treatments-xml/data/0A/3E/26/0A3E2657305CFFB8FF8EFB4BBCA909E0.xml

<document id="5591FD069CEC97E08C7A8C4F54B4728E" ID-DOI="10.5281/zenodo.13799439" ID-ISSN="0068-547X" ID-Zenodo-Dep="13799439" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538212676" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Rainio, Johanna" docDate="2016" docId="0A3E2657305CFFB8FF8EFB4BBCA909E0" docLanguage="en" docName="ProcCalifAcadSci.63.7.201-268.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (7)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Pristacrus ranomafanae Kavanaugh and Rainio 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="241" masterDocId="F6075E2F3079FF90FFF5FF92BD630D1B" masterDocTitle="Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar" masterLastPageNumber="268" masterPageNumber="201" pageNumber="238" updateTime="1726771873529" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="CFA655D069656479868B720A1A83BB42">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
</mods:titleInfo>
<mods:name id="0E998F804307586877877B4CC2BFD36D" type="personal">
<mods:role id="B5ABE2BF348300475CA80EB7F2E34BFE">
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<mods:namePart id="829B0BB6810B62365B5D917B2CE4F03E">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="5CF20DC87DAC37E938528970B48A73F8">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
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<mods:namePart id="63D3577E329CF7F36A2443D5E5AA3B37">Rainio, Johanna</mods:namePart>
<mods:affiliation id="DCBBAEE8E3E94D885460EF98D0DD4ED1">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
</mods:name>
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<mods:title id="E23AFE1C40B856ABA9C9B263B9A5860B">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="46D0F80380FFB1892B521725B61348E4">
<mods:date id="266621E0CC2953FAB718516170B91AD8">2016</mods:date>
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<mods:number id="7DD4C0D03D0EE9789A8D83AFF742575C">2016-04-29</mods:number>
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<mods:number id="B32A0097C404B7AD1B00B666DD54447E">63</mods:number>
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<mods:number id="7903FDAEF26C0B8CDC673E66E7589452">7</mods:number>
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<mods:identifier id="D0E7B3FCB6FBA8A3F193278ABF7DBAA5" type="DOI">10.5281/zenodo.13799439</mods:identifier>
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<treatment id="0A3E2657305CFFB8FF8EFB4BBCA909E0" LSID="urn:lsid:plazi:treatment:0A3E2657305CFFB8FF8EFB4BBCA909E0" httpUri="http://treatment.plazi.org/id/0A3E2657305CFFB8FF8EFB4BBCA909E0" lastPageId="40" lastPageNumber="241" pageId="37" pageNumber="238">
<subSubSection id="CA8DC4CA305CFFB5FF8EFB4BBF9809EA" box="[123,763,1240,1265]" pageId="37" pageNumber="238" type="nomenclature">
<paragraph id="82289741305CFFB5FF8EFB4BBF9809EA" blockId="37.[123,763,1240,1295]" box="[123,763,1240,1265]" pageId="37" pageNumber="238">
<heading id="D960202D305CFFB5FF8EFB4BBF9809EA" box="[123,763,1240,1265]" fontSize="10" level="2" pageId="37" pageNumber="238" reason="4">
<emphasis id="B0E34B53305CFFB5FF8EFB4BBF9809EA" bold="true" box="[123,763,1240,1265]" pageId="37" pageNumber="238">
<taxonomicName id="4597ECC2305CFFB5FF8EFB4BBFFB09EA" ID-CoL="77WH4" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,664,1240,1265]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B53305CFFB5FF8EFB4BBCE409EB" bold="true" box="[123,391,1240,1265]" italics="true" pageId="37" pageNumber="238">Pristacrus ranomafanae</emphasis>
Kavanaugh and Rainio
</taxonomicName>
, 
<taxonomicNameLabel id="ABD0F628305CFFB5FD50FB4BBF9809EA" box="[677,763,1241,1265]" pageId="37" pageNumber="238" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA305CFFB5FF8EFB6BBE010820" pageId="37" pageNumber="238" type="description">
<paragraph id="82289741305CFFB5FF8EFB6BBC660814" blockId="37.[123,763,1240,1295]" box="[123,261,1273,1295]" pageId="37" pageNumber="238">
<figureCitation id="1AAC8BC4305CFFB5FF8EFB6BBD820814" box="[123,225,1273,1295]" captionStart="FIGURE" captionStartId="38.[155,167,1385,1404]" captionTargetBox="[120,1176,216,1353]" captionTargetId="figure-677@38.[120,1176,216,1372]" captionTargetPageId="38" captionText="FIGURE 21. Digital images of holotype male of Pristacrus ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B Labels. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799489" httpUri="https://zenodo.org/record/13799489/files/figure.png" pageId="37" pageNumber="238">Figures 21</figureCitation>
–22
</paragraph>
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<emphasis id="B0E34B53305CFFB5FF8EFAB4BC4F0821" box="[123,300,1317,1339]" italics="true" pageId="37" pageNumber="238">
<taxonomicName id="4597ECC2305CFFB5FF8EFAB4BDBC0820" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[123,223,1318,1339]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="genus" status="n. sp.">Pristacrus</taxonomicName>
<taxonomicNameLabel id="ABD0F628305CFFB5FF10FAB4BC7A0820" box="[229,281,1318,1339]" pageId="37" pageNumber="238" rank="species">n. sp.</taxonomicNameLabel>
1
</emphasis>
; Rainio 2009: 33, Rainio 2012: 73 (informal designation).
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA305CFFB8FF5EFACDBCA909E0" lastPageId="40" lastPageNumber="241" pageId="37" pageNumber="238" type="materials_examined">
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</emphasis>
.— 
<materialsCitation id="32FF9D1C305CFFB5FE77FACDBE670B20" collectingDate="2001-01-02" collectingDateMax="2001-01-22" collectingDateMin="2001-01-02" collectorName="D. H. &amp; K. M Kavanugh &amp; R. L. Brett &amp; E. Elsom &amp; F. Vargas &amp; R. Ranaivosolo &amp; E. F. Randrianirina &amp; N. Rasoamananana &amp; T. J. Ravelomanana &amp; H. C. Raveloson" country="Madagascar" elevation="900" latitude="-21.25041" location="Talatakely area" longLatPrecision="1" longitude="47.41945" municipality="Ranomafana National Park" pageId="37" pageNumber="238" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="holotype">
<typeStatus id="5D2C29E3305CFFB5FE77FACDBC86086C" box="[386,485,1375,1399]" pageId="37" pageNumber="238" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC4305CFFB5FE02FACDBF1A086C" box="[503,633,1375,1400]" captionStart="FIGURE" captionStartId="38.[155,167,1385,1404]" captionTargetBox="[120,1176,216,1353]" captionTargetId="figure-677@38.[120,1176,216,1372]" captionTargetPageId="38" captionText="FIGURE 21. Digital images of holotype male of Pristacrus ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B Labels. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799489" httpUri="https://zenodo.org/record/13799489/files/figure.png" pageId="37" pageNumber="238">Figs.21A–B</figureCitation>
), a male, in CAS, labeled: “CASENT 1003833”/ “ 
<collectingCountry id="FA80D7D1305CFFB5FF73FA12BC270883" box="[134,324,1408,1432]" name="Madagascar" pageId="37" pageNumber="238">MADAGASCAR</collectingCountry>
, 
<collectingRegion id="405359A3305CFFB5FEA0FA12BF240883" box="[341,583,1408,1432]" country="Madagascar" name="Fianarantsoa" pageId="37" pageNumber="238">Fianarantsoa Province</collectingRegion>
, 
<collectingMunicipality id="624C0D3B305CFFB5FDADFA12BEE30883" box="[600,896,1408,1432]" pageId="37" pageNumber="238">Ranomafana National Park</collectingMunicipality>
, 
<location id="8748C19A305CFFB5FC64FA12B955088C" LSID="urn:lsid:plazi:treatment:0A3E2657305CFFB8FF8EFB4BBCA909E0:8748C19A305CFFB5FC64FA12B955088C" box="[913,1078,1408,1432]" country="Madagascar" latitude="-21.25041" longLatPrecision="1" longitude="47.41945" municipality="Ranomafana National Park" name="Talatakely area" pageId="37" pageNumber="238" stateProvince="Fianarantsoa">Talatakely area</location>
, 
<quantity id="456F3AA4305CFFB5FBB2FA12B9EC088C" box="[1095,1167,1408,1432]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="37" pageNumber="238" unit="m" value="900.0">
<elevation id="09BA7072305CFFB5FBB2FA12B9EC088C" box="[1095,1167,1408,1432]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="37" pageNumber="238" unit="m" value="900.0">900 m</elevation>
</quantity>
, mixed tropical forest, 
<date id="F629B181305CFFB5FE9EFA33BF5708A2" box="[363,564,1441,1465]" pageId="37" pageNumber="238" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">
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</date>
,”/ “ 
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/ 
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, Stop# DHK-01-001, 
<collectorName id="2F62F297305CFFB5FB97FA33BC2808C2" pageId="37" pageNumber="238">D.H. &amp; K.M Kavanugh</collectorName>
, 
<collectorName id="2F62F297305CFFB5FE95FA50BCB108C2" box="[352,466,1474,1497]" pageId="37" pageNumber="238">R.L. Brett</collectorName>
, 
<collectorName id="2F62F297305CFFB5FE1DFA50BF3208C2" box="[488,593,1473,1497]" pageId="37" pageNumber="238">E. Elsom</collectorName>
, 
<collectorName id="2F62F297305CFFB5FD93FA50BFAC08C1" box="[614,719,1474,1498]" pageId="37" pageNumber="238">F. Vargas</collectorName>
, 
<collectorName id="2F62F297305CFFB5FD11FA50BEEC08C2" box="[740,911,1473,1497]" pageId="37" pageNumber="238">R. Ranaivosolo</collectorName>
,”/ “ 
<collectorName id="2F62F297305CFFB5FC31FA50B9EC08C2" box="[964,1167,1473,1497]" pageId="37" pageNumber="238">E.F. Randrianirina</collectorName>
, 
<collectorName id="2F62F297305CFFB5FF8EFA70BC2408E2" box="[123,327,1506,1529]" pageId="37" pageNumber="238">N. Rasoamananana</collectorName>
, 
<collectorName id="2F62F297305CFFB5FEA6FA70BF7E08E1" box="[339,541,1506,1530]" pageId="37" pageNumber="238">T.J. Ravelomanana</collectorName>
, and 
<collectorName id="2F62F297305CFFB5FDA3FA70BF9C08E1" box="[598,767,1506,1530]" pageId="37" pageNumber="238">H.C. Raveloson</collectorName>
collectors”/ “collected at night on surface of rotting logs on forest floor or suspended just above it”/ “ 
<typeStatus id="5D2C29E3305CFFB5FC7FF991B9740B00" box="[906,1047,1539,1563]" pageId="37" pageNumber="238" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC2305CFFB5FBD3F991BC990B20" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B53305CFFB5FBD3F991BC6A0B20" italics="true" pageId="37" pageNumber="238">Pristacrus ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628305CFFB5FDF1F9B6BF230B20" box="[516,576,1572,1595]" pageId="37" pageNumber="238" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label]
</materialsCitation>
. 
<materialsCitation id="32FF9D1C305CFFB5FCE1F9B6BC210B47" collectionCode="CAS" country="Madagascar" pageId="37" pageNumber="238" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<typeStatus id="5D2C29E3305CFFB5FCE1F9B6BE1F0B27" box="[788,892,1572,1596]" pageId="37" pageNumber="238" type="paratype">Paratypes</typeStatus>
(total of 19): 
<specimenCount id="94915CC8305CFFB5FBEEF9B1B9050B20" box="[1051,1126,1571,1595]" count="1" pageId="37" pageNumber="238" type="male">1 male</specimenCount>
and 
<specimenCount id="94915CC8305CFFB5FF8EF9D6BDBA0B47" box="[123,217,1604,1628]" count="1" pageId="37" pageNumber="238" type="female">1 female</specimenCount>
(in 
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)
</materialsCitation>
<materialsCitation id="32FF9D1C305CFFB5FEBCF9D6BF0D0B67" collectionCode="CAS" country="Madagascar" pageId="37" pageNumber="238" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa" typeStatus="paratype">
labelled “CASENT 1003832” or “CASENT 1003831”, respectively, otherwise labeled same as 
<typeStatus id="5D2C29E3305CFFB5FEC7F9F6BCEC0B67" box="[306,399,1636,1660]" pageId="37" pageNumber="238" type="holotype">holotype</typeStatus>
; 
<specimenCount id="94915CC8305CFFB5FE55F9F6BF630B67" box="[416,512,1636,1660]" count="1" pageId="37" pageNumber="238" type="female">1 female</specimenCount>
(in 
<collectionCode id="E4860F84305CFFB5FDC4F9F6BF060B67" box="[561,613,1636,1660]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="37" pageNumber="238" type="Museum">CAS</collectionCode>
)
</materialsCitation>
<materialsCitation id="32FF9D1C305CFFB6FD82F9F6B9F60B0C" collectingDate="2001-01-02" collectingDateMax="2001-01-22" collectingDateMin="2001-01-02" collectionCode="CAS" collectorName="D. H. &amp; K. M Kavanugh &amp; R. L. Brett &amp; E. Elsom &amp; F. Vargas &amp; R. Ranaivosolo &amp; E. F. Randrianirina &amp; N. Rasoamananana &amp; T. J. Ravelomanana &amp; H. C. Raveloson" country="Madagascar" elevation="900" lastPageId="38" lastPageNumber="239" latitude="-21.25041" location="Talatakely area" longLatPrecision="1" longitude="47.41945" municipality="Ranomafana National Park" pageId="37" pageNumber="238" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
labeled “CASENT 1004470”/ “ 
<collectingCountry id="FA80D7D1305CFFB5FC24F9F6B9EC0B67" box="[977,1167,1636,1660]" name="Madagascar" pageId="37" pageNumber="238">MADAGASCAR</collectingCountry>
, 
<collectingRegion id="405359A3305CFFB5FF8EF917BC090B86" box="[123,362,1669,1693]" country="Madagascar" name="Fianarantsoa" pageId="37" pageNumber="238">Fianarantsoa Province</collectingRegion>
, 
<collectingMunicipality id="624C0D3B305CFFB5FE8DF917BFFF0B86" box="[376,668,1669,1693]" pageId="37" pageNumber="238">Ranomafana National Park</collectingMunicipality>
, 
<location id="8748C19A305CFFB5FD5CF917BE2F0B87" LSID="urn:lsid:plazi:treatment:0A3E2657305CFFB8FF8EFB4BBCA909E0:8748C19A305CFFB5FD5CF917BE2F0B87" box="[681,844,1669,1693]" country="Madagascar" latitude="-21.25041" longLatPrecision="1" longitude="47.41945" municipality="Ranomafana National Park" name="Talatakely area" pageId="37" pageNumber="238" stateProvince="Fianarantsoa">Talatakely area</location>
, 
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<elevation id="09BA7072305CFFB5FCAEF917BEC30B87" box="[859,928,1669,1693]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="37" pageNumber="238" unit="m" value="900.0">900 m</elevation>
</quantity>
, mixed tropical forest, 
<date id="F629B181305CFFB5FF8EF934BC5C0BA5" box="[123,319,1702,1726]" pageId="37" pageNumber="238" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">
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</date>
,”/ “ 
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, Stop# DHK-01-001, 
<collectorName id="2F62F297305CFFB5FC95F934B9390BA6" box="[864,1114,1701,1726]" pageId="37" pageNumber="238">D.H. &amp; K.M Kavanugh</collectorName>
, 
<collectorName id="2F62F297305CFFB6FB93F934BDD308CD" lastPageId="38" lastPageNumber="239" pageId="37" pageNumber="238">R.L. Brett</collectorName>
, 
<collectorName id="2F62F297305FFFB6FF35FA2DBC4708CC" box="[192,292,1471,1495]" pageId="38" pageNumber="239">E. Elsom</collectorName>
, 
<collectorName id="2F62F297305FFFB6FEC1FA2DBCFB08CC" box="[308,408,1471,1495]" pageId="38" pageNumber="239">F. Vargas</collectorName>
, 
<collectorName id="2F62F297305FFFB6FE52FA2DBF2D08CC" box="[423,590,1471,1495]" pageId="38" pageNumber="239">R. Ranaivosolo</collectorName>
,”/ “ 
<collectorName id="2F62F297305FFFB6FD88FA2DBE2708CC" box="[637,836,1471,1495]" pageId="38" pageNumber="239">E.F. Randrianirina</collectorName>
, 
<collectorName id="2F62F297305FFFB6FCA6FA2DB94008CD" box="[851,1059,1471,1494]" pageId="38" pageNumber="239">N. Rasoamananana</collectorName>
, 
<collectorName id="2F62F297305FFFB6FBC7FA2DBD8D08EC" pageId="38" pageNumber="239">T.J. Ravelomanana</collectorName>
, and 
<collectorName id="2F62F297305FFFB6FED2FA4DBCB308EC" box="[295,464,1503,1527]" pageId="38" pageNumber="239">H.C. Raveloson</collectorName>
collectors”/ “collected in daytime from large accumulation of dead leaf, twig, and branch debris on and extended up to 
<quantity id="456F3AA4305FFFB6FD31FA6DBF920B0D" box="[708,753,1535,1559]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="38" pageNumber="239" unit="m" value="2.0">
<elevation id="09BA7072305FFFB6FD31FA6DBF920B0D" box="[708,753,1535,1559]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="38" pageNumber="239" unit="m" value="2.0">2 m</elevation>
</quantity>
above forest floor”; 
<specimenCount id="94915CC8305FFFB6FC29FA6DB9440B0C" box="[988,1063,1535,1559]" count="1" pageId="38" pageNumber="239" type="male">1 male</specimenCount>
(in 
<collectionCode id="E4860F84305FFFB6FBACFA6DB9EE0B0C" box="[1113,1165,1535,1559]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="38" pageNumber="239" type="Museum">CAS</collectionCode>
)
</materialsCitation>
<materialsCitation id="32FF9D1C305FFFB7FF8EF98DBF0E0DE8" collectingDate="2001-01-02" collectingDateMax="2001-01-22" collectingDateMin="2001-01-02" collectorName="D. H. &amp; K. M Kavanugh &amp; R. L. Brett &amp; E. Elsom &amp; F. Vargas &amp; R. Ranaivosolo &amp; E. F. Randrianirina &amp; N. Rasoamananana &amp; T. J. Ravelomanana &amp; H. C. Raveloson" country="Madagascar" elevation="900" lastPageId="39" lastPageNumber="240" latitude="-21.25041" location="Talatakely area" longLatPrecision="1" longitude="47.41945" municipality="Ranomafana National Park" pageId="38" pageNumber="239" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
labeled “CASENT 1002838”/ “ 
<collectingCountry id="FA80D7D1305FFFB6FE22F98DBFF60B2C" box="[471,661,1567,1591]" name="Madagascar" pageId="38" pageNumber="239">MADAGASCAR</collectingCountry>
, 
<collectingRegion id="405359A3305FFFB6FD53F98DBEF40B2C" box="[678,919,1567,1591]" country="Madagascar" name="Fianarantsoa" pageId="38" pageNumber="239">Fianarantsoa Province</collectingRegion>
, 
<collectingMunicipality id="624C0D3B305FFFB6FC5DF98DBDCF0B4C" pageId="38" pageNumber="239">Ranomafana National Park</collectingMunicipality>
, 
<location id="8748C19A305FFFB6FF36F9ADBC120B4D" LSID="urn:lsid:plazi:treatment:0A3E2657305CFFB8FF8EFB4BBCA909E0:8748C19A305FFFB6FF36F9ADBC120B4D" box="[195,369,1599,1623]" country="Madagascar" latitude="-21.25041" longLatPrecision="1" longitude="47.41945" municipality="Ranomafana National Park" name="Talatakely area" pageId="38" pageNumber="239" stateProvince="Fianarantsoa">Talatakely area</location>
, 
<quantity id="456F3AA4305FFFB6FE7CF9ADBCB40B4D" box="[393,471,1599,1623]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="38" pageNumber="239" unit="m" value="900.0">
<elevation id="09BA7072305FFFB6FE7CF9ADBCB40B4D" box="[393,471,1599,1623]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="38" pageNumber="239" unit="m" value="900.0">900 m</elevation>
</quantity>
, mixed tropical forest, 
<date id="F629B181305FFFB6FD0EF9ADBEBB0B4C" box="[763,984,1599,1623]" pageId="38" pageNumber="239" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">
<collectingDate id="E66D4869305FFFB6FD0EF9ADBEBB0B4C" box="[763,984,1599,1623]" pageId="38" pageNumber="239" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">2-22 January 2001</collectingDate>
</date>
,”/ “ 
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/ 
<geoCoordinate id="E7A3F186305FFFB6FF8EF9CDBD9A0B6D" box="[123,249,1631,1655]" degrees="47. 41945" direction="east" orientation="longitude" pageId="38" pageNumber="239" precision="1" value="47.41945">47.41945°E</geoCoordinate>
, Stop# DHK-01-001, 
<collectorName id="2F62F297305FFFB6FE08F9CDBE6D0B6C" box="[509,782,1631,1655]" pageId="38" pageNumber="239">D.H. &amp; K.M Kavanugh</collectorName>
, 
<collectorName id="2F62F297305FFFB6FCD7F9CDBEF70B6D" box="[802,916,1631,1654]" pageId="38" pageNumber="239">R.L. Brett</collectorName>
, 
<collectorName id="2F62F297305FFFB6FC5CF9CDB9720B6C" box="[937,1041,1631,1655]" pageId="38" pageNumber="239">E. Elsom</collectorName>
, 
<collectorName id="2F62F297305FFFB6FBD3F9CDB9EC0B6C" box="[1062,1167,1631,1655]" pageId="38" pageNumber="239">F. Vargas</collectorName>
, 
<collectorName id="2F62F297305FFFB6FF8EF9EDBC430B8C" box="[123,288,1663,1687]" pageId="38" pageNumber="239">R. Ranaivosolo</collectorName>
,”/ “ 
<collectorName id="2F62F297305FFFB6FEBBF9EDBF700B8C" box="[334,531,1663,1687]" pageId="38" pageNumber="239">E.F. Randrianirina</collectorName>
, 
<collectorName id="2F62F297305FFFB6FDD4F9EDBF930B8D" box="[545,752,1663,1686]" pageId="38" pageNumber="239">N. Rasoamananana</collectorName>
, 
<collectorName id="2F62F297305FFFB6FD0BF9EDBEA90B8C" box="[766,970,1663,1687]" pageId="38" pageNumber="239">T.J. Ravelomanana</collectorName>
, and 
<collectorName id="2F62F297305FFFB6FBF2F9EDBDC30BAD" pageId="38" pageNumber="239">H.C. Raveloson</collectorName>
collectors”/ “collected in daytime under loss bark of rotting log logs on forest floor or suspended just above it”; 
<specimenCount id="94915CC8305EFFB7FE98FF49BCD50DE8" box="[365,438,219,243]" count="1" pageId="39" pageNumber="240" type="male">1 male</specimenCount>
and 
<specimenCount id="94915CC8305EFFB7FE1EFF49BF0E0DE8" box="[491,621,219,243]" count="2" pageId="39" pageNumber="240" type="female">two females</specimenCount>
</materialsCitation>
</paragraph>
<caption id="D6E8C7C9305FFFB6FF6EFAFBBC80088D" ID-DOI="http://doi.org/10.5281/zenodo.13799489" ID-Zenodo-Dep="13799489" httpUri="https://zenodo.org/record/13799489/files/figure.png" pageId="38" pageNumber="239" startId="38.[155,167,1385,1404]" targetBox="[120,1176,216,1353]" targetPageId="38" targetType="figure">
<paragraph id="82289741305FFFB6FF6EFAFBBC80088D" blockId="38.[123,1173,1385,1430]" pageId="38" pageNumber="239">
<smallCapsWord id="84CE019D305FFFB6FF6EFAFBBDBB0861" baselines="1399,1398" box="[155,216,1385,1404]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="38" pageNumber="239">FIGURE</smallCapsWord>
21. Digital images of holotype male of 
<taxonomicName id="4597ECC2305FFFB6FDC4FAFBBEDE0867" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[561,957,1385,1404]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="239" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B53305FFFB6FDC4FAFBBE620867" box="[561,769,1385,1404]" italics="true" pageId="38" pageNumber="239">Pristacrus ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628305FFFB6FC36FAFBB9660867" box="[963,1029,1385,1404]" pageId="38" pageNumber="239" rank="species">sp. nov.</taxonomicNameLabel>
A. Habitus, dorsal aspect; B Labels. Scale line = 1.0 mm.
</paragraph>
</caption>
<paragraph id="82289741305EFFB7FF8EFF69BF0E0C08" blockId="39.[123,1174,219,1715]" box="[123,621,251,275]" pageId="39" pageNumber="240">
(in 
<collectionCode id="E4860F84305EFFB7FF45FF69BD860C08" box="[176,229,251,275]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="39" pageNumber="240" type="Museum">CAS</collectionCode>
and 
<collectionCode id="E4860F84305EFFB7FECCFF69BC120C09" box="[313,369,251,274]" pageId="39" pageNumber="240">MZF</collectionCode>
) labeled “CASENT
</paragraph>
<paragraph id="82289741305EFFB7FF8EFE89BF0D0C28" blockId="39.[123,1174,219,1715]" box="[123,622,283,307]" pageId="39" pageNumber="240">1003752”, “CASENT 1003751” or “CASENT</paragraph>
<paragraph id="82289741305EFFB7FF8EFEA9BF0D0C49" blockId="39.[123,1174,219,1715]" box="[123,622,315,339]" pageId="39" pageNumber="240">
1003753”, respectively/ “ 
<collectingCountry id="FA80D7D1305EFFB7FE53FEA9BF050C48" box="[422,614,315,339]" name="Madagascar" pageId="39" pageNumber="240">MADAGASCAR</collectingCountry>
,
</paragraph>
<paragraph id="82289741305EFFB7FF8EFEC9BF0D0C68" blockId="39.[123,1174,219,1715]" box="[123,622,347,371]" pageId="39" pageNumber="240">
<materialsCitation id="32FF9D1C305EFFB7FF8EFEC9BF0D0C68" box="[123,622,347,371]" location="Ranomafana" pageId="39" pageNumber="240" specimenCount="1" stateProvince="Fianarantsoa">
<collectingRegion id="405359A3305EFFB7FF8EFEC9BC0E0C68" box="[123,365,347,371]" country="Madagascar" name="Fianarantsoa" pageId="39" pageNumber="240">Fianarantsoa Province</collectingRegion>
, 
<location id="8748C19A305EFFB7FE8AFEC9BF650C68" LSID="urn:lsid:plazi:treatment:0A3E2657305CFFB8FF8EFB4BBCA909E0:8748C19A305EFFB7FE8AFEC9BF650C68" box="[383,518,347,371]" name="Ranomafana" pageId="39" pageNumber="240" stateProvince="Fianarantsoa">Ranomafana</location>
National
</materialsCitation>
</paragraph>
<paragraph id="82289741305EFFB7FF8EFEE9BF0E0CA8" blockId="39.[123,1174,219,1715]" pageId="39" pageNumber="240">
<materialsCitation id="32FF9D1C305EFFB7FF8EFEE9BF050CA8" collectingDate="2001-01-02" collectingDateMax="2001-01-22" collectingDateMin="2001-01-02" collectingRegion="Fianarantsoa Province" elevation="900" latitude="-21.25041" location="Park" municipality="Talatakely area" pageId="39" pageNumber="240" specimenCount="1" stateProvince="Fianarantsoa Province">
<location id="8748C19A305EFFB7FF8EFEE9BDCF0C88" LSID="urn:lsid:plazi:treatment:0A3E2657305CFFB8FF8EFB4BBCA909E0:8748C19A305EFFB7FF8EFEE9BDCF0C88" box="[123,172,379,403]" latitude="-21.25041" municipality="Talatakely area" name="Park" pageId="39" pageNumber="240" stateProvince="Fianarantsoa Province">Park</location>
, 
<collectingMunicipality id="624C0D3B305EFFB7FF48FEE9BC000C89" box="[189,355,379,403]" pageId="39" pageNumber="240">Talatakely area</collectingMunicipality>
, 
<quantity id="456F3AA4305EFFB7FE80FEE9BCDE0C89" box="[373,445,379,403]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="240" unit="m" value="900.0">
<elevation id="09BA7072305EFFB7FE80FEE9BCDE0C89" box="[373,445,379,403]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="240" unit="m" value="900.0">900 m</elevation>
</quantity>
, mixed tropical forest, 
<date id="F629B181305EFFB7FF25FE09BCCD0CA8" box="[208,430,411,435]" pageId="39" pageNumber="240" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">
<collectingDate id="E66D4869305EFFB7FF25FE09BCCD0CA8" box="[208,430,411,435]" pageId="39" pageNumber="240" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">2-22 January 2001</collectingDate>
</date>
,”/ “ 
<geoCoordinate id="E7A3F186305EFFB7FE13FE09BF050CA8" box="[486,614,411,435]" degrees="21. 25041" direction="south" orientation="latitude" pageId="39" pageNumber="240" precision="1" value="-21.25041">21.25041°S</geoCoordinate>
</materialsCitation>
/
</paragraph>
<paragraph id="82289741305EFFB7FF8EFE29BF0D0CC9" blockId="39.[123,1174,219,1715]" box="[123,622,443,467]" pageId="39" pageNumber="240">
<geoCoordinate id="E7A3F186305EFFB7FF8EFE29BD9A0CC9" box="[123,249,443,467]" degrees="47. 41945" direction="east" orientation="longitude" pageId="39" pageNumber="240" precision="1" value="47.41945">47.41945°E</geoCoordinate>
, Stop# DHK-01-001, D.H. &amp; K.M
</paragraph>
<paragraph id="82289741305EFFB7FF8EFE49BF0D0CE9" blockId="39.[123,1174,219,1715]" box="[123,622,475,499]" pageId="39" pageNumber="240">Kavanugh, R.L. Brett, E. Elsom, F. Vargas,</paragraph>
<paragraph id="82289741305EFFB7FF8EFE69BF0D0F09" blockId="39.[123,1174,219,1715]" box="[123,622,507,531]" pageId="39" pageNumber="240">R. Ranaivosolo,”/ “E.F. Randrianirina,</paragraph>
<paragraph id="82289741305EFFB7FF8EFD89BF0D0F28" blockId="39.[123,1174,219,1715]" box="[123,622,539,563]" pageId="39" pageNumber="240">N. Rasoamananana, T.J. Ravelomanana, and</paragraph>
<paragraph id="82289741305EFFB7FF8EFDA9BF0D0F88" blockId="39.[123,1174,219,1715]" pageId="39" pageNumber="240">
H.C. Raveloson collectors”/ “collected in daytime on underside of rotting log logs on forest floor or suspended just above it”; 
<specimenCount id="94915CC8305EFFB7FE0FFDE9BF250F88" box="[506,582,635,659]" count="1" pageId="39" pageNumber="240" type="male">1 male</specimenCount>
(in
</paragraph>
<paragraph id="82289741305EFFB7FF8EFD09BE6B0E18" blockId="39.[123,1174,219,1715]" pageId="39" pageNumber="240">
<materialsCitation id="32FF9D1C305EFFB7FF8EFD09BE600E18" collectionCode="CAS" collectorName="Rainio &amp; A. Left &amp; B. Dorsal &amp; Scale" county="Digital" elevation="900" location="Talatakely area" municipality="National Park" pageId="39" pageNumber="240" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<collectionCode id="E4860F84305EFFB7FF8EFD09BDCC0FA8" box="[123,175,667,691]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="39" pageNumber="240" type="Museum">CAS</collectionCode>
) labeled “CASENT 1004313”/ “MADA- 
<smallCapsWord id="84CE019D305EFFB7FD5EFD31BF8B0FAE" baselines="689,689" box="[683,744,675,694]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="39" pageNumber="240">FIGURE</smallCapsWord>
22. 
<collectingCounty id="6B49EFCD305EFFB7FCE3FD31BE320FAD" box="[790,849,675,694]" pageId="39" pageNumber="240">Digital</collectingCounty>
images of aedeagus of genitalia of GASCAR, 
<collectingRegion id="405359A3305EFFB7FF0BFD29BC960FC8" box="[254,501,699,723]" country="Madagascar" name="Fianarantsoa" pageId="39" pageNumber="240">Fianarantsoa Province</collectingRegion>
, Ranoma- male 
<typeStatus id="5D2C29E3305EFFB7FD35FD2EBE680FD4" box="[704,779,700,719]" pageId="39" pageNumber="240" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4597ECC2305EFFB7FCC0FD2FB91A0FD4" authority="Kavanaugh" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[821,1145,700,720]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53305EFFB7FCC0FD2FB96F0FCB" box="[821,1036,701,720]" italics="true" pageId="39" pageNumber="240">Pristacrus ranomafanae</emphasis>
Kavanaugh
</taxonomicName>
&amp; fana 
<collectingMunicipality id="624C0D3B305EFFB7FF40FD49BC2C0FE8" box="[181,335,731,755]" pageId="39" pageNumber="240">National Park</collectingMunicipality>
, 
<location id="8748C19A305EFFB7FE97FD49BF6A0FE9" LSID="urn:lsid:plazi:treatment:0A3E2657305CFFB8FF8EFB4BBCA909E0:8748C19A305EFFB7FE97FD49BF6A0FE9" box="[354,521,731,755]" county="Digital" municipality="National Park" name="Talatakely area" pageId="39" pageNumber="240" stateProvince="Fianarantsoa">Talatakely area</location>
, 
<quantity id="456F3AA4305EFFB7FDE8FD49BF040FE9" box="[541,615,731,755]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="240" unit="m" value="900.0">
<elevation id="09BA7072305EFFB7FDE8FD49BF040FE9" box="[541,615,731,755]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="240" unit="m" value="900.0">900 m</elevation>
</quantity>
, 
<collectorName id="2F62F297305EFFB7FD7EFD44BFA70FF2" box="[651,708,726,745]" pageId="39" pageNumber="240">Rainio</collectorName>
<taxonomicNameLabel id="ABD0F628305EFFB7FD3CFD45BE680FF1" box="[713,779,727,746]" pageId="39" pageNumber="240" rank="species">sp. nov.</taxonomicNameLabel>
<collectorName id="2F62F297305EFFB7FCE5FD44BE2D0FF2" box="[784,846,726,746]" pageId="39" pageNumber="240">A. Left</collectorName>
lateral aspect; 
<collectorName id="2F62F297305EFFB7FC3AFD45B9420FF2" box="[975,1057,726,746]" pageId="39" pageNumber="240">B. Dorsal</collectorName>
aspect. 
<collectorName id="2F62F297305EFFB7FB92FD44B9F60FF2" box="[1127,1173,726,745]" pageId="39" pageNumber="240">Scale</collectorName>
line = 
<quantity id="456F3AA4305EFFB7FD34FD62BE600E18" box="[705,771,752,771]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="39" pageNumber="240" unit="mm" value="0.5">0.5 mm</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="82289741305EFFB7FF8EFD69BF0E0E08" blockId="39.[123,1174,219,1715]" box="[123,621,763,787]" pageId="39" pageNumber="240">
mixed tropical forest, 
<date id="F629B181305EFFB7FE88FD69BF300E08" box="[381,595,763,787]" pageId="39" pageNumber="240" value="2001-01-02" valueMax="2001-01-22" valueMin="2001-01-02">2-22 January 2001</date>
,”/
</paragraph>
<paragraph id="82289741305EFFB7FF8EFC89BE4108A8" blockId="39.[123,1174,219,1715]" pageId="39" pageNumber="240">
“ 
<geoCoordinate id="E7A3F186305EFFB7FF72FC89BC6A0E28" box="[135,265,795,819]" degrees="21. 25041" direction="south" orientation="latitude" pageId="39" pageNumber="240" precision="1" value="-21.25041">21.25041°S</geoCoordinate>
/ 
<geoCoordinate id="E7A3F186305EFFB7FED3FC89BCCA0E29" box="[294,425,795,819]" degrees="47. 41945" direction="east" orientation="longitude" pageId="39" pageNumber="240" precision="1" value="47.41945">47.41945°E</geoCoordinate>
, Stop# DHK-01-001, D.H. &amp; K.M Kavanugh, R.L. Brett, E. Elsom, F. Vargas, R. Ranaivosolo,”/ “E.F. Randrianirina, N. Rasoamananana, T.J. Ravelomanana, and H.C. Raveloson collectors”/ “collected in daytime from leaf litter on forest floor”; 
<materialsCitation id="32FF9D1C305EFFB7FF8EFCE9BC460EE8" collectingDate="1998-04-27" collectionCode="CAS" collectorName="A D. H. Kavanaugh" country="Madagascar" elevation="900" latitude="-21.255" location="Talatakely area" longLatPrecision="126" longitude="47.431667" municipality="Ranomafana National Park" pageId="39" pageNumber="240" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa">
<specimenCount id="94915CC8305EFFB7FF8EFCE9BDBC0E88" box="[123,223,891,915]" count="1" pageId="39" pageNumber="240" type="female">1 female</specimenCount>
(in 
<collectionCode id="E4860F84305EFFB7FEEDFCE9BC2F0E88" box="[280,332,891,915]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="39" pageNumber="240" type="Museum">CAS</collectionCode>
) labeled “CASENT 1051144”/ “ 
<collectingCountry id="FA80D7D1305EFFB7FD3DFCE9BEE50E88" box="[712,902,891,915]" name="Madagascar" pageId="39" pageNumber="240">MADAGASCAR</collectingCountry>
, 
<collectingRegion id="405359A3305EFFB7FC6FFCE9B9EC0E88" box="[922,1167,891,915]" country="Madagascar" name="Fianarantsoa" pageId="39" pageNumber="240">Fianarantsoa Province</collectingRegion>
, 
<collectingMunicipality id="624C0D3B305EFFB7FF8EFC09BCC80EA8" box="[123,427,923,947]" pageId="39" pageNumber="240">Ranomafana National Park</collectingMunicipality>
, 
<location id="8748C19A305EFFB7FE4AFC09BF0B0EA9" LSID="urn:lsid:plazi:treatment:0A3E2657305CFFB8FF8EFB4BBCA909E0:8748C19A305EFFB7FE4AFC09BF0B0EA9" box="[447,616,923,947]" country="Madagascar" latitude="-21.255" longLatPrecision="126" longitude="47.431667" municipality="Ranomafana National Park" name="Talatakely area" pageId="39" pageNumber="240" stateProvince="Fianarantsoa">Talatakely area</location>
, 
<quantity id="456F3AA4305EFFB7FD88FC09BFAB0EA9" box="[637,712,923,947]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="240" unit="m" value="900.0">
<elevation id="09BA7072305EFFB7FD88FC09BFAB0EA9" box="[637,712,923,947]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="240" unit="m" value="900.0">900 m</elevation>
</quantity>
, mixed tropical forest, 
<date id="F629B181305EFFB7FC2AFC09B9E10EA8" box="[991,1154,923,947]" pageId="39" pageNumber="240" value="1998-04-27">
<collectingDate id="E66D4869305EFFB7FC2AFC09B9E10EA8" box="[991,1154,923,947]" pageId="39" pageNumber="240" value="1998-04-27">27 April 1998</collectingDate>
</date>
”/ “ 
<geoCoordinate id="E7A3F186305EFFB7FF72FC29BD910EC8" box="[135,242,955,979]" degrees="21" direction="south" minutes="15. 3" orientation="latitude" pageId="39" pageNumber="240" precision="92" value="-21.255">21°15.3’S</geoCoordinate>
<geoCoordinate id="E7A3F186305EFFB7FF0CFC29BC050EC9" box="[249,358,955,979]" degrees="47" direction="east" minutes="25. 9" orientation="longitude" pageId="39" pageNumber="240" precision="92" value="47.431667">47°25.9’E</geoCoordinate>
, Stop # 98-85 
<collectorName id="2F62F297305EFFB7FE09FC29BFAB0EC8" box="[508,712,955,979]" pageId="39" pageNumber="240">A D.H. Kavanaugh</collectorName>
collector collected in and under rotten logs on forest floor”
</materialsCitation>
; 
<materialsCitation id="32FF9D1C305EFFB7FEC3FC49BF7B0948" collectingDate="2002-04-28" collectingDateMax="2002-05-05" collectingDateMin="2002-04-28" collectionCode="CAS" collectorName="R. Harin'Hala California Acad" country="Madagascar" county="Province" elevation="1020" latitude="-21.2665" location="Belle Vue" longLatPrecision="12" longitude="47.420166" municipality="Ranomafana" pageId="39" pageNumber="240" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa">
<specimenCount id="94915CC8305EFFB7FEC3FC49BCF40EE8" box="[310,407,987,1011]" count="1" pageId="39" pageNumber="240" type="female">1 female</specimenCount>
(in 
<collectionCode id="E4860F84305EFFB7FE32FC49BC980EE8" box="[455,507,987,1011]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="39" pageNumber="240" type="Museum">CAS</collectionCode>
) labeled “CASENT 8068264”/ “ 
<collectingCountry id="FA80D7D1305EFFB7FC9DFC49B9450EE8" box="[872,1062,987,1011]" name="Madagascar" pageId="39" pageNumber="240">MADAGASCAR</collectingCountry>
: 
<collectingCounty id="6B49EFCD305EFFB7FBC2FC49B9F50EE8" box="[1079,1174,987,1011]" pageId="39" pageNumber="240">Province</collectingCounty>
<collectingRegion id="405359A3305EFFB7FF8EFC69BC600908" box="[123,259,1019,1043]" country="Madagascar" name="Fianarantsoa" pageId="39" pageNumber="240">Fianarantsoa</collectingRegion>
, Parc National 
<collectingMunicipality id="624C0D3B305EFFB7FE58FC69BF570908" box="[429,564,1019,1043]" pageId="39" pageNumber="240">Ranomafana</collectingMunicipality>
, 
<location id="8748C19A305EFFB7FDB7FC69BFCE0909" LSID="urn:lsid:plazi:treatment:0A3E2657305CFFB8FF8EFB4BBCA909E0:8748C19A305EFFB7FDB7FC69BFCE0909" box="[578,685,1019,1043]" country="Madagascar" county="Province" latitude="-21.2665" longLatPrecision="12" longitude="47.420166" municipality="Ranomafana" name="Belle Vue" pageId="39" pageNumber="240" stateProvince="Fianarantsoa">Belle Vue</location>
at 
<location id="8748C19A305EFFB7FD3AFC69BE5F0908" LSID="urn:lsid:plazi:treatment:0A3E2657305CFFB8FF8EFB4BBCA909E0:8748C19A305EFFB7FD3AFC69BE5F0908" box="[719,828,1019,1043]" country="Madagascar" county="Province" latitude="-21.2665" longLatPrecision="12" longitude="47.420166" municipality="Ranomafana" name="Talatakely" pageId="39" pageNumber="240" stateProvince="Fianarantsoa">Talatakely</location>
, elev 
<quantity id="456F3AA4305EFFB7FC8AFC69BEB20909" box="[895,977,1019,1043]" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="39" pageNumber="240" unit="m" value="1020.0">
<elevation id="09BA7072305EFFB7FC8AFC69BEB20909" box="[895,977,1019,1043]" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="39" pageNumber="240" unit="m" value="1020.0">1020 m</elevation>
</quantity>
<date id="F629B181305EFFB7FC2CFC69BDD30928" pageId="39" pageNumber="240" value="2002-04-28" valueMax="2002-05-05" valueMin="2002-04-28">
<collectingDate id="E66D4869305EFFB7FC2CFC69BDD30928" pageId="39" pageNumber="240" value="2002-04-28" valueMax="2002-05-05" valueMin="2002-04-28">28 April – 5 May 2002</collectingDate>
</date>
”/ “ 
<geoCoordinate id="E7A3F186305EFFB7FF23FB89BC2C0928" box="[214,335,1051,1075]" degrees="21" direction="south" minutes="15. 99" orientation="latitude" pageId="39" pageNumber="240" precision="9" value="-21.2665">21°15.99’S</geoCoordinate>
<geoCoordinate id="E7A3F186305EFFB7FEA3FB89BCB30929" box="[342,464,1051,1075]" degrees="47" direction="east" minutes="25. 21" orientation="longitude" pageId="39" pageNumber="240" precision="9" value="47.420166">47°25.21’E</geoCoordinate>
collector: 
<collectorName id="2F62F297305EFFB7FDB6FB89BEE90928" box="[579,906,1051,1075]" pageId="39" pageNumber="240">R. Harin’Hala California Acad</collectorName>
of Sciences malaise secondary tropical forest MA-02-09C-27”
</materialsCitation>
; 
<materialsCitation id="32FF9D1C305EFFB7FDD0FBA9BF9A09A8" collectingDate="2001-11-15" collectingDateMax="2001-11-22" collectingDateMin="2001-11-15" collectionCode="CAS" collectorName="R. Harin'Hala California Acad" country="Madagascar" county="Province" elevation="1020" latitude="-21.2665" location="Belle Vue" longLatPrecision="12" longitude="47.420166" municipality="Ranomafana" pageId="39" pageNumber="240" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa">
<specimenCount id="94915CC8305EFFB7FDD0FBA9BFE20948" box="[549,641,1083,1107]" count="1" pageId="39" pageNumber="240" type="female">1 female</specimenCount>
(in 
<collectionCode id="E4860F84305EFFB7FD5FFBA9BFBD0948" box="[682,734,1083,1107]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="39" pageNumber="240" type="Museum">CAS</collectionCode>
) labeled “CASENT 8068264”/ “ 
<collectingCountry id="FA80D7D1305EFFB7FBCEFBA9BD840968" name="Madagascar" pageId="39" pageNumber="240">MADAGASCAR</collectingCountry>
: 
<collectingCounty id="6B49EFCD305EFFB7FF03FBC9BC360968" box="[246,341,1115,1139]" pageId="39" pageNumber="240">Province</collectingCounty>
<collectingRegion id="405359A3305EFFB7FEA8FBC9BC860968" box="[349,485,1115,1139]" country="Madagascar" name="Fianarantsoa" pageId="39" pageNumber="240">Fianarantsoa</collectingRegion>
, Parc National 
<collectingMunicipality id="624C0D3B305EFFB7FD7BFBC9BE760968" box="[654,789,1115,1139]" pageId="39" pageNumber="240">Ranomafana</collectingMunicipality>
, 
<location id="8748C19A305EFFB7FCD1FBC9BEED0969" LSID="urn:lsid:plazi:treatment:0A3E2657305CFFB8FF8EFB4BBCA909E0:8748C19A305EFFB7FCD1FBC9BEED0969" box="[804,910,1115,1139]" country="Madagascar" county="Province" latitude="-21.2665" longLatPrecision="12" longitude="47.420166" municipality="Ranomafana" name="Belle Vue" pageId="39" pageNumber="240" stateProvince="Fianarantsoa">Belle Vue</location>
at 
<location id="8748C19A305EFFB7FC45FBC9B97E0968" LSID="urn:lsid:plazi:treatment:0A3E2657305CFFB8FF8EFB4BBCA909E0:8748C19A305EFFB7FC45FBC9B97E0968" box="[944,1053,1115,1139]" country="Madagascar" county="Province" latitude="-21.2665" longLatPrecision="12" longitude="47.420166" municipality="Ranomafana" name="Talatakely" pageId="39" pageNumber="240" stateProvince="Fianarantsoa">Talatakely</location>
, elev 
<quantity id="456F3AA4305EFFB7FB95FBC9BDF30989" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="39" pageNumber="240" unit="m" value="1020.0">
<elevation id="09BA7072305EFFB7FB95FBC9BDF30989" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="39" pageNumber="240" unit="m" value="1020.0">1020 m</elevation>
</quantity>
<date id="F629B181305EFFB7FF6DFBE9BCED0988" box="[152,398,1147,1171]" pageId="39" pageNumber="240" value="2001-11-15" valueMax="2001-11-22" valueMin="2001-11-15">
<collectingDate id="E66D4869305EFFB7FF6DFBE9BCED0988" box="[152,398,1147,1171]" pageId="39" pageNumber="240" value="2001-11-15" valueMax="2001-11-22" valueMin="2001-11-15">15-22 November 2001</collectingDate>
</date>
”/ “ 
<geoCoordinate id="E7A3F186305EFFB7FE43FBE9BF4C0988" box="[438,559,1147,1171]" degrees="21" direction="south" minutes="15. 99" orientation="latitude" pageId="39" pageNumber="240" precision="9" value="-21.2665">21°15.99’S</geoCoordinate>
<geoCoordinate id="E7A3F186305EFFB7FDCDFBE9BFD10989" box="[568,690,1147,1171]" degrees="47" direction="east" minutes="25. 21" orientation="longitude" pageId="39" pageNumber="240" precision="9" value="47.420166">47°25.21’E</geoCoordinate>
collector: 
<collectorName id="2F62F297305EFFB7FCDCFBE9B9150988" box="[809,1142,1147,1171]" pageId="39" pageNumber="240">R. Harin’Hala California Acad</collectorName>
of Sciences malaise secondary tropical forest MA-02-09C-03”
</materialsCitation>
; 
<materialsCitation id="32FF9D1C305EFFB7FCF2FB09BF8C09E8" collectingDate="1990-02-01" collectingDateMax="1990-02-09" collectingDateMin="1990-02-01" collectionCode="NMNH" collectorName="W. E. Steiner" country="Madagascar" elevation="900" location="Prov." pageId="39" pageNumber="240" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa">
<specimenCount id="94915CC8305EFFB7FCF2FB09BE2C09A8" box="[775,847,1179,1203]" count="1" pageId="39" pageNumber="240" type="male">1 male</specimenCount>
(in 
<collectionCode id="E4860F84305EFFB7FC8FFB09BEA809A9" box="[890,971,1179,1202]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="39" pageNumber="240" type="Museum">NMNH</collectionCode>
) labeled “ 
<collectingCountry id="FA80D7D1305EFFB7FBCEFB09BD8409C8" name="Madagascar" pageId="39" pageNumber="240">MADAGASCAR</collectingCountry>
: 
<location id="8748C19A305EFFB7FF00FB29BC4E09C9" LSID="urn:lsid:plazi:treatment:0A3E2657305CFFB8FF8EFB4BBCA909E0:8748C19A305EFFB7FF00FB29BC4E09C9" box="[245,301,1211,1234]" country="Madagascar" name="Prov." pageId="39" pageNumber="240" stateProvince="Fianarantsoa">Prov.</location>
<collectingRegion id="405359A3305EFFB7FEC6FB29BCD809C8" box="[307,443,1211,1235]" country="Madagascar" name="Fianarantsoa" pageId="39" pageNumber="240">Fianarantsoa</collectingRegion>
, 
<locationDeviation id="939A8E63305EFFB7FE3CFB29BFC809C8" box="[457,683,1211,1235]" location="Ranomafana" pageId="39" pageNumber="240">
<quantity id="456F3AA4305EFFB7FE3CFB29BC9C09C8" box="[457,511,1211,1235]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="39" pageNumber="240" unit="km" value="7.0">7 km</quantity>
W Ranomafana
</locationDeviation>
, 
<quantity id="456F3AA4305EFFB7FD4DFB29BF9F09C9" box="[696,764,1211,1235]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="240" unit="m" value="900.0">
<elevation id="09BA7072305EFFB7FD4DFB29BF9F09C9" box="[696,764,1211,1235]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="240" unit="m" value="900.0">900 m</elevation>
</quantity>
<date id="F629B181305EFFB7FCF7FB29BEAA09C8" box="[770,969,1211,1235]" pageId="39" pageNumber="240" value="1990-02-01" valueMax="1990-02-09" valueMin="1990-02-01">
<collectingDate id="E66D4869305EFFB7FCF7FB29BEAA09C8" box="[770,969,1211,1235]" pageId="39" pageNumber="240" value="1990-02-01" valueMax="1990-02-09" valueMin="1990-02-01">1-9 February 1990</collectingDate>
</date>
<collectorName id="2F62F297305EFFB7FC3AFB29B93E09C8" box="[975,1117,1211,1235]" pageId="39" pageNumber="240">W. E. Steiner</collectorName>
”/ “at black light in montane rainforest near river and stream”
</materialsCitation>
; 
<materialsCitation id="32FF9D1C305EFFB7FCF7FB49BEC00828" collectionCode="CAS" collectorName="Col. K. Will" country="Madagascar" elevation="900" latitude="-21.25041" location="Fianarantsoa" longLatPrecision="1" longitude="47.41945" pageId="39" pageNumber="240" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa">
<specimenCount id="94915CC8305EFFB7FCF7FB49BE2D09E8" box="[770,846,1243,1267]" count="1" pageId="39" pageNumber="240" type="male">1 male</specimenCount>
(in 
<collectionCode id="E4860F84305EFFB7FC77FB49BED509E8" box="[898,950,1243,1267]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="39" pageNumber="240" type="Museum">CAS</collectionCode>
) labeled “CASENT 8005762”/ “ 
<geoCoordinate id="E7A3F186305EFFB7FF0AFB69BC1E0808" box="[255,381,1275,1299]" degrees="21. 25041" direction="south" orientation="latitude" pageId="39" pageNumber="240" precision="1" value="-21.25041">21.25041°S</geoCoordinate>
/ 
<geoCoordinate id="E7A3F186305EFFB7FE79FB69BF680809" box="[396,523,1275,1299]" degrees="47. 41945" direction="east" orientation="longitude" pageId="39" pageNumber="240" precision="1" value="47.41945">47.41945°E</geoCoordinate>
<collectingCountry id="FA80D7D1305EFFB7FDE6FB69BFB20808" box="[531,721,1275,1299]" name="Madagascar" pageId="39" pageNumber="240">MADAGASCAR</collectingCountry>
, 
<collectingRegion id="405359A3305EFFB7FD15FB69BE0B0808" box="[736,872,1275,1299]" country="Madagascar" name="Fianarantsoa" pageId="39" pageNumber="240">Fianarantsoa</collectingRegion>
Prov Ranomafana National Park, Talatakely area, 
<quantity id="456F3AA4305EFFB7FE9CFA89BCC50829" box="[361,422,1307,1331]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="240" unit="m" value="900.0">
<elevation id="09BA7072305EFFB7FE9CFA89BCC50829" box="[361,422,1307,1331]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="240" unit="m" value="900.0">900m</elevation>
</quantity>
, 10:1:2001 
<collectorName id="2F62F297305EFFB7FDD3FA89BFCA0828" box="[550,681,1307,1331]" pageId="39" pageNumber="240">Col. K. Will</collectorName>
headlamp search logs”
</materialsCitation>
; 
<materialsCitation id="32FF9D1C305EFFB7FC44FA89BE7808A8" collectionCode="EMEC, MNHN" collectorName="Col. K. Will &amp; All" country="Madagascar" elevation="900" latitude="-21.25041" location="Fianarantsoa" longLatPrecision="1" longitude="47.41945" pageId="39" pageNumber="240" specimenCount="6" specimenCount-female="1" specimenCount-male="5" stateProvince="Fianarantsoa" typeStatus="paratype">
<specimenCount id="94915CC8305EFFB7FC44FA89B9670828" box="[945,1028,1307,1331]" count="5" pageId="39" pageNumber="240" type="male">5 males</specimenCount>
and 
<specimenCount id="94915CC8305EFFB7FBCDFA89B9F50828" box="[1080,1174,1307,1331]" count="1" pageId="39" pageNumber="240" type="female">1 female</specimenCount>
(in 
<collectionCode id="E4860F84305EFFB7FF6BFAA9BD8B0848" box="[158,232,1339,1363]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="39" pageNumber="240" type="University or college">EMEC</collectionCode>
and 
<collectionCode id="E4860F84305EFFB7FEEFFAA9BC080849" box="[282,363,1339,1362]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="39" pageNumber="240">MNHN</collectionCode>
) labeled “CASENT 8005762”/ “ 
<geoCoordinate id="E7A3F186305EFFB7FD32FAA9BE260848" box="[711,837,1339,1363]" degrees="21. 25041" direction="south" orientation="latitude" pageId="39" pageNumber="240" precision="1" value="-21.25041">21.25041°S</geoCoordinate>
/ 
<geoCoordinate id="E7A3F186305EFFB7FCB9FAA9BEA80849" box="[844,971,1339,1363]" degrees="47. 41945" direction="east" orientation="longitude" pageId="39" pageNumber="240" precision="1" value="47.41945">47.41945°E</geoCoordinate>
<collectingCountry id="FA80D7D1305EFFB7FC24FAA9B9EC0848" box="[977,1167,1339,1363]" name="Madagascar" pageId="39" pageNumber="240">MADAGASCAR</collectingCountry>
, 
<collectingRegion id="405359A3305EFFB7FF8EFAC9BC600868" box="[123,259,1371,1395]" country="Madagascar" name="Fianarantsoa" pageId="39" pageNumber="240">Fianarantsoa</collectingRegion>
Prov Ranomafana National Park, Talatakely area, 
<quantity id="456F3AA4305EFFB7FCECFAC9BE350869" box="[793,854,1371,1395]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="240" unit="m" value="900.0">
<elevation id="09BA7072305EFFB7FCECFAC9BE350869" box="[793,854,1371,1395]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="240" unit="m" value="900.0">900m</elevation>
</quantity>
, 10:1:2001 
<collectorName id="2F62F297305EFFB7FC21FAC9B9370868" box="[980,1108,1371,1395]" pageId="39" pageNumber="240">Col. K. Will</collectorName>
headlamp search logs”. 
<collectorName id="2F62F297305EFFB7FE90FAE9BCE40888" box="[357,391,1403,1427]" pageId="39" pageNumber="240">All</collectorName>
<typeStatus id="5D2C29E3305EFFB7FE6CFAEEBF63088F" box="[409,512,1404,1428]" pageId="39" pageNumber="240" type="paratype">paratypes</typeStatus>
also bear the following label: “ 
<typeStatus id="5D2C29E3305EFFB7FC7AFAE9B9770888" box="[911,1044,1403,1427]" pageId="39" pageNumber="240" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4597ECC2305EFFB7FBD0FAEEBC9708A8" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B53305EFFB7FBD0FAEEBC6A08A8" italics="true" pageId="39" pageNumber="240">Pristacrus ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F628305EFFB7FE0EFA0EBF5508A8" box="[507,566,1436,1459]" pageId="39" pageNumber="240" rank="species">sp. n.</taxonomicNameLabel>
2015” [yellow label]
</materialsCitation>
.
</paragraph>
<paragraph id="82289741305EFFB7FF5EFA29BECA0B09" blockId="39.[123,1174,219,1715]" pageId="39" pageNumber="240">
<materialsCitation id="32FF9D1C305EFFB7FF5EFA29BEC10B08" country="Madagascar" location="Ranomafana National Park" pageId="39" pageNumber="240" specimenCount="2" stateProvince="Fianarantsoa" typeStatus="holotype">
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<typeStatus id="5D2C29E3305EFFB7FF5EFA29BD8708C9" box="[171,228,1467,1491]" pageId="39" pageNumber="240">TYPE</typeStatus>
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<smallCapsWord id="84CE019D305EFFB7FF19FA2CBC3408CA" baselines="1484" box="[236,343,1470,1489]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="39" pageNumber="240">LOCALITY</smallCapsWord>
</emphasis>
.— 
<collectingCountry id="FA80D7D1305EFFB7FE8BFA29BC9C08C8" box="[382,511,1467,1491]" name="Madagascar" pageId="39" pageNumber="240">Madagascar</collectingCountry>
, 
<collectingRegion id="405359A3305EFFB7FDF8FA29BF9F08C8" box="[525,764,1467,1491]" country="Madagascar" name="Fianarantsoa" pageId="39" pageNumber="240">Fianarantsoa Province</collectingRegion>
, 
<location id="8748C19A305EFFB7FCFFFA29B94F08C8" LSID="urn:lsid:plazi:treatment:0A3E2657305CFFB8FF8EFB4BBCA909E0:8748C19A305EFFB7FCFFFA29B94F08C8" box="[778,1068,1467,1491]" country="Madagascar" name="Ranomafana National Park" pageId="39" pageNumber="240" stateProvince="Fianarantsoa">Ranomafana National Park</location>
. 
<emphasis id="B0E34B53305EFFB7FF5EFA49BC9108EA" bold="true" box="[171,498,1499,1523]" pageId="39" pageNumber="240">
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<smallCapsWord id="84CE019D305EFFB7FEC3FA4CBC3108EA" baselines="1516" box="[310,338,1502,1521]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="of" pageId="39" pageNumber="240">OF</smallCapsWord>
<smallCapsWord id="84CE019D305EFFB7FEACFA4CBCCF08EA" baselines="1516" box="[345,428,1502,1521]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="39" pageNumber="240">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D305EFFB7FE40FA4CBC9108EA" baselines="1516" box="[437,498,1502,1521]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="39" pageNumber="240">NAME</smallCapsWord>
</emphasis>
.— The species epithet, 
<taxonomicName id="4597ECC2305EFFB7FD0CFA49BEE408E8" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[761,903,1499,1523]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B53305EFFB7FD0CFA49BEE408E8" box="[761,903,1499,1523]" italics="true" pageId="39" pageNumber="240">ranomafanae</emphasis>
</taxonomicName>
, is a noun in apposition, derived from the name of the national park in which the 
<typeStatus id="5D2C29E3305EFFB7FD29FA69BE690B08" box="[732,778,1531,1555]" pageId="39" pageNumber="240">type</typeStatus>
was collected
</materialsCitation>
.
</paragraph>
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<smallCapsWord id="84CE019D305EFFB7FF5EF989BC210B2A" baselines="1581,1580" box="[171,322,1563,1587]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Recognition" pageId="39" pageNumber="240">
<emphasis id="B0E34B53305EFFB7FF5EF989BC210B2A" bold="true" box="[171,322,1563,1587]" pageId="39" pageNumber="240">RECOGNITION</emphasis>
</smallCapsWord>
.— Size small for genus, SBL of males = 8.3–8. 
<quantity id="456F3AA4305EFFB7FCABF989BEFC0B29" box="[862,927,1563,1587]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="39" pageNumber="240" unit="mm" value="5.0">5 mm</quantity>
, of female = 
<quantity id="456F3AA4305EFFB7FBCDF989B9ED0B29" box="[1080,1166,1563,1587]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="39" pageNumber="240" unit="mm" value="8.6">8.6 mm</quantity>
. Members of this species (
<figureCitation id="1AAC8BC4305EFFB7FE67F9A9BC910B48" box="[402,498,1595,1619]" captionStart="FIGURE" captionStartId="38.[155,167,1385,1404]" captionTargetBox="[120,1176,216,1353]" captionTargetId="figure-677@38.[120,1176,216,1372]" captionTargetPageId="38" captionText="FIGURE 21. Digital images of holotype male of Pristacrus ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B Labels. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799489" httpUri="https://zenodo.org/record/13799489/files/figure.png" pageId="39" pageNumber="240">Fig. 21A</figureCitation>
) are distinguished from those of all other species in the genus except 
<taxonomicName id="4597ECC2305EFFB7FF3CF9CEBF500B68" authority="Klug (1833)" authorityName="Klug" authorityYear="1833" box="[201,563,1627,1651]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="binotatus">
<emphasis id="B0E34B53305EFFB7FF3CF9CEBCC70B68" box="[201,420,1627,1651]" italics="true" pageId="39" pageNumber="240">Pristacrus binotatus</emphasis>
<bibRefCitation id="E606EAB0305EFFB7FE58F9C9BF500B68" author="KLUG, J. C. F." box="[429,563,1627,1651]" pageId="39" pageNumber="240" pagination="91 - 223" refId="ref30967" refString="KLUG, J. C. F. 1833. Bericht uber eine auf Madagascar veranstaltete. Sammlung von Insecten aus der Ordnung Coleoptera. Abhandlungen der Koniglichen Preussischen Akademie der Wissenschaften 1832 - 33: 91 - 223 + 5 pls." type="book chapter" year="1833">Klug (1833)</bibRefCitation>
</taxonomicName>
by their small body size (SBL greater than 10.0 mm in members of the three other species). They differ from members of 
<taxonomicName id="4597ECC2305EFFB7FC94F9EEBD9C0BA8" authority="(Fairmaire 1887)" baseAuthorityName="Fairmaire" baseAuthorityYear="1887" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="semipiceus">
<emphasis id="B0E34B53305EFFB7FC94F9EEB92C0B88" box="[865,1103,1660,1683]" italics="true" pageId="39" pageNumber="240">Pristacrus semipiceus</emphasis>
(
<bibRefCitation id="E606EAB0305EFFB7FB97F9E9BD960BA8" author="FAIRMAIRE, L." pageId="39" pageNumber="240" pagination="56 - 57" refId="ref30177" refString="FAIRMAIRE, L. 1887. Diagnoses du Coleopteres nouveaux de Madagascar. Le Naturaliste (2 e Serie) 1: 56 - 57" type="journal article" year="1887">Fairmaire 1887</bibRefCitation>
)
</taxonomicName>
, in having much smaller body size (SBL = 13.6 to 14. 
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in 
<taxonomicName id="4597ECC2305EFFB7FC48F90EB92D0BA8" baseAuthorityName="Fairmaire" baseAuthorityYear="1887" box="[957,1102,1692,1715]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="semipiceus">
<emphasis id="B0E34B53305EFFB7FC48F90EB92D0BA8" box="[957,1102,1692,1715]" italics="true" pageId="39" pageNumber="240">P. semipiceus</emphasis>
</taxonomicName>
members), a shinier dorsal surface, shorter and broadly rounded mandibles and an impunctate (but longitudinally rugulose) frons (dorsal surface duller, mandibles distinctly longer and more tapered, and frons punctate as well as rugulose in 
<taxonomicName id="4597ECC23051FFB8FDB7FE8EBFB00C28" baseAuthorityName="Fairmaire" baseAuthorityYear="1887" box="[578,723,284,307]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="semipiceus">
<emphasis id="B0E34B533051FFB8FDB7FE8EBFB00C28" box="[578,723,284,307]" italics="true" pageId="40" pageNumber="241">P. semipiceus</emphasis>
</taxonomicName>
members). They differ from members of 
<taxonomicName id="4597ECC23051FFB8FF8EFEAFBD910C4F" baseAuthorityName="Gory and Laporte" baseAuthorityYear="1837" box="[123,242,316,340]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="laticollis">
<emphasis id="B0E34B533051FFB8FF8EFEAFBD910C4F" box="[123,242,316,340]" italics="true" pageId="40" pageNumber="241">P. laticollis</emphasis>
</taxonomicName>
in having slightly smaller body size (SBL = 10.2–11. 
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), slightly shinier dorsal surface, relatively longer and narrower pronotum and elytra that are broadly ovoid with lateral margin arcuate and lateral explanation broad throughout, especially at middle, and elytral apices distinctly angulate and toothed (duller dorsal surface, relatively broader and shorter pronotum and elytra that are nearly parallel-sided at middle with lateral explanation narrower throughout and elytral apices broadly rounded in 
<taxonomicName id="4597ECC23051FFB8FE68FE4CBF750CEE" baseAuthorityName="Gory and Laporte" baseAuthorityYear="1837" box="[413,534,477,501]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="laticollis">
<emphasis id="B0E34B533051FFB8FE68FE4CBF750CEE" box="[413,534,477,501]" italics="true" pageId="40" pageNumber="241">P. laticollis</emphasis>
</taxonomicName>
members). They differ from the unique 
<typeStatus id="5D2C29E33051FFB8FC24FE4CBE9C0CED" box="[977,1023,478,502]" pageId="40" pageNumber="241">type</typeStatus>
of 
<taxonomicName id="4597ECC23051FFB8FBD3FE4CBCCF0F0D" authority="(Fairmaire 1892)" baseAuthorityName="Fairmaire" baseAuthorityYear="1892" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="rotundatus">
<emphasis id="B0E34B533051FFB8FBD3FE4CBD8D0F0E" italics="true" pageId="40" pageNumber="241">Pristacrus rotundatus</emphasis>
(
<bibRefCitation id="E606EAB03051FFB8FF0BFE6FBCC00F0D" author="FAIRMAIRE, L." box="[254,419,509,534]" pageId="40" pageNumber="241" pagination="clxviii - clxxii" refId="ref30226" refString="FAIRMAIRE, L. 1892. Descriptions de quelques Coleopteres de Madagascar, provenant de la region de Diego- Suarez, encore peu exploree, et qui lui ont ete communiques par notre collegue M. Rene Oberthur. Annales de la Societe Entomologique de France 61 (Bulletin Entomologique Seance du 22 Juin 1892): clxviii - clxxii." type="journal article" year="1892">Fairmaire 1892</bibRefCitation>
)
</taxonomicName>
in having smaller body size (SBL = 
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in 
<taxonomicName id="4597ECC23051FFB8FC37FE6CB9330F0E" baseAuthorityName="Fairmaire" baseAuthorityYear="1892" box="[962,1104,509,533]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="rotundatus">
<emphasis id="B0E34B533051FFB8FC37FE6CB9330F0E" box="[962,1104,509,533]" italics="true" pageId="40" pageNumber="241">P. rotundatus</emphasis>
</taxonomicName>
<typeStatus id="5D2C29E33051FFB8FBADFE6CB9E50F0D" box="[1112,1158,510,534]" pageId="40" pageNumber="241">type</typeStatus>
), shinier dorsal surface, elytra less broadly rounded and humeri less anteriorly projected (dorsal surface dull and elytra markedly broad and rounded and humeri more projected anteriorly in 
<taxonomicName id="4597ECC23051FFB8FBCCFDADBDD60F6D" baseAuthorityName="Fairmaire" baseAuthorityYear="1892" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="rotundatus">
<emphasis id="B0E34B533051FFB8FBCCFDADBDD60F6D" italics="true" pageId="40" pageNumber="241">P. rotundatus</emphasis>
</taxonomicName>
<typeStatus id="5D2C29E33051FFB8FF4EFDCDBD8A0F6C" box="[187,233,607,631]" pageId="40" pageNumber="241">type</typeStatus>
). Members of 
<taxonomicName id="4597ECC23051FFB8FE7CFDCDBF520F6D" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[393,561,606,630]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533051FFB8FE7CFDCDBF520F6D" box="[393,561,606,630]" italics="true" pageId="40" pageNumber="241">P. ranomafanae</emphasis>
</taxonomicName>
are very similar to those of 
<taxonomicName id="4597ECC23051FFB8FCAAFDCDBEBF0F6D" authorityName="Klug" authorityYear="1833" box="[863,988,606,630]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="binotatus">
<emphasis id="B0E34B533051FFB8FCAAFDCDBEBF0F6D" box="[863,988,606,630]" italics="true" pageId="40" pageNumber="241">P. binotatus</emphasis>
</taxonomicName>
in size and overall form. Specimens of the former have the dorsal surface slightly shinier, the pronotum relatively broader basally, with subbasal sinuation of the lateral margins absent or only very faint and hind angles less sharply defined, and elytra with striae slightly more deeply impressed and intervals more convex and more roughened at the raised center than in most members of 
<taxonomicName id="4597ECC23051FFB8FC3BFD72B9280FEC" authorityName="Klug" authorityYear="1833" box="[974,1099,735,759]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="binotatus">
<emphasis id="B0E34B533051FFB8FC3BFD72B9280FEC" box="[974,1099,735,759]" italics="true" pageId="40" pageNumber="241">P. binotatus</emphasis>
</taxonomicName>
. However, at least a few specimens of 
<taxonomicName id="4597ECC23051FFB8FE15FC92BF3D0E0C" authorityName="Klug" authorityYear="1833" box="[480,606,767,791]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="binotatus">
<emphasis id="B0E34B533051FFB8FE15FC92BF3D0E0C" box="[480,606,767,791]" italics="true" pageId="40" pageNumber="241">P. binotatus</emphasis>
</taxonomicName>
share one or more of these features with 
<taxonomicName id="4597ECC23051FFB8FBEBFC92BDD50E23" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="ranomofanae">
<emphasis id="B0E34B533051FFB8FBEBFC92BDD50E23" italics="true" pageId="40" pageNumber="241">P. ranomofanae</emphasis>
</taxonomicName>
members. They only features that most reliably distinguish members of 
<taxonomicName id="4597ECC23051FFB8FC41FCB2B9380E23" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[948,1115,800,824]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533051FFB8FC41FCB2B9380E23" box="[948,1115,800,824]" italics="true" pageId="40" pageNumber="241">P. ranomafanae</emphasis>
</taxonomicName>
from those of 
<taxonomicName id="4597ECC23051FFB8FF2BFCD3BC3C0E43" authorityName="Klug" authorityYear="1833" box="[222,351,832,856]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="binotatus">
<emphasis id="B0E34B533051FFB8FF2BFCD3BC3C0E43" box="[222,351,832,856]" italics="true" pageId="40" pageNumber="241">P. binotatus</emphasis>
</taxonomicName>
, as well as from members of the other species, are the male genitalia. In 
<taxonomicName id="4597ECC23051FFB8FF8EFCF3BC470E63" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[123,292,864,888]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533051FFB8FF8EFCF3BC470E63" box="[123,292,864,888]" italics="true" pageId="40" pageNumber="241">P. ranomafanae</emphasis>
</taxonomicName>
males (Fig. 22A–B), the shaft of the median lobe is inflated subbasally but lacks the large, well-defined dorsal bulge seen in males of 
<taxonomicName id="4597ECC23051FFB8FD45FC13BE4E0E83" authorityName="Klug" authorityYear="1833" box="[688,813,896,920]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="binotatus">
<emphasis id="B0E34B533051FFB8FD45FC13BE4E0E83" box="[688,813,896,920]" italics="true" pageId="40" pageNumber="241">P. binotatus</emphasis>
</taxonomicName>
(see Jeannel 1949, Fig. 486) and, in lateral aspect (
<figureCitation id="1AAC8BC43051FFB8FEC7FC33BCF20EA2" box="[306,401,929,953]" captionStart="FIGURE" captionStartId="38.[155,167,1385,1404]" captionTargetBox="[120,1176,216,1353]" captionTargetId="figure-677@38.[120,1176,216,1372]" captionTargetPageId="38" captionText="FIGURE 21. Digital images of holotype male of Pristacrus ranomafanae Kavanaugh &amp; Rainio sp. nov. A. Habitus, dorsal aspect; B Labels. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799489" httpUri="https://zenodo.org/record/13799489/files/figure.png" pageId="40" pageNumber="241">Fig. 21A</figureCitation>
), the apex is distinctly hooked dorsally (apex not hooked in 
<taxonomicName id="4597ECC23051FFB8FBEDFC30B9F60EA2" authorityName="Klug" authorityYear="1833" box="[1048,1173,929,953]" class="Insecta" family="Carabidae" genus="Pristacrus" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="241" phylum="Arthropoda" rank="species" species="binotatus">
<emphasis id="B0E34B533051FFB8FBEDFC30B9F60EA2" box="[1048,1173,929,953]" italics="true" pageId="40" pageNumber="241">P. binotatus</emphasis>
</taxonomicName>
males. Males of the each of the other species also have the shaft of the median lobe uniquely shaped.
</paragraph>
<paragraph id="822897413051FFB8FF5EFB93B9690902" blockId="40.[123,1174,219,1275]" box="[171,1034,1025,1050]" pageId="40" pageNumber="241">
<emphasis id="B0E34B533051FFB8FF5EFB93BC920903" bold="true" box="[171,497,1025,1049]" pageId="40" pageNumber="241">
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<smallCapsWord id="84CE019D3051FFB8FEA8FB97BC920903" baselines="1043" box="[349,497,1029,1048]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="40" pageNumber="241">DISTRIBUTION</smallCapsWord>
</emphasis>
.— At present, known only from the 
<typeStatus id="5D2C29E33051FFB8FC8AFB90BECE0901" box="[895,941,1026,1050]" pageId="40" pageNumber="241">type</typeStatus>
locality.
</paragraph>
<paragraph id="822897413051FFB8FF5EFBB0BCA909E0" blockId="40.[123,1174,219,1275]" pageId="40" pageNumber="241">
<emphasis id="B0E34B533051FFB8FF5EFBB0BCC70923" bold="true" box="[171,420,1058,1082]" pageId="40" pageNumber="241">
<smallCapsWord id="84CE019D3051FFB8FF5EFBB0BC6A0923" baselines="1076,1075" box="[171,265,1058,1082]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Habitat" pageId="40" pageNumber="241">HABITAT</smallCapsWord>
<smallCapsWord id="84CE019D3051FFB8FEE5FBB7BCC70923" baselines="1075" box="[272,420,1061,1080]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="distribution" pageId="40" pageNumber="241">DISTRIBUTION</smallCapsWord>
</emphasis>
.— Specimens of the 
<typeStatus id="5D2C29E33051FFB8FD78FBB0BFD80921" box="[653,699,1058,1082]" pageId="40" pageNumber="241">type</typeStatus>
series were all collected in secondary montane rainforest in the Talatakely area at elevations of 
<quantity id="456F3AA43051FFB8FD47FBD0BE2C0941" box="[690,847,1090,1114]" metricMagnitude="2" metricUnit="m" metricValue="9.6" metricValueMax="10.2" metricValueMin="9.0" pageId="40" pageNumber="241" unit="m" value="960.0" valueMax="1020.0" valueMin="900.0">900 to 1020 m</quantity>
. The area is replete with invasive guava (
<taxonomicName id="4597ECC23051FFB8FEF7FBF0BC820961" box="[258,481,1122,1146]" class="Magnoliopsida" family="Myrtaceae" genus="Psidium" kingdom="Plantae" order="Myrtales" pageId="40" pageNumber="241" phylum="Tracheophyta" rank="species" species="cattleianum">
<emphasis id="B0E34B533051FFB8FEF7FBF0BC820961" box="[258,481,1122,1146]" italics="true" pageId="40" pageNumber="241">Psidium cattleianum</emphasis>
</taxonomicName>
). Specimens were found in daytime in masses of dead leaves, twigs and branch debris accumulated on the forest floor, in leaf litter, and under loose bark or on the underside of rotting logs on the forest floor. Several were found at night running on the surfaces of rotting logs on the forest floor; two were collected in malaise traps and one was attracted to and collected at an ultraviolet light.
</paragraph>
</subSubSection>
</treatment>
</document>