413 lines
49 KiB
XML
413 lines
49 KiB
XML
<document id="60F2A144D7FDD2698961367823A2F3FC" ID-DOI="10.11646/zootaxa.5556.1.19" ID-ISSN="1175-5326" ID-Zenodo-Dep="14595253" ID-ZooBank="B7D2B6BB-62A4-461A-BA26-55A96FA458BA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1735931024094" checkinUser="plazi" docAuthor="Kolesnikov, Vasiliy B., Klimov, Pavel B., Khaustov, Alexander A., Pepato, Almir & Makarova, Olga L." docDate="2024" docId="03B86276FFC1141901DBFED7FF72FE9B" docLanguage="en" docName="zootaxa.5556.1.19.pdf" docOrigin="Zootaxa 5556 (1)" docSource="https://doi.org/10.11646/zootaxa.5556.1.19" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thyreophagus athiasae" docType="treatment" docVersion="2" lastPageNumber="264" masterDocId="FF811A0EFFCB1415014CFFB1FFBAF802" masterDocTitle="A new arenicolous species of the genus Thyreophagus (Acari: Acaridae) from the Caspian sea coast: Taxonomy and ecological insights" masterLastPageNumber="267" masterPageNumber="252" pageNumber="262" updateTime="1735932494398" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="EEDCB1C62DCAE7B266C57FF917624061">A new arenicolous species of the genus Thyreophagus (Acari: Acaridae) from the Caspian sea coast: Taxonomy and ecological insights</mods:title>
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<mods:namePart id="5DA0D9EECCF4928FC0F378ED8672F5B4">Kolesnikov, Vasiliy B.</mods:namePart>
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<mods:affiliation id="09E1341DA85D5CC423DA3FECEFA8A74D">Papanin Institute for Biology of Inland Waters, Russian Academy of Sciences, 152742 Borok, Yaroslavl Province, Russia. & Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, 625003 Tyumen, Russia.</mods:affiliation>
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<mods:namePart id="0138933931411A5F6D8D1269858A9032">Klimov, Pavel B.</mods:namePart>
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<mods:affiliation id="7BF06062FC8E1462B890755A4AF6D427">Department of Biological Sciences, Purdue University, Lilly Hall of Life Sciences, West Lafayette, IN 47907, USA.</mods:affiliation>
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<mods:namePart id="A016C36EDCE274F6A3732293814BCAF6">Khaustov, Alexander A.</mods:namePart>
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<mods:affiliation id="E0BFD24315A30A2133045114007C53EA">Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, 625003 Tyumen, Russia.</mods:affiliation>
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<mods:namePart id="6B24065963AEBA445720D34CB8973EF6">Pepato, Almir</mods:namePart>
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<mods:affiliation id="4D8089288A145D0C35A682AB5CF3FE60">Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, 625003 Tyumen, Russia. & Universidade Federal de Minas Gerais, Instituto de Ciências Biológicas, Departamento de Zoologia, Laboratório de Sistemática eEvolução de Ácaros Acariformes, Av. Antonio Carlos, 6627, Pampulha, Belo Horizonte-MG ZIP: 31270 - 901, Brazil.</mods:affiliation>
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<mods:namePart id="F0BA9751AD4094E9C20F6D29FA1596D3">Makarova, Olga L.</mods:namePart>
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<mods:affiliation id="02AD1A24B1AB08757CCE9E0247B7D5A7">Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Moscow, Russia.</mods:affiliation>
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<mods:title id="C5C937144E81D7C86C8BDF2A179E2D04">Zootaxa</mods:title>
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<mods:date id="19C4E21C655C6884D92C4B0EB7D5871A">2024</mods:date>
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<heading id="D0E6640CFFC1141F01DBFED7FDF5F982" bold="true" box="[151,591,358,385]" fontSize="11" level="1" pageId="10" pageNumber="262" reason="1">
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<taxonomicName id="4C11A8E3FFC1141F01DBFED7FDF5F982" authority="(Fain, 1982)" baseAuthorityName="Fain" baseAuthorityYear="1982" box="[151,591,358,385]" class="Arachnida" family="Acaridae" genus="Thyreophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="262" phylum="Arthropoda" rank="species" species="athiasae">
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<emphasis id="B9650F72FFC1141F01DBFED7FDF5F982" bold="true" box="[151,591,358,385]" pageId="10" pageNumber="262">
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<emphasis id="B9650F72FFC1141F01DBFED7FE08F983" bold="true" box="[151,434,358,385]" italics="true" pageId="10" pageNumber="262">Thyreophagus athiasae</emphasis>
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(
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<bibRefCitation id="EF80AE91FFC1141F008DFED7FDFDF982" author="Fain, A." box="[449,583,358,384]" pageId="10" pageNumber="262" pagination="1 - 47" refId="ref7152" refString="Fain, A. (1982) Revision des genres Thyreophagus Rondani, 1874 et Michaelopus Fain & Johnston, 1974 (Acari, Acaridae) avec description de neuf especes nouvelles. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique Entomologie, 54 (7), 1 - 47." type="journal article" year="1982">Fain, 1982</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30B80EBFFC1141901DBFE3AFF72FE9B" lastPageId="12" lastPageNumber="264" pageId="10" pageNumber="262" type="description">
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<paragraph id="8BAED360FFC1141F01DBFE3AFEB1F9A7" blockId="10.[151,591,358,421]" box="[151,267,395,421]" pageId="10" pageNumber="262">
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(
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<figureCitation id="132ACFE5FFC1141F01D3FE3AFF5DF9A7" box="[159,231,395,421]" captionStart="FIGURE 8" captionStartId="10.[152,255,1978,2002]" captionTargetBox="[154,1432,844,1952]" captionTargetId="figure-244@10.[151,1435,841,1955]" captionTargetPageId="10" captionText="FIGURE 8. Thyreophagus athiasae (Fain, 1982), female holotype: A—tarsus I, posterior view; B—tarsus II, anterior view; C—tarsus III, posterior view; D—tarsus IV, posterior view; E—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14595273" httpUri="https://zenodo.org/record/14595273/files/figure.png" pageId="10" pageNumber="262">Figs 8</figureCitation>
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,
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<figureCitation id="132ACFE5FFC1141F01B8FE3AFEB9F9A7" box="[244,259,395,421]" captionStart="FIGURE 9" captionStartId="11.[152,255,1943,1967]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure-19@11.[178,1409,181,1919]" captionTargetPageId="11" captionText="FIGURE 9. Thyreophagus athiasae (Fain, 1982), female holotype, phase-contrast images: A—ventral view; B—prodorsal shield; C—coxal area I–II, ventral view; D—posterior part of opisthosoma; E—spermatheca; F—tarsus I, posterior view; G— tarsus II, anterior view; H—tarsus III, posterior view; I—tarsus IV, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14595275" httpUri="https://zenodo.org/record/14595275/files/figure.png" pageId="10" pageNumber="262">9</figureCitation>
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)
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</paragraph>
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<paragraph id="8BAED360FFC1141F01DBFE62FDF1FA5B" blockId="10.[151,1436,467,817]" pageId="10" pageNumber="262">
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<emphasis id="B9650F72FFC1141F01DBFE62FE8FF9EF" box="[151,309,467,493]" italics="true" pageId="10" pageNumber="262">Type material</emphasis>
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:
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<materialsCitation id="3B79D93DFFC1141F000BFE62FD19FA37" collectionCode="IRSNB" country="Belgium" location="Royal Belgian Institute of Natural Sciences" municipality="Institut Royal des Sciences" pageId="10" pageNumber="262" specimenCount="1" stateProvince="Region de Bruxelles-Capitale" typeStatus="holotype">
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<typeStatus id="54AA6DC2FFC1141F000BFE62FE13F9EF" box="[327,425,467,493]" pageId="10" pageNumber="262" type="holotype">holotype</typeStatus>
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(female) from
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<location id="8ECE85BBFFC1141F0312FE62FBE3F9EF" LSID="urn:lsid:plazi:treatment:03B86276FFC1141901DBFED7FF72FE9B:8ECE85BBFFC1141F0312FE62FBE3F9EF" box="[606,1113,467,493]" country="Belgium" municipality="Institut Royal des Sciences" name="Royal Belgian Institute of Natural Sciences" pageId="10" pageNumber="262" stateProvince="Region de Bruxelles-Capitale">Royal Belgian Institute of Natural Sciences</location>
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,
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<collectingRegion id="49D51D82FFC1141F0524FE62FB70F9EF" box="[1128,1226,467,493]" country="Belgium" name="Region de Bruxelles-Capitale" pageId="10" pageNumber="262">Brussels</collectingRegion>
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,
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<collectingCountry id="F30693F0FFC1141F0594FE62FA82F9EF" box="[1240,1336,467,493]" name="Belgium" pageId="10" pageNumber="262">Belgium</collectingCountry>
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(
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<collectingMunicipality id="6BCA491AFFC1141F0407FE62FECAFA13" pageId="10" pageNumber="262">Institut Royal des Sciences</collectingMunicipality>
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naturelles de Belgique) (
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<collectionCode id="ED004BA5FFC1141F03C7FE46FD5EFA13" box="[651,740,503,529]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="10" pageNumber="262">IRSNB</collectionCode>
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): from organic residues found at the bottom of a vase that had contained locally produced lentils, in
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<collectingCountry id="F30693F0FFC1141F0307FDAAFD19FA37" box="[587,675,539,565]" name="Algeria" pageId="10" pageNumber="262">Algeria</collectingCountry>
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</materialsCitation>
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.
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<materialsCitation id="3B79D93DFFC1141F03FCFDAAFDFDFA5B" collectingDate="1949-07-20" collectionCode="IRSNB" country="Morocco" location="Coccids" pageId="10" pageNumber="262" specimenCount="1" typeStatus="paratype">
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<typeStatus id="54AA6DC2FFC1141F03FCFDAAFCA8FA37" box="[688,786,539,565]" pageId="10" pageNumber="262" type="paratype">Paratype</typeStatus>
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(female) from
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<collectionCode id="ED004BA5FFC1141F028AFDAAFBA7FA37" box="[966,1053,539,565]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="10" pageNumber="262">IRSNB</collectionCode>
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: on
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<location id="8ECE85BBFFC1141F051DFDAAFB11FA37" LSID="urn:lsid:plazi:treatment:03B86276FFC1141901DBFED7FF72FE9B:8ECE85BBFFC1141F051DFDAAFB11FA37" box="[1105,1195,539,565]" country="Morocco" name="Coccids" pageId="10" pageNumber="262">Coccids</location>
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<taxonomicName id="4C11A8E3FFC1141F05F9FDAAFE82FA5B" authority="Newm., Rabat" authorityName="Newm., Rabat" class="Insecta" family="Diaspididae" genus="Lepidosaphes" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="262" phylum="Arthropoda" rank="species" species="beekii">
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<emphasis id="B9650F72FFC1141F05F9FDAAFA26FA37" box="[1205,1436,539,565]" italics="true" pageId="10" pageNumber="262">Lepidosaphes beekii</emphasis>
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Newm., Rabat
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</taxonomicName>
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in
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<collectingCountry id="F30693F0FFC1141F0010FD8EFE7EFA5B" box="[348,452,575,601]" name="Morocco" pageId="10" pageNumber="262">Morocco</collectingCountry>
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,
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<date id="FFAFF5A0FFC1141F0083FD8EFDFDFA5B" box="[463,583,575,601]" pageId="10" pageNumber="262" value="1949-07-20">
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||
<collectingDate id="EFEB0C48FFC1141F0083FD8EFDFDFA5B" box="[463,583,575,601]" pageId="10" pageNumber="262" value="1949-07-20">20-7-1949</collectingDate>
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</date>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BAED360FFC1141F018BFDD2FF6EFB33" blockId="10.[151,1436,467,817]" pageId="10" pageNumber="262">
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||
<emphasis id="B9650F72FFC1141F018BFDD2FE83FA7E" box="[199,313,611,636]" italics="true" pageId="10" pageNumber="262">Diagnosis</emphasis>
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. Prodorsal shield with small flat protrus in posterior part, smoothly punctate except lineate pattern on posterior 1/3 part of shield. Supracoxal seta
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<emphasis id="B9650F72FFC1141F03C0FD36FD01FAA3" box="[652,699,647,673]" italics="true" pageId="10" pageNumber="262">elcp</emphasis>
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||
present. Idiosomal setae
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||
<emphasis id="B9650F72FFC1141F0296FD36FC52FAA3" box="[986,1000,647,673]" italics="true" pageId="10" pageNumber="262">h</emphasis>
|
||
<subScript id="1795D125FFC1141F02A4FD28FC4AFAA5" attach="left" box="[1000,1008,665,679]" fontSize="6" pageId="10" pageNumber="262">1</subScript>
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,
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||
<emphasis id="B9650F72FFC1141F02B3FD36FBB7FAA3" box="[1023,1037,647,673]" italics="true" pageId="10" pageNumber="262">h</emphasis>
|
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<subScript id="1795D125FFC1141F0541FD28FBAFFAA5" attach="left" box="[1037,1045,665,679]" fontSize="6" pageId="10" pageNumber="262">2</subScript>
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and
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<emphasis id="B9650F72FFC1141F0502FD36FBE6FAA3" box="[1102,1116,647,673]" italics="true" pageId="10" pageNumber="262">h</emphasis>
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||
<subScript id="1795D125FFC1141F0510FD28FBDEFAA5" attach="left" box="[1116,1124,665,679]" fontSize="6" pageId="10" pageNumber="262">3</subScript>
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||
short. Base of spermatheca short, its length is 1.4 times greater than its width, with thickened walls in the conical part where it connects with the inseminatory canal, ½ its part is hidden in the conical base of the sac of spermatheca without “bottleneck” and extended ring. Paired efferent ducts short, their length approximately 1/3 the length of spermatheca base, with short stem.
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</paragraph>
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||
<caption id="DF6E83E8FFC1141F01D4F80BFC1DFFF4" ID-DOI="http://doi.org/10.5281/zenodo.14595273" ID-Zenodo-Dep="14595273" httpUri="https://zenodo.org/record/14595273/files/figure.png" pageId="10" pageNumber="262" startId="10.[152,255,1978,2002]" targetBox="[154,1432,844,1952]" targetPageId="10" targetType="figure">
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<paragraph id="8BAED360FFC1141F01D4F80BFC1DFFF4" blockId="10.[151,1436,1978,2039]" pageId="10" pageNumber="262">
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<emphasis id="B9650F72FFC1141F01D4F80BFEA0FFD0" bold="true" box="[152,282,1978,2002]" pageId="10" pageNumber="262">FIGURE 8.</emphasis>
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||
<taxonomicName id="4C11A8E3FFC1141F006EF80BFD23FFD1" authority="(Fain, 1982)" baseAuthorityName="Fain" baseAuthorityYear="1982" box="[290,665,1978,2003]" class="Arachnida" family="Acaridae" genus="Thyreophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="262" phylum="Arthropoda" rank="species" species="athiasae">
|
||
<emphasis id="B9650F72FFC1141F006EF80BFDAAFFD0" box="[290,528,1978,2002]" italics="true" pageId="10" pageNumber="262">Thyreophagus athiasae</emphasis>
|
||
(
|
||
<bibRefCitation id="EF80AE91FFC1141F0353F80BFD28FFD1" author="Fain, A." box="[543,658,1978,2003]" pageId="10" pageNumber="262" pagination="1 - 47" refId="ref7152" refString="Fain, A. (1982) Revision des genres Thyreophagus Rondani, 1874 et Michaelopus Fain & Johnston, 1974 (Acari, Acaridae) avec description de neuf especes nouvelles. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique Entomologie, 54 (7), 1 - 47." type="journal article" year="1982">Fain, 1982</bibRefCitation>
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||
)
|
||
</taxonomicName>
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||
, female holotype: A—tarsus I, posterior view; B—tarsus II, anterior view; C—tarsus III, posterior view; D—tarsus IV, posterior view; E—spermatheca.
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||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6E83E8FFC0141E01D4F826FC61FFF5" ID-DOI="http://doi.org/10.5281/zenodo.14595275" ID-Zenodo-Dep="14595275" httpUri="https://zenodo.org/record/14595275/files/figure.png" pageId="11" pageNumber="263" startId="11.[152,255,1943,1967]" targetBox="[178,1409,181,1919]" targetPageId="11" targetType="figure">
|
||
<paragraph id="8BAED360FFC0141E01D4F826FC61FFF5" blockId="11.[151,1436,1943,2039]" pageId="11" pageNumber="263">
|
||
<emphasis id="B9650F72FFC0141E01D4F826FEA1FFAD" bold="true" box="[152,283,1943,1967]" pageId="11" pageNumber="263">FIGURE 9.</emphasis>
|
||
<taxonomicName id="4C11A8E3FFC0141E0069F826FD25FFAD" authority="(Fain, 1982)" baseAuthorityName="Fain" baseAuthorityYear="1982" box="[293,671,1943,1967]" class="Arachnida" family="Acaridae" genus="Thyreophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="263" phylum="Arthropoda" rank="species" species="athiasae">
|
||
<emphasis id="B9650F72FFC0141E0069F826FDAEFFAD" box="[293,532,1943,1967]" italics="true" pageId="11" pageNumber="263">Thyreophagus athiasae</emphasis>
|
||
(
|
||
<bibRefCitation id="EF80AE91FFC0141E0368F826FD22FFAD" author="Fain, A." box="[548,664,1943,1967]" pageId="11" pageNumber="263" pagination="1 - 47" refId="ref7152" refString="Fain, A. (1982) Revision des genres Thyreophagus Rondani, 1874 et Michaelopus Fain & Johnston, 1974 (Acari, Acaridae) avec description de neuf especes nouvelles. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique Entomologie, 54 (7), 1 - 47." type="journal article" year="1982">Fain, 1982</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, female holotype, phase-contrast images: A—ventral view; B—prodorsal shield; C—coxal area I–II, ventral view; D—posterior part of opisthosoma; E—spermatheca; F—tarsus I, posterior view; G— tarsus II, anterior view; H—tarsus III, posterior view; I—tarsus IV, posterior view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAED360FFC71419018BFF26FEC9F8B3" blockId="12.[151,1437,151,1689]" box="[199,371,151,177]" pageId="12" pageNumber="264">
|
||
<emphasis id="B9650F72FFC71419018BFF26FED6F8B3" box="[199,364,151,177]" italics="true" pageId="12" pageNumber="264">Re-description</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAED360FFC71419018BFF0AFF4BF8FB" blockId="12.[151,1437,151,1689]" pageId="12" pageNumber="264">
|
||
Female (
|
||
<typeStatus id="54AA6DC2FFC714190067FF0AFE2EF8D7" box="[299,404,187,213]" pageId="12" pageNumber="264" type="holotype">holotype</typeStatus>
|
||
). Idiosoma elongate, 650 × 250 (
|
||
<typeStatus id="54AA6DC2FFC714190258FF0AFCC6F8D7" box="[788,892,187,213]" pageId="12" pageNumber="264" type="holotype">holotype</typeStatus>
|
||
), 2.6 times longer than wide. Idiosomal cuticle smooth.
|
||
</paragraph>
|
||
<paragraph id="8BAED360FFC71419018BFEB2FB90F967" blockId="12.[151,1437,151,1689]" pageId="12" pageNumber="264">
|
||
Subcapitular setae (
|
||
<emphasis id="B9650F72FFC7141900EBFEB2FE0FF91F" box="[423,437,259,285]" italics="true" pageId="12" pageNumber="264">h</emphasis>
|
||
) long, widened basally; palp tibial setae (
|
||
<emphasis id="B9650F72FFC7141902DEFEB2FC1AF91E" box="[914,928,259,284]" italics="true" pageId="12" pageNumber="264">a</emphasis>
|
||
), lateral dorsal palp tibial setae (
|
||
<emphasis id="B9650F72FFC714190454FEB2FA85F91E" box="[1304,1343,259,284]" italics="true" pageId="12" pageNumber="264">sup</emphasis>
|
||
), dorsal palp tarsal seta (
|
||
<emphasis id="B9650F72FFC71419001CFE96FECBF942" box="[336,369,295,320]" italics="true" pageId="12" pageNumber="264">cm</emphasis>
|
||
) filiform; supracoxal seta
|
||
<emphasis id="B9650F72FFC7141903D7FE96FD70F943" box="[667,714,295,321]" italics="true" pageId="12" pageNumber="264">elcp</emphasis>
|
||
present; terminal palp tarsal solenidion ω short; external part of terminal eupathidium
|
||
<emphasis id="B9650F72FFC7141900C2FEFAFE0EF967" box="[398,436,331,357]" italics="true" pageId="12" pageNumber="264">ul”</emphasis>
|
||
dome-shaped; terminal eupathidium
|
||
<emphasis id="B9650F72FFC71419021AFEFAFCCFF967" box="[854,885,331,357]" italics="true" pageId="12" pageNumber="264">ul’</emphasis>
|
||
small, rounded.
|
||
</paragraph>
|
||
<paragraph id="8BAED360FFC71419018BFEDEFAC6FBA7" blockId="12.[151,1437,151,1689]" pageId="12" pageNumber="264">
|
||
Prodorsal shield (
|
||
<figureCitation id="132ACFE5FFC7141900C3FEDEFE5DF98B" box="[399,487,367,393]" captionStart="FIGURE 9" captionStartId="11.[152,255,1943,1967]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure-19@11.[178,1409,181,1919]" captionTargetPageId="11" captionText="FIGURE 9. Thyreophagus athiasae (Fain, 1982), female holotype, phase-contrast images: A—ventral view; B—prodorsal shield; C—coxal area I–II, ventral view; D—posterior part of opisthosoma; E—spermatheca; F—tarsus I, posterior view; G— tarsus II, anterior view; H—tarsus III, posterior view; I—tarsus IV, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14595275" httpUri="https://zenodo.org/record/14595275/files/figure.png" pageId="12" pageNumber="264">Fig. 9B</figureCitation>
|
||
) 130 long, 100 wide, 1.3 times longer than wide, with setae
|
||
<emphasis id="B9650F72FFC7141905EEFEDEFB0CF98A" box="[1186,1206,367,392]" italics="true" pageId="12" pageNumber="264">vi</emphasis>
|
||
(situated at anterior part of shield, alveoli separated at a distance less than their width), rounded anterolateral incisions, and elongate midlateral incisions (insertion points of setae
|
||
<emphasis id="B9650F72FFC7141903E6FE06FD79F9D2" box="[682,707,439,464]" italics="true" pageId="12" pageNumber="264">ve</emphasis>
|
||
). The back of the shield has a small flat protrusion. Prodorsal shield smoothly punctate except lineate pattern on posterior 1/3 part of shield. Grandjean’s organ (GO) with 10 membranous short finger-shaped processes. Supracoxal setae (
|
||
<emphasis id="B9650F72FFC714190217FE4EFCC5FA1A" box="[859,895,511,536]" italics="true" pageId="12" pageNumber="264">scx</emphasis>
|
||
) smooth, sword-shaped, widened and flattened, tapering at tip. Idiosomal setae (
|
||
<emphasis id="B9650F72FFC7141900B3FD92FDA9FA3E" box="[511,531,547,572]" italics="true" pageId="12" pageNumber="264">vi</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC71419036DFD92FD82FA3E" box="[545,568,547,572]" italics="true" pageId="12" pageNumber="264">se</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC71419030BFD92FDE9FA3E" box="[583,595,547,572]" italics="true" pageId="12" pageNumber="264">c</emphasis>
|
||
<subScript id="1795D125FFC71419031FFD84FDE1FA41" attach="left" box="[595,603,565,579]" fontSize="6" pageId="12" pageNumber="264">p</subScript>
|
||
,
|
||
<emphasis id="B9650F72FFC714190325FD92FDCDFA3F" box="[617,631,547,573]" italics="true" pageId="12" pageNumber="264">d</emphasis>
|
||
<subScript id="1795D125FFC71419033BFD84FDC5FA41" attach="left" box="[631,639,565,579]" fontSize="6" pageId="12" pageNumber="264">2</subScript>
|
||
,
|
||
<emphasis id="B9650F72FFC7141903C1FD92FD23FA3E" box="[653,665,547,572]" italics="true" pageId="12" pageNumber="264">e</emphasis>
|
||
<subScript id="1795D125FFC7141903D5FD84FD1BFA41" attach="left" box="[665,673,565,579]" fontSize="6" pageId="12" pageNumber="264">2</subScript>
|
||
,
|
||
<emphasis id="B9650F72FFC7141903E3FD92FD07FA3F" box="[687,701,547,573]" italics="true" pageId="12" pageNumber="264">h</emphasis>
|
||
<subScript id="1795D125FFC7141903F1FD84FD7FFA41" attach="left" box="[701,709,565,579]" fontSize="6" pageId="12" pageNumber="264">1</subScript>
|
||
,
|
||
<emphasis id="B9650F72FFC71419039FFD92FD5BFA3F" box="[723,737,547,573]" italics="true" pageId="12" pageNumber="264">h</emphasis>
|
||
<subScript id="1795D125FFC7141903ADFD84FD53FA41" attach="left" box="[737,745,565,579]" fontSize="6" pageId="12" pageNumber="264">2</subScript>
|
||
,
|
||
<emphasis id="B9650F72FFC7141903BBFD92FCBFFA3F" box="[759,773,547,573]" italics="true" pageId="12" pageNumber="264">h</emphasis>
|
||
<subScript id="1795D125FFC714190249FD84FCB7FA41" attach="left" box="[773,781,565,579]" fontSize="6" pageId="12" pageNumber="264">3</subScript>
|
||
,
|
||
<emphasis id="B9650F72FFC714190250FD92FC8FFA3E" box="[796,821,547,572]" italics="true" pageId="12" pageNumber="264">ps</emphasis>
|
||
<subScript id="1795D125FFC714190278FD84FC86FA41" attach="left" box="[820,828,565,579]" fontSize="6" pageId="12" pageNumber="264">3</subScript>
|
||
) smooth, filiform and short (except long
|
||
<emphasis id="B9650F72FFC714190447FD92FA98FA3E" box="[1291,1314,547,572]" italics="true" pageId="12" pageNumber="264">se</emphasis>
|
||
) (
|
||
<figureCitation id="132ACFE5FFC714190476FD92FF14FA63" captionStart="FIGURE 9" captionStartId="11.[152,255,1943,1967]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure-19@11.[178,1409,181,1919]" captionTargetPageId="11" captionText="FIGURE 9. Thyreophagus athiasae (Fain, 1982), female holotype, phase-contrast images: A—ventral view; B—prodorsal shield; C—coxal area I–II, ventral view; D—posterior part of opisthosoma; E—spermatheca; F—tarsus I, posterior view; G— tarsus II, anterior view; H—tarsus III, posterior view; I—tarsus IV, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14595275" httpUri="https://zenodo.org/record/14595275/files/figure.png" pageId="12" pageNumber="264">Figs 9A, D</figureCitation>
|
||
); opisthosomal gland openings slightly anteriad setal bases
|
||
<emphasis id="B9650F72FFC714190206FDF6FCECFA62" box="[842,854,583,608]" italics="true" pageId="12" pageNumber="264">e</emphasis>
|
||
<subScript id="1795D125FFC71419021AFDE8FCE4FA65" attach="left" box="[854,862,601,615]" fontSize="6" pageId="12" pageNumber="264">2</subScript>
|
||
. Three pairs of fundamental cupules (
|
||
<emphasis id="B9650F72FFC71419044BFDF6FAA7FA62" box="[1287,1309,583,608]" italics="true" pageId="12" pageNumber="264">ia</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC714190467FDF6FAFDFA62" box="[1323,1351,583,608]" italics="true" pageId="12" pageNumber="264">im</emphasis>
|
||
and
|
||
<emphasis id="B9650F72FFC714190431FDF6FA29FA63" box="[1405,1427,583,609]" italics="true" pageId="12" pageNumber="264">ih</emphasis>
|
||
) present,
|
||
<emphasis id="B9650F72FFC7141901BAFDDAFEB6FA86" box="[246,268,619,644]" italics="true" pageId="12" pageNumber="264">ip</emphasis>
|
||
not observed. Ventral idiosoma with four pairs of coxal setae (
|
||
<emphasis id="B9650F72FFC714190281FDDAFC53FA87" box="[973,1001,619,645]" italics="true" pageId="12" pageNumber="264">1a</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC7141902BBFDDAFBA9FA87" box="[1015,1043,619,645]" italics="true" pageId="12" pageNumber="264">3a</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC71419056DFDDAFB87FA87" box="[1057,1085,619,645]" italics="true" pageId="12" pageNumber="264">4a</emphasis>
|
||
and
|
||
<emphasis id="B9650F72FFC714190538FDDAFB2AFA87" box="[1140,1168,619,645]" italics="true" pageId="12" pageNumber="264">4b</emphasis>
|
||
) and one pair of genital setae (
|
||
<emphasis id="B9650F72FFC7141901ADFD3EFF55FAAA" box="[225,239,655,680]" italics="true" pageId="12" pageNumber="264">g</emphasis>
|
||
). Shape of coxal sclerites as in
|
||
<figureCitation id="132ACFE5FFC714190313FD3EFD5AFAAB" box="[607,736,655,681]" captionStart="FIGURE 9" captionStartId="11.[152,255,1943,1967]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure-19@11.[178,1409,181,1919]" captionTargetPageId="11" captionText="FIGURE 9. Thyreophagus athiasae (Fain, 1982), female holotype, phase-contrast images: A—ventral view; B—prodorsal shield; C—coxal area I–II, ventral view; D—posterior part of opisthosoma; E—spermatheca; F—tarsus I, posterior view; G— tarsus II, anterior view; H—tarsus III, posterior view; I—tarsus IV, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14595275" httpUri="https://zenodo.org/record/14595275/files/figure.png" pageId="12" pageNumber="264">Figures 9C</figureCitation>
|
||
. Genital region situated between coxal fields III and IV (
|
||
<figureCitation id="132ACFE5FFC714190423FD3EFF07FACF" captionStart="FIGURE 9" captionStartId="11.[152,255,1943,1967]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure-19@11.[178,1409,181,1919]" captionTargetPageId="11" captionText="FIGURE 9. Thyreophagus athiasae (Fain, 1982), female holotype, phase-contrast images: A—ventral view; B—prodorsal shield; C—coxal area I–II, ventral view; D—posterior part of opisthosoma; E—spermatheca; F—tarsus I, posterior view; G— tarsus II, anterior view; H—tarsus III, posterior view; I—tarsus IV, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14595275" httpUri="https://zenodo.org/record/14595275/files/figure.png" pageId="12" pageNumber="264">Fig. 9A</figureCitation>
|
||
); genital valves form an inverted Y; epigynal and medial apodemes well-developed. Diameter of genital papillae approximately 0.4 the length of genital setae. Anal opening terminal (
|
||
<figureCitation id="132ACFE5FFC7141902D4FD66FC48FAF3" box="[920,1010,727,753]" captionStart="FIGURE 9" captionStartId="11.[152,255,1943,1967]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure-19@11.[178,1409,181,1919]" captionTargetPageId="11" captionText="FIGURE 9. Thyreophagus athiasae (Fain, 1982), female holotype, phase-contrast images: A—ventral view; B—prodorsal shield; C—coxal area I–II, ventral view; D—posterior part of opisthosoma; E—spermatheca; F—tarsus I, posterior view; G— tarsus II, anterior view; H—tarsus III, posterior view; I—tarsus IV, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14595275" httpUri="https://zenodo.org/record/14595275/files/figure.png" pageId="12" pageNumber="264">Fig. 9D</figureCitation>
|
||
). Copulatory tube small. Inseminatory canal (
|
||
<emphasis id="B9650F72FFC7141901AEFD4AFF4CFB16" box="[226,246,763,788]" italics="true" pageId="12" pageNumber="264">ic</emphasis>
|
||
) of spermatheca long, slender tube-like, uniformly wide.Atrium (
|
||
<emphasis id="B9650F72FFC71419029FFD4AFC56FB17" box="[979,1004,763,789]" italics="true" pageId="12" pageNumber="264">hs</emphasis>
|
||
) (
|
||
<figureCitation id="132ACFE5FFC71419054FFD4AFBE6FB17" box="[1027,1116,763,789]" captionStart="FIGURE 8" captionStartId="10.[152,255,1978,2002]" captionTargetBox="[154,1432,844,1952]" captionTargetId="figure-244@10.[151,1435,841,1955]" captionTargetPageId="10" captionText="FIGURE 8. Thyreophagus athiasae (Fain, 1982), female holotype: A—tarsus I, posterior view; B—tarsus II, anterior view; C—tarsus III, posterior view; D—tarsus IV, posterior view; E—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14595273" httpUri="https://zenodo.org/record/14595273/files/figure.png" pageId="12" pageNumber="264">Figs 8E</figureCitation>
|
||
,
|
||
<figureCitation id="132ACFE5FFC71419052BFD4AFB3DFB17" box="[1127,1159,763,789]" captionStart="FIGURE 9" captionStartId="11.[152,255,1943,1967]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure-19@11.[178,1409,181,1919]" captionTargetPageId="11" captionText="FIGURE 9. Thyreophagus athiasae (Fain, 1982), female holotype, phase-contrast images: A—ventral view; B—prodorsal shield; C—coxal area I–II, ventral view; D—posterior part of opisthosoma; E—spermatheca; F—tarsus I, posterior view; G— tarsus II, anterior view; H—tarsus III, posterior view; I—tarsus IV, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14595275" httpUri="https://zenodo.org/record/14595275/files/figure.png" pageId="12" pageNumber="264">9E</figureCitation>
|
||
) of spermatheca present, short, its length 1.4 times longer than its width, with thickened walls in the conical part at point of connection with inseminatory canal.Approximately half of atrium enveloped by spermathecal sac. Sac of spermatheca is not forming a short bottleneck with extended rims (flared bottleneck) around atrium (
|
||
<figureCitation id="132ACFE5FFC714190284FCD6FB9AFB83" box="[968,1056,871,897]" captionStart="FIGURE 8" captionStartId="10.[152,255,1978,2002]" captionTargetBox="[154,1432,844,1952]" captionTargetId="figure-244@10.[151,1435,841,1955]" captionTargetPageId="10" captionText="FIGURE 8. Thyreophagus athiasae (Fain, 1982), female holotype: A—tarsus I, posterior view; B—tarsus II, anterior view; C—tarsus III, posterior view; D—tarsus IV, posterior view; E—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14595273" httpUri="https://zenodo.org/record/14595273/files/figure.png" pageId="12" pageNumber="264">Fig. 8E</figureCitation>
|
||
). Paired sclerites of efferent ducts (
|
||
<emphasis id="B9650F72FFC7141901EDFC3AFF01FBA7" box="[161,187,907,933]" italics="true" pageId="12" pageNumber="264">ed</emphasis>
|
||
) short, their lengths approximately 1/2 the length of spermathecal atrium, V-shaped, with short stem at base.
|
||
</paragraph>
|
||
<paragraph id="8BAED360FFC71419018BFC1EFA8EFDC3" blockId="12.[151,1437,151,1689]" pageId="12" pageNumber="264">
|
||
Legs short, all segments free. Trochanters I–III each with long, filiform seta,
|
||
<emphasis id="B9650F72FFC714190507FC1EFBD0FBCA" box="[1099,1130,943,968]" italics="true" pageId="12" pageNumber="264">pR</emphasis>
|
||
I–II,
|
||
<emphasis id="B9650F72FFC7141905FCFC1EFB76FBCA" box="[1200,1228,943,968]" italics="true" pageId="12" pageNumber="264">sR</emphasis>
|
||
III; trochanter IV without setae. Femoral setation 1-1-0-1; setae
|
||
<emphasis id="B9650F72FFC7141903ECFC62FD04FBEE" box="[672,702,979,1004]" italics="true" pageId="12" pageNumber="264">vF</emphasis>
|
||
I–II and
|
||
<emphasis id="B9650F72FFC71419026AFC62FCF0FBEE" box="[806,842,979,1004]" italics="true" pageId="12" pageNumber="264">wF</emphasis>
|
||
IV long, filiform. Genual setation 2-2-0-0; setae
|
||
<emphasis id="B9650F72FFC714190438FC62FA26FBEF" box="[1396,1436,979,1005]" italics="true" pageId="12" pageNumber="264">mG</emphasis>
|
||
and
|
||
<emphasis id="B9650F72FFC714190189FC46FF5CFC13" box="[197,230,1015,1041]" italics="true" pageId="12" pageNumber="264">cG</emphasis>
|
||
I–II long, filiform; seta
|
||
<emphasis id="B9650F72FFC7141900BCFC46FDA8FC13" box="[496,530,1015,1041]" italics="true" pageId="12" pageNumber="264">nG</emphasis>
|
||
III absent. Tibial setation 2-2-1-1; setae
|
||
<emphasis id="B9650F72FFC71419029DFC46FC55FC13" box="[977,1007,1015,1041]" italics="true" pageId="12" pageNumber="264">hT</emphasis>
|
||
I-II absent; setae
|
||
<emphasis id="B9650F72FFC7141905FEFC46FB6AFC12" box="[1202,1232,1015,1040]" italics="true" pageId="12" pageNumber="264">gT</emphasis>
|
||
I–II and
|
||
<emphasis id="B9650F72FFC71419047FFC46FAF5FC13" box="[1331,1359,1015,1041]" italics="true" pageId="12" pageNumber="264">kT</emphasis>
|
||
III–IV elongate, somewhat spiniform. Tarsal setation 10-10-10-10; pretarsi consist of hooked empodial claws attached to short paired condylophores. Tarsus I and II (
|
||
<figureCitation id="132ACFE5FFC7141903CAFB8EFD47FC5B" box="[646,765,1087,1113]" captionStart="FIGURE 8" captionStartId="10.[152,255,1978,2002]" captionTargetBox="[154,1432,844,1952]" captionTargetId="figure-244@10.[151,1435,841,1955]" captionTargetPageId="10" captionText="FIGURE 8. Thyreophagus athiasae (Fain, 1982), female holotype: A—tarsus I, posterior view; B—tarsus II, anterior view; C—tarsus III, posterior view; D—tarsus IV, posterior view; E—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14595273" httpUri="https://zenodo.org/record/14595273/files/figure.png" pageId="12" pageNumber="264">Figs 8A,B</figureCitation>
|
||
,
|
||
<figureCitation id="132ACFE5FFC714190244FB8EFCFAFC5B" box="[776,832,1087,1113]" captionStart="FIGURE 9" captionStartId="11.[152,255,1943,1967]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure-19@11.[178,1409,181,1919]" captionTargetPageId="11" captionText="FIGURE 9. Thyreophagus athiasae (Fain, 1982), female holotype, phase-contrast images: A—ventral view; B—prodorsal shield; C—coxal area I–II, ventral view; D—posterior part of opisthosoma; E—spermatheca; F—tarsus I, posterior view; G— tarsus II, anterior view; H—tarsus III, posterior view; I—tarsus IV, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14595275" httpUri="https://zenodo.org/record/14595275/files/figure.png" pageId="12" pageNumber="264">9F,G</figureCitation>
|
||
) with setae
|
||
<emphasis id="B9650F72FFC71419028BFB8EFC5AFC5A" box="[967,992,1087,1112]" italics="true" pageId="12" pageNumber="264">ra</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC7141902A1FB8EFBB9FC5B" box="[1005,1027,1087,1113]" italics="true" pageId="12" pageNumber="264">la</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC71419055DFB8EFBA3FC5B" box="[1041,1049,1087,1113]" italics="true" pageId="12" pageNumber="264">f</emphasis>
|
||
and
|
||
<emphasis id="B9650F72FFC714190503FB8EFBE7FC5B" box="[1103,1117,1087,1113]" italics="true" pageId="12" pageNumber="264">d</emphasis>
|
||
filiform,
|
||
<emphasis id="B9650F72FFC714190585FB8EFB6FFC5A" box="[1225,1237,1087,1112]" italics="true" pageId="12" pageNumber="264">e</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC7141905AFFB8EFB4BFC5A" box="[1251,1265,1087,1112]" italics="true" pageId="12" pageNumber="264">u</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC7141905B3FB8EFAB1FC5A" box="[1279,1291,1087,1112]" italics="true" pageId="12" pageNumber="264">v</emphasis>
|
||
spiniform (
|
||
<emphasis id="B9650F72FFC7141904DCFB8EFA26FC5A" box="[1424,1436,1087,1112]" italics="true" pageId="12" pageNumber="264">v</emphasis>
|
||
larger that
|
||
<emphasis id="B9650F72FFC71419005CFBD2FEA4FC7E" box="[272,286,1123,1148]" italics="true" pageId="12" pageNumber="264">u</emphasis>
|
||
),
|
||
<emphasis id="B9650F72FFC714190079FBD2FEF9FC7E" box="[309,323,1123,1148]" italics="true" pageId="12" pageNumber="264">p</emphasis>
|
||
and
|
||
<emphasis id="B9650F72FFC714190035FBD2FE3DFC7E" box="[377,391,1123,1148]" italics="true" pageId="12" pageNumber="264">q</emphasis>
|
||
represented by small spines,
|
||
<emphasis id="B9650F72FFC71419039CFBD2FD61FC7E" box="[720,731,1123,1148]" italics="true" pageId="12" pageNumber="264">s</emphasis>
|
||
small spines (legs I) or flattened, button-shaped (legs II); setae
|
||
<emphasis id="B9650F72FFC7141901DBFB36FF02FCA2" box="[151,184,1159,1184]" italics="true" pageId="12" pageNumber="264">wa</emphasis>
|
||
present, spiniform. Tarsus III and IV (
|
||
<figureCitation id="132ACFE5FFC71419033EFB36FD51FCA3" box="[626,747,1159,1185]" captionStart="FIGURE 8" captionStartId="10.[152,255,1978,2002]" captionTargetBox="[154,1432,844,1952]" captionTargetId="figure-244@10.[151,1435,841,1955]" captionTargetPageId="10" captionText="FIGURE 8. Thyreophagus athiasae (Fain, 1982), female holotype: A—tarsus I, posterior view; B—tarsus II, anterior view; C—tarsus III, posterior view; D—tarsus IV, posterior view; E—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14595273" httpUri="https://zenodo.org/record/14595273/files/figure.png" pageId="12" pageNumber="264">Figs 8C,D</figureCitation>
|
||
,
|
||
<figureCitation id="132ACFE5FFC7141903B4FB36FC96FCA3" box="[760,812,1159,1185]" captionStart="FIGURE 9" captionStartId="11.[152,255,1943,1967]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure-19@11.[178,1409,181,1919]" captionTargetPageId="11" captionText="FIGURE 9. Thyreophagus athiasae (Fain, 1982), female holotype, phase-contrast images: A—ventral view; B—prodorsal shield; C—coxal area I–II, ventral view; D—posterior part of opisthosoma; E—spermatheca; F—tarsus I, posterior view; G— tarsus II, anterior view; H—tarsus III, posterior view; I—tarsus IV, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14595275" httpUri="https://zenodo.org/record/14595275/files/figure.png" pageId="12" pageNumber="264">9H,I</figureCitation>
|
||
) with setae
|
||
<emphasis id="B9650F72FFC7141902F4FB36FC7AFCA3" box="[952,960,1159,1185]" italics="true" pageId="12" pageNumber="264">f</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC71419029CFB36FC64FCA3" box="[976,990,1159,1185]" italics="true" pageId="12" pageNumber="264">d</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC7141902A1FB36FC42FCA2" box="[1005,1016,1159,1184]" italics="true" pageId="12" pageNumber="264">r</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC714190544FB36FBA1FCA2" box="[1032,1051,1159,1184]" italics="true" pageId="12" pageNumber="264">w</emphasis>
|
||
filiform,
|
||
<emphasis id="B9650F72FFC7141905C6FB36FB2CFCA2" box="[1162,1174,1159,1184]" italics="true" pageId="12" pageNumber="264">e</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC7141905EAFB36FB0BFCA2" box="[1190,1201,1159,1184]" italics="true" pageId="12" pageNumber="264">s</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC71419058DFB36FB75FCA2" box="[1217,1231,1159,1184]" italics="true" pageId="12" pageNumber="264">u</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC714190593FB36FB51FCA2" box="[1247,1259,1159,1184]" italics="true" pageId="12" pageNumber="264">v</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC7141905B7FB36FAB3FCA2" box="[1275,1289,1159,1184]" italics="true" pageId="12" pageNumber="264">p</emphasis>
|
||
,
|
||
<emphasis id="B9650F72FFC714190455FB36FA9DFCA2" box="[1305,1319,1159,1184]" italics="true" pageId="12" pageNumber="264">q</emphasis>
|
||
spiniform (
|
||
<emphasis id="B9650F72FFC7141901EDFB1AFF15FCC6" box="[161,175,1195,1220]" italics="true" pageId="12" pageNumber="264">u</emphasis>
|
||
shorter than other ventral spines). Solenidion ω
|
||
<subScript id="1795D125FFC71419039FFB0CFD61FCC9" attach="left" box="[723,731,1213,1227]" fontSize="6" pageId="12" pageNumber="264">1</subScript>
|
||
on tarsus I cylindrical, with clavate apex, straight; solenidion ω
|
||
<subScript id="1795D125FFC7141901E6FB50FF08FCED" attach="left" box="[170,178,1249,1263]" fontSize="6" pageId="12" pageNumber="264">1</subScript>
|
||
on tarsus II simple, cylindrical, with clavate apex, curved, the slightly longer than ω
|
||
<subScript id="1795D125FFC714190521FB50FBCFFCED" attach="left" box="[1133,1141,1249,1263]" fontSize="6" pageId="12" pageNumber="264">1</subScript>
|
||
on tarsus I. Solenidion ω
|
||
<subScript id="1795D125FFC7141904D8FB50FA26FCED" attach="left" box="[1428,1436,1249,1263]" fontSize="6" pageId="12" pageNumber="264">2</subScript>
|
||
on tarsus I shorter than ω
|
||
<subScript id="1795D125FFC7141900F0FAB4FE7EFD11" attach="left" box="[444,452,1285,1299]" fontSize="6" pageId="12" pageNumber="264">1</subScript>
|
||
, cylindrical, with rounded apex, situated slightly distal to ω
|
||
<subScript id="1795D125FFC71419053BFAB4FBC5FD11" attach="right" box="[1143,1151,1285,1299]" fontSize="6" pageId="12" pageNumber="264">1</subScript>
|
||
. Solenidion ω
|
||
<subScript id="1795D125FFC714190468FAB4FA96FD11" attach="left" box="[1316,1324,1285,1299]" fontSize="6" pageId="12" pageNumber="264">3</subScript>
|
||
on tarsus I cylindrical, with rounded tip, slightly shorter than ω
|
||
<subScript id="1795D125FFC7141903B1FA98FCBFFD35" attach="left" box="[765,773,1321,1335]" fontSize="6" pageId="12" pageNumber="264">1</subScript>
|
||
and slightly longer than ω
|
||
<subScript id="1795D125FFC714190575FA98FBFBFD35" attach="left" box="[1081,1089,1321,1335]" fontSize="6" pageId="12" pageNumber="264">2</subScript>
|
||
. Famulus (ε) of tarsus I wide, short, with broadly rounded apex, widest at middle. Solenidia φ of tibiae I–III elongate, tapering, well extending beyond apices of respective tarsi with ambulacra; solenidion φ IV shorter, shorter than tarsus IV (with ambulacrum). Solenidia σ’ and σ’’ on genu I elongate, tapering, σ’ is longer than σ’’, slightly not reaching base of φ I. Solenidion σ on genu II more than 5–6 times longer than its width, with rounded tip. Solenidion σ of genu III absent.
|
||
</paragraph>
|
||
<paragraph id="8BAED360FFC71419018BFA7AFD27FDE7" blockId="12.[151,1437,151,1689]" box="[199,669,1483,1509]" pageId="12" pageNumber="264">Male and phoretic deutonymph unknown.</paragraph>
|
||
<paragraph id="8BAED360FFC71419018BFA5EFF72FE9B" blockId="12.[151,1437,151,1689]" pageId="12" pageNumber="264">
|
||
<emphasis id="B9650F72FFC71419018BFA5EFE93FE0B" box="[199,297,1519,1545]" italics="true" pageId="12" pageNumber="264">Remarks</emphasis>
|
||
. The
|
||
<typeStatus id="54AA6DC2FFC714190024FA5EFE70FE0B" box="[360,458,1519,1545]" pageId="12" pageNumber="264" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C11A8E3FFC7141900A3FA5EFDC3FE0B" baseAuthorityName="Fain" baseAuthorityYear="1982" box="[495,633,1519,1545]" class="Arachnida" family="Acaridae" genus="Thyreophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="264" phylum="Arthropoda" rank="species" species="athiasae">
|
||
<emphasis id="B9650F72FFC7141900A3FA5EFDC3FE0B" box="[495,633,1519,1545]" italics="true" pageId="12" pageNumber="264">Th. athiasae</emphasis>
|
||
</taxonomicName>
|
||
was well-illustrated and described in
|
||
<bibRefCitation id="EF80AE91FFC714190553FA5EFB1FFE0B" author="Fain, A." box="[1055,1189,1519,1545]" pageId="12" pageNumber="264" pagination="1 - 47" refId="ref7152" refString="Fain, A. (1982) Revision des genres Thyreophagus Rondani, 1874 et Michaelopus Fain & Johnston, 1974 (Acari, Acaridae) avec description de neuf especes nouvelles. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique Entomologie, 54 (7), 1 - 47." type="journal article" year="1982">Fain (1982)</bibRefCitation>
|
||
. However, illustration in
|
||
<bibRefCitation id="EF80AE91FFC7141901FBF9A2FE8DFE2F" author="Fain, A." box="[183,311,1555,1581]" pageId="12" pageNumber="264" pagination="1 - 47" refId="ref7152" refString="Fain, A. (1982) Revision des genres Thyreophagus Rondani, 1874 et Michaelopus Fain & Johnston, 1974 (Acari, Acaridae) avec description de neuf especes nouvelles. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique Entomologie, 54 (7), 1 - 47." type="journal article" year="1982">Fain (1982</bibRefCitation>
|
||
, fig. 58) show that tarsi I and II lack
|
||
<emphasis id="B9650F72FFC7141903BCF9A2FD44FE2E" box="[752,766,1555,1580]" italics="true" pageId="12" pageNumber="264">p</emphasis>
|
||
and
|
||
<emphasis id="B9650F72FFC714190276F9A2FCF2FE2E" box="[826,840,1555,1580]" italics="true" pageId="12" pageNumber="264">q</emphasis>
|
||
, while
|
||
<emphasis id="B9650F72FFC7141902EDF9A2FC16FE2E" box="[929,940,1555,1580]" italics="true" pageId="12" pageNumber="264">s</emphasis>
|
||
is spiniform. However, a closer inspection revealed the presence of well-developed, albeit small,
|
||
<emphasis id="B9650F72FFC7141903B1F986FCB1FE52" box="[765,779,1591,1616]" italics="true" pageId="12" pageNumber="264">p</emphasis>
|
||
and
|
||
<emphasis id="B9650F72FFC71419020FF986FCEBFE52" box="[835,849,1591,1616]" italics="true" pageId="12" pageNumber="264">q</emphasis>
|
||
spines on tarsi I–II (
|
||
<figureCitation id="132ACFE5FFC714190570F986FB01FE53" box="[1084,1211,1591,1617]" captionStart="FIGURE 8" captionStartId="10.[152,255,1978,2002]" captionTargetBox="[154,1432,844,1952]" captionTargetId="figure-244@10.[151,1435,841,1955]" captionTargetPageId="10" captionText="FIGURE 8. Thyreophagus athiasae (Fain, 1982), female holotype: A—tarsus I, posterior view; B—tarsus II, anterior view; C—tarsus III, posterior view; D—tarsus IV, posterior view; E—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14595273" httpUri="https://zenodo.org/record/14595273/files/figure.png" pageId="12" pageNumber="264">Figs 8A, B</figureCitation>
|
||
,
|
||
<figureCitation id="132ACFE5FFC714190584F986FAB0FE53" box="[1224,1290,1591,1617]" captionStart="FIGURE 9" captionStartId="11.[152,255,1943,1967]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure-19@11.[178,1409,181,1919]" captionTargetPageId="11" captionText="FIGURE 9. Thyreophagus athiasae (Fain, 1982), female holotype, phase-contrast images: A—ventral view; B—prodorsal shield; C—coxal area I–II, ventral view; D—posterior part of opisthosoma; E—spermatheca; F—tarsus I, posterior view; G— tarsus II, anterior view; H—tarsus III, posterior view; I—tarsus IV, posterior view." figureDoi="http://doi.org/10.5281/zenodo.14595275" httpUri="https://zenodo.org/record/14595275/files/figure.png" pageId="12" pageNumber="264">9F, G</figureCitation>
|
||
). Spine
|
||
<emphasis id="B9650F72FFC71419042AF986FACBFE52" box="[1382,1393,1591,1616]" italics="true" pageId="12" pageNumber="264">s</emphasis>
|
||
I is short and spiniform on tarsus I (as originally described), however, on tarsus II
|
||
<emphasis id="B9650F72FFC7141902B3F9EAFBB0FE76" box="[1023,1034,1627,1652]" italics="true" pageId="12" pageNumber="264">s</emphasis>
|
||
is flattened and button-shaped (
|
||
<figureCitation id="132ACFE5FFC714190423F9EAFF01FE9B" captionStart="FIGURE 8" captionStartId="10.[152,255,1978,2002]" captionTargetBox="[154,1432,844,1952]" captionTargetId="figure-244@10.[151,1435,841,1955]" captionTargetPageId="10" captionText="FIGURE 8. Thyreophagus athiasae (Fain, 1982), female holotype: A—tarsus I, posterior view; B—tarsus II, anterior view; C—tarsus III, posterior view; D—tarsus IV, posterior view; E—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14595273" httpUri="https://zenodo.org/record/14595273/files/figure.png" pageId="12" pageNumber="264">Fig. 8B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |