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<document id="42FB979C8047D28A81E1195FE2E301B0" ID-DOI="10.5281/zenodo.13799439" ID-ISSN="0068-547X" ID-Zenodo-Dep="13799439" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538212676" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Rainio, Johanna" docDate="2016" docId="0A3E2657306DFF86FF8EFB54BE4608B9" docLanguage="en" docName="ProcCalifAcadSci.63.7.201-268.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (7)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Chlaenius robertae Kavanaugh and Rainio 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="223" masterDocId="F6075E2F3079FF90FFF5FF92BD630D1B" masterDocTitle="Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar" masterLastPageNumber="268" masterPageNumber="201" pageNumber="221" updateTime="1726771873529" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="71149B4C49FDBACE299571938477CB75" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3D8EC67964C4E809F3CACECA24861E0F">
<mods:title id="77B7887F9BBBC2624CC0435F27E8453C">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
</mods:titleInfo>
<mods:name id="6D372FD8DA5BE091498CFB259363C928" type="personal">
<mods:role id="52BDDFBFFC03382C0656700815B3DE21">
<mods:roleTerm id="327B0D79296EB14DAFD86AA4E1B66D0C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6626785107CDDD7B908EAFD8C0295FA0">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="D1982D88DEBAF9DB5D347EC875BD8785">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
</mods:name>
<mods:name id="EA4D4FAEA34B94FD078B4CE9B7726BD3" type="personal">
<mods:role id="1754073987E828D991837FFF6C8284A1">
<mods:roleTerm id="1C6F384C49C1B9CDB63D401EEA42F097">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="127E57F8F5F013ADC182DE302BC27CB2">Rainio, Johanna</mods:namePart>
<mods:affiliation id="E669017F49B68E3E99B1B579C425EAB3">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
</mods:name>
<mods:typeOfResource id="772BBA4522545A9BD32FE1246FDFBEDD">text</mods:typeOfResource>
<mods:relatedItem id="32AFA98EAA753B719FDD98D9B6628205" type="host">
<mods:titleInfo id="80A0E732142A3212902F89D502EB511F">
<mods:title id="616A6ADBB7E51AAB66D70C1BF86BC7B0">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="66F3938462006A4CB322A5A089C6CEE8">
<mods:date id="93DDB53C71B3416D8185E79E8CE59F4E">2016</mods:date>
<mods:detail id="BD5D197A2D8FCE2734EB545D144F4F2A" type="pubDate">
<mods:number id="C2E864725AD3895586C4A62ED8489C2C">2016-04-29</mods:number>
</mods:detail>
<mods:detail id="4E46603AB6443246BEF0825C5DAD3AF6" type="volume">
<mods:number id="D5D4C4C2664E8CC06FFA5D52560ADB83">63</mods:number>
</mods:detail>
<mods:detail id="4A00B1D7DF9CCD7567481741E9A33D98" type="issue">
<mods:number id="2E4436BBFE6F51300C26A2F9396524B6">7</mods:number>
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<mods:start id="5E5F8314633898778C5B819006AED898">201</mods:start>
<mods:end id="840DDF90CF808A3D61A2506C272E5B02">268</mods:end>
</mods:extent>
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<mods:classification id="FE322FB8DD3611A66EB6DB81724378B3">journal article</mods:classification>
<mods:identifier id="AF096607A437D5804C650AF42EA8E1CD" type="DOI">10.5281/zenodo.13799439</mods:identifier>
<mods:identifier id="D53E66F081A0E2BF4CC8DF081208CF94" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="8E8C0EAD3D716CCDD50D38DFBAC7FA4B" type="Zenodo-Dep">13799439</mods:identifier>
</mods:mods>
<treatment id="0A3E2657306DFF86FF8EFB54BE4608B9" LSID="urn:lsid:plazi:treatment:0A3E2657306DFF86FF8EFB54BE4608B9" httpUri="http://treatment.plazi.org/id/0A3E2657306DFF86FF8EFB54BE4608B9" lastPageId="22" lastPageNumber="223" pageId="20" pageNumber="221">
<subSubSection id="CA8DC4CA306DFF84FF8EFB54BFA009C4" box="[123,707,1222,1247]" pageId="20" pageNumber="221" type="nomenclature">
<paragraph id="82289741306DFF84FF8EFB54BFA009C4" blockId="20.[123,707,1222,1277]" box="[123,707,1222,1247]" pageId="20" pageNumber="221">
<heading id="D960202D306DFF84FF8EFB54BFA009C4" box="[123,707,1222,1247]" fontSize="10" level="2" pageId="20" pageNumber="221" reason="4">
<emphasis id="B0E34B53306DFF84FF8EFB54BFA009C4" bold="true" box="[123,707,1222,1247]" pageId="20" pageNumber="221">
<taxonomicName id="4597ECC2306DFF84FF8EFB54BF3C09C4" ID-CoL="69VV5" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,607,1222,1247]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="robertae" status="sp. nov.">
<emphasis id="B0E34B53306DFF84FF8EFB54BC2C09C5" bold="true" box="[123,335,1222,1246]" italics="true" pageId="20" pageNumber="221">Chlaenius robertae</emphasis>
Kavanaugh and Rainio
</taxonomicName>
,
<taxonomicNameLabel id="ABD0F628306DFF84FD98FB55BFA009C4" box="[621,707,1223,1247]" pageId="20" pageNumber="221" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA306DFF84FF8EFB75BDBB09E7" box="[123,216,1255,1277]" pageId="20" pageNumber="221" type="description">
<paragraph id="82289741306DFF84FF8EFB75BDBB09E7" blockId="20.[123,707,1222,1277]" box="[123,216,1255,1277]" pageId="20" pageNumber="221">
<figureCitation id="1AAC8BC4306DFF84FF8EFB75BDBB09E7" box="[123,216,1255,1277]" captionStart="FIGURE" captionStartId="21.[155,167,1641,1660]" captionTargetBox="[130,1176,216,1618]" captionTargetId="figure-67@21.[120,1176,216,1618]" captionTargetPageId="21" captionText="FIGURE 12. Digital images of holotype male of Chlaenius robertae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799469" httpUri="https://zenodo.org/record/13799469/files/figure.png" pageId="20" pageNumber="221">Figure 12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA306DFF86FF5EFAB3BE4608B9" lastPageId="22" lastPageNumber="223" pageId="20" pageNumber="221" type="materials_examined">
<paragraph id="82289741306DFF84FF5EFAB3BFEF0B67" blockId="20.[123,1174,1313,1726]" pageId="20" pageNumber="221">
<emphasis id="B0E34B53306DFF84FF5EFAB3BC39082C" bold="true" box="[171,346,1313,1337]" pageId="20" pageNumber="221">
<smallCapsWord id="84CE019D306DFF84FF5EFAB3BD870823" baselines="1331,1331" box="[171,228,1313,1337]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="20" pageNumber="221">TYPE</smallCapsWord>
<smallCapsWord id="84CE019D306DFF84FF18FAB6BC39082C" baselines="1330" box="[237,346,1316,1335]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="20" pageNumber="221">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C306DFF84FE77FAB3BC2808E0" collectingDate="2001-01-02" collectingDateMax="2001-01-22" collectingDateMin="2001-01-02" collectorName="D. H. &amp; K. M. Kavanaugh &amp; R. L. Brett &amp; E. Elsom &amp; F. Vargas &amp; E. F. Randrianifirina &amp; N. Rasoamananana &amp; T. J. Ravelomanana &amp; H. C. Raveloson" country="Madagascar" elevation="1150" latitude="-21.24032" location="Vohiparara area" longLatPrecision="1" longitude="47.39399" municipality="Ranomafana National Park" pageId="20" pageNumber="221" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="5D2C29E3306DFF84FE77FAB3BC860822" box="[386,485,1313,1337]" pageId="20" pageNumber="221" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC4306DFF84FE02FAB3BF1A0823" box="[503,633,1313,1337]" captionStart="FIGURE" captionStartId="21.[155,167,1641,1660]" captionTargetBox="[130,1176,216,1618]" captionTargetId="figure-67@21.[120,1176,216,1618]" captionTargetPageId="21" captionText="FIGURE 12. Digital images of holotype male of Chlaenius robertae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799469" httpUri="https://zenodo.org/record/13799469/files/figure.png" pageId="20" pageNumber="221">Figs.12A–B</figureCitation>
), a male, in CAS, labeled: “CASENT 1002781”/ “
<collectingCountry id="FA80D7D1306DFF84FF73FAD3BC270842" box="[134,324,1345,1369]" name="Madagascar" pageId="20" pageNumber="221">MADAGASCAR</collectingCountry>
,
<collectingRegion id="405359A3306DFF84FEA6FAD3BF200842" box="[339,579,1345,1369]" country="Madagascar" name="Fianarantsoa" pageId="20" pageNumber="221">Fianarantsoa Province</collectingRegion>
,
<collectingMunicipality id="624C0D3B306DFF84FDA7FAD3BE150842" box="[594,886,1345,1369]" pageId="20" pageNumber="221">Ranomafana National Park</collectingMunicipality>
,
<location id="8748C19A306DFF84FC70FAD3B94D0842" LSID="urn:lsid:plazi:treatment:0A3E2657306DFF86FF8EFB54BE4608B9:8748C19A306DFF84FC70FAD3B94D0842" box="[901,1070,1345,1369]" country="Madagascar" latitude="-21.24032" longLatPrecision="1" longitude="47.39399" municipality="Ranomafana National Park" name="Vohiparara area" pageId="20" pageNumber="221" stateProvince="Fianarantsoa">Vohiparara area</location>
,
<quantity id="456F3AA4306DFF84FBC8FAD3B9EC0842" box="[1085,1167,1345,1369]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="20" pageNumber="221" unit="m" value="1150.0">
<elevation id="09BA7072306DFF84FBC8FAD3B9EC0842" box="[1085,1167,1345,1369]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="20" pageNumber="221" unit="m" value="1150.0">1150 m</elevation>
</quantity>
, mixed tropical forest,
<date id="F629B181306DFF84FE9EFAF0BF570861" box="[363,564,1378,1402]" pageId="20" pageNumber="221" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">
<collectingDate id="E66D4869306DFF84FE9EFAF0BF570861" box="[363,564,1378,1402]" pageId="20" pageNumber="221" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">2-22 January 2001</collectingDate>
</date>
,”/ “
<geoCoordinate id="E7A3F186306DFF84FD96FAF0BF830861" box="[611,736,1378,1402]" degrees="21. 24032" direction="south" orientation="latitude" pageId="20" pageNumber="221" precision="1" value="-21.24032">21.24032°S</geoCoordinate>
/
<geoCoordinate id="E7A3F186306DFF84FD05FAF0BE0C0862" box="[752,879,1378,1402]" degrees="47. 39399" direction="east" orientation="longitude" pageId="20" pageNumber="221" precision="1" value="47.39399">47.39399°E</geoCoordinate>
, Stop# DHK-01-002,
<collectorName id="2F62F297306DFF84FB94FAF0BC3D0881" pageId="20" pageNumber="221">D.H. &amp; K.M. Kavanaugh</collectorName>
,
<collectorName id="2F62F297306DFF84FE86FA10BC850882" box="[371,486,1410,1433]" pageId="20" pageNumber="221">R.L. Brett</collectorName>
,
<collectorName id="2F62F297306DFF84FE09FA10BF060881" box="[508,613,1410,1434]" pageId="20" pageNumber="221">E. Elsom</collectorName>
,
<collectorName id="2F62F297306DFF84FD8EFA10BF870881" box="[635,740,1410,1434]" pageId="20" pageNumber="221">F. Vargas</collectorName>
, Ranaivosolo,”/ “
<collectorName id="2F62F297306DFF84FC46FA10B9EC0881" box="[947,1167,1410,1434]" pageId="20" pageNumber="221">E.F. Randrianifirina</collectorName>
,
<collectorName id="2F62F297306DFF84FF8EFA31BC2A08A1" box="[123,329,1443,1466]" pageId="20" pageNumber="221">N. Rasoamananana</collectorName>
,
<collectorName id="2F62F297306DFF84FEA3FA31BF4208A1" box="[342,545,1442,1466]" pageId="20" pageNumber="221">T.J. Ravelomanana</collectorName>
, and
<collectorName id="2F62F297306DFF84FDA9FA31BE6408A1" box="[604,775,1442,1466]" pageId="20" pageNumber="221">H.C. Raveloson</collectorName>
collectors”/ “collected in daytime by treading marsh vegetation and shore”/ “
<typeStatus id="5D2C29E3306DFF84FDD1FA51BFD208C0" box="[548,689,1475,1499]" pageId="20" pageNumber="221" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC2306DFF84FD4DFA51B91308C0" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[696,1136,1475,1499]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="robertae" status="sp. nov.">
<emphasis id="B0E34B53306DFF84FD4DFA51BEE508C0" box="[696,902,1475,1499]" italics="true" pageId="20" pageNumber="221">Chlaenius robertae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628306DFF84FB82FA51BDEC08E0" pageId="20" pageNumber="221" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label]
</materialsCitation>
.
<materialsCitation id="32FF9D1C306DFF84FEADFA76BFE60B67" collectingDate="1998-04-27" collectionCode="CAS" collectorName="D. H. Kavanaugh" country="Madagascar" elevation="1050" latitude="-21.226667" location="Vohiparara area" longLatPrecision="126" longitude="47.383335" municipality="Ranomafana National Park" pageId="20" pageNumber="221" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="paratype">
<typeStatus id="5D2C29E3306DFF84FEADFA76BCA308E7" box="[344,448,1508,1532]" pageId="20" pageNumber="221" type="paratype">Paratypes</typeStatus>
(only 1): a female (in
<collectionCode id="E4860F84306DFF84FD5BFA71BF8108E0" box="[686,738,1507,1531]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="20" pageNumber="221" type="Museum">CAS</collectionCode>
) labeled “CASENT1049009”/ “
<collectingCountry id="FA80D7D1306DFF84FBCEFA71BD840B07" name="Madagascar" pageId="20" pageNumber="221">MADAGASCAR</collectingCountry>
,
<collectingRegion id="405359A3306DFF84FF0DF991BC8A0B00" box="[248,489,1539,1563]" country="Madagascar" name="Fianarantsoa" pageId="20" pageNumber="221">Fianarantsoa Province</collectingRegion>
,
<collectingMunicipality id="624C0D3B306DFF84FE0FF991BE410B00" box="[506,802,1539,1563]" pageId="20" pageNumber="221">Ranomafana National Park</collectingMunicipality>
,
<location id="8748C19A306DFF84FCC7F991BEBF0B00" LSID="urn:lsid:plazi:treatment:0A3E2657306DFF86FF8EFB54BE4608B9:8748C19A306DFF84FCC7F991BEBF0B00" box="[818,988,1539,1563]" country="Madagascar" latitude="-21.226667" longLatPrecision="126" longitude="47.383335" municipality="Ranomafana National Park" name="Vohiparara area" pageId="20" pageNumber="221" stateProvince="Fianarantsoa">Vohiparara area</location>
,
<quantity id="456F3AA4306DFF84FC1BF991B9210B00" box="[1006,1090,1539,1563]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="20" pageNumber="221" unit="m" value="1050.0">
<elevation id="09BA7072306DFF84FC1BF991B9210B00" box="[1006,1090,1539,1563]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="20" pageNumber="221" unit="m" value="1050.0">1050 m</elevation>
</quantity>
, mixed tropical forest,
<date id="F629B181306DFF84FEEAF9B6BCD70B27" box="[287,436,1572,1596]" pageId="20" pageNumber="221" value="1998-04-27">
<collectingDate id="E66D4869306DFF84FEEAF9B6BCD70B27" box="[287,436,1572,1596]" pageId="20" pageNumber="221" value="1998-04-27">27 April 1998</collectingDate>
</date>
,”/ “
<geoCoordinate id="E7A3F186306DFF84FE17F9B6BF2D0B27" box="[482,590,1572,1596]" degrees="21" direction="south" minutes="13. 6" orientation="latitude" pageId="20" pageNumber="221" precision="92" value="-21.226667">21°13.6’S</geoCoordinate>
/
<geoCoordinate id="E7A3F186306DFF84FDA9F9B6BFAA0B20" box="[604,713,1572,1596]" degrees="47" direction="east" minutes="23. 0" orientation="longitude" pageId="20" pageNumber="221" precision="92" value="47.383335">47°23.0’E</geoCoordinate>
,”/ “Stop # 98-91,
<collectorName id="2F62F297306DFF84FC65F9B6B9260B27" box="[912,1093,1572,1596]" pageId="20" pageNumber="221">D.H. Kavanaugh</collectorName>
collector collected on bare soil at night in abandoned rice paddy area”/ “
<typeStatus id="5D2C29E3306DFF84FCC8F9D6BEA20B47" box="[829,961,1604,1628]" pageId="20" pageNumber="221" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4597ECC2306DFF84FC3DF9D6BC3C0B66" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="robertae" status="sp. nov.">
<emphasis id="B0E34B53306DFF84FC3DF9D6B9F60B47" box="[968,1173,1604,1628]" italics="true" pageId="20" pageNumber="221">Chlaenius robertae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628306DFF84FE93F9F7BCC30B67" box="[358,416,1637,1660]" pageId="20" pageNumber="221" rank="species">sp. n.</taxonomicNameLabel>
2015” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="82289741306DFF84FF5EF917B9500B86" blockId="20.[123,1174,1313,1726]" box="[171,1075,1669,1693]" pageId="20" pageNumber="221">
<materialsCitation id="32FF9D1C306DFF84FF5EF917B94F0B86" box="[171,1068,1669,1693]" country="Madagascar" location="Ranomafana National Park" pageId="20" pageNumber="221" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B53306DFF84FF5EF917BC340B80" bold="true" box="[171,343,1669,1693]" pageId="20" pageNumber="221">
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<typeStatus id="5D2C29E3306DFF84FF5EF917BD870B87" box="[171,228,1669,1693]" pageId="20" pageNumber="221">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D306DFF84FF19F91ABC340B80" baselines="1686" box="[236,343,1672,1691]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="20" pageNumber="221">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D1306DFF84FE8BF917BC9C0B86" box="[382,511,1669,1693]" name="Madagascar" pageId="20" pageNumber="221">Madagascar</collectingCountry>
,
<collectingRegion id="405359A3306DFF84FDF8F917BF9F0B86" box="[525,764,1669,1693]" country="Madagascar" name="Fianarantsoa" pageId="20" pageNumber="221">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A306DFF84FCFFF917B94F0B86" LSID="urn:lsid:plazi:treatment:0A3E2657306DFF86FF8EFB54BE4608B9:8748C19A306DFF84FCFFF917B94F0B86" box="[778,1068,1669,1693]" country="Madagascar" name="Ranomafana National Park" pageId="20" pageNumber="221" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="82289741306DFF84FF5EF934B9F50BA6" blockId="20.[123,1174,1313,1726]" box="[171,1174,1701,1726]" pageId="20" pageNumber="221">
<emphasis id="B0E34B53306DFF84FF5EF934BC8D0BA7" bold="true" box="[171,494,1702,1726]" pageId="20" pageNumber="221">
<smallCapsWord id="84CE019D306DFF84FF5EF934BC4D0BA7" baselines="1720,1719" box="[171,302,1702,1726]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="20" pageNumber="221">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D306DFF84FEC1F93BBC330BA7" baselines="1719" box="[308,336,1705,1724]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="20" pageNumber="221">OF</smallCapsWord>
<smallCapsWord id="84CE019D306DFF84FEA3F93BBCCA0BA7" baselines="1719" box="[342,425,1705,1724]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="20" pageNumber="221">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D306DFF84FE44F93BBC8D0BA7" baselines="1719" box="[433,494,1705,1724]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="20" pageNumber="221">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC2306DFF84FD05F937BE280BA6" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[752,843,1701,1725]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="robertae">
<emphasis id="B0E34B53306DFF84FD05F937BE280BA6" box="[752,843,1701,1725]" italics="true" pageId="20" pageNumber="221">robertae</emphasis>
</taxonomicName>
, is a noun in the genitive case,
</paragraph>
<caption id="D6E8C7C9306CFF85FF6EF9FBB9450B8E" ID-DOI="http://doi.org/10.5281/zenodo.13799469" ID-Zenodo-Dep="13799469" httpUri="https://zenodo.org/record/13799469/files/figure.png" pageId="21" pageNumber="222" startId="21.[155,167,1641,1660]" targetBox="[130,1176,216,1618]" targetPageId="21" targetType="figure">
<paragraph id="82289741306CFF85FF6EF9FBB9450B8E" blockId="21.[123,1173,1640,1685]" pageId="21" pageNumber="222">
<smallCapsWord id="84CE019D306CFF85FF6EF9FBBDBB0B61" baselines="1655,1654" box="[155,216,1641,1660]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="21" pageNumber="222">FIGURE</smallCapsWord>
12. Digital images of holotype male of
<taxonomicName id="4597ECC2306CFF85FDC8F9FABEC60B60" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[573,933,1640,1659]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="222" phylum="Arthropoda" rank="species" species="robertae" status="sp. nov.">
<emphasis id="B0E34B53306CFF85FDC8F9FABF870B60" box="[573,740,1640,1659]" italics="true" pageId="21" pageNumber="222">Chlaenius robertae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628306CFF85FC59F9FBBE8C0B67" box="[940,1007,1641,1660]" pageId="21" pageNumber="222" rank="species">sp. nov.</taxonomicNameLabel>
A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 1.0 mm.
</paragraph>
</caption>
<paragraph id="82289741306FFF86FF8EFF49BFCB0C08" blockId="22.[123,1174,219,1443]" pageId="22" pageNumber="223">
derived from the given name of Roberta L. Brett, former graduate student and technical assistant of DHK and one of the collectors of the
<typeStatus id="5D2C29E3306FFF86FDDAFF6EBF3E0C0F" box="[559,605,252,276]" pageId="22" pageNumber="223">type</typeStatus>
series.
</paragraph>
<paragraph id="82289741306FFF86FF5EFE8EBE0F081B" blockId="22.[123,1174,219,1443]" pageId="22" pageNumber="223">
<smallCapsWord id="84CE019D306FFF86FF5EFE8EBC210C29" baselines="302,301" box="[171,322,284,308]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="22" pageNumber="223">
<emphasis id="B0E34B53306FFF86FF5EFE8EBC210C29" bold="true" box="[171,322,284,308]" pageId="22" pageNumber="223">RECOGNITION</emphasis>
</smallCapsWord>
.— Sized moderate for genus, SBL of male =
<quantity id="456F3AA4306FFF86FCCCFE8EBEFA0C28" box="[825,921,284,308]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="22" pageNumber="223" unit="mm" value="12.9">12.9 mm</quantity>
, of female =
<quantity id="456F3AA4306FFF86FBDBFE8EB9ED0C28" box="[1070,1166,284,308]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="22" pageNumber="223" unit="mm" value="13.3">13.3 mm</quantity>
. Members of this species (
<figureCitation id="1AAC8BC4306FFF86FE69FEAEBC9D0C4F" box="[412,510,316,340]" captionStart="FIGURE" captionStartId="21.[155,167,1641,1660]" captionTargetBox="[130,1176,216,1618]" captionTargetId="figure-67@21.[120,1176,216,1618]" captionTargetPageId="21" captionText="FIGURE 12. Digital images of holotype male of Chlaenius robertae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799469" httpUri="https://zenodo.org/record/13799469/files/figure.png" pageId="22" pageNumber="223">Fig. 12A</figureCitation>
) exhibit the following features, which place them in genus
<taxonomicName id="4597ECC2306FFF86FF8EFECFBC860C6E" authority="sensu Jeannel (1949)" authorityName="sensu Jeannel" authorityYear="1949" box="[123,485,349,373]" class="Insecta" family="Carabidae" genus="Chlaenites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B53306FFF86FF8EFECFBD8E0C6E" box="[123,237,349,373]" italics="true" pageId="22" pageNumber="223">Chlaenites</emphasis>
sensu Jeannel (1949)
</taxonomicName>
: antennomere 3 longer than antennomere 4; maxillary palpi setose; penultimate labial palpomeres 4 or 5 setose; pronotum with basolateral setae inserted distinctly anterior to the hind angles; elytra with pubescence restricted to near the striae on disk (i.e., not present on the centers of intervals), except more generally and densely present on lateral intervals and near apices; and tarsi asetose dorsally, tarsomeres 5 with two rows of stout setae ventrally. In Jeannel’s (1949) key to subgenera of
<taxonomicName id="4597ECC2306FFF86FDB1FE6DBFD50F0C" authorityName="sensu Jeannel" authorityYear="1949" box="[580,694,511,535]" class="Insecta" family="Carabidae" genus="Chlaenites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B53306FFF86FDB1FE6DBFD50F0C" box="[580,694,511,535]" italics="true" pageId="22" pageNumber="223">Chlaenites</emphasis>
</taxonomicName>
, members of this species key to his new subgenus
<taxonomicName id="4597ECC2306FFF86FF34FD8DBC320F2C" box="[193,337,543,567]" class="Insecta" family="Carabidae" genus="Chlaenitidius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B53306FFF86FF34FD8DBC320F2C" box="[193,337,543,567]" italics="true" pageId="22" pageNumber="223">Chlaenitidius</emphasis>
</taxonomicName>
, which is now considered a junior synonym of subgenus
<taxonomicName id="4597ECC2306FFF86FC34FD8DBC2D0F43" authority="Laferte-Senectere (1851)" authorityName="Laferte-Senectere" authorityYear="1851" class="Insecta" family="Carabidae" genus="Amblygenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B53306FFF86FC34FD8DB9280F2C" box="[961,1099,543,567]" italics="true" pageId="22" pageNumber="223">Amblygenius</emphasis>
<bibRefCitation id="E606EAB0306FFF86FBA6FD8DBC2D0F43" author="LAFERTE-SENECTERE, F. T." pageId="22" pageNumber="223" pagination="209 - 294" refId="ref31239" refString="LAFERTE-SENECTERE, F. T. 1851. Revision de la tribu des Pelmatellimanes de Dejean, coleopteres pentameres de la famille des Carabiques. Annales de la Societe Entomologique de France (2) 9: 209 - 294." type="journal article" year="1851">Laferté-Sénectère (1851)</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="E606EAB0306FFF86FEAAFDD2BC8B0F4C" author="LORENZ, W." box="[351,488,575,599]" pageId="22" pageNumber="223" refId="ref31666" refString="LORENZ, W. 2005. Systematic List of Extant Ground Beetles of the World (Insecta Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second edition. W. Lorenz, Tutzing. 530 pp." type="book" year="2005">Lorenz 2005</bibRefCitation>
), in having the pronotum relatively broad and the setose punctures along the lateral margins of the elytral intervals slightly foveate, especially in the apical half apically. Member s of
<taxonomicName id="4597ECC2306FFF86FE9EFD13BC850F83" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[363,486,640,664]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="robertae">
<emphasis id="B0E34B53306FFF86FE9EFD13BC850F83" box="[363,486,640,664]" italics="true" pageId="22" pageNumber="223">C. robertae</emphasis>
</taxonomicName>
differ distinctly from those of the each of the four species Jeannel included in this subgenus. They differ from members of
<taxonomicName id="4597ECC2306FFF86FCF0FD33B9F60FA2" authority="Faimaire (1901)" authorityName="Faimaire" authorityYear="1901" box="[773,1173,673,697]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="cupreolus">
<emphasis id="B0E34B53306FFF86FCF0FD33BE830FA2" box="[773,992,673,697]" italics="true" pageId="22" pageNumber="223">Chlaenius cupreolus</emphasis>
Faimaire (1901)
</taxonomicName>
in having striae deeply impressed and intervals moderately convex (striae superficial only and intervals flat in
<taxonomicName id="4597ECC2306FFF86FEDFFD70BCD70FE1" authorityName="Faimaire" authorityYear="1901" box="[298,436,738,762]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="cupreolus">
<emphasis id="B0E34B53306FFF86FEDFFD70BCD70FE1" box="[298,436,738,762]" italics="true" pageId="22" pageNumber="223">C. cupreolus</emphasis>
</taxonomicName>
members), head with metallic green reflection but pronotum black without a trace of metallic reflection (both head and pronotum with green metallic reflection in
<taxonomicName id="4597ECC2306FFF86FF8EFCB1BC620E20" authorityName="Faimaire" authorityYear="1901" box="[123,257,803,827]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="cupreolus">
<emphasis id="B0E34B53306FFF86FF8EFCB1BC620E20" box="[123,257,803,827]" italics="true" pageId="22" pageNumber="223">C. cupreolus</emphasis>
</taxonomicName>
members), and elytra without a pale lateral and apical margin (a thin pale margin present in
<taxonomicName id="4597ECC2306FFF86FF35FCD1BC2B0E40" authorityName="Faimaire" authorityYear="1901" box="[192,328,835,859]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="cupreolus">
<emphasis id="B0E34B53306FFF86FF35FCD1BC2B0E40" box="[192,328,835,859]" italics="true" pageId="22" pageNumber="223">C. cupreolus</emphasis>
</taxonomicName>
members). They differ from members of
<taxonomicName id="4597ECC2306FFF86FCFAFCD1B9EC0E40" authority="Alluaud (1919)" authorityName="Alluaud" authorityYear="1919" box="[783,1167,835,859]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="allacteus">
<emphasis id="B0E34B53306FFF86FCFAFCD1BE800E40" box="[783,995,835,859]" italics="true" pageId="22" pageNumber="223">Chlaenius allacteus</emphasis>
<bibRefCitation id="E606EAB0306FFF86FC1FFCD1B9EC0E40" author="ALLUAUD, C." box="[1002,1167,835,859]" pageId="22" pageNumber="223" pagination="50 - 53" refId="ref28857" refString="ALLUAUD, C. 1919. Contributions a l'etude des Carabiques d'Afrique et de Madagascar [Col.] XXII. Descriptions de quatre especes et d'un genre nouveaux de Chleniens. Bulletin de la Societe Entomologique de France 1919: 50 - 53." type="journal article" year="1919">Alluaud (1919)</bibRefCitation>
</taxonomicName>
, which is current classified in subgenus
<taxonomicName id="4597ECC2306FFF86FDDDFCF1BE380E67" authority="Alluaud (1933)" authorityName="Alluaud" authorityYear="1933" box="[552,859,867,892]" class="Insecta" family="Carabidae" genus="Oochlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B53306FFF86FDDDFCF1BFCC0E60" box="[552,687,867,891]" italics="true" pageId="22" pageNumber="223">Oochlaenius</emphasis>
<bibRefCitation id="E606EAB0306FFF86FD40FCF1BE380E67" author="ALLUAUD, C." box="[693,859,867,892]" pageId="22" pageNumber="223" pagination="10 - 20" refId="ref28924" refString="ALLUAUD, C. 1933 II. Chlaeniinae d'Afrique tropicale. Afra, Cahiers d'Entomologie 6: 10 - 20." type="journal article" year="1933">Alluaud (1933)</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="E606EAB0306FFF86FC99FCF6BE960E60" author="LORENZ, W." box="[876,1013,867,891]" pageId="22" pageNumber="223" refId="ref31666" refString="LORENZ, W. 2005. Systematic List of Extant Ground Beetles of the World (Insecta Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second edition. W. Lorenz, Tutzing. 530 pp." type="book" year="2005">Lorenz 2005</bibRefCitation>
), in having all elytral interval similarly and moderately convex (intervals 1, 3, 5 and 7 more convex and intervals 2, 4, 6 and 8 flat in
<taxonomicName id="4597ECC2306FFF86FEBBFC37BCAE0EA7" authorityName="Alluaud" authorityYear="1919" box="[334,461,932,956]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="allacteus">
<emphasis id="B0E34B53306FFF86FEBBFC37BCAE0EA7" box="[334,461,932,956]" italics="true" pageId="22" pageNumber="223">C. allacteus</emphasis>
</taxonomicName>
members) and, again, elytra without a pale lateral and apical margin (a wide pale margin present in
<taxonomicName id="4597ECC2306FFF86FE18FC57BF0F0EC6" authorityName="Alluaud" authorityYear="1919" box="[493,620,965,989]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="allacteus">
<emphasis id="B0E34B53306FFF86FE18FC57BF0F0EC6" box="[493,620,965,989]" italics="true" pageId="22" pageNumber="223">C. allacteus</emphasis>
</taxonomicName>
members). They differ from members of
<taxonomicName id="4597ECC2306FFF86FBDCFC57BCC50EE6" authority="Faimaire (1901)" authorityName="Faimaire" authorityYear="1901" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="B0E34B53306FFF86FBDCFC57BD890EE6" italics="true" pageId="22" pageNumber="223">Chlaenius inaequalis</emphasis>
Faimaire (1901)
</taxonomicName>
in having the pronotum black without metallic reflection (metallic green reflection present on pronotum in
<taxonomicName id="4597ECC2306FFF86FDC9FB94BFAE0905" authorityName="Faimaire" authorityYear="1901" box="[572,717,1030,1054]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="B0E34B53306FFF86FDC9FB94BFAE0905" box="[572,717,1030,1054]" italics="true" pageId="22" pageNumber="223">C. inaequalis</emphasis>
</taxonomicName>
members), a longer and narrower elytral form (shorter and broader elytra in
<taxonomicName id="4597ECC2306FFF86FE0AFBB4BFF30925" authorityName="Faimaire" authorityYear="1901" box="[511,656,1062,1086]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="B0E34B53306FFF86FE0AFBB4BFF30925" box="[511,656,1062,1086]" italics="true" pageId="22" pageNumber="223">C. inaequalis</emphasis>
</taxonomicName>
members) and elytral epipleurae black (pale in
<taxonomicName id="4597ECC2306FFF86FF8EFBD5BC6E0945" authorityName="Faimaire" authorityYear="1901" box="[123,269,1094,1118]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="B0E34B53306FFF86FF8EFBD5BC6E0945" box="[123,269,1094,1118]" italics="true" pageId="22" pageNumber="223">C. inaequalis</emphasis>
</taxonomicName>
members). Overall, members of
<taxonomicName id="4597ECC2306FFF86FD74FBD5BF9C0945" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[641,767,1094,1118]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="robertae">
<emphasis id="B0E34B53306FFF86FD74FBD5BF9C0945" box="[641,767,1094,1118]" italics="true" pageId="22" pageNumber="223">C. robertae</emphasis>
</taxonomicName>
are most similar to those the fourth species,
<taxonomicName id="4597ECC2306FFF86FF22FBF5BF270964" authority="Jeannel (1949)" authorityName="Jeannel" authorityYear="1949" box="[215,580,1127,1151]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="lyperus">
<emphasis id="B0E34B53306FFF86FF22FBF5BCF90964" box="[215,410,1127,1151]" italics="true" pageId="22" pageNumber="223">Chlaenius lyperus</emphasis>
Jeannel (1949)
</taxonomicName>
, with which they share similar elytral shape, depth of striae and convexity of intervals. However, they differ from
<taxonomicName id="4597ECC2306FFF86FD0DFB1ABE070984" authorityName="Jeannel" authorityYear="1949" box="[760,868,1159,1183]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="lyperus">
<emphasis id="B0E34B53306FFF86FD0DFB1ABE070984" box="[760,868,1159,1183]" italics="true" pageId="22" pageNumber="223">C. lyperus</emphasis>
</taxonomicName>
members in having a pronotum broader basally than the latter and legs that are pale in color (dark in
<taxonomicName id="4597ECC2306FFF86FC5EFB3AB97809DB" authorityName="Jeannel" authorityYear="1949" box="[939,1051,1192,1216]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="lyperus">
<emphasis id="B0E34B53306FFF86FC5EFB3AB97809DB" box="[939,1051,1192,1216]" italics="true" pageId="22" pageNumber="223">C. lyperus</emphasis>
</taxonomicName>
members). Males of
<taxonomicName id="4597ECC2306FFF86FF17FB5BBC3D09FB" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[226,350,1224,1248]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="robertae">
<emphasis id="B0E34B53306FFF86FF17FB5BBC3D09FB" box="[226,350,1224,1248]" italics="true" pageId="22" pageNumber="223">C. robertae</emphasis>
</taxonomicName>
also have genitalia (
<figureCitation id="1AAC8BC4306FFF86FDB5FB5ABFA909FB" box="[576,714,1224,1248]" captionStart="FIGURE" captionStartId="21.[155,167,1641,1660]" captionTargetBox="[130,1176,216,1618]" captionTargetId="figure-67@21.[120,1176,216,1618]" captionTargetPageId="21" captionText="FIGURE 12. Digital images of holotype male of Chlaenius robertae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799469" httpUri="https://zenodo.org/record/13799469/files/figure.png" pageId="22" pageNumber="223">Figs. 12C–D</figureCitation>
) that are markedly different in form those of the other four species (compare with Jeannel’s (1949) Fig. 386a-d).
</paragraph>
<paragraph id="82289741306FFF86FF5EFA9BB969083B" blockId="22.[123,1174,219,1443]" box="[171,1034,1289,1313]" pageId="22" pageNumber="223">
<emphasis id="B0E34B53306FFF86FF5EFA9BBC920804" bold="true" box="[171,497,1289,1313]" pageId="22" pageNumber="223">
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.— At present, known only from the
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locality.
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.— Both specimens of the
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series were collected in open habitats at the edges of mixed tropical forest in the Vohiparara area at elevations ranging from
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. One was collected in daytime by treading down vegetation at the edge of a marsh, the other was found at night running on bare soil in an abandoned rice paddy.
</paragraph>
</subSubSection>
</treatment>
</document>
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