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<mods:title id="E6C47E73427576ED5ED5F36C6784BD35">Megarthrus of China. Part 2. Megarthrus basicornis Fauvel, 1904 with description of a new species from Mount Emei (Coleoptera: Staphylinidae: Proteininae)</mods:title>
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<paragraph id="8BF4F7026471FF8D3B8FFF259C683C89" blockId="2.[151,592,151,213]" box="[151,592,151,178]" pageId="2" pageNumber="126">
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<taxonomicName id="4C4B8C816471FF8D3B8FFF259C683C89" ID-CoL="3YX8N" authority="Fauvel, 1904" authorityName="Fauvel" authorityYear="1904" box="[151,592,151,178]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="126" phylum="Arthropoda" rank="species" species="basicornis">
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<emphasis id="B93F2B106471FF8D3B8FFF259F923C8A" bold="true" box="[151,426,151,178]" italics="true" pageId="2" pageNumber="126">Megarthrus basicornis</emphasis>
<bibRefCitation id="EFDA8AF36471FF8D3AA9FF259C683C89" author="Fauvel, A." box="[433,592,151,177]" pageId="2" pageNumber="126" pagination="76 - 112" refId="ref2837" refString="Fauvel, A. (1904) Staphylinides exotiques nouveaux, 2. Revue d'entomologie, 23, 76 - 112." type="journal article" year="1904">Fauvel, 1904</bibRefCitation>
</emphasis>
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<subSubSection id="C351A4896471FF8D3B8FFF099F473CED" box="[151,383,187,213]" pageId="2" pageNumber="126" type="description">
<paragraph id="8BF4F7026471FF8D3B8FFF099F473CED" blockId="2.[151,592,151,213]" box="[151,383,187,213]" pageId="2" pageNumber="126">
(
<figureCitation id="1370EB876471FF8D3B87FF099EC63CED" box="[159,254,187,213]" captionStart-0="FIGURES 1-3" captionStart-1="FIGURES 4-5" captionStart-2="FIGURES 6-9" captionStartId-0="1.[151,264,1857,1881]" captionStartId-1="2.[151,264,1943,1967]" captionStartId-2="3.[151,264,1906,1930]" captionTargetBox-0="[170,1424,283,1824]" captionTargetBox-1="[161,1428,857,1910]" captionTargetBox-2="[158,1436,181,1882]" captionTargetId-0="figure-85@1.[151,1436,273,1834]" captionTargetId-1="figure-245@2.[151,1436,844,1920]" captionTargetId-2="figure-34@3.[151,1436,181,1882]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURES 1-3. Megarthrus basicornis Fauvel, habitus, male: dorsal (1), ventral (2), and lateral (3) views. Scale bar = 0.5 mm." captionText-1="FIGURES 4-5. Megarthrus basicornis Fauvel, male: aedeagus in lateral (4) and ventral (5) views. Scale bar = 0.35 mm." captionText-2="FIGURES 6-9. Megarthrus basicornis Fauvel (6-7) and M. budai sp. n. (8-9): prothorax in dorsal (6, 8) and ventral (7, 9) views. Scale bar = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4321567" figureDoi-1="http://doi.org/10.5281/zenodo.4321569" figureDoi-2="http://doi.org/10.5281/zenodo.4321574" httpUri-0="https://zenodo.org/record/4321567/files/figure.png" httpUri-1="https://zenodo.org/record/4321569/files/figure.png" httpUri-2="https://zenodo.org/record/4321574/files/figure.png" pageId="2" pageNumber="126">Figs 1-7</figureCitation>
,
<figureCitation id="1370EB876471FF8D3A13FF099F753CED" box="[267,333,187,213]" captionStart="FIGURES 10-13" captionStartId="4.[151,264,1812,1836]" captionTargetBox="[151,1436,848,1788]" captionTargetId="figure-317@4.[151,1436,848,1788]" captionTargetPageId="4" captionText="FIGURES 10-13. Megarthrus basicornis Fauvel (10-11) and M. budai sp. n. (12-13): Portarsus of male in ventrolateral (10, 12) and ventral (11, 13) views. Ovals indicate absence (red) or presence (green) of ventral patch of modified adhesive setae (green arrows). Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4321576" httpUri="https://zenodo.org/record/4321576/files/figure.png" pageId="2" pageNumber="126">10-11</figureCitation>
,
<figureCitation id="1370EB876471FF8D3A41FF099F4E3CED" box="[345,374,187,213]" captionStart="FIGURE 39" captionStartId="9.[151,250,1121,1145]" captionTargetBox="[151,1436,345,1061]" captionTargetId="figure-102@9.[151,1436,345,1061]" captionTargetPageId="9" captionText="FIGURE 39. Distribution map of Megarthrus basicornis Fauvel (pink squares) and Megarthrus budai sp. n. (red circle). Scale bar =200 km." figureDoi="http://doi.org/10.5281/zenodo.4321586" httpUri="https://zenodo.org/record/4321586/files/figure.png" pageId="2" pageNumber="126">39</figureCitation>
)
</paragraph>
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<subSubSection id="C351A4896471FF8D3B8FFEB29DFA3D20" box="[151,962,255,280]" pageId="2" pageNumber="126" type="reference_group">
<paragraph id="8BF4F7026471FF8D3B8FFEB29DFA3D20" blockId="2.[151,962,255,280]" box="[151,962,255,280]" pageId="2" pageNumber="126">
<treatmentCitationGroup id="AB5BD02C6471FF8D3B8FFEB29DFA3D20" box="[151,962,255,280]" pageId="2" pageNumber="126">
<treatmentCitation id="0AEAD1136471FF8D3B8FFEB29C0B3D20" author="Fauvel, A." box="[151,563,256,280]" page="87" pageId="2" pageNumber="126" year="1904">
<taxonomicName id="4C4B8C816471FF8D3B8FFEB29C0B3D20" ID-CoL="3YX8N" authority="Fauvel, 1904: 87" authorityName="Fauvel" authorityPageNumber="87" authorityYear="1904" box="[151,563,256,280]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="126" phylum="Arthropoda" rank="species" species="basicornis">
<emphasis id="B93F2B106471FF8D3B8FFEB29F473D20" box="[151,383,256,280]" italics="true" pageId="2" pageNumber="126">Megarthrus basicornis</emphasis>
<bibRefCitation id="EFDA8AF36471FF8D3A9DFEB29C0B3D20" author="Fauvel, A." box="[389,563,256,280]" pageId="2" pageNumber="126" pagination="76 - 112" refId="ref2837" refString="Fauvel, A. (1904) Staphylinides exotiques nouveaux, 2. Revue d'entomologie, 23, 76 - 112." type="journal article" year="1904">Fauvel, 1904: 87</bibRefCitation>
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;
<treatmentCitation id="0AEAD1136471FF8D3925FEB29D273D2F" author="Cuccodoro, G." box="[573,799,255,280]" page="366" pageId="2" pageNumber="126" year="2003">
<bibRefCitation id="EFDA8AF36471FF8D3925FEB29D273D2F" author="Cuccodoro, G." box="[573,799,255,280]" pageId="2" pageNumber="126" refId="ref2715" refString="Cuccodoro, G. (2003) A revision of Megarthrus Curtis of India, Nepal and Sri Lanka (Coleoptera, Staphylinidae, Proteininae). In: Cuccodoro, G. &amp; Leschen, R. A. B. (Eds.), Systematics of Coleoptera: Papers celebrating the retirement of Ivan Lobl. Memoirs on Entomology. Vol. 17. International, Gainesville, pp. 359 - 455." type="book" year="2003">Cuccodoro, 2003: 366</bibRefCitation>
</treatmentCitation>
(redescription).
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</paragraph>
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<subSubSection id="C351A4896471FF8D3B8FFEF49DAF3F0C" pageId="2" pageNumber="126" type="diagnosis">
<paragraph id="8BF4F7026471FF8D3B8FFEF49FF93EB8" blockId="2.[151,1437,326,820]" pageId="2" pageNumber="126">
Diagnosis. For detailed morphology see
<bibRefCitation id="EFDA8AF36471FF8D394FFEF49D1E3D58" author="Cuccodoro, G." box="[599,806,326,352]" pageId="2" pageNumber="126" refId="ref2715" refString="Cuccodoro, G. (2003) A revision of Megarthrus Curtis of India, Nepal and Sri Lanka (Coleoptera, Staphylinidae, Proteininae). In: Cuccodoro, G. &amp; Leschen, R. A. B. (Eds.), Systematics of Coleoptera: Papers celebrating the retirement of Ivan Lobl. Memoirs on Entomology. Vol. 17. International, Gainesville, pp. 359 - 455." type="book" year="2003">Cuccodoro (2003)</bibRefCitation>
. Habitus as in
<figureCitation id="1370EB876471FF8D38DDFEF49A1A3D58" box="[965,1058,326,352]" captionStart="FIGURES 1-3" captionStartId="1.[151,264,1857,1881]" captionTargetBox="[170,1424,283,1824]" captionTargetId="figure-85@1.[151,1436,273,1834]" captionTargetPageId="1" captionText="FIGURES 1-3. Megarthrus basicornis Fauvel, habitus, male: dorsal (1), ventral (2), and lateral (3) views. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4321567" httpUri="https://zenodo.org/record/4321567/files/figure.png" pageId="2" pageNumber="126">Figs 1-3</figureCitation>
. Combined length of head, pronotum, and elytra
<quantity id="4CB35AE76471FF8D3A50FED89FF63DBC" box="[328,462,362,388]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="2" pageNumber="126" unit="mm" value="1.65" valueMax="1.7" valueMin="1.6">1.6-1.7 mm</quantity>
; maximal pronotal width =
<quantity id="4CB35AE76471FF8D381EFED89DB33DBC" box="[774,907,362,388]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="2" pageNumber="126" unit="mm" value="0.85" valueMax="0.9" valueMin="0.8">0.8-0.9 mm</quantity>
. Body uniformly dark brown, with appendages slightly paler; antennomeres 1-4 paler than antennomeres 5-11. Dorsal pubescence uniform; pubescence on frons converging, with medial setae directed backward; metaventral setae shorter than proventral setae, fairly uniform; antenna with short and dense pubescence present only on antennomeres 5-11; abdominal pubescence converging on tergites IV-V; each sternite bearing a pair of long posteromedial setae. Frons above clypeus forming sharp ridge, the latter not carinate; temples abruptly narrowed just behind eyes, almost smooth; occipital ridge indistinct; pronotum (
<figureCitation id="1370EB876471FF8D3BB8FDF09EDC3E65" box="[160,228,578,605]" captionStart="FIGURES 6-9" captionStartId="3.[151,264,1906,1930]" captionTargetBox="[158,1436,181,1882]" captionTargetId="figure-34@3.[151,1436,181,1882]" captionTargetPageId="3" captionText="FIGURES 6-9. Megarthrus basicornis Fauvel (6-7) and M. budai sp. n. (8-9): prothorax in dorsal (6, 8) and ventral (7, 9) views. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4321574" httpUri="https://zenodo.org/record/4321574/files/figure.png" pageId="2" pageNumber="126">Fig. 6</figureCitation>
) only slightly larger than elytra at base; prohypomeron punctate, lacking transverse ridge (
<figureCitation id="1370EB876471FF8D3FF7FDF09B0D3E64" box="[1263,1333,578,604]" captionStart="FIGURES 6-9" captionStartId="3.[151,264,1906,1930]" captionTargetBox="[158,1436,181,1882]" captionTargetId="figure-34@3.[151,1436,181,1882]" captionTargetPageId="3" captionText="FIGURES 6-9. Megarthrus basicornis Fauvel (6-7) and M. budai sp. n. (8-9): prothorax in dorsal (6, 8) and ventral (7, 9) views. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4321574" httpUri="https://zenodo.org/record/4321574/files/figure.png" pageId="2" pageNumber="126">Fig. 7</figureCitation>
); elytron abruptly narrowed basally.
</paragraph>
<paragraph id="8BF4F7026471FF8D3BDFFD389A6B3F29" blockId="2.[151,1437,326,820]" pageId="2" pageNumber="126">
<emphasis id="B93F2B106471FF8D3BDFFD389F393E9C" box="[199,257,650,676]" italics="true" pageId="2" pageNumber="126">Male</emphasis>
. Frontoclypeal area, protarsomere 5, metasternum and abdominal sternites IV-VI unmodified. Protarsomere 1 without ventral patch of modified adhesive setae (
<figureCitation id="1370EB876471FF8D380FFD1C9DAB3EF0" box="[791,915,686,712]" captionStart="FIGURES 10-13" captionStartId="4.[151,264,1812,1836]" captionTargetBox="[151,1436,848,1788]" captionTargetId="figure-317@4.[151,1436,848,1788]" captionTargetPageId="4" captionText="FIGURES 10-13. Megarthrus basicornis Fauvel (10-11) and M. budai sp. n. (12-13): Portarsus of male in ventrolateral (10, 12) and ventral (11, 13) views. Ovals indicate absence (red) or presence (green) of ventral patch of modified adhesive setae (green arrows). Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4321576" httpUri="https://zenodo.org/record/4321576/files/figure.png" pageId="2" pageNumber="126">Figs 10-11</figureCitation>
). Mesofemur about as long as metafemur. Mesotibia about as long as metatibia. Peg-like setae absent from protibia, metatrochanter and metafemur, and arranged in a single row on mesotrochanter, mesotibia, and metatibia. Aedeagus as in
<figureCitation id="1370EB876471FF8D38E8FD449A683F29" box="[1008,1104,758,785]" captionStart="FIGURES 4-5" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[161,1428,857,1910]" captionTargetId="figure-245@2.[151,1436,844,1920]" captionTargetPageId="2" captionText="FIGURES 4-5. Megarthrus basicornis Fauvel, male: aedeagus in lateral (4) and ventral (5) views. Scale bar = 0.35 mm." figureDoi="http://doi.org/10.5281/zenodo.4321569" httpUri="https://zenodo.org/record/4321569/files/figure.png" pageId="2" pageNumber="126">Figs 4-5</figureCitation>
.
</paragraph>
<paragraph id="8BF4F7026471FF8D3BDFFCA89DAF3F0C" blockId="2.[151,1437,326,820]" box="[199,919,794,820]" pageId="2" pageNumber="126">
<emphasis id="B93F2B106471FF8D3BDFFCA89F233F0C" box="[199,283,794,820]" italics="true" pageId="2" pageNumber="126">Female</emphasis>
. Abdominal tergite VIII without medioapical projection.
</paragraph>
</subSubSection>
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<paragraph id="8BF4F7026471FF8D3B8FF8259B663B97" blockId="2.[151,1374,1943,1968]" box="[151,1374,1943,1968]" pageId="2" pageNumber="126">
<emphasis id="B93F2B106471FF8D3B8FF8259F0E3B88" bold="true" box="[151,310,1943,1968]" pageId="2" pageNumber="126">FIGURES 4-5.</emphasis>
<taxonomicName id="4C4B8C816471FF8D3A25F82A9C4A3B88" authority="Fauvel" authorityName="Fauvel" authorityYear="1904" box="[317,626,1944,1968]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="126" phylum="Arthropoda" rank="species" species="basicornis">
<emphasis id="B93F2B106471FF8D3A25F82A9C1D3B88" box="[317,549,1944,1968]" italics="true" pageId="2" pageNumber="126">Megarthrus basicornis</emphasis>
Fauvel
</taxonomicName>
, male: aedeagus in lateral (4) and ventral (5) views. Scale bar = 0.35 mm.
</paragraph>
</caption>
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<paragraph id="8BF4F7026470FF8C3B8FF8C09F5C3B95" blockId="3.[151,1437,1906,1966]" pageId="3" pageNumber="127">
<emphasis id="B93F2B106470FF8C3B8FF8C09F0D3BB2" bold="true" box="[151,309,1906,1930]" pageId="3" pageNumber="127">FIGURES 6-9.</emphasis>
<taxonomicName id="4C4B8C816470FF8C3A22F8C09C543BB2" authority="Fauvel" authorityName="Fauvel" authorityYear="1904" box="[314,620,1906,1930]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="127" phylum="Arthropoda" rank="species" species="basicornis">
<emphasis id="B93F2B106470FF8C3A22F8C09C1A3BB2" box="[314,546,1906,1930]" italics="true" pageId="3" pageNumber="127">Megarthrus basicornis</emphasis>
Fauvel
</taxonomicName>
(6-7) and
<taxonomicName id="4C4B8C816470FF8C39CBF8C19D163BB2" authority="Liu &amp; Cuccodoro, 2020" authorityName="Liu &amp; Cuccodoro" authorityYear="2020" box="[723,814,1906,1930]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="127" phylum="Arthropoda" rank="species" species="budai" status="sp. nov.">
<emphasis id="B93F2B106470FF8C39CBF8C19D163BB2" box="[723,814,1906,1930]" italics="true" pageId="3" pageNumber="127">M. budai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20C966B6470FF8C382AF8C09D503BB1" box="[818,872,1906,1929]" pageId="3" pageNumber="127" rank="species">sp. n.</taxonomicNameLabel>
(8-9): prothorax in dorsal (6, 8) and ventral (7, 9) views. Scale bar = 0.2 mm.
</paragraph>
</caption>
<subSubSection id="C351A4896477FF8B3BDFFF249C763D24" pageId="4" pageNumber="128" type="materials_examined">
<paragraph id="8BF4F7026477FF8B3BDFFF249C763D24" blockId="4.[151,1436,150,825]" pageId="4" pageNumber="128">
<emphasis id="B93F2B106477FF8B3BDFFF249C0F3C88" bold="true" box="[199,567,150,176]" pageId="4" pageNumber="128">Additional material examined</emphasis>
(
<specimenCount id="9D4D3C8B6477FF8B3954FF249CD83C89" box="[588,736,150,177]" pageId="4" pageNumber="128" type="generic">2 specimens</specimenCount>
).
<materialsCitation id="3B23FD5F6477FF8B39EFFF249C723D24" ID-GBIF-Occurrence="2994520304" collectingDate="2006-06-04" collectingDateMax="2009-09-14" collectingDateMin="2006-06-04" collectionCode="NMPC" collectorName="M. Schulke" country="China" elevation="2480" latitude="25.17097" location="Wuliang Shan" longLatPrecision="1" longitude="100.23939" pageId="4" pageNumber="128" specimenCount="2" specimenCount-male="2" stateProvince="Yunnan">
<collectingCountry id="F35CB7926477FF8B3819FF249D713C88" box="[769,841,150,176]" name="China" pageId="4" pageNumber="128">China</collectingCountry>
,
<collectingRegion id="498F39E06477FF8B3841FF259D8A3C89" box="[857,946,151,177]" country="China" name="Yunnan" pageId="4" pageNumber="128">Yunnan</collectingRegion>
, Dali Bai Aut. Pref.,
<location id="8E94A1D96477FF8B3FACFF249B623C88" LSID="urn:lsid:plazi:treatment:03E246146471FF8B3B8FFF259C303F01:8E94A1D96477FF8B3FACFF249B623C88" box="[1204,1370,150,176]" country="China" latitude="25.17097" longLatPrecision="1" longitude="100.23939" name="Wuliang Shan" pageId="4" pageNumber="128" stateProvince="Yunnan">Wuliang Shan</location>
,
<locationDeviation id="9A46EE206477FF8B3E72FF249F163CEC" location="Weishan" pageId="4" pageNumber="128">
<quantity id="4CB35AE76477FF8B3E72FF249BA43C88" box="[1386,1436,150,176]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="128" unit="km" value="9.0">9km</quantity>
SW Weishan
</locationDeviation>
,
<geoCoordinate id="EE7F91C56477FF8B3A24FF089FD93CED" box="[316,481,186,213]" degrees="25" direction="north" minutes="10" orientation="latitude" pageId="4" pageNumber="128" precision="1" seconds="15.5" value="25.17097">25°1015.5N</geoCoordinate>
,
<geoCoordinate id="EE7F91C56477FF8B3AE8FF089C983CEC" box="[496,672,186,212]" degrees="100" direction="east" minutes="14" orientation="longitude" pageId="4" pageNumber="128" precision="1" seconds="21.8" value="100.23939">100°1421.8E</geoCoordinate>
,
<quantity id="4CB35AE76477FF8B39B7FF089D383CEC" box="[687,768,186,212]" metricMagnitude="3" metricUnit="m" metricValue="2.48" pageId="4" pageNumber="128" unit="m" value="2480.0">
<elevation id="006610316477FF8B39B7FF089D383CEC" box="[687,768,186,212]" metricMagnitude="3" metricUnit="m" metricValue="2.48" pageId="4" pageNumber="128" unit="m" value="2480.0">2480m</elevation>
</quantity>
, scrub with (oak, alder, pine), litter &amp; mushrooms sifted,
<date id="FFF5D1C26477FF8B3B8FFF6C9F213CC0" box="[151,281,222,248]" pageId="4" pageNumber="128" value="2009-09-14">
<collectingDate id="EFB1282A6477FF8B3B8FFF6C9F213CC0" box="[151,281,222,248]" pageId="4" pageNumber="128" value="2009-09-14">14.IX.2009</collectingDate>
</date>
, leg.,
<collectorName id="26BE92D46477FF8B3A78FF6D9FDB3CC0" box="[352,483,222,249]" pageId="4" pageNumber="128">M. Schülke</collectorName>
[CH09-51]”,
<specimenCount id="9D4D3C8B6477FF8B399BFF6C9CE83CC0" box="[643,720,222,248]" pageId="4" pageNumber="128" type="male">1 male</specimenCount>
in cSch; “
<collectingCountry id="F35CB7926477FF8B3851FF6C9DA93CC0" box="[841,913,222,248]" name="China" pageId="4" pageNumber="128">China</collectingCountry>
,
<collectingRegion id="498F39E06477FF8B3885FF6D9DCE3CC1" box="[925,1014,223,249]" country="China" name="Yunnan" pageId="4" pageNumber="128">Yunnan</collectingRegion>
, Gaoligong Shan, pass SW Baoshan,
<date id="FFF5D1C26477FF8B3B8FFEB09F113D24" box="[151,297,258,284]" pageId="4" pageNumber="128" value="2006-06-04" valueMax="2006-06-06" valueMin="2006-06-04">
<collectingDate id="EFB1282A6477FF8B3B8FFEB09F113D24" box="[151,297,258,284]" pageId="4" pageNumber="128" value="2006-06-04" valueMax="2006-06-06" valueMin="2006-06-04">4.-6.VI.2006</collectingDate>
</date>
, Jeniš”,
<specimenCount id="9D4D3C8B6477FF8B3A9FFEB09FEA3D24" box="[391,466,258,284]" pageId="4" pageNumber="128" type="male">1 male</specimenCount>
in
<collectionCode id="ED5A6FC76477FF8B3AEFFEB09C723D24" box="[503,586,258,284]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="4" pageNumber="128">NMPC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C351A4896477FF8B3BDFFE949C303F01" pageId="4" pageNumber="128" type="distribution">
<paragraph id="8BF4F7026477FF8B3BDFFE949AAA3D94" blockId="4.[151,1436,150,825]" pageId="4" pageNumber="128">
<emphasis id="B93F2B106477FF8B3BDFFE949C703D78" bold="true" box="[199,584,294,320]" pageId="4" pageNumber="128">Distribution and natural history.</emphasis>
The species was known so far only from in
<collectingCountry id="F35CB7926477FF8B3F3EFE949A583D78" box="[1062,1120,294,320]" name="India" pageId="4" pageNumber="128">India</collectingCountry>
(
<collectingRegion id="498F39E06477FF8B3F74FE949B043D78" box="[1132,1340,294,320]" country="India" name="Himachal Pradesh" pageId="4" pageNumber="128">Himachal Pradesh</collectingRegion>
,
<collectingRegion id="498F39E06477FF8B3E5DFE949EF03D5C" country="India" name="Meghalaya" pageId="4" pageNumber="128">Meghalaya</collectingRegion>
, and
<collectingRegion id="498F39E06477FF8B3A1DFEF99FAD3D5D" box="[261,405,330,357]" country="India" name="West Bengal" pageId="4" pageNumber="128">West Bengal</collectingRegion>
) and
<collectingCountry id="F35CB7926477FF8B3ACDFEF89C203D5C" box="[469,536,330,356]" name="Nepal" pageId="4" pageNumber="128">Nepal</collectingCountry>
(Mid-Western, Western, and Central Regions) (
<bibRefCitation id="EFDA8AF36477FF8B3F2CFEF89AFD3D5C" author="Fauvel, A." box="[1076,1221,330,356]" pageId="4" pageNumber="128" pagination="76 - 112" refId="ref2837" refString="Fauvel, A. (1904) Staphylinides exotiques nouveaux, 2. Revue d'entomologie, 23, 76 - 112." type="journal article" year="1904">Fauvel 1904</bibRefCitation>
,
<bibRefCitation id="EFDA8AF36477FF8B3FC9FEF89BA83D5C" author="Cuccodoro, G." box="[1233,1424,330,356]" pageId="4" pageNumber="128" refId="ref2715" refString="Cuccodoro, G. (2003) A revision of Megarthrus Curtis of India, Nepal and Sri Lanka (Coleoptera, Staphylinidae, Proteininae). In: Cuccodoro, G. &amp; Leschen, R. A. B. (Eds.), Systematics of Coleoptera: Papers celebrating the retirement of Ivan Lobl. Memoirs on Entomology. Vol. 17. International, Gainesville, pp. 359 - 455." type="book" year="2003">Cuccodoro 2003</bibRefCitation>
), and it is reported here for the first time from
<collectingCountry id="F35CB7926477FF8B399FFEDC9CF23DB0" box="[647,714,366,392]" name="China" pageId="4" pageNumber="128">China</collectingCountry>
(
<collectingRegion id="498F39E06477FF8B39C0FEDC9DA33DB1" box="[728,923,366,393]" country="China" name="Yunnan" pageId="4" pageNumber="128">Yunnan Province</collectingRegion>
) (
<figureCitation id="1370EB876477FF8B38A9FEDC9DC63DB0" box="[945,1022,366,392]" captionStart="FIGURE 39" captionStartId="9.[151,250,1121,1145]" captionTargetBox="[151,1436,345,1061]" captionTargetId="figure-102@9.[151,1436,345,1061]" captionTargetPageId="9" captionText="FIGURE 39. Distribution map of Megarthrus basicornis Fauvel (pink squares) and Megarthrus budai sp. n. (red circle). Scale bar =200 km." figureDoi="http://doi.org/10.5281/zenodo.4321586" httpUri="https://zenodo.org/record/4321586/files/figure.png" pageId="4" pageNumber="128">Fig 39</figureCitation>
). It occurs at elevations ranging from
<quantity id="4CB35AE76477FF8B3B8FFE209F703D95" box="[151,328,402,429]" metricMagnitude="3" metricUnit="m" metricValue="2.35" metricValueMax="2.95" metricValueMin="1.75" pageId="4" pageNumber="128" unit="m" value="2350.0" valueMax="2950.0" valueMin="1750.0">1750 to 2950 m</quantity>
a.s.l., mainly in samples of forest leaf litter with decaying wood and fungi.
</paragraph>
<paragraph id="8BF4F7026477FF8B3BDFFE049F2A3E04" blockId="4.[151,1436,150,825]" pageId="4" pageNumber="128">
<emphasis id="B93F2B106477FF8B3BDFFE049C5B3DE8" bold="true" box="[199,611,438,464]" pageId="4" pageNumber="128">Comparisons and diagnostic notes.</emphasis>
<taxonomicName id="4C4B8C816477FF8B3971FE049D503DE8" authorityName="Fauvel" authorityYear="1904" box="[617,872,438,464]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="128" phylum="Arthropoda" rank="species" species="basicornis">
<emphasis id="B93F2B106477FF8B3971FE049D503DE8" box="[617,872,438,464]" italics="true" pageId="4" pageNumber="128">Megarthrus basicornis</emphasis>
</taxonomicName>
may be easily distinguished from the other
<taxonomicName id="4C4B8C816477FF8B3E51FE059EE93DCC" baseAuthorityName="Cuccodoro" baseAuthorityYear="2011" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B93F2B106477FF8B3E51FE059EE93DCC" italics="true" pageId="4" pageNumber="128">Megarthrus</emphasis>
</taxonomicName>
species known so far to occur in
<collectingCountry id="F35CB7926477FF8B395AFE689CBD3DCC" box="[578,645,474,500]" name="China" pageId="4" pageNumber="128">China</collectingCountry>
by its uniformly dark brown body color, in combination with the elytra abruptly narrowed basally, and the abdominal pubescence converging on tergites IV-V. Its sexual characters are also diagnostic.
</paragraph>
<paragraph id="8BF4F7026477FF8B3BDFFDF49C303F01" blockId="4.[151,1436,150,825]" pageId="4" pageNumber="128">
This species is particularly notable because on the one hand it possesses many features found elsewhere in the genus almost exclusively in members of the
<taxonomicName id="4C4B8C816477FF8B399FFDD99D213EBC" baseAuthorityName="Paykull" baseAuthorityYear="1789" box="[647,793,618,644]" class="Insecta" family="Staphylinidae" genus="Megarthrus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="128" phylum="Arthropoda" rank="species" species="depressus">
<emphasis id="B93F2B106477FF8B399FFDD99D213EBC" box="[647,793,618,644]" italics="true" pageId="4" pageNumber="128">M. depressus</emphasis>
</taxonomicName>
supergroup of species, such as the proventral pubescence longer than the metaventral pubescence and the presence of a pair of long posteromedial setae on each abdominal sternite in combination with the absence of a prohypomeral ridge, but on the other hand it lacks a ventral patch of modified adhesive setae on the male first protarsomere (
<figureCitation id="1370EB876477FF8B3800FD649DAE3EC8" box="[792,918,726,752]" captionStart="FIGURES 10-13" captionStartId="4.[151,264,1812,1836]" captionTargetBox="[151,1436,848,1788]" captionTargetId="figure-317@4.[151,1436,848,1788]" captionTargetPageId="4" captionText="FIGURES 10-13. Megarthrus basicornis Fauvel (10-11) and M. budai sp. n. (12-13): Portarsus of male in ventrolateral (10, 12) and ventral (11, 13) views. Ovals indicate absence (red) or presence (green) of ventral patch of modified adhesive setae (green arrows). Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4321576" httpUri="https://zenodo.org/record/4321576/files/figure.png" pageId="4" pageNumber="128">Figs 10-11</figureCitation>
), which is the only character uniquely shared by all the members of this supergroup of species. The presence of a tuft of long setae at base of the aedeagal dorsal valve is unique within the genus.
</paragraph>
</subSubSection>
</treatment>
</document>