treatments-xml/data/03/82/E7/0382E754E764FFB3FF22F90BFDE4FD1C.xml

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<mods:title id="1C78537FE8A205B7AD52ECA3B46E6132">A review of Peruvian diving beetles of the genus Hydaticus Leach, 1817, with description of Hydaticus (Prodaticus) panguana sp. nov. and notes on other Neotropical species (Coleoptera: Dytiscidae)</mods:title>
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<mods:namePart id="5FCEE29D18420A2A718C7991FE77C157">Megna, Yoandri S.</mods:namePart>
<mods:affiliation id="C8B1C108B286B44B7965911E2A6E3ED5">SNSB-Zoologische Staatssammlung München, Münchhausenstrasse 21, 81247 München, Germany &amp; Present address: Departamento de Biología, Universidad de Oriente. Patricio Lumumba s / n, Santiago, Santiago de Cuba, Cuba. Emil-Überall-Str. 29, 01159 Dresden, Germany</mods:affiliation>
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<mods:namePart id="DC9061E8D2CB00A295A57830F474D94F">Balke, Michael</mods:namePart>
<mods:affiliation id="E1E32AE8701067D7C585E5057078F416">SNSB-Zoologische Staatssammlung München, Münchhausenstrasse 21, 81247 München, Germany</mods:affiliation>
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<mods:namePart id="CEEEFF6052F27FC564E3204798011B11">Apenborn, Rico</mods:namePart>
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<mods:namePart id="48FE10D53A204D76CF55D4EFC56E0556">Hendrich, Lars</mods:namePart>
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<heading id="D0DCE12EE764FFB1FF22F90BFD70FEEE" bold="true" box="[151,661,1762,1789]" fontSize="11" level="1" pageId="1" pageNumber="114" reason="1">
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<taxonomicName id="4C2B2DC1E764FFB1FF22F90BFDD5FEEE" ID-CoL="8LZ8X" authority="Megna &amp; Balke &amp; Apenborn &amp; Hendrich, 2019" authorityName="Megna &amp; Balke &amp; Apenborn &amp; Hendrich" authorityYear="2019" box="[151,560,1762,1789]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="114" phylum="Arthropoda" rank="species" species="panguana" status="sp. nov." subGenus="Prodaticus">
<emphasis id="B95F8A50E764FFB1FF22F90BFEF4FEEE" bold="true" box="[151,273,1762,1789]" italics="true" pageId="1" pageNumber="114">Hydaticus</emphasis>
(
<emphasis id="B95F8A50E764FFB1FE96F90BFE40FEEE" bold="true" box="[291,421,1762,1789]" italics="true" pageId="1" pageNumber="114">Prodaticus</emphasis>
) 
<emphasis id="B95F8A50E764FFB1FE03F90AFDD5FEEE" bold="true" box="[438,560,1763,1789]" italics="true" pageId="1" pageNumber="114">panguana</emphasis>
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(
<figureCitation id="13104AC7E764FFB1FF2AF8EEFF19FF32" box="[159,252,1799,1825]" captionStart="FIGURE 2" captionStartId="6.[151,250,1970,1995]" captionTargetBox="[196,1395,185,1946]" captionTargetId="figure@6.[186,1401,181,1951]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of H. panguana sp. nov. (A), H. lateralis (B), H. subfasciatus (C) and H. xanthomelas (D). All specimens collected in Panguana, Peru. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995402" httpUri="https://zenodo.org/record/3995402/files/figure.png" pageId="1" pageNumber="114">Figs 2A</figureCitation>
, 
<figureCitation id="13104AC7E764FFB1FEBDF8EEFEFDFF32" box="[264,280,1799,1825]" captionStart="FIGURES 4–7" captionStartId="8.[151,264,1972,1997]" captionTargetBox="[173,1415,181,1952]" captionTargetId="figure@8.[173,1415,181,1952]" captionTargetPageId="8" captionText="FIGURES 4–7. Median lobe of aedeagus in dorsal (A) and lateral view (B): H. panguana sp. nov. (4), H. lateralis (5), H. subfasciatus (6), and H. xanthomelas (7). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3995406" httpUri="https://zenodo.org/record/3995406/files/figure.png" pageId="1" pageNumber="114">4</figureCitation>
, 
<figureCitation id="13104AC7E764FFB1FE91F8EEFED6FF32" box="[292,307,1799,1825]" captionStart="FIGURE 8" captionStartId="11.[151,250,1243,1268]" captionText="FIGURE 8. All known records of Hydaticus in Peru." pageId="1" pageNumber="114">8</figureCitation>
, 
<figureCitation id="13104AC7E764FFB1FEF5F8EEFE81FF32" box="[320,356,1799,1825]" captionStart="FIGURE 9" captionStartId="12.[151,250,1958,1983]" captionTargetBox="[213,1375,181,1938]" captionTargetId="figure@12.[213,1375,181,1938]" captionTargetPageId="12" captionText="FIGURE 9. Habitats of H. panguana sp. nov. and H. lateralis (A), and H. subfasciatus and H. lateralis (B) in Panguana, Huànuco, Peru. Shallow and temporary forest pool with rotten leaves and wooden debris (A), and Aguajal forest pond (B)." figureDoi="http://doi.org/10.5281/zenodo.3995408" httpUri="https://zenodo.org/record/3995408/files/figure.png" pageId="1" pageNumber="114">9A</figureCitation>
)
</paragraph>
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<paragraph id="8B945642E764FFB1FF22F8A6FF3BFF9D" blockId="1.[151,1437,1870,2041]" pageId="1" pageNumber="114">
<emphasis id="B95F8A50E764FFB1FF22F8A6FED1FF7A" bold="true" box="[151,308,1870,1897]" pageId="1" pageNumber="114">Type locality.</emphasis>
<materialsCitation id="3B435C1FE764FFB1FE8FF8A6FA79FF7A" ID-GBIF-Occurrence="2847257304" box="[314,1436,1871,1897]" country="Peru" county="Department" elevation="230" latitude="-9.62" location="River Yuyapichis" longLatPrecision="779" longitude="-74.93" municipality="Panguana" pageId="1" pageNumber="114" specimenCount="1" stateProvince="Huanuco">
<collectingCountry id="F33C16D2E764FFB1FE8FF8A6FE95FF7A" box="[314,368,1871,1897]" name="Peru" pageId="1" pageNumber="114">Peru</collectingCountry>
, 
<collectingRegion id="49EF98A0E764FFB1FECFF8A6FE3AFF7A" box="[378,479,1871,1897]" country="Peru" name="Huanuco" pageId="1" pageNumber="114">Huánuco</collectingRegion>
<collectingCounty id="62F52ECEE764FFB1FE50F8A6FD89FF7A" box="[485,620,1871,1897]" pageId="1" pageNumber="114">Department</collectingCounty>
, 
<collectingMunicipality id="6BF0CC38E764FFB1FDC2F8A6FD02FF7A" box="[631,743,1871,1897]" pageId="1" pageNumber="114">Panguana</collectingMunicipality>
, 
<location id="8EF40099E764FFB1FD45F8A6FC56FF7A" LSID="urn:lsid:plazi:treatment:0382E754E764FFB3FF22F90BFDE4FD1C:8EF40099E764FFB1FD45F8A6FC56FF7A" box="[752,947,1871,1897]" country="Peru" county="Department" latitude="-9.62" longLatPrecision="779" longitude="-74.93" municipality="Panguana" name="River Yuyapichis" pageId="1" pageNumber="114" stateProvince="Huanuco">River Yuyapichis</location>
; at 
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<geoCoordinate id="EE1F3085E764FFB1FBABF8A6FB9DFF7A" box="[1054,1144,1871,1897]" degrees="74.93" direction="west" orientation="longitude" pageId="1" pageNumber="114" precision="555" value="-74.93">74.93W</geoCoordinate>
, ca. 
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above sea level
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(a.s.l).
</paragraph>
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, male (
</emphasis>
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<emphasis id="B95F8A50E764FFB1FE57F87FFE17FFA2" bold="true" box="[482,498,1942,1969]" pageId="1" pageNumber="114">).</emphasis>
“ 
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, 
<collectingCounty id="62F52ECEE764FFB1FDEEF87EFCF0FFA2" box="[603,789,1943,1969]" pageId="1" pageNumber="114">Yuyapichis river</collectingCounty>
, 
<collectingMunicipality id="6BF0CC38E764FFB1FC95F87EFC75FFA2" box="[800,912,1943,1969]" pageId="1" pageNumber="114">Panguana</collectingMunicipality>
, 
<collectingRegion id="49EF98A0E764FFB1FC2EF87EFBE5FFA2" box="[923,1024,1943,1969]" country="Peru" name="Huanuco" pageId="1" pageNumber="114">Huánuco</collectingRegion>
<location id="8EF40099E764FFB1FBB2F87EFB6BFFA2" LSID="urn:lsid:plazi:treatment:0382E754E764FFB3FF22F90BFDE4FD1C:8EF40099E764FFB1FBB2F87EFB6BFFA2" box="[1031,1166,1943,1969]" country="Peru" county="Yuyapichis river" latitude="-9.6167" longLatPrecision="7" longitude="-74.9333" municipality="Panguana" name="Department" pageId="1" pageNumber="114" stateProvince="Huanuco">Department</location>
, 
<location id="8EF40099E764FFB1FB2FF87EFB35FFA2" LSID="urn:lsid:plazi:treatment:0382E754E764FFB3FF22F90BFDE4FD1C:8EF40099E764FFB1FB2FF87EFB35FFA2" box="[1178,1232,1943,1969]" country="Peru" county="Yuyapichis river" latitude="-9.6167" longLatPrecision="7" longitude="-74.9333" municipality="Panguana" name="Peru" pageId="1" pageNumber="114" stateProvince="Huanuco">Peru</location>
, 
<date id="FF957082E764FFB1FB6EF87FFA72FFA2" box="[1243,1431,1942,1969]" pageId="1" pageNumber="114" value="2013-05-10" valueMax="2013-07-25" valueMin="2013-05-10">
<collectingDate id="EFD1896AE764FFB1FB6EF87FFA72FFA2" box="[1243,1431,1942,1969]" pageId="1" pageNumber="114" value="2013-05-10" valueMax="2013-07-25" valueMin="2013-05-10">10.v-25.vii.2013</collectingDate>
</date>
, - 
<geoCoordinate id="EE1F3085E764FFB1FF2AF852FEE5FFC6" box="[159,256,1979,2005]" degrees="9.6167" direction="south" orientation="latitude" pageId="1" pageNumber="114" precision="5" value="-9.6167">9.6167S</geoCoordinate>
, - 
<geoCoordinate id="EE1F3085E764FFB1FEA1F852FE69FFC6" box="[276,396,1979,2005]" degrees="74.9333" direction="west" orientation="longitude" pageId="1" pageNumber="114" precision="5" value="-74.9333">74.9333W</geoCoordinate>
, ca. 
<quantity id="4CD3FBA7E764FFB1FE77F852FDE8FFC6" box="[450,525,1979,2005]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="114" unit="m" value="230.0">
<elevation id="0006B171E764FFB1FE77F852FDE8FFC6" box="[450,525,1979,2005]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="114" unit="m" value="230.0">230 m</elevation>
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, leg. 
<collectorName id="26DE3394E764FFB1FDF8F852FD01FFC6" box="[589,740,1979,2005]" pageId="1" pageNumber="114">R. Apenborn</collectorName>
”, / “ 
<typeStatus id="5490E8E0E764FFB1FCA1F852FC9AFFC6" box="[788,895,1979,2005]" pageId="1" pageNumber="114" type="holotype">Holotype</typeStatus>
<taxonomicName id="4C2B2DC1E764FFB1FC3CF852FB96FFC6" authority="Megna &amp; Balke &amp; Apenborn &amp; Hendrich, 2019" authorityName="Megna &amp; Balke &amp; Apenborn &amp; Hendrich" authorityYear="2019" box="[905,1139,1979,2005]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="114" phylum="Arthropoda" rank="species" species="panguana" status="sp. nov.">
<emphasis id="B95F8A50E764FFB1FC3CF852FB96FFC6" box="[905,1139,1979,2005]" italics="true" pageId="1" pageNumber="114">Hydaticus panguana</emphasis>
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<taxonomicNameLabel id="A26C372BE764FFB1FBC9F852FB32FFC6" box="[1148,1239,1979,2005]" pageId="1" pageNumber="114" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<determinerName id="7E48BA86E764FFB1FB53F852FA92FFC6" box="[1254,1399,1979,2005]" pageId="1" pageNumber="114">Y. S. Megna</determinerName>
, 
<determinerName id="7E48BA86E764FFB1FA31F852FF1AFFEA" pageId="1" pageNumber="114">L. Hendrich</determinerName>
&amp; 
<determinerName id="7E48BA86E764FFB1FE92F836FE74FFEA" box="[295,401,2015,2041]" pageId="1" pageNumber="114">M. Balke</determinerName>
det. 2018” [red printed label]
</materialsCitation>
. 
<materialsCitation id="3B435C1FE764FFB2FD4CF836FF3BF8A5" ID-GBIF-Occurrence="2847257375" collectionCode="MUSM, ZSM" country="Peru" lastPageId="2" lastPageNumber="115" pageId="1" pageNumber="114" specimenCount="4" stateProvince="Huanuco" typeStatus="paratype">
<emphasis id="B95F8A50E764FFB1FD4CF836FC94FFEA" bold="true" box="[761,881,2015,2041]" pageId="1" pageNumber="114">
<typeStatus id="5490E8E0E764FFB1FD4CF836FC94FFEA" box="[761,881,2015,2041]" pageId="1" pageNumber="114" type="paratype">Paratypes</typeStatus>
</emphasis>
: 
<specimenCount id="9D2D9DCBE764FFB1FC34F836FBEBFFEA" box="[897,1038,2015,2041]" pageId="1" pageNumber="114" type="generic">4 specimens</specimenCount>
with the same label data (
<collectionCode id="ED3ACE87E764FFB1FA8BF836FA7DFFEA" box="[1342,1432,2015,2041]" pageId="1" pageNumber="114">MUSM</collectionCode>
, 
<collectionCode id="ED3ACE87E767FFB2FF22FF75FF30F8A5" box="[151,213,156,182]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="2" pageNumber="115" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B435C1FE767FFB2FF5CFF75FBAEF8C9" ID-GBIF-Occurrence="2847257312" collectingDate="2017-10-10" collectionCode="ZSM" collectorName="R. Yuyapichis &amp; Burmeister" country="Peru" latitude="-9.616667" location="Wald Station" longLatPrecision="1299" longitude="-74.583336" municipality="Peru Prov." pageId="2" pageNumber="115" specimenCount="1" stateProvince="Huanuco" typeStatus="paratype">
<specimenCount id="9D2D9DCBE767FFB2FF5CFF75FEC4F8A5" box="[233,289,156,182]" pageId="2" pageNumber="115" type="generic">1 ex.</specimenCount>
, “ 
<collectingMunicipality id="6BF0CC38E767FFB2FE83FF75FE4BF8A5" box="[310,430,156,182]" pageId="2" pageNumber="115">Peru Prov.</collectingMunicipality>
<collectingRegion id="49EF98A0E767FFB2FE01FF75FDFCF8A5" box="[436,537,156,182]" country="Peru" name="Huanuco" pageId="2" pageNumber="115">Huánuco</collectingRegion>
<collectorName id="26DE3394E767FFB2FD95FF75FD5BF8A5" box="[544,702,156,182]" pageId="2" pageNumber="115">R. Yuyapichis</collectorName>
, Stat. Panguana, 
<geoCoordinate id="EE1F3085E767FFB2FCC8FF75FC2FF8A5" box="[893,970,152,183]" degrees="9" direction="south" minutes="37" orientation="latitude" pageId="2" pageNumber="115" precision="925" value="-9.616667">9°37ʹS</geoCoordinate>
, 
<geoCoordinate id="EE1F3085E767FFB2FC63FF72FBD9F8A5" box="[982,1084,152,183]" degrees="74" direction="west" minutes="35" orientation="longitude" pageId="2" pageNumber="115" precision="925" value="-74.583336">74°35ʹW</geoCoordinate>
, 
<location id="8EF40099E767FFB2FBF3FF75FB33F8A5" LSID="urn:lsid:plazi:treatment:0382E754E764FFB3FF22F90BFDE4FD1C:8EF40099E767FFB2FBF3FF75FB33F8A5" box="[1094,1238,156,182]" country="Peru" latitude="-9.616667" longLatPrecision="1299" longitude="-74.583336" municipality="Peru Prov." name="Wald Station" pageId="2" pageNumber="115" stateProvince="Huanuco">Wald Station</location>
27.9.- 
<date id="FF957082E767FFB2FAAFFF75FA79F8A5" box="[1306,1436,156,182]" pageId="2" pageNumber="115" value="2017-10-10">
<collectingDate id="EFD1896AE767FFB2FAAFFF75FA79F8A5" box="[1306,1436,156,182]" pageId="2" pageNumber="115" value="2017-10-10">10.10.2017</collectingDate>
</date>
leg. 
<collectorName id="26DE3394E767FFB2FF72FF29FEA2F8C9" box="[199,327,192,218]" pageId="2" pageNumber="115">Burmeister</collectorName>
” (
<collectionCode id="ED3ACE87E767FFB2FEEAFF29FE7BF8C9" box="[351,414,192,218]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="2" pageNumber="115" type="Museum">ZSM</collectionCode>
). Each 
<typeStatus id="5490E8E0E767FFB2FE45FF29FDB5F8C9" box="[496,592,192,218]" pageId="2" pageNumber="115" type="paratype">paratype</typeStatus>
is provided with a red printed 
<typeStatus id="5490E8E0E767FFB2FC1CFF29FBECF8C9" box="[937,1033,192,218]" pageId="2" pageNumber="115" type="paratype">paratype</typeStatus>
label
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33105C9E767FFB3FF72FF0AFEA7FAE0" lastPageId="3" lastPageNumber="116" pageId="2" pageNumber="115" type="description">
<paragraph id="8B945642E767FFB2FF72FF0AFE43F955" blockId="2.[151,1437,152,470]" pageId="2" pageNumber="115">
<emphasis id="B95F8A50E767FFB2FF72FF0AFE02F8ED" bold="true" box="[199,487,227,254]" pageId="2" pageNumber="115">
Description of 
<typeStatus id="5490E8E0E767FFB2FECCFF0DFE07F8ED" box="[377,482,228,254]" pageId="2" pageNumber="115" type="holotype">holotype</typeStatus>
.
</emphasis>
TL = 14.0 mm; TL-H = 
<quantity id="4CD3FBA7E767FFB2FCB6FF0DFC8EF8ED" box="[771,875,228,254]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="2" pageNumber="115" unit="mm" value="13.3">13.3 mm</quantity>
; MW = 
<quantity id="4CD3FBA7E767FFB2FC7EFF0DFBC0F8ED" box="[971,1061,228,254]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="2" pageNumber="115" unit="mm" value="7.2">7.2 mm</quantity>
. Body oval, elongate, slightly attenuated posteriorly, dorsoventrally depressed; subparallel medially, widest just behind middle, lateral outline continuous in dorsal aspect.
</paragraph>
<paragraph id="8B945642E767FFB2FF72FEA6FC4AF9C5" blockId="2.[151,1437,152,470]" pageId="2" pageNumber="115">
<emphasis id="B95F8A50E767FFB2FF72FEA6FEC5F979" bold="true" box="[199,288,335,362]" pageId="2" pageNumber="115">Colour.</emphasis>
Dorsal surface predominantly black. Head capsule black with clypeus and frons testaceous (
<figureCitation id="13104AC7E767FFB2FA80FEB9FA75F979" box="[1333,1424,336,362]" captionStart="FIGURE 2" captionStartId="6.[151,250,1970,1995]" captionTargetBox="[196,1395,185,1946]" captionTargetId="figure@6.[186,1401,181,1951]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of H. panguana sp. nov. (A), H. lateralis (B), H. subfasciatus (C) and H. xanthomelas (D). All specimens collected in Panguana, Peru. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995402" httpUri="https://zenodo.org/record/3995402/files/figure.png" pageId="2" pageNumber="115">Fig. 2A</figureCitation>
). Pronotum black with lateral margins testaceous. Elytron black, with two lateral testaceous spots, one basal and another just behind middle, lateral margin of apex largely irrorated. Ventral surface predominantly rufous, antenna, prosternum, mouthparts, pro-, meso- and metathoracic legs testaceous.
</paragraph>
<caption id="DF5406CAE767FFB2FF22F899FC27FFE6" ID-DOI="http://doi.org/10.5281/zenodo.3995400" ID-Zenodo-Dep="3995400" httpUri="https://zenodo.org/record/3995400/files/figure.png" pageId="2" pageNumber="115" startId="2.[151,250,1904,1929]" targetBox="[391,1187,591,1847]" targetPageId="2">
<paragraph id="8B945642E767FFB2FF22F899FC27FFE6" blockId="2.[151,1437,1904,2037]" pageId="2" pageNumber="115">
<emphasis id="B95F8A50E767FFB2FF22F899FEF7FF9A" bold="true" box="[151,274,1904,1929]" pageId="2" pageNumber="115">FIGURE 1.</emphasis>
Dorsal view of the colour pattern of a generalized 
<taxonomicName id="4C2B2DC1E767FFB2FCA1F898FC9EFF9A" authorityName="Leach" authorityYear="1817" box="[788,891,1905,1929]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis id="B95F8A50E767FFB2FCA1F898FC9EFF9A" box="[788,891,1905,1929]" italics="true" pageId="2" pageNumber="115">Hydaticus</emphasis>
</taxonomicName>
for the Neotropical region. Head (He): Ptbb-posterior transverse black band. Pronotum (Pr): Abb- anterior black band; Pbb-posterior black band; Plaa- posterior lateral angle acute. Scutellum (Sc). Elytron (El): Ai- apex irrorated; Bs- basal spot; Ls- lateral yellow stripes; Ms- middle spot; Sls- Subbasal and lateral yellow stripe; Tbf- transverse basal fasciae; Tpi- third posterior irrorated.
</paragraph>
</caption>
<paragraph id="8B945642E766FFB3FF72FF71FC48F8C4" blockId="3.[151,1437,152,1295]" pageId="3" pageNumber="116">
<emphasis id="B95F8A50E766FFB3FF72FF71FE19F8A0" bold="true" box="[199,508,152,179]" pageId="3" pageNumber="116">Sculpture and punctation.</emphasis>
Dosal sculpture consisting of very fine, dense micropunctures and fine microreticulation. Protarsomeres II with strong reticulations on the dorsal surface.
</paragraph>
<paragraph id="8B945642E766FFB3FF72FF09FE5BF9E4" blockId="3.[151,1437,152,1295]" pageId="3" pageNumber="116">
<emphasis id="B95F8A50E766FFB3FF72FF09FEA4F8E8" bold="true" box="[199,321,224,251]" pageId="3" pageNumber="116">Structure.</emphasis>
Pronotum with posterolateral angle produced and acute, posterior margin broadly sinuate and recurved lateral to basomedial lobe. Prosternal process almost flat. Metaventral wing relatively broad basally, narrow laterally with anterolateral margin straight. Protarsomere I with a series of spines across the dorsal surface. Protibia with stout pegs on the apicodorsal surface. Mesotarsomeres with a small basal brush of setae ventrally. Metatibial spurs acuminate apically. Posterior surface of metafemur with few large punctures on anterior surface and without interspersed small punctures. Metatibia without punctures on anterior surface; posterior surface with dorsal series of seven bifid setae curved ventro-basally, not parallel to dorsal margin. Metatarsal claws unequal, anterior claw shorter than posterior one.
</paragraph>
<paragraph id="8B945642E766FFB3FF72FDE8FE84FA70" blockId="3.[151,1437,152,1295]" pageId="3" pageNumber="116">
<emphasis id="B95F8A50E766FFB3FF72FDE8FE9FFA08" bold="true" box="[199,378,512,539]" pageId="3" pageNumber="116">Male genitalia.</emphasis>
Median lobe of aedeagus narrow in ventral view; in lateral view enlarged in apical third and with apex rounded (
<figureCitation id="13104AC7E766FFB3FEC2FDCCFE58FA2C" box="[375,445,549,575]" captionStart="FIGURES 4–7" captionStartId="8.[151,264,1972,1997]" captionTargetBox="[173,1415,181,1952]" captionTargetId="figure@8.[173,1415,181,1952]" captionTargetPageId="8" captionText="FIGURES 4–7. Median lobe of aedeagus in dorsal (A) and lateral view (B): H. panguana sp. nov. (4), H. lateralis (5), H. subfasciatus (6), and H. xanthomelas (7). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3995406" httpUri="https://zenodo.org/record/3995406/files/figure.png" pageId="3" pageNumber="116">Fig. 4</figureCitation>
). Parameres are not illustrated because by no means useful in identification of most species of 
<taxonomicName id="4C2B2DC1E766FFB3FF5DFDA0FEBFFA70" authorityName="Leach" authorityYear="1817" box="[232,346,585,611]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B95F8A50E766FFB3FF5DFDA0FEBFFA70" box="[232,346,585,611]" italics="true" pageId="3" pageNumber="116">Hydaticus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B945642E766FFB3FF72FD85FEA7FAE0" blockId="3.[151,1437,152,1295]" pageId="3" pageNumber="116">
<emphasis id="B95F8A50E766FFB3FF72FD85FE52FA94" bold="true" box="[199,439,620,647]" pageId="3" pageNumber="116">Sexual dimorphism.</emphasis>
Male with protarsomeres I-III enlarged to form a large oval palette which ventrally bears 22 rounded suckers; mesotarsomeres I-III moderately enlarged, ventrally with 14 rounded suckers. Female with TL = 13.0– 
<quantity id="4CD3FBA7E766FFB3FF58FD5CFEB6FADC" box="[237,339,693,719]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="3" pageNumber="116" unit="mm" value="13.2">13.2 mm</quantity>
; TL-H = 12.3–12.5; MW = 
<quantity id="4CD3FBA7E766FFB3FD3CFD5CFCF7FADC" box="[649,786,693,719]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="7.1" metricValueMin="6.9" pageId="3" pageNumber="116" unit="mm" value="7.0" valueMax="7.1" valueMin="6.9">6.9–7.1 mm</quantity>
; metatarsomeres of female with natatory setae on posteroventral margin.
</paragraph>
</subSubSection>
<subSubSection id="C33105C9E766FFB3FF72FD15FB4EFC00" pageId="3" pageNumber="116" type="diagnosis">
<paragraph id="8B945642E766FFB3FF72FD15FE5CFBB4" blockId="3.[151,1437,152,1295]" pageId="3" pageNumber="116">
<emphasis id="B95F8A50E766FFB3FF72FD15FEA7FB04" bold="true" box="[199,322,764,791]" pageId="3" pageNumber="116">Diagnosis.</emphasis>
TL = 13.0–14.0 mm; TL-H = 
<quantity id="4CD3FBA7E766FFB3FD3AFD14FCD1FB04" box="[655,820,765,791]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" metricValueMax="1.33" metricValueMin="1.23" pageId="3" pageNumber="116" unit="mm" value="12.8" valueMax="13.3" valueMin="12.3">12.3–13.3 mm</quantity>
; MW = 
<quantity id="4CD3FBA7E766FFB3FC38FD15FBC1FB04" box="[909,1060,764,791]" metricMagnitude="-3" metricUnit="m" metricValue="7.025" metricValueMax="7.15" metricValueMin="6.9" pageId="3" pageNumber="116" unit="mm" value="7.025" valueMax="7.15" valueMin="6.9">6.9–7.15 mm</quantity>
; pronotum black with lateral margins testaceous; elytron with two lateral testaceous spots, one basal and another just behind middle, lateral margin of apex largely irrorated; protarsomere I with a series of spines across the dorsal surface; metatibia with bifid setae curved ventro-basally on posterior surface; median lobe of aedeagus with apex rounded and curved toward the dorsal margin in lateral view.
</paragraph>
<paragraph id="8B945642E766FFB3FF72FC59FB4EFC00" blockId="3.[151,1437,152,1295]" pageId="3" pageNumber="116">
<emphasis id="B95F8A50E766FFB3FF72FC59FE28FBD8" bold="true" box="[199,461,944,971]" pageId="3" pageNumber="116">Differential diagnosis.</emphasis>
The shape of the median lobe of 
<taxonomicName id="4C2B2DC1E766FFB3FC8FFC5BFC29FBD8" authority="Megna &amp; Balke &amp; Apenborn &amp; Hendrich, 2019" authorityName="Megna &amp; Balke &amp; Apenborn &amp; Hendrich" authorityYear="2019" box="[826,972,946,971]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="panguana" status="sp. nov.">
<emphasis id="B95F8A50E766FFB3FC8FFC5BFC29FBD8" box="[826,972,946,971]" italics="true" pageId="3" pageNumber="116">H. panguana</emphasis>
</taxonomicName>
<emphasis id="B95F8A50E766FFB3FC67FC58FBCFFBD8" bold="true" box="[978,1066,945,971]" pageId="3" pageNumber="116">
<taxonomicNameLabel id="A26C372BE766FFB3FC67FC58FBCFFBD8" box="[978,1066,945,971]" pageId="3" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to that of 
<taxonomicName id="4C2B2DC1E766FFB3FAB0FC5BFF5DFBFC" baseAuthorityName="Brulle" baseAuthorityYear="1837" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="xanthomelas">
<emphasis id="B95F8A50E766FFB3FAB0FC5BFF5DFBFC" italics="true" pageId="3" pageNumber="116">H. xanthomelas</emphasis>
</taxonomicName>
(
<figureCitation id="13104AC7E766FFB3FF73FC3CFEEDFBFC" box="[198,264,981,1007]" captionStart="FIGURES 4–7" captionStartId="8.[151,264,1972,1997]" captionTargetBox="[173,1415,181,1952]" captionTargetId="figure@8.[173,1415,181,1952]" captionTargetPageId="8" captionText="FIGURES 4–7. Median lobe of aedeagus in dorsal (A) and lateral view (B): H. panguana sp. nov. (4), H. lateralis (5), H. subfasciatus (6), and H. xanthomelas (7). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3995406" httpUri="https://zenodo.org/record/3995406/files/figure.png" pageId="3" pageNumber="116">Fig. 7</figureCitation>
) but both species can be easily separated by its different size (TL= 13.0–14.0 mm in 
<taxonomicName id="4C2B2DC1E766FFB3FB18FC3FFADBFBFC" authority="Megna &amp; Balke &amp; Apenborn &amp; Hendrich, 2019" authorityName="Megna &amp; Balke &amp; Apenborn &amp; Hendrich" authorityYear="2019" box="[1197,1342,982,1007]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="panguana" status="sp. nov.">
<emphasis id="B95F8A50E766FFB3FB18FC3FFADBFBFC" box="[1197,1342,982,1007]" italics="true" pageId="3" pageNumber="116">H. panguana</emphasis>
</taxonomicName>
<emphasis id="B95F8A50E766FFB3FAF1FC3CFA79FBFC" bold="true" box="[1348,1436,981,1007]" pageId="3" pageNumber="116">
<taxonomicNameLabel id="A26C372BE766FFB3FAF1FC3CFA79FBFC" box="[1348,1436,981,1007]" pageId="3" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and TL = 
<quantity id="4CD3FBA7E766FFB3FEB0FC10FE7FFC00" box="[261,410,1017,1043]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.04" metricValueMin="0.96" pageId="3" pageNumber="116" unit="mm" value="10.0" valueMax="10.4" valueMin="9.6">9.6–10.4 mm</quantity>
in 
<taxonomicName id="4C2B2DC1E766FFB3FE08FC13FD8BFC00" baseAuthorityName="Brulle" baseAuthorityYear="1837" box="[445,622,1017,1043]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="xanthomelas">
<emphasis id="B95F8A50E766FFB3FE08FC13FD8BFC00" box="[445,622,1017,1043]" italics="true" pageId="3" pageNumber="116">H. xanthomelas</emphasis>
</taxonomicName>
) and the colour pattern of the elytra (
<figureCitation id="13104AC7E766FFB3FBA5FC10FB7AFC00" box="[1040,1183,1017,1043]" captionStart="FIGURE 2" captionStartId="6.[151,250,1970,1995]" captionTargetBox="[196,1395,185,1946]" captionTargetId="figure@6.[186,1401,181,1951]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of H. panguana sp. nov. (A), H. lateralis (B), H. subfasciatus (C) and H. xanthomelas (D). All specimens collected in Panguana, Peru. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995402" httpUri="https://zenodo.org/record/3995402/files/figure.png" pageId="3" pageNumber="116">Figs 2A, 2D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33105C9E766FFB3FF72FBF4FB53FC48" pageId="3" pageNumber="116" type="etymology">
<paragraph id="8B945642E766FFB3FF72FBF4FB53FC48" blockId="3.[151,1437,152,1295]" pageId="3" pageNumber="116">
<emphasis id="B95F8A50E766FFB3FF72FBF4FEABFC24" bold="true" box="[199,334,1053,1079]" pageId="3" pageNumber="116">Etymology.</emphasis>
This species is named in reference to the “Área de Conservación Privada Panguana”, where the species was discovered. The name is a noun in the nominative singular standing in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C33105C9E766FFB3FF72FB8DFB1DFCB0" pageId="3" pageNumber="116" type="distribution">
<paragraph id="8B945642E766FFB3FF72FB8DFB1DFCB0" blockId="3.[151,1437,152,1295]" pageId="3" pageNumber="116">
<emphasis id="B95F8A50E766FFB3FF72FB8DFE84FC6C" bold="true" box="[199,353,1124,1151]" pageId="3" pageNumber="116">Distribution.</emphasis>
So far 
<taxonomicName id="4C2B2DC1E766FFB3FE0DFB8CFD47FC6C" authority="Megna &amp; Balke &amp; Apenborn &amp; Hendrich, 2019" authorityName="Megna &amp; Balke &amp; Apenborn &amp; Hendrich" authorityYear="2019" box="[440,674,1125,1151]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="panguana" status="sp. nov.">
<emphasis id="B95F8A50E766FFB3FE0DFB8CFD47FC6C" box="[440,674,1125,1151]" italics="true" pageId="3" pageNumber="116">Hydaticus panguana</emphasis>
</taxonomicName>
<emphasis id="B95F8A50E766FFB3FD1EFB8CFCE3FC6C" bold="true" box="[683,774,1125,1151]" pageId="3" pageNumber="116">
<taxonomicNameLabel id="A26C372BE766FFB3FD1EFB8CFCE3FC6C" box="[683,774,1125,1151]" pageId="3" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is only known from the 
<typeStatus id="5490E8E0E766FFB3FB91FB8CFBB1FC6C" box="[1060,1108,1125,1151]" pageId="3" pageNumber="116">type</typeStatus>
locality in 
<collectingRegion id="49EF98A0E766FFB3FB6CFB8CFADBFC6C" box="[1241,1342,1125,1151]" country="Peru" name="Huanuco" pageId="3" pageNumber="116">Huánuco</collectingRegion>
Department, Panguana Field Station, 
<collectingCountry id="F33C16D2E766FFB3FE5AFB60FDC7FCB0" box="[495,546,1161,1187]" name="Peru" pageId="3" pageNumber="116">Peru</collectingCountry>
(
<figureCitation id="13104AC7E766FFB3FD87FB60FD93FCB0" box="[562,630,1161,1187]" captionStart="FIGURE 8" captionStartId="11.[151,250,1243,1268]" captionText="FIGURE 8. All known records of Hydaticus in Peru." pageId="3" pageNumber="116">Fig. 8</figureCitation>
). Probably more widespread in the Amazonian lowlands.
</paragraph>
</subSubSection>
<subSubSection id="C33105C9E766FFB3FF72FB44FDE4FD1C" pageId="3" pageNumber="116" type="biology_ecology">
<paragraph id="8B945642E766FFB3FF72FB44FDE4FD1C" blockId="3.[151,1437,152,1295]" pageId="3" pageNumber="116">
<emphasis id="B95F8A50E766FFB3FF72FB44FEC9FCD4" bold="true" box="[199,300,1197,1223]" pageId="3" pageNumber="116">Ecology.</emphasis>
Most specimens of 
<taxonomicName id="4C2B2DC1E766FFB3FDA4FB44FD1EFCD4" authority="Megna &amp; Balke &amp; Apenborn &amp; Hendrich, 2019" authorityName="Megna &amp; Balke &amp; Apenborn &amp; Hendrich" authorityYear="2019" box="[529,763,1197,1223]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="panguana" status="sp. nov.">
<emphasis id="B95F8A50E766FFB3FDA4FB44FD1EFCD4" box="[529,763,1197,1223]" italics="true" pageId="3" pageNumber="116">Hydaticus panguana</emphasis>
</taxonomicName>
<emphasis id="B95F8A50E766FFB3FCB6FB44FCB8FCD4" bold="true" box="[771,861,1197,1223]" pageId="3" pageNumber="116">
<taxonomicNameLabel id="A26C372BE766FFB3FCB6FB44FCB8FCD4" box="[771,861,1197,1223]" pageId="3" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
were collected in a partly shaded, shallow, temporary, ditch-like forest pool, rich in decaying leaves and twigs (
<figureCitation id="13104AC7E766FFB3FCF6FB38FC79FCF8" box="[835,924,1233,1259]" captionStart="FIGURE 9" captionStartId="12.[151,250,1958,1983]" captionTargetBox="[213,1375,181,1938]" captionTargetId="figure@12.[213,1375,181,1938]" captionTargetPageId="12" captionText="FIGURE 9. Habitats of H. panguana sp. nov. and H. lateralis (A), and H. subfasciatus and H. lateralis (B) in Panguana, Huànuco, Peru. Shallow and temporary forest pool with rotten leaves and wooden debris (A), and Aguajal forest pond (B)." figureDoi="http://doi.org/10.5281/zenodo.3995408" httpUri="https://zenodo.org/record/3995408/files/figure.png" pageId="3" pageNumber="116">Fig. 9A</figureCitation>
). The species is sympatric if not syntopic with 
<taxonomicName id="4C2B2DC1E766FFB3FF22FB1FFEF3FD1C" authorityName="Laporte" authorityYear="1835" box="[151,278,1269,1295]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="B95F8A50E766FFB3FF22FB1FFEF3FD1C" box="[151,278,1269,1295]" italics="true" pageId="3" pageNumber="116">H. lateralis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2B2DC1E766FFB3FEF8FB1FFE1CFD1C" authorityName="Laporte" authorityYear="1835" box="[333,505,1269,1295]" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="116" phylum="Arthropoda" rank="species" species="subfasciatus">
<emphasis id="B95F8A50E766FFB3FEF8FB1FFE1CFD1C" box="[333,505,1269,1295]" italics="true" pageId="3" pageNumber="116">H. subfasciatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>