213 lines
26 KiB
XML
213 lines
26 KiB
XML
<document id="54A72A14EBBF1B45480FA1D8BF9E22AA" ID-DOI="10.5281/zenodo.213507" ID-GBIF-Dataset="38892d63-388e-47c6-b23f-e18e16a1af14" ID-ISSN="1175-5326" ID-Zenodo-Dep="213507" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460475539065" checkinUser="plazi" docAuthor="Pérez-González, Sergio & Zaballos, Juan P." docDate="2012" docId="0A5887DCFF85FA39FF68ED1E91FBAB31" docLanguage="en" docName="zt03279p062.pdf" docOrigin="Zootaxa 3279" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Typhlocharis quadridentata Coiffait 1969" docType="treatment" docVersion="8" lastPageNumber="56" masterDocId="F661FFA4FF82FA33FFFFEB72923AA91C" masterDocTitle="Re-description of two species of Typhlocharis (Coleoptera: Carabidae: Anillini) and revision of the models of female genitalia within the genus" masterLastPageNumber="62" masterPageNumber="46" pageNumber="53" updateTime="1698306628870" updateUser="plazi">
|
||
<mods:mods id="658D9015903C64598DF2A7219F230F06" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="95DBFB56708ACCADD7DC6C5DB786A9F0">
|
||
<mods:title id="4EF23CCC027F8A36C916487C212A53C0">Re-description of two species of Typhlocharis (Coleoptera: Carabidae: Anillini) and revision of the models of female genitalia within the genus</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="8CD40D6280CADDE191E15587BF577D2B" type="personal">
|
||
<mods:role id="74BF72E304D3983135F01BFA4B9929E1">
|
||
<mods:roleTerm id="CF308A0F57344E34BA143C225590A1CC">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="C43CF49EE23537026BA12F67AF246937">Pérez-González, Sergio</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="7A44724074F9CC14892FC7BDF4819E67" type="personal">
|
||
<mods:role id="60703BF8F13D712011CFE7AEBE4C695A">
|
||
<mods:roleTerm id="FF83916090E19A6564D309A493EF40B2">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="24D8879E2B95226CA9B39174E2FB651E">Zaballos, Juan P.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="1CBE922BC861C2C5ACC58F5817EA1AA2">text</mods:typeOfResource>
|
||
<mods:relatedItem id="7A6A6DB6D1E8ED42FABE8FB873D5D5A6" type="host">
|
||
<mods:titleInfo id="9D350DB3779B95A98A3C4F9861D6C7B0">
|
||
<mods:title id="BBF0ED6E213CD5EF2FD0B174DE3AF619">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="C075ADB05D5DB9696E401AE1F8AB4B41">
|
||
<mods:date id="A6BB38495ECBE58CF20277479A02E0F9">2012</mods:date>
|
||
<mods:detail id="641CBB0FF00849282C20787B7D671932" type="volume">
|
||
<mods:number id="94A0AC2DBF4475360E0A521FBBE6F827">3279</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="4D5CB1C40876C4A3DD01190360B2998C" unit="page">
|
||
<mods:start id="7C53B73FEE8C02B3712B23EC3BA28881">46</mods:start>
|
||
<mods:end id="2F5FB2E647CED1E7F97347828EBC384E">62</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="F50F9E7E3CEAAF788ABE8B4E42C8DA27">journal article</mods:classification>
|
||
<mods:identifier id="4B80EE0DDD4C11E4562A01FB06495272" type="DOI">10.5281/zenodo.213507</mods:identifier>
|
||
<mods:identifier id="BA9A601BB6ACCB0D5F50C350194C1E69" type="GBIF-Dataset">38892d63-388e-47c6-b23f-e18e16a1af14</mods:identifier>
|
||
<mods:identifier id="C84C5AED910188ECF0C4A28D8FF590FE" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="6E8F2AA3AB3D09E0DA4E202684BAEC54" type="Zenodo-Dep">213507</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="0A5887DCFF85FA39FF68ED1E91FBAB31" ID-DOI="http://doi.org/10.5281/zenodo.5623765" ID-GBIF-Taxon="119516184" ID-Zenodo-Dep="5623765" LSID="urn:lsid:plazi:treatment:0A5887DCFF85FA39FF68ED1E91FBAB31" httpUri="http://treatment.plazi.org/id/0A5887DCFF85FA39FF68ED1E91FBAB31" lastPageId="10" lastPageNumber="56" pageId="7" pageNumber="53">
|
||
<subSubSection id="CAEB6541FF85FA34FF68ED1E92DFAFB4" pageId="7" pageNumber="53" type="nomenclature">
|
||
<paragraph id="824E36CAFF85FA34FF68ED1E90A8AF99" blockId="7.[151,658,1644,1704]" box="[151,658,1644,1669]" pageId="7" pageNumber="53">
|
||
<heading id="D90681A6FF85FA34FF68ED1E90A8AF99" bold="true" box="[151,658,1644,1669]" fontSize="10" level="2" pageId="7" pageNumber="53" reason="2">
|
||
<taxonomicName id="45F14D49FF85FA34FF68ED1E90A8AF99" ID-CoL="59VKJ" authority="Coiffait, 1969" authorityName="Coiffait" authorityYear="1969" box="[151,658,1644,1669]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="53" phylum="Arthropoda" rank="species" species="quadridentata">
|
||
<emphasis id="B085EAD8FF85FA34FF68ED1E90A8AF99" bold="true" box="[151,658,1644,1669]" pageId="7" pageNumber="53">
|
||
<emphasis id="B085EAD8FF85FA34FF68ED1E93ECAF99" bold="true" box="[151,470,1644,1669]" italics="true" pageId="7" pageNumber="53">Typhlocharis quadridentata</emphasis>
|
||
(
|
||
<bibRefCitation id="E6604B3BFF85FA34FE1BED1E90B3AF99" author="Coiffait" box="[484,649,1644,1669]" pageId="7" pageNumber="53" refString="Coiffait, H. (1969) Nouveaux Anillini du Maroc et du Sud de la peninsule iberique. Bulletin de la Societe des Sciences Naturelles et Physiques du Maroc, 48, 55 - 56." type="journal article" year="1969">Coiffait, 1969</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="824E36CAFF85FA34FF68EDE292DFAFB4" blockId="7.[151,658,1644,1704]" box="[151,229,1680,1704]" pageId="7" pageNumber="53">(fig. 6)</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CAEB6541FF85FA34FF68EDAA93ADAE9C" pageId="7" pageNumber="53" type="materials_examined">
|
||
<paragraph id="824E36CAFF85FA34FF68EDAA93ADAE9C" blockId="7.[151,1436,1752,2029]" pageId="7" pageNumber="53">
|
||
<emphasis id="B085EAD8FF85FA34FF68EDAA933CAFED" bold="true" box="[151,262,1752,1777]" pageId="7" pageNumber="53">Material.</emphasis>
|
||
<typeStatus id="5D4A8868FF85FA34FEEFEDAA9392AFEC" box="[272,424,1752,1776]" pageId="7" pageNumber="53" type="holotype">HOLOTYPE</typeStatus>
|
||
: 1 3
|
||
<collectingCountry id="FAE6765AFF85FA34FE12EDAA9075AFEC" box="[493,591,1752,1776]" name="Portugal" pageId="7" pageNumber="53">Portugal</collectingCountry>
|
||
, Algarve, Estoï, H. Coiffait 3-64 (
|
||
<collectionCode id="E4E0AE0FFF85FA34FC20EDAA9671AFEC" box="[991,1099,1752,1776]" pageId="7" pageNumber="53">MNHNP</collectionCode>
|
||
);
|
||
<typeStatus id="5D4A8868FF85FA34FB9EEDAB96CBAFED" box="[1121,1265,1753,1777]" pageId="7" pageNumber="53" type="paratype">PARATYPE</typeStatus>
|
||
: 1 3
|
||
<collectingCountry id="FAE6765AFF85FA34FAC9EDAA97A2AFEC" box="[1334,1432,1752,1776]" name="Portugal" pageId="7" pageNumber="53">Portugal</collectingCountry>
|
||
, Algarve, Alportel, H. Coiffait 3-64 (
|
||
<collectionCode id="E4E0AE0FFF85FA34FDCFED8E90A6AE08" box="[560,668,1788,1812]" pageId="7" pageNumber="53">MNHNP</collectionCode>
|
||
).
|
||
<date id="F64F100AFF85FA34FD50ED8E9131AE08" box="[687,779,1788,1813]" pageId="7" pageNumber="53" value="1933-04-02">4 33, 2</date>
|
||
Ƥ Ƥ
|
||
<collectingCountry id="FAE6765AFF85FA34FCB2ED8E9195AE08" box="[845,943,1788,1812]" name="Portugal" pageId="7" pageNumber="53">Portugal</collectingCountry>
|
||
, Algarve, Sta. Barbara de Nexe, H. Coiffait
|
||
<date id="F64F100AFF85FA34FF68EC5292D5AE24" box="[151,239,1824,1848]" pageId="7" pageNumber="53" value="1970-02-19">19-2-70</date>
|
||
(
|
||
<collectionCode id="E4E0AE0FFF85FA34FF01EC529350AE24" box="[254,362,1824,1848]" pageId="7" pageNumber="53">MNHNP</collectionCode>
|
||
); 1 3, 2 Ƥ Ƥ
|
||
<collectingCountry id="FAE6765AFF85FA34FDFCEC52905EAE24" box="[515,612,1824,1848]" name="Portugal" pageId="7" pageNumber="53">Portugal</collectingCountry>
|
||
, Algarve, Boliquiéme, H. Coiffait, 11-70 (
|
||
<collectionCode id="E4E0AE0FFF85FA34FBBFEC529696AE24" box="[1088,1196,1824,1848]" pageId="7" pageNumber="53">MNHNP</collectionCode>
|
||
); 1 3 Sta. Barbara de Nexe
|
||
<date id="F64F100AFF85FA34FF25EC369368AE40" box="[218,338,1860,1884]" pageId="7" pageNumber="53" value="1970-02-12">12-2-1970</date>
|
||
, H. Coiffait, Coll. Zaballos (
|
||
<collectionCode id="E4E0AE0FFF85FA34FD66EC3690E5AE40" LSID="urn:lsid:biocol.org:col:34854" box="[665,735,1860,1884]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="7" pageNumber="53">UCM</collectionCode>
|
||
); 1 Ƥ
|
||
<collectingCountry id="FAE6765AFF85FA34FCD8EC3691B3AE40" box="[807,905,1860,1884]" name="Portugal" pageId="7" pageNumber="53">Portugal</collectingCountry>
|
||
, Algarve, Moncarapacho 12-1965, H. Coiffait, Coll. Zaballos (
|
||
<collectionCode id="E4E0AE0FFF85FA34FEBAEC1A93B1AE9C" LSID="urn:lsid:biocol.org:col:34854" box="[325,395,1896,1920]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="7" pageNumber="53">UCM</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CAEB6541FF85FA3BFF36ECFE9198A803" lastPageId="8" lastPageNumber="54" pageId="7" pageNumber="53" type="diagnosis">
|
||
<paragraph id="824E36CAFF85FA3BFF36ECFE9198A803" blockId="7.[151,1436,1752,2029]" lastBlockId="8.[151,1437,151,826]" lastPageId="8" lastPageNumber="54" pageId="7" pageNumber="53">
|
||
<emphasis id="B085EAD8FF85FA34FF36ECFE9379AEB9" bold="true" box="[201,323,1932,1957]" pageId="7" pageNumber="53">Diagnosis.</emphasis>
|
||
Anopthtalmous, very small endogean beetle. Parallel body, subrectangular and narrow with pale brown, microreticulated tegument and scattered pubescence. Vertex without
|
||
<emphasis id="B085EAD8FF85FA34FC02ECC096ABAED5" box="[1021,1169,1969,1993]" italics="true" pageId="7" pageNumber="53">pars stridens</emphasis>
|
||
. Subtrapezoidal pronotum narrower in posterior region. Apical edge of elytra with four strongly developed teeth, a pair at the end of the 7th stria and a pair near the suture. Umbilicate series with six setae (4+2) and one pair of scutellar setae. No discal setae. Abdominal sternum II with slight foveae in both sexes and a pair of lateral teeth in the last ventrite (fig. 6). The scaly microsculpture “belt” is diffuse. Sickle-shaped, robust aedeagus (fig. 7b). Female genitalia with short, angled unguiform-like gonocoxites; subspheric spermatheca (fig. 7c).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CAEB6541FF8AFA39FF36EA5891FBAB31" lastPageId="10" lastPageNumber="56" pageId="8" pageNumber="54" type="description">
|
||
<paragraph id="824E36CAFF8AFA3BFF36EA5891DFA85E" blockId="8.[151,1437,151,826]" box="[201,997,298,323]" pageId="8" pageNumber="54">
|
||
<emphasis id="B085EAD8FF8AFA3BFF36EA589361A85F" bold="true" box="[201,347,298,323]" pageId="8" pageNumber="54">Description.</emphasis>
|
||
Length:
|
||
<quantity id="45099B2FFF8AFA3BFE40EA58905BA85E" box="[447,609,298,322]" metricMagnitude="-3" metricUnit="m" metricValue="1.04" metricValueMax="1.11" metricValueMin="0.97" pageId="8" pageNumber="54" unit="mm" value="1.04" valueMax="1.11" valueMin="0.97">0.97–1.11 mm</quantity>
|
||
(males), 1.00–
|
||
<quantity id="45099B2FFF8AFA3BFCF5EA589157A85E" box="[778,877,298,322]" metricMagnitude="-3" metricUnit="m" metricValue="1.07" pageId="8" pageNumber="54" unit="mm" value="1.07">1.07 mm</quantity>
|
||
(females).
|
||
</paragraph>
|
||
<paragraph id="824E36CAFF8AFA3BFF36EA3D9156A8EB" blockId="8.[151,1437,151,826]" pageId="8" pageNumber="54">
|
||
Head wider (
|
||
<quantity id="45099B2FFF8AFA3BFEA7EA3D93C7A87B" box="[344,509,335,359]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" metricValueMax="2.6" metricValueMin="2.2" pageId="8" pageNumber="54" unit="mm" value="0.24" valueMax="0.26" valueMin="0.22">0.22–0.26 mm</quantity>
|
||
) than long (
|
||
<quantity id="45099B2FFF8AFA3BFD7DEA3D9110A87B" box="[642,810,335,359]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="8" pageNumber="54" unit="mm" value="0.22" valueMax="0.24" valueMin="0.2">0.20–0.24 mm</quantity>
|
||
), covered with subhexagonal microreticulation. There is no stridulatory organ in the vertex region. Posterolateral region of the head with semilunar notch. Labrum with rounded edges, slightly depressed in the middle of the anterior margin, with a triangle-shaped region and a subcircular button of thicker cuticle. Clypeus with straight anterior margin. Moniliform antennae, with 11 antennomeres, progressively more square-shaped, except the last one, pyriform.
|
||
</paragraph>
|
||
<paragraph id="824E36CAFF8AFA3BFF36E97091EDAA26" blockId="8.[151,1437,151,826]" pageId="8" pageNumber="54">
|
||
Right mandible with two teeth along the internal margin. Left mandible with a prominent edge in the anterior region. Labium with a low middle tooth. Ligula with prominent, convex middle lobe and long paraglossae. Wide gula, partially fused to the cephalic capsule (suture is only visible in the anterior region). Cephalic chaetotaxy: 5 pairs of labral setae (
|
||
<emphasis id="B085EAD8FF8AFA3BFE75E91D90BDAB9B" bold="true" box="[394,647,622,647]" pageId="8" pageNumber="54">s-l-m-s-m / m-s-m-l-s</emphasis>
|
||
), 2 pairs of clypeal setae (
|
||
<emphasis id="B085EAD8FF8AFA3BFC44E91D9637AB9B" bold="true" box="[955,1037,622,647]" pageId="8" pageNumber="54">l-s / s-l</emphasis>
|
||
), 1 pair of frontal setae, 2 pairs of supraocular setae (anterior and posterior), 1 pair of supra-antennal setae, 3 pairs of occipital setae and 1 pair of genal setae, as well as scattered pubescence. Labium with a pair of setae near the base of the middle tooth, a pair of long setae near the base of the epilobes and a pair of very short setae near the posterior suture (sometimes there is another pair of small setigerous pores). Prebasilar with a lateral pair of setae near the anterior margin, a pair of short setae in the mid region and two pairs of setae in the posterior region.
|
||
</paragraph>
|
||
<caption id="D68E6642FF8AFA3BFF68ECD4932CAEC7" httpUri="https://zenodo.org/record/213513/files/figure.png" pageId="8" pageNumber="54" targetBox="[169,1419,898,1934]" targetPageId="8">
|
||
<paragraph id="824E36CAFF8AFA3BFF68ECD4932CAEC7" blockId="8.[151,1436,1958,2011]" pageId="8" pageNumber="54">
|
||
<emphasis id="B085EAD8FF8AFA3BFF68ECD4932FAEA0" bold="true" box="[151,277,1958,1981]" pageId="8" pageNumber="54">FIGURE 6.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="45F14D49FF8AFA3BFE6AECD59165AEA0" authority="Coiffait, 1969" authorityName="Coiffait" authorityYear="1969" box="[405,863,1958,1981]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="54" phylum="Arthropoda" rank="species" species="quadridentata">
|
||
<emphasis id="B085EAD8FF8AFA3BFE6AECD59088AEA1" box="[405,690,1959,1981]" italics="true" pageId="8" pageNumber="54">Typhlocharis quadridentata</emphasis>
|
||
(Coiffait, 1969)
|
||
</taxonomicName>
|
||
. Ventral (left half: male, right half: female) and dorsal (male) view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="D68E6642FF8BFA3AFF68E8109079AA84" httpUri="https://zenodo.org/record/213514/files/figure.png" pageId="9" pageNumber="55" targetBox="[195,1406,205,846]" targetPageId="9">
|
||
<paragraph id="824E36CAFF8BFA3AFF68E8109079AA84" blockId="9.[151,1436,866,920]" pageId="9" pageNumber="55">
|
||
<emphasis id="B085EAD8FF8BFA3AFF68E810932FAA64" bold="true" box="[151,277,866,889]" pageId="9" pageNumber="55">FIGURE 7.</emphasis>
|
||
<taxonomicName id="45F14D49FF8BFA3AFEE0E8109001AA64" box="[287,571,866,888]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="55" phylum="Arthropoda" rank="species" species="quadridentata">
|
||
<emphasis id="B085EAD8FF8BFA3AFEE0E8109001AA64" box="[287,571,866,888]" italics="true" pageId="9" pageNumber="55">Typhlocharis quadridentata</emphasis>
|
||
</taxonomicName>
|
||
. Aedeagus, (a) median lobe (dorsal view), (b) median lobe and parameres (lateral view). Female genitalia (ventral view) (c).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="824E36CAFF8BFA3AFF36E8CC9741ADD4" blockId="9.[151,1437,957,2032]" pageId="9" pageNumber="55">
|
||
Pronotum slightly longer (
|
||
<quantity id="45099B2FFF8BFA3AFE0FE8CF90B2AAC9" box="[496,648,957,981]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" metricValueMax="3.0" metricValueMin="2.6" pageId="9" pageNumber="55" unit="mm" value="0.28" valueMax="0.3" valueMin="0.26">0.26–0.3 mm</quantity>
|
||
) than wide (
|
||
<quantity id="45099B2FFF8BFA3AFCEBE8CF9186AAC9" box="[788,956,957,981]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" metricValueMax="2.9" metricValueMin="2.5" pageId="9" pageNumber="55" unit="mm" value="0.27" valueMax="0.29" valueMin="0.25">0.25–0.29 mm</quantity>
|
||
), subtrapezoidal, narrowed in the posterior region. Anterior margin almost straight, slightly crenulated, with medial hiatus, approximately as long as two adjacent intersetae spaces. Lateral margins with 5–6 low, irregular denticles, preceding the posterior angles. Surface covered with subhexagonal microreticulation. Flattened disc, with a medial line and two pairs of faint lateral sulci. Chaetotaxy: 1 pair of long setae in the first third of the lateral margins, 1 pair of long setae in the posterior angles, a row of 6 pairs of setae
|
||
<emphasis id="B085EAD8FF8BFA3AFE64EF189045AD9F" bold="true" box="[411,639,1130,1155]" pageId="9" pageNumber="55">(l-l-l-l-l-l / l-l-l-l-l-l)</emphasis>
|
||
parallel to the anterior margin, 4 pairs of setae parallel to the posterior margin (
|
||
<emphasis id="B085EAD8FF8BFA3AFF0AEFFF93F9ADB9" bold="true" box="[245,451,1165,1190]" pageId="9" pageNumber="55">s-m-l-m / m-l-m-s</emphasis>
|
||
), a row of small, thin and regularly placed setae in anterior and posterior margins, lateral margins with a row of short setae. Disc covered by approximately 8 longitudinal rows of pubescence in the disc.
|
||
</paragraph>
|
||
<paragraph id="824E36CAFF8BFA3AFF36EFA0976EAC2C" blockId="9.[151,1437,957,2032]" pageId="9" pageNumber="55">
|
||
Proepisternal suture is almost invisible. Prosternal apophysis rounded. Anterior margin of prosternum with a row of long and thin setae and 5–6 pairs of short setae parallel to it [
|
||
<emphasis id="B085EAD8FF8BFA3AFC6BEF879695AC12" bold="true" box="[916,1199,1269,1294]" pageId="9" pageNumber="55">s-(s)-s-s-s-s / s-s-s-s-(s)-s</emphasis>
|
||
] (fig. 6). Prosternum covered in scattered pubescence, homogeneously distributed, without distinction of glabrous proepisternum.
|
||
</paragraph>
|
||
<paragraph id="824E36CAFF8BFA3AFF36EE489681AFA4" blockId="9.[151,1437,957,2032]" pageId="9" pageNumber="55">
|
||
Elytra approximately 2.1 times longer (
|
||
<quantity id="45099B2FFF8BFA3AFD77EE48910AAC4E" box="[648,816,1338,1362]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" metricValueMax="5.699999999999999" metricValueMin="5.1" pageId="9" pageNumber="55" unit="mm" value="0.54" valueMax="0.57" valueMin="0.51">0.51–0.57 mm</quantity>
|
||
) than wide (
|
||
<quantity id="45099B2FFF8BFA3AFC40EE489653AC4E" box="[959,1129,1338,1362]" metricMagnitude="-4" metricUnit="m" metricValue="2.55" metricValueMax="2.8" metricValueMin="2.3" pageId="9" pageNumber="55" unit="mm" value="0.255" valueMax="0.28" valueMin="0.23">0.23–0.28 mm</quantity>
|
||
), with subparallel margins. Lateral margins with 17–20 denticles, slightly marked and progressively smoother towards posterior. Apical margin with four strongly marked teeth, two lateral, associated to the end of the 7th stria and two submedial, near the suture. Disc flattened, with a longitudinal carinae associated to the 7th stria, typical in the genus. Surface covered with subhexagonal to irregular microreticulation. Scutellar and disc region punctured, the pores are small and extend through the 7th stria, from the shoulder region to approximately a half of the length of the elytra. Transverse scutellar organ and a pair of “buttonholes” are present. Chaetotaxy: umbilicate series composed by an anterior group of 4 setae and a posterior group of 2 setae (fig. 6). There is 1 pair of scutellar setae. There are no discal setae differentiated from the surrounding pubescence, which is relatively long and distributed in 5–6 irregular longitudinal rows. There are 3 pairs of apical setae near the end of the 7th stria and numerous pubescence setae bordering the apical margin. A short seta is associated to every denticle forming a row in the lateral margins.
|
||
</paragraph>
|
||
<paragraph id="824E36CAFF8BFA3AFF36EDB691E0AFE2" blockId="9.[151,1437,957,2032]" pageId="9" pageNumber="55">Legs are similar in both sexes. Very narrow intermetacoxal space. Inner edge of metafemora is slightly angular. Rounded metatrochanters. Metatibia with slightly dilated distal region.</paragraph>
|
||
<paragraph id="824E36CAFF8BFA3AFF36EC7A96D6AE94" blockId="9.[151,1437,957,2032]" pageId="9" pageNumber="55">
|
||
Abdomen with irregular pattern of microreticulation, except the last segment, with faint, scaly and imbricate microsculpture, but not forming the clearly defined “belt” observed in other species. First ventrite (sternum II) with slight posterolateral foveae in both sexes. Last ventrite with a pair of lateral teeth associated with a notch and 6–7 pairs of setae in the posterior margin, with sexual dimorphism:
|
||
<emphasis id="B085EAD8FF8BFA3AFCA3EC0296AAAE95" bold="true" box="[860,1168,1904,1929]" pageId="9" pageNumber="55">l-(s)-s-s-l-s-s / s-l-l-s-s-(s)-l</emphasis>
|
||
(fig. 6).
|
||
</paragraph>
|
||
<paragraph id="824E36CAFF8BFA39FF36ECE196D6A9CE" blockId="9.[151,1437,957,2032]" lastBlockId="10.[151,1436,151,557]" lastPageId="10" lastPageNumber="56" pageId="9" pageNumber="55">
|
||
Median lobe of aedeagus (length:
|
||
<quantity id="45099B2FFF8BFA3AFDAEECE19083AEB7" box="[593,697,1939,1963]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="9" pageNumber="55" unit="mm" value="0.16">0.16 mm</quantity>
|
||
) sickle-shaped and robust. Basal lamina almost straight, slightly curved downwards in the apical region (fig. 7b). Apex subtriangular, with a prominent edge in the right margin, giving an asymmetric “arrowhead” shape. Median lobe of edeagus straight in dorsal view (fig. 7a). Internal sac with two irregular, “stick-shaped” sclerites in lateral view. Left paramere is triangular, with two long apical setae; right paramere is subtriangular, shorter and narrower, also ending in two long apical setae (fig. 7b).
|
||
</paragraph>
|
||
<paragraph id="824E36CAFF88FA39FF36EBAF9397A8BE" blockId="10.[151,1436,151,557]" pageId="10" pageNumber="56">
|
||
Female genitalia do not conform to the models described for the genus (
|
||
<bibRefCitation id="E6604B3BFF88FA39FBE9EBAF9737A9E9" author="Vigna-Taglianti" box="[1046,1293,221,245]" pageId="10" pageNumber="56" refString="Vigna-Taglianti, A. (1972) Un nuovo Typhlocharis di Spagna. (Col., Carab.). Bolletino della Societa Entomologica Italiana. Societa Entomologica Italiana, 104, 148 - 156." type="journal article" year="1972">Vigna-Taglianti 1972</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="E6604B3BFF88FA39FAE3EBAF931BA80B" author="Zaballos" pageId="10" pageNumber="56" refString="Zaballos J. P. & Wrase, D. (1998) Tres nuevos Typhlocharis Dieck, 1869 (Coleoptera, Caraboidea, Trechidae) de Navarra (Espana). Graellsia, 54, 43 - 52." type="journal article" year="1998">Zaballos & Wrase 1998</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="E6604B3BFF88FA39FED1EB8D9035A80B" author="Andujar" box="[302,527,255,280]" pageId="10" pageNumber="56" refString="Andujar, C., Grebennikov, V., Lencina, J. L. & Serrano, J. (2010) A new Typhlocharis (Coleoptera: Carabidae: Anillina) from Spain: combining adult and larval morphological data with DNA information. Zootaxa, 2485, 47 - 57." type="journal article" year="2010">
|
||
Andújar
|
||
<emphasis id="B085EAD8FF88FA39FE6EEA7393F9A804" box="[401,451,256,280]" italics="true" pageId="10" pageNumber="56">et al</emphasis>
|
||
. 2010
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="E6604B3BFF88FA39FDE5EB8D9114A80B" author="Ortuno" box="[538,814,255,279]" pageId="10" pageNumber="56" refString="Ortuno, V. M. & Gilgado, J. D. (2011) Historical perspective, new contributions and an enlightening dispersal mechanism for the endogean genus Typhlocharis Dieck 1869 (Coleoptera: Carabidae: Trechinae). Journal of Natural History, 45, 1233 - 1256." type="journal article" year="2011">Ortuño & Gilgado 2011</bibRefCitation>
|
||
). Gonocoxites are short and robust, strongly bended in the apical region, almost forming a right angle (fig. 7c), with a subapical seta and a short associated seta. Gonoduct is progressively thicker towards the distal region, but not forming two differentiated regions. Subspheric spermatheca (diameter:
|
||
<quantity id="45099B2FFF88FA39FEC1EA15938FA863" box="[318,437,359,383]" metricMagnitude="-5" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="56" unit="mm" value="0.012">0.012 mm</quantity>
|
||
). Spermathecal gland tubular to conical in shape (length:
|
||
<quantity id="45099B2FFF88FA39FBC2EA15968EA863" box="[1085,1204,359,383]" metricMagnitude="-5" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="56" unit="mm" value="0.024">0.024 mm</quantity>
|
||
), slender and sclerotized in the distal region.
|
||
</paragraph>
|
||
<paragraph id="824E36CAFF88FA39FF36EADF91FBAB31" blockId="10.[151,1436,151,557]" pageId="10" pageNumber="56">
|
||
<emphasis id="B085EAD8FF88FA39FF36EADF9316A8DA" bold="true" box="[201,300,429,454]" pageId="10" pageNumber="56">Habitat.</emphasis>
|
||
<bibRefCitation id="E6604B3BFF88FA39FECCEADF93D8A8D9" box="[307,482,429,453]" pageId="10" pageNumber="56" refString="Coiffait, H. (1969) Nouveaux Anillini du Maroc et du Sud de la peninsule iberique. Bulletin de la Societe des Sciences Naturelles et Physiques du Maroc, 48, 55 - 56." type="journal article">Coiffait, (1969)</bibRefCitation>
|
||
does not provide data about the sort of habitat where the species was collected, cited by this author in localities far away from each other (south of the Sierra de Caldeirao and Monchique). Some data in the labels are shared (
|
||
<bibRefCitation id="E6604B3BFF88FA39FE52EA80910FAB16" author="Zaballos" box="[429,821,498,522]" pageId="10" pageNumber="56" refString="Zaballos, J. P. & Perez-Gonzalez, S. (2010 a) Typhlocharis algarvensis Coiffait, 1971: nueva descripcion y afinidades (Coleoptera, Caraboidea, Trechidae). Graellsia, 66, 149 - 156." type="journal article" year="2010" yearSuffix="a">Zaballos & Pérez-González 2010a</bibRefCitation>
|
||
) with some
|
||
<typeStatus id="5D4A8868FF88FA39FC3CEA8191C9AB17" box="[963,1011,499,523]" pageId="10" pageNumber="56">type</typeStatus>
|
||
specimens of
|
||
<taxonomicName id="45F14D49FF88FA39FB67EA8692EBAB31" authority="Coiffait, 1971" authorityName="Coiffait" authorityYear="1971" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="56" phylum="Arthropoda" rank="species" species="algarvensis">
|
||
<emphasis id="B085EAD8FF88FA39FB67EA86970CAB17" box="[1176,1334,499,523]" italics="true" pageId="10" pageNumber="56">T. algarvensis</emphasis>
|
||
Coiffait, 1971
|
||
</taxonomicName>
|
||
, thus inferring a similar habitat for both species (clayey, wet soils).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |