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<document id="9076AA8606C750CEAF2892DA03B2DFAA" ID-DOI="10.5281/zenodo.13799439" ID-ISSN="0068-547X" ID-Zenodo-Dep="13799439" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538212676" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Rainio, Johanna" docDate="2016" docId="0A3E2657306BFF84FF8EFBE3BF8A0985" docLanguage="en" docName="ProcCalifAcadSci.63.7.201-268.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (7)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Chlaenius kathrynae Kavanaugh and Rainio 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="221" masterDocId="F6075E2F3079FF90FFF5FF92BD630D1B" masterDocTitle="Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar" masterLastPageNumber="268" masterPageNumber="201" pageNumber="219" updateTime="1726771873529" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="12D80E5BC6E2EE884FA2B6182FDB738E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F87242116344C1F2FDBEE32CC01164C6">
<mods:title id="67CDFEAC3F66A4C3AE4C02DFFEE3FF39">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
</mods:titleInfo>
<mods:name id="5C94FF4996A92C33301871EF9FFF4938" type="personal">
<mods:role id="44B77DB3400F84B33BEEC4D6C11930A6">
<mods:roleTerm id="17CA1DB7F50B69D5F3E9980BCBB3BF42">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="056E053ACE383D59B155C8E4FE307409">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="E6C8DD5E6A19084AC3ED157AF7F4EBD2">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
</mods:name>
<mods:name id="5652FF2EDB34C19EB045CC33A18257AC" type="personal">
<mods:role id="B0931C9B0C2A8DD48EA0E5CA2EE1DB9C">
<mods:roleTerm id="47874CBEC1A1CB72668D9F05C7B6D4B5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C1BA80B8C318A868C6F6697495CC31BD">Rainio, Johanna</mods:namePart>
<mods:affiliation id="F56E0C99FD0CACB56598D4DB980D7DD9">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
</mods:name>
<mods:typeOfResource id="8F481E1F9FF11ED843FCE5279D23F8D7">text</mods:typeOfResource>
<mods:relatedItem id="7B52B15158F066E2B368231536321D2C" type="host">
<mods:titleInfo id="5667D12CC2799D3E947197EA47773FB9">
<mods:title id="2541B8E91401427B5A0ACEE019AD89DF">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="2DBDB8A9E989B5AEAB55E92774906A87">
<mods:date id="16239678B4D05F3DFA7FCB9D57DBBF12">2016</mods:date>
<mods:detail id="5D7182F00B36AE290A6C6F9E5D95616A" type="pubDate">
<mods:number id="D6FD9F018CC3F61928146A99AD8A7C9E">2016-04-29</mods:number>
</mods:detail>
<mods:detail id="12F17EA491D5F4D7CFB0D234AF4D05BF" type="volume">
<mods:number id="846F0A64266045EA9136BD3B551AFAEE">63</mods:number>
</mods:detail>
<mods:detail id="C565A0211ACF53834ABBC3ABEF6E96D0" type="issue">
<mods:number id="8F5D59731BE863F53AB10A85F63C7AD4">7</mods:number>
</mods:detail>
<mods:extent id="DFD4EC7ED4473B5BFF259A401A04EF08" unit="page">
<mods:start id="45D7B234134E6930526AF9083C42BA63">201</mods:start>
<mods:end id="8606FB43D69EC1F738D64425A3ECC2B8">268</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="992C033DDC7DB5415E884E2FE00297D0">journal article</mods:classification>
<mods:identifier id="89107FB216BC289D974B99E94F60494F" type="DOI">10.5281/zenodo.13799439</mods:identifier>
<mods:identifier id="D634FF3AEEF2404ECE1863F8324F6B59" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="E09A0AC4CE3806D234CE4FBC7F98FA2B" type="Zenodo-Dep">13799439</mods:identifier>
</mods:mods>
<treatment id="0A3E2657306BFF84FF8EFBE3BF8A0985" ID-DOI="http://doi.org/10.5281/zenodo.13799467" ID-Zenodo-Dep="13799467" LSID="urn:lsid:plazi:treatment:0A3E2657306BFF84FF8EFBE3BF8A0985" httpUri="http://treatment.plazi.org/id/0A3E2657306BFF84FF8EFBE3BF8A0985" lastPageId="20" lastPageNumber="221" pageId="18" pageNumber="219">
<subSubSection id="CA8DC4CA306BFF82FF8EFBE3BFB50991" box="[123,726,1137,1162]" pageId="18" pageNumber="219" type="nomenclature">
<paragraph id="82289741306BFF82FF8EFBE3BFB50991" blockId="18.[123,726,1137,1192]" box="[123,726,1137,1162]" pageId="18" pageNumber="219">
<heading id="D960202D306BFF82FF8EFBE3BFB50991" box="[123,726,1137,1162]" fontSize="10" level="2" pageId="18" pageNumber="219" reason="4">
<emphasis id="B0E34B53306BFF82FF8EFBE3BFB50991" bold="true" box="[123,726,1137,1162]" pageId="18" pageNumber="219">
<taxonomicName id="4597ECC2306BFF82FF8EFBE3BF100991" ID-CoL="5XVVN" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,627,1137,1162]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="kathrynae" status="sp. nov.">
<emphasis id="B0E34B53306BFF82FF8EFBE3BC010992" bold="true" box="[123,354,1137,1161]" italics="true" pageId="18" pageNumber="219">Chlaenius kathrynae</emphasis>
Kavanaugh and Rainio
</taxonomicName>
,
<taxonomicNameLabel id="ABD0F628306BFF82FD75FBE0BFB50991" box="[640,726,1138,1162]" pageId="18" pageNumber="219" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA306BFF82FF8EFB00BDB409BC" box="[123,215,1170,1192]" pageId="18" pageNumber="219" type="description">
<paragraph id="82289741306BFF82FF8EFB00BDB409BC" blockId="18.[123,726,1137,1192]" box="[123,215,1170,1192]" pageId="18" pageNumber="219">
<figureCitation id="1AAC8BC4306BFF82FF8EFB00BDB409BC" box="[123,215,1170,1192]" captionStart="FIGURE" captionStartId="19.[155,167,1621,1640]" captionTargetBox="[152,1175,216,1604]" captionTargetId="figure-67@19.[120,1176,216,1618]" captionTargetPageId="19" captionText="FIGURE 11. Digital images of holotype male of Chlaenius kathrynae Kavanaugh &amp; Rainio sp. nov. A Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799465" httpUri="https://zenodo.org/record/13799465/files/figure.png" pageId="18" pageNumber="219">Figure 11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA306BFF84FF5EFB57BF8A0985" lastPageId="20" lastPageNumber="221" pageId="18" pageNumber="219" type="materials_examined">
<paragraph id="82289741306BFF82FF5EFB57BF680BA6" blockId="18.[123,1174,1221,1726]" pageId="18" pageNumber="219">
<emphasis id="B0E34B53306BFF82FF5EFB57BC3809C7" bold="true" box="[171,347,1221,1245]" pageId="18" pageNumber="219">
<smallCapsWord id="84CE019D306BFF82FF5EFB57BD8709C7" baselines="1239,1239" box="[171,228,1221,1245]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="18" pageNumber="219">TYPE</smallCapsWord>
<smallCapsWord id="84CE019D306BFF82FF1BFB5BBC3809C7" baselines="1239" box="[238,347,1225,1244]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="18" pageNumber="219">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C306BFF82FE77FB57BC160886" collectingDate="2001-01-02" collectingDateMax="2001-01-22" collectingDateMin="2001-01-02" collectorName="D. H. &amp; K. M. Kavanaugh &amp; R. L. Brett &amp; E. Elsom &amp; F. Vargas &amp; E. F. Randrianifirina &amp; N. Rasoamananana &amp; T. J. Ravelomanana &amp; H. C. Raveloson" country="Madagascar" elevation="1150" latitude="-21.24032" location="Vohiparara area" longLatPrecision="1" longitude="47.39399" municipality="Ranomafana National Park" pageId="18" pageNumber="219" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="5D2C29E3306BFF82FE77FB57BC8609C6" box="[386,485,1221,1245]" pageId="18" pageNumber="219" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC4306BFF82FE02FB57BF1B09C6" box="[503,632,1221,1246]" captionStart="FIGURE" captionStartId="19.[155,167,1621,1640]" captionTargetBox="[152,1175,216,1604]" captionTargetId="figure-67@19.[120,1176,216,1618]" captionTargetPageId="19" captionText="FIGURE 11. Digital images of holotype male of Chlaenius kathrynae Kavanaugh &amp; Rainio sp. nov. A Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799465" httpUri="https://zenodo.org/record/13799465/files/figure.png" pageId="18" pageNumber="219">Figs.11A–B</figureCitation>
), a male, in CAS, labeled: “CASENT 1002782”/ “
<collectingCountry id="FA80D7D1306BFF82FF72FB77BC2609E6" box="[135,325,1253,1277]" name="Madagascar" pageId="18" pageNumber="219">MADAGASCAR</collectingCountry>
,
<collectingRegion id="405359A3306BFF82FEA6FB77BF2009E6" box="[339,579,1253,1277]" country="Madagascar" name="Fianarantsoa" pageId="18" pageNumber="219">Fianarantsoa Province</collectingRegion>
,
<collectingMunicipality id="624C0D3B306BFF82FDA7FB77BE1509E6" box="[594,886,1253,1277]" pageId="18" pageNumber="219">Ranomafana National Park</collectingMunicipality>
,
<location id="8748C19A306BFF82FC70FB77B94D09E6" LSID="urn:lsid:plazi:treatment:0A3E2657306BFF84FF8EFBE3BF8A0985:8748C19A306BFF82FC70FB77B94D09E6" box="[901,1070,1253,1277]" country="Madagascar" latitude="-21.24032" longLatPrecision="1" longitude="47.39399" municipality="Ranomafana National Park" name="Vohiparara area" pageId="18" pageNumber="219" stateProvince="Fianarantsoa">Vohiparara area</location>
,
<quantity id="456F3AA4306BFF82FBC8FB77B9EC09E6" box="[1085,1167,1253,1277]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="18" pageNumber="219" unit="m" value="1150.0">
<elevation id="09BA7072306BFF82FBC8FB77B9EC09E6" box="[1085,1167,1253,1277]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="18" pageNumber="219" unit="m" value="1150.0">1150 m</elevation>
</quantity>
, mixed tropical forest,
<date id="F629B181306BFF82FE9EFA97BF570806" box="[363,564,1285,1310]" pageId="18" pageNumber="219" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">
<collectingDate id="E66D4869306BFF82FE9EFA97BF570806" box="[363,564,1285,1310]" pageId="18" pageNumber="219" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">2-22 January 2001</collectingDate>
</date>
,”/ “
<geoCoordinate id="E7A3F186306BFF82FD96FA97BF830806" box="[611,736,1285,1309]" degrees="21. 24032" direction="south" orientation="latitude" pageId="18" pageNumber="219" precision="1" value="-21.24032">21.24032°S</geoCoordinate>
/
<geoCoordinate id="E7A3F186306BFF82FD05FA97BE0C0806" box="[752,879,1285,1309]" degrees="47. 39399" direction="east" orientation="longitude" pageId="18" pageNumber="219" precision="1" value="47.39399">47.39399°E</geoCoordinate>
, Stop# DHK-01-002,
<collectorName id="2F62F297306BFF82FB97FA94BC3D0826" pageId="18" pageNumber="219">D.H. &amp; K.M. Kavanaugh</collectorName>
,
<collectorName id="2F62F297306BFF82FE86FAB4BC850826" box="[371,486,1318,1341]" pageId="18" pageNumber="219">R.L. Brett</collectorName>
,
<collectorName id="2F62F297306BFF82FE09FAB4BF060826" box="[508,613,1317,1341]" pageId="18" pageNumber="219">E. Elsom</collectorName>
,
<collectorName id="2F62F297306BFF82FD8EFAB4BF870825" box="[635,740,1318,1342]" pageId="18" pageNumber="219">F. Vargas</collectorName>
, Ranaivosolo,”/ “
<collectorName id="2F62F297306BFF82FC46FAB4B9EC0826" box="[947,1167,1317,1341]" pageId="18" pageNumber="219">E.F. Randrianifirina</collectorName>
,
<collectorName id="2F62F297306BFF82FF8EFAD4BC2A0846" box="[123,329,1350,1373]" pageId="18" pageNumber="219">N. Rasoamananana</collectorName>
,
<collectorName id="2F62F297306BFF82FEA3FAD4BF420846" box="[342,545,1349,1373]" pageId="18" pageNumber="219">T.J. Ravelomanana</collectorName>
, and
<collectorName id="2F62F297306BFF82FDA9FAD4BE640846" box="[604,775,1349,1373]" pageId="18" pageNumber="219">H.C. Raveloson</collectorName>
collectors”/ “collected in daytime by treading marsh vegetation and shore”/ “
<typeStatus id="5D2C29E3306BFF82FDDAFAF4BFDF0865" box="[559,700,1382,1406]" pageId="18" pageNumber="219" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC2306BFF82FD30FAF7B9F60866" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[709,1173,1381,1406]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="kathrynae" status="sp. nov.">
<emphasis id="B0E34B53306BFF82FD30FAF7BEC60866" box="[709,933,1381,1405]" italics="true" pageId="18" pageNumber="219">Chlaenius kathrynae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628306BFF82FF8EFA14BDD60886" box="[123,181,1414,1437]" pageId="18" pageNumber="219" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label]
</materialsCitation>
.
<materialsCitation id="32FF9D1C306BFF82FE71FA14B9530886" box="[388,1072,1413,1438]" collectionCode="CAS, MNHN, MZF" country="Madagascar" pageId="18" pageNumber="219" specimenCount="4" specimenCount-male="4" stateProvince="Fianarantsoa" typeStatus="paratype">
<typeStatus id="5D2C29E3306BFF82FE71FA14BC8F0885" box="[388,492,1414,1438]" pageId="18" pageNumber="219" type="paratype">Paratypes</typeStatus>
(total of 8):
<specimenCount id="94915CC8306BFF82FD82FA17BFA90886" box="[631,714,1413,1437]" count="4" pageId="18" pageNumber="219" type="male">4 males</specimenCount>
(in
<collectionCode id="E4860F84306BFF82FD0DFA17BE4F0886" box="[760,812,1413,1437]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="18" pageNumber="219" type="Museum">CAS</collectionCode>
,
<collectionCode id="E4860F84306BFF82FCCFFA14BEE80886" box="[826,907,1414,1437]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="18" pageNumber="219">MNHN</collectionCode>
and
<collectionCode id="E4860F84306BFF82FC37FA14BE9A0886" box="[962,1017,1414,1437]" pageId="18" pageNumber="219">MZF</collectionCode>
) and
</materialsCitation>
<materialsCitation id="32FF9D1C306BFF82FBCDFA17BEF008C6" collectionCode="CAS" country="Madagascar" pageId="18" pageNumber="219" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<specimenCount id="94915CC8306BFF82FBCDFA17B9F50886" box="[1080,1174,1413,1437]" count="1" pageId="18" pageNumber="219" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F84306BFF82FF6AFA37BDB008A6" box="[159,211,1445,1469]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="18" pageNumber="219" type="Museum">CAS</collectionCode>
) labeled same as
<typeStatus id="5D2C29E3306BFF82FE64FA37BC8D08A6" box="[401,494,1445,1469]" pageId="18" pageNumber="219" type="holotype">holotype</typeStatus>
except “CASENT 1002783”, “CASENT 1002784”, “CASENT 1002786”, “CASENT 1002787”, and “CASENT 1002785”, respectively
</materialsCitation>
;
<materialsCitation id="32FF9D1C306BFF82FC51FA57BCD40B45" collectingDate="1998-04-23" collectionCode="CAS" collectorName="D. H. Kavanaugh" country="Madagascar" elevation="1050" latitude="-21.226667" location="Vohiparara area" longLatPrecision="126" longitude="47.383335" municipality="Ranomafana National Park" pageId="18" pageNumber="219" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<specimenCount id="94915CC8306BFF82FC51FA57BE8C08C6" box="[932,1007,1477,1501]" count="1" pageId="18" pageNumber="219" type="male">1 male</specimenCount>
(
<collectionCode id="E4860F84306BFF82FBF7FA57B95508C6" box="[1026,1078,1477,1501]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="18" pageNumber="219" type="Museum">CAS</collectionCode>
) labeled “CASENT 1049008”/ “
<collectingCountry id="FA80D7D1306BFF82FE70FA77BF2008E6" box="[389,579,1509,1533]" name="Madagascar" pageId="18" pageNumber="219">MADAGASCAR</collectingCountry>
,
<collectingRegion id="405359A3306BFF82FDA2FA77BE2F08E6" box="[599,844,1509,1533]" country="Madagascar" name="Fianarantsoa" pageId="18" pageNumber="219">Fianarantsoa Province</collectingRegion>
,
<collectingMunicipality id="624C0D3B306BFF82FC95FA77B9EC08E6" box="[864,1167,1509,1533]" pageId="18" pageNumber="219">Ranomafana National Park</collectingMunicipality>
,
<location id="8748C19A306BFF82FF8EF997BC410B06" LSID="urn:lsid:plazi:treatment:0A3E2657306BFF84FF8EFBE3BF8A0985:8748C19A306BFF82FF8EF997BC410B06" box="[123,290,1541,1565]" country="Madagascar" latitude="-21.226667" longLatPrecision="126" longitude="47.383335" municipality="Ranomafana National Park" name="Vohiparara area" pageId="18" pageNumber="219" stateProvince="Fianarantsoa">Vohiparara area</location>
,
<quantity id="456F3AA4306BFF82FEC5F997BCE20B06" box="[304,385,1541,1565]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="18" pageNumber="219" unit="m" value="1050.0">
<elevation id="09BA7072306BFF82FEC5F997BCE20B06" box="[304,385,1541,1565]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="18" pageNumber="219" unit="m" value="1050.0">1050 m</elevation>
</quantity>
, mixed tropical forest,
<date id="F629B181306BFF82FD8EF994BE6C0B05" box="[635,783,1541,1566]" pageId="18" pageNumber="219" value="1998-04-23">
<collectingDate id="E66D4869306BFF82FD8EF994BE6C0B05" box="[635,783,1541,1566]" pageId="18" pageNumber="219" value="1998-04-23">23 April 1998</collectingDate>
</date>
,”/ “
<geoCoordinate id="E7A3F186306BFF82FCC9F994BECB0B05" box="[828,936,1541,1566]" degrees="21" direction="south" minutes="13. 6" orientation="latitude" pageId="18" pageNumber="219" precision="92" value="-21.226667">21°13.6’S</geoCoordinate>
/
<geoCoordinate id="E7A3F186306BFF82FC43F994B9400B06" box="[950,1059,1542,1566]" degrees="47" direction="east" minutes="23. 0" orientation="longitude" pageId="18" pageNumber="219" precision="92" value="47.383335">47°23.0’E</geoCoordinate>
,”/ “Stop # 98-71,
<collectorName id="2F62F297306BFF82FF32F9B4BC190B26" box="[199,378,1573,1597]" pageId="18" pageNumber="219">D.H. Kavanaugh</collectorName>
collector collected in soil cracks, under dirt clods, and under root clods in abandoned rice paddy area”
</materialsCitation>
;
<materialsCitation id="32FF9D1C306BFF82FE3EF9D4BF670BA6" collectingDate="1994-11-25" collectionCode="MTEC" collectorName="M. A. Ivie &amp; D. A. Pollack" country="Madagascar" latitude="-21.256666" location="Fianaran. Pr. Ranomafana" longLatPrecision="21" longitude="47.420834" pageId="18" pageNumber="219" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="94915CC8306BFF82FE3EF9D4BF5B0B46" box="[459,568,1605,1630]" count="2" pageId="18" pageNumber="219" type="female">2 females</specimenCount>
(in
<collectionCode id="E4860F84306BFF82FD85F9D4BFD90B45" box="[624,698,1606,1630]" country="USA" httpUri="http://grbio.org/cool/zm6z-4gpy" name="Montana State Entomology Collection" pageId="18" pageNumber="219" type="Museum">MTEC</collectionCode>
) labeled “
<collectingCountry id="FA80D7D1306BFF82FCC0F9D4BE900B45" box="[821,1011,1606,1630]" name="Madagascar" pageId="18" pageNumber="219">MADAGASCAR</collectingCountry>
:
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N.P. HQ. area
<geoCoordinate id="E7A3F186306BFF82FE50F9F4BF400B65" box="[421,547,1637,1662]" degrees="21" direction="south" minutes="15" orientation="latitude" pageId="18" pageNumber="219" precision="15" seconds="24" value="-21.256666">21°15’24”S</geoCoordinate>
<geoCoordinate id="E7A3F186306BFF82FDDFF9F4BFC90B66" box="[554,682,1637,1662]" degrees="47" direction="east" minutes="25" orientation="longitude" pageId="18" pageNumber="219" precision="15" seconds="15" value="47.420834">47°25’15”E</geoCoordinate>
<date id="F629B181306BFF82FD44F9F7BE580B65" box="[689,827,1637,1662]" pageId="18" pageNumber="219" value="1994-11-25">
<collectingDate id="E66D4869306BFF82FD44F9F7BE580B65" box="[689,827,1637,1662]" pageId="18" pageNumber="219" value="1994-11-25">25NOV1994</collectingDate>
</date>
, at night
<collectorName id="2F62F297306BFF82FC54F9F4B9690B66" box="[929,1034,1637,1662]" pageId="18" pageNumber="219">M.A. Ivie</collectorName>
&amp;
<collectorName id="2F62F297306BFF82FBDBF9F4BDC40B86" pageId="18" pageNumber="219">D.A. Pollack</collectorName>
”. All
<typeStatus id="5D2C29E3306BFF82FF1FF914BC330B85" box="[234,336,1670,1694]" pageId="18" pageNumber="219" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="5D2C29E3306BFF82FD5CF914BE4E0B85" box="[681,813,1670,1694]" pageId="18" pageNumber="219" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4597ECC2306BFF82FCC0F917BDBD0BA6" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="kathrynae" status="sp. nov.">
<emphasis id="B0E34B53306BFF82FCC0F917B9770B86" box="[821,1044,1669,1693]" italics="true" pageId="18" pageNumber="219">Chlaenius kathrynae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628306BFF82FF10F934BC7C0BA6" box="[229,287,1702,1725]" pageId="18" pageNumber="219" rank="species">sp. n.</taxonomicNameLabel>
2015” [yellow label]
</materialsCitation>
.
</paragraph>
<caption id="D6E8C7C9306AFF83FF6EF9C7B9450B99" ID-DOI="http://doi.org/10.5281/zenodo.13799465" ID-Zenodo-Dep="13799465" httpUri="https://zenodo.org/record/13799465/files/figure.png" pageId="19" pageNumber="220" startId="19.[155,167,1621,1640]" targetBox="[152,1175,216,1604]" targetPageId="19" targetType="figure">
<paragraph id="82289741306AFF83FF6EF9C7B9450B99" blockId="19.[123,1173,1620,1666]" pageId="19" pageNumber="220">
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11. Digital images of holotype male of
<taxonomicName id="4597ECC2306AFF83FDCFF9C6BECE0B7C" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[570,941,1620,1640]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="220" phylum="Arthropoda" rank="species" species="kathrynae" status="sp. nov.">
<emphasis id="B0E34B53306AFF83FDCFF9C6BF8E0B7C" box="[570,749,1620,1639]" italics="true" pageId="19" pageNumber="220">Chlaenius kathrynae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628306AFF83FC41F9C7BE940B73" box="[948,1015,1621,1640]" pageId="19" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
A Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 1.0 mm.
</paragraph>
</caption>
<paragraph id="82289741306DFF84FF5EFF49B9500DE9" blockId="20.[123,1174,219,1182]" box="[171,1075,219,243]" pageId="20" pageNumber="221">
<materialsCitation id="32FF9D1C306DFF84FF5EFF49B94F0DE8" box="[171,1068,219,243]" country="Madagascar" location="Ranomafana National Park" pageId="20" pageNumber="221" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B53306DFF84FF5EFF49BC340DEA" bold="true" box="[171,343,219,243]" pageId="20" pageNumber="221">
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<typeStatus id="5D2C29E3306DFF84FF5EFF49BD870DE9" box="[171,228,219,243]" pageId="20" pageNumber="221">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D306DFF84FF19FF4CBC340DEA" baselines="236" box="[236,343,222,241]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="20" pageNumber="221">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D1306DFF84FE8BFF49BC9C0DE8" box="[382,511,219,243]" name="Madagascar" pageId="20" pageNumber="221">Madagascar</collectingCountry>
,
<collectingRegion id="405359A3306DFF84FDF8FF49BF9F0DE8" box="[525,764,219,243]" country="Madagascar" name="Fianarantsoa" pageId="20" pageNumber="221">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A306DFF84FCFFFF49B94F0DE8" LSID="urn:lsid:plazi:treatment:0A3E2657306BFF84FF8EFBE3BF8A0985:8748C19A306DFF84FCFFFF49B94F0DE8" box="[778,1068,219,243]" country="Madagascar" name="Ranomafana National Park" pageId="20" pageNumber="221" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="82289741306DFF84FF5EFF6EBCCB0C4F" blockId="20.[123,1174,219,1182]" pageId="20" pageNumber="221">
<emphasis id="B0E34B53306DFF84FF5EFF6EBC890C09" bold="true" box="[171,490,252,276]" pageId="20" pageNumber="221">
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<smallCapsWord id="84CE019D306DFF84FEC6FF6DBC2C0C09" baselines="269" box="[307,335,255,274]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="20" pageNumber="221">OF</smallCapsWord>
<smallCapsWord id="84CE019D306DFF84FEA1FF6DBCC40C09" baselines="269" box="[340,423,255,274]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="20" pageNumber="221">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D306DFF84FE58FF6DBC890C09" baselines="269" box="[429,490,255,274]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="20" pageNumber="221">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC2306DFF84FD1CFF69BE370C08" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[745,852,251,275]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="kathrynae">
<emphasis id="B0E34B53306DFF84FD1CFF69BE370C08" box="[745,852,251,275]" italics="true" pageId="20" pageNumber="221">kathrynae</emphasis>
</taxonomicName>
, is a noun in the genitive case, derived from the first given name of Kathryn May Kavanaugh, daughter of DHK and one of the collectors of the
<typeStatus id="5D2C29E3306DFF84FEDAFEAFBC3E0C4E" box="[303,349,317,341]" pageId="20" pageNumber="221">type</typeStatus>
series.
</paragraph>
<paragraph id="82289741306DFF84FF5EFECFBF930EE7" blockId="20.[123,1174,219,1182]" pageId="20" pageNumber="221">
<smallCapsWord id="84CE019D306DFF84FF5EFECFBC210C68" baselines="367,366" box="[171,322,349,373]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="20" pageNumber="221">
<emphasis id="B0E34B53306DFF84FF5EFECFBC210C68" bold="true" box="[171,322,349,373]" pageId="20" pageNumber="221">RECOGNITION</emphasis>
</smallCapsWord>
.— Size slightly below average for genus, SBL of males = 10.2–10.
<quantity id="456F3AA4306DFF84FBDBFECFB90D0C6F" box="[1070,1134,349,373]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="20" pageNumber="221" unit="mm" value="8.0">8 mm</quantity>
, of females 9.8–11.
<quantity id="456F3AA4306DFF84FED2FEEFBC0B0C8F" box="[295,360,381,405]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="221" unit="mm" value="1.0">1 mm</quantity>
. Members of this species (
<figureCitation id="1AAC8BC4306DFF84FD6FFEEFBF980C8E" box="[666,763,381,405]" captionStart="FIGURE" captionStartId="19.[155,167,1621,1640]" captionTargetBox="[152,1175,216,1604]" captionTargetId="figure-67@19.[120,1176,216,1618]" captionTargetPageId="19" captionText="FIGURE 11. Digital images of holotype male of Chlaenius kathrynae Kavanaugh &amp; Rainio sp. nov. A Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799465" httpUri="https://zenodo.org/record/13799465/files/figure.png" pageId="20" pageNumber="221">Fig. 11A</figureCitation>
) share the following features, which place them in genus
<taxonomicName id="4597ECC2306DFF84FEAEFE0FBCAE0CAE" authorityName="Motschulsky" authorityYear="1860" box="[347,461,413,437]" class="Insecta" family="Carabidae" genus="Chlaenites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B53306DFF84FEAEFE0FBCAE0CAE" box="[347,461,413,437]" italics="true" pageId="20" pageNumber="221">Chlaenites</emphasis>
</taxonomicName>
sense Jeannel (1948): antennomere 3 longer than antennomere 4; maxillary palpi setose, penultimate labial palpomeres 4 or 5 setose; pronotum with basolateral setae inserted distinctly anterior to the hind angles; elytra with pubescence restricted to near the striae on disk (
<emphasis id="B0E34B53306DFF84FF43FE6DBDB40F0D" box="[182,215,511,534]" italics="true" pageId="20" pageNumber="221">i.e.</emphasis>
, not present on the centers of intervals), except more generally and densely present on lateral intervals and near apices; and tarsi asetose dorsally, tarsomeres 5 with two rows of stout setae ventrally. In Jeannel’s (1949) key to subgenera of
<taxonomicName id="4597ECC2306DFF84FD56FDADBE760F4C" authorityName="Motschulsky" authorityYear="1860" box="[675,789,575,599]" class="Insecta" family="Carabidae" genus="Chlaenites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B53306DFF84FD56FDADBE760F4C" box="[675,789,575,599]" italics="true" pageId="20" pageNumber="221">Chlaenites</emphasis>
</taxonomicName>
, members of this species key best, although not fully, to subgenus
<taxonomicName id="4597ECC2306DFF84FE20FDF2BE4F0F63" authority="Kuntzen (1919)" authorityName="Kuntzen" authorityYear="1919" box="[469,812,608,632]" class="Insecta" family="Carabidae" genus="Chlaeniostenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B53306DFF84FE20FDF2BF1A0F63" box="[469,633,608,632]" italics="true" pageId="20" pageNumber="221">Chlaeniostenus</emphasis>
<bibRefCitation id="E606EAB0306DFF84FD77FDF2BE4F0F63" author="KUNTZEN, H." box="[642,812,608,632]" pageId="20" pageNumber="221" pagination="1918 - 19" refId="ref31152" refString="KUNTZEN, H. 1919. Die Carabinenfaune Deutsch-Sudwestafrikas. Mitteilungen aus dem Zoologischen Museum in Berlin 9 [1918 - 19]: 91 - 156." type="journal article" year="1919">Kuntzen (1919)</bibRefCitation>
</taxonomicName>
, which is now considered a subgenus of
<taxonomicName id="4597ECC2306DFF84FF2BFD12BC880F83" authority="(Lorenz 2005)" baseAuthorityName="Lorenz" baseAuthorityYear="2005" box="[222,491,640,664]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B53306DFF84FF2BFD12BC290F83" box="[222,330,640,664]" italics="true" pageId="20" pageNumber="221">Chlaenius</emphasis>
(
<bibRefCitation id="E606EAB0306DFF84FEAFFD12BC810F83" author="LORENZ, W." box="[346,482,640,664]" pageId="20" pageNumber="221" refId="ref31666" refString="LORENZ, W. 2005. Systematic List of Extant Ground Beetles of the World (Insecta Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second edition. W. Lorenz, Tutzing. 530 pp." type="book" year="2005">Lorenz 2005</bibRefCitation>
)
</taxonomicName>
. Shared with other Malagasy members of this subgenus are the following features: pronotum relatively narrow, more or less cordiform, narrowed basally, with the anterior angles not projected anterior of the basal margin and the anterolateral areas of the pronotum distinctly curved ventrally; and the setose punctures along the lateral margins of the elytral intervals not or only faintly evident, most clearly evident in the apical half only. Jeannel included three other species in this subgenus; namely,
<taxonomicName id="4597ECC2306DFF84FD92FCB0BEB40E21" authority="Klug (1833)" authorityName="Klug" authorityYear="1833" box="[615,983,802,826]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="attenuatus">
<emphasis id="B0E34B53306DFF84FD92FCB0BE290E21" box="[615,842,802,826]" italics="true" pageId="20" pageNumber="221">Chlaenius attenuatus</emphasis>
<bibRefCitation id="E606EAB0306DFF84FCA7FCB0BEB40E21" author="KLUG, J. C. F." box="[850,983,802,826]" pageId="20" pageNumber="221" pagination="91 - 223" refId="ref30967" refString="KLUG, J. C. F. 1833. Bericht uber eine auf Madagascar veranstaltete. Sammlung von Insecten aus der Ordnung Coleoptera. Abhandlungen der Koniglichen Preussischen Akademie der Wissenschaften 1832 - 33: 91 - 223 + 5 pls." type="book chapter" year="1833">Klug (1833)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4597ECC2306DFF84FC13FCB0BC110E41" authority="Chaudoir (1876)" authorityName="Chaudoir" authorityYear="1876" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="subovatus">
<emphasis id="B0E34B53306DFF84FC13FCB0BDD00E41" italics="true" pageId="20" pageNumber="221">Chlaenius subovatus</emphasis>
<bibRefCitation id="E606EAB0306DFF84FF48FCD0BC110E41" author="CHAUDOIR, M." box="[189,370,834,858]" pageId="20" pageNumber="221" pagination="3 - 315" refId="ref29840" refString="CHAUDOIR, M. 1876. Monographie des chleniens. Annali del Museo Civico di Storia Naturelle di Genova 8: 3 - 315." type="journal article" year="1876">Chaudoir (1876)</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4597ECC2306DFF84FE41FCD0BE400E41" authority="Dejean (1831)" authorityName="Dejean" authorityYear="1831" box="[436,803,834,858]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="sellatus">
<emphasis id="B0E34B53306DFF84FE41FCD0BF180E41" box="[436,635,834,858]" italics="true" pageId="20" pageNumber="221">Chlaenius sellatus</emphasis>
<bibRefCitation id="E606EAB0306DFF84FD73FCD0BE400E41" author="DEJEAN, P. F. M. A." box="[646,803,834,858]" pageId="20" pageNumber="221" refId="ref29934" refString="DEJEAN, P. F. M. A. 1831. Species general des coleopteres, de la collection de M. le Comte Dejean. Tome cinquieme. Mequignon-Marvis, Paris, France. viii + 883 pp." type="book" year="1831">Dejean (1831)</bibRefCitation>
</taxonomicName>
. Members of
<taxonomicName id="4597ECC2306DFF84FC36FCD1B9320E41" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[963,1105,834,858]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="kathrynae">
<emphasis id="B0E34B53306DFF84FC36FCD1B9320E41" box="[963,1105,834,858]" italics="true" pageId="20" pageNumber="221">C. kathrynae</emphasis>
</taxonomicName>
differ from those of the other species in lacking any trace of the pale lateral and apical elytral border distinct in members of the other three species. They are also smaller than members of the other species (SBL in the latter ranges from 11.9 to 15.
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), have slightly broader and less cordiform pronota and males have genitalia (
<figureCitation id="1AAC8BC4306DFF84FE6EFC56BF470EC0" box="[411,548,964,988]" captionStart="FIGURE" captionStartId="19.[155,167,1621,1640]" captionTargetBox="[152,1175,216,1604]" captionTargetId="figure-67@19.[120,1176,216,1618]" captionTargetPageId="19" captionText="FIGURE 11. Digital images of holotype male of Chlaenius kathrynae Kavanaugh &amp; Rainio sp. nov. A Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799465" httpUri="https://zenodo.org/record/13799465/files/figure.png" pageId="20" pageNumber="221">Figs. 11C–D</figureCitation>
) that are very different in form from those of any of the other species (compare with Jeannel’s (1949) Figs. 388c-i)
</paragraph>
<paragraph id="82289741306DFF84FF5EFB96B9690907" blockId="20.[123,1174,219,1182]" box="[171,1034,1028,1053]" pageId="20" pageNumber="221">
<emphasis id="B0E34B53306DFF84FF5EFB96BC920900" bold="true" box="[171,497,1028,1052]" pageId="20" pageNumber="221">
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<smallCapsWord id="84CE019D306DFF84FEA8FB9ABC920900" baselines="1046" box="[349,497,1032,1051]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="20" pageNumber="221">DISTRIBUTION</smallCapsWord>
</emphasis>
.— At present, known only from the
<typeStatus id="5D2C29E3306DFF84FC8AFB97BECE0906" box="[895,941,1029,1053]" pageId="20" pageNumber="221">type</typeStatus>
locality.
</paragraph>
<paragraph id="82289741306DFF84FF5EFBB7BF8A0985" blockId="20.[123,1174,219,1182]" pageId="20" pageNumber="221">
<emphasis id="B0E34B53306DFF84FF5EFBB7BCC60920" bold="true" box="[171,421,1061,1085]" pageId="20" pageNumber="221">
<smallCapsWord id="84CE019D306DFF84FF5EFBB7BC6A0920" baselines="1079,1078" box="[171,265,1061,1085]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Habitat" pageId="20" pageNumber="221">HABITAT</smallCapsWord>
<smallCapsWord id="84CE019D306DFF84FEE4FBBABCC60920" baselines="1078" box="[273,421,1064,1083]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="20" pageNumber="221">DISTRIBUTION</smallCapsWord>
</emphasis>
.— Most specimens of the
<typeStatus id="5D2C29E3306DFF84FD3FFBB7BF9B0926" box="[714,760,1061,1085]" pageId="20" pageNumber="221">type</typeStatus>
series were collected in open habitats at the edges of mixed tropical forest in the Vohiparara area at an elevation of
<quantity id="456F3AA4306DFF84FC58FBD7BE9D0946" box="[941,1022,1093,1117]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="20" pageNumber="221" unit="m" value="1050.0">1050 m</quantity>
. Several were collected by treading down vegetation at the edge of a marsh, and
<specimenCount id="94915CC8306DFF84FCBDFBF4BEB80965" box="[840,987,1126,1150]" count="1" pageId="20" pageNumber="221" type="generic">one specimen</specimenCount>
was found under loose dirt and root clods in an abandoned rice paddy area.
</paragraph>
</subSubSection>
</treatment>
</document>
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