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<document id="AA1689BBC8E78D260C0E59D30E80BB1D" ID-DOI="10.5281/zenodo.13799439" ID-ISSN="0068-547X" ID-Zenodo-Dep="13799439" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538212676" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Rainio, Johanna" docDate="2016" docId="0A3E26573060FF8BFF8EFF49BEB20CA8" docLanguage="en" docName="ProcCalifAcadSci.63.7.201-268.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (7)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Perigona (Ripogena) deuvei Kavanaugh and Rainio 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="228" masterDocId="F6075E2F3079FF90FFF5FF92BD630D1B" masterDocTitle="Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar" masterLastPageNumber="268" masterPageNumber="201" pageNumber="226" updateTime="1726771873529" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="296AAE7ECB713CE3010A1DAFF5CD669A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5EC68CC7E26CDCFDC7324168D235866A">
<mods:title id="03D903BCC59E7E95F4FFF253479B80B1">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
</mods:titleInfo>
<mods:name id="458AF69656C396C42F8C77F050570E96" type="personal">
<mods:role id="9B0959156639E56A28B079162C2B779E">
<mods:roleTerm id="C602C6AAB31938D7038F7A6A57BF40A8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CF7EB2114E5246DF61D57E0353ED3E35">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="FC34716A21989435B70D065BF1487DF2">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
</mods:name>
<mods:name id="826828FFDC52EB1302AB20F8299B47B9" type="personal">
<mods:role id="4286DD4EF1D3328CBCC887624E618A74">
<mods:roleTerm id="E31EC30DA576901394A55753786C659F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="77F53A2A5A8B9D0B9B775D3AB22C2448">Rainio, Johanna</mods:namePart>
<mods:affiliation id="4B47BF0C7F162271990C6E7F4B63EADD">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
</mods:name>
<mods:typeOfResource id="7776839B71C93CF89347DB18222E8D14">text</mods:typeOfResource>
<mods:relatedItem id="BEBB4FC9B4CD041C0496906872A77DC1" type="host">
<mods:titleInfo id="7597BF9F9840CF0ED8C52172EB998E79">
<mods:title id="641AF31B4869FA7D0AEC8AD45BFA413D">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="39464A4CD517A298AA4433254A2BC292">
<mods:date id="8B10D68958B41CD61F2C5A992B3A3ECB">2016</mods:date>
<mods:detail id="64D8BFAADEF11D55C4B2E707BD982967" type="pubDate">
<mods:number id="1555FA9A0C145BE84E553BC1400139C1">2016-04-29</mods:number>
</mods:detail>
<mods:detail id="444054B6CF5C3D1EFCE796D3DA935474" type="volume">
<mods:number id="575A05C59BF97BDB58B33F8C954B5539">63</mods:number>
</mods:detail>
<mods:detail id="D576B084BD8810520261AC81A461B126" type="issue">
<mods:number id="C5B2F8095E21996C2F2FA955A83E3576">7</mods:number>
</mods:detail>
<mods:extent id="00169686BD555E0290786F00BC09BB68" unit="page">
<mods:start id="04253025E8B5935BB9A69360D2FF85C8">201</mods:start>
<mods:end id="3134F2FFFFBBF6E79308AF4FF213212C">268</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="2B80737AAE32F1220B259C8548980C02">journal article</mods:classification>
<mods:identifier id="D0621A87117868150388370107D91FFA" type="DOI">10.5281/zenodo.13799439</mods:identifier>
<mods:identifier id="5B14E987AE2B1FEA691E9FC81EF9B264" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="42E84DCC75250878839C1924AD72A586" type="Zenodo-Dep">13799439</mods:identifier>
</mods:mods>
<treatment id="0A3E26573060FF8BFF8EFF49BEB20CA8" LSID="urn:lsid:plazi:treatment:0A3E26573060FF8BFF8EFF49BEB20CA8" httpUri="http://treatment.plazi.org/id/0A3E26573060FF8BFF8EFF49BEB20CA8" lastPageId="27" lastPageNumber="228" pageId="25" pageNumber="226">
<subSubSection id="CA8DC4CA3060FF89FF8EFF49BE460DE8" box="[123,805,219,243]" pageId="25" pageNumber="226" type="nomenclature">
<paragraph id="822897413060FF89FF8EFF49BE460DE8" blockId="25.[123,805,219,273]" box="[123,805,219,243]" pageId="25" pageNumber="226">
<heading id="D960202D3060FF89FF8EFF49BE460DE8" box="[123,805,219,243]" fontSize="10" level="2" pageId="25" pageNumber="226" reason="4">
<taxonomicName id="4597ECC23060FF89FF8EFF49BFA20DE8" ID-CoL="775R9" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,705,219,243]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="deuvei" status="sp. nov." subGenus="Ripogena">
<emphasis id="B0E34B533060FF89FF8EFF49BC3A0DE8" bold="true" box="[123,345,219,243]" pageId="25" pageNumber="226">
<emphasis id="B0E34B533060FF89FF8EFF49BD830DE8" bold="true" box="[123,224,219,243]" italics="true" pageId="25" pageNumber="226">Perigona</emphasis>
(
<emphasis id="B0E34B533060FF89FF05FF49BC3A0DE8" bold="true" box="[240,345,219,243]" italics="true" pageId="25" pageNumber="226">Ripogena</emphasis>
</emphasis>
)
<emphasis id="B0E34B533060FF89FE9CFF49BFA20DE8" bold="true" box="[361,705,219,243]" pageId="25" pageNumber="226">
<emphasis id="B0E34B533060FF89FE9CFF49BCD30DE8" bold="true" box="[361,432,219,243]" italics="true" pageId="25" pageNumber="226">deuvei</emphasis>
Kavanaugh and Rainio
</emphasis>
</taxonomicName>
<emphasis id="B0E34B533060FF89FD34FF49BE460DE8" bold="true" box="[705,805,219,243]" pageId="25" pageNumber="226">
,
<taxonomicNameLabel id="ABD0F6283060FF89FD3AFF49BE460DE8" box="[719,805,219,243]" pageId="25" pageNumber="226" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3060FF89FF8EFF69BEA90C47" pageId="25" pageNumber="226" type="description">
<paragraph id="822897413060FF89FF8EFF69BDBB0C0B" blockId="25.[123,805,219,273]" box="[123,216,251,273]" pageId="25" pageNumber="226">
<figureCitation id="1AAC8BC43060FF89FF8EFF69BDBB0C0B" box="[123,216,251,273]" captionStart="FIGURE" captionStartId="26.[155,167,1478,1497]" captionTargetBox="[139,1176,216,1455]" captionTargetId="figure-53@26.[120,1176,216,1455]" captionTargetPageId="26" captionText="FIGURE 14. Digital images of holotype male of Perigona (Ripogena) deuvei Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13799473" httpUri="https://zenodo.org/record/13799473/files/figure.png" pageId="25" pageNumber="226">Figure 14</figureCitation>
</paragraph>
<paragraph id="822897413060FF89FF8EFEB5BFD20C20" blockId="25.[123,970,294,348]" box="[123,689,294,316]" pageId="25" pageNumber="226">
<emphasis id="B0E34B533060FF89FF8EFEB5BC470C20" box="[123,292,294,316]" italics="true" pageId="25" pageNumber="226">
<taxonomicName id="4597ECC23060FF89FF8EFEB5BDB40C27" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[123,215,295,316]" pageId="25" pageNumber="224" rank="subGenus" status="n. sp." subGenus="Ripogena">Ripogena</taxonomicName>
<taxonomicNameLabel id="ABD0F6283060FF89FF28FEB5BC720C27" box="[221,273,295,316]" pageId="25" pageNumber="226" rank="species">n. sp.</taxonomicNameLabel>
1
</emphasis>
; Rainio 2009: 33 (informal designation).
</paragraph>
<paragraph id="822897413060FF89FF8EFED5BEA90C47" blockId="25.[123,970,294,348]" box="[123,970,326,348]" pageId="25" pageNumber="226">
<taxonomicName id="4597ECC23060FF89FF8EFED5BC240C40" authorityName="Jeannel" authorityYear="1941" box="[123,327,326,348]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="subGenus" subGenus="Ripogena">
<emphasis id="B0E34B533060FF89FF8EFED5BDB70C47" box="[123,212,327,348]" italics="true" pageId="25" pageNumber="226">Perigona</emphasis>
(
<emphasis id="B0E34B533060FF89FF16FED5BC5C0C47" box="[227,319,327,348]" italics="true" pageId="25" pageNumber="226">Ripogena</emphasis>
)
</taxonomicName>
<emphasis id="B0E34B533060FF89FEB8FED5BCF70C40" box="[333,404,326,348]" italics="true" pageId="25" pageNumber="226">n. sp. 1</emphasis>
; Rainio 2012: 73, Rainio 2013: 96 (informal designation).
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3060FF8BFF5EFEEDBEB20CA8" lastPageId="27" lastPageNumber="228" pageId="25" pageNumber="226" type="materials_examined">
<paragraph id="822897413060FF89FF5EFEEDBE3B0E22" blockId="25.[123,1174,383,1726]" pageId="25" pageNumber="226">
<emphasis id="B0E34B533060FF89FF5EFEEDBC3D0C8D" bold="true" box="[171,350,383,407]" pageId="25" pageNumber="226">
<smallCapsWord id="84CE019D3060FF89FF5EFEEDBD870C8D" baselines="401,401" box="[171,228,383,407]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="25" pageNumber="226">TYPE</smallCapsWord>
<smallCapsWord id="84CE019D3060FF89FF04FE11BC3D0C8D" baselines="401" box="[241,350,387,406]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="25" pageNumber="226">MATERIAL</smallCapsWord>
</emphasis>
.—
<materialsCitation id="32FF9D1C3060FF89FE73FEEDBCA30CEC" collectingDate="2005-07-24" collectorName="F. Ratalata &amp; J. Rainio" country="Madagascar" elevation="350" location="Vatoharanana" pageId="25" pageNumber="226" specimenCount="2" typeStatus="holotype">
<typeStatus id="5D2C29E33060FF89FE73FEEDBC8A0C8C" box="[390,489,383,407]" pageId="25" pageNumber="226" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC43060FF89FE0BFEEDBFE30C8C" box="[510,640,383,408]" captionStart="FIGURE" captionStartId="26.[155,167,1478,1497]" captionTargetBox="[139,1176,216,1455]" captionTargetId="figure-53@26.[120,1176,216,1455]" captionTargetPageId="26" captionText="FIGURE 14. Digital images of holotype male of Perigona (Ripogena) deuvei Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13799473" httpUri="https://zenodo.org/record/13799473/files/figure.png" pageId="25" pageNumber="226">Figs.14A–B</figureCitation>
), a male, in CAS, labeled: “
<collectingCountry id="FA80D7D13060FF89FC24FEEDB9EC0C8C" box="[977,1167,383,407]" name="Madagascar" pageId="25" pageNumber="226">MADAGASCAR</collectingCountry>
, Ranomafana NP.,
<location id="8748C19A3060FF89FECBFE0DBCAC0CAC" LSID="urn:lsid:plazi:treatment:0A3E26573060FF8BFF8EFF49BEB20CA8:8748C19A3060FF89FECBFE0DBCAC0CAC" box="[318,463,415,439]" country="Madagascar" name="Vatoharanana" pageId="25" pageNumber="226">Vatoharanana</location>
, Trail K at
<quantity id="456F3AA43060FF89FDBAFE0DBFF60CAC" box="[591,661,415,439]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="25" pageNumber="226" unit="m" value="350.0">
<elevation id="09BA70723060FF89FDBAFE0DBFF60CAC" box="[591,661,415,439]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="25" pageNumber="226" unit="m" value="350.0">350 m</elevation>
</quantity>
mark,
<date id="F629B1813060FF89FD17FE32BE390CAC" box="[738,858,415,440]" pageId="25" pageNumber="226" value="2005-07-24">
<collectingDate id="E66D48693060FF89FD17FE32BE390CAC" box="[738,858,415,440]" pageId="25" pageNumber="226" value="2005-07-24">24.07.2005</collectingDate>
</date>
”/ “collected by hand in daytime,
<collectorName id="2F62F2973060FF89FF42FE52BC4B0CC3" box="[183,296,448,472]" pageId="25" pageNumber="226">F. Ratalata</collectorName>
&amp;
<collectorName id="2F62F2973060FF89FEBCFE52BCC40CC3" box="[329,423,448,472]" pageId="25" pageNumber="226">J. Rainio</collectorName>
collectors”/ “
<typeStatus id="5D2C29E33060FF89FDCEFE52BFAB0CC3" box="[571,712,448,472]" pageId="25" pageNumber="226" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC23060FF89FD3BFE52BDA10CE3" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="deuvei" status="sp. nov." subGenus="Ripogena">
<emphasis id="B0E34B533060FF89FD3BFE52BE520CCC" box="[718,817,448,471]" italics="true" pageId="25" pageNumber="226">Perigona</emphasis>
(
<emphasis id="B0E34B533060FF89FCB5FE52BEC50CCC" box="[832,934,448,471]" italics="true" pageId="25" pageNumber="226">Ripogena</emphasis>
)
<emphasis id="B0E34B533060FF89FC40FE52BE980CC3" box="[949,1019,448,472]" italics="true" pageId="25" pageNumber="226">deuvei</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283060FF89FF3CFE72BC610CEC" box="[201,258,480,503]" pageId="25" pageNumber="226" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label]
</materialsCitation>
.
<materialsCitation id="32FF9D1C3060FF89FE38FE72BE010F23" collectingDate="2005-10-06" collectionCode="MNHN" collectorName="F. Ratalata &amp; J. Rainio" country="Madagascar" elevation="50" location="Vatoharanana" municipality="Trail" pageId="25" pageNumber="226" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="5D2C29E33060FF89FE38FE72BF560CE3" box="[461,565,480,504]" pageId="25" pageNumber="226" type="paratype">Paratypes</typeStatus>
(total of 4):
<specimenCount id="94915CC83060FF89FD49FE72BE790CE3" box="[700,794,480,504]" count="1" pageId="25" pageNumber="226" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F843060FF89FCB0FE72BEF50CEC" box="[837,918,480,503]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="226">MNHN</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13060FF89FBF3FE72BDD10F03" name="Madagascar" pageId="25" pageNumber="226">MADAGASCAR</collectingCountry>
, Ranomafana NP.,
<location id="8748C19A3060FF89FE88FD92BF6D0F03" LSID="urn:lsid:plazi:treatment:0A3E26573060FF8BFF8EFF49BEB20CA8:8748C19A3060FF89FE88FD92BF6D0F03" box="[381,526,512,536]" country="Madagascar" municipality="Trail" name="Vatoharanana" pageId="25" pageNumber="226">Vatoharanana</location>
,
<collectingMunicipality id="624C0D3B3060FF89FDEFFD92BF2E0F03" box="[538,589,512,536]" pageId="25" pageNumber="226">Trail</collectingMunicipality>
J at
<quantity id="456F3AA43060FF89FD89FD92BFD10F0C" box="[636,690,512,536]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="226" unit="m" value="50.0">
<elevation id="09BA70723060FF89FD89FD92BFD10F0C" box="[636,690,512,536]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="25" pageNumber="226" unit="m" value="50.0">50 m</elevation>
</quantity>
mark,
<date id="F629B1813060FF89FD0EFD92BE060F03" box="[763,869,512,536]" pageId="25" pageNumber="226" value="2005-10-06">
<collectingDate id="E66D48693060FF89FD0EFD92BE060F03" box="[763,869,512,536]" pageId="25" pageNumber="226" value="2005-10-06">6.10.2005</collectingDate>
</date>
”/ “collected by hand in daytime,
<collectorName id="2F62F2973060FF89FF4FFDB3BC4D0F23" box="[186,302,544,568]" pageId="25" pageNumber="226">F. Ratalata</collectorName>
&amp;
<collectorName id="2F62F2973060FF89FEA0FDB3BCD40F23" box="[341,439,544,568]" pageId="25" pageNumber="226">J. Rainio</collectorName>
collectors”/ “239.” [handwritten label]
</materialsCitation>
;
<materialsCitation id="32FF9D1C3060FF89FC86FDB2B9720F63" collectingDate="2005-09-09" collectionCode="CAS" collectorName="F. Ratalata &amp; J. Rainio" country="Madagascar" elevation="600" location="Vatoharanana" municipality="Trail" pageId="25" pageNumber="226" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="94915CC83060FF89FC86FDB2BEB00F23" box="[883,979,544,568]" count="1" pageId="25" pageNumber="226" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F843060FF89FBF7FDB2B9550F23" box="[1026,1078,544,568]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="25" pageNumber="226" type="Museum">CAS</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13060FF89FF72FDD2BC260F43" box="[135,325,576,600]" name="Madagascar" pageId="25" pageNumber="226">MADAGASCAR</collectingCountry>
, Ranomafana NP.,
<location id="8748C19A3060FF89FDE0FDD2BFC50F43" LSID="urn:lsid:plazi:treatment:0A3E26573060FF8BFF8EFF49BEB20CA8:8748C19A3060FF89FDE0FDD2BFC50F43" box="[533,678,576,600]" country="Madagascar" municipality="Trail" name="Vatoharanana" pageId="25" pageNumber="226">Vatoharanana</location>
,
<collectingMunicipality id="624C0D3B3060FF89FD41FDD2BF840F43" box="[692,743,576,600]" pageId="25" pageNumber="226">Trail</collectingMunicipality>
S at
<quantity id="456F3AA43060FF89FCD5FDD2BE070F43" box="[800,868,576,600]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="25" pageNumber="226" unit="m" value="600.0">
<elevation id="09BA70723060FF89FCD5FDD2BE070F43" box="[800,868,576,600]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="25" pageNumber="226" unit="m" value="600.0">600 m</elevation>
</quantity>
mark,
<date id="F629B1813060FF89FC44FDD2B96D0F43" box="[945,1038,576,600]" pageId="25" pageNumber="226" value="2005-09-09">
<collectingDate id="E66D48693060FF89FC44FDD2B96D0F43" box="[945,1038,576,600]" pageId="25" pageNumber="226" value="2005-09-09">9.9.2005</collectingDate>
</date>
”/ “collected by hand in daytime,
<collectorName id="2F62F2973060FF89FE96FDF3BCB40F63" box="[355,471,608,632]" pageId="25" pageNumber="226">F. Ratalata</collectorName>
&amp;
<collectorName id="2F62F2973060FF89FDF5FDF3BF010F63" box="[512,610,608,632]" pageId="25" pageNumber="226">J. Rainio</collectorName>
collectors”/ “239.” [handwritten label]
</materialsCitation>
;
<materialsCitation id="32FF9D1C3060FF89FBD7FDF2BD830FC3" collectingDate="2005-07-24" collectionCode="MZF" collectorName="F. Ratalata &amp; J. Rainio" country="Madagascar" elevation="600" location="Vatoharanana" municipality="Trail" pageId="25" pageNumber="226" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="94915CC83060FF89FBD7FDF2B90D0F63" box="[1058,1134,608,632]" count="1" pageId="25" pageNumber="226" type="male">1 male</specimenCount>
(in
<collectionCode id="E4860F843060FF89FF8EFD13BDD10F83" box="[123,178,641,664]" pageId="25" pageNumber="226">MZF</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13060FF89FEC4FD13BC8C0F82" box="[305,495,641,665]" name="Madagascar" pageId="25" pageNumber="226">MADAGASCAR</collectingCountry>
, Ranomafana NP.,
<location id="8748C19A3060FF89FD20FD12BE050F83" LSID="urn:lsid:plazi:treatment:0A3E26573060FF8BFF8EFF49BEB20CA8:8748C19A3060FF89FD20FD12BE050F83" box="[725,870,640,664]" country="Madagascar" municipality="Trail" name="Vatoharanana" pageId="25" pageNumber="226">Vatoharanana</location>
,
<collectingMunicipality id="624C0D3B3060FF89FC8EFD12BECD0F83" box="[891,942,640,664]" pageId="25" pageNumber="226">Trail</collectingMunicipality>
S at
<quantity id="456F3AA43060FF89FC08FD12B92A0F83" box="[1021,1097,640,664]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="25" pageNumber="226" unit="m" value="600.0">
<elevation id="09BA70723060FF89FC08FD12B92A0F83" box="[1021,1097,640,664]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="25" pageNumber="226" unit="m" value="600.0">600 m</elevation>
</quantity>
mark,
<date id="F629B1813060FF89FF8EFD33BD860FA3" box="[123,229,672,697]" pageId="25" pageNumber="226" value="2005-07-24">
<collectingDate id="E66D48693060FF89FF8EFD33BD860FA3" box="[123,229,672,697]" pageId="25" pageNumber="226" value="2005-07-24">24.7.2005</collectingDate>
</date>
”/ “collected by hand in daytime,
<collectorName id="2F62F2973060FF89FDA1FD33BFA40FA2" box="[596,711,673,697]" pageId="25" pageNumber="226">F. Ratalata</collectorName>
&amp;
<collectorName id="2F62F2973060FF89FD19FD33BE2F0FA2" box="[748,844,673,697]" pageId="25" pageNumber="226">J. Rainio</collectorName>
collectors”/ “253.” [handwritten label]
</materialsCitation>
;
<materialsCitation id="32FF9D1C3060FF89FF1BFD53BE320E22" collectingDate="2005-10-06" collectionCode="MZF" collectorName="F. Ratalata &amp; J. Rainio" country="Madagascar" elevation="600" location="Vatoharanana" municipality="Trail" pageId="25" pageNumber="226" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="94915CC83060FF89FF1BFD53BC2F0FC2" box="[238,332,705,729]" count="1" pageId="25" pageNumber="226" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F843060FF89FE8DFD53BCCC0FC3" box="[376,431,705,728]" pageId="25" pageNumber="226">MZF</collectionCode>
) labeled “
<collectingCountry id="FA80D7D13060FF89FDD5FD53BFBD0FC2" box="[544,734,705,729]" name="Madagascar" pageId="25" pageNumber="226">MADAGASCAR</collectingCountry>
, Ranomafana NP.,
<location id="8748C19A3060FF89FC58FD53B95D0FC2" LSID="urn:lsid:plazi:treatment:0A3E26573060FF8BFF8EFF49BEB20CA8:8748C19A3060FF89FC58FD53B95D0FC2" box="[941,1086,705,729]" country="Madagascar" municipality="Trail" name="Vatoharanana" pageId="25" pageNumber="226">Vatoharanana</location>
,
<collectingMunicipality id="624C0D3B3060FF89FBB9FD53B91C0FC2" box="[1100,1151,705,729]" pageId="25" pageNumber="226">Trail</collectingMunicipality>
S at
<quantity id="456F3AA43060FF89FF6DFD73BDBD0FE3" box="[152,222,737,761]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="25" pageNumber="226" unit="m" value="600.0">
<elevation id="09BA70723060FF89FF6DFD73BDBD0FE3" box="[152,222,737,761]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="25" pageNumber="226" unit="m" value="600.0">600 m</elevation>
</quantity>
mark,
<date id="F629B1813060FF89FEDAFD73BCFA0FE2" box="[303,409,737,761]" pageId="25" pageNumber="226" value="2005-10-06">
<collectingDate id="E66D48693060FF89FEDAFD73BCFA0FE2" box="[303,409,737,761]" pageId="25" pageNumber="226" value="2005-10-06">6.10.2005</collectingDate>
</date>
”/ “collected by hand in daytime,
<collectorName id="2F62F2973060FF89FCE6FD73BEE40FE2" box="[787,903,737,761]" pageId="25" pageNumber="226">F. Ratalata</collectorName>
&amp;
<collectorName id="2F62F2973060FF89FC5AFD73B9720FE2" box="[943,1041,737,761]" pageId="25" pageNumber="226">J. Rainio</collectorName>
collectors”/ “245.” [handwritten label]. All
<typeStatus id="5D2C29E33060FF89FE22FC90BF5E0E01" box="[471,573,770,794]" pageId="25" pageNumber="226" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="5D2C29E33060FF89FC56FC93B9440E02" box="[931,1063,769,793]" pageId="25" pageNumber="226" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4597ECC23060FF89FBC7FC90BF480E22" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="deuvei" status="sp. nov." subGenus="Ripogena">
<emphasis id="B0E34B533060FF89FBC7FC90B9F60E02" box="[1074,1173,770,793]" italics="true" pageId="25" pageNumber="226">Perigona</emphasis>
(
<emphasis id="B0E34B533060FF89FF71FCB0BD890E22" box="[132,234,802,825]" italics="true" pageId="25" pageNumber="226">Ripogena</emphasis>
)
<emphasis id="B0E34B533060FF89FF0FFCB3BC230E22" box="[250,320,801,825]" italics="true" pageId="25" pageNumber="226">deuvei</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283060FF89FDC7FCB0BF0F0E22" box="[562,620,802,825]" pageId="25" pageNumber="226" rank="species">sp. n.</taxonomicNameLabel>
2015” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="822897413060FF89FF5EFCD0B9500E42" blockId="25.[123,1174,383,1726]" box="[171,1075,833,858]" pageId="25" pageNumber="226">
<materialsCitation id="32FF9D1C3060FF89FF5EFCD0B94F0E42" box="[171,1068,833,858]" country="Madagascar" location="Ranomafana National Park" pageId="25" pageNumber="226" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B0E34B533060FF89FF5EFCD0BC340E43" bold="true" box="[171,343,834,858]" pageId="25" pageNumber="226">
<smallCapsWord id="84CE019D3060FF89FF5EFCD0BD870E43" baselines="852,851" box="[171,228,834,858]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="25" pageNumber="226">
<typeStatus id="5D2C29E33060FF89FF5EFCD0BD870E43" box="[171,228,834,858]" pageId="25" pageNumber="226">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="84CE019D3060FF89FF19FCD7BC340E43" baselines="851" box="[236,343,837,856]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="25" pageNumber="226">LOCALITY</smallCapsWord>
</emphasis>
.—
<collectingCountry id="FA80D7D13060FF89FE8BFCD3BC9C0E42" box="[382,511,833,857]" name="Madagascar" pageId="25" pageNumber="226">Madagascar</collectingCountry>
,
<collectingRegion id="405359A33060FF89FDF8FCD3BF9F0E42" box="[525,764,833,857]" country="Madagascar" name="Fianarantsoa" pageId="25" pageNumber="226">Fianarantsoa Province</collectingRegion>
,
<location id="8748C19A3060FF89FCFFFCD3B94F0E42" LSID="urn:lsid:plazi:treatment:0A3E26573060FF8BFF8EFF49BEB20CA8:8748C19A3060FF89FCFFFCD3B94F0E42" box="[778,1068,833,857]" country="Madagascar" name="Ranomafana National Park" pageId="25" pageNumber="226" stateProvince="Fianarantsoa">Ranomafana National Park</location>
</materialsCitation>
.
</paragraph>
<paragraph id="822897413060FF89FF5EFCF0BD8B0EE1" blockId="25.[123,1174,383,1726]" pageId="25" pageNumber="226">
<emphasis id="B0E34B533060FF89FF5EFCF0BC8D0E63" bold="true" box="[171,494,866,890]" pageId="25" pageNumber="226">
<smallCapsWord id="84CE019D3060FF89FF5EFCF0BC4D0E63" baselines="884,883" box="[171,302,866,890]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Derivation" pageId="25" pageNumber="226">DERIVATION</smallCapsWord>
<smallCapsWord id="84CE019D3060FF89FEC0FCF7BC320E63" baselines="883" box="[309,337,869,888]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="25" pageNumber="226">OF</smallCapsWord>
<smallCapsWord id="84CE019D3060FF89FEA2FCF7BCC90E63" baselines="883" box="[343,426,869,888]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="25" pageNumber="226">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D3060FF89FE44FCF7BC8D0E63" baselines="883" box="[433,494,869,888]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="25" pageNumber="226">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC23060FF89FD04FCF0BE540E61" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[753,823,866,890]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="deuvei">
<emphasis id="B0E34B533060FF89FD04FCF0BE540E61" box="[753,823,866,890]" italics="true" pageId="25" pageNumber="226">deuvei</emphasis>
</taxonomicName>
, is a noun in apposition, derived from the surname of Dr. Thierry Deuve of the Laboratoire d’Entomologie at the Muséum National d’Histoire Naturelle in Paris. We are pleased to name this new species in honor of Dr. Deuve in thanks for his friendship and kind hospitality during our several visits to MNHN in the course of this study.
</paragraph>
<paragraph id="822897413060FF8BFF5EFB90BE880C48" blockId="25.[123,1174,383,1726]" lastBlockId="27.[123,1174,219,436]" lastPageId="27" lastPageNumber="228" pageId="25" pageNumber="226">
<smallCapsWord id="84CE019D3060FF89FF5EFB90BC210902" baselines="1044,1044" box="[171,322,1026,1050]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="25" pageNumber="226">
<emphasis id="B0E34B533060FF89FF5EFB90BC210902" bold="true" box="[171,322,1026,1050]" pageId="25" pageNumber="226">RECOGNITION</emphasis>
</smallCapsWord>
.— Size average for subgenus, SBL of males = 3.5–
<quantity id="456F3AA43060FF89FC9BFB90BEA30901" box="[878,960,1026,1050]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="25" pageNumber="226" unit="mm" value="3.8">3.8 mm</quantity>
, of females 3.8–
<quantity id="456F3AA43060FF89FB81FB90BDC70921" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="25" pageNumber="226" unit="mm" value="3.9">3.9 mm</quantity>
. Members of this species (
<figureCitation id="1AAC8BC43060FF89FE32FBB0BF440920" box="[455,551,1058,1083]" captionStart="FIGURE" captionStartId="26.[155,167,1478,1497]" captionTargetBox="[139,1176,216,1455]" captionTargetId="figure-53@26.[120,1176,216,1455]" captionTargetPageId="26" captionText="FIGURE 14. Digital images of holotype male of Perigona (Ripogena) deuvei Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13799473" httpUri="https://zenodo.org/record/13799473/files/figure.png" pageId="25" pageNumber="226">Fig. 14A</figureCitation>
) share with those of
<taxonomicName id="4597ECC23060FF89FCFEFBB1B9F60920" authority="Alluaud (1936)" authorityName="Alluaud" authorityYear="1936" box="[779,1173,1058,1083]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="heterodera">
<emphasis id="B0E34B533060FF89FCFEFBB1BE890921" box="[779,1002,1058,1082]" italics="true" pageId="25" pageNumber="226">Perigona heterodera</emphasis>
Alluaud (1936)
</taxonomicName>
and
<taxonomicName id="4597ECC23060FF89FF5DFBD1BF750940" authority="Alluaud (1936)" authorityName="Alluaud" authorityYear="1936" box="[168,534,1091,1115]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="prasinus">
<emphasis id="B0E34B533060FF89FF5DFBD1BC0E0941" box="[168,365,1091,1114]" italics="true" pageId="25" pageNumber="226">Perigona prasinus</emphasis>
Alluaud (1936)
</taxonomicName>
the following features: pronotum with lateral margins without sinuation anterior to hind angles, posterior margin slightly and smoothly convex, and hind angles either obtuse or broadly rounded. These features distinguish members of these three species from those of
<taxonomicName id="4597ECC23060FF89FEE2FB36BFDD09A0" authority="Alluaud (1936)" authorityName="Alluaud" authorityYear="1936" box="[279,702,1187,1211]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="bembidioides">
<emphasis id="B0E34B533060FF89FEE2FB36BF7209A0" box="[279,529,1187,1211]" italics="true" pageId="25" pageNumber="226">Perigona bembidioides</emphasis>
Alluaud (1936)
</taxonomicName>
and
<taxonomicName id="4597ECC23060FF89FD00FB36B9EC09A0" authority="Jeannel (1948)" authorityName="Jeannel" authorityYear="1948" box="[757,1167,1187,1211]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="viridimicans">
<emphasis id="B0E34B533060FF89FD00FB36BE8509A0" box="[757,998,1187,1211]" italics="true" pageId="25" pageNumber="226">Perigona viridimicans</emphasis>
Jeannel (1948)
</taxonomicName>
, which have distinct sinuation of the lateral margins anterior to rectangular hind angles and a straight basal margin. Members of
<taxonomicName id="4597ECC23060FF89FE19FB76BF2F09E0" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[492,588,1251,1275]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="deuvei">
<emphasis id="B0E34B533060FF89FE19FB76BF2F09E0" box="[492,588,1251,1275]" italics="true" pageId="25" pageNumber="226">P. deuvei</emphasis>
</taxonomicName>
differ from those of
<taxonomicName id="4597ECC23060FF89FCD2FB76BEFF09E0" authorityName="Alluaud" authorityYear="1936" box="[807,924,1252,1275]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="prasinus">
<emphasis id="B0E34B533060FF89FCD2FB76BEFF09E0" box="[807,924,1252,1275]" italics="true" pageId="25" pageNumber="226">P. prasinus</emphasis>
</taxonomicName>
in having smaller body size (SBL =
<quantity id="456F3AA43060FF89FEF7FA96BC360800" box="[258,341,1284,1308]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="25" pageNumber="226" unit="mm" value="4.7">4.7 mm</quantity>
in
<taxonomicName id="4597ECC23060FF89FE8FFA96BC910800" authorityName="Alluaud" authorityYear="1936" box="[378,498,1284,1307]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="prasinus">
<emphasis id="B0E34B533060FF89FE8FFA96BC910800" box="[378,498,1284,1307]" italics="true" pageId="25" pageNumber="226">P. prasinus</emphasis>
</taxonomicName>
), narrower body form, and dorsum without metallic reflection (dorsum with metallic green reflection in
<taxonomicName id="4597ECC23060FF89FDC9FAB6BFD10820" authorityName="Alluaud" authorityYear="1936" box="[572,690,1316,1339]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="prasinus">
<emphasis id="B0E34B533060FF89FDC9FAB6BFD10820" box="[572,690,1316,1339]" italics="true" pageId="25" pageNumber="226">P. prasinus</emphasis>
</taxonomicName>
) but elytra with moderate but distinct iridescence generated by microsculpture comprised of transverse microlines and marjkedly transverse sculpticells. They are most similar to members of
<taxonomicName id="4597ECC23060FF89FD6AFAF7BE4C0867" authorityName="Alluaud" authorityYear="1936" box="[671,815,1380,1404]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="heterodera">
<emphasis id="B0E34B533060FF89FD6AFAF7BE4C0867" box="[671,815,1380,1404]" italics="true" pageId="25" pageNumber="226">P. heterodera</emphasis>
</taxonomicName>
in body form and size but differ from them in having the pronotum proportionately longer and male genitalia (
<figureCitation id="1AAC8BC43060FF89FC3EFA16B9360887" box="[971,1109,1412,1436]" captionStart="FIGURE" captionStartId="26.[155,167,1478,1497]" captionTargetBox="[139,1176,216,1455]" captionTargetId="figure-53@26.[120,1176,216,1455]" captionTargetPageId="26" captionText="FIGURE 14. Digital images of holotype male of Perigona (Ripogena) deuvei Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13799473" httpUri="https://zenodo.org/record/13799473/files/figure.png" pageId="25" pageNumber="226">Figs. 14C–D</figureCitation>
) with shaft markedly narrowed basally, markedly inflated in apical two-thirds, and apex longer, slender and ventrally bent in lateral view (
<figureCitation id="1AAC8BC43060FF89FE0DFA56BF3B08C7" box="[504,600,1476,1500]" captionStart="FIGURE" captionStartId="26.[155,167,1478,1497]" captionTargetBox="[139,1176,216,1455]" captionTargetId="figure-53@26.[120,1176,216,1455]" captionTargetPageId="26" captionText="FIGURE 14. Digital images of holotype male of Perigona (Ripogena) deuvei Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13799473" httpUri="https://zenodo.org/record/13799473/files/figure.png" pageId="25" pageNumber="226">Fig. 14C</figureCitation>
), slightly deflected left and slightly longer in dorsal view (
<figureCitation id="1AAC8BC43060FF89FF4AFA76BC7C08E7" box="[191,287,1508,1533]" captionStart="FIGURE" captionStartId="26.[155,167,1478,1497]" captionTargetBox="[139,1176,216,1455]" captionTargetId="figure-53@26.[120,1176,216,1455]" captionTargetPageId="26" captionText="FIGURE 14. Digital images of holotype male of Perigona (Ripogena) deuvei Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13799473" httpUri="https://zenodo.org/record/13799473/files/figure.png" pageId="25" pageNumber="226">Fig. 14D</figureCitation>
) than in
<taxonomicName id="4597ECC23060FF89FE77FA77BF7208E7" authorityName="Alluaud" authorityYear="1936" box="[386,529,1508,1532]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="heterodera">
<emphasis id="B0E34B533060FF89FE77FA77BF7208E7" box="[386,529,1508,1532]" italics="true" pageId="25" pageNumber="226">P. heterodera</emphasis>
</taxonomicName>
(see Figs. 351b-c in Jeannel 1948). They differ from members of
<taxonomicName id="4597ECC23060FF89FF3AF997BF1C0B06" authority="(Deuve 1998)" baseAuthorityName="Deuve" baseAuthorityYear="1998" box="[207,639,1541,1565]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="descarpentriesi">
<emphasis id="B0E34B533060FF89FF3AF997BC830B06" box="[207,480,1541,1565]" italics="true" pageId="25" pageNumber="226">Perigona descarpentriesi</emphasis>
(
<bibRefCitation id="E606EAB03060FF89FE07F997BF150B06" author="DEUVE, T." box="[498,630,1541,1565]" pageId="25" pageNumber="226" pagination="2 - 4" refId="ref30013" refString="DEUVE, T. 1998. Description d'un Perigoninae microphtalme de Madagascar [Coleoptera, Harpalidea]. Revue Francaise d'Entomologie (N. S.) 20: 2 - 4." type="journal article" year="1998">Deuve 1998</bibRefCitation>
)
</taxonomicName>
in having moderately large eyes (eyes markedly reduced eyes in size in
<taxonomicName id="4597ECC23060FF89FE84F9B4BF4C0B26" baseAuthorityName="Deuve" baseAuthorityYear="1998" box="[369,559,1573,1597]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="descarpentriesi">
<emphasis id="B0E34B533060FF89FE84F9B4BF4C0B26" box="[369,559,1573,1597]" italics="true" pageId="25" pageNumber="226">P. descarpentriesi</emphasis>
</taxonomicName>
), pronotum distinctly narrower and with less rounded lateral margins, and elytra with only stria 1 deeply impressed, stria 2 and 3 shallowly impressed but evident on disk, striae 4 to 7 effaced, and stria 8 impressed only in apical one-third (striae 1–7 shallowly impressed but evident at least on disc and stria 8 deeply impressed throughout in
<taxonomicName id="4597ECC23060FF89FBDFF914BDBB0BA6" baseAuthorityName="Deuve" baseAuthorityYear="1998" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="descarpentriesi">
<emphasis id="B0E34B533060FF89FBDFF914BDBB0BA6" italics="true" pageId="25" pageNumber="226">P. descarpentriesi</emphasis>
</taxonomicName>
members). Also, the apex of the median lobe of the male genitalia is narrower in lateral aspect in
<taxonomicName id="4597ECC23062FF8BFF15FF4EBC230DE8" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[224,320,219,243]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="deuvei">
<emphasis id="B0E34B533062FF8BFF15FF4EBC230DE8" box="[224,320,219,243]" italics="true" pageId="27" pageNumber="228">P. deuvei</emphasis>
</taxonomicName>
males (Fig. 143C) than in those of
<taxonomicName id="4597ECC23062FF8BFD4EFF4EBE1A0DE8" baseAuthorityName="Deuve" baseAuthorityYear="1998" box="[699,889,219,243]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="descarpentriesi">
<emphasis id="B0E34B533062FF8BFD4EFF4EBE1A0DE8" box="[699,889,219,243]" italics="true" pageId="27" pageNumber="228">P. descarpentriesi</emphasis>
</taxonomicName>
(see
<bibRefCitation id="E606EAB03062FF8BFC47FF4EB9500DE8" author="DEUVE, T." box="[946,1075,219,243]" pageId="27" pageNumber="228" pagination="2 - 4" refId="ref30013" refString="DEUVE, T. 1998. Description d'un Perigoninae microphtalme de Madagascar [Coleoptera, Harpalidea]. Revue Francaise d'Entomologie (N. S.) 20: 2 - 4." type="journal article" year="1998">Deuve 1998</bibRefCitation>
,
<figureCitation id="1AAC8BC43062FF8BFBB4FF49B91C0DE8" box="[1089,1151,219,243]" captionStart="FIGURE" captionStartId="5.[155,167,1152,1171]" captionTargetBox="[133,1174,245,1135]" captionTargetId="figure-997@5.[120,1174,216,1150]" captionTargetPageId="5" captionText="FIGURE 2. Digital images of holotype female of Brachypelus ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799443" httpUri="https://zenodo.org/record/13799443/files/figure.png" pageId="27" pageNumber="228">Fig. 2</figureCitation>
.). Finally,
<taxonomicName id="4597ECC23062FF8BFF26FF6EBC570C08" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[211,308,251,275]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="deuvei">
<emphasis id="B0E34B533062FF8BFF26FF6EBC570C08" box="[211,308,251,275]" italics="true" pageId="27" pageNumber="228">P. deuvei</emphasis>
</taxonomicName>
members differ from those of the new species described below in having small- er body size, pronotum with lateral margination very narrow throughout, fewer elytral striae evident, and median lobe of the male genitalia with a longer and ventrally bent apex.
</paragraph>
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<paragraph id="822897413063FF8AFF6EFA54BC680B17" blockId="26.[123,1173,1478,1548]" pageId="26" pageNumber="227">
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14. Digital images of holotype male of
<taxonomicName id="4597ECC23063FF8AFDC1FA54BE8608C2" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[564,997,1478,1497]" class="Insecta" family="Carabidae" genus="Perigona" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="deuvei" status="sp. nov." subGenus="Ripogena">
<emphasis id="B0E34B533063FF8AFDC1FA54BFE008C3" box="[564,643,1478,1496]" italics="true" pageId="26" pageNumber="227">Perigona</emphasis>
(
<emphasis id="B0E34B533063FF8AFD64FA54BF8008C3" box="[657,739,1478,1496]" italics="true" pageId="26" pageNumber="227">Ripogena</emphasis>
)
<emphasis id="B0E34B533063FF8AFD05FA54BE4B08C2" box="[752,808,1478,1497]" italics="true" pageId="26" pageNumber="227">deuvei</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283063FF8AFC1EFA54B94E08C2" box="[1003,1069,1478,1497]" pageId="26" pageNumber="227" rank="species">sp. nov.</taxonomicNameLabel>
A. Habitus, dorsal aspect; B Labels; C–D. Aedeagus of genitalia; C. Left lateral aspect; D. Dorsal aspect. Scale lines A = 1.0 mm, C–D = 0.25 mm.
</paragraph>
</caption>
<paragraph id="822897413062FF8BFF5EFEC9B9690C68" blockId="27.[123,1174,219,436]" box="[171,1034,347,372]" pageId="27" pageNumber="228">
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<smallCapsWord id="84CE019D3062FF8BFEA8FECDBC920C69" baselines="365" box="[349,497,351,370]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="27" pageNumber="228">DISTRIBUTION</smallCapsWord>
</emphasis>
.— At present, known only from the
<typeStatus id="5D2C29E33062FF8BFC8AFECEBECE0C6F" box="[895,941,348,372]" pageId="27" pageNumber="228">type</typeStatus>
locality.
</paragraph>
<paragraph id="822897413062FF8BFF5EFEEEBEB20CA8" blockId="27.[123,1174,219,436]" pageId="27" pageNumber="228">
<emphasis id="B0E34B533062FF8BFF5EFEEEBCC70C89" bold="true" box="[171,420,380,404]" pageId="27" pageNumber="228">
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</emphasis>
.— All specimens of this species were collected by hand in daytime in primary montane rainforest in the Vatoharanana area at an elevation of
<quantity id="456F3AA43062FF8BFC8DFE0EBEA90CA8" box="[888,970,412,436]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="27" pageNumber="228" unit="m" value="1200.0">1200 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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