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<document id="86A5DDCD8CA0838E379D54F2CA327BE5" ID-DOI="10.5281/zenodo.13799439" ID-ISSN="0068-547X" ID-Zenodo-Dep="13799439" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538212676" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Rainio, Johanna" docDate="2016" docId="0A3E26573069FF82FF8EFD07BCD40E48" docLanguage="en" docName="ProcCalifAcadSci.63.7.201-268.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (7)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Trichinillus (Mallopelmus) ranomafanae Kavanaugh and Rainio 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="219" masterDocId="F6075E2F3079FF90FFF5FF92BD630D1B" masterDocTitle="Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar" masterLastPageNumber="268" masterPageNumber="201" pageNumber="217" updateTime="1726771873529" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="2AF0B3AB1979700C94A1205964B9DF04" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C7B6BA384153615B10D6F4A03DABDF6A">
<mods:title id="9CC9CD558011D63FF2179FCE2CCB190B">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
</mods:titleInfo>
<mods:name id="8CCCAA8B02AE09C3A063E63C6927D662" type="personal">
<mods:role id="05525E2519D1F8CDB8B840E1C7966243">
<mods:roleTerm id="DCF0B5DBDF6F40CFD3F6A18B482BD145">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4846C64F009FDB9CE9829C609B5B35FC">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="5E4836D6E5EE6CBCC0812043E4CCA76F">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
</mods:name>
<mods:name id="A3593560A5DC4CD195DCBA1407103D7D" type="personal">
<mods:role id="B9FAAA694650BFDEF87F38D630BE2B28">
<mods:roleTerm id="A71A8351D324CB7BE06D6B2A38A16AED">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0F52AF84B91DF25630A398B8940305AA">Rainio, Johanna</mods:namePart>
<mods:affiliation id="7191BB43A5EDB28426E94C7A5743D8B8">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
</mods:name>
<mods:typeOfResource id="AB10C90BE71C047FB7FFFEF1BE2EE42C">text</mods:typeOfResource>
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<mods:titleInfo id="77F0B2E3CF90C32888A1982387323EF2">
<mods:title id="A980F94A3E5D5815C732FA91C9A4CC88">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="DD635B2969BE068BF59F9CF2ECCB435F">
<mods:date id="218340E7761409E15D3C1289A467DEB8">2016</mods:date>
<mods:detail id="017FD084FECFE5113BA193C5797CAA11" type="pubDate">
<mods:number id="866C76875A664E4DA9F11CF16B7E7807">2016-04-29</mods:number>
</mods:detail>
<mods:detail id="A320B6A9F90052CE78D0D6FAA21E81F6" type="volume">
<mods:number id="6216756222E77AEB5CAD8D966C5EA780">63</mods:number>
</mods:detail>
<mods:detail id="013D11F4252CE6CED1484075E7013973" type="issue">
<mods:number id="A72656A6ABFC545D4EAD2F6A296F77C3">7</mods:number>
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<mods:start id="039133B27A3BD323D78BDA0BBCECAA0A">201</mods:start>
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<mods:classification id="1CD521EBA0A8F4AF72A7D72BF7DB7DB2">journal article</mods:classification>
<mods:identifier id="9DAA986F52828F580E096C1C3E9B911B" type="DOI">10.5281/zenodo.13799439</mods:identifier>
<mods:identifier id="8AE2193CE6C231B2DA8CB8E36A17AD49" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="4DDAFC08F7274D720B875EBB716F5004" type="Zenodo-Dep">13799439</mods:identifier>
</mods:mods>
<treatment id="0A3E26573069FF82FF8EFD07BCD40E48" ID-DOI="http://doi.org/10.5281/zenodo.13799457" ID-Zenodo-Dep="13799457" LSID="urn:lsid:plazi:treatment:0A3E26573069FF82FF8EFD07BCD40E48" httpUri="http://treatment.plazi.org/id/0A3E26573069FF82FF8EFD07BCD40E48" lastPageId="18" lastPageNumber="219" pageId="16" pageNumber="217">
<subSubSection id="CA8DC4CA3069FF80FF8EFD07BED60FB5" box="[123,949,661,686]" pageId="16" pageNumber="217" type="nomenclature">
<paragraph id="822897413069FF80FF8EFD07BED60FB5" blockId="16.[123,949,661,716]" box="[123,949,661,686]" pageId="16" pageNumber="217">
<heading id="D960202D3069FF80FF8EFD07BED60FB5" box="[123,949,661,686]" fontSize="10" level="2" pageId="16" pageNumber="217" reason="4">
<emphasis id="B0E34B533069FF80FF8EFD07BED60FB5" bold="true" box="[123,949,661,686]" pageId="16" pageNumber="217">
<taxonomicName id="4597ECC23069FF80FF8EFD07BE320FB5" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,849,661,686]" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov." subGenus="Mallopelmus">
<emphasis id="B0E34B533069FF80FF8EFD07BD9F0FB6" bold="true" box="[123,252,661,685]" italics="true" pageId="16" pageNumber="217">Trichinillus</emphasis>
(
<emphasis id="B0E34B533069FF80FEF9FD07BCFF0FB6" bold="true" box="[268,412,661,685]" italics="true" pageId="16" pageNumber="217">Mallopelmus</emphasis>
)
<emphasis id="B0E34B533069FF80FE59FD07BF230FB6" bold="true" box="[428,576,661,685]" italics="true" pageId="16" pageNumber="217">ranomafanae</emphasis>
Kavanaugh and Rainio
</taxonomicName>
,
<taxonomicNameLabel id="ABD0F6283069FF80FCAAFD04BED60FB5" box="[863,949,662,686]" pageId="16" pageNumber="217" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3069FF80FF8EFD24BE160FEC" pageId="16" pageNumber="217" type="description">
<paragraph id="822897413069FF80FF8EFD24BDBB0FD0" blockId="16.[123,949,661,716]" box="[123,216,694,716]" pageId="16" pageNumber="217">
<figureCitation id="1AAC8BC43069FF80FF8EFD24BDBB0FD0" box="[123,216,694,716]" captionStart="FIGURE" captionStartId="17.[155,167,1571,1590]" captionTargetBox="[139,1176,235,1551]" captionTargetId="figure-68@17.[120,1176,216,1569]" captionTargetPageId="17" captionText="FIGURE 10. Digital images of holotype male of Trichinillus (Mallopelmus) ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799463" httpUri="https://zenodo.org/record/13799463/files/figure.png" pageId="16" pageNumber="217">Figure 10</figureCitation>
</paragraph>
<paragraph id="822897413069FF80FF8EFD73BE160FEC" blockId="16.[123,885,737,759]" box="[123,885,737,759]" pageId="16" pageNumber="217">
<emphasis id="B0E34B533069FF80FF8EFD73BC5C0FEC" box="[123,319,737,759]" italics="true" pageId="16" pageNumber="217">
<taxonomicName id="4597ECC23069FF80FF8EFD73BD9B0FED" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[123,248,737,758]" class="Insecta" family="Carabidae" genus="Mallopelmus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="genus" status="n. sp.">Mallopelmus</taxonomicName>
<taxonomicNameLabel id="ABD0F6283069FF80FF0AFD70BC500FEC" box="[255,307,738,759]" pageId="16" pageNumber="217" rank="species">n. sp.</taxonomicNameLabel>
1
</emphasis>
; Rainio 2009: 32, Rainio 2012: 74 (informal designation).
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3069FF82FF5EFC89BCD40E48" lastPageId="18" lastPageNumber="219" pageId="16" pageNumber="217" type="materials_examined">
<paragraph id="822897413069FF80FF5EFC89BE3E0EEE" blockId="16.[123,1174,795,1726]" pageId="16" pageNumber="217">
<emphasis id="B0E34B533069FF80FF5EFC89BC3E0E29" bold="true" box="[171,349,795,819]" pageId="16" pageNumber="217">
<smallCapsWord id="84CE019D3069FF80FF5EFC89BD870E29" baselines="813,813" box="[171,228,795,819]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="16" pageNumber="217">TYPE</smallCapsWord>
<smallCapsWord id="84CE019D3069FF80FF05FC8DBC3E0E29" baselines="813" box="[240,349,799,818]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="16" pageNumber="217">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C3069FF80FE70FC89BCAB0E8F" collectingDate="2002-10-14" collectorName="F. Ratalata &amp; J. Rainio" country="Madagascar" location="Buffer" pageId="16" pageNumber="217" specimenCount="2" typeStatus="holotype">
<typeStatus id="5D2C29E33069FF80FE70FC89BC8B0E28" box="[389,488,795,819]" pageId="16" pageNumber="217" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC43069FF80FE08FC89BFE80E28" box="[509,651,795,819]" captionStart="FIGURE" captionStartId="17.[155,167,1571,1590]" captionTargetBox="[139,1176,235,1551]" captionTargetId="figure-68@17.[120,1176,216,1569]" captionTargetPageId="17" captionText="FIGURE 10. Digital images of holotype male of Trichinillus (Mallopelmus) ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799463" httpUri="https://zenodo.org/record/13799463/files/figure.png" pageId="16" pageNumber="217">Figs. 10A–B</figureCitation>
), a male, in CAS, labeled: “
<collectingCountry id="FA80D7D13069FF80FC2DFC89B9F50E28" box="[984,1174,795,819]" name="Madagascar" pageId="16" pageNumber="217">MADAGASCAR</collectingCountry>
Ranomafana NP.
<location id="8748C19A3069FF80FEC6FCA9BC1B0E48" LSID="urn:lsid:plazi:treatment:0A3E26573069FF82FF8EFD07BCD40E48:8748C19A3069FF80FEC6FCA9BC1B0E48" box="[307,376,827,851]" country="Madagascar" name="Buffer" pageId="16" pageNumber="217">Buffer</location>
zone, degraded forest
<date id="F629B1813069FF80FD92FCAEBFBC0E4F" box="[615,735,828,852]" pageId="16" pageNumber="217" value="2002-10-14">
<collectingDate id="E66D48693069FF80FD92FCAEBFBC0E4F" box="[615,735,828,852]" pageId="16" pageNumber="217" value="2002-10-14">14.10.2002</collectingDate>
</date>
”/ “collected by hand in daytime,
<collectorName id="2F62F2973069FF80FBB7FCAEBDC20E6F" pageId="16" pageNumber="217">F. Ratalata</collectorName>
&amp;
<collectorName id="2F62F2973069FF80FF36FCCEBC420E6F" box="[195,289,860,884]" pageId="16" pageNumber="217">J. Rainio</collectorName>
collectors”/ “
<typeStatus id="5D2C29E33069FF80FE40FCCEBF210E6F" box="[437,578,860,884]" pageId="16" pageNumber="217" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC23069FF80FDBDFCCEBDA10E8F" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov." subGenus="Mallopelmus">
<emphasis id="B0E34B533069FF80FDBDFCCEBFA00E6F" box="[584,707,860,884]" italics="true" pageId="16" pageNumber="217">Trichinillus</emphasis>
(
<emphasis id="B0E34B533069FF80FD27FCCEBE3E0E6F" box="[722,861,860,884]" italics="true" pageId="16" pageNumber="217">Mallopelmus</emphasis>
)
<emphasis id="B0E34B533069FF80FC99FCCEBE990E6F" box="[876,1018,860,884]" italics="true" pageId="16" pageNumber="217">ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283069FF80FF3FFCEFBC650E8F" box="[202,262,893,916]" pageId="16" pageNumber="217" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label]
</materialsCitation>
.
<materialsCitation id="32FF9D1C3069FF80FE22FCEFBE350EEE" collectingDate="2004-03-05" collectionCode="MNHN" collectorName="F. Ratalata &amp; J. Rainio" country="Madagascar" elevation="600" location="Talatakely" pageId="16" pageNumber="217" specimenCount="600" typeStatus="paratype">
<typeStatus id="5D2C29E33069FF80FE22FCEFBF5C0E8E" box="[471,575,893,917]" pageId="16" pageNumber="217" type="paratype">Paratypes</typeStatus>
(only 1): a female (in
<collectionCode id="E4860F843069FF80FCCFFCEFBEE80E8F" box="[826,907,893,916]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="16" pageNumber="217">MNHN</collectionCode>
) labelled “
<collectingCountry id="FA80D7D13069FF80FBF3FCEEBDD10EAE" name="Madagascar" pageId="16" pageNumber="217">MADAGASCAR</collectingCountry>
Ranomafana NP.
<location id="8748C19A3069FF80FE84FC0EBCBD0EAF" LSID="urn:lsid:plazi:treatment:0A3E26573069FF82FF8EFD07BCD40E48:8748C19A3069FF80FE84FC0EBCBD0EAF" box="[369,478,924,948]" country="Madagascar" name="Talatakely" pageId="16" pageNumber="217">Talatakely</location>
, trail B,
<specimenCount id="94915CC83069FF80FDCEFC0EBFE60EAF" box="[571,645,924,948]" count="600" pageId="16" pageNumber="217" type="generic">
<quantity id="456F3AA43069FF80FDCEFC0EBF1D0EAF" box="[571,638,924,948]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="16" pageNumber="217" unit="m" value="600.0">
<elevation id="09BA70723069FF80FDCEFC0EBF1D0EAF" box="[571,638,924,948]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="16" pageNumber="217" unit="m" value="600.0">600 m</elevation>
</quantity>
.
</specimenCount>
<date id="F629B1813069FF80FD7EFC0EBE600EAE" box="[651,771,924,949]" pageId="16" pageNumber="217" value="2004-03-05">
<collectingDate id="E66D48693069FF80FD7EFC0EBE600EAE" box="[651,771,924,949]" pageId="16" pageNumber="217" value="2004-03-05">05.03.2004</collectingDate>
</date>
”/ “collected by hand in daytime from dead leaves in bushes,
<collectorName id="2F62F2973069FF80FE81FC2FBC850ECE" box="[372,486,957,981]" pageId="16" pageNumber="217">F. Ratalata</collectorName>
and
<collectorName id="2F62F2973069FF80FDE8FC2FBF1E0ECE" box="[541,637,957,981]" pageId="16" pageNumber="217">J. Rainio</collectorName>
collectors”/ “
<typeStatus id="5D2C29E33069FF80FCE1FC2FBEFB0ECE" box="[788,920,957,981]" pageId="16" pageNumber="217" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4597ECC23069FF80FC55FC2FBF4C0EEE" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov." subGenus="Mallopelmus">
<emphasis id="B0E34B533069FF80FC55FC2FB9780ECE" box="[928,1051,957,981]" italics="true" pageId="16" pageNumber="217">Trichinillus</emphasis>
(
<emphasis id="B0E34B533069FF80FBD9FC2FBDC50EEE" italics="true" pageId="16" pageNumber="217">Mallopelmus</emphasis>
)
<emphasis id="B0E34B533069FF80FF43FC4FBC270EEE" box="[182,324,989,1013]" italics="true" pageId="16" pageNumber="217">ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283069FF80FDC2FC4CBF120EEE" box="[567,625,990,1013]" pageId="16" pageNumber="217" rank="species">sp. n.</taxonomicNameLabel>
2015” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="822897413069FF80FF5EFC6CB950090E" blockId="16.[123,1174,795,1726]" box="[171,1075,1022,1046]" pageId="16" pageNumber="217">
<materialsCitation id="32FF9D1C3069FF80FF5EFC6CB94F090D" box="[171,1068,1022,1046]" country="Madagascar" location="Ranomafana National Park" pageId="16" pageNumber="217" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B533069FF80FF5EFC6CBC34090F" bold="true" box="[171,343,1022,1046]" pageId="16" pageNumber="217">
<smallCapsWord id="84CE019D3069FF80FF5EFC6CBD87090F" baselines="1040,1039" box="[171,228,1022,1046]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="16" pageNumber="217">
<typeStatus id="5D2C29E33069FF80FF5EFC6CBD87090F" box="[171,228,1022,1046]" pageId="16" pageNumber="217">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D3069FF80FF19FB93BC34090F" baselines="1039" box="[236,343,1025,1044]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="16" pageNumber="217">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D13069FF80FE8BFC6CBC9C090D" box="[382,511,1022,1046]" name="Madagascar" pageId="16" pageNumber="217">Madagascar</collectingCountry>
,
<collectingRegion id="405359A33069FF80FDF8FC6CBF9F090D" box="[525,764,1022,1046]" country="Madagascar" name="Fianarantsoa" pageId="16" pageNumber="217">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A3069FF80FCFFFC6CB94F090D" LSID="urn:lsid:plazi:treatment:0A3E26573069FF82FF8EFD07BCD40E48:8748C19A3069FF80FCFFFC6CB94F090D" box="[778,1068,1022,1046]" country="Madagascar" name="Ranomafana National Park" pageId="16" pageNumber="217" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="822897413069FF80FF5EFB8CBECA094D" blockId="16.[123,1174,795,1726]" pageId="16" pageNumber="217">
<emphasis id="B0E34B533069FF80FF5EFB8CBC91092F" bold="true" box="[171,498,1054,1078]" pageId="16" pageNumber="217">
<smallCapsWord id="84CE019D3069FF80FF5EFB8CBC4D092F" baselines="1072,1071" box="[171,302,1054,1078]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="16" pageNumber="217">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D3069FF80FEC3FBB3BC31092F" baselines="1071" box="[310,338,1057,1076]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="16" pageNumber="217">OF</smallCapsWord>
<smallCapsWord id="84CE019D3069FF80FEACFBB3BCCF092F" baselines="1071" box="[345,428,1057,1076]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="16" pageNumber="217">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D3069FF80FE40FBB3BC91092F" baselines="1071" box="[437,498,1057,1076]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="16" pageNumber="217">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC23069FF80FD0CFB8CBEE4092D" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[761,903,1054,1078]" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533069FF80FD0CFB8CBEE4092D" box="[761,903,1054,1078]" italics="true" pageId="16" pageNumber="217">ranomafanae</emphasis>
</taxonomicName>
, is a noun in apposition, derived from the name of the national park in which the
<typeStatus id="5D2C29E33069FF80FD29FBADBE69094C" box="[732,778,1087,1111]" pageId="16" pageNumber="217">type</typeStatus>
was collected.
</paragraph>
<paragraph id="822897413069FF82FF5EFBCDBCB30F68" blockId="16.[123,1174,795,1726]" lastBlockId="18.[123,1174,219,852]" lastPageId="18" lastPageNumber="219" pageId="16" pageNumber="217">
<smallCapsWord id="84CE019D3069FF80FF5EFBCDBC21096E" baselines="1137,1136" box="[171,322,1119,1143]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="16" pageNumber="217">
<emphasis id="B0E34B533069FF80FF5EFBCDBC21096E" bold="true" box="[171,322,1119,1143]" pageId="16" pageNumber="217">RECOGNITION</emphasis>
</smallCapsWord>
.— Slightly smaller than average for subgenus, SBL male =
<quantity id="456F3AA43069FF80FC21FBCDB944096D" box="[980,1063,1119,1143]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="16" pageNumber="217" unit="mm" value="7.2">7.2 mm</quantity>
, female =
<quantity id="456F3AA43069FF80FF8EFB12BDAE098C" box="[123,205,1151,1175]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="16" pageNumber="217" unit="mm" value="7.5">7.5 mm</quantity>
. The following combination of features identify members of this species as coelostomine pterostichines of the genus
<taxonomicName id="4597ECC23069FF80FE47FB0DBF4E09AC" authorityName="Straneo" authorityYear="1938" box="[434,557,1183,1207]" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B533069FF80FE47FB0DBF4E09AC" box="[434,557,1183,1207]" italics="true" pageId="16" pageNumber="217">Trichinillus</emphasis>
</taxonomicName>
, subgenus
<taxonomicName id="4597ECC23069FF80FD5AFB0DBE5909AC" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[687,826,1183,1207]" class="Insecta" family="Carabidae" genus="Mallopelmus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B533069FF80FD5AFB0DBE5909AC" box="[687,826,1183,1207]" italics="true" pageId="16" pageNumber="217">Mallopelmus</emphasis>
</taxonomicName>
: antennae with antennomere 3 attached symmetrically or nearly symmetrically to antennomere 2; mentum with deep emargination, apex of median mental tooth distinctly posterior to apices of epilobes; elytron with epipleuron interrupted by internal plica subapically; parascutellar stria short but evident; striae smooth, impunctate; basal setiferious puncture situated at base of stria 2 + parascutellar stria; tarsomeres 1 to 4 of all legs with dense pads of setae ventrally; front and middle tarsomeres 1 to 3 with more or less faintly-defined longitudinal groove medially and depressed areas paralaterally on dorsal surface; male front tarsomeres 1 to 4 wide, dilated, not latero- or medioapically toothed; aedeagus of male with right face dorsal in repose, right paramere conchoid, left paramere reduced, short, apically digitiform. Of the three previously described species represented in Jeannel’s (1948) key to
<taxonomicName id="4597ECC23069FF80FF8EFA71BC3D08E0" box="[123,350,1507,1531]" pageId="16" pageNumber="217">
<emphasis id="B0E34B533069FF80FF8EFA71BC6508E0" box="[123,262,1507,1531]" italics="true" pageId="16" pageNumber="217">Mallopelmus</emphasis>
species
</taxonomicName>
, members of
<taxonomicName id="4597ECC23069FF80FE0AFA76BFCF08E0" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[511,684,1507,1531]" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533069FF80FE0AFA76BFCF08E0" box="[511,684,1507,1531]" italics="true" pageId="16" pageNumber="217">T. ranomafanae</emphasis>
</taxonomicName>
(
<figureCitation id="1AAC8BC43069FF80FD34FA71BE4708E0" box="[705,804,1507,1531]" captionStart="FIGURE" captionStartId="17.[155,167,1571,1590]" captionTargetBox="[139,1176,235,1551]" captionTargetId="figure-68@17.[120,1176,216,1569]" captionTargetPageId="17" captionText="FIGURE 10. Digital images of holotype male of Trichinillus (Mallopelmus) ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799463" httpUri="https://zenodo.org/record/13799463/files/figure.png" pageId="16" pageNumber="217">Fig. 10A</figureCitation>
) are most similar to
<taxonomicName id="4597ECC23069FF80FBEFFA71BCC50B07" authority="(Alluaud 1936)" baseAuthorityName="Alluaud" baseAuthorityYear="1936" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="abacetoides">
<emphasis id="B0E34B533069FF80FBEFFA71BD990B07" italics="true" pageId="16" pageNumber="217">Trichinillus abacetoides</emphasis>
(Alluaud 1936)
</taxonomicName>
members based on the following shared features: head with temporal area long, nearly as long as the diameter of the eye, and joined to the neck region at a very obtuse (about 165°) angle; eyes moderately convex and projected; pronotum with basal area distinctly and abruptly depressed relative to disc; and elytral base without margination medial to base of stria 6. Members of the other two species,
<taxonomicName id="4597ECC23069FF80FDF6F917BE820B86" authority="(Alluaud 1936)" baseAuthorityName="Alluaud" baseAuthorityYear="1936" box="[515,993,1669,1693]" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="dactyleuryoides">
<emphasis id="B0E34B533069FF80FDF6F917BE530B86" box="[515,816,1669,1693]" italics="true" pageId="16" pageNumber="217">Trichinillus dactyleuryoides</emphasis>
(Alluaud 1936)
</taxonomicName>
and
<taxonomicName id="4597ECC23069FF80FBEFF917BC110BA5" authority="(Jeannel 1948)" baseAuthorityName="Jeannel" baseAuthorityYear="1948" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="perrieri">
<emphasis id="B0E34B533069FF80FBEFF917BDAD0BA6" italics="true" pageId="16" pageNumber="217">Trichinillus perrieri</emphasis>
(Jeannel 1948)
</taxonomicName>
, differ in having the head with the temporal area shorter, about one-third as long as the diameter of the eye, and joined to the neck region at a moderately obtuse (about 135°) angle, eyes more convex and projected, pronotum not or less depressed, especially medially, and margination of the elytra base more varied, extended medially to the base of stria
<quantity id="456F3AA4306BFF82FC00FE89B97C0C28" box="[1013,1055,283,307]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="18" pageNumber="219" unit="in" value="3.0">3 in</quantity>
most individuals). The apex of the median lobe of the male genitalia is also differently shaped in each of these species (compare with Figs. 191c-d and 191f-g, respectively, in Jeannel 1948). Members of
<taxonomicName id="4597ECC2306BFF82FF8EFEEEBC470C88" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[123,292,379,403]" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53306BFF82FF8EFEEEBC470C88" box="[123,292,379,403]" italics="true" pageId="18" pageNumber="219">T. ranomafanae</emphasis>
</taxonomicName>
differ from those of
<taxonomicName id="4597ECC2306BFF82FDFDFEEEBFC10C88" baseAuthorityName="Alluaud" baseAuthorityYear="1936" box="[520,674,379,403]" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="abacetoides">
<emphasis id="B0E34B53306BFF82FDFDFEEEBFC10C88" box="[520,674,379,403]" italics="true" pageId="18" pageNumber="219">T. abacetoides</emphasis>
</taxonomicName>
in having: slightly larger size (SBL =
<quantity id="456F3AA4306BFF82FBB6FEE9B9F60C88" box="[1091,1173,379,403]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="18" pageNumber="219" unit="mm" value="6.4">6.4 mm</quantity>
in
<taxonomicName id="4597ECC2306BFF82FF6FFE0EBC540CA8" baseAuthorityName="Alluaud" baseAuthorityYear="1936" box="[154,311,411,435]" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="abacetoides">
<emphasis id="B0E34B53306BFF82FF6FFE0EBC540CA8" box="[154,311,411,435]" italics="true" pageId="18" pageNumber="219">T. abacetoides</emphasis>
</taxonomicName>
); pronotum with shallow but distinct subbasal sinuation of the lateral margin (absent from
<taxonomicName id="4597ECC2306BFF82FEFAFE2EBCC80CC8" baseAuthorityName="Alluaud" baseAuthorityYear="1936" box="[271,427,443,467]" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="abacetoides">
<emphasis id="B0E34B53306BFF82FEFAFE2EBCC80CC8" box="[271,427,443,467]" italics="true" pageId="18" pageNumber="219">T. abacetoides</emphasis>
</taxonomicName>
members); elytra relatively shorter and wider, especially across the base, and humeri broadly rounded and not at all sloped (humeri more slope in
<taxonomicName id="4597ECC2306BFF82FC32FE4EB9030CE8" baseAuthorityName="Alluaud" baseAuthorityYear="1936" box="[967,1120,475,499]" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="abacetoides">
<emphasis id="B0E34B53306BFF82FC32FE4EB9030CE8" box="[967,1120,475,499]" italics="true" pageId="18" pageNumber="219">T. abacetoides</emphasis>
</taxonomicName>
, and median lobe of male genitalia with apex straighter, slightly bent ventrally toward apex in lateral aspect (
<figureCitation id="1AAC8BC4306BFF82FF3AFD89BC4C0F28" box="[207,303,539,563]" captionStart="FIGURE" captionStartId="17.[155,167,1571,1590]" captionTargetBox="[139,1176,235,1551]" captionTargetId="figure-68@17.[120,1176,216,1569]" captionTargetPageId="17" captionText="FIGURE 10. Digital images of holotype male of Trichinillus (Mallopelmus) ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799463" httpUri="https://zenodo.org/record/13799463/files/figure.png" pageId="18" pageNumber="219">Fig. 10C</figureCitation>
) and slightly deflect right in dorsal view (
<figureCitation id="1AAC8BC4306BFF82FCF6FD89BE070F28" box="[771,868,539,563]" captionStart="FIGURE" captionStartId="17.[155,167,1571,1590]" captionTargetBox="[139,1176,235,1551]" captionTargetId="figure-68@17.[120,1176,216,1569]" captionTargetPageId="17" captionText="FIGURE 10. Digital images of holotype male of Trichinillus (Mallopelmus) ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13799463" httpUri="https://zenodo.org/record/13799463/files/figure.png" pageId="18" pageNumber="219">Fig. 10D</figureCitation>
) (not recurved dorsally and then ventrally in lateral aspect and straight in dorsal aspect as in
<taxonomicName id="4597ECC2306BFF82FCB3FDAEBE810F48" baseAuthorityName="Alluaud" baseAuthorityYear="1936" box="[838,994,571,595]" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="abacetoides">
<emphasis id="B0E34B53306BFF82FCB3FDAEBE810F48" box="[838,994,571,595]" italics="true" pageId="18" pageNumber="219">T. abacetoides</emphasis>
</taxonomicName>
males; compare with Fig. 191e in Jeannel 1948)
</paragraph>
<caption id="D6E8C7C93068FF81FF6EF9B1BC2E0B72" ID-DOI="http://doi.org/10.5281/zenodo.13799463" ID-Zenodo-Dep="13799463" httpUri="https://zenodo.org/record/13799463/files/figure.png" pageId="17" pageNumber="218" startId="17.[155,167,1571,1590]" targetBox="[139,1176,235,1551]" targetPageId="17" targetType="figure">
<paragraph id="822897413068FF81FF6EF9B1BC2E0B72" blockId="17.[123,1173,1571,1641]" pageId="17" pageNumber="218">
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10. Digital images of holotype male of
<taxonomicName id="4597ECC23068FF81FDC7F9B1B92E0B2D" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[562,1101,1571,1590]" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov." subGenus="Mallopelmus">
<emphasis id="B0E34B533068FF81FDC7F9B1BFF70B2D" box="[562,660,1571,1590]" italics="true" pageId="17" pageNumber="218">Trichinillus</emphasis>
(
<emphasis id="B0E34B533068FF81FD57F9B1BE720B2D" box="[674,785,1571,1590]" italics="true" pageId="17" pageNumber="218">Mallopelmus</emphasis>
)
<emphasis id="B0E34B533068FF81FCEBF9B1BEF30B2D" box="[798,912,1571,1590]" italics="true" pageId="17" pageNumber="218">ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283068FF81FBA6F9B1B9F60B2D" box="[1107,1173,1571,1590]" pageId="17" pageNumber="218" rank="species">sp. nov.</taxonomicNameLabel>
A. Habitus, dorsal aspect; B. Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.5 mm.
</paragraph>
</caption>
<paragraph id="82289741306BFF82FF5EFDE9B9690F88" blockId="18.[123,1174,219,852]" box="[171,1034,635,660]" pageId="18" pageNumber="219">
<emphasis id="B0E34B53306BFF82FF5EFDE9BC920F8A" bold="true" box="[171,497,635,659]" pageId="18" pageNumber="219">
<smallCapsWord id="84CE019D306BFF82FF5EFDE9BC350F8A" baselines="653,652" box="[171,342,635,659]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Geographical" pageId="18" pageNumber="219">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="84CE019D306BFF82FEA8FDECBC920F8A" baselines="652" box="[349,497,638,657]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="18" pageNumber="219">DISTRIBUTION</smallCapsWord>
</emphasis>
.— At present, known only from the
<typeStatus id="5D2C29E3306BFF82FC8AFDEEBECE0F8F" box="[895,941,636,660]" pageId="18" pageNumber="219">type</typeStatus>
locality.
</paragraph>
<paragraph id="82289741306BFF82FF5EFD09BCD40E48" blockId="18.[123,1174,219,852]" pageId="18" pageNumber="219">
<emphasis id="B0E34B53306BFF82FF5EFD09BCC50FA9" bold="true" box="[171,422,667,691]" pageId="18" pageNumber="219">
<smallCapsWord id="84CE019D306BFF82FF5EFD09BC6A0FA9" baselines="685,685" box="[171,265,667,691]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Habitat" pageId="18" pageNumber="219">HABITAT</smallCapsWord>
<smallCapsWord id="84CE019D306BFF82FEE7FD0DBCC50FA9" baselines="685" box="[274,422,671,690]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="18" pageNumber="219">DISTRIBUTION</smallCapsWord>
</emphasis>
.— The male
<typeStatus id="5D2C29E3306BFF82FDCBFD09BFF80FA8" box="[574,667,667,691]" pageId="18" pageNumber="219" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4597ECC2306BFF82FD31FD0EBE0C0FA8" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[708,879,667,691]" class="Insecta" family="Carabidae" genus="Trichinillus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53306BFF82FD31FD0EBE0C0FA8" box="[708,879,667,691]" italics="true" pageId="18" pageNumber="219">T. ranomafanae</emphasis>
</taxonomicName>
was collected in degraded secondary montane rainforest at an elevation of
<quantity id="456F3AA4306BFF82FD65FD29BFB50FC8" box="[656,726,699,723]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="18" pageNumber="219" unit="m" value="800.0">800 m</quantity>
, near a village in the peripheral zone of Ranomafana National Park about1000 meters from the main gate. The female
<typeStatus id="5D2C29E3306BFF82FC20FD4EB9520FEF" box="[981,1073,732,756]" pageId="18" pageNumber="219" type="paratype">paratype</typeStatus>
was collected in secondary montane rainforest in the Talatakely area, at an elevation of
<quantity id="456F3AA4306BFF82FC2DFD69B97E0E08" box="[984,1053,763,787]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="18" pageNumber="219" unit="m" value="900.0">900 m</quantity>
, at the
<quantity id="456F3AA4306BFF82FB98FD69BDF30E28" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="219" unit="m" value="600.0">600 m</quantity>
mark along Trail B. Vegetation in this area was dominated by young trees (DBH &lt;
<quantity id="456F3AA4306BFF82FBEEFC89B93C0E28" box="[1051,1119,795,819]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="219" unit="cm" value="10.0">10 cm</quantity>
) and guava (
<taxonomicName id="4597ECC2306BFF82FF3EFCA9BCC40E48" box="[203,423,827,851]" class="Magnoliopsida" family="Myrtaceae" genus="Psidium" kingdom="Plantae" order="Myrtales" pageId="18" pageNumber="219" phylum="Tracheophyta" rank="species" species="cattleianum">
<emphasis id="B0E34B53306BFF82FF3EFCA9BCC40E48" box="[203,423,827,851]" italics="true" pageId="18" pageNumber="219">Psidium cattleianum</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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