758 lines
100 KiB
XML
758 lines
100 KiB
XML
<document id="99F81F874D178078F1C0BA2A523BF3ED" ID-DOI="10.11646/zootaxa.5556.1.19" ID-ISSN="1175-5326" ID-Zenodo-Dep="14595253" ID-ZooBank="B7D2B6BB-62A4-461A-BA26-55A96FA458BA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1735931024094" checkinUser="plazi" docAuthor="Kolesnikov, Vasiliy B., Klimov, Pavel B., Khaustov, Alexander A., Pepato, Almir & Makarova, Olga L." docDate="2024" docId="03B86276FFC8141F01DBFF6EFCE0F91F" docLanguage="en" docName="zootaxa.5556.1.19.pdf" docOrigin="Zootaxa 5556 (1)" docSource="https://doi.org/10.11646/zootaxa.5556.1.19" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.23:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="23" docTitle="Thyreophagus subiasi Kolesnikov, Klimov, Khaustov, Pepato & Makarova, 2024, sp. nov." docType="treatment" docVersion="4" lastPageNumber="260" masterDocId="FF811A0EFFCB1415014CFFB1FFBAF802" masterDocTitle="A new arenicolous species of the genus Thyreophagus (Acari: Acaridae) from the Caspian sea coast: Taxonomy and ecological insights" masterLastPageNumber="267" masterPageNumber="252" pageNumber="255" updateTime="1736011682392" updateUser="felipe" zenodo-license-document="CLOSED">
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<mods:title id="ABA74C4602187F9FAA3E1F5C9C8D3982">A new arenicolous species of the genus Thyreophagus (Acari: Acaridae) from the Caspian sea coast: Taxonomy and ecological insights</mods:title>
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<mods:roleTerm id="7157118D084DE96E58E11C6C92F45E23">Author</mods:roleTerm>
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<mods:namePart id="7B529FCDC7219B85AAAB44574732A1B8">Kolesnikov, Vasiliy B.</mods:namePart>
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<mods:affiliation id="B08D77DBFB068D12B45FA18C4C14FED3">Papanin Institute for Biology of Inland Waters, Russian Academy of Sciences, 152742 Borok, Yaroslavl Province, Russia. & Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, 625003 Tyumen, Russia.</mods:affiliation>
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</mods:name>
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<mods:name id="2C35D8F09674E5019C5D93340F8F5C88" type="personal">
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<mods:roleTerm id="8C4C9F587C2F12268309999847D81E2C">Author</mods:roleTerm>
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<mods:namePart id="77E88DB8902AACF624476EC575F954E6">Klimov, Pavel B.</mods:namePart>
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<mods:affiliation id="B0D8C54E62F6769CF819DE4311F2736C">Department of Biological Sciences, Purdue University, Lilly Hall of Life Sciences, West Lafayette, IN 47907, USA.</mods:affiliation>
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<mods:roleTerm id="3B8923D798300C2447A8B6742BAFE0A7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B3B8833A7359B96CAE148E14FD4A10B1">Khaustov, Alexander A.</mods:namePart>
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<mods:affiliation id="270704B69E35576C451377C39BBA17DA">Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, 625003 Tyumen, Russia.</mods:affiliation>
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</mods:name>
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<mods:namePart id="232C06E27EA63D44652AC37BF3A21A71">Pepato, Almir</mods:namePart>
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<mods:affiliation id="7A5BAAC859A6E6D05B1D855FCD8D84AB">Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, 625003 Tyumen, Russia. & Universidade Federal de Minas Gerais, Instituto de Ciências Biológicas, Departamento de Zoologia, Laboratório de Sistemática eEvolução de Ácaros Acariformes, Av. Antonio Carlos, 6627, Pampulha, Belo Horizonte-MG ZIP: 31270 - 901, Brazil.</mods:affiliation>
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<mods:roleTerm id="A201F8EE104E0194FB9DBDE54B3320CD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="44F4C86C77A9F7BC29193CBCD54FA131">Makarova, Olga L.</mods:namePart>
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<mods:affiliation id="1E865EA3D218FBF74B01599A37CDB140">Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Moscow, Russia.</mods:affiliation>
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<mods:typeOfResource id="955C1C9A35D12D8B22886013EC08A911">text</mods:typeOfResource>
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<mods:title id="A6B34F939CF84D9128923B954918C746">Zootaxa</mods:title>
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<mods:part id="15791DA8B5608A60EF14A6859C683C82">
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<mods:date id="7A179F8E969E305F7A4ED1CA9D67F96B">2024</mods:date>
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<mods:number id="B8AC48B376111C2AF46D2AB3E4EDE416">2024-12-23</mods:number>
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<mods:number id="7CAA85EDF0A170522F6BD6FF72A413DF">5556</mods:number>
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<mods:url id="CFCF8799EB08D289B5A41A37F5065053">https://doi.org/10.11646/zootaxa.5556.1.19</mods:url>
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<mods:classification id="09CB5673C8446A0E9EEA3DA8535BF01C">journal article</mods:classification>
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<mods:identifier id="900805C8E0711018C8F3A539BA2B8F3F" type="DOI">10.11646/zootaxa.5556.1.19</mods:identifier>
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<treatment id="03B86276FFC8141F01DBFF6EFCE0F91F" ID-DOI="http://doi.org/10.5281/zenodo.14596200" ID-Zenodo-Dep="14596200" LSID="urn:lsid:plazi:treatment:03B86276FFC8141F01DBFF6EFCE0F91F" httpUri="http://treatment.plazi.org/id/03B86276FFC8141F01DBFF6EFCE0F91F" lastPageId="10" lastPageNumber="260" pageId="3" pageNumber="255">
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<subSubSection id="C30B80EBFFC8141601DBFF6EFDBDF8FB" box="[151,519,223,250]" pageId="3" pageNumber="255" type="nomenclature">
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<paragraph id="8BAED360FFC8141601DBFF6EFDBDF8FB" blockId="3.[151,519,223,285]" box="[151,519,223,250]" pageId="3" pageNumber="255">
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<heading id="D0E6640CFFC8141601DBFF6EFDBDF8FB" bold="true" box="[151,519,223,250]" fontSize="11" level="1" pageId="3" pageNumber="255" reason="1">
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<emphasis id="B9650F72FFC8141601DBFF6EFDBDF8FB" bold="true" box="[151,519,223,250]" pageId="3" pageNumber="255">
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<taxonomicName id="4C11A8E3FFC8141601DBFF6EFE18F8F8" authorityName="Kolesnikov & Klimov & Khaustov & Pepato & Makarova" authorityYear="2024" box="[151,418,223,250]" class="Arachnida" family="Acaridae" genus="Thyreophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="255" phylum="Arthropoda" rank="species" species="subiasi" status="sp. nov.">
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<emphasis id="B9650F72FFC8141601DBFF6EFE18F8F8" bold="true" box="[151,418,223,250]" italics="true" pageId="3" pageNumber="255">Thyreophagus subiasi</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A256B209FFC8141600E5FF6EFDBDF8FB" box="[425,519,223,249]" pageId="3" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C30B80EBFFC8141601DBFEB2FEB1F91F" box="[151,267,259,285]" pageId="3" pageNumber="255" type="description">
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<paragraph id="8BAED360FFC8141601DBFEB2FEB1F91F" blockId="3.[151,519,223,285]" box="[151,267,259,285]" pageId="3" pageNumber="255">
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(
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||
<figureCitation id="132ACFE5FFC8141601D3FEB2FEBEF91F" box="[159,260,259,285]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStartId-0="4.[152,255,1957,1981]" captionStartId-1="5.[152,255,1625,1649]" captionStartId-2="6.[152,255,1875,1899]" captionStartId-3="7.[152,255,1060,1084]" captionStartId-4="8.[152,255,1798,1822]" captionStartId-5="9.[152,255,1943,1967]" captionTargetBox-0="[154,1432,384,1931]" captionTargetBox-1="[154,1432,184,1598]" captionTargetBox-2="[154,1432,184,1848]" captionTargetBox-3="[154,1432,184,1033]" captionTargetBox-4="[151,1436,181,1774]" captionTargetBox-5="[178,1409,181,1919]" captionTargetId-0="figure-115@4.[151,1435,381,1934]" captionTargetId-1="figure-19@5.[151,1435,181,1601]" captionTargetId-2="figure-19@6.[151,1435,181,1851]" captionTargetId-3="figure-19@7.[151,1435,181,1036]" captionTargetId-4="figure-19@8.[151,1436,181,1775]" captionTargetId-5="figure-19@9.[178,1409,181,1919]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionText-0="FIGURE 2. Thyreophagus subiasi sp. nov., female holotype: A—dorsal view; B—ventral view. Chaetotaxy of legs and palps is omitted." captionText-1="FIGURE 3. Thyreophagus subiasi sp. nov., male paratype: A—dorsal view; B—ventral view. Chaetotaxy of legs and palps is omitted." captionText-2="FIGURE 4. Thyreophagus subiasi sp. nov., female holotype:A—leg I, dorsal view; B—tarsus I, ventral view; C—leg II, dorsal view; D—tarsus II, ventral view; E—leg III, dorsal view; F—tarsus III, ventral view; G—leg IV, dorsal view; H—tarsus IV, ventral view." captionText-3="FIGURE 5. Thyreophagus subiasi sp. nov., female holotype (A, B) and male, paratype (C–G):A—subcapitulum, ventral view; B—spermatheca; C—male genitalia, ventral view; D—leg I, dorsal view; E—tarsus I, ventral view; F—leg IV, anterior view; G—tarsus IV, posterior view." captionText-4="FIGURE 6. Thyreophagus subiasi sp. nov., female holotype, DIC images: A—dorsal view; B—ventral view." captionText-5="FIGURE 7. Thyreophagus subiasi sp. nov., male paratype (A–C, E, F) and female, holotype (D), DIC images:A—dorsal view; B—ventral view; C—prodorsal shield; D—spermatheca; E—posterior part of opisthosoma, dorsal view; G—posterior part of opisthosoma, ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.14595257" figureDoi-1="http://doi.org/10.5281/zenodo.14595259" figureDoi-2="http://doi.org/10.5281/zenodo.14595261" figureDoi-3="http://doi.org/10.5281/zenodo.14595265" figureDoi-4="http://doi.org/10.5281/zenodo.14595267" figureDoi-5="http://doi.org/10.5281/zenodo.14595271" httpUri-0="https://zenodo.org/record/14595257/files/figure.png" httpUri-1="https://zenodo.org/record/14595259/files/figure.png" httpUri-2="https://zenodo.org/record/14595261/files/figure.png" httpUri-3="https://zenodo.org/record/14595265/files/figure.png" httpUri-4="https://zenodo.org/record/14595267/files/figure.png" httpUri-5="https://zenodo.org/record/14595271/files/figure.png" pageId="3" pageNumber="255">Figs 2–7</figureCitation>
|
||
)
|
||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30B80EBFFC8141601DBFEFDFCD5FA63" pageId="3" pageNumber="255" type="materials_examined">
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<paragraph id="8BAED360FFC8141601DBFEFDFDB3F9AF" blockId="3.[151,1437,331,2020]" pageId="3" pageNumber="255">
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<emphasis id="B9650F72FFC8141601DBFEFDFE88F967" box="[151,306,331,357]" italics="true" pageId="3" pageNumber="255">Type material</emphasis>
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:
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||
<materialsCitation id="3B79D93DFFC81416000DFEFAFDB0F967" box="[321,522,331,357]" collectedFrom="shore debris (decomposing land grasses, driftwood)" collectingDate="2021-04-04" collectorName="O. Makarova" country="Russia" latitude="41.87636" location="sandy beach" longLatPrecision="1" longitude="48.548897" municipality="Samur Forest" pageId="3" pageNumber="255" specimenCount="1" specimenCount-female="1" stateProvince="Dagestan" typeStatus="holotype">
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<typeStatus id="54AA6DC2FFC81416000DFEFAFE19F967" box="[321,419,331,357]" pageId="3" pageNumber="255" type="holotype">holotype</typeStatus>
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(
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<specimenCount id="9D1718E9FFC8141600FEFEFAFDB8F967" box="[434,514,331,357]" pageId="3" pageNumber="255" type="female">female</specimenCount>
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)
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</materialsCitation>
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and
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||
<materialsCitation id="3B79D93DFFC81416030DFEFDFDB3F9AF" collectedFrom="shore debris (decomposing land grasses, driftwood)" collectingDate="2021-04-04" collectionCode="ZISP" collectorName="O. Makarova" country="Russia" latitude="41.87636" location="sandy beach" longLatPrecision="1" longitude="48.548897" municipality="Samur Forest" pageId="3" pageNumber="255" specimenCode="ZISP-8446" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Dagestan" typeStatus="paratype">
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||
<typeStatus id="54AA6DC2FFC81416030DFEFDFD16F964" box="[577,684,332,358]" pageId="3" pageNumber="255" type="paratype">paratypes</typeStatus>
|
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(
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<specimenCount id="9D1718E9FFC8141603F7FEFAFCFFF967" box="[699,837,331,357]" count="2" pageId="3" pageNumber="255" type="female">two females</specimenCount>
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and
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<specimenCount id="9D1718E9FFC814160231FEFDFC4AF967" box="[893,1008,331,358]" count="2" pageId="3" pageNumber="255" type="male">two males</specimenCount>
|
||
)—
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<collectingCountry id="F30693F0FFC81416055AFEFAFBC7F967" box="[1046,1149,331,357]" name="Russia" pageId="3" pageNumber="255">RUSSIA</collectingCountry>
|
||
:
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||
<collectingRegion id="49D51D82FFC8141605C4FEFDFB49F964" box="[1160,1267,332,358]" country="Russia" name="Dagestan" pageId="3" pageNumber="255">Dagestan</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BCA491AFFC8141605B3FEFAFA22F964" box="[1279,1432,331,358]" country="Russia" latitude="41.87636" longLatPrecision="1" longitude="48.548897" name="Samur Forest" pageId="3" pageNumber="255" stateProvince="Dagestan">Samur Forest</collectingMunicipality>
|
||
,
|
||
<location id="8ECE85BBFFC8141601DBFEDEFE9EF98B" LSID="urn:lsid:plazi:treatment:03B86276FFC8141F01DBFF6EFCE0F91F:8ECE85BBFFC8141601DBFEDEFE9EF98B" box="[151,292,367,393]" country="Russia" latitude="41.87636" longLatPrecision="1" longitude="48.548897" municipality="Samur Forest" name="sandy beach" pageId="3" pageNumber="255" stateProvince="Dagestan">sandy beach</location>
|
||
,
|
||
<collectedFrom id="380D909DFFC81416007CFEDEFCC2F98B" box="[304,888,367,394]" pageId="3" pageNumber="255">shore debris (decomposing land grasses, driftwood)</collectedFrom>
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,
|
||
<geoCoordinate id="EE25B5A7FFC8141602C8FEDEFB9EF98B" box="[900,1060,367,393]" degrees="41.876362" direction="north" orientation="latitude" pageId="3" pageNumber="255" precision="1" value="41.87636">41.876362 °N</geoCoordinate>
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,
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<geoCoordinate id="EE25B5A7FFC81416057CFEDEFB7CF988" box="[1072,1222,367,394]" degrees="48.548897" direction="east" orientation="longitude" pageId="3" pageNumber="255" precision="1" value="48.548897">48.548897°E</geoCoordinate>
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,
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<date id="FFAFF5A0FFC81416059EFEDEFADDF98B" box="[1234,1383,367,393]" pageId="3" pageNumber="255" value="2021-04-04">
|
||
<collectingDate id="EFEB0C48FFC81416059EFEDEFADDF98B" box="[1234,1383,367,393]" pageId="3" pageNumber="255" value="2021-04-04">04 Apr. 2021</collectingDate>
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||
</date>
|
||
, leg.
|
||
<collectorName id="26E4B6B6FFC8141601DBFE22FE91F9AF" box="[151,299,403,429]" pageId="3" pageNumber="255">O. Makarova</collectorName>
|
||
(Hoyer’s medium).
|
||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8BAED360FFC81416018BFE09FD1AFA1B" blockId="3.[151,1437,331,2020]" pageId="3" pageNumber="255">
|
||
<emphasis id="B9650F72FFC81416018BFE09FE28F9D3" box="[199,402,439,465]" italics="true" pageId="3" pageNumber="255">Non-type material</emphasis>
|
||
(specimens not mounted)—
|
||
<materialsCitation id="3B79D93DFFC814160381FE06FD26FA1B" latitude="41.881485" longitude="48.588768" pageId="3" pageNumber="255">
|
||
<collectingCountry id="F30693F0FFC814160381FE06FC88F9D3" box="[717,818,439,465]" name="Russia" pageId="3" pageNumber="255">RUSSIA</collectingCountry>
|
||
:
|
||
<collectingRegion id="49D51D82FFC814160276FE09FC1FF9D0" box="[826,933,440,466]" country="Russia" name="Dagestan" pageId="3" pageNumber="255">Dagestan</collectingRegion>
|
||
,
|
||
<location id="8ECE85BBFFC8141602E3FE06FBFCF9D0" LSID="urn:lsid:plazi:treatment:03B86276FFC8141F01DBFF6EFCE0F91F:8ECE85BBFFC8141602E3FE06FBFCF9D0" box="[943,1094,439,466]" country="Russia" latitude="41.881485" longLatPrecision="1" longitude="48.53611" name="Samur Forest" pageId="3" pageNumber="255" stateProvince="Dagestan">Samur Forest</location>
|
||
, sandy beach, in stands of reed, cattails and blackberry bushes,
|
||
<geoCoordinate id="EE25B5A7FFC8141600BEFE6AFD3DF9F4" box="[498,647,475,502]" degrees="41.844918" direction="north" orientation="latitude" pageId="3" pageNumber="255" precision="1" value="41.844917">41.844918 N</geoCoordinate>
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,
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<geoCoordinate id="EE25B5A7FFC8141603DDFE6AFC98F9F4" box="[657,802,475,502]" degrees="48.588768" direction="east" orientation="longitude" pageId="3" pageNumber="255" precision="1" value="48.588768">48.588768 E</geoCoordinate>
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;
|
||
<geoCoordinate id="EE25B5A7FFC814160261FE6AFC7BF9F4" box="[813,961,475,502]" degrees="41.844460" direction="north" orientation="latitude" pageId="3" pageNumber="255" precision="1" value="41.84446">41.844460 N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE25B5A7FFC814160287FE6AFBE7F9F4" box="[971,1117,475,502]" degrees="48.589462" direction="east" orientation="longitude" pageId="3" pageNumber="255" precision="1" value="48.589462">48.589462 E</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="EE25B5A7FFC814160524FE6AFB46F9F4" box="[1128,1276,475,502]" degrees="41.854107" direction="north" orientation="latitude" pageId="3" pageNumber="255" precision="1" value="41.854107">41.854107 N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE25B5A7FFC81416044AFE6AFA2DF9F4" box="[1286,1431,475,502]" degrees="48.571503" direction="east" orientation="longitude" pageId="3" pageNumber="255" precision="1" value="48.571503">48.571503 E</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="EE25B5A7FFC8141601DBFE4EFE96FA18" box="[151,300,511,538]" degrees="41.881485" direction="north" orientation="latitude" pageId="3" pageNumber="255" precision="1" value="41.881485">41.881485 N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE25B5A7FFC81416007BFE4EFE72FA18" box="[311,456,511,538]" degrees="48.536109" direction="east" orientation="longitude" pageId="3" pageNumber="255" precision="1" value="48.53611">48.536109 E</geoCoordinate>
|
||
, leg. O. Makarova
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAED360FFC81416018BFD95FCD5FA63" blockId="3.[151,1437,331,2020]" pageId="3" pageNumber="255">
|
||
<emphasis id="B9650F72FFC81416018BFD95FEF9FA3F" box="[199,323,548,573]" italics="true" pageId="3" pageNumber="255">Depository</emphasis>
|
||
. The slide-mounted
|
||
<typeStatus id="54AA6DC2FFC814160365FD92FD31FA3F" box="[553,651,547,573]" pageId="3" pageNumber="255" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="54AA6DC2FFC8141603F3FD95FC90FA3C" box="[703,810,548,574]" pageId="3" pageNumber="255" type="paratype">paratypes</typeStatus>
|
||
(
|
||
<specimenCode id="DBB77B1BFFC814160275FD92FC0DFA3F" box="[825,951,547,573]" pageId="3" pageNumber="255">ZISP-8446</specimenCode>
|
||
) are deposited in the
|
||
<location id="8ECE85BBFFC8141605EFFD92FAC5FA3F" LSID="urn:lsid:plazi:treatment:03B86276FFC8141F01DBFF6EFCE0F91F:8ECE85BBFFC8141605EFFD92FAC5FA3F" box="[1187,1407,547,573]" country="Russia" municipality="Russian Academy of Sciences" name="Zoological Institute" pageId="3" pageNumber="255" stateProvince="Saint Petersburg">Zoological Institute</location>
|
||
of the
|
||
<collectingMunicipality id="6BCA491AFFC81416018CFDF6FDAEFA63" box="[192,532,583,609]" pageId="3" pageNumber="255">Russian Academy of Sciences</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49D51D82FFC814160353FDF6FD04FA63" box="[543,702,583,609]" country="Russia" name="Saint Petersburg" pageId="3" pageNumber="255">St. Petersburg</collectingRegion>
|
||
,
|
||
<collectingCountry id="F30693F0FFC814160386FDF6FCAFFA63" box="[714,789,583,609]" name="Russia" pageId="3" pageNumber="255">Russia</collectingCountry>
|
||
(
|
||
<collectionCode id="ED004BA5FFC81416026FFDF6FCD8FA63" box="[803,866,583,609]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="3" pageNumber="255">ZISP</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30B80EBFFC81416018BFDDDFDA4FB14" pageId="3" pageNumber="255" type="diagnosis">
|
||
<paragraph id="8BAED360FFC81416018BFDDDFDA4FB14" blockId="3.[151,1437,331,2020]" pageId="3" pageNumber="255">
|
||
<emphasis id="B9650F72FFC81416018BFDDDFE83FA87" box="[199,313,620,645]" italics="true" pageId="3" pageNumber="255">Diagnosis</emphasis>
|
||
. Prodorsal shield with small flat protrus in posterior part, smoothly punctate except lineate pattern on posterior 1/3 part of shield. Supracoxal seta
|
||
<emphasis id="B9650F72FFC8141603C0FD3EFD01FAAB" box="[652,699,655,681]" italics="true" pageId="3" pageNumber="255">elcp</emphasis>
|
||
present. Idiosomal setae
|
||
<emphasis id="B9650F72FFC814160296FD3EFC52FAAB" box="[986,1000,655,681]" italics="true" pageId="3" pageNumber="255">h</emphasis>
|
||
<subScript id="1795D125FFC8141602A4FD13FC4AFAB2" attach="left" box="[1000,1008,674,688]" fontSize="6" pageId="3" pageNumber="255">1</subScript>
|
||
,
|
||
<emphasis id="B9650F72FFC8141602B3FD3EFBB7FAAB" box="[1023,1037,655,681]" italics="true" pageId="3" pageNumber="255">h</emphasis>
|
||
<subScript id="1795D125FFC814160541FD13FBAFFAB2" attach="left" box="[1037,1045,674,688]" fontSize="6" pageId="3" pageNumber="255">2</subScript>
|
||
and
|
||
<emphasis id="B9650F72FFC814160502FD3EFBE6FAAB" box="[1102,1116,655,681]" italics="true" pageId="3" pageNumber="255">h</emphasis>
|
||
<subScript id="1795D125FFC814160510FD13FBDEFAB2" attach="left" box="[1116,1124,674,688]" fontSize="6" pageId="3" pageNumber="255">3</subScript>
|
||
short. Base of spermatheca narrow, its most part is hidden in the spermathecal sac, surrounding the base of spermatheca in a “bottleneck” shape with an extended ring at the top. Paired efferent ducts short, their length approximately 1/3 the length of spermatheca base, with short stem.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30B80EBFFC81412018BFC91FD81FE36" lastPageId="7" lastPageNumber="259" pageId="3" pageNumber="255" type="description">
|
||
<paragraph id="8BAED360FFC81416018BFC91FEE8FB38" blockId="3.[151,1437,331,2020]" box="[199,338,800,826]" pageId="3" pageNumber="255">
|
||
<emphasis id="B9650F72FFC81416018BFC91FEF1FB3B" box="[199,331,800,825]" italics="true" pageId="3" pageNumber="255">Description</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAED360FFC81416018BFCF2FE00FB83" blockId="3.[151,1437,331,2020]" pageId="3" pageNumber="255">
|
||
Female (
|
||
<figureCitation id="132ACFE5FFC814160067FCF2FECDFB5F" box="[299,375,835,861]" captionStart="FIGURE 2" captionStartId="4.[152,255,1957,1981]" captionTargetBox="[154,1432,384,1931]" captionTargetId="figure-115@4.[151,1435,381,1934]" captionTargetPageId="4" captionText="FIGURE 2. Thyreophagus subiasi sp. nov., female holotype: A—dorsal view; B—ventral view. Chaetotaxy of legs and palps is omitted." figureDoi="http://doi.org/10.5281/zenodo.14595257" httpUri="https://zenodo.org/record/14595257/files/figure.png" pageId="3" pageNumber="255">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="132ACFE5FFC8141600C9FCF2FE2FFB5F" box="[389,405,835,861]" captionStart="FIGURE 4" captionStartId="6.[152,255,1875,1899]" captionTargetBox="[154,1432,184,1848]" captionTargetId="figure-19@6.[151,1435,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Thyreophagus subiasi sp. nov., female holotype:A—leg I, dorsal view; B—tarsus I, ventral view; C—leg II, dorsal view; D—tarsus II, ventral view; E—leg III, dorsal view; F—tarsus III, ventral view; G—leg IV, dorsal view; H—tarsus IV, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14595261" httpUri="https://zenodo.org/record/14595261/files/figure.png" pageId="3" pageNumber="255">4</figureCitation>
|
||
,
|
||
<figureCitation id="132ACFE5FFC8141600E8FCF2FE56FB5C" box="[420,492,835,862]" captionStart="FIGURE 5" captionStartId="7.[152,255,1060,1084]" captionTargetBox="[154,1432,184,1033]" captionTargetId="figure-19@7.[151,1435,181,1036]" captionTargetPageId="7" captionText="FIGURE 5. Thyreophagus subiasi sp. nov., female holotype (A, B) and male, paratype (C–G):A—subcapitulum, ventral view; B—spermatheca; C—male genitalia, ventral view; D—leg I, dorsal view; E—tarsus I, ventral view; F—leg IV, anterior view; G—tarsus IV, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14595265" httpUri="https://zenodo.org/record/14595265/files/figure.png" pageId="3" pageNumber="255">5A, B</figureCitation>
|
||
,
|
||
<figureCitation id="132ACFE5FFC8141600B6FCF2FDB0FB5F" box="[506,522,835,861]" captionStart="FIGURE 6" captionStartId="8.[152,255,1798,1822]" captionTargetBox="[151,1436,181,1774]" captionTargetId="figure-19@8.[151,1436,181,1775]" captionTargetPageId="8" captionText="FIGURE 6. Thyreophagus subiasi sp. nov., female holotype, DIC images: A—dorsal view; B—ventral view." figureDoi="http://doi.org/10.5281/zenodo.14595267" httpUri="https://zenodo.org/record/14595267/files/figure.png" pageId="3" pageNumber="255">6</figureCitation>
|
||
,
|
||
<figureCitation id="132ACFE5FFC814160355FCF2FDD8FB5F" box="[537,610,835,861]" captionStart="FIGURE 7" captionStartId="9.[152,255,1943,1967]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure-19@9.[178,1409,181,1919]" captionTargetPageId="9" captionText="FIGURE 7. Thyreophagus subiasi sp. nov., male paratype (A–C, E, F) and female, holotype (D), DIC images:A—dorsal view; B—ventral view; C—prodorsal shield; D—spermatheca; E—posterior part of opisthosoma, dorsal view; G—posterior part of opisthosoma, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14595271" httpUri="https://zenodo.org/record/14595271/files/figure.png" pageId="3" pageNumber="255">7C, D</figureCitation>
|
||
). Idiosoma elongate, 430 × 220 (
|
||
<typeStatus id="54AA6DC2FFC8141602AFFCF2FBF1FB5F" box="[995,1099,835,861]" pageId="3" pageNumber="255" type="holotype">holotype</typeStatus>
|
||
), 1.9 times longer than wide. Idiosomal cuticle smooth.
|
||
</paragraph>
|
||
<paragraph id="8BAED360FFC81416018BFC3AFB8FFBEC" blockId="3.[151,1437,331,2020]" pageId="3" pageNumber="255">
|
||
Subcapitular setae (
|
||
<emphasis id="B9650F72FFC8141600E6FC3AFE02FBA7" box="[426,440,907,933]" italics="true" pageId="3" pageNumber="255">h</emphasis>
|
||
) setiform; palp tibial setae (
|
||
<emphasis id="B9650F72FFC814160249FC3DFCA9FBA7" box="[773,787,908,933]" italics="true" pageId="3" pageNumber="255">a</emphasis>
|
||
) lateral, dorsal palp tibial setae (
|
||
<emphasis id="B9650F72FFC8141605D9FC3DFB06FBA7" box="[1173,1212,908,933]" italics="true" pageId="3" pageNumber="255">sup</emphasis>
|
||
) dorsal, palp tarsal seta (
|
||
<emphasis id="B9650F72FFC814160199FC01FF4CFBCB" box="[213,246,944,969]" italics="true" pageId="3" pageNumber="255">cm</emphasis>
|
||
) filiform; supracoxal seta
|
||
<emphasis id="B9650F72FFC81416036BFC1EFDECFBCB" box="[551,598,943,969]" italics="true" pageId="3" pageNumber="255">elcp</emphasis>
|
||
present; terminal palp tarsal solenidion ω short; external part of terminal eupathidium
|
||
<emphasis id="B9650F72FFC814160066FC62FEEAFBEF" box="[298,336,979,1005]" italics="true" pageId="3" pageNumber="255">ul”</emphasis>
|
||
dome-shaped; terminal eupathidium
|
||
<emphasis id="B9650F72FFC8141603BEFC62FCABFBEF" box="[754,785,979,1005]" italics="true" pageId="3" pageNumber="255">ul’</emphasis>
|
||
small, rounded (
|
||
<figureCitation id="132ACFE5FFC814160282FC62FB92FBEC" box="[974,1064,979,1006]" captionStart="FIGURE 5" captionStartId="7.[152,255,1060,1084]" captionTargetBox="[154,1432,184,1033]" captionTargetId="figure-19@7.[151,1435,181,1036]" captionTargetPageId="7" captionText="FIGURE 5. Thyreophagus subiasi sp. nov., female holotype (A, B) and male, paratype (C–G):A—subcapitulum, ventral view; B—spermatheca; C—male genitalia, ventral view; D—leg I, dorsal view; E—tarsus I, ventral view; F—leg IV, anterior view; G—tarsus IV, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14595265" httpUri="https://zenodo.org/record/14595265/files/figure.png" pageId="3" pageNumber="255">Fig. 5A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAED360FFC81416018BFC46FD09FE50" blockId="3.[151,1437,331,2020]" pageId="3" pageNumber="255">
|
||
Prodorsal shield (
|
||
<figureCitation id="132ACFE5FFC8141600C7FC46FE5DFC13" box="[395,487,1015,1041]" captionStart="FIGURE 2" captionStartId="4.[152,255,1957,1981]" captionTargetBox="[154,1432,384,1931]" captionTargetId="figure-115@4.[151,1435,381,1934]" captionTargetPageId="4" captionText="FIGURE 2. Thyreophagus subiasi sp. nov., female holotype: A—dorsal view; B—ventral view. Chaetotaxy of legs and palps is omitted." figureDoi="http://doi.org/10.5281/zenodo.14595257" httpUri="https://zenodo.org/record/14595257/files/figure.png" pageId="3" pageNumber="255">Figs 2A</figureCitation>
|
||
,
|
||
<figureCitation id="132ACFE5FFC8141600BFFC46FDAFFC13" box="[499,533,1015,1041]" captionStart="FIGURE 7" captionStartId="9.[152,255,1943,1967]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure-19@9.[178,1409,181,1919]" captionTargetPageId="9" captionText="FIGURE 7. Thyreophagus subiasi sp. nov., male paratype (A–C, E, F) and female, holotype (D), DIC images:A—dorsal view; B—ventral view; C—prodorsal shield; D—spermatheca; E—posterior part of opisthosoma, dorsal view; G—posterior part of opisthosoma, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14595271" httpUri="https://zenodo.org/record/14595271/files/figure.png" pageId="3" pageNumber="255">7C</figureCitation>
|
||
) 100 long, 82 wide, 1.2 times longer than wide, with setae
|
||
<emphasis id="B9650F72FFC8141605E4FC49FB06FC13" box="[1192,1212,1016,1041]" italics="true" pageId="3" pageNumber="255">vi</emphasis>
|
||
(situated at anterior part of shield, alveoli separated at a distance less than their width), rounded anterolateral incisions, and elongate midlateral incisions (insertion points of setae
|
||
<emphasis id="B9650F72FFC8141603D0FBF1FD0FFC5B" box="[668,693,1088,1113]" italics="true" pageId="3" pageNumber="255">ve</emphasis>
|
||
). The back of the shield has a small flat protrus. Prodorsal shield smoothly punctate except lineate pattern on posterior 1/3 part of shield. Grandjean’s organ (GO) with 14 membranous short finger-shaped processes. Supracoxal setae (
|
||
<emphasis id="B9650F72FFC8141603F7FB39FD65FCA3" box="[699,735,1160,1185]" italics="true" pageId="3" pageNumber="255">scx</emphasis>
|
||
) smooth, sword-shaped, widened and flattened, tapering at tip. Idiosomal setae (
|
||
<emphasis id="B9650F72FFC81416001AFB1DFED0FCC7" box="[342,362,1196,1221]" italics="true" pageId="3" pageNumber="255">vi</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC814160034FB1DFE35FCC7" box="[376,399,1196,1221]" italics="true" pageId="3" pageNumber="255">se</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC8141600D1FB1DFE13FCC7" box="[413,425,1196,1221]" italics="true" pageId="3" pageNumber="255">c</emphasis>
|
||
<subScript id="1795D125FFC8141600E5FB0FFE0BFCCE" attach="left" box="[425,433,1214,1228]" fontSize="6" pageId="3" pageNumber="255">p</subScript>
|
||
,
|
||
<emphasis id="B9650F72FFC8141600F3FB1AFE77FCC7" box="[447,461,1195,1221]" italics="true" pageId="3" pageNumber="255">d</emphasis>
|
||
<subScript id="1795D125FFC814160081FB0FFE6FFCCE" attach="left" box="[461,469,1214,1228]" fontSize="6" pageId="3" pageNumber="255">2</subScript>
|
||
,
|
||
<emphasis id="B9650F72FFC8141600AFFB1DFE55FCC7" box="[483,495,1196,1221]" italics="true" pageId="3" pageNumber="255">e</emphasis>
|
||
<subScript id="1795D125FFC8141600A3FB0FFE4DFCCE" attach="left" box="[495,503,1214,1228]" fontSize="6" pageId="3" pageNumber="255">2</subScript>
|
||
,
|
||
<emphasis id="B9650F72FFC814160349FB1AFDA9FCC7" box="[517,531,1195,1221]" italics="true" pageId="3" pageNumber="255">h</emphasis>
|
||
<subScript id="1795D125FFC81416035FFB0FFDA1FCCE" attach="left" box="[531,539,1214,1228]" fontSize="6" pageId="3" pageNumber="255">1</subScript>
|
||
,
|
||
<emphasis id="B9650F72FFC814160365FB1AFD8DFCC7" box="[553,567,1195,1221]" italics="true" pageId="3" pageNumber="255">h</emphasis>
|
||
<subScript id="1795D125FFC81416037BFB0FFD85FCCE" attach="left" box="[567,575,1214,1228]" fontSize="6" pageId="3" pageNumber="255">2</subScript>
|
||
,
|
||
<emphasis id="B9650F72FFC814160300FB1AFDE0FCC7" box="[588,602,1195,1221]" italics="true" pageId="3" pageNumber="255">h</emphasis>
|
||
<subScript id="1795D125FFC814160316FB0FFDD8FCCE" attach="left" box="[602,610,1214,1228]" fontSize="6" pageId="3" pageNumber="255">3</subScript>
|
||
,
|
||
<emphasis id="B9650F72FFC81416033CFB1DFD33FCC7" box="[624,649,1196,1221]" italics="true" pageId="3" pageNumber="255">ps</emphasis>
|
||
<subScript id="1795D125FFC8141603C5FB0FFD2BFCCE" attach="left" box="[649,657,1214,1228]" fontSize="6" pageId="3" pageNumber="255">3</subScript>
|
||
) smooth (
|
||
<figureCitation id="132ACFE5FFC81416024EFB1AFCF0FCC7" box="[770,842,1195,1221]" captionStart="FIGURE 2" captionStartId="4.[152,255,1957,1981]" captionTargetBox="[154,1432,384,1931]" captionTargetId="figure-115@4.[151,1435,381,1934]" captionTargetPageId="4" captionText="FIGURE 2. Thyreophagus subiasi sp. nov., female holotype: A—dorsal view; B—ventral view. Chaetotaxy of legs and palps is omitted." figureDoi="http://doi.org/10.5281/zenodo.14595257" httpUri="https://zenodo.org/record/14595257/files/figure.png" pageId="3" pageNumber="255">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="132ACFE5FFC81416021AFB1AFCDCFCC7" box="[854,870,1195,1221]" captionStart="FIGURE 6" captionStartId="8.[152,255,1798,1822]" captionTargetBox="[151,1436,181,1774]" captionTargetId="figure-19@8.[151,1436,181,1775]" captionTargetPageId="8" captionText="FIGURE 6. Thyreophagus subiasi sp. nov., female holotype, DIC images: A—dorsal view; B—ventral view." figureDoi="http://doi.org/10.5281/zenodo.14595267" httpUri="https://zenodo.org/record/14595267/files/figure.png" pageId="3" pageNumber="255">6</figureCitation>
|
||
), filiform and short (except long
|
||
<emphasis id="B9650F72FFC814160594FB1DFB55FCC7" box="[1240,1263,1196,1221]" italics="true" pageId="3" pageNumber="255">se</emphasis>
|
||
); opisthosomal gland openings slightly anteriad setal bases
|
||
<emphasis id="B9650F72FFC8141603C1FB61FD23FCEB" box="[653,665,1232,1257]" italics="true" pageId="3" pageNumber="255">e</emphasis>
|
||
<subScript id="1795D125FFC8141603D5FB53FD1BFCF2" attach="left" box="[665,673,1250,1264]" fontSize="6" pageId="3" pageNumber="255">2</subScript>
|
||
. Three pairs of fundamental cupules (
|
||
<emphasis id="B9650F72FFC81416051AFB61FBD6FCEB" box="[1110,1132,1232,1257]" italics="true" pageId="3" pageNumber="255">ia</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC814160530FB61FB22FCEB" box="[1148,1176,1232,1257]" italics="true" pageId="3" pageNumber="255">im</emphasis>
|
||
and
|
||
<emphasis id="B9650F72FFC81416059EFB7EFB52FCEB" box="[1234,1256,1231,1257]" italics="true" pageId="3" pageNumber="255">ih</emphasis>
|
||
) present,
|
||
<emphasis id="B9650F72FFC814160416FB61FACAFCEB" box="[1370,1392,1232,1257]" italics="true" pageId="3" pageNumber="255">ip</emphasis>
|
||
not observed (
|
||
<figureCitation id="132ACFE5FFC814160042FB42FEECFD0F" box="[270,342,1267,1293]" captionStart="FIGURE 2" captionStartId="4.[152,255,1957,1981]" captionTargetBox="[154,1432,384,1931]" captionTargetId="figure-115@4.[151,1435,381,1934]" captionTargetPageId="4" captionText="FIGURE 2. Thyreophagus subiasi sp. nov., female holotype: A—dorsal view; B—ventral view. Chaetotaxy of legs and palps is omitted." figureDoi="http://doi.org/10.5281/zenodo.14595257" httpUri="https://zenodo.org/record/14595257/files/figure.png" pageId="3" pageNumber="255">Fig. 2</figureCitation>
|
||
). Ventral idiosoma (
|
||
<figureCitation id="132ACFE5FFC814160373FB42FD2DFD0F" box="[575,663,1267,1293]" captionStart="FIGURE 2" captionStartId="4.[152,255,1957,1981]" captionTargetBox="[154,1432,384,1931]" captionTargetId="figure-115@4.[151,1435,381,1934]" captionTargetPageId="4" captionText="FIGURE 2. Thyreophagus subiasi sp. nov., female holotype: A—dorsal view; B—ventral view. Chaetotaxy of legs and palps is omitted." figureDoi="http://doi.org/10.5281/zenodo.14595257" httpUri="https://zenodo.org/record/14595257/files/figure.png" pageId="3" pageNumber="255">Fig. 2B</figureCitation>
|
||
) with four pairs of coxal setae (
|
||
<emphasis id="B9650F72FFC814160541FB42FB90FD0F" box="[1037,1066,1267,1293]" italics="true" pageId="3" pageNumber="255">1a</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC814160576FB42FBECFD0F" box="[1082,1110,1267,1293]" italics="true" pageId="3" pageNumber="255">3a</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC81416052AFB42FB39FD0F" box="[1126,1155,1267,1293]" italics="true" pageId="3" pageNumber="255">4a</emphasis>
|
||
and
|
||
<emphasis id="B9650F72FFC8141605F2FB42FB60FD0F" box="[1214,1242,1267,1293]" italics="true" pageId="3" pageNumber="255">4b</emphasis>
|
||
) and one pair of genital setae (
|
||
<emphasis id="B9650F72FFC814160078FAA9FEF8FD33" box="[308,322,1304,1329]" italics="true" pageId="3" pageNumber="255">g</emphasis>
|
||
). Shape of coxal sclerites as in
|
||
<figureCitation id="132ACFE5FFC8141603E4FAA6FC9EFD33" box="[680,804,1303,1329]" captionStart="FIGURE 1" captionStartId="2.[152,255,1888,1912]" captionTargetBox="[151,1436,417,1865]" captionTargetId="figure-103@2.[151,1436,417,1865]" captionTargetPageId="2" captionText="FIGURE 1. Habitats of Thyreophagus subiasi sp. nov. on the shore of the protected Samur Forest: A—general view of a pebble beach; B—pebble beach with Typha laxmannii; C—sandy beach with Phragmites australis and Typha australis; D— reed thicket (Phragmites australis, Rubus caesius, Elaeagnus angustifolia) near abandoned fish ponds; E—driftwood and sea debris; F—grassy sea debris (Dagestan, Caspian Sea, April 2021)." figureDoi="http://doi.org/10.5281/zenodo.14595255" httpUri="https://zenodo.org/record/14595255/files/figure.png" pageId="3" pageNumber="255">Figures 1B</figureCitation>
|
||
and
|
||
<figureCitation id="132ACFE5FFC814160217FAA6FCC5FD30" box="[859,895,1303,1330]" captionStart="FIGURE 5" captionStartId="7.[152,255,1060,1084]" captionTargetBox="[154,1432,184,1033]" captionTargetId="figure-19@7.[151,1435,181,1036]" captionTargetPageId="7" captionText="FIGURE 5. Thyreophagus subiasi sp. nov., female holotype (A, B) and male, paratype (C–G):A—subcapitulum, ventral view; B—spermatheca; C—male genitalia, ventral view; D—leg I, dorsal view; E—tarsus I, ventral view; F—leg IV, anterior view; G—tarsus IV, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14595265" httpUri="https://zenodo.org/record/14595265/files/figure.png" pageId="3" pageNumber="255">5B</figureCitation>
|
||
. Genital region situated between coxal fields III and IV; genital valves form an inverted Y; epigynal and medial apodemes well-developed (
|
||
<figureCitation id="132ACFE5FFC8141605E9FA8AFB44FD57" box="[1189,1278,1339,1365]" captionStart="FIGURE 2" captionStartId="4.[152,255,1957,1981]" captionTargetBox="[154,1432,384,1931]" captionTargetId="figure-115@4.[151,1435,381,1934]" captionTargetPageId="4" captionText="FIGURE 2. Thyreophagus subiasi sp. nov., female holotype: A—dorsal view; B—ventral view. Chaetotaxy of legs and palps is omitted." figureDoi="http://doi.org/10.5281/zenodo.14595257" httpUri="https://zenodo.org/record/14595257/files/figure.png" pageId="3" pageNumber="255">Fig. 2B</figureCitation>
|
||
). Diameter of genital papillae approximately 0.3–0.4 the length of genital setae. Anal opening terminal. Copulatory tube small. Inseminatory canal (
|
||
<emphasis id="B9650F72FFC8141600CCFA35FE2EFD9F" box="[384,404,1412,1437]" italics="true" pageId="3" pageNumber="255">ic</emphasis>
|
||
) of spermatheca long, slender tube-like, uniformly wide. Atrium of spermatheca (
|
||
<emphasis id="B9650F72FFC81416047BFA32FAEAFD9F" box="[1335,1360,1411,1437]" italics="true" pageId="3" pageNumber="255">hs</emphasis>
|
||
) vase-shaped, narrow, with thin walls, slightly widened at the junction with the inseminatory canal, distinctly widened at point of connection with spermatheca, the length of the base of spermatheca is 2.5 times longer than its width in the middle part. Atrium is enveloped by spermatheca, forming a short bottleneck with extended rims (flared bottleneck) (
|
||
<figureCitation id="132ACFE5FFC8141601D3F9A2FF40FE2C" box="[159,250,1555,1582]" captionStart="FIGURE 5" captionStartId="7.[152,255,1060,1084]" captionTargetBox="[154,1432,184,1033]" captionTargetId="figure-19@7.[151,1435,181,1036]" captionTargetPageId="7" captionText="FIGURE 5. Thyreophagus subiasi sp. nov., female holotype (A, B) and male, paratype (C–G):A—subcapitulum, ventral view; B—spermatheca; C—male genitalia, ventral view; D—leg I, dorsal view; E—tarsus I, ventral view; F—leg IV, anterior view; G—tarsus IV, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14595265" httpUri="https://zenodo.org/record/14595265/files/figure.png" pageId="3" pageNumber="255">Figs 5B</figureCitation>
|
||
,
|
||
<figureCitation id="132ACFE5FFC81416004AF9A2FE96FE2F" box="[262,300,1555,1581]" captionStart="FIGURE 7" captionStartId="9.[152,255,1943,1967]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure-19@9.[178,1409,181,1919]" captionTargetPageId="9" captionText="FIGURE 7. Thyreophagus subiasi sp. nov., male paratype (A–C, E, F) and female, holotype (D), DIC images:A—dorsal view; B—ventral view; C—prodorsal shield; D—spermatheca; E—posterior part of opisthosoma, dorsal view; G—posterior part of opisthosoma, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14595271" httpUri="https://zenodo.org/record/14595271/files/figure.png" pageId="3" pageNumber="255">7D</figureCitation>
|
||
). Sac of spermatheca rounded, folded. Paired efferent ducts (
|
||
<emphasis id="B9650F72FFC814160294F9A2FC48FE2F" box="[984,1010,1555,1581]" italics="true" pageId="3" pageNumber="255">ed</emphasis>
|
||
) short, their length approximately 1/3 the length of spermatheca base, with short stem.
|
||
</paragraph>
|
||
<paragraph id="8BAED360FFC81411018BF9EDFA9AF91F" blockId="3.[151,1437,331,2020]" lastBlockId="4.[151,1437,151,357]" lastPageId="4" lastPageNumber="256" pageId="3" pageNumber="255">
|
||
Legs (
|
||
<figureCitation id="132ACFE5FFC814160040F9EAFEE9FE77" box="[268,339,1627,1653]" captionStart="FIGURE 4" captionStartId="6.[152,255,1875,1899]" captionTargetBox="[154,1432,184,1848]" captionTargetId="figure-19@6.[151,1435,181,1851]" captionTargetPageId="6" captionText="FIGURE 4. Thyreophagus subiasi sp. nov., female holotype:A—leg I, dorsal view; B—tarsus I, ventral view; C—leg II, dorsal view; D—tarsus II, ventral view; E—leg III, dorsal view; F—tarsus III, ventral view; G—leg IV, dorsal view; H—tarsus IV, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14595261" httpUri="https://zenodo.org/record/14595261/files/figure.png" pageId="3" pageNumber="255">Figs 4</figureCitation>
|
||
) short, all segments free. Trochanters I–III each with long, filiform seta,
|
||
<emphasis id="B9650F72FFC8141605CCF9EDFB25FE77" box="[1152,1183,1628,1653]" italics="true" pageId="3" pageNumber="255">pR</emphasis>
|
||
I–II,
|
||
<emphasis id="B9650F72FFC814160592F9EDFB40FE77" box="[1246,1274,1628,1653]" italics="true" pageId="3" pageNumber="255">sR</emphasis>
|
||
III; trochanter IV without setae. Femoral setation 1-1-0-1; setae
|
||
<emphasis id="B9650F72FFC814160385F931FD5DFE9B" box="[713,743,1664,1689]" italics="true" pageId="3" pageNumber="255">vF</emphasis>
|
||
I–II and
|
||
<emphasis id="B9650F72FFC81416021DF931FCCFFE9B" box="[849,885,1664,1689]" italics="true" pageId="3" pageNumber="255">wF</emphasis>
|
||
IV long, filiform. Genual setation 2-2-0-0; setae
|
||
<emphasis id="B9650F72FFC8141601DBF915FF05FEBC" box="[151,191,1700,1726]" italics="true" pageId="3" pageNumber="255">mG</emphasis>
|
||
and
|
||
<emphasis id="B9650F72FFC8141601B6F915FEA1FEBC" box="[250,283,1700,1726]" italics="true" pageId="3" pageNumber="255">cG</emphasis>
|
||
I–II long, filiform; seta
|
||
<emphasis id="B9650F72FFC81416037FF915FDEFFEBC" box="[563,597,1700,1726]" italics="true" pageId="3" pageNumber="255">nG</emphasis>
|
||
III absent. Tibial setation 2-2-1-1; setae
|
||
<emphasis id="B9650F72FFC814160565F912FBFDFEBF" box="[1065,1095,1699,1725]" italics="true" pageId="3" pageNumber="255">hT</emphasis>
|
||
I–II absent; setae
|
||
<emphasis id="B9650F72FFC814160457F915FA83FEBF" box="[1307,1337,1700,1725]" italics="true" pageId="3" pageNumber="255">gT</emphasis>
|
||
I–II and
|
||
<emphasis id="B9650F72FFC8141601DBF976FF09FEE3" box="[151,179,1735,1761]" italics="true" pageId="3" pageNumber="255">kT</emphasis>
|
||
III–IV elongate, somewhat spiniform. Tarsal setation 10-10-10-10; pretarsi consists of hooked empodial claws attached to short paired condylophores. Tarsus I and II with setae
|
||
<emphasis id="B9650F72FFC814160237F95DFC2EFF07" box="[891,916,1772,1797]" italics="true" pageId="3" pageNumber="255">ra</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC8141602EEF95AFC02FF07" box="[930,952,1771,1797]" italics="true" pageId="3" pageNumber="255">la</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC81416028AF95AFC74FF07" box="[966,974,1771,1797]" italics="true" pageId="3" pageNumber="255">f</emphasis>
|
||
and
|
||
<emphasis id="B9650F72FFC814160549F95AFBA9FF07" box="[1029,1043,1771,1797]" italics="true" pageId="3" pageNumber="255">d</emphasis>
|
||
filiform,
|
||
<emphasis id="B9650F72FFC814160533F95DFB31FF07" box="[1151,1163,1772,1797]" italics="true" pageId="3" pageNumber="255">e</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC8141605D6F95DFB12FF07" box="[1178,1192,1772,1797]" italics="true" pageId="3" pageNumber="255">u</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC8141605FAF95DFB78FF07" box="[1206,1218,1772,1797]" italics="true" pageId="3" pageNumber="255">v</emphasis>
|
||
spiniform (
|
||
<emphasis id="B9650F72FFC814160404F95DFAEEFF07" box="[1352,1364,1772,1797]" italics="true" pageId="3" pageNumber="255">v</emphasis>
|
||
larger that
|
||
<emphasis id="B9650F72FFC814160184F8A1FF6CFF2B" box="[200,214,1808,1833]" italics="true" pageId="3" pageNumber="255">u</emphasis>
|
||
),
|
||
<emphasis id="B9650F72FFC8141601A1F8A1FF41FF2B" box="[237,251,1808,1833]" italics="true" pageId="3" pageNumber="255">p</emphasis>
|
||
and
|
||
<emphasis id="B9650F72FFC81416007EF8A1FEFAFF2B" box="[306,320,1808,1833]" italics="true" pageId="3" pageNumber="255">q</emphasis>
|
||
represented by small triangular rudiments,
|
||
<emphasis id="B9650F72FFC81416026BF8A1FC88FF2B" box="[807,818,1808,1833]" italics="true" pageId="3" pageNumber="255">s</emphasis>
|
||
flattened, button-shaped or minute spiniform; setae
|
||
<emphasis id="B9650F72FFC814160430F8A1FA27FF2B" box="[1404,1437,1808,1833]" italics="true" pageId="3" pageNumber="255">wa</emphasis>
|
||
present, spiniform. Tarsus III and IV with setae
|
||
<emphasis id="B9650F72FFC8141603E1F882FD0FFF4F" box="[685,693,1843,1869]" italics="true" pageId="3" pageNumber="255">f</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC81416038FF882FD6BFF4F" box="[707,721,1843,1869]" italics="true" pageId="3" pageNumber="255">d</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC814160393F885FD50FF4F" box="[735,746,1844,1869]" italics="true" pageId="3" pageNumber="255">r</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC8141603BBF885FCB0FF4F" box="[759,778,1844,1869]" italics="true" pageId="3" pageNumber="255">w</emphasis>
|
||
filiform,
|
||
<emphasis id="B9650F72FFC81416023AF885FC38FF4F" box="[886,898,1844,1869]" italics="true" pageId="3" pageNumber="255">e</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC8141602DCF885FC21FF4F" box="[912,923,1844,1869]" italics="true" pageId="3" pageNumber="255">s</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC8141602E5F885FC0DFF4F" box="[937,951,1844,1869]" italics="true" pageId="3" pageNumber="255">u</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC814160289F885FC6BFF4F" box="[965,977,1844,1869]" italics="true" pageId="3" pageNumber="255">v</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC814160293F885FC57FF4F" box="[991,1005,1844,1869]" italics="true" pageId="3" pageNumber="255">p</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC8141602B7F885FBB3FF4F" box="[1019,1033,1844,1869]" italics="true" pageId="3" pageNumber="255">q</emphasis>
|
||
spiniform (
|
||
<emphasis id="B9650F72FFC8141605C1F885FB21FF4F" box="[1165,1179,1844,1869]" italics="true" pageId="3" pageNumber="255">p</emphasis>
|
||
and
|
||
<emphasis id="B9650F72FFC81416059DF885FB67FF4F" box="[1233,1245,1844,1869]" italics="true" pageId="3" pageNumber="255">v</emphasis>
|
||
larger than other ventral spines,
|
||
<emphasis id="B9650F72FFC814160073F8E9FEF7FF73" box="[319,333,1880,1905]" italics="true" pageId="3" pageNumber="255">u</emphasis>
|
||
shorter than other ventral spines). Solenidion ω
|
||
<subScript id="1795D125FFC81416022EF8DBFCD0FF7A" attach="left" box="[866,874,1898,1912]" fontSize="6" pageId="3" pageNumber="255">1</subScript>
|
||
on tarsus I cylindrical, with clavate apex, straight; solenidion ω
|
||
<subScript id="1795D125FFC81416006FF83FFE91FF9E" attach="left" box="[291,299,1934,1948]" fontSize="6" pageId="3" pageNumber="255">1</subScript>
|
||
on tarsus II simple, cylindrical, with clavate apex, straight, the same length as ω
|
||
<subScript id="1795D125FFC8141605D4F83FFB1AFF9E" attach="left" box="[1176,1184,1934,1948]" fontSize="6" pageId="3" pageNumber="255">1</subScript>
|
||
on tarsus I. Solenidion ω
|
||
<subScript id="1795D125FFC8141601E6F803FF08FFC2" attach="left" box="[170,178,1970,1984]" fontSize="6" pageId="3" pageNumber="255">2</subScript>
|
||
on tarsus I shorter than ω
|
||
<subScript id="1795D125FFC81416009CF803FE62FFC2" attach="left" box="[464,472,1970,1984]" fontSize="6" pageId="3" pageNumber="255">1</subScript>
|
||
, cylindrical, with rounded apex, situated slightly distal to ω
|
||
<subScript id="1795D125FFC81416053EF803FBC0FFC2" attach="right" box="[1138,1146,1970,1984]" fontSize="6" pageId="3" pageNumber="255">1</subScript>
|
||
. Solenidion ω
|
||
<subScript id="1795D125FFC814160456F803FA98FFC2" attach="left" box="[1306,1314,1970,1984]" fontSize="6" pageId="3" pageNumber="255">3</subScript>
|
||
on tarsus I cylindrical, with rounded tip, slightly shorter than ω
|
||
<subScript id="1795D125FFC81416039BF867FD65FFE6" attach="left" box="[727,735,2006,2020]" fontSize="6" pageId="3" pageNumber="255">1</subScript>
|
||
and slightly longer than ω
|
||
<subScript id="1795D125FFC814160549F867FBB7FFE6" attach="right" box="[1029,1037,2006,2020]" fontSize="6" pageId="3" pageNumber="255">2</subScript>
|
||
. Famulus (ε) of tarsus I wide, short, with broadly rounded apex, widest at middle. Solenidia φ of tibiae I–III elongate, tapering, well extending beyond apices of respective tarsi with ambulacra; solenidion φ IV shorter, shorter than tarsus IV (with ambulacra). Solenidia σ’ and σ” on genu I elongate, tapering, σ’ is slightly longer than σ”, slightly not reaching bases of φ I. Solenidion σ on genu II more than 5–6 times longer than its width) with rounded tip. Solenidion σ of genu III absent.
|
||
</paragraph>
|
||
<paragraph id="8BAED360FFCF1411018BFE96FF4BF967" blockId="4.[151,1437,151,357]" pageId="4" pageNumber="256">
|
||
Male (
|
||
<figureCitation id="132ACFE5FFCF14110043FE96FEEDF943" box="[271,343,295,321]" captionStart="FIGURE 3" captionStartId="5.[152,255,1625,1649]" captionTargetBox="[154,1432,184,1598]" captionTargetId="figure-19@5.[151,1435,181,1601]" captionTargetPageId="5" captionText="FIGURE 3. Thyreophagus subiasi sp. nov., male paratype: A—dorsal view; B—ventral view. Chaetotaxy of legs and palps is omitted." figureDoi="http://doi.org/10.5281/zenodo.14595259" httpUri="https://zenodo.org/record/14595259/files/figure.png" pageId="4" pageNumber="256">Figs 3</figureCitation>
|
||
,
|
||
<figureCitation id="132ACFE5FFCF1411002FFE97FE12F943" box="[355,424,294,321]" captionStart="FIGURE 5" captionStartId="7.[152,255,1060,1084]" captionTargetBox="[154,1432,184,1033]" captionTargetId="figure-19@7.[151,1435,181,1036]" captionTargetPageId="7" captionText="FIGURE 5. Thyreophagus subiasi sp. nov., female holotype (A, B) and male, paratype (C–G):A—subcapitulum, ventral view; B—spermatheca; C—male genitalia, ventral view; D—leg I, dorsal view; E—tarsus I, ventral view; F—leg IV, anterior view; G—tarsus IV, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14595265" httpUri="https://zenodo.org/record/14595265/files/figure.png" pageId="4" pageNumber="256">5C–G</figureCitation>
|
||
,
|
||
<figureCitation id="132ACFE5FFCF141100F8FE96FD82F943" box="[436,568,295,321]" captionStart="FIGURE 7" captionStartId="9.[152,255,1943,1967]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure-19@9.[178,1409,181,1919]" captionTargetPageId="9" captionText="FIGURE 7. Thyreophagus subiasi sp. nov., male paratype (A–C, E, F) and female, holotype (D), DIC images:A—dorsal view; B—ventral view; C—prodorsal shield; D—spermatheca; E—posterior part of opisthosoma, dorsal view; G—posterior part of opisthosoma, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14595271" httpUri="https://zenodo.org/record/14595271/files/figure.png" pageId="4" pageNumber="256">7A–B, E–F</figureCitation>
|
||
). Idiosoma elongate, 380 × 140, 2.7 times longer than wide. Idiosomal cuticle smooth.
|
||
</paragraph>
|
||
<caption id="DF6E83E8FFCF141101D4F814FEB9FFE3" ID-DOI="http://doi.org/10.5281/zenodo.14595257" ID-Zenodo-Dep="14595257" httpUri="https://zenodo.org/record/14595257/files/figure.png" pageId="4" pageNumber="256" startId="4.[152,255,1957,1981]" targetBox="[154,1432,384,1931]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BAED360FFCF141101D4F814FEB9FFE3" blockId="4.[151,1436,1957,2017]" pageId="4" pageNumber="256">
|
||
<emphasis id="B9650F72FFCF141101D4F814FEA3FFBF" bold="true" box="[152,281,1957,1981]" pageId="4" pageNumber="256">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C11A8E3FFCF14110053F814FE44FFBF" authorityName="Kolesnikov & Klimov & Khaustov & Pepato & Makarova" authorityYear="2024" box="[287,510,1957,1981]" class="Arachnida" family="Acaridae" genus="Thyreophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="256" phylum="Arthropoda" rank="species" species="subiasi" status="sp. nov.">
|
||
<emphasis id="B9650F72FFCF14110053F814FE44FFBF" box="[287,510,1957,1981]" italics="true" pageId="4" pageNumber="256">Thyreophagus subiasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9650F72FFCF14110348F814FDEFFFBF" bold="true" box="[516,597,1957,1981]" pageId="4" pageNumber="256">
|
||
<taxonomicNameLabel id="A256B209FFCF14110348F814FDEFFFBF" box="[516,597,1957,1981]" pageId="4" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female holotype: A—dorsal view; B—ventral view. Chaetotaxy of legs and palps is omitted.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6E83E8FFCE141001D4F9E8FF56FE97" ID-DOI="http://doi.org/10.5281/zenodo.14595259" ID-Zenodo-Dep="14595259" httpUri="https://zenodo.org/record/14595259/files/figure.png" pageId="5" pageNumber="257" startId="5.[152,255,1625,1649]" targetBox="[154,1432,184,1598]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8BAED360FFCE141001D4F9E8FF56FE97" blockId="5.[151,1436,1625,1685]" pageId="5" pageNumber="257">
|
||
<emphasis id="B9650F72FFCE141001D4F9E8FEA2FE73" bold="true" box="[152,280,1625,1649]" pageId="5" pageNumber="257">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C11A8E3FFCE14100053F9E8FE44FE73" authorityName="Kolesnikov & Klimov & Khaustov & Pepato & Makarova" authorityYear="2024" box="[287,510,1625,1649]" class="Arachnida" family="Acaridae" genus="Thyreophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="257" phylum="Arthropoda" rank="species" species="subiasi" status="sp. nov.">
|
||
<emphasis id="B9650F72FFCE14100053F9E8FE44FE73" box="[287,510,1625,1649]" italics="true" pageId="5" pageNumber="257">Thyreophagus subiasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9650F72FFCE14100348F9E8FDEFFE73" bold="true" box="[516,597,1625,1649]" pageId="5" pageNumber="257">
|
||
<taxonomicNameLabel id="A256B209FFCE14100348F9E8FDEFFE73" box="[516,597,1625,1649]" pageId="5" pageNumber="257" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male paratype: A—dorsal view; B—ventral view. Chaetotaxy of legs and palps is omitted.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAED360FFCE1410018BF956FE51FF03" blockId="5.[151,1437,1767,2010]" box="[199,491,1767,1794]" pageId="5" pageNumber="257">Gnathosoma as in female.</paragraph>
|
||
<paragraph id="8BAED360FFCE1412018BF8BAFAC7FD83" blockId="5.[151,1437,1767,2010]" lastBlockId="7.[151,1437,1238,2021]" lastPageId="7" lastPageNumber="259" pageId="5" pageNumber="257">
|
||
Prodorsal shield (
|
||
<figureCitation id="132ACFE5FFCE141000CBF8BAFE66FF24" box="[391,476,1803,1830]" captionStart="FIGURE 3" captionStartId="5.[152,255,1625,1649]" captionTargetBox="[154,1432,184,1598]" captionTargetId="figure-19@5.[151,1435,181,1601]" captionTargetPageId="5" captionText="FIGURE 3. Thyreophagus subiasi sp. nov., male paratype: A—dorsal view; B—ventral view. Chaetotaxy of legs and palps is omitted." figureDoi="http://doi.org/10.5281/zenodo.14595259" httpUri="https://zenodo.org/record/14595259/files/figure.png" pageId="5" pageNumber="257">Fig. 3A</figureCitation>
|
||
) 80 long, 62 wide, 1.3 times longer than wide, with setae
|
||
<emphasis id="B9650F72FFCE1410050BF8BDFBE1FF27" box="[1095,1115,1804,1829]" italics="true" pageId="5" pageNumber="257">vi</emphasis>
|
||
, incisions and ornamented as in female (sometimes lineate pattern is weakly developed). Grandjean’s organ (GO) with 14 membranous processes. Supracoxal seta (
|
||
<emphasis id="B9650F72FFCE14100019F8E5FEC3FF6F" box="[341,377,1876,1901]" italics="true" pageId="5" pageNumber="257">scx</emphasis>
|
||
) as in female. Idiosomal setae (
|
||
<emphasis id="B9650F72FFCE1410039AF8E5FD50FF6F" box="[726,746,1876,1901]" italics="true" pageId="5" pageNumber="257">vi</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFCE141003BBF8E5FCB4FF6F" box="[759,782,1876,1901]" italics="true" pageId="5" pageNumber="257">se</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFCE14100257F8E5FC9DFF6F" box="[795,807,1876,1901]" italics="true" pageId="5" pageNumber="257">c</emphasis>
|
||
<subScript id="1795D125FFCE1410026BF8D7FC95FF76" attach="left" box="[807,815,1894,1908]" fontSize="6" pageId="5" pageNumber="257">p</subScript>
|
||
,
|
||
<emphasis id="B9650F72FFCE14100270F8E2FCF0FF6F" box="[828,842,1875,1901]" italics="true" pageId="5" pageNumber="257">d</emphasis>
|
||
<subScript id="1795D125FFCE14100206F8D7FCE8FF76" attach="left" box="[842,850,1894,1908]" fontSize="6" pageId="5" pageNumber="257">2</subScript>
|
||
,
|
||
<emphasis id="B9650F72FFCE14100213F8E5FCD1FF6F" box="[863,875,1876,1901]" italics="true" pageId="5" pageNumber="257">e</emphasis>
|
||
<subScript id="1795D125FFCE14100227F8D7FCC9FF76" attach="left" box="[875,883,1894,1908]" fontSize="6" pageId="5" pageNumber="257">2</subScript>
|
||
,
|
||
<emphasis id="B9650F72FFCE141002CCF8E2FC34FF6F" box="[896,910,1875,1901]" italics="true" pageId="5" pageNumber="257">h</emphasis>
|
||
<subScript id="1795D125FFCE141002C2F8D7FC2CFF76" attach="left" box="[910,918,1894,1908]" fontSize="6" pageId="5" pageNumber="257">1</subScript>
|
||
,
|
||
<emphasis id="B9650F72FFCE141002EFF8E2FC0BFF6F" box="[931,945,1875,1901]" italics="true" pageId="5" pageNumber="257">h</emphasis>
|
||
<subScript id="1795D125FFCE141002FDF8D7FC03FF76" attach="left" box="[945,953,1894,1908]" fontSize="6" pageId="5" pageNumber="257">2</subScript>
|
||
,
|
||
<emphasis id="B9650F72FFCE1410028AF8E2FC6EFF6F" box="[966,980,1875,1901]" italics="true" pageId="5" pageNumber="257">h</emphasis>
|
||
<subScript id="1795D125FFCE14100298F8D7FC66FF76" attach="left" box="[980,988,1894,1908]" fontSize="6" pageId="5" pageNumber="257">3</subScript>
|
||
) smooth, filiform and short (except long
|
||
<emphasis id="B9650F72FFCE141001DBF8C9FF14FF93" box="[151,174,1912,1937]" italics="true" pageId="5" pageNumber="257">se</emphasis>
|
||
) (
|
||
<figureCitation id="132ACFE5FFCE1410018BF8C6FEB4FF90" box="[199,270,1911,1938]" captionStart="FIGURE 3" captionStartId="5.[152,255,1625,1649]" captionTargetBox="[154,1432,184,1598]" captionTargetId="figure-19@5.[151,1435,181,1601]" captionTargetPageId="5" captionText="FIGURE 3. Thyreophagus subiasi sp. nov., male paratype: A—dorsal view; B—ventral view. Chaetotaxy of legs and palps is omitted." figureDoi="http://doi.org/10.5281/zenodo.14595259" httpUri="https://zenodo.org/record/14595259/files/figure.png" pageId="5" pageNumber="257">Figs 3</figureCitation>
|
||
,
|
||
<figureCitation id="132ACFE5FFCE14100057F8C9FEDBFF90" box="[283,353,1912,1938]" captionStart="FIGURE 7" captionStartId="9.[152,255,1943,1967]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure-19@9.[178,1409,181,1919]" captionTargetPageId="9" captionText="FIGURE 7. Thyreophagus subiasi sp. nov., male paratype (A–C, E, F) and female, holotype (D), DIC images:A—dorsal view; B—ventral view; C—prodorsal shield; D—spermatheca; E—posterior part of opisthosoma, dorsal view; G—posterior part of opisthosoma, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14595271" httpUri="https://zenodo.org/record/14595271/files/figure.png" pageId="5" pageNumber="257">7A, B</figureCitation>
|
||
); opisthosomal gland openings slightly anteriad setal bases
|
||
<emphasis id="B9650F72FFCE141002B2F8C9FBB0FF93" box="[1022,1034,1912,1937]" italics="true" pageId="5" pageNumber="257">e</emphasis>
|
||
<subScript id="1795D125FFCE14100546F83BFBA8FF9A" attach="left" box="[1034,1042,1930,1944]" fontSize="6" pageId="5" pageNumber="257">2</subScript>
|
||
. Two pairs of fundamental cupules (
|
||
<emphasis id="B9650F72FFCE141001EDF82DFF0DFFB7" box="[161,183,1948,1973]" italics="true" pageId="5" pageNumber="257">ia</emphasis>
|
||
and
|
||
<emphasis id="B9650F72FFCE141001A3F82AFEBFFFB7" box="[239,261,1947,1973]" italics="true" pageId="5" pageNumber="257">ih</emphasis>
|
||
) present,
|
||
<emphasis id="B9650F72FFCE1410003AF82DFE28FFB7" box="[374,402,1948,1973]" italics="true" pageId="5" pageNumber="257">im</emphasis>
|
||
and
|
||
<emphasis id="B9650F72FFCE14100086F82DFE5AFFB7" box="[458,480,1948,1973]" italics="true" pageId="5" pageNumber="257">ip</emphasis>
|
||
not observed (
|
||
<figureCitation id="132ACFE5FFCE141003C6F82AFD75FFB4" box="[650,719,1947,1974]" captionStart="FIGURE 3" captionStartId="5.[152,255,1625,1649]" captionTargetBox="[154,1432,184,1598]" captionTargetId="figure-19@5.[151,1435,181,1601]" captionTargetPageId="5" captionText="FIGURE 3. Thyreophagus subiasi sp. nov., male paratype: A—dorsal view; B—ventral view. Chaetotaxy of legs and palps is omitted." figureDoi="http://doi.org/10.5281/zenodo.14595259" httpUri="https://zenodo.org/record/14595259/files/figure.png" pageId="5" pageNumber="257">Fig. 3</figureCitation>
|
||
). Opisthonotal shield (
|
||
<figureCitation id="132ACFE5FFCE1410029DF82AFB95FFB4" box="[977,1071,1947,1974]" captionStart="FIGURE 3" captionStartId="5.[152,255,1625,1649]" captionTargetBox="[154,1432,184,1598]" captionTargetId="figure-19@5.[151,1435,181,1601]" captionTargetPageId="5" captionText="FIGURE 3. Thyreophagus subiasi sp. nov., male paratype: A—dorsal view; B—ventral view. Chaetotaxy of legs and palps is omitted." figureDoi="http://doi.org/10.5281/zenodo.14595259" httpUri="https://zenodo.org/record/14595259/files/figure.png" pageId="5" pageNumber="257">Figs 3A</figureCitation>
|
||
,
|
||
<figureCitation id="132ACFE5FFCE14100570F82DFBDAFFB4" box="[1084,1120,1948,1974]" captionStart="FIGURE 7" captionStartId="9.[152,255,1943,1967]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure-19@9.[178,1409,181,1919]" captionTargetPageId="9" captionText="FIGURE 7. Thyreophagus subiasi sp. nov., male paratype (A–C, E, F) and female, holotype (D), DIC images:A—dorsal view; B—ventral view; C—prodorsal shield; D—spermatheca; E—posterior part of opisthosoma, dorsal view; G—posterior part of opisthosoma, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14595271" httpUri="https://zenodo.org/record/14595271/files/figure.png" pageId="5" pageNumber="257">7A</figureCitation>
|
||
) smoothly punctate; ventral part extends to anal suckers. Posterior region of idiosoma with a large rounded lobe extending the body backwards (56 × 75) (
|
||
<figureCitation id="132ACFE5FFCC14120042FB67FEEFFCF3" box="[270,341,1238,1265]" captionStart="FIGURE 3" captionStartId="5.[152,255,1625,1649]" captionTargetBox="[154,1432,184,1598]" captionTargetId="figure-19@5.[151,1435,181,1601]" captionTargetPageId="5" captionText="FIGURE 3. Thyreophagus subiasi sp. nov., male paratype: A—dorsal view; B—ventral view. Chaetotaxy of legs and palps is omitted." figureDoi="http://doi.org/10.5281/zenodo.14595259" httpUri="https://zenodo.org/record/14595259/files/figure.png" pageId="7" pageNumber="259">
|
||
Figs
|
||
<geoCoordinate id="EE25B5A7FFCC1412000AFB66FEEFFCF3" box="[326,341,1239,1265]" degrees="3, 7" direction="east" orientation="longitude" pageId="7" pageNumber="259" precision="5555" value="3.7">3</geoCoordinate>
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="132ACFE5FFCC1412002EFB66FE18FCF3" box="[354,418,1239,1265]" captionStart="FIGURE 7" captionStartId="9.[152,255,1943,1967]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure-19@9.[178,1409,181,1919]" captionTargetPageId="9" captionText="FIGURE 7. Thyreophagus subiasi sp. nov., male paratype (A–C, E, F) and female, holotype (D), DIC images:A—dorsal view; B—ventral view; C—prodorsal shield; D—spermatheca; E—posterior part of opisthosoma, dorsal view; G—posterior part of opisthosoma, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14595271" httpUri="https://zenodo.org/record/14595271/files/figure.png" pageId="7" pageNumber="259">7E, F</figureCitation>
|
||
). Ventral idiosoma with four pairs of coxal setae (
|
||
<emphasis id="B9650F72FFCC14120283FB66FC56FCF3" box="[975,1004,1239,1265]" italics="true" pageId="7" pageNumber="259">1a</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFCC141202B5FB66FBAFFCF3" box="[1017,1045,1239,1265]" italics="true" pageId="7" pageNumber="259">3a</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFCC1412056FFB66FB85FCF3" box="[1059,1087,1239,1265]" italics="true" pageId="7" pageNumber="259">4a</emphasis>
|
||
and
|
||
<emphasis id="B9650F72FFCC14120539FB67FB2BFCF2" box="[1141,1169,1238,1264]" italics="true" pageId="7" pageNumber="259">4b</emphasis>
|
||
) and one pair of genital setae (
|
||
<emphasis id="B9650F72FFCC141201ADFB4AFF55FD16" box="[225,239,1275,1300]" italics="true" pageId="7" pageNumber="259">g</emphasis>
|
||
) (
|
||
<figureCitation id="132ACFE5FFCC14120045FB4BFEDCFD17" box="[265,358,1274,1301]" captionStart="FIGURE 3" captionStartId="5.[152,255,1625,1649]" captionTargetBox="[154,1432,184,1598]" captionTargetId="figure-19@5.[151,1435,181,1601]" captionTargetPageId="5" captionText="FIGURE 3. Thyreophagus subiasi sp. nov., male paratype: A—dorsal view; B—ventral view. Chaetotaxy of legs and palps is omitted." figureDoi="http://doi.org/10.5281/zenodo.14595259" httpUri="https://zenodo.org/record/14595259/files/figure.png" pageId="7" pageNumber="259">Figs 3B</figureCitation>
|
||
). Shape of coxal sclerites on
|
||
<figureCitation id="132ACFE5FFCC141203FBFB4BFC8EFD17" box="[695,820,1274,1301]" captionStart="FIGURE 3" captionStartId="5.[152,255,1625,1649]" captionTargetBox="[154,1432,184,1598]" captionTargetId="figure-19@5.[151,1435,181,1601]" captionTargetPageId="5" captionText="FIGURE 3. Thyreophagus subiasi sp. nov., male paratype: A—dorsal view; B—ventral view. Chaetotaxy of legs and palps is omitted." figureDoi="http://doi.org/10.5281/zenodo.14595259" httpUri="https://zenodo.org/record/14595259/files/figure.png" pageId="7" pageNumber="259">Figures 3B</figureCitation>
|
||
and
|
||
<figureCitation id="132ACFE5FFCC14120222FB4AFC2BFD17" box="[878,913,1275,1301]" captionStart="FIGURE 7" captionStartId="9.[152,255,1943,1967]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure-19@9.[178,1409,181,1919]" captionTargetPageId="9" captionText="FIGURE 7. Thyreophagus subiasi sp. nov., male paratype (A–C, E, F) and female, holotype (D), DIC images:A—dorsal view; B—ventral view; C—prodorsal shield; D—spermatheca; E—posterior part of opisthosoma, dorsal view; G—posterior part of opisthosoma, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14595271" httpUri="https://zenodo.org/record/14595271/files/figure.png" pageId="7" pageNumber="259">7B</figureCitation>
|
||
. Genital region between coxisternal fields IV; arms of genital capsule rounded; aedeagus short, not protruding beyond anterior edge of genital capsule, dorsal supporting sclerite wider that genital capsule (
|
||
<figureCitation id="132ACFE5FFCC141203D6FAF3FD4FFD5F" box="[666,757,1346,1373]" captionStart="FIGURE 3" captionStartId="5.[152,255,1625,1649]" captionTargetBox="[154,1432,184,1598]" captionTargetId="figure-19@5.[151,1435,181,1601]" captionTargetPageId="5" captionText="FIGURE 3. Thyreophagus subiasi sp. nov., male paratype: A—dorsal view; B—ventral view. Chaetotaxy of legs and palps is omitted." figureDoi="http://doi.org/10.5281/zenodo.14595259" httpUri="https://zenodo.org/record/14595259/files/figure.png" pageId="7" pageNumber="259">Figs 3B</figureCitation>
|
||
,
|
||
<figureCitation id="132ACFE5FFCC1412024CFAF3FC9FFD5F" box="[768,805,1346,1373]" captionStart="FIGURE 5" captionStartId="7.[152,255,1060,1084]" captionTargetBox="[154,1432,184,1033]" captionTargetId="figure-19@7.[151,1435,181,1036]" captionTargetPageId="7" captionText="FIGURE 5. Thyreophagus subiasi sp. nov., female holotype (A, B) and male, paratype (C–G):A—subcapitulum, ventral view; B—spermatheca; C—male genitalia, ventral view; D—leg I, dorsal view; E—tarsus I, ventral view; F—leg IV, anterior view; G—tarsus IV, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14595265" httpUri="https://zenodo.org/record/14595265/files/figure.png" pageId="7" pageNumber="259">5C</figureCitation>
|
||
). Diameter of genital papillae approximately 0.3–0.4 the length of coxal and genital setae. Anal suckers rounded in outline (
|
||
<figureCitation id="132ACFE5FFCC141202CEFAD7FC67FD83" box="[898,989,1382,1409]" captionStart="FIGURE 3" captionStartId="5.[152,255,1625,1649]" captionTargetBox="[154,1432,184,1598]" captionTargetId="figure-19@5.[151,1435,181,1601]" captionTargetPageId="5" captionText="FIGURE 3. Thyreophagus subiasi sp. nov., male paratype: A—dorsal view; B—ventral view. Chaetotaxy of legs and palps is omitted." figureDoi="http://doi.org/10.5281/zenodo.14595259" httpUri="https://zenodo.org/record/14595259/files/figure.png" pageId="7" pageNumber="259">Figs 3B</figureCitation>
|
||
,
|
||
<figureCitation id="132ACFE5FFCC141202A5FAD6FBB0FD83" box="[1001,1034,1383,1409]" captionStart="FIGURE 7" captionStartId="9.[152,255,1943,1967]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure-19@9.[178,1409,181,1919]" captionTargetPageId="9" captionText="FIGURE 7. Thyreophagus subiasi sp. nov., male paratype (A–C, E, F) and female, holotype (D), DIC images:A—dorsal view; B—ventral view; C—prodorsal shield; D—spermatheca; E—posterior part of opisthosoma, dorsal view; G—posterior part of opisthosoma, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14595271" httpUri="https://zenodo.org/record/14595271/files/figure.png" pageId="7" pageNumber="259">7F</figureCitation>
|
||
). Setae
|
||
<emphasis id="B9650F72FFCC1412052EFAD6FBC1FD82" box="[1122,1147,1383,1408]" italics="true" pageId="7" pageNumber="259">ps</emphasis>
|
||
<subScript id="1795D125FFCC14120537FAC8FB29FD85" attach="left" box="[1147,1171,1401,1415]" fontSize="6" pageId="7" pageNumber="259">1–3</subScript>
|
||
very short (
|
||
<figureCitation id="132ACFE5FFCC14120456FAD7FAD5FD83" box="[1306,1391,1382,1409]" captionStart="FIGURE 7" captionStartId="9.[152,255,1943,1967]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure-19@9.[178,1409,181,1919]" captionTargetPageId="9" captionText="FIGURE 7. Thyreophagus subiasi sp. nov., male paratype (A–C, E, F) and female, holotype (D), DIC images:A—dorsal view; B—ventral view; C—prodorsal shield; D—spermatheca; E—posterior part of opisthosoma, dorsal view; G—posterior part of opisthosoma, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14595271" httpUri="https://zenodo.org/record/14595271/files/figure.png" pageId="7" pageNumber="259">Fig. 7F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF6E83E8FFCD141301D4F8E2FEA1FFB1" ID-DOI="http://doi.org/10.5281/zenodo.14595261" ID-Zenodo-Dep="14595261" httpUri="https://zenodo.org/record/14595261/files/figure.png" pageId="6" pageNumber="258" startId="6.[152,255,1875,1899]" targetBox="[154,1432,184,1848]" targetPageId="6" targetType="figure">
|
||
<paragraph id="8BAED360FFCD141301D4F8E2FEA1FFB1" blockId="6.[151,1437,1875,1971]" pageId="6" pageNumber="258">
|
||
<emphasis id="B9650F72FFCD141301D4F8E2FEADFF69" bold="true" box="[152,279,1875,1899]" pageId="6" pageNumber="258">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C11A8E3FFCD14130050F8E2FE40FF69" authorityName="Kolesnikov & Klimov & Khaustov & Pepato & Makarova" authorityYear="2024" box="[284,506,1875,1899]" class="Arachnida" family="Acaridae" genus="Thyreophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="258" phylum="Arthropoda" rank="species" species="subiasi" status="sp. nov.">
|
||
<emphasis id="B9650F72FFCD14130050F8E2FE40FF69" box="[284,506,1875,1899]" italics="true" pageId="6" pageNumber="258">Thyreophagus subiasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9650F72FFCD141300B3F8E2FDF4FF69" bold="true" box="[511,590,1875,1899]" pageId="6" pageNumber="258">
|
||
<taxonomicNameLabel id="A256B209FFCD141300B3F8E2FDF4FF69" box="[511,590,1875,1899]" pageId="6" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female holotype: A—leg I, dorsal view; B—tarsus I, ventral view; C—leg II, dorsal view; D—tarsus II, ventral view; E—leg III, dorsal view; F—tarsus III, ventral view; G—leg IV, dorsal view; H—tarsus IV, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6E83E8FFCC141201D4FB95FE7AFC86" ID-DOI="http://doi.org/10.5281/zenodo.14595265" ID-Zenodo-Dep="14595265" httpUri="https://zenodo.org/record/14595265/files/figure.png" pageId="7" pageNumber="259" startId="7.[152,255,1060,1084]" targetBox="[154,1432,184,1033]" targetPageId="7" targetType="figure">
|
||
<paragraph id="8BAED360FFCC141201D4FB95FE7AFC86" blockId="7.[151,1436,1060,1156]" pageId="7" pageNumber="259">
|
||
<emphasis id="B9650F72FFCC141201D4FB95FEADFC3E" bold="true" box="[152,279,1060,1084]" pageId="7" pageNumber="259">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C11A8E3FFCC14120051FB95FE40FC3E" authorityName="Kolesnikov & Klimov & Khaustov & Pepato & Makarova" authorityYear="2024" box="[285,506,1060,1084]" class="Arachnida" family="Acaridae" genus="Thyreophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="259" phylum="Arthropoda" rank="species" species="subiasi" status="sp. nov.">
|
||
<emphasis id="B9650F72FFCC14120051FB95FE40FC3E" box="[285,506,1060,1084]" italics="true" pageId="7" pageNumber="259">Thyreophagus subiasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9650F72FFCC1412034CFB95FDF5FC3E" bold="true" box="[512,591,1060,1084]" pageId="7" pageNumber="259">
|
||
<taxonomicNameLabel id="A256B209FFCC1412034CFB95FDF5FC3E" box="[512,591,1060,1084]" pageId="7" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female holotype (A, B) and male, paratype (C–G): A—subcapitulum, ventral view; B—spermatheca; C—male genitalia, ventral view; D—leg I, dorsal view; E—tarsus I, ventral view; F—leg IV, anterior view; G—tarsus IV, posterior view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAED360FFCC1412018BFA3AFADCFE12" blockId="7.[151,1437,1238,2021]" pageId="7" pageNumber="259">
|
||
Legs I–III as female, except solenidion ω
|
||
<subScript id="1795D125FFCC141203D6FA2CFD18FDA9" attach="left" box="[666,674,1437,1451]" fontSize="6" pageId="7" pageNumber="259">3</subScript>
|
||
on tarsus I very short, truncated (
|
||
<figureCitation id="132ACFE5FFCC1412056DFA3BFB1AFDA7" box="[1057,1184,1418,1445]" captionStart="FIGURE 5" captionStartId="7.[152,255,1060,1084]" captionTargetBox="[154,1432,184,1033]" captionTargetId="figure-19@7.[151,1435,181,1036]" captionTargetPageId="7" captionText="FIGURE 5. Thyreophagus subiasi sp. nov., female holotype (A, B) and male, paratype (C–G):A—subcapitulum, ventral view; B—spermatheca; C—male genitalia, ventral view; D—leg I, dorsal view; E—tarsus I, ventral view; F—leg IV, anterior view; G—tarsus IV, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14595265" httpUri="https://zenodo.org/record/14595265/files/figure.png" pageId="7" pageNumber="259">Figs 5D, E</figureCitation>
|
||
). Trochanter and genu IV without setae, femur IV with
|
||
<emphasis id="B9650F72FFCC1412034DFA1EFD9FFDCA" box="[513,549,1455,1480]" italics="true" pageId="7" pageNumber="259">wF</emphasis>
|
||
IV filiform, tibia IV with
|
||
<emphasis id="B9650F72FFCC1412020AFA1FFCD8FDCA" box="[838,866,1454,1480]" italics="true" pageId="7" pageNumber="259">kT</emphasis>
|
||
IV elongate, somewhat spiniform (
|
||
<figureCitation id="132ACFE5FFCC141205A0FA1FFAFBFDCB" box="[1260,1345,1454,1481]" captionStart="FIGURE 5" captionStartId="7.[152,255,1060,1084]" captionTargetBox="[154,1432,184,1033]" captionTargetId="figure-19@7.[151,1435,181,1036]" captionTargetPageId="7" captionText="FIGURE 5. Thyreophagus subiasi sp. nov., female holotype (A, B) and male, paratype (C–G):A—subcapitulum, ventral view; B—spermatheca; C—male genitalia, ventral view; D—leg I, dorsal view; E—tarsus I, ventral view; F—leg IV, anterior view; G—tarsus IV, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14595265" httpUri="https://zenodo.org/record/14595265/files/figure.png" pageId="7" pageNumber="259">Fig. 5F</figureCitation>
|
||
). Tarsus IV with 10 setae, of them
|
||
<emphasis id="B9650F72FFCC141200F7FA62FE79FDEF" box="[443,451,1491,1517]" italics="true" pageId="7" pageNumber="259">f</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFCC1412009DFA62FE66FDEE" box="[465,476,1491,1516]" italics="true" pageId="7" pageNumber="259">r</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFCC141200A6FA62FE47FDEE" box="[490,509,1491,1516]" italics="true" pageId="7" pageNumber="259">w</emphasis>
|
||
filiform,
|
||
<emphasis id="B9650F72FFCC14120325FA63FDCDFDEE" box="[617,631,1490,1516]" italics="true" pageId="7" pageNumber="259">d</emphasis>
|
||
and
|
||
<emphasis id="B9650F72FFCC141203E1FA62FD03FDEE" box="[685,697,1491,1516]" italics="true" pageId="7" pageNumber="259">e</emphasis>
|
||
represented by suckers,
|
||
<emphasis id="B9650F72FFCC14120282FA62FC63FDEE" box="[974,985,1491,1516]" italics="true" pageId="7" pageNumber="259">s</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFCC141202ABFA62FC4FFDEE" box="[999,1013,1491,1516]" italics="true" pageId="7" pageNumber="259">u</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFCC1412054FFA62FBB5FDEE" box="[1027,1039,1491,1516]" italics="true" pageId="7" pageNumber="259">v</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFCC14120552FA62FB96FDEE" box="[1054,1068,1491,1516]" italics="true" pageId="7" pageNumber="259">p</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFCC14120576FA62FBF2FDEE" box="[1082,1096,1491,1516]" italics="true" pageId="7" pageNumber="259">q</emphasis>
|
||
spiniform (
|
||
<emphasis id="B9650F72FFCC14120580FA62FB60FDEE" box="[1228,1242,1491,1516]" italics="true" pageId="7" pageNumber="259">p</emphasis>
|
||
and
|
||
<emphasis id="B9650F72FFCC1412045DFA62FAA7FDEE" box="[1297,1309,1491,1516]" italics="true" pageId="7" pageNumber="259">v</emphasis>
|
||
larger than other ventral spines,
|
||
<emphasis id="B9650F72FFCC141200CCFA46FE34FE12" box="[384,398,1527,1552]" italics="true" pageId="7" pageNumber="259">u</emphasis>
|
||
shorter than other ventral spines) (
|
||
<figureCitation id="132ACFE5FFCC14120259FA47FC35FE13" box="[789,911,1526,1553]" captionStart="FIGURE 5" captionStartId="7.[152,255,1060,1084]" captionTargetBox="[154,1432,184,1033]" captionTargetId="figure-19@7.[151,1435,181,1036]" captionTargetPageId="7" captionText="FIGURE 5. Thyreophagus subiasi sp. nov., female holotype (A, B) and male, paratype (C–G):A—subcapitulum, ventral view; B—spermatheca; C—male genitalia, ventral view; D—leg I, dorsal view; E—tarsus I, ventral view; F—leg IV, anterior view; G—tarsus IV, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14595265" httpUri="https://zenodo.org/record/14595265/files/figure.png" pageId="7" pageNumber="259">Figs 5F, G</figureCitation>
|
||
). Solenidion φ on tibia IV short and wide.
|
||
</paragraph>
|
||
<paragraph id="8BAED360FFCC1412018BF9ABFD81FE36" blockId="7.[151,1437,1238,2021]" box="[199,571,1562,1589]" pageId="7" pageNumber="259">
|
||
<emphasis id="B9650F72FFCC1412018BF9ABFE0DFE36" box="[199,439,1562,1588]" italics="true" pageId="7" pageNumber="259">Phoretic deutonymph</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30B80EBFFCC1412018BF98FFC22FFE7" pageId="7" pageNumber="259" type="discussion">
|
||
<paragraph id="8BAED360FFCC1412018BF98FFC22FFE7" blockId="7.[151,1437,1238,2021]" pageId="7" pageNumber="259">
|
||
<emphasis id="B9650F72FFCC1412018BF98FFE93FE5A" box="[199,297,1598,1624]" italics="true" pageId="7" pageNumber="259">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="4C11A8E3FFCC14120077F98FFD8FFE5A" authorityName="Kolesnikov & Klimov & Khaustov & Pepato & Makarova" authorityYear="2024" box="[315,565,1598,1624]" class="Arachnida" family="Acaridae" genus="Thyreophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="259" phylum="Arthropoda" rank="species" species="subiasi" status="sp. nov.">
|
||
<emphasis id="B9650F72FFCC14120077F98FFD8FFE5A" box="[315,565,1598,1624]" italics="true" pageId="7" pageNumber="259">Thyreophagus subiasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9650F72FFCC1412030CF98EFD24FE5B" bold="true" box="[576,670,1599,1625]" pageId="7" pageNumber="259">
|
||
<taxonomicNameLabel id="A256B209FFCC1412030CF98EFD24FE5B" box="[576,670,1599,1625]" pageId="7" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is morphologically close to
|
||
<taxonomicName id="4C11A8E3FFCC141202BDF98FFB3AFE5A" baseAuthorityName="Fain" baseAuthorityYear="1982" box="[1009,1152,1598,1624]" class="Arachnida" family="Acaridae" genus="Thyreophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="259" phylum="Arthropoda" rank="species" species="athiasae">
|
||
<emphasis id="B9650F72FFCC141202BDF98FFB3AFE5A" box="[1009,1152,1598,1624]" italics="true" pageId="7" pageNumber="259">Th. athiasae</emphasis>
|
||
</taxonomicName>
|
||
, as they share a similar prodorsal shield (with a small flat protrus and lineate pattern in posterior part), gnathosoma (with
|
||
<emphasis id="B9650F72FFCC141205A1F9D3FAA6FE7E" box="[1261,1308,1634,1660]" italics="true" pageId="7" pageNumber="259">elcp</emphasis>
|
||
setae), and leg chaetotaxy (except for the shape of
|
||
<emphasis id="B9650F72FFCC1412031CF936FDE4FEA2" box="[592,606,1671,1696]" italics="true" pageId="7" pageNumber="259">p</emphasis>
|
||
and
|
||
<emphasis id="B9650F72FFCC141203D8F936FD18FEA2" box="[660,674,1671,1696]" italics="true" pageId="7" pageNumber="259">q</emphasis>
|
||
I–II, with spiniform
|
||
<emphasis id="B9650F72FFCC141202C2F936FC20FEA2" box="[910,922,1671,1696]" italics="true" pageId="7" pageNumber="259">v</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFCC141202E4F936FC09FEA2" box="[936,947,1671,1696]" italics="true" pageId="7" pageNumber="259">s</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFCC1412028DF936FC75FEA2" box="[961,975,1671,1696]" italics="true" pageId="7" pageNumber="259">u</emphasis>
|
||
and setiform
|
||
<emphasis id="B9650F72FFCC14120526F936FBC7FEA2" box="[1130,1149,1671,1696]" italics="true" pageId="7" pageNumber="259">w</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFCC141205C6F936FB2FFEA2" box="[1162,1173,1671,1696]" italics="true" pageId="7" pageNumber="259">r</emphasis>
|
||
on III–IV tarsi, present
|
||
<emphasis id="B9650F72FFCC141201DBF91AFF02FEC6" box="[151,184,1707,1732]" italics="true" pageId="7" pageNumber="259">wa</emphasis>
|
||
I–II and absent
|
||
<emphasis id="B9650F72FFCC1412003DF91BFE35FEC6" box="[369,399,1706,1732]" italics="true" pageId="7" pageNumber="259">hT</emphasis>
|
||
I–II), and the length of idiosomal setae (setae
|
||
<emphasis id="B9650F72FFCC141202ECF91BFC14FEC6" box="[928,942,1706,1732]" italics="true" pageId="7" pageNumber="259">h</emphasis>
|
||
<subScript id="1795D125FFCC141202E2F90CFC79FEC9" attach="left" box="[942,963,1725,1739]" fontSize="6" pageId="7" pageNumber="259">1-3</subScript>
|
||
short, not longer than length of prodorsal shield). Both species have similar spermathecas: sclerotized atrium of spermatheca present, vase-shaped, partially enveloped by the spermathecal sac; atrium is expanded at the point of connection with the spermatheca. However,
|
||
<taxonomicName id="4C11A8E3FFCC141201DBF8A7FEAEFF32" authorityName="Kolesnikov & Klimov & Khaustov & Pepato & Makarova" authorityYear="2024" box="[151,276,1814,1840]" class="Arachnida" family="Acaridae" genus="Thyreophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="259" phylum="Arthropoda" rank="species" species="subiasi" status="sp. nov.">
|
||
<emphasis id="B9650F72FFCC141201DBF8A7FEAEFF32" box="[151,276,1814,1840]" italics="true" pageId="7" pageNumber="259">Th. subiasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9650F72FFCC14120051F8A6FEC2FF33" bold="true" box="[285,376,1815,1841]" pageId="7" pageNumber="259">
|
||
<taxonomicNameLabel id="A256B209FFCC14120051F8A6FEC2FF33" box="[285,376,1815,1841]" pageId="7" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C11A8E3FFCC141200FEF8A7FD85FF32" baseAuthorityName="Fain" baseAuthorityYear="1982" box="[434,575,1814,1840]" class="Arachnida" family="Acaridae" genus="Thyreophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="259" phylum="Arthropoda" rank="species" species="athiasae">
|
||
<emphasis id="B9650F72FFCC141200FEF8A7FD85FF32" box="[434,575,1814,1840]" italics="true" pageId="7" pageNumber="259">Th. athiasae</emphasis>
|
||
</taxonomicName>
|
||
differ in several key features: (i) the shape of the atrium of spermatheca at the point of connection with the inseminatory canal: thin-walled, cylindrical, with rectangular shape in the cross-section in
|
||
<taxonomicName id="4C11A8E3FFCC14120043F8EFFE36FF7A" authorityName="Kolesnikov & Klimov & Khaustov & Pepato & Makarova" authorityYear="2024" box="[271,396,1886,1912]" class="Arachnida" family="Acaridae" genus="Thyreophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="259" phylum="Arthropoda" rank="species" species="subiasi" status="sp. nov.">
|
||
<emphasis id="B9650F72FFCC14120043F8EFFE36FF7A" box="[271,396,1886,1912]" italics="true" pageId="7" pageNumber="259">Th. subiasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9650F72FFCC141200D9F8EEFE4BFF7B" bold="true" box="[405,497,1887,1913]" pageId="7" pageNumber="259">
|
||
<taxonomicNameLabel id="A256B209FFCC141200D9F8EEFE4BFF7B" box="[405,497,1887,1913]" pageId="7" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="132ACFE5FFCC1412034EF8EFFDE5FF7B" box="[514,607,1886,1913]" captionStart="FIGURE 5" captionStartId="7.[152,255,1060,1084]" captionTargetBox="[154,1432,184,1033]" captionTargetId="figure-19@7.[151,1435,181,1036]" captionTargetPageId="7" captionText="FIGURE 5. Thyreophagus subiasi sp. nov., female holotype (A, B) and male, paratype (C–G):A—subcapitulum, ventral view; B—spermatheca; C—male genitalia, ventral view; D—leg I, dorsal view; E—tarsus I, ventral view; F—leg IV, anterior view; G—tarsus IV, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14595265" httpUri="https://zenodo.org/record/14595265/files/figure.png" pageId="7" pageNumber="259">Figs 5B</figureCitation>
|
||
,
|
||
<figureCitation id="132ACFE5FFCC14120322F8EEFD28FF7B" box="[622,658,1887,1913]" captionStart="FIGURE 7" captionStartId="9.[152,255,1943,1967]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure-19@9.[178,1409,181,1919]" captionTargetPageId="9" captionText="FIGURE 7. Thyreophagus subiasi sp. nov., male paratype (A–C, E, F) and female, holotype (D), DIC images:A—dorsal view; B—ventral view; C—prodorsal shield; D—spermatheca; E—posterior part of opisthosoma, dorsal view; G—posterior part of opisthosoma, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14595271" httpUri="https://zenodo.org/record/14595271/files/figure.png" pageId="7" pageNumber="259">7D</figureCitation>
|
||
) vs. with tip conical thick-walled, narrowing in the cross-section in
|
||
<taxonomicName id="4C11A8E3FFCC141201DBF833FE98FF9E" baseAuthorityName="Fain" baseAuthorityYear="1982" box="[151,290,1922,1948]" class="Arachnida" family="Acaridae" genus="Thyreophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="259" phylum="Arthropoda" rank="species" species="athiasae">
|
||
<emphasis id="B9650F72FFCC141201DBF833FE98FF9E" box="[151,290,1922,1948]" italics="true" pageId="7" pageNumber="259">Th. athiasae</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<figureCitation id="132ACFE5FFCC14120075F833FE29FF9F" box="[313,403,1922,1949]" captionStart="FIGURE 8" captionStartId="10.[152,255,1978,2002]" captionTargetBox="[154,1432,844,1952]" captionTargetId="figure-244@10.[151,1435,841,1955]" captionTargetPageId="10" captionText="FIGURE 8. Thyreophagus athiasae (Fain, 1982), female holotype: A—tarsus I, posterior view; B—tarsus II, anterior view; C—tarsus III, posterior view; D—tarsus IV, posterior view; E—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14595273" httpUri="https://zenodo.org/record/14595273/files/figure.png" pageId="7" pageNumber="259">Figs 8E</figureCitation>
|
||
,
|
||
<figureCitation id="132ACFE5FFCC141200D3F832FE7BFF9F" box="[415,449,1923,1949]" captionStart="FIGURE 9" captionStartId="11.[152,255,1943,1967]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure-19@11.[178,1409,181,1919]" captionTargetPageId="11" captionText="FIGURE 9. Thyreophagus athiasae (Fain, 1982), female holotype, phase-contrast images: A—ventral view; B—prodorsal shield; C—coxal area I–II, ventral view; D—posterior part of opisthosoma; E—spermatheca; F—tarsus I, posterior view; G— tarsus II, anterior view; H—tarsus III, posterior view; I—tarsus IV, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14595275" httpUri="https://zenodo.org/record/14595275/files/figure.png" pageId="7" pageNumber="259">9E</figureCitation>
|
||
); (ii) the shape of spermathecal sac enveloping the atrium: forms a short bottleneck with expanded rims (“flared bottleneck”) in
|
||
<taxonomicName id="4C11A8E3FFCC1412031CF817FD71FFC2" authorityName="Kolesnikov & Klimov & Khaustov & Pepato & Makarova" authorityYear="2024" box="[592,715,1958,1984]" class="Arachnida" family="Acaridae" genus="Thyreophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="259" phylum="Arthropoda" rank="species" species="subiasi" status="sp. nov.">
|
||
<emphasis id="B9650F72FFCC1412031CF817FD71FFC2" box="[592,715,1958,1984]" italics="true" pageId="7" pageNumber="259">Th. subiasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9650F72FFCC1412039EF816FC96FFC3" bold="true" box="[722,812,1959,1985]" pageId="7" pageNumber="259">
|
||
<taxonomicNameLabel id="A256B209FFCC1412039EF816FC96FFC3" box="[722,812,1959,1985]" pageId="7" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
vs. simple in
|
||
<taxonomicName id="4C11A8E3FFCC14120286F817FBEFFFC2" baseAuthorityName="Fain" baseAuthorityYear="1982" box="[970,1109,1958,1984]" class="Arachnida" family="Acaridae" genus="Thyreophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="259" phylum="Arthropoda" rank="species" species="athiasae">
|
||
<emphasis id="B9650F72FFCC14120286F817FBEFFFC2" box="[970,1109,1958,1984]" italics="true" pageId="7" pageNumber="259">Th. athiasae</emphasis>
|
||
</taxonomicName>
|
||
; (iii) tarsal setae
|
||
<emphasis id="B9650F72FFCC14120454F816FA9CFFC2" box="[1304,1318,1959,1984]" italics="true" pageId="7" pageNumber="259">p</emphasis>
|
||
and
|
||
<emphasis id="B9650F72FFCC14120411F816FAD1FFC2" box="[1373,1387,1959,1984]" italics="true" pageId="7" pageNumber="259">q</emphasis>
|
||
I–II are rudimentary in
|
||
<taxonomicName id="4C11A8E3FFCC14120021F87BFE52FFE6" authorityName="Kolesnikov & Klimov & Khaustov & Pepato & Makarova" authorityYear="2024" box="[365,488,1994,2020]" class="Arachnida" family="Acaridae" genus="Thyreophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="259" phylum="Arthropoda" rank="species" species="subiasi" status="sp. nov.">
|
||
<emphasis id="B9650F72FFCC14120021F87BFE52FFE6" box="[365,488,1994,2020]" italics="true" pageId="7" pageNumber="259">Th. subiasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9650F72FFCC141200A3F87AFDF2FFE7" bold="true" box="[495,584,1995,2021]" pageId="7" pageNumber="259">
|
||
<taxonomicNameLabel id="A256B209FFCC141200A3F87AFDF2FFE7" box="[495,584,1995,2021]" pageId="7" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
vs. spiniform in
|
||
<taxonomicName id="4C11A8E3FFCC1412024BF87BFC28FFE6" baseAuthorityName="Fain" baseAuthorityYear="1982" box="[775,914,1994,2020]" class="Arachnida" family="Acaridae" genus="Thyreophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="259" phylum="Arthropoda" rank="species" species="athiasae">
|
||
<emphasis id="B9650F72FFCC1412024BF87BFC28FFE6" box="[775,914,1994,2020]" italics="true" pageId="7" pageNumber="259">Th. athiasae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF6E83E8FFC3141D01D4F8B7FB49FF1C" ID-DOI="http://doi.org/10.5281/zenodo.14595267" ID-Zenodo-Dep="14595267" box="[152,1267,1798,1823]" httpUri="https://zenodo.org/record/14595267/files/figure.png" pageId="8" pageNumber="260" startId="8.[152,255,1798,1822]" targetBox="[151,1436,181,1774]" targetPageId="8" targetType="figure">
|
||
<paragraph id="8BAED360FFC3141D01D4F8B7FB49FF1C" blockId="8.[152,1267,1798,1823]" box="[152,1267,1798,1823]" pageId="8" pageNumber="260">
|
||
<emphasis id="B9650F72FFC3141D01D4F8B7FEA2FF1C" bold="true" box="[152,280,1798,1822]" pageId="8" pageNumber="260">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C11A8E3FFC3141D0053F8B7FE47FF1C" authorityName="Kolesnikov & Klimov & Khaustov & Pepato & Makarova" authorityYear="2024" box="[287,509,1798,1822]" class="Arachnida" family="Acaridae" genus="Thyreophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="260" phylum="Arthropoda" rank="species" species="subiasi" status="sp. nov.">
|
||
<emphasis id="B9650F72FFC3141D0053F8B7FE47FF1C" box="[287,509,1798,1822]" italics="true" pageId="8" pageNumber="260">Thyreophagus subiasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9650F72FFC3141D034FF8B6FDEEFF1D" bold="true" box="[515,596,1799,1823]" pageId="8" pageNumber="260">
|
||
<taxonomicNameLabel id="A256B209FFC3141D034FF8B6FDEEFF1D" box="[515,596,1799,1823]" pageId="8" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female holotype, DIC images: A—dorsal view; B—ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C30B80EBFFC3141D018BF8C0FBA9FFAD" pageId="8" pageNumber="260" type="etymology">
|
||
<paragraph id="8BAED360FFC3141D018BF8C0FBA9FFAD" blockId="8.[151,1437,1905,2039]" pageId="8" pageNumber="260">
|
||
<emphasis id="B9650F72FFC3141D018BF8C0FE85FF89" box="[199,319,1905,1931]" italics="true" pageId="8" pageNumber="260">Etymology</emphasis>
|
||
. The new species is named in honor of Luis S. Subias, the renowned acarologist and specialist on oribatid mites, in recognition of his profound contribution to the study of mites.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30B80EBFFC3141F018BF808FCE0F91F" lastPageId="10" lastPageNumber="262" pageId="8" pageNumber="260" type="biology_ecology">
|
||
<paragraph id="8BAED360FFC3141F018BF808FCE0F91F" blockId="8.[151,1437,1905,2039]" lastBlockId="10.[151,1436,151,285]" lastPageId="10" lastPageNumber="262" pageId="8" pageNumber="260">
|
||
<emphasis id="B9650F72FFC3141D018BF808FEA7FFD1" box="[199,285,1977,2003]" italics="true" pageId="8" pageNumber="260">Habitat</emphasis>
|
||
.
|
||
<taxonomicName id="4C11A8E3FFC3141D0064F808FDA0FFD1" authorityName="Kolesnikov & Klimov & Khaustov & Pepato & Makarova" authorityYear="2024" box="[296,538,1977,2003]" class="Arachnida" family="Acaridae" genus="Thyreophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="260" phylum="Arthropoda" rank="species" species="subiasi" status="sp. nov.">
|
||
<emphasis id="B9650F72FFC3141D0064F808FDA0FFD1" box="[296,538,1977,2003]" italics="true" pageId="8" pageNumber="260">Thyreophagus subiasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9650F72FFC3141D0353F808FDCCFFD1" bold="true" box="[543,630,1977,2003]" pageId="8" pageNumber="260">
|
||
<taxonomicNameLabel id="A256B209FFC3141D0353F808FDCCFFD1" box="[543,630,1977,2003]" pageId="8" pageNumber="260" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has been repeatedly found across several biotopes along the Caspian coast in the Samur Forest. Most mites were collected from wet, sandy soil with cattails and reeds (
|
||
<figureCitation id="132ACFE5FFC3141D05DDF86CFAB1FFF5" box="[1169,1291,2013,2039]" captionStart="FIGURE 1" captionStartId="2.[152,255,1888,1912]" captionTargetBox="[151,1436,417,1865]" captionTargetId="figure-103@2.[151,1436,417,1865]" captionTargetPageId="2" captionText="FIGURE 1. Habitats of Thyreophagus subiasi sp. nov. on the shore of the protected Samur Forest: A—general view of a pebble beach; B—pebble beach with Typha laxmannii; C—sandy beach with Phragmites australis and Typha australis; D— reed thicket (Phragmites australis, Rubus caesius, Elaeagnus angustifolia) near abandoned fish ponds; E—driftwood and sea debris; F—grassy sea debris (Dagestan, Caspian Sea, April 2021)." figureDoi="http://doi.org/10.5281/zenodo.14595255" httpUri="https://zenodo.org/record/14595255/files/figure.png" pageId="8" pageNumber="260">Fig. 1A–D</figureCitation>
|
||
), where their abundance was estimated at 240–480 individuals per square meter.
|
||
<taxonomicName id="4C11A8E3FFC1141F02CEFF26FC41F8B3" authorityName="Kolesnikov & Klimov & Khaustov & Pepato & Makarova" authorityYear="2024" box="[898,1019,151,177]" class="Arachnida" family="Acaridae" genus="Thyreophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="262" phylum="Arthropoda" rank="species" species="subiasi" status="sp. nov.">
|
||
<emphasis id="B9650F72FFC1141F02CEFF26FC41F8B3" box="[898,1019,151,177]" italics="true" pageId="10" pageNumber="262">Th. subiasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9650F72FFC1141F054EFF26FBE1F8B3" bold="true" box="[1026,1115,151,177]" pageId="10" pageNumber="262">
|
||
<taxonomicNameLabel id="A256B209FFC1141F054EFF26FBE1F8B3" box="[1026,1115,151,177]" pageId="10" pageNumber="262" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
was particularly abundant in marine debris, consisting of decomposing land grass and driftwood on wet sand (
|
||
<figureCitation id="132ACFE5FFC1141F055FFF0AFB3EF8D7" box="[1043,1156,187,213]" captionStart="FIGURE 1" captionStartId="2.[152,255,1888,1912]" captionTargetBox="[151,1436,417,1865]" captionTargetId="figure-103@2.[151,1436,417,1865]" captionTargetPageId="2" captionText="FIGURE 1. Habitats of Thyreophagus subiasi sp. nov. on the shore of the protected Samur Forest: A—general view of a pebble beach; B—pebble beach with Typha laxmannii; C—sandy beach with Phragmites australis and Typha australis; D— reed thicket (Phragmites australis, Rubus caesius, Elaeagnus angustifolia) near abandoned fish ponds; E—driftwood and sea debris; F—grassy sea debris (Dagestan, Caspian Sea, April 2021)." figureDoi="http://doi.org/10.5281/zenodo.14595255" httpUri="https://zenodo.org/record/14595255/files/figure.png" pageId="10" pageNumber="262">Fig. 1E, F</figureCitation>
|
||
), with an average density of 32 individuals per liter. In all these biotopes, not only adults but also protonymphs and tritonymphs of
|
||
<taxonomicName id="4C11A8E3FFC1141F046EFF6EFA26F8FB" authorityName="Kolesnikov & Klimov & Khaustov & Pepato & Makarova" authorityYear="2024" box="[1314,1436,223,249]" class="Arachnida" family="Acaridae" genus="Thyreophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="262" phylum="Arthropoda" rank="species" species="subiasi" status="sp. nov.">
|
||
<emphasis id="B9650F72FFC1141F046EFF6EFA26F8FB" box="[1314,1436,223,249]" italics="true" pageId="10" pageNumber="262">Th. subiasi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9650F72FFC1141F01DBFEB2FF4BF91F" bold="true" box="[151,241,259,285]" pageId="10" pageNumber="262">
|
||
<taxonomicNameLabel id="A256B209FFC1141F01DBFEB2FF4BF91F" box="[151,241,259,285]" pageId="10" pageNumber="262" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
were found (larvae and deutonymphs were not found).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |