treatments-xml/data/A3/12/19/A312190FEC13FF85509F3CCDBF0CFAE6.xml
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<document id="ACEFC593F1F8C1FB3998CA97EF2D1623" ID-CLB-Dataset="32076" ID-DOI="10.11646/zootaxa.4320.3.6" ID-GBIF-Dataset="e80fc464-0c2a-4003-a867-ab8b5b3b399e" ID-ISSN="1175-5326" ID-Zenodo-Dep="1051951" ID-ZooBank="Ca50E5A5-B417-4000-B67D-5F6028Da9154" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1505712517045" checkinUser="plazi" docAuthor="Bocak, Ladislav" docDate="2017" docId="A312190FEC13FF85509F3CCDBF0CFAE6" docLanguage="en" docName="zootaxa.4320.3.6.pdf" docOrigin="Zootaxa 4320 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cladophorus craterensis Bocak 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="511" masterDocId="5F2B6177EC15FF8350083D11BB5CFFD4" masterDocTitle="Species delimitation of colour polymorphic Cladophorus (Coleoptera: Lycidae) from New Guinea" masterLastPageNumber="522" masterPageNumber="505" pageNumber="511" updateTime="1698468596363" updateUser="ExternalLinkService">
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<mods:title id="0B3F4D5EE81239145DA9716AB063E710">Species delimitation of colour polymorphic Cladophorus (Coleoptera: Lycidae) from New Guinea</mods:title>
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<mods:namePart id="C2801DB48D7AFA9CA8B73D4B2269455B">Bocak, Ladislav</mods:namePart>
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<mods:title id="502421672623C929B803AC022CB0D76A">Zootaxa</mods:title>
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<mods:date id="435E560E4738FA61D519698260106ACC">2017</mods:date>
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<mods:number id="F79BE50381B55B40056BEDC0E0D8EBF3">2017-09-18</mods:number>
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<treatment id="A312190FEC13FF85509F3CCDBF0CFAE6" ID-DOI="http://doi.org/10.5281/zenodo.6018022" ID-GBIF-Taxon="134090577" ID-Zenodo-Dep="6018022" LSID="urn:lsid:plazi:treatment:A312190FEC13FF85509F3CCDBF0CFAE6" httpUri="http://treatment.plazi.org/id/A312190FEC13FF85509F3CCDBF0CFAE6" lastPageNumber="511" pageId="6" pageNumber="511">
<subSubSection id="63A1FB92EC13FF85509F3CCDBAC2FDC3" pageId="6" pageNumber="511" type="nomenclature">
<paragraph id="2B04A819EC13FF85509F3CCDB97FFE22" blockId="6.[151,547,476,535]" box="[151,547,476,502]" pageId="6" pageNumber="511">
<heading id="704C1F75EC13FF85509F3CCDB97FFE22" bold="true" box="[151,547,476,502]" fontSize="11" level="1" pageId="6" pageNumber="511" reason="1">
<emphasis id="19CF740BEC13FF85509F3CCDB97FFE22" bold="true" box="[151,547,476,502]" pageId="6" pageNumber="511">
<taxonomicName id="ECBBD39AEC13FF85509F3CCDBAE3FE22" authority="Bocak, 2017" authorityName="Bocak" authorityYear="2017" box="[151,447,476,502]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="511" phylum="Arthropoda" rank="species" species="craterensis" status="sp. nov.">
<emphasis id="19CF740BEC13FF85509F3CCDBAE3FE22" bold="true" box="[151,447,476,502]" italics="true" pageId="6" pageNumber="511">Cladophorus craterensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="02FCC970EC13FF8551CD3CCDB97FFE22" box="[453,547,476,502]" pageId="6" pageNumber="511" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="2B04A819EC13FF85509F3CEFBAC2FDC3" blockId="6.[151,547,476,535]" box="[151,414,510,535]" pageId="6" pageNumber="511">
(
<figureCitation id="B380B49CEC13FF8550973CEFBA45FDC3" box="[159,281,510,535]" captionStart="FIGURES 6 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="6" pageNumber="511">Figs. 8, 22</figureCitation>
,
<figureCitation id="B380B49CEC13FF85512D3CEEBA1FFDC3" box="[293,323,511,535]" captionStart="FIGURES 33 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 52. Elytron. 33 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="6" pageNumber="511">38</figureCitation>
,
<figureCitation id="B380B49CEC13FF8551463CEFBACAFDC3" box="[334,406,510,535]" captionStart="FIGURES 53 78" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[174,1418,196,1884]" captionTargetId="figure@13.[171,1423,193,1887]" captionTargetPageId="13" captionText="FIGURES 53 78. Male genitalia ventrally and laterally. 53 54, Cladophorus bicolor Kleine; 55 56, C. boceki sp. nov.; 57 58, C. craterensis sp. nov.; 59 60, C. haiaensis sp. nov.; 61 62, C. humeralis Kleine; 63 64, C. kailakiensis sp. nov.; 65 66, C. manokwarensis sp. nov.; 67 68, C. mindikensis sp. nov.; 69 70, C. motykai sp. nov.; 71 72, C. pallescens sp. nov.; 73 74, C. praecipuus Kleine; 75 76, C. riedeli sp. nov.; 77 78, C. wasiorensis sp. nov. Scales 0.5 mm." httpUri="https://zenodo.org/record/1051961/files/figure.png" pageId="6" pageNumber="511">5758</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC13FF85509F3F57BBA4FD57" pageId="6" pageNumber="511" type="materials_examined">
<paragraph id="2B04A819EC13FF85509F3F57BBA4FD57" blockId="6.[151,1437,582,1330]" pageId="6" pageNumber="511">
<materialsCitation id="9BD3A244EC13FF85509F3F57BBA4FD57" ID-GBIF-Occurrence="1638540615" collectionCode="LMBC" country="Papua New Guinea" elevation="780" latitude="-6.7" location="Crater Mt." longLatPrecision="1304" longitude="145.0" municipality="Haia" pageId="6" pageNumber="511" specimenCount="2" typeStatus="holotype">
<emphasis id="19CF740BEC13FF85509F3F57BA16FD8B" bold="true" box="[151,330,582,607]" pageId="6" pageNumber="511">
<typeStatus id="F40016BBEC13FF85509F3F57BB8EFD8B" box="[151,210,582,607]" pageId="6" pageNumber="511">Type</typeStatus>
material.
</emphasis>
<typeStatus id="F40016BBEC13FF85515F3F57BA9FFD8B" box="[343,451,582,607]" pageId="6" pageNumber="511" type="holotype">Holotype</typeStatus>
. Male.
<collectingCountry id="53ACE889EC13FF8552293F56B854FD8B" box="[545,776,582,607]" name="Papua New Guinea" pageId="6" pageNumber="511">Papua New Guinea</collectingCountry>
,
<location id="2E64FEC2EC13FF8553113F56B8CAFD8A" LSID="urn:lsid:plazi:treatment:A312190FEC13FF85509F3CCDBF0CFAE6:2E64FEC2EC13FF8553113F56B8CAFD8A" box="[793,918,583,607]" country="Papua New Guinea" latitude="-6.7" longLatPrecision="1304" longitude="145.0" municipality="Haia" name="Crater Mt." pageId="6" pageNumber="511">Crater Mt.</location>
,
<collectingMunicipality id="CB603263EC13FF8553AF3F57B883FD8B" box="[935,991,582,607]" pageId="6" pageNumber="511">Haia</collectingMunicipality>
,
<geoCoordinate id="4E8FCEDEEC13FF8553F83F57BF17FD8B" box="[1008,1099,582,607]" direction="south" orientation="latitude" pageId="6" pageNumber="511" precision="925" value="-6.7">06°42´S</geoCoordinate>
<geoCoordinate id="4E8FCEDEEC13FF8554513F57BF9BFD8B" box="[1113,1223,582,607]" direction="east" orientation="longitude" pageId="6" pageNumber="511" precision="925" value="145.0">145°00´E</geoCoordinate>
,
<elevation id="A0964F2AEC13FF8554D03F56BE79FD8A" box="[1240,1317,583,607]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="6" pageNumber="511" unit="m" value="780.0">
<quantity id="EC4305FCEC13FF8554D03F56BE79FD8A" box="[1240,1317,583,607]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="6" pageNumber="511" unit="m" value="780.0">780 m</quantity>
</elevation>
(
<accessionNumber id="34E835FAEC13FF8555313F56BEC4FD8B" box="[1337,1432,583,607]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A00171" pageId="6" pageNumber="511" type="EnaNcbi">A00171</accessionNumber>
,
<collectionCode id="4DAA30DCEC13FF85509F3F7ABBB0FD57" box="[151,236,619,643]" pageId="6" pageNumber="511">LMBC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC13FF8550CF3F9FBADAFCE3" pageId="6" pageNumber="511" type="diagnosis">
<paragraph id="2B04A819EC13FF8550CF3F9FBADAFCE3" blockId="6.[151,1437,582,1330]" pageId="6" pageNumber="511">
<emphasis id="19CF740BEC13FF8550CF3F9FBA1DFD73" bold="true" box="[199,321,654,679]" pageId="6" pageNumber="511">Diagnosis.</emphasis>
<taxonomicName id="ECBBD39AEC13FF8551423F9FB93CFD72" authorityName="Bocak" authorityYear="2017" box="[330,608,654,678]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="511" phylum="Arthropoda" rank="species" species="craterensis">
<emphasis id="19CF740BEC13FF8551423F9FB93CFD72" box="[330,608,654,678]" italics="true" pageId="6" pageNumber="511">Cladophorus craterensis</emphasis>
</taxonomicName>
belongs to the group of mostly light-yellow coloured species such as
<taxonomicName id="ECBBD39AEC13FF85558A3F9EBA4FFD1E" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="511" phylum="Arthropoda" rank="species" species="praecipuus">
<emphasis id="19CF740BEC13FF85558A3F9EBA4FFD1E" italics="true" pageId="6" pageNumber="511">C. praecipuus</emphasis>
</taxonomicName>
,
<taxonomicName id="ECBBD39AEC13FF8551293FA2BAD7FD1F" authorityName="Bocak" authorityYear="2017" box="[289,395,691,715]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="511" phylum="Arthropoda" rank="species" species="riedeli">
<emphasis id="19CF740BEC13FF8551293FA2BAD7FD1F" box="[289,395,691,715]" italics="true" pageId="6" pageNumber="511">C. riedeli</emphasis>
</taxonomicName>
, and
<taxonomicName id="ECBBD39AEC13FF8551C23FA2B91FFD1F" authorityName="Bocak" authorityYear="2017" box="[458,579,691,715]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="511" phylum="Arthropoda" rank="species" species="motykai">
<emphasis id="19CF740BEC13FF8551C23FA2B91FFD1F" box="[458,579,691,715]" italics="true" pageId="6" pageNumber="511">C. motykai</emphasis>
</taxonomicName>
. The phylogenetic analysis showed that these species are not closely related (
<figureCitation id="B380B49CEC13FF8550973FC7BBBEFD3B" box="[159,226,726,751]" captionStart="FIGURES 4 5. 4" captionStartId="3.[151,264,1220,1243]" captionTargetBox="[159,1417,255,1198]" captionTargetId="figure@3.[151,1436,255,1210]" captionTargetPageId="3" captionText="FIGURES 4 5. 4, Phylogenetic hypothesis of Papuan Cladophorus inferred from the cox 1 mtDNA dataset using maximum likelihood criterion. 5, Cladophorus sp., general appearance." httpUri="https://zenodo.org/record/1051955/files/figure.png" pageId="6" pageNumber="511">Fig. 4</figureCitation>
) despite the quite similar shape of their male genitalia (
<figureCitation id="B380B49CEC13FF8553453FC7BF2FFD3B" box="[845,1139,726,751]" captionStart="FIGURES 53 78" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[174,1418,196,1884]" captionTargetId="figure@13.[171,1423,193,1887]" captionTargetPageId="13" captionText="FIGURES 53 78. Male genitalia ventrally and laterally. 53 54, Cladophorus bicolor Kleine; 55 56, C. boceki sp. nov.; 57 58, C. craterensis sp. nov.; 59 60, C. haiaensis sp. nov.; 61 62, C. humeralis Kleine; 63 64, C. kailakiensis sp. nov.; 65 66, C. manokwarensis sp. nov.; 67 68, C. mindikensis sp. nov.; 69 70, C. motykai sp. nov.; 71 72, C. pallescens sp. nov.; 73 74, C. praecipuus Kleine; 75 76, C. riedeli sp. nov.; 77 78, C. wasiorensis sp. nov. Scales 0.5 mm." httpUri="https://zenodo.org/record/1051961/files/figure.png" pageId="6" pageNumber="511">Fig. 5758, 6970, 7376</figureCitation>
). The closest relative of
<taxonomicName id="ECBBD39AEC13FF85558A3FC6BA4FFCC6" authorityName="Bocak" authorityYear="2017" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="511" phylum="Arthropoda" rank="species" species="craterensis">
<emphasis id="19CF740BEC13FF85558A3FC6BA4FFCC6" italics="true" pageId="6" pageNumber="511">C. craterensis</emphasis>
</taxonomicName>
is
<taxonomicName id="ECBBD39AEC13FF85513A3FEABAF7FCC7" authorityName="Bocak" authorityYear="2017" box="[306,427,763,787]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="511" phylum="Arthropoda" rank="species" species="motykai">
<emphasis id="19CF740BEC13FF85513A3FEABAF7FCC7" box="[306,427,763,787]" italics="true" pageId="6" pageNumber="511">C. motykai</emphasis>
</taxonomicName>
and these two species differ in short lamellae of
<taxonomicName id="ECBBD39AEC13FF8553C43FEABF3BFCC6" authorityName="Bocak" authorityYear="2017" box="[972,1127,763,786]" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="511" phylum="Arthropoda" rank="species" species="craterensis">
<emphasis id="19CF740BEC13FF8553C43FEABF3BFCC6" box="[972,1127,763,786]" italics="true" pageId="6" pageNumber="511">C. craterensis</emphasis>
</taxonomicName>
and the long lamellae of
<taxonomicName id="ECBBD39AEC13FF85558A3FEABBACFCE2" authorityName="Bocak" authorityYear="2017" class="Insecta" family="Lycidae" genus="Cladophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="511" phylum="Arthropoda" rank="species" species="motykai">
<emphasis id="19CF740BEC13FF85558A3FEABBACFCE2" italics="true" pageId="6" pageNumber="511">C. motykai</emphasis>
</taxonomicName>
(
<figureCitation id="B380B49CEC13FF8550F73E0FBA25FCE3" box="[255,377,798,823]" captionStart="FIGURES 6 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="6" pageNumber="511">Figs. 8, 14</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC13FF8550CF3E53B84EFB3D" pageId="6" pageNumber="511" type="description">
<paragraph id="2B04A819EC13FF8550CF3E53BF3EFB4B" blockId="6.[151,1437,582,1330]" pageId="6" pageNumber="511">
<emphasis id="19CF740BEC13FF8550CF3E53BA05FC8F" bold="true" box="[199,345,834,859]" pageId="6" pageNumber="511">Description.</emphasis>
Male. Body medium-sized,
<quantity id="EC4305FCEC13FF8552AE3E52B852FC8E" box="[678,782,835,859]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="6" pageNumber="511" unit="mm" value="13.3">13.3 mm</quantity>
long, dark brown to black, pronotum and elytra almost completely uniformly yellow, only very narrow part of humeral margin and the apex slightly darkened (
<figureCitation id="B380B49CEC13FF8555323E77BED3FCAB" box="[1338,1423,870,895]" captionStart="FIGURES 33 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 52. Elytron. 33 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="6" pageNumber="511">Fig. 38</figureCitation>
), scutellum black. Head small, with small hemispherically prominent eyes, frontal interocular distance 1.78 times maximum eye diameter; antennae long, reaching two thirds of elytral length, flabellate, antennomere 3 with lamella 1.4 times longer than stem of antennomere, lamella attached to apical part of antennomere (
<figureCitation id="B380B49CEC13FF8554C43EC3BE48FC3F" box="[1228,1300,978,1003]" captionStart="FIGURES 6 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="6" pageNumber="511">Fig. 8</figureCitation>
). Pronotum transverse, 1.5 times wider than long at midline, lateral margins conspicuously elevated, frontal margin slightly rounded, frontal angles obtuse, inconspicuous, lateral margins straight, posterior angles prominent, postero-lateral carinae weak in middle of pronotal disc, median anterior carina weak (
<figureCitation id="B380B49CEC13FF8553B3392FBF4DFB83" box="[955,1041,1086,1111]" captionStart="FIGURES 6 32" captionStartId="11.[151,264,1702,1725]" captionTargetBox="[184,1399,341,1662]" captionTargetId="figure@11.[181,1405,339,1690]" captionTargetPageId="11" captionText="FIGURES 6 32. Basal antennomeres, males. 6, Cladophorus bicolor Kleine; 7, C. boceki sp. nov.; 8, C. craterensis sp. nov.; 9, C. haiaensis sp. nov.; 10, C. humeralis Kleine; 11, C. kailakiensis sp. nov.; 12, C. manokwarensis sp. nov.; 13, C. mindikensis sp. nov.; 14, C. praecipuus Kleine; 15, C. motykai sp. nov.; 16, C. riedeli sp. nov.; 17, C. pallescens sp. nov.; 18, C. wasiorensis sp. nov. Terminal antennomeres, male. 19, C. haiaensis sp. nov. Pronotum. 20, C. bicolor Kleine; 21, C. boceki sp. nov., 22, C. craterensis sp. nov.; 23, C. haiaensis sp. nov.; 24, C. humeralis Kleine; 25, C. kailakiensis sp. nov.; 26, C. manokwarensis sp. nov.; 27, C. mindikensis sp. nov.; 28, C. motykai sp. nov.; 29, C. pallescens sp. nov.; 30, C. praecipuus Kleine; 31, C. riedeli sp. nov.; 32, C. wasiorensis sp. nov. a, length of lamella; b, length of the stem of antennomere 3. Scales: antennae 1.5 mm; pronota 0.5 mm." httpUri="https://zenodo.org/record/1051957/files/figure.png" pageId="6" pageNumber="511">Fig. 22</figureCitation>
). Elytra flat, very slightly widened posteriorly, transverse costae weak, partly covered by pubescence, elytral areoles slightly transverse (
<figureCitation id="B380B49CEC13FF85553F3973BED3FBAF" box="[1335,1423,1122,1147]" captionStart="FIGURES 33 52" captionStartId="12.[151,264,1876,1899]" captionTargetBox="[174,1412,193,1843]" captionTargetId="figure@12.[174,1418,193,1871]" captionTargetPageId="12" captionText="FIGURES 33 52. Elytron. 33 36, Cladophorus bicolor Kleine; 37, C. boceki sp. nov.; 38, C. craterensis sp. nov.; 39, C. haiaensis sp. nov.; 40, C. humeralis Kleine; 41, C. kailakiensis sp. nov.; 42, C. manokwarensis sp. nov.; 43, C. mindikensis sp. nov.; 44 45, C. motykai sp. nov.; 46, C. pallescens sp. nov.; 47, C. praecipuus Kleine; 48, C. riedeli sp. nov.; 49, C. wasiorensis sp. nov. Structure of elytral costae in C. bicolor Kleine. 50, humeral part of the elytron; 51, transition between the dark and light colored part of the elytron; 52, apical part of the elytron. Scales: elytron 1.5 mm; elytra detail 0.2 mm." httpUri="https://zenodo.org/record/1051959/files/figure.png" pageId="6" pageNumber="511">Fig. 38</figureCitation>
). Phallus moderately slender, parallel-sided, gradually acuminate apically (
<figureCitation id="B380B49CEC13FF8553C43997BF0AFB4B" box="[972,1110,1158,1183]" captionStart="FIGURES 53 78" captionStartId="13.[151,264,1895,1918]" captionTargetBox="[174,1418,196,1884]" captionTargetId="figure@13.[171,1423,193,1887]" captionTargetPageId="13" captionText="FIGURES 53 78. Male genitalia ventrally and laterally. 53 54, Cladophorus bicolor Kleine; 55 56, C. boceki sp. nov.; 57 58, C. craterensis sp. nov.; 59 60, C. haiaensis sp. nov.; 61 62, C. humeralis Kleine; 63 64, C. kailakiensis sp. nov.; 65 66, C. manokwarensis sp. nov.; 67 68, C. mindikensis sp. nov.; 69 70, C. motykai sp. nov.; 71 72, C. pallescens sp. nov.; 73 74, C. praecipuus Kleine; 75 76, C. riedeli sp. nov.; 77 78, C. wasiorensis sp. nov. Scales 0.5 mm." httpUri="https://zenodo.org/record/1051961/files/figure.png" pageId="6" pageNumber="511">Figs. 5758</figureCitation>
).
</paragraph>
<paragraph id="2B04A819EC13FF8550CF39BBB84EFB3D" blockId="6.[151,1437,582,1330]" pageId="6" pageNumber="511">
<emphasis id="19CF740BEC13FF8550CF39BBBA21FB17" bold="true" box="[199,381,1194,1219]" pageId="6" pageNumber="511">Measurements.</emphasis>
BL
<quantity id="EC4305FCEC13FF8551A739BAB949FB16" box="[431,533,1195,1219]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="6" pageNumber="511" unit="mm" value="13.3">13.3 mm</quantity>
, PL
<quantity id="EC4305FCEC13FF85524039BAB9F3FB16" box="[584,687,1195,1219]" metricMagnitude="-3" metricUnit="m" metricValue="1.74" pageId="6" pageNumber="511" unit="mm" value="1.74">1.74 mm</quantity>
, PW
<quantity id="EC4305FCEC13FF8552E339BBB80EFB16" box="[747,850,1194,1219]" metricMagnitude="-3" metricUnit="m" metricValue="2.58" pageId="6" pageNumber="511" unit="mm" value="2.58">2.58 mm</quantity>
, WH
<quantity id="EC4305FCEC13FF85539B39BBB8A5FB16" box="[915,1017,1194,1219]" metricMagnitude="-3" metricUnit="m" metricValue="1.56" pageId="6" pageNumber="511" unit="mm" value="1.56">1.56 mm</quantity>
, Edist
<quantity id="EC4305FCEC13FF85544E39BBBFF0FB16" box="[1094,1196,1194,1219]" metricMagnitude="-4" metricUnit="m" metricValue="9.6" pageId="6" pageNumber="511" unit="mm" value="0.96">0.96 mm</quantity>
, Ediam
<quantity id="EC4305FCEC13FF85550039BBBE32FB16" box="[1288,1390,1194,1219]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="6" pageNumber="511" unit="mm" value="0.54">0.54 mm</quantity>
, LE
<quantity id="EC4305FCEC13FF85509F39C3BBA0FB3D" box="[151,252,1234,1258]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="6" pageNumber="511" unit="mm" value="11.1">11.1 mm</quantity>
, antennomere 3: stem
<quantity id="EC4305FCEC13FF8551F039C3B90CFB3D" box="[504,592,1234,1258]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="511" unit="mm" value="1.2">1.2 mm</quantity>
, lamella
<quantity id="EC4305FCEC13FF8552BD39C0B852FB3D" box="[693,782,1233,1258]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="511" unit="mm" value="1.6">1.6 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC13FF8550CF39E4B8DEFADA" box="[199,898,1269,1294]" pageId="6" pageNumber="511" type="distribution">
<paragraph id="2B04A819EC13FF8550CF39E4B8DEFADA" blockId="6.[151,1437,582,1330]" box="[199,898,1269,1294]" pageId="6" pageNumber="511">
<emphasis id="19CF740BEC13FF8550CF39E4BA3CFADA" bold="true" box="[199,352,1269,1294]" pageId="6" pageNumber="511">Distribution.</emphasis>
Eastern New
<collectingCountry id="53ACE889EC13FF8551F639E4B90EFADA" box="[510,594,1269,1294]" name="Guinea-Bissau" pageId="6" pageNumber="511">Guinea</collectingCountry>
,
<collectingRegion id="E97F66FBEC13FF85525539E4B87CFADA" box="[605,800,1269,1294]" country="Papua New Guinea" name="Chimbu" pageId="6" pageNumber="511">Chimbu province</collectingRegion>
(
<figureCitation id="B380B49CEC13FF85532739E4B828FADA" box="[815,884,1269,1294]" captionStart="FIGURES 1 3. 1 2" captionStartId="2.[151,264,1396,1419]" captionTargetBox="[151,1436,339,1375]" captionTargetId="figure@2.[151,1436,339,1375]" captionTargetPageId="2" captionText="FIGURES 1 3. 1 2, General appearance of Cladophorus bicolor in the Central mountains of New Guinea. 3, Sampled localities and the distribution of Cladophorus species" httpUri="https://zenodo.org/record/1051953/files/figure.png" pageId="6" pageNumber="511">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="63A1FB92EC13FF8550CF3808BF0CFAE6" box="[199,1104,1305,1330]" pageId="6" pageNumber="511" type="etymology">
<paragraph id="2B04A819EC13FF8550CF3808BF0CFAE6" blockId="6.[151,1437,582,1330]" box="[199,1104,1305,1330]" pageId="6" pageNumber="511">
<emphasis id="19CF740BEC13FF8550CF3808BA12FAE6" bold="true" box="[199,334,1305,1330]" pageId="6" pageNumber="511">Etymology.</emphasis>
The epithet refers to the Crater Mts., the
<typeStatus id="F40016BBEC13FF855315380BB811FAE6" box="[797,845,1306,1330]" pageId="6" pageNumber="511">type</typeStatus>
locality of the species.
</paragraph>
</subSubSection>
</treatment>
</document>