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<document id="A703E57A9582FE6EF0E6FD7C6EF0818F" ID-DOI="10.5281/zenodo.13799439" ID-ISSN="0068-547X" ID-Zenodo-Dep="13799439" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722538212676" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Rainio, Johanna" docDate="2016" docId="0A3E26573044FFAFFF8EF9B3BE7F0E68" docLanguage="en" docName="ProcCalifAcadSci.63.7.201-268.pdf" docOrigin="Proceedings of the California Academy of Sciences 63 (7)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Erephognathus ranomafanae Kavanaugh and Rainio 2016, sp. nov." docType="treatment" docVersion="2" lastPageNumber="264" masterDocId="F6075E2F3079FF90FFF5FF92BD630D1B" masterDocTitle="Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar" masterLastPageNumber="268" masterPageNumber="201" pageNumber="262" updateTime="1726771873529" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="876C4F278C148FDE7B731CA19B6F256D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2DAE3D58A474A619AA361D7D04FA9641">
<mods:title id="B2AFB64A668B48528D1449804CC1FC3B">Twenty-six New Species of Predaceous Ground Beetles (Coleoptera: Adephaga: Carabidae) from Ranomafana National Park, Madagascar</mods:title>
</mods:titleInfo>
<mods:name id="174E1F795010CC376261793BF46C2840" type="personal">
<mods:role id="8A98FC6E9A3C3EAB3014E136E798F963">
<mods:roleTerm id="DB5446F068B4A45F6F4C1FE00E429F29">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="35C3D299B879E916ED9ADCBA92A3EA7C">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="DD81AC17A6FD9CB04AB96B299A366B83">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA; Email: dkavanaugh @ calacademy. org; &amp; Corresponding author: David H. Kavanaugh (dkavanaugh @ calacademy. org)</mods:affiliation>
</mods:name>
<mods:name id="CAEE6A96D2BCAD4DE4A30D8EAEE9EECA" type="personal">
<mods:role id="F293F819A7EC260F971FA77BFAA0087D">
<mods:roleTerm id="845B6BB8651C9A5B6F41E837C49DC092">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9FA12F1F84E84DA8567F38E755721F29">Rainio, Johanna</mods:namePart>
<mods:affiliation id="D064CEB0030F5820C443040683804C81">Department of Ecology and Systematics, University of Helsinki, P. O. Box 65, FIN- 00014 Helsinki, Finland; Email: kjohannarainio @ gmail. com</mods:affiliation>
</mods:name>
<mods:typeOfResource id="29288224A48B00BDB8293E08AF453440">text</mods:typeOfResource>
<mods:relatedItem id="F448B19A0387459C9A547150E3DA2007" type="host">
<mods:titleInfo id="C6D37968A14913AD2276F1D0257D8965">
<mods:title id="3628751EA964FEFE118F4708B50277C7">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="90F3E88C62E665485D724D2E347C1AEE">
<mods:date id="C33BB3C28531C478688808B92684F6FD">2016</mods:date>
<mods:detail id="FCA85D52C7C7B7610380A99E36EEE336" type="pubDate">
<mods:number id="C730E4560D3F7AD241C4C5EB18CF50F1">2016-04-29</mods:number>
</mods:detail>
<mods:detail id="FBC5F9FA7B1015AB9CEF65A08B24CC6A" type="volume">
<mods:number id="69CDB88AA90C3EC97EA913E642816D68">63</mods:number>
</mods:detail>
<mods:detail id="1563EC4DD498E3CF0B85CC0A0B8BF9E1" type="issue">
<mods:number id="A7BD88A5AD09C429F4E542DA773FD11B">7</mods:number>
</mods:detail>
<mods:extent id="0C2AF243DFB5EBD22AF7BF46C72B51F3" unit="page">
<mods:start id="3B8E5D6103A2F9B483416F6D9D295338">201</mods:start>
<mods:end id="E7499D55E5D505B7CB61E96F2746715C">268</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="0BEC68EE80B1AAC54AEE51AD1DDA68DA">journal article</mods:classification>
<mods:identifier id="732E456D19F331686AA57257EF047116" type="DOI">10.5281/zenodo.13799439</mods:identifier>
<mods:identifier id="D40900111C2E17BD1244EA65E0048F92" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="8EE4DC7B41D83BE451C97C6BC23527C5" type="Zenodo-Dep">13799439</mods:identifier>
</mods:mods>
<treatment id="0A3E26573044FFAFFF8EF9B3BE7F0E68" LSID="urn:lsid:plazi:treatment:0A3E26573044FFAFFF8EF9B3BE7F0E68" httpUri="http://treatment.plazi.org/id/0A3E26573044FFAFFF8EF9B3BE7F0E68" lastPageId="63" lastPageNumber="264" pageId="61" pageNumber="262">
<subSubSection id="CA8DC4CA3044FFADFF8EF9B3BE560B21" box="[123,821,1569,1594]" pageId="61" pageNumber="262" type="nomenclature">
<paragraph id="822897413044FFADFF8EF9B3BE560B21" blockId="61.[123,821,1569,1624]" box="[123,821,1569,1594]" pageId="61" pageNumber="262">
<heading id="D960202D3044FFADFF8EF9B3BE560B21" box="[123,821,1569,1594]" fontSize="10" level="2" pageId="61" pageNumber="262" reason="4">
<emphasis id="B0E34B533044FFADFF8EF9B3BE560B21" bold="true" box="[123,821,1569,1594]" pageId="61" pageNumber="262">
<taxonomicName id="4597ECC23044FFADFF8EF9B3BFB20B22" ID-CoL="6GH2T" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[123,721,1569,1594]" class="Insecta" family="Carabidae" genus="Erephognathus" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="262" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B533044FFADFF8EF9B3BCA30B22" bold="true" box="[123,448,1569,1593]" italics="true" pageId="61" pageNumber="262">Erephognathus ranomafanae</emphasis>
Kavanaugh and Rainio
</taxonomicName>
,
<taxonomicNameLabel id="ABD0F6283044FFADFD2AF9B0BE560B21" box="[735,821,1570,1594]" pageId="61" pageNumber="262" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3044FFADFF8EF9D0BDBB0B4C" box="[123,216,1602,1624]" pageId="61" pageNumber="262" type="description">
<paragraph id="822897413044FFADFF8EF9D0BDBB0B4C" blockId="61.[123,821,1569,1624]" box="[123,216,1602,1624]" pageId="61" pageNumber="262">
<figureCitation id="1AAC8BC43044FFADFF8EF9D0BDBB0B4C" box="[123,216,1602,1624]" captionStart="FIGURE" captionStartId="62.[155,167,1459,1478]" captionTargetBox="[152,1176,222,1433]" captionTargetId="figure-507@62.[120,1176,216,1456]" captionTargetPageId="62" captionText="FIGURE 38. Digital images of holotype female of Erephognathus ranomafanae Kavanaugh and Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799525" httpUri="https://zenodo.org/record/13799525/files/figure.png" pageId="61" pageNumber="262">Figure 38</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="CA8DC4CA3044FFAFFF5EF9E9BE7F0E68" lastPageId="63" lastPageNumber="264" pageId="61" pageNumber="262" type="materials_examined">
<paragraph id="822897413044FFADFF5EF9E9B9F60B88" blockId="61.[123,1173,1658,1715]" box="[171,1173,1658,1683]" pageId="61" pageNumber="262">
<emphasis id="B0E34B533044FFADFF5EF9E9BC3B0B8A" bold="true" box="[171,344,1659,1683]" pageId="61" pageNumber="262">
<smallCapsWord id="84CE019D3044FFADFF5EF9E9BD870B8A" baselines="1677,1676" box="[171,228,1659,1683]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="61" pageNumber="262">TYPE</smallCapsWord>
<smallCapsWord id="84CE019D3044FFADFF1EF9ECBC3B0B8A" baselines="1676" box="[235,344,1662,1681]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="61" pageNumber="262">MATERIAL</smallCapsWord>
</emphasis>
.—
<typeStatus id="5D2C29E33044FFADFE75F9E8BC800B89" box="[384,483,1658,1682]" pageId="61" pageNumber="262" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1AAC8BC43044FFADFE06F9E8BF170B89" box="[499,628,1658,1683]" captionStart="FIGURE" captionStartId="62.[155,167,1459,1478]" captionTargetBox="[152,1176,222,1433]" captionTargetId="figure-507@62.[120,1176,216,1456]" captionTargetPageId="62" captionText="FIGURE 38. Digital images of holotype female of Erephognathus ranomafanae Kavanaugh and Rainio sp. nov. A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13799525" httpUri="https://zenodo.org/record/13799525/files/figure.png" pageId="61" pageNumber="262">Figs.38A–B</figureCitation>
), a female, in CAS, labeled: “CASENT 8068531”/
</paragraph>
<paragraph id="822897413044FFAFFF8EF909BE130C28" blockId="61.[123,1173,1658,1715]" lastBlockId="63.[123,1174,219,883]" lastPageId="63" lastPageNumber="264" pageId="61" pageNumber="262">
“
<materialsCitation id="32FF9D1C3044FFAFFF72F909BE0A0C28" collectingDate="2000-12-05" collectingDateMax="2001-11-15" collectingDateMin="2000-12-05" collectionCode="CAS" collectorName="R. Harin'Hala California Acad &amp; Fisher &amp; Both" country="Madagascar" county="Province" elevation="1110" lastPageId="63" lastPageNumber="264" latitude="-21.226166" location="Vohiparara" longLatPrecision="12" longitude="47.369835" municipality="Parc National Ranomafana" pageId="61" pageNumber="262" specimenCount="2" specimenCount-female="2" stateProvince="Fianarantsoa" typeStatus="paratype">
<collectingCountry id="FA80D7D13044FFADFF72F909BC260BA8" box="[135,325,1691,1715]" name="Madagascar" pageId="61" pageNumber="262">MADAGASCAR</collectingCountry>
:
<collectingCounty id="6B49EFCD3044FFADFEACF908BCDB0BA9" box="[345,440,1690,1714]" pageId="61" pageNumber="262">Province</collectingCounty>
<collectingRegion id="405359A33044FFADFE30F908BF2E0BA9" box="[453,589,1690,1714]" country="Madagascar" name="Fianarantsoa" pageId="61" pageNumber="262">Fianarantsoa</collectingRegion>
,
<collectingMunicipality id="624C0D3B3044FFADFD97F909BEEC0BA9" box="[610,911,1690,1714]" pageId="61" pageNumber="262">Parc National Ranomafana</collectingMunicipality>
,
<location id="8748C19A3044FFADFC57F908B9750BA9" LSID="urn:lsid:plazi:treatment:0A3E26573044FFAFFF8EF9B3BE7F0E68:8748C19A3044FFADFC57F908B9750BA9" box="[930,1046,1690,1714]" country="Madagascar" county="Province" latitude="-21.226166" longLatPrecision="12" longitude="47.369835" municipality="Parc National Ranomafana" name="Vohiparara" pageId="61" pageNumber="262" stateProvince="Fianarantsoa">Vohiparara</location>
, at broken bridge, el
<quantity id="456F3AA43047FFAEFF12F996BC550B07" box="[231,310,1540,1564]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="62" pageNumber="263" unit="m" value="1110.0">
<elevation id="09BA70723047FFAEFF12F996BC550B07" box="[231,310,1540,1564]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="62" pageNumber="263" unit="m" value="1110.0">1110 m</elevation>
</quantity>
<date id="F629B1813047FFAEFEC8F996BF420B07" box="[317,545,1540,1564]" pageId="62" pageNumber="263" value="2001-11-08" valueMax="2001-11-15" valueMin="2001-11-08">
<collectingDate id="E66D48693047FFAEFEC8F996BF420B07" box="[317,545,1540,1564]" pageId="62" pageNumber="263" value="2001-11-08" valueMax="2001-11-15" valueMin="2001-11-08">8-15 November 2001</collectingDate>
</date>
”/ “
<geoCoordinate id="E7A3F1863047FFAEFDBDF996BFA20B07" box="[584,705,1540,1564]" degrees="21" direction="south" minutes="13. 57" orientation="latitude" pageId="62" pageNumber="263" precision="9" value="-21.226166">21°13.57’S</geoCoordinate>
,
<geoCoordinate id="E7A3F1863047FFAEFD3BF996BE350B07" box="[718,854,1540,1564]" degrees="047" direction="east" minutes="22. 19" orientation="longitude" pageId="62" pageNumber="263" precision="9" value="47.369835">047°22.19’E</geoCoordinate>
collector:
<collectorName id="2F62F2973047FFAEFC3CF997BC630B27" pageId="62" pageNumber="263">
R.
<location id="8748C19A3047FFAEFC1DF996BC630B27" LSID="urn:lsid:plazi:treatment:0A3E26573044FFAFFF8EF9B3BE7F0E68:8748C19A3047FFAEFC1DF996BC630B27" country="Madagascar" county="Province" latitude="-21.226166" longLatPrecision="12" longitude="47.369835" municipality="Parc National Ranomafana" name="Harin'Hala California Acad of Sciences" pageId="62" pageNumber="263" stateProvince="Fianarantsoa">Harin’Hala California Acad</location>
</collectorName>
of Sciences malaise trap in high altitude rainforest, MA-02-09A-02”/ “
<typeStatus id="5D2C29E33047FFAEFBFCF9B7B9F50B26" box="[1033,1174,1573,1597]" pageId="62" pageNumber="263" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4597ECC23047FFAEFF8EF9D7BFFE0B46" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" box="[123,669,1605,1629]" class="Insecta" family="Carabidae" genus="Erephognathus" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="263" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B533047FFAEFF8EF9D7BCD10B46" box="[123,434,1605,1629]" italics="true" pageId="62" pageNumber="263">Erephognathus ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283047FFAEFD50F9D7BFBC0B47" box="[677,735,1605,1628]" pageId="62" pageNumber="263" rank="species">sp. n.</taxonomicNameLabel>
2015” [red label].
<typeStatus id="5D2C29E33047FFAEFC5EF9D7B9700B46" box="[939,1043,1605,1629]" pageId="62" pageNumber="263" type="paratype">Paratypes</typeStatus>
(total of 2):
<specimenCount id="94915CC83047FFAEFF8EF9F7BDB80B66" box="[123,219,1637,1661]" count="1" pageId="62" pageNumber="263" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F843047FFAEFEFBF9F7BC210B66" box="[270,322,1637,1661]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="62" pageNumber="263" type="Museum">CAS</collectionCode>
) labeled same as
<typeStatus id="5D2C29E33047FFAEFDFBF9F7BF080B66" box="[526,619,1637,1661]" pageId="62" pageNumber="263" type="holotype">holotype</typeStatus>
, except “CASENT 8068532”;
<specimenCount id="94915CC83047FFAEFC33F9F7B9450B66" box="[966,1062,1637,1661]" count="1" pageId="62" pageNumber="263" type="female">1 female</specimenCount>
(in
<collectionCode id="E4860F843047FFAEFBACF9F7B9EE0B66" box="[1113,1165,1637,1661]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="62" pageNumber="263" type="Museum">CAS</collectionCode>
) labeled “CASENT 8068672”/; “
<collectingCountry id="FA80D7D13047FFAEFE20F917BFF00B86" box="[469,659,1669,1693]" name="Madagascar" pageId="62" pageNumber="263">MADAGASCAR</collectingCountry>
:
<collectingRegion id="405359A33047FFAEFD54F917BECD0B86" box="[673,942,1669,1693]" country="Madagascar" name="Antananarivo" pageId="62" pageNumber="263">Province d’Antananarivo</collectingRegion>
<locationDeviation id="939A8E633047FFAEFC43F917BDA10BA5" location="Andranomay" pageId="62" pageNumber="263">
<quantity id="456F3AA43047FFAEFC43F917BE8F0B86" box="[950,1004,1669,1693]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="62" pageNumber="263" unit="km" value="3.0">3 km</quantity>
41° NE Andranomay
</locationDeviation>
,
<locationDeviation id="939A8E633047FFAEFF2DF934BF210BA6" box="[216,578,1701,1726]" location="Anjozorobe" pageId="62" pageNumber="263">
<quantity id="456F3AA43047FFAEFF2DF934BC540BA6" box="[216,311,1701,1726]" metricMagnitude="4" metricUnit="m" metricValue="1.15" pageId="62" pageNumber="263" unit="km" value="11.5">11.5 km</quantity>
147° SSE Anjozorobe
</locationDeviation>
elev
<quantity id="456F3AA43047FFAEFD78F934BF840BA6" box="[653,743,1702,1726]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="62" pageNumber="263" unit="m" value="1300.0">
<elevation id="09BA70723047FFAEFD78F934BF840BA6" box="[653,743,1702,1726]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="62" pageNumber="263" unit="m" value="1300.0">1300 m</elevation>
</quantity>
<date id="F629B1813047FFAEFD03F937BE8B0BA5" box="[758,1000,1701,1726]" pageId="62" pageNumber="263" value="2000-12-05" valueMax="2000-12-13" valueMin="2000-12-05">
<collectingDate id="E66D48693047FFAEFD03F937BE8B0BA5" box="[758,1000,1701,1726]" pageId="62" pageNumber="263" value="2000-12-05" valueMax="2000-12-13" valueMin="2000-12-05">5-13 December 2000</collectingDate>
</date>
”/ “
<geoCoordinate id="E7A3F1863047FFAEFBE2F934B9F50BA5" box="[1047,1174,1702,1726]" degrees="18" direction="south" minutes="28" orientation="latitude" pageId="62" pageNumber="263" precision="15" seconds="24" value="-18.473333">18°28’24”S</geoCoordinate>
<geoCoordinate id="E7A3F1863046FFAFFF8EFF49BD990DE9" box="[123,250,219,243]" degrees="47" direction="east" minutes="57" orientation="longitude" pageId="63" pageNumber="264" precision="15" seconds="38" value="47.960556">47°57’38”E</geoCoordinate>
coll.
<collectorName id="2F62F2973046FFAFFECDFF49BC190DE8" box="[312,378,219,243]" pageId="63" pageNumber="264">Fisher</collectorName>
, Griswold et al.
<location id="8748C19A3046FFAFFDC5FF49BE070DE8" LSID="urn:lsid:plazi:treatment:0A3E26573044FFAFFF8EF9B3BE7F0E68:8748C19A3046FFAFFDC5FF49BE070DE8" box="[560,868,219,243]" country="Madagascar" county="Province" latitude="-21.226166" longLatPrecision="12" longitude="47.369835" municipality="Parc National Ranomafana" name="California Acad. of Sciences" pageId="63" pageNumber="264" stateProvince="Fianarantsoa">California Acad. of Sciences</location>
montane rainforest,
<collectingMethod id="5BD6EF563046FFAFFBB1FF49BDC70C08" pageId="63" pageNumber="264">malaise trap</collectingMethod>
, coll. code: BLF2375”.
<collectorName id="2F62F2973046FFAFFE59FF69BC830C08" box="[428,480,251,275]" pageId="63" pageNumber="264">Both</collectorName>
<typeStatus id="5D2C29E33046FFAFFE12FF6EBF2E0C0F" box="[487,589,252,276]" pageId="63" pageNumber="264" type="paratype">paratypes</typeStatus>
also bear the following label: “
<typeStatus id="5D2C29E33046FFAFFC50FF69B94A0C08" box="[933,1065,251,275]" pageId="63" pageNumber="264" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4597ECC23046FFAFFBC7FF69BF200C28" authority="Kavanaugh &amp; Rainio" authorityName="Kavanaugh &amp; Rainio" authorityYear="2016" class="Insecta" family="Carabidae" genus="Erephognathus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="264" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B533046FFAFFBC7FF69BC340C28" italics="true" pageId="63" pageNumber="264">Erephognathus ranomafanae</emphasis>
Kavanaugh &amp; Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283046FFAFFDBFFE8EBFE70C28" box="[586,644,284,307]" pageId="63" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
2015” [yellow label]
</materialsCitation>
.
</paragraph>
<caption id="D6E8C7C93047FFAEFF6EFA21BF0E08FB" ID-DOI="http://doi.org/10.5281/zenodo.13799525" ID-Zenodo-Dep="13799525" httpUri="https://zenodo.org/record/13799525/files/figure.png" pageId="62" pageNumber="263" startId="62.[155,167,1459,1478]" targetBox="[152,1176,222,1433]" targetPageId="62" targetType="figure">
<paragraph id="822897413047FFAEFF6EFA21B9F608DD" blockId="62.[123,1173,1459,1504]" box="[155,1173,1459,1478]" pageId="62" pageNumber="263">
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38. Digital images of holotype female of
<taxonomicName id="4597ECC23047FFAEFD9EFA21B92008DD" authority="Kavanaugh and Rainio" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[619,1091,1459,1478]" class="Insecta" family="Carabidae" genus="Erephognathus" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="263" phylum="Arthropoda" rank="species" species="ranomafanae" status="sp. nov.">
<emphasis id="B0E34B533047FFAEFD9EFA21BE0A08DD" box="[619,873,1459,1478]" italics="true" pageId="62" pageNumber="263">Erephognathus ranomafanae</emphasis>
Kavanaugh and Rainio
</taxonomicName>
<taxonomicNameLabel id="ABD0F6283047FFAEFBBBFA21B9F608DD" box="[1102,1173,1459,1478]" pageId="62" pageNumber="263" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="822897413047FFAEFF8EFA5FBF0E08FB" blockId="62.[123,1173,1459,1504]" box="[123,621,1484,1504]" pageId="62" pageNumber="263">A. Habitus, dorsal aspect; B. Labels. Scale line = 1.0 mm.</paragraph>
</caption>
<paragraph id="822897413046FFAFFF5EFEA9B9500C48" blockId="63.[123,1174,219,883]" box="[171,1075,315,339]" pageId="63" pageNumber="264">
<materialsCitation id="32FF9D1C3046FFAFFF5EFEA9B94F0C48" box="[171,1068,315,339]" country="Madagascar" location="Ranomafana National Park" pageId="63" pageNumber="264" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
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<typeStatus id="5D2C29E33046FFAFFF5EFEA9BD870C49" box="[171,228,315,339]" pageId="63" pageNumber="264">TYPE</typeStatus>
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</emphasis>
.—
<collectingCountry id="FA80D7D13046FFAFFE8BFEA9BC9C0C48" box="[382,511,315,339]" name="Madagascar" pageId="63" pageNumber="264">Madagascar</collectingCountry>
,
<collectingRegion id="405359A33046FFAFFDF8FEA9BF9F0C48" box="[525,764,315,339]" country="Madagascar" name="Fianarantsoa" pageId="63" pageNumber="264">Fianarantsoa Province</collectingRegion>
,
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</materialsCitation>
.
</paragraph>
<paragraph id="822897413046FFAFFF5EFEC9BECA0C88" blockId="63.[123,1174,219,883]" pageId="63" pageNumber="264">
<emphasis id="B0E34B533046FFAFFF5EFEC9BC910C69" bold="true" box="[171,498,347,371]" pageId="63" pageNumber="264">
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<smallCapsWord id="84CE019D3046FFAFFEACFECCBCCF0C6A" baselines="364" box="[345,428,350,369]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="species" pageId="63" pageNumber="264">SPECIES</smallCapsWord>
<smallCapsWord id="84CE019D3046FFAFFE40FECDBC910C69" baselines="365" box="[437,498,351,370]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="name" pageId="63" pageNumber="264">NAME</smallCapsWord>
</emphasis>
.— The species epithet,
<taxonomicName id="4597ECC23046FFAFFD0CFEC9BEE40C68" authorityName="Kavanaugh and Rainio" authorityYear="2016" box="[761,903,347,371]" class="Insecta" family="Carabidae" genus="Erephognathus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="264" phylum="Arthropoda" rank="species" species="ranomafanae">
<emphasis id="B0E34B533046FFAFFD0CFEC9BEE40C68" box="[761,903,347,371]" italics="true" pageId="63" pageNumber="264">ranomafanae</emphasis>
</taxonomicName>
, is a noun in apposition, derived from the name of the national park in which the
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was collected.
</paragraph>
<paragraph id="822897413046FFAFFF5EFE09BF4C0F68" blockId="63.[123,1174,219,883]" pageId="63" pageNumber="264">
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<emphasis id="B0E34B533046FFAFFF5EFE09BC210CAA" bold="true" box="[171,322,411,435]" pageId="63" pageNumber="264">RECOGNITION</emphasis>
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.— Size moderate for genus, SBL of females = 7.5
<emphasis id="B0E34B533046FFAFFCAAFE09BE0B0CA8" bold="true" box="[863,872,411,435]" pageId="63" pageNumber="264">-</emphasis>
8.
<quantity id="456F3AA43046FFAFFC74FE09BEDD0CA8" box="[897,958,411,435]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="63" pageNumber="264" unit="mm" value="9.0">9 mm</quantity>
. In Jeannel’s (1949) key to species of
<taxonomicName id="4597ECC23046FFAFFECFFE29BCBF0CC8" authorityName="Alluaud" authorityYear="1932" box="[314,476,443,467]" class="Insecta" family="Carabidae" genus="Erephognathus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="B0E34B533046FFAFFECFFE29BCBF0CC8" box="[314,476,443,467]" italics="true" pageId="63" pageNumber="264">Erephognathus</emphasis>
</taxonomicName>
, members of
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<emphasis id="B0E34B533046FFAFFD86FE2EBE430CC8" box="[627,800,443,467]" italics="true" pageId="63" pageNumber="264">E. ranomafanae</emphasis>
</taxonomicName>
key to
<taxonomicName id="4597ECC23046FFAFFC81FE2EBDDA0CE8" authority="(Fairmaire 1903)" baseAuthorityName="Fairmaire" baseAuthorityYear="1903" class="Insecta" family="Carabidae" genus="Erephognathus" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="264" phylum="Arthropoda" rank="species" species="coerulescens">
<emphasis id="B0E34B533046FFAFFC81FE2EB97E0CC8" box="[884,1053,443,467]" italics="true" pageId="63" pageNumber="264">E. coerulescens</emphasis>
(
<bibRefCitation id="E606EAB03046FFAFFBDBFE29BDD30CE8" author="FAIRMAIRE, L." pageId="63" pageNumber="264" pagination="181 - 259" refId="ref30378" refString="FAIRMAIRE, L. 1903. Materiaux pour la faune coleopterique de la region Malagache (16 e Note). Annales de la Societe Entomologique de France 72: 181 - 259." type="journal article" year="1903">Fairmaire 1903</bibRefCitation>
)
</taxonomicName>
based on their short antennae (extended posteriorly only to the basal one-fourth of the elytra or less) and the absence of the tuft of setae from the apex of antennomere 11 (terminal antennomere) found in
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<emphasis id="B0E34B533046FFAFFEC9FD8EBC800F28" box="[316,483,539,563]" italics="true" pageId="63" pageNumber="264">E. margarithrix</emphasis>
<bibRefCitation id="E606EAB03046FFAFFE1DFD89BFFB0F28" author="ALLUAUD, C." box="[488,664,539,563]" pageId="63" pageNumber="264" pagination="1 - 13" refId="ref28980" refString="ALLUAUD, C. 1936 a. Carabidae recueillis a Madagascar par MM. Syrig, G. Olsoufieff, Vadon, R. Catala, etc. Afra, Cahiers d'Entomologie 11: 1 - 13." type="journal article" year="1936">Alluaud (1936a)</bibRefCitation>
</taxonomicName>
members. The pronotum (
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) is distinctly longer, narrower and less convex and with anterior angles more projected, narrowed and angulate than in members of
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<emphasis id="B0E34B533046FFAFFE75FDCEBF4A0F68" box="[384,553,603,627]" italics="true" pageId="63" pageNumber="264">E. coerulescens</emphasis>
</taxonomicName>
.
</paragraph>
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</emphasis>
.—
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We have examined specimens from only two areas.
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specimens were collected in
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, the
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locality, on the eastern slope off the central plateau in the northeastern part of the southern half of the island of
<collectingCountry id="FA80D7D13046FFAFFBD9FD29BDFF0FE8" name="Madagascar" pageId="63" pageNumber="264">Madagascar</collectingCountry>
. A third specimen, also a female, was collected near
<collectingMunicipality id="624C0D3B3046FFAFFD1EFD49BE0A0FE8" box="[747,873,731,755]" pageId="63" pageNumber="264">Anjozorobe</collectingMunicipality>
, in
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, slightly north of the center of the island and just over
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north of
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along the eastern slope off the central plateau.
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specimens have been recorded from intervening areas
</materialsCitation>
.
</paragraph>
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</emphasis>
.— Specimens of the
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series were all collected in malaise traps in montane rainforest at elevations ranging from
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.
</paragraph>
</subSubSection>
</treatment>
</document>
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