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<document id="E1A52F3C7F4C84896FA334C95D755B4D" ID-DOI="10.5281/zenodo.13799439" ID-ISSN="0068-547X" ID-Zenodo-Dep="13799439" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538212676" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Rainio, Johanna" docDate="2016" docId="0A3E26573062FF8CFF8EFE4EBC080B0A" docLanguage="en" docName="ProcCalifAcadSci.63.7.201-268.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (7)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Perigona (Ripogena) ranomafanae Kavanaugh and Rainio 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="228" masterDocId="F6075E2F3079FF90FFF5FF92BD630D1B" masterDocTitle="Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar" masterLastPageNumber="268" masterPageNumber="201" pageNumber="228" updateTime="1726771873529" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="331BF332A9CB4BA02555011B839B43F4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A0C4A35251E0B97F71002F5014EC61B6">
<mods:title id="6A95786E3D7E58B1B7F9F8307FAACA31">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
</mods:titleInfo>
<mods:name id="CBA6BA2ECA6685FEDDCE27F27BA3996F" type="personal">
<mods:role id="0D5F36B58E4437A4E3ACEABAA7CECC6B">
<mods:roleTerm id="C6F2C030F8D9F3503DD812690A6F60F1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1B63B5B81C7249264C10717043B2E44E">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="07B10B653256E263550AF87E87FC967D">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
</mods:name>
<mods:name id="B9E13B7FC9FFB27480755F2A246F8BC7" type="personal">
<mods:role id="79000922D333218C8E2242036559C30C">
<mods:roleTerm id="DD5C02C0DC6618157456CAA33382FB1B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F77F4158EFE9AA83A24628A9A2A02DD2">Rainio, Johanna</mods:namePart>
<mods:affiliation id="8BBFF75A73D9C5615F545CB0F85EFC02">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
</mods:name>
<mods:typeOfResource id="B764007B51CFA1AAA29718C268B20131">text</mods:typeOfResource>
<mods:relatedItem id="2BB1C9F874C3F9C1C9A3C3C7E18DFAC7" type="host">
<mods:titleInfo id="AA5599165CA9EF93C8BCFFE7C500EA42">
<mods:title id="961F6BD120AA54C0C9E26DFC5E6EDE6E">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="39E56E05BA138DD1B352CA773471DBD7">
<mods:date id="DEFE4394DB5699D7B975EA55167970D7">2016</mods:date>
<mods:detail id="EBCF3648E7B2E003F361910BEA55DE05" type="pubDate">
<mods:number id="0A38AC72D95B44FFE44D928BCB255FEB">2016-04-29</mods:number>
</mods:detail>
<mods:detail id="C4E42B880548B22D0FBA36BF00711335" type="volume">
<mods:number id="21D6904DB4ECA908F91D5E26B15820D1">63</mods:number>
</mods:detail>
<mods:detail id="7FA814337F8FB54E634C7967FD1B5BAF" type="issue">
<mods:number id="62592E0F260590B43F0E00084F723E17">7</mods:number>
</mods:detail>
<mods:extent id="EBA75A4874F393407B70B4A75C6F40B6" unit="page">
<mods:start id="72485D205AED9B96D5E2AF522CE6D9F4">201</mods:start>
<mods:end id="0A829FBCF115F2D2DB23D8EB098F101E">268</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="D9F5DF03AE05CFB285893CB1E0696D7D">journal article</mods:classification>
<mods:identifier id="BFA3818A5C2D05344837574CF5045169" type="DOI">10.5281/zenodo.13799439</mods:identifier>
<mods:identifier id="306F9014B5F418E7E231551249BC0DE7" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="27B57A66BB3A759A077E034D3524FD6D" type="Zenodo-Dep">13799439</mods:identifier>
</mods:mods>
<treatment id="0A3E26573062FF8CFF8EFE4EBC080B0A" LSID="urn:lsid:plazi:treatment:0A3E26573062FF8CFF8EFE4EBC080B0A" httpUri="http://treatment.plazi.org/id/0A3E26573062FF8CFF8EFE4EBC080B0A" lastPageId="28" lastPageNumber="228" pageId="27" pageNumber="228">
<subSubSection id="CA8DC4CA3062FF8BFF8EFE4EBE110CEF" box="[123,882,476,500]" pageId="27" pageNumber="228" type="nomenclature">
<paragraph id="822897413062FF8BFF8EFE4EBE110CEF" blockId="27.[123,882,476,530]" box="[123,882,476,500]" pageId="27" pageNumber="228">
<heading id="D960202D3062FF8BFF8EFE4EBE110CEF" box="[123,882,476,500]" fontSize="10" level="2" pageId="27" pageNumber="228" reason="4">
<taxonomicName id="4597ECC23062FF8BFF8EFE4EBE6D0CEF" ID-CoL="775QY" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,782,476,500]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov." subGenus="Ripogena">
<emphasis id="B0E34B533062FF8BFF8EFE4EBC3A0CEF" bold="true" box="[123,345,476,500]" pageId="27" pageNumber="228">
<emphasis id="B0E34B533062FF8BFF8EFE4EBD830CEF" bold="true" box="[123,224,476,500]" italics="true" pageId="27" pageNumber="228">Perigona</emphasis>
(
<emphasis id="B0E34B533062FF8BFF05FE4EBC3A0CEF" bold="true" box="[240,345,476,500]" italics="true" pageId="27" pageNumber="228">Ripogena</emphasis>
</emphasis>
)
<emphasis id="B0E34B533062FF8BFE9CFE4EBE6D0CEF" bold="true" box="[361,782,476,500]" pageId="27" pageNumber="228">
<emphasis id="B0E34B533062FF8BFE9CFE4EBC9E0CEF" bold="true" box="[361,509,476,500]" italics="true" pageId="27" pageNumber="228">ranomafanae</emphasis>
Kavanaugh and Rainio
</emphasis>
</taxonomicName>
<emphasis id="B0E34B533062FF8BFCFBFE4EBE110CEF" bold="true" box="[782,882,476,500]" pageId="27" pageNumber="228">
,
<taxonomicNameLabel id="ABD0F6283062FF8BFCE9FE4EBE110CEF" box="[796,882,476,500]" pageId="27" pageNumber="228" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3062FF8BFF8EFE6EBEA90F46" pageId="27" pageNumber="228" type="description">
<paragraph id="822897413062FF8BFF8EFE6EBDBB0F09" blockId="27.[123,882,476,530]" box="[123,216,508,530]" pageId="27" pageNumber="228">
<figureCitation id="1AAC8BC43062FF8BFF8EFE6EBDBB0F09" box="[123,216,508,530]" captionStart="FIGURE" captionStartId="28.[155,167,1395,1414]" captionTargetBox="[148,1176,233,1374]" captionTargetId="figure-419@28.[120,1176,216,1392]" captionTargetPageId="28" captionText="FIGURE 15. Digital images of holotype male of Perigona (Ripogena) ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B Labels; C. Aedeagus of genitalia, left lateral aspect [dissection not fully cleaned due to teneral nature of holotype]; D. Dorsal aspect. Scale lines A = 1.0 mm, C = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13799475" httpUri="https://zenodo.org/record/13799475/files/figure.png" pageId="27" pageNumber="228">Figure 15</figureCitation>
</paragraph>
<paragraph id="822897413062FF8BFF8EFDBABFD20F26" blockId="27.[123,970,551,605]" box="[123,689,551,573]" pageId="27" pageNumber="228">
<emphasis id="B0E34B533062FF8BFF8EFDBABC470F27" box="[123,292,551,573]" italics="true" pageId="27" pageNumber="228">
<taxonomicName id="4597ECC23062FF8BFF8EFDBABDB40F26" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[123,215,552,573]" pageId="27" pageNumber="224" rank="subGenus" status="n. sp." subGenus="Ripogena">Ripogena</taxonomicName>
<taxonomicNameLabel id="ABD0F6283062FF8BFF28FDBABC720F26" box="[221,273,552,573]" pageId="27" pageNumber="228" rank="species">n. sp.</taxonomicNameLabel>
2
</emphasis>
; Rainio 2009: 33 (informal designation).
</paragraph>
<paragraph id="822897413062FF8BFF8EFDDABEA90F46" blockId="27.[123,970,551,605]" box="[123,970,583,605]" pageId="27" pageNumber="228">
<taxonomicName id="4597ECC23062FF8BFF8EFDDABC240F47" authorityName="Jeannel" authorityYear="1941" box="[123,327,583,605]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="subGenus" subGenus="Ripogena">
<emphasis id="B0E34B533062FF8BFF8EFDDABDB70F46" box="[123,212,584,605]" italics="true" pageId="27" pageNumber="228">Perigona</emphasis>
(
<emphasis id="B0E34B533062FF8BFF16FDDABC5C0F46" box="[227,319,584,605]" italics="true" pageId="27" pageNumber="228">Ripogena</emphasis>
)
</taxonomicName>
<emphasis id="B0E34B533062FF8BFEB8FDDABCF70F47" box="[333,404,583,605]" italics="true" pageId="27" pageNumber="228">n. sp. 2</emphasis>
; Rainio 2012: 73, Rainio 2013: 96 (informal designation).
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3062FF8CFF5EFD13BC080B0A" lastPageId="28" lastPageNumber="229" pageId="27" pageNumber="228" type="materials_examined">
<paragraph id="822897413062FF8BFF5EFD13BE190E02" blockId="27.[123,1174,640,1726]" pageId="27" pageNumber="228">
<emphasis id="B0E34B533062FF8BFF5EFD13BC3A0F8C" bold="true" box="[171,345,641,665]" pageId="27" pageNumber="228">
<smallCapsWord id="84CE019D3062FF8BFF5EFD13BD870F8C" baselines="659,658" box="[171,228,641,665]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="27" pageNumber="228">TYPE</smallCapsWord>
<smallCapsWord id="84CE019D3062FF8BFF19FD16BC3A0F8C" baselines="658" box="[236,345,644,663]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="27" pageNumber="228">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C3062FF8BFE74FD12BE190E02" collectingDate="1998-04-29" collectorName="D. H. Kavanaugh" country="Madagascar" county="Ranomafana National Park" elevation="1050" latitude="-21.278334" location="Abotovory" longLatPrecision="126" longitude="47.435" municipality="Vatoharanana area" pageId="27" pageNumber="228" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="5D2C29E33062FF8BFE74FD12BC870F83" box="[385,484,640,664]" pageId="27" pageNumber="228" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC43062FF8BFE00FD12BF150F83" box="[501,630,640,665]" captionStart="FIGURE" captionStartId="28.[155,167,1395,1414]" captionTargetBox="[148,1176,233,1374]" captionTargetId="figure-419@28.[120,1176,216,1392]" captionTargetPageId="28" captionText="FIGURE 15. Digital images of holotype male of Perigona (Ripogena) ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B Labels; C. Aedeagus of genitalia, left lateral aspect [dissection not fully cleaned due to teneral nature of holotype]; D. Dorsal aspect. Scale lines A = 1.0 mm, C = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13799475" httpUri="https://zenodo.org/record/13799475/files/figure.png" pageId="27" pageNumber="228">Figs.15A–B</figureCitation>
), a male, in CAS, labeled: ““CASENT 1049017”/ “
<collectingCountry id="FA80D7D13062FF8BFF72FD33BC260FA2" box="[135,325,673,697]" name="Madagascar" pageId="27" pageNumber="228">MADAGASCAR</collectingCountry>
,
<collectingRegion id="405359A33062FF8BFEA1FD33BF270FA2" box="[340,580,673,697]" country="Madagascar" name="Fianarantsoa" pageId="27" pageNumber="228">Fianarantsoa Province</collectingRegion>
,
<collectingCounty id="6B49EFCD3062FF8BFDA1FD33BE190FA2" box="[596,890,673,697]" pageId="27" pageNumber="228">Ranomafana National Park</collectingCounty>
,
<collectingMunicipality id="624C0D3B3062FF8BFC7CFD33B9330FA3" box="[905,1104,673,697]" pageId="27" pageNumber="228">Vatoharanana area</collectingMunicipality>
,
<quantity id="456F3AA43062FF8BFB95FD32BDF30FC3" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="27" pageNumber="228" unit="m" value="1050.0">
<elevation id="09BA70723062FF8BFB95FD32BDF30FC3" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="27" pageNumber="228" unit="m" value="1050.0">1050 m</elevation>
</quantity>
,
<location id="8748C19A3062FF8BFF6EFD53BC6C0FC2" LSID="urn:lsid:plazi:treatment:0A3E26573062FF8CFF8EFE4EBC080B0A:8748C19A3062FF8BFF6EFD53BC6C0FC2" box="[155,271,705,729]" country="Madagascar" county="Ranomafana National Park" latitude="-21.278334" longLatPrecision="126" longitude="47.435" municipality="Vatoharanana area" name="Abotovory" pageId="27" pageNumber="228" stateProvince="Fianarantsoa">Abotovory</location>
stream,
<date id="F629B1813062FF8BFE9DFD53BC990FC2" box="[360,506,705,729]" pageId="27" pageNumber="228" value="1998-04-29">
<collectingDate id="E66D48693062FF8BFE9DFD53BC990FC2" box="[360,506,705,729]" pageId="27" pageNumber="228" value="1998-04-29">29 April 1998</collectingDate>
</date>
”/ “
<geoCoordinate id="E7A3F1863062FF8BFDEAFD53BFE80FC2" box="[543,651,705,729]" degrees="21" direction="south" minutes="16. 7" orientation="latitude" pageId="27" pageNumber="228" precision="92" value="-21.278334">21°16.7’S</geoCoordinate>
<geoCoordinate id="E7A3F1863062FF8BFD64FD53BE670FC3" box="[657,772,705,729]" degrees="47" direction="east" minutes="26. 1" orientation="longitude" pageId="27" pageNumber="228" precision="92" value="47.435">47° 26.1’E</geoCoordinate>
, Stop # 98-100,
<collectorName id="2F62F2973062FF8BFC46FD53B9050FC2" box="[947,1126,705,729]" pageId="27" pageNumber="228">D.H. Kavanaugh</collectorName>
collector collected in large rotting log just under loose bark with cavities”/ “
<typeStatus id="5D2C29E33062FF8BFC68FD73B9490FE2" box="[925,1066,737,761]" pageId="27" pageNumber="228" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC23062FF8BFBC7FD70BF100E02" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov." subGenus="Ripogena">
<emphasis id="B0E34B533062FF8BFBC7FD70B9F60FE2" box="[1074,1173,738,761]" italics="true" pageId="27" pageNumber="228">Perigona</emphasis>
(
<emphasis id="B0E34B533062FF8BFF71FC90BD890E02" box="[132,234,770,793]" italics="true" pageId="27" pageNumber="228">Ripogena</emphasis>
)
<emphasis id="B0E34B533062FF8BFF0FFC93BCEB0E02" box="[250,392,769,793]" italics="true" pageId="27" pageNumber="228">ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283062FF8BFD8EFC90BFD60E02" box="[635,693,770,793]" pageId="27" pageNumber="228" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label].
</materialsCitation>
</paragraph>
<paragraph id="822897413062FF8BFF5EFCB3B9500E22" blockId="27.[123,1174,640,1726]" box="[171,1075,801,825]" pageId="27" pageNumber="228">
<materialsCitation id="32FF9D1C3062FF8BFF5EFCB3B94F0E22" box="[171,1068,801,825]" country="Madagascar" location="Ranomafana National Park" pageId="27" pageNumber="228" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B533062FF8BFF5EFCB3BC340E23" bold="true" box="[171,343,801,825]" pageId="27" pageNumber="228">
<smallCapsWord id="84CE019D3062FF8BFF5EFCB3BD870E23" baselines="819,819" box="[171,228,801,825]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="27" pageNumber="228">
<typeStatus id="5D2C29E33062FF8BFF5EFCB3BD870E23" box="[171,228,801,825]" pageId="27" pageNumber="228">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D3062FF8BFF19FCB7BC340E23" baselines="819" box="[236,343,805,824]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="27" pageNumber="228">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D13062FF8BFE8BFCB3BC9C0E22" box="[382,511,801,825]" name="Madagascar" pageId="27" pageNumber="228">Madagascar</collectingCountry>
,
<collectingRegion id="405359A33062FF8BFDF8FCB3BF9F0E22" box="[525,764,801,825]" country="Madagascar" name="Fianarantsoa" pageId="27" pageNumber="228">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A3062FF8BFCFFFCB3B94F0E22" LSID="urn:lsid:plazi:treatment:0A3E26573062FF8CFF8EFE4EBC080B0A:8748C19A3062FF8BFCFFFCB3B94F0E22" box="[778,1068,801,825]" country="Madagascar" name="Ranomafana National Park" pageId="27" pageNumber="228" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="822897413062FF8BFF5EFCD0BECA0E62" blockId="27.[123,1174,640,1726]" pageId="27" pageNumber="228">
<emphasis id="B0E34B533062FF8BFF5EFCD0BC910E43" bold="true" box="[171,498,834,858]" pageId="27" pageNumber="228">
<smallCapsWord id="84CE019D3062FF8BFF5EFCD0BC4D0E43" baselines="852,851" box="[171,302,834,858]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="27" pageNumber="228">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D3062FF8BFEC3FCD7BC310E43" baselines="851" box="[310,338,837,856]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="27" pageNumber="228">OF</smallCapsWord>
<smallCapsWord id="84CE019D3062FF8BFEACFCD7BCCF0E43" baselines="851" box="[345,428,837,856]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="27" pageNumber="228">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D3062FF8BFE40FCD7BC910E43" baselines="851" box="[437,498,837,856]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="27" pageNumber="228">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC23062FF8BFD0CFCD3BEE40E42" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[761,903,833,857]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533062FF8BFD0CFCD3BEE40E42" box="[761,903,833,857]" italics="true" pageId="27" pageNumber="228">ranomafanae</emphasis>
</taxonomicName>
, is a noun in apposition, derived from the name of the national park in which the
<typeStatus id="5D2C29E33062FF8BFD29FCF0BE690E61" box="[732,778,866,890]" pageId="27" pageNumber="228">type</typeStatus>
was collected.
</paragraph>
<paragraph id="822897413062FF8BFF5EFC10BFD50B66" blockId="27.[123,1174,640,1726]" pageId="27" pageNumber="228">
<smallCapsWord id="84CE019D3062FF8BFF5EFC10BC210E83" baselines="916,915" box="[171,322,898,922]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="27" pageNumber="228">
<emphasis id="B0E34B533062FF8BFF5EFC10BC210E83" bold="true" box="[171,322,898,922]" pageId="27" pageNumber="228">RECOGNITION</emphasis>
</smallCapsWord>
.— Size larger than average for subgenus, SBL =
<quantity id="456F3AA43062FF8BFCA6FC10BEC70E82" box="[851,932,898,922]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="27" pageNumber="228" unit="mm" value="4.4">4.4 mm</quantity>
. The
<typeStatus id="5D2C29E33062FF8BFC14FC10B95D0E81" box="[993,1086,898,922]" pageId="27" pageNumber="228" type="holotype">holotype</typeStatus>
male of this species (
<figureCitation id="1AAC8BC43062FF8BFEFEFC30BC0F0EA1" box="[267,364,930,954]" captionStart="FIGURE" captionStartId="28.[155,167,1395,1414]" captionTargetBox="[148,1176,233,1374]" captionTargetId="figure-419@28.[120,1176,216,1392]" captionTargetPageId="28" captionText="FIGURE 15. Digital images of holotype male of Perigona (Ripogena) ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B Labels; C. Aedeagus of genitalia, left lateral aspect [dissection not fully cleaned due to teneral nature of holotype]; D. Dorsal aspect. Scale lines A = 1.0 mm, C = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13799475" httpUri="https://zenodo.org/record/13799475/files/figure.png" pageId="27" pageNumber="228">Fig. 15A</figureCitation>
) shares with members of
<taxonomicName id="4597ECC23062FF8BFD65FC31B9410EA1" authority="Alluaud (1936)" authorityName="Alluaud" authorityYear="1936" box="[656,1058,930,954]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="heterodera">
<emphasis id="B0E34B533062FF8BFD65FC31BE120EA1" box="[656,881,930,954]" italics="true" pageId="27" pageNumber="228">Perigona heterodera</emphasis>
Alluaud (1936)
</taxonomicName>
,
<taxonomicName id="4597ECC23062FF8BFBC7FC31BCE20EC1" authority="Alluaud (1936)" authorityName="Alluaud" authorityYear="1936" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="prasinus">
<emphasis id="B0E34B533062FF8BFBC7FC31BDB40EC1" italics="true" pageId="27" pageNumber="228">Perigona prasinus</emphasis>
Alluaud (1936)
</taxonomicName>
and
<taxonomicName id="4597ECC23062FF8BFE43FC51BF740EC1" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[438,535,962,986]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="deuvei" status="sp. nov.">
<emphasis id="B0E34B533062FF8BFE43FC51BF740EC1" box="[438,535,962,986]" italics="true" pageId="27" pageNumber="228">P. deuvei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283062FF8BFDEBFC51BF120EC1" box="[542,625,963,986]" pageId="27" pageNumber="228" rank="species">sp. nov.</taxonomicNameLabel>
the following features: pronotum with lateral margins without sinuation anterior to hind angles, posterior margin slightly and smoothly convex, and hind angles either obtuse or broadly rounded. These features distinguish members of these four species from those of
<taxonomicName id="4597ECC23062FF8BFE98FBB1BE770920" authority="Alluaud (1936)" authorityName="Alluaud" authorityYear="1936" box="[365,788,1058,1083]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="bembidioides">
<emphasis id="B0E34B533062FF8BFE98FBB1BF040921" box="[365,615,1058,1082]" italics="true" pageId="27" pageNumber="228">Perigona bembidioides</emphasis>
Alluaud (1936)
</taxonomicName>
and
<taxonomicName id="4597ECC23062FF8BFCBEFBB1BDA10940" authority="Jeannel (1948)" authorityName="Jeannel" authorityYear="1948" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="viridimicans">
<emphasis id="B0E34B533062FF8BFCBEFBB1B95F0921" box="[843,1084,1058,1082]" italics="true" pageId="27" pageNumber="228">Perigona viridimicans</emphasis>
Jeannel (1948)
</taxonomicName>
, which have distinct sinuation of the lateral margins anterior to rectangular hind angles and a straight basal margin. Members of
<taxonomicName id="4597ECC23062FF8BFDE6FBF6BFDC0960" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[531,703,1123,1147]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533062FF8BFDE6FBF6BFDC0960" box="[531,703,1123,1147]" italics="true" pageId="27" pageNumber="228">P. ranomafanae</emphasis>
</taxonomicName>
differ from those of
<taxonomicName id="4597ECC23062FF8BFC59FBF6B9460961" authorityName="Alluaud" authorityYear="1936" box="[940,1061,1123,1147]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="prasinus">
<emphasis id="B0E34B533062FF8BFC59FBF6B9460961" box="[940,1061,1123,1147]" italics="true" pageId="27" pageNumber="228">P. prasinus</emphasis>
</taxonomicName>
in having slightly smaller body size and dorsum without metallic reflection (dorsum with metallic green reflection in
<taxonomicName id="4597ECC23062FF8BFEF6FB36BC1909A0" authorityName="Alluaud" authorityYear="1936" box="[259,378,1188,1211]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="prasinus">
<emphasis id="B0E34B533062FF8BFEF6FB36BC1909A0" box="[259,378,1188,1211]" italics="true" pageId="27" pageNumber="228">P. prasinus</emphasis>
</taxonomicName>
). They differ from members of
<taxonomicName id="4597ECC23062FF8BFD3AFB36BE3E09A0" authorityName="Alluaud" authorityYear="1936" box="[719,861,1187,1211]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="heterodera">
<emphasis id="B0E34B533062FF8BFD3AFB36BE3E09A0" box="[719,861,1187,1211]" italics="true" pageId="27" pageNumber="228">P. heterodera</emphasis>
</taxonomicName>
and
<taxonomicName id="4597ECC23062FF8BFC67FB36BE9109A0" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[914,1010,1187,1211]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="deuvei">
<emphasis id="B0E34B533062FF8BFC67FB36BE9109A0" box="[914,1010,1187,1211]" italics="true" pageId="27" pageNumber="228">P. deuvei</emphasis>
</taxonomicName>
in having larg- er body size (SBL =
<quantity id="456F3AA43062FF8BFEAAFB51BCD009C0" box="[351,435,1219,1243]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="27" pageNumber="228" unit="mm" value="3.4">3.4 mm</quantity>
in
<taxonomicName id="4597ECC23062FF8BFE2FFB56BF0909C0" authorityName="Alluaud" authorityYear="1936" box="[474,618,1219,1243]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="heterodera">
<emphasis id="B0E34B533062FF8BFE2FFB56BF0909C0" box="[474,618,1219,1243]" italics="true" pageId="27" pageNumber="228">P. heterodera</emphasis>
</taxonomicName>
, 3.5 to
<quantity id="456F3AA43062FF8BFD37FB51BE7609C0" box="[706,789,1219,1243]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="27" pageNumber="228" unit="mm" value="3.9">3.9 mm</quantity>
in
<taxonomicName id="4597ECC23062FF8BFCC9FB56BEFC09C0" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[828,927,1219,1243]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="deuvei">
<emphasis id="B0E34B533062FF8BFCC9FB56BEFC09C0" box="[828,927,1219,1243]" italics="true" pageId="27" pageNumber="228">P. deuvei</emphasis>
</taxonomicName>
), pronotal hind angles more sharply angulate, lateral explanation of pronotum markedly widened and flattened in region of hind angles (pronotal hind angles rounded or at least less sharply angulate and lateral explanation only slightly wider near hind angles than anteriorly in
<taxonomicName id="4597ECC23062FF8BFD1CFAB6BE150827" authorityName="Alluaud" authorityYear="1936" box="[745,886,1316,1340]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="heterodera">
<emphasis id="B0E34B533062FF8BFD1CFAB6BE150827" box="[745,886,1316,1340]" italics="true" pageId="27" pageNumber="228">P. heterodera</emphasis>
</taxonomicName>
and
<taxonomicName id="4597ECC23062FF8BFC5CFAB6B96B0827" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[937,1032,1316,1340]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="deuvei">
<emphasis id="B0E34B533062FF8BFC5CFAB6B96B0827" box="[937,1032,1316,1340]" italics="true" pageId="27" pageNumber="228">P. deuvei</emphasis>
</taxonomicName>
). The unique male
<typeStatus id="5D2C29E33062FF8BFF43FAD6BC700847" box="[182,275,1348,1372]" pageId="27" pageNumber="228" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4597ECC23062FF8BFEC2FAD7BC830847" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[311,480,1348,1372]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533062FF8BFEC2FAD7BC830847" box="[311,480,1348,1372]" italics="true" pageId="27" pageNumber="228">P. ranomafanae</emphasis>
</taxonomicName>
is slightly teneral and the genitalia are very soft and only lightly sclerotized. However, the apex is sclerotized sufficiently to allow comparisons with those of
<taxonomicName id="4597ECC23062FF8BFBA7FAF7BDAD0887" authorityName="Alluaud" authorityYear="1936" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="heterodera">
<emphasis id="B0E34B533062FF8BFBA7FAF7BDAD0887" italics="true" pageId="27" pageNumber="228">P. heterodera</emphasis>
</taxonomicName>
and
<taxonomicName id="4597ECC23062FF8BFEF1FA17BC050887" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[260,358,1412,1436]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="deuvei">
<emphasis id="B0E34B533062FF8BFEF1FA17BC050887" box="[260,358,1412,1436]" italics="true" pageId="27" pageNumber="228">P. deuvei</emphasis>
</taxonomicName>
males. In lateral view (
<figureCitation id="1AAC8BC43062FF8BFD92FA16BFA50887" box="[615,710,1412,1436]" captionStart="FIGURE" captionStartId="28.[155,167,1395,1414]" captionTargetBox="[148,1176,233,1374]" captionTargetId="figure-419@28.[120,1176,216,1392]" captionTargetPageId="28" captionText="FIGURE 15. Digital images of holotype male of Perigona (Ripogena) ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B Labels; C. Aedeagus of genitalia, left lateral aspect [dissection not fully cleaned due to teneral nature of holotype]; D. Dorsal aspect. Scale lines A = 1.0 mm, C = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13799475" httpUri="https://zenodo.org/record/13799475/files/figure.png" pageId="27" pageNumber="228">Fig. 15C</figureCitation>
), the apex is slightly longer than in
<taxonomicName id="4597ECC23062FF8BFBA5FA17BDAD08A7" authorityName="Alluaud" authorityYear="1936" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="heterodera">
<emphasis id="B0E34B533062FF8BFBA5FA17BDAD08A7" italics="true" pageId="27" pageNumber="228">P. heterodera</emphasis>
</taxonomicName>
males (see Jeannel 1948, Fig. 351b) and distinctly shorter and straighter than in
<taxonomicName id="4597ECC23062FF8BFBC0FA37B9F508A7" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[1077,1174,1444,1468]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="deuvei">
<emphasis id="B0E34B533062FF8BFBC0FA37B9F508A7" box="[1077,1174,1444,1468]" italics="true" pageId="27" pageNumber="228">P. deuvei</emphasis>
</taxonomicName>
males. Finally, they differ from members of
<taxonomicName id="4597ECC23062FF8BFD9CFA57BE4908C7" baseAuthorityName="Deuve" baseAuthorityYear="1998" box="[617,810,1476,1500]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="descarpentriesi">
<emphasis id="B0E34B533062FF8BFD9CFA57BE4908C7" box="[617,810,1476,1500]" italics="true" pageId="27" pageNumber="228">P. descarpentriesi</emphasis>
</taxonomicName>
in having moderately large eyes (eyes markedly reduced eyes in size in
<taxonomicName id="4597ECC23062FF8BFDEEFA77BFBA08E6" baseAuthorityName="Deuve" baseAuthorityYear="1998" box="[539,729,1509,1533]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="descarpentriesi">
<emphasis id="B0E34B533062FF8BFDEEFA77BFBA08E6" box="[539,729,1509,1533]" italics="true" pageId="27" pageNumber="228">P. descarpentriesi</emphasis>
</taxonomicName>
), pronotum widest anterior to middle and lateral margins nearly straight in posterior half (pronotum widest at middle and lateral margins evenly arcuate throughout
<taxonomicName id="4597ECC23062FF8BFE6CF9B4BF3B0B26" baseAuthorityName="Deuve" baseAuthorityYear="1998" box="[409,600,1573,1597]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="descarpentriesi">
<emphasis id="B0E34B533062FF8BFE6CF9B4BF3B0B26" box="[409,600,1573,1597]" italics="true" pageId="27" pageNumber="228">P. descarpentriesi</emphasis>
</taxonomicName>
). Also, the apex of the median lobe of the male genitalia is narrower and more pointed in lateral aspect in
<taxonomicName id="4597ECC23062FF8BFD26F9D4BE1D0B46" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[723,894,1605,1629]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533062FF8BFD26F9D4BE1D0B46" box="[723,894,1605,1629]" italics="true" pageId="27" pageNumber="228">P. ranomafanae</emphasis>
</taxonomicName>
males (
<figureCitation id="1AAC8BC43062FF8BFC22F9D7B95B0B46" box="[983,1080,1605,1629]" captionStart="FIGURE" captionStartId="28.[155,167,1395,1414]" captionTargetBox="[148,1176,233,1374]" captionTargetId="figure-419@28.[120,1176,216,1392]" captionTargetPageId="28" captionText="FIGURE 15. Digital images of holotype male of Perigona (Ripogena) ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B Labels; C. Aedeagus of genitalia, left lateral aspect [dissection not fully cleaned due to teneral nature of holotype]; D. Dorsal aspect. Scale lines A = 1.0 mm, C = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13799475" httpUri="https://zenodo.org/record/13799475/files/figure.png" pageId="27" pageNumber="228">Fig. 15C</figureCitation>
) than in those of
<taxonomicName id="4597ECC23062FF8BFF2DF9F4BCF40B66" baseAuthorityName="Deuve" baseAuthorityYear="1998" box="[216,407,1637,1661]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="descarpentriesi">
<emphasis id="B0E34B533062FF8BFF2DF9F4BCF40B66" box="[216,407,1637,1661]" italics="true" pageId="27" pageNumber="228">P. descarpentriesi</emphasis>
</taxonomicName>
(see
<bibRefCitation id="E606EAB03062FF8BFE24F9F4BF300B66" author="DEUVE, T." box="[465,595,1637,1661]" pageId="27" pageNumber="228" pagination="2 - 4" refId="ref30013" refString="DEUVE, T. 1998. Description d'un Perigoninae microphtalme de Madagascar [Coleoptera, Harpalidea]. Revue Francaise d'Entomologie (N. S.) 20: 2 - 4." type="journal article" year="1998">Deuve 1998</bibRefCitation>
,
<figureCitation id="1AAC8BC43062FF8BFD94F9F7BFC30B66" box="[609,672,1637,1661]" captionStart="FIGURE" captionStartId="5.[155,167,1152,1171]" captionTargetBox="[133,1174,245,1135]" captionTargetId="figure-997@5.[120,1174,216,1150]" captionTargetPageId="5" captionText="FIGURE 2. Digital images of holotype female of Brachypelus ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799443" httpUri="https://zenodo.org/record/13799443/files/figure.png" pageId="27" pageNumber="228">Fig. 2</figureCitation>
.).
</paragraph>
<paragraph id="822897413062FF8BFF5EF917B9690B86" blockId="27.[123,1174,640,1726]" box="[171,1034,1669,1694]" pageId="27" pageNumber="228">
<emphasis id="B0E34B533062FF8BFF5EF917BC920B87" bold="true" box="[171,497,1669,1693]" pageId="27" pageNumber="228">
<smallCapsWord id="84CE019D3062FF8BFF5EF917BC350B87" baselines="1687,1687" box="[171,342,1669,1693]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Geographical" pageId="27" pageNumber="228">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="84CE019D3062FF8BFEA8F91BBC920B87" baselines="1687" box="[349,497,1673,1692]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="27" pageNumber="228">DISTRIBUTION</smallCapsWord>
</emphasis>
.— At present, known only from the
<typeStatus id="5D2C29E33062FF8BFC8AF914BECE0B85" box="[895,941,1670,1694]" pageId="27" pageNumber="228">type</typeStatus>
locality.
</paragraph>
<paragraph id="822897413062FF8CFF5EF934BC080B0A" blockId="27.[123,1174,640,1726]" lastBlockId="28.[123,1174,1497,1553]" lastPageId="28" lastPageNumber="229" pageId="27" pageNumber="228">
<emphasis id="B0E34B533062FF8BFF5EF934BCC40BA7" bold="true" box="[171,423,1702,1726]" pageId="27" pageNumber="228">
<smallCapsWord id="84CE019D3062FF8BFF5EF934BC6A0BA7" baselines="1720,1719" box="[171,265,1702,1726]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Habitat" pageId="27" pageNumber="228">HABITAT</smallCapsWord>
<smallCapsWord id="84CE019D3062FF8BFEE6F93BBCC40BA7" baselines="1719" box="[275,423,1705,1724]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="27" pageNumber="228">DISTRIBUTION</smallCapsWord>
</emphasis>
.— The unique
<typeStatus id="5D2C29E33062FF8BFDA0F937BFD10BA6" box="[597,690,1701,1725]" pageId="27" pageNumber="228" type="holotype">holotype</typeStatus>
was collected by hand in primary montane rainforest in the Vatoharanana area at an elevation of
<quantity id="456F3AA43065FF8CFD4FFA4BBE6808EB" box="[698,779,1497,1521]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="28" pageNumber="229" unit="m" value="1050.0">1050 m</quantity>
from under the loose bark of a large log on the forest floor.
</paragraph>
</subSubSection>
</treatment>
</document>
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