
|
|
<mods:number>34</mods:number>
|
|
</mods:detail>
|
|
<mods:extent unit="page">
|
|
<mods:start>77</mods:start>
|
|
<mods:end>104</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:classification>journal article</mods:classification>
|
|
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.34.268</mods:identifier>
|
|
<mods:identifier type="ZooBank">553C91CE-FEA1-4867-929B-2865647E4B2E</mods:identifier>
|
|
</mods:mods>
|
|
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6210813" ID-GBIF-Taxon="114923704" ID-Zenodo-Dep="6210813" LSID="urn:lsid:zoobank.org:act:E8ACB7E2-9473-4521-B654-F8A923F10AF0" httpUri="http://treatment.plazi.org/id/038951095C1A3D0466B6FF63ABB6FD92" lastPageId="24" lastPageNumber="101" pageId="18" pageNumber="95">
|
|
<subSubSection box="[140,488,167,193]" pageId="18" pageNumber="95" type="nomenclature">
|
|
<paragraph box="[140,488,167,193]" pageId="18" pageNumber="95">
|
|
<heading bold="true" box="[140,488,167,193]" fontSize="11" level="1" pageId="18" pageNumber="95" reason="6">
|
|
<emphasis bold="true" box="[140,488,167,193]" pageId="18" pageNumber="95">
|
|
<emphasis bold="true" box="[140,416,167,193]" italics="true" pageId="18" pageNumber="95">
|
|
<taxonomicName LSID-ZBK="urn:lsid:zoobank.org:act:E8ACB7E2-9473-4521-B654-F8A923F10AF0 " box="[140,416,167,193]" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="95" phylum="Arthropoda" rank="species" species="tsurui" status="sp. n.">Pterorthochaetes tsurui</taxonomicName>
|
|
</emphasis>
|
|
<taxonomicNameLabel box="[423,488,167,193]" pageId="18" pageNumber="95" rank="species">sp. n.</taxonomicNameLabel>
|
|
</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<paragraph pageId="18" pageNumber="95">
|
|
<subSubSection box="[140,959,201,228]" pageId="18" pageNumber="95" type="description">urn:lsid:zoobank.org:act:E8ACB7E2-9473-4521-B654-F8A923F10AF0</subSubSection>
|
|
<subSubSection box="[140,281,237,263]" pageId="18" pageNumber="95" type="description">
|
|
(
|
|
<figureCitation box="[148,214,237,263]" captionStart="Figure 3" captionStartId="18.[140,211,1590,1612]" captionTargetBox="[146,1100,502,1555]" captionTargetId="figure@18.[140,1108,487,1570]" captionTargetPageId="18" captionText="Figure 3. Ceratocanthinae of Ulu Gombak. A Pterorthochaetes insularis habitus dorsal B Pterorthochaetes sp. habitus dorsal C Pterorthochaetes tsurui sp. n. habitus dorsal D idem habitus lateral E Cyphopisthes sp. habitus dorsal." pageId="18" pageNumber="95">Fig. 3</figureCitation>
|
|
C, D)
|
|
</subSubSection>
|
|
</paragraph>
|
|
<subSubSection lastPageId="19" lastPageNumber="96" pageId="18" pageNumber="95" type="materials_examined">
|
|
<paragraph box="[140,990,307,334]" pageId="18" pageNumber="95">
|
|
<emphasis bold="true" box="[140,301,307,334]" pageId="18" pageNumber="95">Type locality:</emphasis>
|
|
West Malaysia: Ulu Gombak (Selangor), 3°19
|
|
<emphasis box="[827,833,308,334]" italics="true" pageId="18" pageNumber="95">'</emphasis>
|
|
N 101°45
|
|
<emphasis box="[952,959,308,334]" italics="true" pageId="18" pageNumber="95">'</emphasis>
|
|
E.
|
|
</paragraph>
|
|
<paragraph lastPageId="19" lastPageNumber="96" pageId="18" pageNumber="95">
|
|
<emphasis bold="true" box="[185,417,343,369]" pageId="18" pageNumber="95">Material examined.</emphasis>
|
|
<emphasis box="[426,521,342,369]" italics="true" pageId="18" pageNumber="95">Holotype</emphasis>
|
|
, male: West Malaysia, Selangor, Ulu Gombak, 250 mt., 29.IV.2007, Leg. M. Maruyama /
|
|
<taxonomicName box="[600,848,377,404]" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="95" phylum="Arthropoda" rank="species" species="tsurui" status="sp. n.">
|
|
<emphasis box="[600,848,377,404]" italics="true" pageId="18" pageNumber="95">Pterorthochaetes tsurui</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel box="[858,922,378,404]" pageId="18" pageNumber="95" rank="species">sp. n.</taxonomicNameLabel>
|
|
det. Ballerio & Maruyama 2009 Holotypus (KUM). [distended specimen glued on a card, dissected, genitalia mounted in DMHF resin on a separate card, same pin].
|
|
<emphasis box="[887,987,1294,1319]" italics="true" pageId="19" pageNumber="96">Paratypes</emphasis>
|
|
[one male and four females dissected]: 15.IV.2007, M. Maruyama (1 male); 26.IV.2007, M. Maruyama (1 female and 1 male); 28. IV. 2007, M. Maruyama (1 female); 29. IV. 2007, M. Maruyama (1 female); 30. IV. 2007, M. Maruyama (1 male and female); 3. V. 2007, M. Maruyama (1 female); 4. V. 2007, M. Maruyama (1 female). All from termite nest(s). (2 ABCB, 7 KUM)
|
|
</paragraph>
|
|
</subSubSection>
|
|
<caption pageId="18" pageNumber="95" targetBox="[146,1100,502,1555]" targetPageId="18">
|
|
<paragraph pageId="18" pageNumber="95">
|
|
<emphasis bold="true" box="[140,238,1590,1612]" pageId="18" pageNumber="95">Figure 3.</emphasis>
|
|
Ceratocanthinae of Ulu Gombak.
|
|
<emphasis bold="true" box="[565,584,1590,1612]" pageId="18" pageNumber="95">A</emphasis>
|
|
<emphasis box="[590,814,1590,1612]" italics="true" pageId="18" pageNumber="95">
|
|
<taxonomicName box="[590,814,1590,1612]" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="95" phylum="Arthropoda" rank="species" species="insularis">Pterorthochaetes insularis</taxonomicName>
|
|
</emphasis>
|
|
habitus dorsal
|
|
<emphasis bold="true" box="[956,973,1590,1612]" pageId="18" pageNumber="95">B</emphasis>
|
|
<emphasis italics="true" pageId="18" pageNumber="95">
|
|
<taxonomicName class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="95" phylum="Arthropoda" rank="genus">Pterorthochaetes</taxonomicName>
|
|
sp.
|
|
</emphasis>
|
|
habitus dorsal
|
|
<emphasis bold="true" box="[326,345,1622,1644]" pageId="18" pageNumber="95">C</emphasis>
|
|
<emphasis box="[349,548,1622,1644]" italics="true" pageId="18" pageNumber="95">
|
|
<taxonomicName box="[349,548,1622,1644]" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="95" phylum="Arthropoda" rank="species" species="tsurui" status="sp. n.">Pterorthochaetes tsurui</taxonomicName>
|
|
</emphasis>
|
|
<taxonomicNameLabel box="[553,602,1623,1644]" pageId="18" pageNumber="95" rank="species">sp. n.</taxonomicNameLabel>
|
|
habitus dorsal
|
|
<emphasis bold="true" box="[739,758,1622,1644]" pageId="18" pageNumber="95">D</emphasis>
|
|
idem habitus lateral
|
|
<emphasis bold="true" box="[948,963,1622,1644]" pageId="18" pageNumber="95">E</emphasis>
|
|
<emphasis box="[967,1109,1622,1644]" italics="true" pageId="18" pageNumber="95">
|
|
<taxonomicName box="[967,1086,1622,1644]" class="Insecta" family="Hybosoridae" genus="Cyphopisthes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="95" phylum="Arthropoda" rank="genus">Cyphopisthes</taxonomicName>
|
|
sp.
|
|
</emphasis>
|
|
habitus dorsal.
|
|
</paragraph>
|
|
</caption>
|
|
<caption pageId="19" pageNumber="96" targetBox="[156,1100,180,1155]" targetPageId="19">
|
|
<paragraph pageId="19" pageNumber="96">
|
|
<emphasis bold="true" box="[140,237,1182,1204]" pageId="19" pageNumber="96">Figure 4.</emphasis>
|
|
Ceratocanthinae of Ulu Gombak.
|
|
<emphasis bold="true" box="[560,579,1182,1204]" pageId="19" pageNumber="96">A</emphasis>
|
|
<emphasis box="[585,765,1182,1204]" italics="true" pageId="19" pageNumber="96">
|
|
<taxonomicName box="[585,720,1182,1204]" class="Insecta" family="Hybosoridae" genus="Eusphaeropeltis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="96" phylum="Arthropoda" rank="genus">Eusphaeropeltis</taxonomicName>
|
|
sp. a
|
|
</emphasis>
|
|
habitus dorsal
|
|
<emphasis bold="true" box="[905,921,1182,1204]" pageId="19" pageNumber="96">B</emphasis>
|
|
<emphasis box="[927,1107,1182,1204]" italics="true" pageId="19" pageNumber="96">
|
|
<taxonomicName box="[927,1062,1182,1204]" class="Insecta" family="Hybosoridae" genus="Eusphaeropeltis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="96" phylum="Arthropoda" rank="genus">Eusphaeropeltis</taxonomicName>
|
|
sp. b
|
|
</emphasis>
|
|
habitus dorsal
|
|
<emphasis bold="true" box="[275,293,1214,1236]" pageId="19" pageNumber="96">C</emphasis>
|
|
<emphasis box="[299,477,1214,1236]" italics="true" pageId="19" pageNumber="96">
|
|
<taxonomicName box="[299,434,1214,1236]" class="Insecta" family="Hybosoridae" genus="Eusphaeropeltis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="96" phylum="Arthropoda" rank="genus">Eusphaeropeltis</taxonomicName>
|
|
sp. c
|
|
</emphasis>
|
|
habitus dorsal
|
|
<emphasis bold="true" box="[618,637,1214,1236]" pageId="19" pageNumber="96">D</emphasis>
|
|
<emphasis box="[643,824,1214,1236]" italics="true" pageId="19" pageNumber="96">
|
|
<taxonomicName box="[643,824,1214,1236]" class="Insecta" family="Hybosoridae" genus="Ebbrittoniella" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="96" phylum="Arthropoda" rank="species" species="ignita">Ebbrittoniella ignita</taxonomicName>
|
|
</emphasis>
|
|
habitus dorsal.
|
|
</paragraph>
|
|
</caption>
|
|
<subSubSection lastPageId="23" lastPageNumber="100" pageId="19" pageNumber="96" type="description">
|
|
<paragraph pageId="19" pageNumber="96">
|
|
<emphasis bold="true" box="[185,335,1504,1530]" pageId="19" pageNumber="96">Description.</emphasis>
|
|
HL = 0.69 mm; HW = 1.44 mm; PL = 1.38 mm; PW = 2.33 mm; EL = 2.36 mm; EW = 2.27 mm.
|
|
</paragraph>
|
|
<paragraph pageId="19" pageNumber="96">
|
|
Medium sized
|
|
<emphasis box="[354,528,1574,1601]" italics="true" pageId="19" pageNumber="96">
|
|
<taxonomicName box="[354,528,1574,1601]" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="96" phylum="Arthropoda" rank="genus">Pterorthochaetes</taxonomicName>
|
|
</emphasis>
|
|
, surface shiny, setose; “rolling up” coaptations perfect; volant.
|
|
</paragraph>
|
|
<paragraph box="[185,988,1645,1671]" pageId="19" pageNumber="96">Dorsum brown, setation yellowish, sternum reddish, antennae reddish.</paragraph>
|
|
<caption pageId="20" pageNumber="97" targetBox="[140,1107,165,1070]" targetPageId="20">
|
|
<paragraph pageId="20" pageNumber="97">
|
|
<emphasis bold="true" box="[140,237,1093,1115]" pageId="20" pageNumber="97">Figure 5.</emphasis>
|
|
SEM photographs of:
|
|
<emphasis bold="true" box="[444,463,1093,1115]" pageId="20" pageNumber="97">A</emphasis>
|
|
<emphasis box="[468,644,1093,1115]" italics="true" pageId="20" pageNumber="97">
|
|
<taxonomicName box="[468,573,1093,1115]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="97" phylum="Arthropoda" rank="genus">Madrasostes</taxonomicName>
|
|
clypeale
|
|
</emphasis>
|
|
: male pronotum
|
|
<emphasis bold="true" box="[804,820,1093,1115]" pageId="20" pageNumber="97">B</emphasis>
|
|
detail of male pronotum puncture
|
|
<emphasis bold="true" box="[184,203,1125,1147]" pageId="20" pageNumber="97">C</emphasis>
|
|
female pronotum
|
|
<emphasis bold="true" box="[380,400,1125,1147]" pageId="20" pageNumber="97">D</emphasis>
|
|
detail of female pronotum punctures
|
|
<emphasis bold="true" box="[761,776,1125,1147]" pageId="20" pageNumber="97">E</emphasis>
|
|
and
|
|
<emphasis bold="true" box="[825,840,1125,1147]" pageId="20" pageNumber="97">F</emphasis>
|
|
<taxonomicName box="[847,1048,1125,1147]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="97" phylum="Arthropoda" rank="species" species="mirificum" status="sp.n.">
|
|
<emphasis box="[847,1048,1125,1147]" italics="true" pageId="20" pageNumber="97">Madrasostes mirificum</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel box="[1056,1108,1126,1147]" pageId="20" pageNumber="97">sp.n.</taxonomicNameLabel>
|
|
mouthparts: labium
|
|
<emphasis bold="true" box="[331,350,1157,1179]" pageId="20" pageNumber="97">G</emphasis>
|
|
epipharynx
|
|
<emphasis bold="true" box="[466,486,1157,1179]" pageId="20" pageNumber="97">H</emphasis>
|
|
mandibles
|
|
<emphasis bold="true" box="[593,601,1157,1179]" pageId="20" pageNumber="97">I</emphasis>
|
|
detail of galear brush
|
|
<emphasis bold="true" box="[806,814,1157,1179]" pageId="20" pageNumber="97">J</emphasis>
|
|
maxilla.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="20" pageNumber="97">
|
|
<emphasis box="[185,251,1222,1249]" italics="true" pageId="20" pageNumber="97">Head:</emphasis>
|
|
subrectangular, wider than long, fore margin irregularly sinuated, serrated, tip acute, interocular distance about 9 times the maximum width of dorsal ocular area, dorsal ocular area large, sculpure distally made of very coarse and deep wrinkles and proximally of impressed small dense horseshoe-shaped punctures, each one having a pore in the middle bearing an erect simple seta.
|
|
</paragraph>
|
|
<paragraph pageId="20" pageNumber="97">
|
|
<emphasis box="[185,301,1399,1425]" italics="true" pageId="20" pageNumber="97">Pronotum:</emphasis>
|
|
wider than long (W/L ratio= 1.68), fore angles normally shaped, elytral lateral margins fringed with a row of long simple setae, spaced out by an interval about half of their length, the whole pronotal surface covered by medium sized round, shallow, ocellate punctures, each one having a pore in the middle bearing a long simple erect seta; puncturation relatively dense, the distance between punctures being inferior to their diameter, denser on disc and at sides, only at sides the ocellate punctures are replaced by horseshoe-shaped punctures with openings outwards. Pronotal pubescence relatively long, approximately as long as marginal setae.
|
|
</paragraph>
|
|
<caption pageId="21" pageNumber="98" targetBox="[140,1107,165,1186]" targetPageId="21">
|
|
<paragraph pageId="21" pageNumber="98">
|
|
<emphasis bold="true" box="[140,238,1206,1228]" pageId="21" pageNumber="98">Figure 6.</emphasis>
|
|
Photos of alive specimens taken in habitat at Ulu Gombak (photos © M. Maruyama, 2008).
|
|
<emphasis bold="true" box="[140,159,1238,1260]" pageId="21" pageNumber="98">A</emphasis>
|
|
<emphasis box="[165,374,1238,1260]" italics="true" pageId="21" pageNumber="98">
|
|
<taxonomicName box="[165,270,1238,1260]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="98" phylum="Arthropoda" rank="genus">Madrasostes</taxonomicName>
|
|
variolosum
|
|
</emphasis>
|
|
<emphasis bold="true" box="[381,397,1238,1260]" pageId="21" pageNumber="98">B</emphasis>
|
|
<taxonomicName box="[404,572,1238,1260]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="98" phylum="Arthropoda" rank="species" species="agostii">
|
|
<emphasis box="[404,572,1238,1260]" italics="true" pageId="21" pageNumber="98">Madrasostes agostii</emphasis>
|
|
</taxonomicName>
|
|
<emphasis bold="true" box="[579,597,1238,1260]" pageId="21" pageNumber="98">C</emphasis>
|
|
<emphasis box="[604,783,1238,1260]" italics="true" pageId="21" pageNumber="98">
|
|
<taxonomicName box="[604,709,1238,1260]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="98" phylum="Arthropoda" rank="genus">Madrasostes</taxonomicName>
|
|
clypeale
|
|
</emphasis>
|
|
male
|
|
<emphasis bold="true" box="[840,860,1238,1260]" pageId="21" pageNumber="98">D</emphasis>
|
|
idem female
|
|
<emphasis bold="true" box="[987,1003,1238,1260]" pageId="21" pageNumber="98">E</emphasis>
|
|
<emphasis italics="true" pageId="21" pageNumber="98">
|
|
<taxonomicName class="Insecta" family="Hybosoridae" genus="Eusphaeropeltis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="98" phylum="Arthropoda" rank="genus">Eusphaeropeltis</taxonomicName>
|
|
sp. b
|
|
</emphasis>
|
|
<emphasis bold="true" box="[237,251,1270,1292]" pageId="21" pageNumber="98">F</emphasis>
|
|
<emphasis box="[257,435,1270,1292]" italics="true" pageId="21" pageNumber="98">
|
|
<taxonomicName box="[257,392,1270,1292]" class="Insecta" family="Hybosoridae" genus="Eusphaeropeltis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="98" phylum="Arthropoda" rank="genus">Eusphaeropeltis</taxonomicName>
|
|
sp. c
|
|
</emphasis>
|
|
.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph box="[185,830,1328,1355]" pageId="21" pageNumber="98">
|
|
<emphasis box="[185,300,1328,1355]" italics="true" pageId="21" pageNumber="98">Scutellum:</emphasis>
|
|
punctures horseshoe-shaped, thick and coarse.
|
|
</paragraph>
|
|
<paragraph pageId="21" pageNumber="98">Elytra: shape oval, longer than wide (W/L ratio= 0.96); elytral surface covered by medium sized shallow horseshoe-shaped punctures with opening directed backwards, spaced out by an interval inferior to their diameter. Each horseshoe-shaped puncture containing a pore bearing a long erect simple seta.</paragraph>
|
|
<paragraph pageId="21" pageNumber="98">
|
|
<emphasis box="[185,357,1504,1531]" italics="true" pageId="21" pageNumber="98">Male genitalia:</emphasis>
|
|
parameres about as long as basal piece, slightly asymmetrical (
|
|
<figureCitation box="[149,211,1539,1566]" captionStart="Figure 7" captionStartId="22.[140,211,1590,1612]" captionTargetBox="[231,1090,199,1518]" captionTargetId="figure@22.[140,1108,162,1568]" captionTargetPageId="22" captionText="Figure 7. Pterorthochaetes tsurui sp. n.: A parameres in lateral view (scale bar: 0, 1 mm) B parameres in dorsal view (scale bar: 0, 1 mm) C female bursal sclerites (scale bar: 0, 1 mm) D, sclerites of the internal sac of aedeagus (scale bar: 0, 1 mm) E male genital segment (scale bar: 0, 1 mm)." pageId="21" pageNumber="98">Fig.7</figureCitation>
|
|
A, B), apex distinctly bent, internal sac armed with longitudinal subrectangular sclerotization about as long as the basal piece with three short distal sclerites (
|
|
<figureCitation box="[1003,1068,1574,1601]" captionStart="Figure 7" captionStartId="22.[140,211,1590,1612]" captionTargetBox="[231,1090,199,1518]" captionTargetId="figure@22.[140,1108,162,1568]" captionTargetPageId="22" captionText="Figure 7. Pterorthochaetes tsurui sp. n.: A parameres in lateral view (scale bar: 0, 1 mm) B parameres in dorsal view (scale bar: 0, 1 mm) C female bursal sclerites (scale bar: 0, 1 mm) D, sclerites of the internal sac of aedeagus (scale bar: 0, 1 mm) E male genital segment (scale bar: 0, 1 mm)." pageId="21" pageNumber="98">Fig. 7</figureCitation>
|
|
D), genital segment with manubrium long but strongly bent/twisted, so that in dorsal view it appears very short (
|
|
<figureCitation box="[383,449,1645,1671]" captionStart="Figure 7" captionStartId="22.[140,211,1590,1612]" captionTargetBox="[231,1090,199,1518]" captionTargetId="figure@22.[140,1108,162,1568]" captionTargetPageId="22" captionText="Figure 7. Pterorthochaetes tsurui sp. n.: A parameres in lateral view (scale bar: 0, 1 mm) B parameres in dorsal view (scale bar: 0, 1 mm) C female bursal sclerites (scale bar: 0, 1 mm) D, sclerites of the internal sac of aedeagus (scale bar: 0, 1 mm) E male genital segment (scale bar: 0, 1 mm)." pageId="21" pageNumber="98">Fig. 7</figureCitation>
|
|
E).
|
|
</paragraph>
|
|
<caption pageId="22" pageNumber="99" targetBox="[231,1090,199,1518]" targetPageId="22">
|
|
<paragraph pageId="22" pageNumber="99">
|
|
<emphasis bold="true" box="[140,239,1590,1612]" pageId="22" pageNumber="99">Figure 7.</emphasis>
|
|
<taxonomicName box="[245,446,1590,1612]" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="99" phylum="Arthropoda" rank="species" species="tsurui" status="sp. n.">
|
|
<emphasis box="[245,446,1590,1612]" italics="true" pageId="22" pageNumber="99">Pterorthochaetes tsurui</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel box="[453,506,1591,1612]" pageId="22" pageNumber="99" rank="species">sp. n.</taxonomicNameLabel>
|
|
:
|
|
<emphasis bold="true" box="[516,535,1590,1612]" pageId="22" pageNumber="99">A</emphasis>
|
|
parameres in lateral view (scale bar: 0,1 mm)
|
|
<emphasis bold="true" box="[966,982,1590,1612]" pageId="22" pageNumber="99">B</emphasis>
|
|
parameres in dorsal view (scale bar: 0,1 mm)
|
|
<emphasis bold="true" box="[429,448,1622,1644]" pageId="22" pageNumber="99">C</emphasis>
|
|
female bursal sclerites (scale bar: 0,1 mm)
|
|
<emphasis bold="true" box="[838,857,1622,1644]" pageId="22" pageNumber="99">D</emphasis>
|
|
, sclerites of the internal sac of aedeagus (scale bar: 0,1 mm)
|
|
<emphasis bold="true" box="[437,452,1654,1676]" pageId="22" pageNumber="99">E</emphasis>
|
|
male genital segment (scale bar: 0,1 mm).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph box="[185,1043,166,193]" pageId="23" pageNumber="100">
|
|
<emphasis box="[185,370,166,193]" italics="true" pageId="23" pageNumber="100">Female genitalia:</emphasis>
|
|
bursal sclerites weakly sclerotized and shaped as in
|
|
<figureCitation box="[951,1017,166,193]" captionStart="Figure 7" captionStartId="22.[140,211,1590,1612]" captionTargetBox="[231,1090,199,1518]" captionTargetId="figure@22.[140,1108,162,1568]" captionTargetPageId="22" captionText="Figure 7. Pterorthochaetes tsurui sp. n.: A parameres in lateral view (scale bar: 0, 1 mm) B parameres in dorsal view (scale bar: 0, 1 mm) C female bursal sclerites (scale bar: 0, 1 mm) D, sclerites of the internal sac of aedeagus (scale bar: 0, 1 mm) E male genital segment (scale bar: 0, 1 mm)." pageId="23" pageNumber="100">Fig. 7</figureCitation>
|
|
C.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection lastPageId="24" lastPageNumber="101" pageId="23" pageNumber="100" type="diagnosis">
|
|
<paragraph lastPageId="24" lastPageNumber="101" pageId="23" pageNumber="100">
|
|
<emphasis bold="true" box="[185,356,202,228]" pageId="23" pageNumber="100">Identification.</emphasis>
|
|
This species is characterized by the following combination of characters: a) pronotal puncturation made of ocellate, shallow, medium sized punctures, b) lateral margins of pronotum fringed by long, simple setae, setae relatively spaced out, c) elytral puncturation made of medium sized horseshoe-shaped punctures with opening backwards and bearing a long erect seta, d) shape of parameres (about as long as basal piece and with bent apex, e) shape of genital segment of males, with manubrium strongly twisted, f) shape of female bursal sclerites and, g) head subrectangular, with fore margin somewhat sinuated in a way very unusual for
|
|
<emphasis box="[790,963,237,263]" italics="true" pageId="24" pageNumber="101">
|
|
<taxonomicName box="[790,963,237,263]" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="101" phylum="Arthropoda" rank="genus">Pterorthochaetes</taxonomicName>
|
|
</emphasis>
|
|
. Perhaps the species closest to it could be
|
|
<emphasis box="[461,763,272,299]" italics="true" pageId="24" pageNumber="101">
|
|
<taxonomicName box="[461,763,272,299]" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="101" phylum="Arthropoda" rank="species" species="brevisetosus">Pterorthochaetes brevisetosus</taxonomicName>
|
|
</emphasis>
|
|
, which, however, has pronotal and elytral puncturation much less dense and shorter setae.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<caption pageId="23" pageNumber="100" targetBox="[145,1078,415,1513]" targetPageId="23">
|
|
<paragraph pageId="23" pageNumber="100">
|
|
<emphasis bold="true" box="[140,240,1558,1580]" pageId="23" pageNumber="100">Figure 8.</emphasis>
|
|
<emphasis box="[247,448,1558,1580]" italics="true" pageId="23" pageNumber="100">
|
|
<taxonomicName box="[247,448,1558,1580]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="100" phylum="Arthropoda" rank="species" species="mirificum" status="sp. n.">Madrasostes mirificum</taxonomicName>
|
|
</emphasis>
|
|
<taxonomicNameLabel box="[456,510,1559,1580]" pageId="23" pageNumber="100" rank="species">sp. n.</taxonomicNameLabel>
|
|
:
|
|
<emphasis bold="true" box="[521,540,1558,1580]" pageId="23" pageNumber="100">A</emphasis>
|
|
male genital segment (scale bar: 0,2 mm)
|
|
<emphasis bold="true" box="[944,961,1558,1580]" pageId="23" pageNumber="100">B</emphasis>
|
|
aedeagus (scale bar: 0,1 mm)
|
|
<emphasis bold="true" box="[270,289,1590,1612]" pageId="23" pageNumber="100">C</emphasis>
|
|
parameres in dorsal view (scale bar: 0,0 5 mm)
|
|
<emphasis bold="true" box="[729,749,1590,1612]" pageId="23" pageNumber="100">D</emphasis>
|
|
<taxonomicName box="[755,925,1590,1612]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="100" phylum="Arthropoda" rank="species" species="hasimi" status="sp. n.">
|
|
<emphasis box="[755,925,1590,1612]" italics="true" pageId="23" pageNumber="100">Madrasostes hasimi</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel box="[932,985,1591,1612]" pageId="23" pageNumber="100" rank="species">sp. n.</taxonomicNameLabel>
|
|
: male genital segment (scale bar: 0,1 mm)
|
|
<emphasis bold="true" box="[409,425,1622,1644]" pageId="23" pageNumber="100">E</emphasis>
|
|
parameres (scale bar: 0,1 mm)
|
|
<emphasis bold="true" box="[717,731,1622,1644]" pageId="23" pageNumber="100">F</emphasis>
|
|
sclerite of internal sac of aedeagus (scale bar: 0,1 mm)
|
|
<emphasis bold="true" box="[269,288,1654,1676]" pageId="23" pageNumber="100">G</emphasis>
|
|
<taxonomicName box="[294,494,1654,1676]" class="Insecta" family="Hybosoridae" genus="Madrasostes" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="100" phylum="Arthropoda" rank="species" species="depressum">
|
|
<emphasis box="[294,494,1654,1676]" italics="true" pageId="23" pageNumber="100">Madrasostes depressum</emphasis>
|
|
</taxonomicName>
|
|
: sclerite of internal sac of aedeagus (scale bar: 0,1 mm).
|
|
</paragraph>
|
|
</caption>
|
|
<subSubSection pageId="24" pageNumber="101" type="discussion">
|
|
<paragraph pageId="24" pageNumber="101">
|
|
<emphasis bold="true" box="[185,315,343,369]" pageId="24" pageNumber="101">Variability.</emphasis>
|
|
The type series shows a strong uniformity as for size, shape, setation and microsculpturing.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="24" pageNumber="101" type="etymology">
|
|
<paragraph pageId="24" pageNumber="101">
|
|
<emphasis bold="true" box="[185,320,413,439]" pageId="24" pageNumber="101">Etymology.</emphasis>
|
|
Dedicated to Mr. Tomoyuki Tsuru, who found the huge termite nest from which the junior author collected more than 400 Ceratocanthinae, among which
|
|
<taxonomicName box="[140,229,484,510]" class="Insecta" family="Hybosoridae" genus="Pterorthochaetes" genus.bestMatchDistance="1276" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="101" phylum="Arthropoda" rank="species" species="tsurui" status="sp. n.">
|
|
<emphasis box="[140,229,484,510]" italics="true" pageId="24" pageNumber="101">P. tsurui</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel box="[236,298,484,510]" pageId="24" pageNumber="101" rank="species">sp. n.</taxonomicNameLabel>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="24" pageNumber="101" type="distribution">
|
|
<paragraph pageId="24" pageNumber="101">
|
|
<emphasis bold="true" box="[185,491,519,545]" pageId="24" pageNumber="101">Distribution and habitat.</emphasis>
|
|
This new species is know only from the type locality. For details on collecting circumstances see the introductory paragraphs.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |