518 lines
78 KiB
XML
518 lines
78 KiB
XML
<document ID-DOI="10.11646/zootaxa.5264.1.9" ID-GBIF-Dataset="75639f12-bb22-4f92-9964-d92af5f7fab0" ID-ISSN="1175-5326" ID-Zenodo-Dep="7836296" ID-ZooBank="AD7AABA8-17CB-4AA3-A180-8781C4924ECA" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1681726657704" checkinUser="plazi" docAuthor="Shapovalov, Andrey M. & Skrylnyk, Yuriy E." docDate="2023" docId="E21C5A06FFA15853FEECFE63AEB5FE27" docLanguage="en" docName="zootaxa.5264.1.9.pdf" docOrigin="Zootaxa 5264 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5264.1.9" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calydus yaroslavi Shapovalov & Skrylnyk 2023, new species" docType="treatment" docVersion="4" lastPageNumber="136" masterDocId="1E25227EFFA05854FE7BFFF1AA2DFFAE" masterDocTitle="The occurrence of the genus Calydus Reitter, 1896 (Coleoptera, Meloidae) in Afghanistan with the description of a new species" masterLastPageNumber="136" masterPageNumber="129" pageNumber="130" updateTime="1681752524762" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The occurrence of the genus Calydus Reitter, 1896 (Coleoptera, Meloidae) in Afghanistan with the description of a new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shapovalov, Andrey M.</mods:namePart>
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<mods:affiliation>Institute of Zoology RK, 93 Al-Farabi Str., 050060, Almaty, Kazakhstan.</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Skrylnyk, Yuriy E.</mods:namePart>
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<mods:affiliation>Institute of Zoology RK, 93 Al-Farabi Str., 050060, Almaty, Kazakhstan.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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||
<mods:titleInfo>
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||
<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-04-12</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5264</mods:number>
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</mods:detail>
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||
<mods:detail type="issue">
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<mods:number>1</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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<mods:start>129</mods:start>
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<mods:end>136</mods:end>
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</mods:extent>
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</mods:part>
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||
</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5264.1.9</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5264.1.9</mods:identifier>
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<mods:identifier type="GBIF-Dataset">75639f12-bb22-4f92-9964-d92af5f7fab0</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7836296</mods:identifier>
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<mods:identifier type="ZooBank">AD7AABA8-17CB-4AA3-A180-8781C4924ECA</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7836298" ID-Zenodo-Dep="7836298" LSID="urn:lsid:plazi:treatment:E21C5A06FFA15853FEECFE63AEB5FE27" httpUri="http://treatment.plazi.org/id/E21C5A06FFA15853FEECFE63AEB5FE27" lastPageId="7" lastPageNumber="136" pageId="1" pageNumber="130">
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<subSubSection box="[151,517,402,429]" pageId="1" pageNumber="130" type="nomenclature">
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||
<paragraph blockId="1.[151,567,402,465]" box="[151,517,402,429]" pageId="1" pageNumber="130">
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||
<heading bold="true" box="[151,517,402,429]" fontSize="11" level="1" pageId="1" pageNumber="130" reason="1">
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||
<emphasis bold="true" box="[151,517,402,429]" pageId="1" pageNumber="130">
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||
<taxonomicName authority="Shapovalov & Skrylnyk, 2023" authorityName="Shapovalov & Skrylnyk" authorityYear="2023" box="[151,365,402,429]" class="Insecta" family="Meloidae" genus="Calydus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="130" phylum="Arthropoda" rank="species" species="yaroslavi" status="new species">
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<emphasis bold="true" box="[151,365,402,429]" italics="true" pageId="1" pageNumber="130">Calydus yaroslavi</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel box="[373,517,402,429]" pageId="1" pageNumber="130" rank="species">new species</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection box="[151,567,438,465]" pageId="1" pageNumber="130" type="description">
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||
<paragraph blockId="1.[151,567,402,465]" box="[151,567,438,465]" pageId="1" pageNumber="130">
|
||
(
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||
<figureCitation box="[159,267,439,465]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[321,1266,192,1911]" captionTargetId="figure-17@3.[307,1281,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Calydus spp., general view. 1—C. yaroslavi new species (holotype, male); 2—C. yaroslavi new species (paratype, female); 3—C. semenovi (Escherich, 1896) (male, Iran, West Azerbaijan Prov.); 4—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.)." figureDoi="http://doi.org/10.5281/zenodo.7836300" httpUri="https://zenodo.org/record/7836300/files/figure.png" pageId="1" pageNumber="130">Figs. 1, 2</figureCitation>
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,
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<figureCitation box="[279,558,438,465]" captionStart="FIGURES 5–19" captionStartId="4.[151,264,1807,1831]" captionTargetBox="[249,1334,191,1779]" captionTargetId="figure-27@4.[249,1338,181,1784]" captionTargetPageId="4" captionText="FIGURES 5–19. Calydus spp. 5, 7, 9, 11, 13, 15–18—C. yaroslavi new species (5, 7, 9, 11, 17–19—holotype, male; 13— paratype, male; 15—paratype, female); 6, 8, 10, 12, 16—C. semenovi (Escherich, 1896) (6, 8, 10, 12—male, 16—female; Iran, West Azerbaijan Prov.); 14—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.). 5, 6—head; 7, 8—right mandible; 9, 10 pronotum; 11, 12—antennae; 13 (the arrow indicates medial elevation), 14 (the arrow indicates elevated ventrolateral margin of pronotum)—prosternum; 15, 16—mesoventrite; 17—tegmen, ventral view; 18—tegmen, lateral view; 19—aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7838276" httpUri="https://zenodo.org/record/7838276/files/figure.png" pageId="1" pageNumber="130">5, 7, 9, 11, 13, 15, 17–18</figureCitation>
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||
)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="1" pageNumber="130" type="materials_examined">
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||
<paragraph blockId="1.[151,1437,510,2013]" pageId="1" pageNumber="130">
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||
<emphasis bold="true" box="[151,314,510,537]" pageId="1" pageNumber="130">
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||
<typeStatus box="[151,210,511,537]" pageId="1" pageNumber="130">Type</typeStatus>
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locality.
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||
</emphasis>
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||
<collectingCounty box="[325,407,511,537]" pageId="1" pageNumber="130">Central</collectingCounty>
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||
<collectingCountry box="[417,557,511,537]" name="Afghanistan" pageId="1" pageNumber="130">Afghanistan</collectingCountry>
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||
,
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||
<collectingRegion box="[572,779,511,537]" country="Afghanistan" name="Bamyan" pageId="1" pageNumber="130">Bamyan Province</collectingRegion>
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||
,
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||
<collectingMunicipality box="[794,995,511,537]" pageId="1" pageNumber="130">Yakawlang Distr.</collectingMunicipality>
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||
,
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||
<location LSID="urn:lsid:plazi:treatment:E21C5A06FFA15853FEECFE63AEB5FE27:6F6ABDCBFFA15855FD89FE0EAF06FDB7" box="[1010,1323,511,537]" country="Afghanistan" county="Central" latitude="34.79952" longLatPrecision="1" longitude="67.18088" municipality="Yakawlang Distr." name="Bande-Amir National Park" pageId="1" pageNumber="130" stateProvince="Bamyan">Bande-Amir National Park</location>
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||
,
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||
<locationDeviation location="Sabzel" pageId="1" pageNumber="130">
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||
<quantity box="[1338,1400,511,537]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="130" unit="km" value="3.0">3 km</quantity>
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||
W Sabzel
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||
</locationDeviation>
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||
,
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||
<geoCoordinate box="[240,411,548,575]" degrees="34" direction="north" minutes="47" orientation="latitude" pageId="1" pageNumber="130" precision="1" seconds="58.27" value="34.79952">34°47′58.27″N</geoCoordinate>
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||
,
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||
<geoCoordinate box="[422,590,548,575]" degrees="67" direction="east" minutes="10" orientation="longitude" pageId="1" pageNumber="130" precision="1" seconds="51.17" value="67.18088">67°10′51.17″E</geoCoordinate>
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||
,
|
||
<elevation box="[601,746,547,573]" metricMagnitude="3" metricUnit="m" metricValue="2.903" pageId="1" pageNumber="130" unit="m" value="2903.0">
|
||
<quantity box="[601,686,547,573]" metricMagnitude="3" metricUnit="m" metricValue="2.903" pageId="1" pageNumber="130" unit="m" value="2903.0">2903 m</quantity>
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||
a.s.l.
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||
</elevation>
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||
</paragraph>
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||
<paragraph blockId="1.[151,1437,510,2013]" pageId="1" pageNumber="130">
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||
<emphasis bold="true" box="[199,367,582,609]" pageId="1" pageNumber="130">Type material</emphasis>
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||
(
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||
<specimenCount box="[385,530,583,609]" pageId="1" pageNumber="130" type="generic">8 specimens</specimenCount>
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||
).
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||
<materialsCitation collectingDate="2009-07-06" collectingDateMax="2009-07-08" collectingDateMin="2009-07-06" collectionCode="ZIN" collectorName="Yu. E. Skrylnyk" country="Afghanistan" elevation="2903" latitude="34.79952" location="BandeAmir National Park" longLatPrecision="1" longitude="67.18088" municipality="Yakawlang Distr." pageId="1" pageNumber="130" specimenCount="1" stateProvince="Bamyan Province" typeStatus="holotype">
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||
<typeStatus box="[549,656,583,609]" pageId="1" pageNumber="130">Holotype</typeStatus>
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||
, ♂ (
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||
<collectionCode box="[706,758,583,609]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="130" type="Museum">ZIN</collectionCode>
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||
),
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||
<collectingCountry box="[777,917,583,609]" name="Afghanistan" pageId="1" pageNumber="130">Afghanistan</collectingCountry>
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||
,
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||
<collectingRegion box="[930,1133,583,609]" country="Afghanistan" name="Bamyan" pageId="1" pageNumber="130">Bamyan Province</collectingRegion>
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||
,
|
||
<collectingMunicipality box="[1145,1343,583,609]" pageId="1" pageNumber="130">Yakawlang Distr.</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:E21C5A06FFA15853FEECFE63AEB5FE27:6F6ABDCBFFA15855FB30FDB6AB5EFD2B" country="Afghanistan" latitude="34.79952" longLatPrecision="1" longitude="67.18088" municipality="Yakawlang Distr." name="BandeAmir National Park" pageId="1" pageNumber="130" stateProvince="Bamyan Province">BandeAmir National Park</location>
|
||
,
|
||
<locationDeviation box="[381,548,619,645]" location="Sabzel" pageId="1" pageNumber="130">
|
||
<quantity box="[381,436,619,645]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="130" unit="km" value="3.0">3 km</quantity>
|
||
W Sabzel
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||
</locationDeviation>
|
||
,
|
||
<geoCoordinate box="[558,729,620,647]" degrees="34" direction="north" minutes="47" orientation="latitude" pageId="1" pageNumber="130" precision="1" seconds="58.27" value="34.79952">34°47′58.27″N</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[739,906,620,647]" degrees="67" direction="east" minutes="10" orientation="longitude" pageId="1" pageNumber="130" precision="1" seconds="51.17" value="67.18088">67°10′51.17″E</geoCoordinate>
|
||
, h =
|
||
<elevation box="[956,1098,619,645]" metricMagnitude="3" metricUnit="m" metricValue="2.903" pageId="1" pageNumber="130" unit="m" value="2903.0">
|
||
<quantity box="[956,1039,619,645]" metricMagnitude="3" metricUnit="m" metricValue="2.903" pageId="1" pageNumber="130" unit="m" value="2903.0">2903 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date box="[1109,1257,619,645]" pageId="1" pageNumber="130" value="2009-07-06" valueMax="2009-07-08" valueMin="2009-07-06">
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||
<collectingDate box="[1109,1257,619,645]" pageId="1" pageNumber="130" value="2009-07-06" valueMax="2009-07-08" valueMin="2009-07-06">6-8.VII.2009</collectingDate>
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||
</date>
|
||
,
|
||
<collectorName box="[1266,1437,619,645]" pageId="1" pageNumber="130">Yu.E. Skrylnyk</collectorName>
|
||
leg.
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||
</materialsCitation>
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||
;
|
||
<materialsCitation box="[207,815,655,683]" collectingDate="2009-07-06" collectingDateMax="2009-07-08" collectingDateMin="2009-07-06" collectionCode="ZIN" collectorName="Yu. E. Skrylnyk" country="Afghanistan" elevation="2903" latitude="34.79952" location="BandeAmir National Park" longLatPrecision="1" longitude="67.18088" municipality="Yakawlang Distr." pageId="1" pageNumber="130" specimenCount="9" specimenCount-female="4" specimenCount-male="5" stateProvince="Bamyan" typeStatus="paratype">
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||
<typeStatus box="[207,319,655,681]" pageId="1" pageNumber="130">Paratypes</typeStatus>
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||
,
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||
<specimenCount box="[330,393,655,683]" pageId="1" pageNumber="130" type="male">4 ♂♂</specimenCount>
|
||
and
|
||
<specimenCount box="[447,512,655,681]" pageId="1" pageNumber="130" type="female">3 ♀♀</specimenCount>
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||
:
|
||
<specimenCount box="[524,566,655,683]" pageId="1" pageNumber="130" type="male">1 ♂</specimenCount>
|
||
<specimenCount box="[573,615,655,681]" pageId="1" pageNumber="130" type="female">1 ♀</specimenCount>
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||
, same data (
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<collectionCode box="[755,807,655,681]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="130" type="Museum">ZIN</collectionCode>
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||
)
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||
</materialsCitation>
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||
;
|
||
<materialsCitation box="[827,1106,655,683]" collectingDate="2009-07-06" collectingDateMax="2009-07-08" collectingDateMin="2009-07-06" collectionCode="ASPC" collectorName="Yu. E. Skrylnyk" country="Afghanistan" elevation="2903" latitude="34.79952" location="BandeAmir National Park" longLatPrecision="1" longitude="67.18088" municipality="Yakawlang Distr." pageId="1" pageNumber="130" specimenCount="3" specimenCount-male="3" stateProvince="Bamyan" typeStatus="paratype">
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||
<specimenCount box="[827,889,655,683]" pageId="1" pageNumber="130" type="male">3 ♂♂</specimenCount>
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||
same data (
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||
<collectionCode box="[1022,1098,655,681]" collectionName="Arnd Schroeder" pageId="1" pageNumber="130">ASPC</collectionCode>
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||
)
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||
</materialsCitation>
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||
;
|
||
<materialsCitation box="[1117,1376,655,681]" collectingDate="2009-07-06" collectingDateMax="2009-07-08" collectingDateMin="2009-07-06" collectionCode="YSPC" collectorName="Yu. E. Skrylnyk" country="Afghanistan" elevation="2903" latitude="34.79952" location="BandeAmir National Park" longLatPrecision="1" longitude="67.18088" municipality="Yakawlang Distr." pageId="1" pageNumber="130" specimenCount="1" specimenCount-female="1" stateProvince="Bamyan" typeStatus="paratype">
|
||
<specimenCount box="[1117,1159,655,681]" pageId="1" pageNumber="130" type="female">1 ♀</specimenCount>
|
||
same data (
|
||
<collectionCode box="[1292,1368,655,681]" collectionName="YSPC" pageId="1" pageNumber="130">YSPC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation collectingDate="2012-06-12" collectionCode="ASPC" collectorName="I. G. Pljushtch" country="Afghanistan" elevation="2900" location="Khushkak" pageId="1" pageNumber="130" specimenCount="1" specimenCount-female="1" stateProvince="Bamyan Distr." typeStatus="paratype">
|
||
<specimenCount box="[1387,1429,655,681]" pageId="1" pageNumber="130" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectingCountry box="[151,291,691,717]" name="Afghanistan" pageId="1" pageNumber="130">Afghanistan</collectingCountry>
|
||
,
|
||
<collectingRegion box="[303,470,691,717]" country="Afghanistan" name="Bamyan" pageId="1" pageNumber="130">Bamyan Distr.</collectingRegion>
|
||
,
|
||
<locationDeviation box="[482,683,691,717]" location="Bamyan" pageId="1" pageNumber="130">
|
||
<quantity box="[482,555,691,717]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="130" unit="km" value="10.0">10 km</quantity>
|
||
S Bamyan
|
||
</locationDeviation>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:E21C5A06FFA15853FEECFE63AEB5FE27:6F6ABDCBFFA15855FCC3FD42A907FD63" box="[696,810,691,717]" country="Afghanistan" name="Khushkak" pageId="1" pageNumber="130" stateProvince="Bamyan Distr.">Khushkak</location>
|
||
env., h =
|
||
<elevation box="[925,1073,691,717]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="1" pageNumber="130" unit="m" value="2900.0">
|
||
<quantity box="[925,1012,691,717]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="1" pageNumber="130" unit="m" value="2900.0">2900 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date box="[1087,1218,691,717]" pageId="1" pageNumber="130" value="2012-06-12">
|
||
<collectingDate box="[1087,1218,691,717]" pageId="1" pageNumber="130" value="2012-06-12">12.VI.2012</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName box="[1230,1386,691,717]" pageId="1" pageNumber="130">I.G. Pljushtch</collectorName>
|
||
leg. (
|
||
<collectionCode box="[159,235,727,753]" collectionName="Arnd Schroeder" pageId="1" pageNumber="130">ASPC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="1" pageNumber="130" type="diagnosis">
|
||
<paragraph blockId="1.[151,1437,510,2013]" pageId="1" pageNumber="130">
|
||
<emphasis bold="true" box="[199,322,762,788]" pageId="1" pageNumber="130">Diagnosis.</emphasis>
|
||
Body black, elytra with faint purple metallic tinge, pronotum orange with black margins and a pair of paramedian round black spots on disc, frontal red spot not distinctly swollen. Head not distinctly depressed dorsally; temples relatively widened and progressively rounded posteriorly; submentum almost flat, only with weak longitudinal paramedian impressions; mandibles robust, relatively wide, external outline strongly curved inward at apical third; antennae moniliform, rather thickened distally, distal antennomeres not approached, distinctly extending beyond the base of elytra; pronotum slightly (about 1.1 times) wider than long, subglobular, surface almost smooth with only fine scattered punctures; ventrolateral margin of pronotum almost evenly merging with margin of prosternum, not forming suture reaching beyond anterior outline of procoxal cavities; prosternum before the middle with sharp laterally flattened medial elevation between procoxal cavities; shoulders nearly rectangular; elytra with faint purple tinge; legs and tarsi slender; body with comparatively long pubescence; head, pronotum and elytra with distinct erect hairs; parameres relatively narrow with dorsal hook before apex.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="2" lastPageNumber="131" pageId="1" pageNumber="130" type="description">
|
||
<paragraph blockId="1.[151,1437,510,2013]" box="[199,1225,1158,1187]" pageId="1" pageNumber="130">
|
||
<emphasis bold="true" box="[199,531,1158,1185]" pageId="1" pageNumber="130">
|
||
Description. Male (
|
||
<typeStatus box="[427,531,1158,1184]" pageId="1" pageNumber="130">holotype</typeStatus>
|
||
</emphasis>
|
||
♂). Body length 14.5 mm, elytral width at shoulders 3.9 mm.
|
||
</paragraph>
|
||
<paragraph blockId="1.[151,1437,510,2013]" pageId="1" pageNumber="130">
|
||
<emphasis box="[199,257,1195,1221]" italics="true" pageId="1" pageNumber="130">Body</emphasis>
|
||
(
|
||
<figureCitation box="[271,334,1195,1221]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[321,1266,192,1911]" captionTargetId="figure-17@3.[307,1281,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Calydus spp., general view. 1—C. yaroslavi new species (holotype, male); 2—C. yaroslavi new species (paratype, female); 3—C. semenovi (Escherich, 1896) (male, Iran, West Azerbaijan Prov.); 4—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.)." figureDoi="http://doi.org/10.5281/zenodo.7836300" httpUri="https://zenodo.org/record/7836300/files/figure.png" pageId="1" pageNumber="130">Fig. 1</figureCitation>
|
||
) black, shiny, elytra with faint purple metallic tinge, pronotum red with black margins and a pair of paramedian round black spots on disc. Body pubescence unicolourous black; hairs erect or suberect and relatively long on ventral side of body, head, pronotum (but central part of disc almost glabrous), abdomen, external margin and base of elytra, femora; antennomeres 1–2 with rather long and erect setation, antennomeres 3–6 with short recumbent pubescence, antennomeres 7–11 with very short and dense recumbent pubescence; underside of tarsi with numerous, but not dense black setae, not forming distinct pads.
|
||
</paragraph>
|
||
<paragraph blockId="1.[151,1437,510,2013]" lastBlockId="2.[151,1437,151,2013]" lastPageId="2" lastPageNumber="131" pageId="1" pageNumber="130">
|
||
<emphasis box="[199,260,1411,1437]" italics="true" pageId="1" pageNumber="130">Head</emphasis>
|
||
capsule (
|
||
<figureCitation box="[365,430,1411,1437]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[321,1266,192,1911]" captionTargetId="figure-17@3.[307,1281,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Calydus spp., general view. 1—C. yaroslavi new species (holotype, male); 2—C. yaroslavi new species (paratype, female); 3—C. semenovi (Escherich, 1896) (male, Iran, West Azerbaijan Prov.); 4—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.)." figureDoi="http://doi.org/10.5281/zenodo.7836300" httpUri="https://zenodo.org/record/7836300/files/figure.png" pageId="1" pageNumber="130">Fig. 1</figureCitation>
|
||
,
|
||
<figureCitation box="[444,458,1410,1437]" captionStart="FIGURES 5–19" captionStartId="4.[151,264,1807,1831]" captionTargetBox="[249,1334,191,1779]" captionTargetId="figure-27@4.[249,1338,181,1784]" captionTargetPageId="4" captionText="FIGURES 5–19. Calydus spp. 5, 7, 9, 11, 13, 15–18—C. yaroslavi new species (5, 7, 9, 11, 17–19—holotype, male; 13— paratype, male; 15—paratype, female); 6, 8, 10, 12, 16—C. semenovi (Escherich, 1896) (6, 8, 10, 12—male, 16—female; Iran, West Azerbaijan Prov.); 14—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.). 5, 6—head; 7, 8—right mandible; 9, 10 pronotum; 11, 12—antennae; 13 (the arrow indicates medial elevation), 14 (the arrow indicates elevated ventrolateral margin of pronotum)—prosternum; 15, 16—mesoventrite; 17—tegmen, ventral view; 18—tegmen, lateral view; 19—aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7838276" httpUri="https://zenodo.org/record/7838276/files/figure.png" pageId="1" pageNumber="130">5</figureCitation>
|
||
) subquadrate, 1.2 times as wide as long at temples level, at temples 1.8 times as wide as at frons between eyes, temples relatively widened and rounded posteriorly, length of temples 1.4 times as great as transverse diameter of eye, longitudinal diameter of eyes 1.8 times as great as transverse diameter, distance between eyes 2.2 times as great as transverse eye diameter and 1.2 times as great as longitudinal eye diameter; frons flattened and weakly depressed proximally, in the middle with suboval depression and red spot; median sulcus narrow and partly shallow, weakly depressed on vertex; head dorsally with dull area between eyes and from anterior margin of head capsule almost to vertex with dense microsculpture and strongly smoothed fine punctures, outside this area surface of head shiny, with more deep punctures, interspace between punctures almost more than punctures. Clypeus transverse, convex, anteriorly depressed; fronto-clypeal suture clearly visible; labrum slightly shorter and narrower than clypeus, at sides arcuate, anterior margin slightly sinuate. Mandibles (
|
||
<figureCitation box="[1087,1152,1735,1761]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[321,1266,192,1911]" captionTargetId="figure-17@3.[307,1281,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Calydus spp., general view. 1—C. yaroslavi new species (holotype, male); 2—C. yaroslavi new species (paratype, female); 3—C. semenovi (Escherich, 1896) (male, Iran, West Azerbaijan Prov.); 4—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.)." figureDoi="http://doi.org/10.5281/zenodo.7836300" httpUri="https://zenodo.org/record/7836300/files/figure.png" pageId="1" pageNumber="130">Fig. 1</figureCitation>
|
||
,
|
||
<figureCitation box="[1165,1180,1735,1761]" captionStart="FIGURES 5–19" captionStartId="4.[151,264,1807,1831]" captionTargetBox="[249,1334,191,1779]" captionTargetId="figure-27@4.[249,1338,181,1784]" captionTargetPageId="4" captionText="FIGURES 5–19. Calydus spp. 5, 7, 9, 11, 13, 15–18—C. yaroslavi new species (5, 7, 9, 11, 17–19—holotype, male; 13— paratype, male; 15—paratype, female); 6, 8, 10, 12, 16—C. semenovi (Escherich, 1896) (6, 8, 10, 12—male, 16—female; Iran, West Azerbaijan Prov.); 14—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.). 5, 6—head; 7, 8—right mandible; 9, 10 pronotum; 11, 12—antennae; 13 (the arrow indicates medial elevation), 14 (the arrow indicates elevated ventrolateral margin of pronotum)—prosternum; 15, 16—mesoventrite; 17—tegmen, ventral view; 18—tegmen, lateral view; 19—aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7838276" httpUri="https://zenodo.org/record/7838276/files/figure.png" pageId="1" pageNumber="130">7</figureCitation>
|
||
) robust, with bidentate apex, interiorly with tooth proximally of the middle, external outline strongly curved from apical third. Submentum almost flat, only with weak paramedian longitudinal impressions. Last maxillary palpomere apically thickened and rounded at apex, external margin weakly emarginate beyond apex. Antennae (
|
||
<figureCitation box="[1014,1080,1843,1869]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[321,1266,192,1911]" captionTargetId="figure-17@3.[307,1281,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Calydus spp., general view. 1—C. yaroslavi new species (holotype, male); 2—C. yaroslavi new species (paratype, female); 3—C. semenovi (Escherich, 1896) (male, Iran, West Azerbaijan Prov.); 4—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.)." figureDoi="http://doi.org/10.5281/zenodo.7836300" httpUri="https://zenodo.org/record/7836300/files/figure.png" pageId="1" pageNumber="130">Fig. 1</figureCitation>
|
||
,
|
||
<figureCitation box="[1093,1120,1843,1869]" captionStart="FIGURES 5–19" captionStartId="4.[151,264,1807,1831]" captionTargetBox="[249,1334,191,1779]" captionTargetId="figure-27@4.[249,1338,181,1784]" captionTargetPageId="4" captionText="FIGURES 5–19. Calydus spp. 5, 7, 9, 11, 13, 15–18—C. yaroslavi new species (5, 7, 9, 11, 17–19—holotype, male; 13— paratype, male; 15—paratype, female); 6, 8, 10, 12, 16—C. semenovi (Escherich, 1896) (6, 8, 10, 12—male, 16—female; Iran, West Azerbaijan Prov.); 14—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.). 5, 6—head; 7, 8—right mandible; 9, 10 pronotum; 11, 12—antennae; 13 (the arrow indicates medial elevation), 14 (the arrow indicates elevated ventrolateral margin of pronotum)—prosternum; 15, 16—mesoventrite; 17—tegmen, ventral view; 18—tegmen, lateral view; 19—aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7838276" httpUri="https://zenodo.org/record/7838276/files/figure.png" pageId="1" pageNumber="130">11</figureCitation>
|
||
) moniliform, comparatively long, extending behind posterior margin of pronotum by apical part of antennomere 9, distal antennomeres distinctly wider than proximal ones, antennomeres 1–6 shiny, antennomeres 7–11 matt with dense sculpture; scape three times as long as pedicel; antennomeres 3–5 greatly elongate, subequal in length and 2.5 times as long as wide, antennomere 3 cylindrical, antennomeres 4–5 subcylindrical (only slightly thickened apically); antennomere 6 short and wider than preceding ones; antennomeres 7–11 rather wide, antennomere 7 slightly wider than 6 and slightly narrower than 8, antennomeres 8–10 wider than others (1.5 times as long as wide and 1.5 times wider than antennomere 3), antennomere 11 slightly narrower than 10, elongate and narrowing apically.
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="131">
|
||
<emphasis box="[199,310,259,284]" italics="true" pageId="2" pageNumber="131">Pronotum</emphasis>
|
||
(
|
||
<figureCitation box="[323,388,259,285]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[321,1266,192,1911]" captionTargetId="figure-17@3.[307,1281,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Calydus spp., general view. 1—C. yaroslavi new species (holotype, male); 2—C. yaroslavi new species (paratype, female); 3—C. semenovi (Escherich, 1896) (male, Iran, West Azerbaijan Prov.); 4—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.)." figureDoi="http://doi.org/10.5281/zenodo.7836300" httpUri="https://zenodo.org/record/7836300/files/figure.png" pageId="2" pageNumber="131">Fig. 1</figureCitation>
|
||
,
|
||
<figureCitation box="[399,414,259,285]" captionStart="FIGURES 5–19" captionStartId="4.[151,264,1807,1831]" captionTargetBox="[249,1334,191,1779]" captionTargetId="figure-27@4.[249,1338,181,1784]" captionTargetPageId="4" captionText="FIGURES 5–19. Calydus spp. 5, 7, 9, 11, 13, 15–18—C. yaroslavi new species (5, 7, 9, 11, 17–19—holotype, male; 13— paratype, male; 15—paratype, female); 6, 8, 10, 12, 16—C. semenovi (Escherich, 1896) (6, 8, 10, 12—male, 16—female; Iran, West Azerbaijan Prov.); 14—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.). 5, 6—head; 7, 8—right mandible; 9, 10 pronotum; 11, 12—antennae; 13 (the arrow indicates medial elevation), 14 (the arrow indicates elevated ventrolateral margin of pronotum)—prosternum; 15, 16—mesoventrite; 17—tegmen, ventral view; 18—tegmen, lateral view; 19—aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7838276" httpUri="https://zenodo.org/record/7838276/files/figure.png" pageId="2" pageNumber="131">9</figureCitation>
|
||
) 1.1 times wider than long, widest near the middle, narrowed anteriorly, convex, subglobular, with moderately depressed middle-line in centre of disc; surface almost smooth with only fine scattered punctures. Ventrolateral margin of pronotum (
|
||
<figureCitation box="[546,627,331,357]" captionStart="FIGURES 5–19" captionStartId="4.[151,264,1807,1831]" captionTargetBox="[249,1334,191,1779]" captionTargetId="figure-27@4.[249,1338,181,1784]" captionTargetPageId="4" captionText="FIGURES 5–19. Calydus spp. 5, 7, 9, 11, 13, 15–18—C. yaroslavi new species (5, 7, 9, 11, 17–19—holotype, male; 13— paratype, male; 15—paratype, female); 6, 8, 10, 12, 16—C. semenovi (Escherich, 1896) (6, 8, 10, 12—male, 16—female; Iran, West Azerbaijan Prov.); 14—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.). 5, 6—head; 7, 8—right mandible; 9, 10 pronotum; 11, 12—antennae; 13 (the arrow indicates medial elevation), 14 (the arrow indicates elevated ventrolateral margin of pronotum)—prosternum; 15, 16—mesoventrite; 17—tegmen, ventral view; 18—tegmen, lateral view; 19—aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7838276" httpUri="https://zenodo.org/record/7838276/files/figure.png" pageId="2" pageNumber="131">Fig. 13</figureCitation>
|
||
) almost evenly merging with margin of prosternum, not forming suture reaching beyond anterior outline of procoxal cavities. Prosternum (
|
||
<figureCitation box="[917,999,367,393]" captionStart="FIGURES 5–19" captionStartId="4.[151,264,1807,1831]" captionTargetBox="[249,1334,191,1779]" captionTargetId="figure-27@4.[249,1338,181,1784]" captionTargetPageId="4" captionText="FIGURES 5–19. Calydus spp. 5, 7, 9, 11, 13, 15–18—C. yaroslavi new species (5, 7, 9, 11, 17–19—holotype, male; 13— paratype, male; 15—paratype, female); 6, 8, 10, 12, 16—C. semenovi (Escherich, 1896) (6, 8, 10, 12—male, 16—female; Iran, West Azerbaijan Prov.); 14—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.). 5, 6—head; 7, 8—right mandible; 9, 10 pronotum; 11, 12—antennae; 13 (the arrow indicates medial elevation), 14 (the arrow indicates elevated ventrolateral margin of pronotum)—prosternum; 15, 16—mesoventrite; 17—tegmen, ventral view; 18—tegmen, lateral view; 19—aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7838276" httpUri="https://zenodo.org/record/7838276/files/figure.png" pageId="2" pageNumber="131">Fig. 13</figureCitation>
|
||
) transverse, near the middle of length with elevated transversal ridge, proximally of the middle with sharp laterally flattened medial elevation between procoxal cavities.
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="131">
|
||
<emphasis box="[199,345,475,500]" italics="true" pageId="2" pageNumber="131">Mesoventrite</emphasis>
|
||
(
|
||
<figureCitation box="[361,443,474,501]" captionStart="FIGURES 5–19" captionStartId="4.[151,264,1807,1831]" captionTargetBox="[249,1334,191,1779]" captionTargetId="figure-27@4.[249,1338,181,1784]" captionTargetPageId="4" captionText="FIGURES 5–19. Calydus spp. 5, 7, 9, 11, 13, 15–18—C. yaroslavi new species (5, 7, 9, 11, 17–19—holotype, male; 13— paratype, male; 15—paratype, female); 6, 8, 10, 12, 16—C. semenovi (Escherich, 1896) (6, 8, 10, 12—male, 16—female; Iran, West Azerbaijan Prov.); 14—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.). 5, 6—head; 7, 8—right mandible; 9, 10 pronotum; 11, 12—antennae; 13 (the arrow indicates medial elevation), 14 (the arrow indicates elevated ventrolateral margin of pronotum)—prosternum; 15, 16—mesoventrite; 17—tegmen, ventral view; 18—tegmen, lateral view; 19—aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7838276" httpUri="https://zenodo.org/record/7838276/files/figure.png" pageId="2" pageNumber="131">Fig. 15</figureCitation>
|
||
) arcuate anteriorly, without modified area (“scutum”), middle part smooth and glabrous (without punctures and hairs); mesepisterna with furrowed anterior border. Abdominal ventrite VI not deeply angularly emarginate in middle of posterior margin, punctuation of abdomen fine and not dense.
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="131">
|
||
<emphasis box="[199,269,583,609]" italics="true" pageId="2" pageNumber="131">Elytra</emphasis>
|
||
(
|
||
<figureCitation box="[287,353,583,609]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[321,1266,192,1911]" captionTargetId="figure-17@3.[307,1281,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Calydus spp., general view. 1—C. yaroslavi new species (holotype, male); 2—C. yaroslavi new species (paratype, female); 3—C. semenovi (Escherich, 1896) (male, Iran, West Azerbaijan Prov.); 4—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.)." figureDoi="http://doi.org/10.5281/zenodo.7836300" httpUri="https://zenodo.org/record/7836300/files/figure.png" pageId="2" pageNumber="131">Fig. 1</figureCitation>
|
||
) at the base 1.6 times as wide as pronotum, shoulders distinct and nearly rectangular. Surface shiny, finely rugose with small and partly merged punctures.
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="131">
|
||
<emphasis box="[199,252,655,680]" italics="true" pageId="2" pageNumber="131">Legs</emphasis>
|
||
(
|
||
<figureCitation box="[272,342,655,681]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[321,1266,192,1911]" captionTargetId="figure-17@3.[307,1281,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Calydus spp., general view. 1—C. yaroslavi new species (holotype, male); 2—C. yaroslavi new species (paratype, female); 3—C. semenovi (Escherich, 1896) (male, Iran, West Azerbaijan Prov.); 4—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.)." figureDoi="http://doi.org/10.5281/zenodo.7836300" httpUri="https://zenodo.org/record/7836300/files/figure.png" pageId="2" pageNumber="131">Fig. 1</figureCitation>
|
||
) slender, tarsomeres distinctly longer than wide in lateral aspect; tarsomeres 1-4 moderately thickened apically in lateral aspect, tarsomere 5 subcylindrical, moderately widened in dorsal aspect; pro- and mesotarsi nearly as long as corresponding tibiae, tarsomere 5 longest, tarsomere 1 slightly shorter than 5, tarsomeres 2 and 3 subequal and shorter than 1, tarsomere 4 shortest; metatarsi 0.9 times the length of metatibia with longest tarsomere 1; pro- and mesotibial spurs simple, apically pointed, the inner metatibial spur slightly flattened and rounded at apex, the external spur pointed; claws slightly curved, with ventral blade fused to dorsal blade proximally and not attaining apex (equal to ¾ of length) of dorsal blade, ventral blade diverging at middle of dorsal blade.
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="131">
|
||
<emphasis box="[199,363,907,933]" italics="true" pageId="2" pageNumber="131">Male genitalia</emphasis>
|
||
(
|
||
<figureCitation box="[376,513,907,933]" captionStart="FIGURES 5–19" captionStartId="4.[151,264,1807,1831]" captionTargetBox="[249,1334,191,1779]" captionTargetId="figure-27@4.[249,1338,181,1784]" captionTargetPageId="4" captionText="FIGURES 5–19. Calydus spp. 5, 7, 9, 11, 13, 15–18—C. yaroslavi new species (5, 7, 9, 11, 17–19—holotype, male; 13— paratype, male; 15—paratype, female); 6, 8, 10, 12, 16—C. semenovi (Escherich, 1896) (6, 8, 10, 12—male, 16—female; Iran, West Azerbaijan Prov.); 14—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.). 5, 6—head; 7, 8—right mandible; 9, 10 pronotum; 11, 12—antennae; 13 (the arrow indicates medial elevation), 14 (the arrow indicates elevated ventrolateral margin of pronotum)—prosternum; 15, 16—mesoventrite; 17—tegmen, ventral view; 18—tegmen, lateral view; 19—aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7838276" httpUri="https://zenodo.org/record/7838276/files/figure.png" pageId="2" pageNumber="131">Figs. 17–19</figureCitation>
|
||
). Parameres relatively narrow with dorsal hook before apex, lobes with dorsal hook before apex and with long setae basal to hook; penis with one apical hook.
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="131">
|
||
<emphasis bold="true" box="[199,445,978,1005]" pageId="2" pageNumber="131">Variability in males.</emphasis>
|
||
Body length 10.9–14.5 mm, width elytra in shoulders 2.6–3.9 mm. Disc of pronotum often with pair of paramedian impressions proximal of the middle.
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="131">
|
||
<emphasis bold="true" box="[199,297,1051,1077]" pageId="2" pageNumber="131">Females</emphasis>
|
||
(
|
||
<figureCitation box="[316,388,1051,1077]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[321,1266,192,1911]" captionTargetId="figure-17@3.[307,1281,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Calydus spp., general view. 1—C. yaroslavi new species (holotype, male); 2—C. yaroslavi new species (paratype, female); 3—C. semenovi (Escherich, 1896) (male, Iran, West Azerbaijan Prov.); 4—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.)." figureDoi="http://doi.org/10.5281/zenodo.7836300" httpUri="https://zenodo.org/record/7836300/files/figure.png" pageId="2" pageNumber="131">Fig. 2</figureCitation>
|
||
,
|
||
<figureCitation box="[403,434,1050,1077]" captionStart="FIGURES 5–19" captionStartId="4.[151,264,1807,1831]" captionTargetBox="[249,1334,191,1779]" captionTargetId="figure-27@4.[249,1338,181,1784]" captionTargetPageId="4" captionText="FIGURES 5–19. Calydus spp. 5, 7, 9, 11, 13, 15–18—C. yaroslavi new species (5, 7, 9, 11, 17–19—holotype, male; 13— paratype, male; 15—paratype, female); 6, 8, 10, 12, 16—C. semenovi (Escherich, 1896) (6, 8, 10, 12—male, 16—female; Iran, West Azerbaijan Prov.); 14—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.). 5, 6—head; 7, 8—right mandible; 9, 10 pronotum; 11, 12—antennae; 13 (the arrow indicates medial elevation), 14 (the arrow indicates elevated ventrolateral margin of pronotum)—prosternum; 15, 16—mesoventrite; 17—tegmen, ventral view; 18—tegmen, lateral view; 19—aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7838276" httpUri="https://zenodo.org/record/7838276/files/figure.png" pageId="2" pageNumber="131">15</figureCitation>
|
||
). Body length: 10.8–17.3 mm. Head without dull area between eyes, with rather deep uniform punctures, shiny, without dense microsculpture; antennae slightly shorter than in males, with antennomere 11 attaining the base of elytra, antennomeres 1–5 shiny, 1–2 with dense and rather long erect pubescence, 3–5 with comparatively long and dense semierect pubescence, antennomeres 6–11 matt with dense sculpture and dense and very short recumbent pubescence. Ventral blade of claws slightly shorter than in males and equal nearly to 2/3 of length of dorsal blade.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="2" pageNumber="131" type="distribution">
|
||
<paragraph blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="131">
|
||
<emphasis bold="true" box="[199,353,1266,1292]" pageId="2" pageNumber="131">Distribution.</emphasis>
|
||
The new species was recorded only from the
|
||
<collectingRegion box="[908,1107,1267,1293]" country="Afghanistan" name="Bamyan" pageId="2" pageNumber="131">Bamian Province</collectingRegion>
|
||
in central
|
||
<collectingCountry box="[1238,1379,1267,1293]" name="Afghanistan" pageId="2" pageNumber="131">Afghanistan</collectingCountry>
|
||
: two localities on the northern slope of Koh-I Baba (or Baba) Mountain Range in the Hindu Kush mountain system (see map,
|
||
<figureCitation box="[213,297,1339,1365]" captionStart="FIGURES 22–23" captionStartId="6.[151,264,1913,1937]" captionTargetBox="[174,1413,182,1875]" captionTargetId="figure-27@6.[174,1413,181,1888]" captionTargetPageId="6" captionText="FIGURES 22–23. Biotope and distribution of Calydus spp. 22—map of distribution Calydus yaroslavi new species (1—type locality, 3 km W Sabzel; 2–10 km S Bamyan); 23—biotope of the Calydus semenovi (Escherich, 1896) in Iran, West Azerbaijan, Takht-e Soleyman, 24.V.2017, 36°37′15.46″N 47°18′36.94″E (photo by A. Zubov)." figureDoi="http://doi.org/10.5281/zenodo.7836304" httpUri="https://zenodo.org/record/7836304/files/figure.png" pageId="2" pageNumber="131">Fig. 22</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="131">
|
||
<emphasis bold="true" box="[199,516,1374,1401]" pageId="2" pageNumber="131">Comparison and remarks.</emphasis>
|
||
The shape of claws of the new species (with ventral blade fused to dorsal blade) and genitalia (lobes of parameres hooked dorsally before apex, penis with one apical hook) are typical of the genus
|
||
<taxonomicName authorityName="Reitter" authorityYear="1896" box="[225,317,1447,1473]" class="Insecta" family="Meloidae" genus="Calydus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[225,317,1447,1473]" italics="true" pageId="2" pageNumber="131">Calydus</emphasis>
|
||
</taxonomicName>
|
||
. However
|
||
<taxonomicName authority="Shapovalov & Skrylnyk, 2023" authorityName="Shapovalov & Skrylnyk" authorityYear="2023" box="[443,580,1447,1473]" class="Insecta" family="Meloidae" genus="Calydus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="yaroslavi" status="new species">
|
||
<emphasis box="[443,580,1447,1473]" italics="true" pageId="2" pageNumber="131">C. yaroslavi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[589,725,1447,1473]" pageId="2" pageNumber="131" rank="species">new species</taxonomicNameLabel>
|
||
is very distinct from all congeners in the shape of the temples (
|
||
<figureCitation box="[159,227,1482,1509]" captionStart="FIGURES 5–19" captionStartId="4.[151,264,1807,1831]" captionTargetBox="[249,1334,191,1779]" captionTargetId="figure-27@4.[249,1338,181,1784]" captionTargetPageId="4" captionText="FIGURES 5–19. Calydus spp. 5, 7, 9, 11, 13, 15–18—C. yaroslavi new species (5, 7, 9, 11, 17–19—holotype, male; 13— paratype, male; 15—paratype, female); 6, 8, 10, 12, 16—C. semenovi (Escherich, 1896) (6, 8, 10, 12—male, 16—female; Iran, West Azerbaijan Prov.); 14—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.). 5, 6—head; 7, 8—right mandible; 9, 10 pronotum; 11, 12—antennae; 13 (the arrow indicates medial elevation), 14 (the arrow indicates elevated ventrolateral margin of pronotum)—prosternum; 15, 16—mesoventrite; 17—tegmen, ventral view; 18—tegmen, lateral view; 19—aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7838276" httpUri="https://zenodo.org/record/7838276/files/figure.png" pageId="2" pageNumber="131">Fig. 5</figureCitation>
|
||
), head dorsally not depressed, mandibles (
|
||
<figureCitation box="[708,775,1483,1509]" captionStart="FIGURES 5–19" captionStartId="4.[151,264,1807,1831]" captionTargetBox="[249,1334,191,1779]" captionTargetId="figure-27@4.[249,1338,181,1784]" captionTargetPageId="4" captionText="FIGURES 5–19. Calydus spp. 5, 7, 9, 11, 13, 15–18—C. yaroslavi new species (5, 7, 9, 11, 17–19—holotype, male; 13— paratype, male; 15—paratype, female); 6, 8, 10, 12, 16—C. semenovi (Escherich, 1896) (6, 8, 10, 12—male, 16—female; Iran, West Azerbaijan Prov.); 14—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.). 5, 6—head; 7, 8—right mandible; 9, 10 pronotum; 11, 12—antennae; 13 (the arrow indicates medial elevation), 14 (the arrow indicates elevated ventrolateral margin of pronotum)—prosternum; 15, 16—mesoventrite; 17—tegmen, ventral view; 18—tegmen, lateral view; 19—aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7838276" httpUri="https://zenodo.org/record/7838276/files/figure.png" pageId="2" pageNumber="131">Fig. 7</figureCitation>
|
||
) with external outline strongly curved inward at the apical third, longer and moniliform antennae (
|
||
<figureCitation box="[602,685,1519,1545]" captionStart="FIGURES 5–19" captionStartId="4.[151,264,1807,1831]" captionTargetBox="[249,1334,191,1779]" captionTargetId="figure-27@4.[249,1338,181,1784]" captionTargetPageId="4" captionText="FIGURES 5–19. Calydus spp. 5, 7, 9, 11, 13, 15–18—C. yaroslavi new species (5, 7, 9, 11, 17–19—holotype, male; 13— paratype, male; 15—paratype, female); 6, 8, 10, 12, 16—C. semenovi (Escherich, 1896) (6, 8, 10, 12—male, 16—female; Iran, West Azerbaijan Prov.); 14—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.). 5, 6—head; 7, 8—right mandible; 9, 10 pronotum; 11, 12—antennae; 13 (the arrow indicates medial elevation), 14 (the arrow indicates elevated ventrolateral margin of pronotum)—prosternum; 15, 16—mesoventrite; 17—tegmen, ventral view; 18—tegmen, lateral view; 19—aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7838276" httpUri="https://zenodo.org/record/7838276/files/figure.png" pageId="2" pageNumber="131">Fig. 11</figureCitation>
|
||
), subglobular and almost smooth pronotum (
|
||
<figureCitation box="[1198,1268,1519,1545]" captionStart="FIGURES 5–19" captionStartId="4.[151,264,1807,1831]" captionTargetBox="[249,1334,191,1779]" captionTargetId="figure-27@4.[249,1338,181,1784]" captionTargetPageId="4" captionText="FIGURES 5–19. Calydus spp. 5, 7, 9, 11, 13, 15–18—C. yaroslavi new species (5, 7, 9, 11, 17–19—holotype, male; 13— paratype, male; 15—paratype, female); 6, 8, 10, 12, 16—C. semenovi (Escherich, 1896) (6, 8, 10, 12—male, 16—female; Iran, West Azerbaijan Prov.); 14—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.). 5, 6—head; 7, 8—right mandible; 9, 10 pronotum; 11, 12—antennae; 13 (the arrow indicates medial elevation), 14 (the arrow indicates elevated ventrolateral margin of pronotum)—prosternum; 15, 16—mesoventrite; 17—tegmen, ventral view; 18—tegmen, lateral view; 19—aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7838276" httpUri="https://zenodo.org/record/7838276/files/figure.png" pageId="2" pageNumber="131">Fig. 9</figureCitation>
|
||
), distinct erect pubescence of body including pronotum and elytra (
|
||
<figureCitation box="[744,811,1555,1581]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[321,1266,192,1911]" captionTargetId="figure-17@3.[307,1281,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Calydus spp., general view. 1—C. yaroslavi new species (holotype, male); 2—C. yaroslavi new species (paratype, female); 3—C. semenovi (Escherich, 1896) (male, Iran, West Azerbaijan Prov.); 4—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.)." figureDoi="http://doi.org/10.5281/zenodo.7836300" httpUri="https://zenodo.org/record/7836300/files/figure.png" pageId="2" pageNumber="131">Fig. 1</figureCitation>
|
||
). Body coloration is similar only to that of
|
||
<taxonomicName authorityName="Kaszab" authorityYear="1960" box="[1309,1436,1555,1581]" class="Insecta" family="Meloidae" genus="Calydus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="syriacus">
|
||
<emphasis box="[1309,1436,1555,1581]" italics="true" pageId="2" pageNumber="131">C. syriacus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[159,229,1591,1617]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[321,1266,192,1911]" captionTargetId="figure-17@3.[307,1281,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Calydus spp., general view. 1—C. yaroslavi new species (holotype, male); 2—C. yaroslavi new species (paratype, female); 3—C. semenovi (Escherich, 1896) (male, Iran, West Azerbaijan Prov.); 4—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.)." figureDoi="http://doi.org/10.5281/zenodo.7836300" httpUri="https://zenodo.org/record/7836300/files/figure.png" pageId="2" pageNumber="131">Fig. 4</figureCitation>
|
||
), but in latter the elytra totally lack a metallic tinge. Other
|
||
<taxonomicName box="[901,1082,1591,1617]" pageId="2" pageNumber="131">
|
||
<emphasis box="[901,993,1591,1617]" italics="true" pageId="2" pageNumber="131">Calydus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
differ from the new species by the body colouration, having elytra which are totally black or with red, brown-red (
|
||
<figureCitation box="[1092,1159,1627,1653]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[321,1266,192,1911]" captionTargetId="figure-17@3.[307,1281,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Calydus spp., general view. 1—C. yaroslavi new species (holotype, male); 2—C. yaroslavi new species (paratype, female); 3—C. semenovi (Escherich, 1896) (male, Iran, West Azerbaijan Prov.); 4—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.)." figureDoi="http://doi.org/10.5281/zenodo.7836300" httpUri="https://zenodo.org/record/7836300/files/figure.png" pageId="2" pageNumber="131">Fig. 3</figureCitation>
|
||
) or bicoloured. Also, all the previously known species of
|
||
<taxonomicName authorityName="Reitter" authorityYear="1896" box="[523,615,1663,1689]" class="Insecta" family="Meloidae" genus="Calydus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[523,615,1663,1689]" italics="true" pageId="2" pageNumber="131">Calydus</emphasis>
|
||
</taxonomicName>
|
||
are characterised by the temples progressively widened and not rounded posteriorly (
|
||
<figureCitation box="[292,364,1699,1725]" captionStart="FIGURES 5–19" captionStartId="4.[151,264,1807,1831]" captionTargetBox="[249,1334,191,1779]" captionTargetId="figure-27@4.[249,1338,181,1784]" captionTargetPageId="4" captionText="FIGURES 5–19. Calydus spp. 5, 7, 9, 11, 13, 15–18—C. yaroslavi new species (5, 7, 9, 11, 17–19—holotype, male; 13— paratype, male; 15—paratype, female); 6, 8, 10, 12, 16—C. semenovi (Escherich, 1896) (6, 8, 10, 12—male, 16—female; Iran, West Azerbaijan Prov.); 14—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.). 5, 6—head; 7, 8—right mandible; 9, 10 pronotum; 11, 12—antennae; 13 (the arrow indicates medial elevation), 14 (the arrow indicates elevated ventrolateral margin of pronotum)—prosternum; 15, 16—mesoventrite; 17—tegmen, ventral view; 18—tegmen, lateral view; 19—aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7838276" httpUri="https://zenodo.org/record/7838276/files/figure.png" pageId="2" pageNumber="131">Fig. 6</figureCitation>
|
||
), rather short antennae not attaining elytral base and weakly widened distally, antennae not moniliform with antennomeres closely articulated (
|
||
<figureCitation box="[744,848,1735,1761]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[321,1266,192,1911]" captionTargetId="figure-17@3.[307,1281,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Calydus spp., general view. 1—C. yaroslavi new species (holotype, male); 2—C. yaroslavi new species (paratype, female); 3—C. semenovi (Escherich, 1896) (male, Iran, West Azerbaijan Prov.); 4—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.)." figureDoi="http://doi.org/10.5281/zenodo.7836300" httpUri="https://zenodo.org/record/7836300/files/figure.png" pageId="2" pageNumber="131">Fig. 3, 4</figureCitation>
|
||
,
|
||
<figureCitation box="[864,895,1735,1761]" captionStart="FIGURES 5–19" captionStartId="4.[151,264,1807,1831]" captionTargetBox="[249,1334,191,1779]" captionTargetId="figure-27@4.[249,1338,181,1784]" captionTargetPageId="4" captionText="FIGURES 5–19. Calydus spp. 5, 7, 9, 11, 13, 15–18—C. yaroslavi new species (5, 7, 9, 11, 17–19—holotype, male; 13— paratype, male; 15—paratype, female); 6, 8, 10, 12, 16—C. semenovi (Escherich, 1896) (6, 8, 10, 12—male, 16—female; Iran, West Azerbaijan Prov.); 14—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.). 5, 6—head; 7, 8—right mandible; 9, 10 pronotum; 11, 12—antennae; 13 (the arrow indicates medial elevation), 14 (the arrow indicates elevated ventrolateral margin of pronotum)—prosternum; 15, 16—mesoventrite; 17—tegmen, ventral view; 18—tegmen, lateral view; 19—aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7838276" httpUri="https://zenodo.org/record/7838276/files/figure.png" pageId="2" pageNumber="131">12</figureCitation>
|
||
); pronotum almost campaniform with the disc flattened or only weakly convex, integument with rather deep distinct punctuation (
|
||
<figureCitation box="[1083,1179,1771,1797]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[321,1266,192,1911]" captionTargetId="figure-17@3.[307,1281,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Calydus spp., general view. 1—C. yaroslavi new species (holotype, male); 2—C. yaroslavi new species (paratype, female); 3—C. semenovi (Escherich, 1896) (male, Iran, West Azerbaijan Prov.); 4—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.)." figureDoi="http://doi.org/10.5281/zenodo.7836300" httpUri="https://zenodo.org/record/7836300/files/figure.png" pageId="2" pageNumber="131">Fig. 3, 4</figureCitation>
|
||
,
|
||
<figureCitation box="[1191,1222,1771,1797]" captionStart="FIGURES 5–19" captionStartId="4.[151,264,1807,1831]" captionTargetBox="[249,1334,191,1779]" captionTargetId="figure-27@4.[249,1338,181,1784]" captionTargetPageId="4" captionText="FIGURES 5–19. Calydus spp. 5, 7, 9, 11, 13, 15–18—C. yaroslavi new species (5, 7, 9, 11, 17–19—holotype, male; 13— paratype, male; 15—paratype, female); 6, 8, 10, 12, 16—C. semenovi (Escherich, 1896) (6, 8, 10, 12—male, 16—female; Iran, West Azerbaijan Prov.); 14—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.). 5, 6—head; 7, 8—right mandible; 9, 10 pronotum; 11, 12—antennae; 13 (the arrow indicates medial elevation), 14 (the arrow indicates elevated ventrolateral margin of pronotum)—prosternum; 15, 16—mesoventrite; 17—tegmen, ventral view; 18—tegmen, lateral view; 19—aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7838276" httpUri="https://zenodo.org/record/7838276/files/figure.png" pageId="2" pageNumber="131">10</figureCitation>
|
||
); legs more robust, tarsomeres shorter and in lateral view rather widened (
|
||
<figureCitation box="[764,862,1807,1833]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[321,1266,192,1911]" captionTargetId="figure-17@3.[307,1281,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Calydus spp., general view. 1—C. yaroslavi new species (holotype, male); 2—C. yaroslavi new species (paratype, female); 3—C. semenovi (Escherich, 1896) (male, Iran, West Azerbaijan Prov.); 4—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.)." figureDoi="http://doi.org/10.5281/zenodo.7836300" httpUri="https://zenodo.org/record/7836300/files/figure.png" pageId="2" pageNumber="131">Fig. 3, 4</figureCitation>
|
||
); body pubescence short and recumbent, pronotum and elytra with poorly visible, very short recumbent pubescence (
|
||
<figureCitation box="[894,993,1843,1869]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[321,1266,192,1911]" captionTargetId="figure-17@3.[307,1281,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Calydus spp., general view. 1—C. yaroslavi new species (holotype, male); 2—C. yaroslavi new species (paratype, female); 3—C. semenovi (Escherich, 1896) (male, Iran, West Azerbaijan Prov.); 4—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.)." figureDoi="http://doi.org/10.5281/zenodo.7836300" httpUri="https://zenodo.org/record/7836300/files/figure.png" pageId="2" pageNumber="131">Fig. 3, 4</figureCitation>
|
||
,
|
||
<figureCitation box="[1006,1109,1843,1869]" captionStart="FIGURES 5–19" captionStartId="4.[151,264,1807,1831]" captionTargetBox="[249,1334,191,1779]" captionTargetId="figure-27@4.[249,1338,181,1784]" captionTargetPageId="4" captionText="FIGURES 5–19. Calydus spp. 5, 7, 9, 11, 13, 15–18—C. yaroslavi new species (5, 7, 9, 11, 17–19—holotype, male; 13— paratype, male; 15—paratype, female); 6, 8, 10, 12, 16—C. semenovi (Escherich, 1896) (6, 8, 10, 12—male, 16—female; Iran, West Azerbaijan Prov.); 14—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.). 5, 6—head; 7, 8—right mandible; 9, 10 pronotum; 11, 12—antennae; 13 (the arrow indicates medial elevation), 14 (the arrow indicates elevated ventrolateral margin of pronotum)—prosternum; 15, 16—mesoventrite; 17—tegmen, ventral view; 18—tegmen, lateral view; 19—aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7838276" httpUri="https://zenodo.org/record/7838276/files/figure.png" pageId="2" pageNumber="131">6, 10, 14</figureCitation>
|
||
). In the examined species
|
||
<taxonomicName baseAuthorityName="Escherich" baseAuthorityYear="1896" class="Insecta" family="Meloidae" genus="Calydus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="semenovi">
|
||
<emphasis italics="true" pageId="2" pageNumber="131">C. semenovi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Kaszab" authorityYear="1960" box="[310,471,1879,1905]" class="Insecta" family="Meloidae" genus="Calydus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="syriacus">
|
||
<emphasis box="[310,335,1879,1905]" italics="true" pageId="2" pageNumber="131">C.</emphasis>
|
||
cf.
|
||
<emphasis box="[378,471,1879,1904]" italics="true" pageId="2" pageNumber="131">syriacus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[487,583,1879,1905]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[321,1266,192,1911]" captionTargetId="figure-17@3.[307,1281,181,1919]" captionTargetPageId="3" captionText="FIGURES 1–4. Calydus spp., general view. 1—C. yaroslavi new species (holotype, male); 2—C. yaroslavi new species (paratype, female); 3—C. semenovi (Escherich, 1896) (male, Iran, West Azerbaijan Prov.); 4—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.)." figureDoi="http://doi.org/10.5281/zenodo.7836300" httpUri="https://zenodo.org/record/7836300/files/figure.png" pageId="2" pageNumber="131">Fig. 3, 4</figureCitation>
|
||
,
|
||
<figureCitation box="[595,805,1879,1905]" captionStart="FIGURES 5–19" captionStartId="4.[151,264,1807,1831]" captionTargetBox="[249,1334,191,1779]" captionTargetId="figure-27@4.[249,1338,181,1784]" captionTargetPageId="4" captionText="FIGURES 5–19. Calydus spp. 5, 7, 9, 11, 13, 15–18—C. yaroslavi new species (5, 7, 9, 11, 17–19—holotype, male; 13— paratype, male; 15—paratype, female); 6, 8, 10, 12, 16—C. semenovi (Escherich, 1896) (6, 8, 10, 12—male, 16—female; Iran, West Azerbaijan Prov.); 14—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.). 5, 6—head; 7, 8—right mandible; 9, 10 pronotum; 11, 12—antennae; 13 (the arrow indicates medial elevation), 14 (the arrow indicates elevated ventrolateral margin of pronotum)—prosternum; 15, 16—mesoventrite; 17—tegmen, ventral view; 18—tegmen, lateral view; 19—aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7838276" httpUri="https://zenodo.org/record/7838276/files/figure.png" pageId="2" pageNumber="131">6, 8, 10, 12, 14, 16</figureCitation>
|
||
) mandibles are narrower with external outline gradually and comparatively weakly curved inward (
|
||
<figureCitation box="[628,695,1915,1941]" captionStart="FIGURES 5–19" captionStartId="4.[151,264,1807,1831]" captionTargetBox="[249,1334,191,1779]" captionTargetId="figure-27@4.[249,1338,181,1784]" captionTargetPageId="4" captionText="FIGURES 5–19. Calydus spp. 5, 7, 9, 11, 13, 15–18—C. yaroslavi new species (5, 7, 9, 11, 17–19—holotype, male; 13— paratype, male; 15—paratype, female); 6, 8, 10, 12, 16—C. semenovi (Escherich, 1896) (6, 8, 10, 12—male, 16—female; Iran, West Azerbaijan Prov.); 14—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.). 5, 6—head; 7, 8—right mandible; 9, 10 pronotum; 11, 12—antennae; 13 (the arrow indicates medial elevation), 14 (the arrow indicates elevated ventrolateral margin of pronotum)—prosternum; 15, 16—mesoventrite; 17—tegmen, ventral view; 18—tegmen, lateral view; 19—aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7838276" httpUri="https://zenodo.org/record/7838276/files/figure.png" pageId="2" pageNumber="131">Fig. 8</figureCitation>
|
||
), submentum widely impressed, ventrolateral margin of pronotum not evenly merging with margin of prosternum and forming a distinct elevated suture extending beyond anterior outline of procoxal cavities, prosternum proximal to the middle with a wide obtuse elevation (
|
||
<figureCitation box="[1204,1287,1987,2013]" captionStart="FIGURES 5–19" captionStartId="4.[151,264,1807,1831]" captionTargetBox="[249,1334,191,1779]" captionTargetId="figure-27@4.[249,1338,181,1784]" captionTargetPageId="4" captionText="FIGURES 5–19. Calydus spp. 5, 7, 9, 11, 13, 15–18—C. yaroslavi new species (5, 7, 9, 11, 17–19—holotype, male; 13— paratype, male; 15—paratype, female); 6, 8, 10, 12, 16—C. semenovi (Escherich, 1896) (6, 8, 10, 12—male, 16—female; Iran, West Azerbaijan Prov.); 14—C. cf. syriacus Kaszab, 1960 (female, Iran, Lorestan Prov.). 5, 6—head; 7, 8—right mandible; 9, 10 pronotum; 11, 12—antennae; 13 (the arrow indicates medial elevation), 14 (the arrow indicates elevated ventrolateral margin of pronotum)—prosternum; 15, 16—mesoventrite; 17—tegmen, ventral view; 18—tegmen, lateral view; 19—aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7838276" httpUri="https://zenodo.org/record/7838276/files/figure.png" pageId="2" pageNumber="131">Fig. 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7836300" ID-Zenodo-Dep="7836300" httpUri="https://zenodo.org/record/7836300/files/figure.png" pageId="3" pageNumber="132" startId="3.[151,264,1943,1967]" targetBox="[321,1266,192,1911]" targetPageId="3">
|
||
<paragraph blockId="3.[151,1437,1943,2040]" pageId="3" pageNumber="132">
|
||
<emphasis bold="true" box="[151,319,1943,1967]" pageId="3" pageNumber="132">FIGURES 1–4.</emphasis>
|
||
<taxonomicName box="[330,467,1944,1968]" class="Insecta" family="Meloidae" genus="Calydus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="132" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis box="[330,413,1944,1968]" italics="true" pageId="3" pageNumber="132">Calydus</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
, general view. 1—
|
||
<taxonomicName authority="Shapovalov & Skrylnyk, 2023" authorityName="Shapovalov & Skrylnyk" authorityYear="2023" box="[670,797,1944,1968]" class="Insecta" family="Meloidae" genus="Calydus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="132" phylum="Arthropoda" rank="species" species="yaroslavi" status="new species">
|
||
<emphasis box="[670,797,1944,1968]" italics="true" pageId="3" pageNumber="132">C. yaroslavi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[808,934,1944,1968]" pageId="3" pageNumber="132" rank="species">new species</taxonomicNameLabel>
|
||
(holotype, male); 2—
|
||
<taxonomicName authority="Shapovalov & Skrylnyk, 2023" authorityName="Shapovalov & Skrylnyk" authorityYear="2023" box="[1172,1299,1944,1968]" class="Insecta" family="Meloidae" genus="Calydus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="132" phylum="Arthropoda" rank="species" species="yaroslavi" status="new species">
|
||
<emphasis box="[1172,1299,1944,1968]" italics="true" pageId="3" pageNumber="132">C. yaroslavi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1310,1436,1944,1968]" pageId="3" pageNumber="132" rank="species">new species</taxonomicNameLabel>
|
||
(paratype, female); 3—
|
||
<taxonomicName authority="(Escherich, 1896)" baseAuthorityName="Escherich" baseAuthorityYear="1896" box="[393,709,1979,2004]" class="Insecta" family="Meloidae" genus="Calydus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="132" phylum="Arthropoda" rank="species" species="semenovi">
|
||
<emphasis box="[393,518,1980,2003]" italics="true" pageId="3" pageNumber="132">C. semenovi</emphasis>
|
||
(Escherich, 1896)
|
||
</taxonomicName>
|
||
(male, Iran, West Azerbaijan Prov.); 4—
|
||
<taxonomicName authority="Kaszab, 1960" authorityName="Kaszab" authorityYear="1960" box="[1137,1437,1979,2004]" class="Insecta" family="Meloidae" genus="Calydus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="132" phylum="Arthropoda" rank="species" species="syriacus">
|
||
<emphasis box="[1137,1161,1980,2003]" italics="true" pageId="3" pageNumber="132">C.</emphasis>
|
||
cf.
|
||
<emphasis box="[1203,1287,1980,2003]" italics="true" pageId="3" pageNumber="132">syriacus</emphasis>
|
||
<bibRefCitation author="Kaszab, Z." box="[1296,1437,1979,2004]" pageId="3" pageNumber="132" pagination="125 - 134" refId="ref3623" refString="Kaszab, Z. (1960) Die Arten und systematische Stellung der Meloiden-Gattung Calydus Reitter (Coleoptera). Acta zoologica Academiae scientiarum Hungaricae, 6, 125 - 134." type="journal article" year="1960">Kaszab, 1960</bibRefCitation>
|
||
</taxonomicName>
|
||
(female, Iran, Lorestan Prov.).
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7838276" ID-Zenodo-Dep="7838276" httpUri="https://zenodo.org/record/7838276/files/figure.png" pageId="4" pageNumber="133" startId="4.[151,264,1807,1831]" targetBox="[249,1334,191,1779]" targetPageId="4">
|
||
<paragraph blockId="4.[151,1437,1807,2012]" pageId="4" pageNumber="133">
|
||
<emphasis bold="true" box="[151,330,1807,1831]" pageId="4" pageNumber="133">FIGURES 5–19.</emphasis>
|
||
<taxonomicName box="[338,494,1807,1832]" class="Insecta" family="Meloidae" genus="Calydus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="133" phylum="Arthropoda" rank="species" species="undefined-5">
|
||
<emphasis box="[338,421,1808,1832]" italics="true" pageId="4" pageNumber="133">Calydus</emphasis>
|
||
spp. 5
|
||
</taxonomicName>
|
||
, 7, 9, 11, 13, 15–18—
|
||
<taxonomicName authority="Shapovalov & Skrylnyk, 2023" authorityName="Shapovalov & Skrylnyk" authorityYear="2023" box="[731,856,1808,1832]" class="Insecta" family="Meloidae" genus="Calydus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="133" phylum="Arthropoda" rank="species" species="yaroslavi" status="new species">
|
||
<emphasis box="[731,856,1808,1832]" italics="true" pageId="4" pageNumber="133">C. yaroslavi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[865,989,1808,1832]" pageId="4" pageNumber="133" rank="species">new species</taxonomicNameLabel>
|
||
(5, 7, 9, 11, 17–19—
|
||
<typeStatus box="[1215,1308,1808,1832]" pageId="4" pageNumber="133">holotype</typeStatus>
|
||
, male; 13—
|
||
<typeStatus box="[151,241,1844,1868]" pageId="4" pageNumber="133">paratype</typeStatus>
|
||
, male; 15—
|
||
<typeStatus box="[363,454,1843,1867]" pageId="4" pageNumber="133">paratype</typeStatus>
|
||
, female); 6, 8, 10, 12, 16—
|
||
<taxonomicName authority="(Escherich, 1896)" baseAuthorityName="Escherich" baseAuthorityYear="1896" box="[731,1041,1844,1868]" class="Insecta" family="Meloidae" genus="Calydus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="133" phylum="Arthropoda" rank="species" species="semenovi">
|
||
<emphasis box="[731,855,1844,1867]" italics="true" pageId="4" pageNumber="133">C. semenovi</emphasis>
|
||
(Escherich, 1896)
|
||
</taxonomicName>
|
||
(6, 8, 10, 12—male, 16—female;
|
||
<collectingCountry box="[1389,1432,1844,1867]" name="Iran" pageId="4" pageNumber="133">Iran</collectingCountry>
|
||
,
|
||
<collectingRegion box="[151,382,1880,1904]" country="Iran" name="Azarbayjan-e Gharbi" pageId="4" pageNumber="133">West Azerbaijan Prov.</collectingRegion>
|
||
); 14—
|
||
<taxonomicName authority="Kaszab, 1960" authorityName="Kaszab" authorityYear="1960" box="[451,743,1880,1904]" class="Insecta" family="Meloidae" genus="Calydus" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="133" phylum="Arthropoda" rank="species" species="syriacus">
|
||
<emphasis box="[451,596,1880,1904]" italics="true" pageId="4" pageNumber="133">C. cf. syriacus</emphasis>
|
||
<bibRefCitation author="Kaszab, Z." box="[604,743,1880,1904]" pageId="4" pageNumber="133" pagination="125 - 134" refId="ref3623" refString="Kaszab, Z. (1960) Die Arten und systematische Stellung der Meloiden-Gattung Calydus Reitter (Coleoptera). Acta zoologica Academiae scientiarum Hungaricae, 6, 125 - 134." type="journal article" year="1960">Kaszab, 1960</bibRefCitation>
|
||
</taxonomicName>
|
||
(female,
|
||
<collectingCountry box="[840,883,1880,1903]" name="Iran" pageId="4" pageNumber="133">Iran</collectingCountry>
|
||
,
|
||
<collectingRegion box="[894,1047,1880,1904]" country="Iran" name="Lorestan" pageId="4" pageNumber="133">Lorestan Prov.</collectingRegion>
|
||
). 5, 6—head; 7, 8—right mandible; 9, 10 pronotum; 11, 12—antennae; 13 (the arrow indicates medial elevation), 14 (the arrow indicates elevated ventrolateral margin of pronotum)—prosternum; 15, 16—mesoventrite; 17—tegmen, ventral view; 18—tegmen, lateral view; 19—aedeagus, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7836302" ID-Zenodo-Dep="7836302" box="[151,1248,1998,2022]" httpUri="https://zenodo.org/record/7836302/files/figure.png" pageId="5" pageNumber="134" startId="5.[151,250,1998,2022]" targetBox="[173,1414,181,1963]" targetPageId="5">
|
||
<paragraph blockId="5.[151,1248,1998,2022]" box="[151,1248,1998,2022]" pageId="5" pageNumber="134">
|
||
<emphasis bold="true" box="[151,322,1998,2022]" pageId="5" pageNumber="134">FIGURE 20-21.</emphasis>
|
||
The type locality of the
|
||
<taxonomicName authority="Shapovalov & Skrylnyk, 2023" authorityName="Shapovalov & Skrylnyk" authorityYear="2023" box="[570,752,1998,2022]" class="Insecta" family="Meloidae" genus="Calydus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="134" phylum="Arthropoda" rank="species" species="yaroslavi" status="new species">
|
||
<emphasis box="[570,752,1998,2022]" italics="true" pageId="5" pageNumber="134">Calydus yaroslavi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[759,884,1998,2022]" pageId="5" pageNumber="134" rank="species">new species</taxonomicNameLabel>
|
||
. 20—view of locality, 21—biotope.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7836304" ID-Zenodo-Dep="7836304" httpUri="https://zenodo.org/record/7836304/files/figure.png" pageId="6" pageNumber="135" startId="6.[151,264,1913,1937]" targetBox="[174,1413,182,1875]" targetPageId="6">
|
||
<paragraph blockId="6.[151,1437,1913,2011]" pageId="6" pageNumber="135">
|
||
<emphasis bold="true" box="[151,341,1913,1937]" pageId="6" pageNumber="135">FIGURES 22–23.</emphasis>
|
||
Biotope and distribution of
|
||
<taxonomicName box="[632,765,1913,1937]" class="Insecta" family="Meloidae" genus="Calydus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis box="[632,715,1913,1937]" italics="true" pageId="6" pageNumber="135">Calydus</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
22—map of distribution
|
||
<taxonomicName authority="Shapovalov & Skrylnyk, 2023" authorityName="Shapovalov & Skrylnyk" authorityYear="2023" box="[1025,1209,1913,1937]" class="Insecta" family="Meloidae" genus="Calydus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="yaroslavi" status="new species">
|
||
<emphasis box="[1025,1209,1913,1937]" italics="true" pageId="6" pageNumber="135">Calydus yaroslavi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1216,1339,1913,1937]" pageId="6" pageNumber="135" rank="species">new species</taxonomicNameLabel>
|
||
(1—type locality, 3 km W Sabzel; 2–10 km S Bamyan); 23—biotope of the
|
||
<taxonomicName authority="(Escherich, 1896)" baseAuthorityName="Escherich" baseAuthorityYear="1896" box="[815,1182,1949,1973]" class="Insecta" family="Meloidae" genus="Calydus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="semenovi">
|
||
<emphasis box="[815,997,1949,1973]" italics="true" pageId="6" pageNumber="135">Calydus semenovi</emphasis>
|
||
(Escherich, 1896)
|
||
</taxonomicName>
|
||
in Iran, West Azerbaijan, Takht-e Soleyman, 24.V.2017, 36°37′15.46″N 47°18′36.94″E (photo by A. Zubov).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="7" pageNumber="136" type="etymology">
|
||
<paragraph blockId="7.[151,1436,151,393]" pageId="7" pageNumber="136">
|
||
<emphasis bold="true" box="[199,334,151,177]" pageId="7" pageNumber="136">Etymology.</emphasis>
|
||
The new species is named after Yaroslav Skrylnyk, the younger son of the second author of this paper.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="7" pageNumber="136" type="biology_ecology">
|
||
<paragraph blockId="7.[151,1436,151,393]" pageId="7" pageNumber="136">
|
||
<emphasis bold="true" box="[199,300,223,249]" pageId="7" pageNumber="136">Ecology.</emphasis>
|
||
Specimens were collected from June, 12 to July, 8. Both known localities are situated at an altitude of about
|
||
<quantity box="[254,340,259,285]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="7" pageNumber="136" unit="m" value="2900.0">2900 m</quantity>
|
||
a.s.l. In the
|
||
<typeStatus box="[483,531,259,285]" pageId="7" pageNumber="136">type</typeStatus>
|
||
locality (
|
||
<quantity box="[642,700,259,285]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="136" unit="km" value="3.0">3 km</quantity>
|
||
W Sabzel) the new species was found in a localised mesophytic meadow biotope with shrubs in the floodplain of the Bande-Amir River surrounded by very dry hilly landscapes (
|
||
<figureCitation box="[159,295,331,357]" captionStart="FIGURE 20-21" captionStartId="5.[151,250,1998,2022]" captionTargetBox="[173,1414,181,1963]" captionTargetId="figure-17@5.[173,1414,181,1974]" captionTargetPageId="5" captionText="FIGURE 20-21. The type locality of the Calydus yaroslavi new species. 20—view of locality, 21—biotope." figureDoi="http://doi.org/10.5281/zenodo.7836302" httpUri="https://zenodo.org/record/7836302/files/figure.png" pageId="7" pageNumber="136">Figs. 20, 21</figureCitation>
|
||
). It is interesting to note that in the same biotope was found also the recently described
|
||
<taxonomicName authority="A. Shapovalov, 2016" authorityName="A. Shapovalov" authorityYear="2016" class="Insecta" family="Meloidae" genus="Lytta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="skrylniki" subGenus="Eolytta">
|
||
<emphasis box="[1273,1330,331,356]" italics="true" pageId="7" pageNumber="136">Lytta</emphasis>
|
||
(
|
||
<emphasis box="[1346,1427,331,357]" italics="true" pageId="7" pageNumber="136">Eolytta</emphasis>
|
||
)
|
||
<emphasis box="[151,247,367,393]" italics="true" pageId="7" pageNumber="136">skrylniki</emphasis>
|
||
A.
|
||
<bibRefCitation author="Shapovalov, A. M." box="[287,488,367,393]" pageId="7" pageNumber="136" pagination="99 - 108" refId="ref3732" refString="Shapovalov, A. M. (2016) Remarks on taxonomy of the genus Lytta Fabricius, 1775 (Coleoptera: Meloidae) with description of a new subgenus and two new species from Asia. Caucasian Entomological Bulletin, 12 (1), 99 - 108. [in Russian, English abstract] https: // doi. org / 10.23885 / 1814 - 3326 - 2016 - 12 - 1 - 99 - 108" type="journal article" year="2016">Shapovalov, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="Waterhouse, 1889" authorityName="Waterhouse" authorityYear="1889" box="[542,942,367,393]" class="Insecta" family="Meloidae" genus="Lytta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="laeta" subGenus="Eolytta">
|
||
<emphasis box="[542,558,367,392]" italics="true" pageId="7" pageNumber="136">L</emphasis>
|
||
. (
|
||
<emphasis box="[581,662,367,393]" italics="true" pageId="7" pageNumber="136">Eolytta</emphasis>
|
||
)
|
||
<emphasis box="[678,734,367,393]" italics="true" pageId="7" pageNumber="136">laeta</emphasis>
|
||
Waterhouse, 1889
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Shapovalov, A. M." box="[957,1164,367,393]" pageId="7" pageNumber="136" pagination="99 - 108" refId="ref3732" refString="Shapovalov, A. M. (2016) Remarks on taxonomy of the genus Lytta Fabricius, 1775 (Coleoptera: Meloidae) with description of a new subgenus and two new species from Asia. Caucasian Entomological Bulletin, 12 (1), 99 - 108. [in Russian, English abstract] https: // doi. org / 10.23885 / 1814 - 3326 - 2016 - 12 - 1 - 99 - 108" type="journal article" year="2016">Shapovalov, 2016</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |