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<document id="C34BAF8074DDBD35136D45F9C3AD8344" ID-DOI="10.5281/zenodo.3745504" ID-GBIF-Dataset="6ac2f34f-6521-4e02-b7ba-9151c40fdf61" ID-ISSN="0253-116X" ID-Zenodo-Dep="3745504" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1586373739657" checkinUser="valdenar" docAuthor="Skalický, Stanislav" docDate="2019" docId="515087FAFF91FFA58BAE077514542D76" docLanguage="en" docName="Linzer.biol.Beitr.51.2.1421-1428.pdf.imf" docOrigin="Linzer biologische Beiträge 51 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Augyles davranogloui Skalický 2019, nov.sp." docType="treatment" docVersion="7" lastPageNumber="1424" masterDocId="AD69FF82FF93FFA68B39045E151A2E3A" masterDocTitle="New species of Heteroceridae from Kenya and Zambia (Insecta, Coleoptera)" masterLastPageNumber="1428" masterPageNumber="1421" pageNumber="1423" updateTime="1698736669822" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="E85884A45DE32B38F98691E5865DE381">
<mods:title id="77BD5E6279E4204A23723E099D668C4A">New species of Heteroceridae from Kenya and Zambia (Insecta, Coleoptera)</mods:title>
</mods:titleInfo>
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<mods:namePart id="2AAA989D81B310118225B269A349A0A1">Skalický, Stanislav</mods:namePart>
</mods:name>
<mods:typeOfResource id="4986309665CE35872465C23E5BD28A5B">text</mods:typeOfResource>
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<mods:title id="FDF20662E326F781E0D3AB87CBE979F6">Linzer biologische Beiträge</mods:title>
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<mods:date id="480531C939CD8AD23C47D63CD3D18900">2019</mods:date>
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<mods:number id="F54FE9E8D681505F79344C2F4DE4CA3C">2019-12-20</mods:number>
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<mods:classification id="508C2139754843527998397BCD76062C">journal article</mods:classification>
<mods:identifier id="78DE50D13EDF2B1FB1EB25AFA5FF253D" type="DOI">10.5281/zenodo.3745504</mods:identifier>
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<treatment id="515087FAFF91FFA58BAE077514542D76" ID-DOI="http://doi.org/10.5281/zenodo.3809421" ID-GBIF-Taxon="163540364" ID-Zenodo-Dep="3809421" LSID="urn:lsid:plazi:treatment:515087FAFF91FFA58BAE077514542D76" httpUri="http://treatment.plazi.org/id/515087FAFF91FFA58BAE077514542D76" lastPageId="3" lastPageNumber="1424" pageId="2" pageNumber="1423">
<subSubSection id="91E36567FF91FFA48BAE077514D52D7B" box="[151,463,811,833]" pageId="2" pageNumber="1423" type="nomenclature">
<paragraph id="D94636ECFF91FFA48BAE077514D52D7B" blockId="2.[151,463,811,833]" box="[151,463,811,833]" pageId="2" pageNumber="1423">
<heading id="820E8180FF91FFA48BAE077514D52D7B" box="[151,463,811,833]" fontSize="8" level="2" pageId="2" pageNumber="1423" reason="2">
<emphasis id="EB8DEAFEFF91FFA48BAE077514D52D7B" bold="true" box="[151,463,811,833]" pageId="2" pageNumber="1423">
<taxonomicName id="1EF94D6FFF91FFA48BAE077514662D7B" authority="Skalický, 2019" authorityName="Skalický" authorityYear="2019" box="[151,380,811,833]" class="Insecta" family="Heteroceridae" genus="Augyles" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1423" phylum="Arthropoda" rank="species" species="davranogloui" status="sp. nov.">
<emphasis id="EB8DEAFEFF91FFA48BAE077514662D7B" bold="true" box="[151,380,811,833]" italics="true" pageId="2" pageNumber="1423">Augyles davranogloui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0BE5785FF91FFA48ABB077214D52D7B" box="[386,463,812,833]" pageId="2" pageNumber="1423" rank="species">nov.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="91E36567FF91FFA48BAE070C14162DE5" pageId="2" pageNumber="1423" type="materials_examined">
<paragraph id="D94636ECFF91FFA48BAE070C14162DE5" blockId="2.[151,1059,848,991]" pageId="2" pageNumber="1423">
<materialsCitation id="69913CB1FF91FFA48BAE070C168C2D45" ID-GBIF-Occurrence="2597468615" collectingDate="2011-11-27" country="Kenya" elevation="600" location="Snizek" municipality="Mwingi Sosoma" pageId="2" pageNumber="1423" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
T y p e m a t e r i a l:
<typeStatus id="0642884EFF91FFA48AB0070C14C02D52" box="[393,474,850,872]" pageId="2" pageNumber="1423" type="holotype">Holotype</typeStatus>
<specimenCount id="CFFFFD65FF91FFA48AD9070E14E82D5D" box="[480,498,848,871]" pageId="2" pageNumber="1423" type="male">♂</specimenCount>
: &quot;
<collectingCountry id="A1EE767CFF91FFA4893A070C17242D5D" box="[515,574,850,871]" name="Kenya" pageId="2" pageNumber="1423">Kenya</collectingCountry>
E,
<quantity id="1E019B09FF91FFA48965070C17942D52" box="[604,654,850,872]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="1423" unit="m" value="600.0">
<elevation id="52D4D1DFFF91FFA48965070C17942D52" box="[604,654,850,872]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="1423" unit="m" value="600.0">600m</elevation>
</quantity>
E of
<collectingMunicipality id="3922AC96FF91FFA48986070C164A2D5D" box="[703,848,850,872]" pageId="2" pageNumber="1423">Mwingi Sosoma</collectingMunicipality>
,
<date id="AD47102CFF91FFA48863070C16A62D5D" box="[858,956,850,871]" pageId="2" pageNumber="1423" value="2011-11-27">
<collectingDate id="BD03E9C4FF91FFA48863070C16A62D5D" box="[858,956,850,871]" pageId="2" pageNumber="1423" value="2011-11-27">27.11.2011</collectingDate>
</date>
<location id="DC266037FF91FFA488F8070C16E62D52" LSID="urn:lsid:plazi:treatment:515087FAFF91FFA58BAE077514542D76:DC266037FF91FFA488F8070C16E62D52" box="[961,1020,850,872]" country="Kenya" municipality="Mwingi Sosoma" name="Snizek" pageId="2" pageNumber="1423">Snížek</location>
lgt&quot; // &quot;
<typeStatus id="0642884EFF91FFA48BE4073414492D45" box="[221,339,874,895]" pageId="2" pageNumber="1423" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="1EF94D6FFF91FFA48A61073417E32D45" authority="Skal. Det. Skalicky 2019" authorityName="Skal. Det. Skalicky" authorityYear="2019" box="[344,761,874,896]" class="Insecta" family="Heteroceridae" genus="Augyles" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1423" phylum="Arthropoda" rank="species" species="davranogloui">
<location id="DC266037FF91FFA48A61073417DC2DBA" LSID="urn:lsid:plazi:treatment:515087FAFF91FFA58BAE077514542D76:DC266037FF91FFA48A61073417DC2DBA" box="[344,710,874,896]" country="Kenya" municipality="Mwingi Sosoma" name="Augyles davranogloui Skal. Det. Skalicky" pageId="2" pageNumber="1423">Augyles davranogloui Skal. Det. Skalický</location>
2019
</taxonomicName>
&quot; [red label] CSU.
</materialsCitation>
<materialsCitation id="69913CB1FF91FFA488A3073417532D95" ID-GBIF-Occurrence="2597468607" location="Augyles davranogloui Skal. Det. Skalicky" municipality="Allotype" pageId="2" pageNumber="1423" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<collectingMunicipality id="3922AC96FF91FFA488A3073416FC2DBA" box="[922,998,874,896]" pageId="2" pageNumber="1423">Allotype</collectingMunicipality>
<specimenCount id="CFFFFD65FF91FFA488D5073416E42D45" box="[1004,1022,874,895]" pageId="2" pageNumber="1423" type="female">♀</specimenCount>
: the same data as
<typeStatus id="0642884EFF91FFA48A7C07DC148B2DA2" box="[325,401,898,920]" pageId="2" pageNumber="1423" type="holotype">holotype</typeStatus>
only, second determinate label is: &quot;
<typeStatus id="0642884EFF91FFA489E607DC16492DAD" box="[735,851,898,919]" pageId="2" pageNumber="1423" type="allotype">ALLOTYPE</typeStatus>
<taxonomicName id="1EF94D6FFF91FFA4886407DC14BF2D95" authority="Skal. Det. Skalicky 2019" authorityName="Skal. Det. Skalicky" authorityYear="2019" class="Insecta" family="Heteroceridae" genus="Augyles" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1423" phylum="Arthropoda" rank="species" species="davranogloui">
<location id="DC266037FF91FFA4886407DC146A2D8A" LSID="urn:lsid:plazi:treatment:515087FAFF91FFA58BAE077514542D76:DC266037FF91FFA4886407DC146A2D8A" municipality="Allotype" name="Augyles davranogloui Skal. Det. Skalicky" pageId="2" pageNumber="1423">Augyles davranogloui Skal. Det. Skalický</location>
2019
</taxonomicName>
&quot; [red label] CSU,
</materialsCitation>
<materialsCitation id="69913CB1FF91FFA4896907C414162DE5" ID-GBIF-Occurrence="2597468617" collectionCode="NHML" location="Augyles davranogloui Skal. Det. Skalicky" pageId="2" pageNumber="1423" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="0642884EFF91FFA4896907C417862D95" box="[592,668,922,943]" pageId="2" pageNumber="1423" type="paratype">Paratype</typeStatus>
<specimenCount id="CFFFFD65FF91FFA4899D07C417AC2D95" box="[676,694,922,943]" pageId="2" pageNumber="1423" type="female">♀</specimenCount>
: the same data as
<typeStatus id="0642884EFF91FFA4886707C416B02D8A" box="[862,938,922,944]" pageId="2" pageNumber="1423" type="holotype">holotype</typeStatus>
only, second determined label is: &quot;
<typeStatus id="0642884EFF91FFA48ABF07EC14E02DFD" box="[390,506,946,967]" pageId="2" pageNumber="1423" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="1EF94D6FFF91FFA4893A07EC16AF2DFD" authority="Skal. Det. Skalicky 2019" authorityName="Skal. Det. Skalicky" authorityYear="2019" box="[515,949,946,968]" class="Insecta" family="Heteroceridae" genus="Augyles" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1423" phylum="Arthropoda" rank="species" species="davranogloui">
<location id="DC266037FF91FFA4893A07EC16652DF2" LSID="urn:lsid:plazi:treatment:515087FAFF91FFA58BAE077514542D76:DC266037FF91FFA4893A07EC16652DF2" box="[515,895,946,968]" name="Augyles davranogloui Skal. Det. Skalicky" pageId="2" pageNumber="1423">Augyles davranogloui Skal. Det. Skalický</location>
2019
</taxonomicName>
&quot; [red label]
<collectionCode id="BFE8AE29FF91FFA48BFD079414122DE5" box="[196,264,970,991]" collectionName="Libya, Tripoli, Natural History Museum" pageId="2" pageNumber="1423">NHML</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="91E36567FF91FFA58BAE07B114542D76" lastPageId="3" lastPageNumber="1424" pageId="2" pageNumber="1423" type="description">
<paragraph id="D94636ECFF91FFA58BAE07B1140B2FA2" blockId="2.[151,1058,1004,1585]" lastBlockId="3.[151,1059,211,409]" lastPageId="3" lastPageNumber="1424" pageId="2" pageNumber="1423">
D e s c r i p t i o n:
<typeStatus id="0642884EFF91FFA48A4D07B014C82A3E" box="[372,466,1006,1028]" pageId="2" pageNumber="1423" type="holotype">Holotype</typeStatus>
♂: Total length
<quantity id="1E019B09FF91FFA4894707B017C22A3E" box="[638,728,1006,1028]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="2" pageNumber="1423" unit="mm" value="2.9">2.90 mm</quantity>
(to apex of labrum); elytra
<quantity id="1E019B09FF91FFA488CF07B015A42A1B" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="1423" unit="mm" value="1.8">1.80 mm</quantity>
long,
<quantity id="1E019B09FF91FFA48A3A005514452A1B" box="[259,351,1035,1057]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="1423" unit="mm" value="0.7">0.70 mm</quantity>
wide across shoulders. Ground colour light chestnut brown without any distinct pattern, labrum darker, eyes black. Labrum (Fig. 9) short, visible part wider than long (ratio 1.35: 1), lateral margins almost parallel; surface finely granulated, setae irregular semi-erect, yellowish and quite long. Mandibles (Fig. 9) short and robust, dorsal subapical tooth short, rounded, dental lobe with two rouned spines. Prostheca with teeth on dental lobe, without prosthecal notch. Antennae 11-segmented, with a 7- segmented club, pedicel oval, scape and pedicel with sparse long erect setae. Head very finely granulated, setae as in labrum. Clypeus (Fig. 9) without pair of anterior horns, anterior margin deeply emarginate, setae as in head. Pronotum as wide as the base of elytra, oval, pronotal base completely rimmed; surface densely regularly granular, punctures approximately as large as 1.5 times eye facets; setae sparse, short, erect, light brown. Scutellum pointed, triangular, lateral margins straight. Elytra with feebly-marked three longitudinal furrows in each elytron (they are clearly visible in wet specimens), without scutellar depressions, humeral depressions shallow and short. Surface of elytra microgranular, with intermixed coarse punctures approximately 2.5 as large as eye facets. Coarse punctures irregular. Setae on elytra light brown, short, erect without longer setae. Epipleural ridges absent. Metaventrite with post-mesocoxal ridge. Mesoventrite with three small rounded spines placed far apart in front of each mesocoxa. Ventral surface sparsely setose. Protibia with 10 stout spines along its lateral margin and two short spines at its inner apical angle, mesotibia with 8 weak and metatibia with 7 thin spines. Tibiae with dense, long setae. Abdomen with complete post-metacoxal ridge. Stridulatory arch marked, without striae. Spiculum gastrale
<quantity id="1E019B09FF90FFA5883A055016452F1E" box="[771,863,270,292]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="1424" unit="mm" value="0.6">0.60 mm</quantity>
long, V-shaped as in Fig. 10, arms narrow, firmly connected apically. Aedeagus
<quantity id="1E019B09FF90FFA5887B057516BB2F7B" box="[834,929,299,321]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="1424" unit="mm" value="0.5">0.50 mm</quantity>
long, wellsclerotized; parameres short and completely fused together, firmly connected with phallobasis, penis without internal sac and processus accessories; shape of aedeagus as in Figs. 11-14.
</paragraph>
<paragraph id="D94636ECFF90FFA58BAE05F714752FE6" blockId="3.[151,1058,425,476]" pageId="3" pageNumber="1424">
<typeStatus id="0642884EFF90FFA58BAE05F715EA2F85" box="[151,240,425,447]" pageId="3" pageNumber="1424" type="allotype">Allotype</typeStatus>
<specimenCount id="CFFFFD65FF90FFA58BCE05F714162F84" box="[247,268,425,446]" pageId="3" pageNumber="1424" type="female">♀</specimenCount>
: The same diameter as in
<typeStatus id="0642884EFF90FFA5892E05F717692F85" box="[535,627,425,447]" pageId="3" pageNumber="1424" type="holotype">holotype</typeStatus>
. Elytra without longitudinal ridges. Externally similar to male.
</paragraph>
<paragraph id="D94636ECFF90FFA58BAE05B216F32CD6" blockId="3.[151,1059,491,748]" pageId="3" pageNumber="1424">
D i f f e r e n t i a l d i a g n o s i s: The shape of the aedeagus, the presence of a complete post-metacoxal ridge, antenomere number and the absence of clypeal horns in the male of
<taxonomicName id="1EF94D6FFF90FFA58A23067814D82C01" authority="Skalický, 2019" authorityName="Skalický" authorityYear="2019" box="[282,450,550,571]" class="Insecta" family="Heteroceridae" genus="Augyles" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1424" phylum="Arthropoda" rank="species" species="davranogloui" status="sp. nov.">
<emphasis id="EB8DEAFEFF90FFA58A23067814D82C01" box="[282,450,550,571]" italics="true" pageId="3" pageNumber="1424">A. davranogloui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F0BE5785FF90FFA58AF50679170C2C06" box="[460,534,551,572]" pageId="3" pageNumber="1424" rank="species">nov.sp.</taxonomicNameLabel>
belongs to the
<emphasis id="EB8DEAFEFF90FFA589860679161E2C06" box="[703,772,551,572]" italics="true" pageId="3" pageNumber="1424">parvus</emphasis>
group sensu
<bibRefCitation id="BD684B1DFF90FFA588AC067815C42C63" author="CHARPENTIER R" pageId="3" pageNumber="1424" pagination="214 - 343" refId="ref4565" refString="CHARPENTIER R. (1965): A monograph of the family Heteroceridae (Coleoptera) of the Ethiopian Region. - South Afr. Anim. Life 11: 214 - 343." type="journal article" year="1965">CHARPENTIER (1965)</bibRefCitation>
. In the Ethiopian region, only two widely distributed representatives of this group are known:
<emphasis id="EB8DEAFEFF90FFA58A36063F14042C4C" box="[271,286,609,630]" italics="true" pageId="3" pageNumber="1424">A</emphasis>
.
<emphasis id="EB8DEAFEFF90FFA58A15063F146B2C4C" box="[300,369,609,630]" italics="true" pageId="3" pageNumber="1424">parvus</emphasis>
(GROUVELLE, 1909) and
<emphasis id="EB8DEAFEFF90FFA5894F063F179F2C4C" box="[630,645,609,630]" italics="true" pageId="3" pageNumber="1424">A</emphasis>
.
<emphasis id="EB8DEAFEFF90FFA589AA063F16092C4C" box="[659,787,609,630]" italics="true" pageId="3" pageNumber="1424">breviusculus</emphasis>
(CHARPENTIER, 1962) and the new species is probably related to
<emphasis id="EB8DEAFEFF90FFA5891A062017282CA9" box="[547,562,638,659]" italics="true" pageId="3" pageNumber="1424">A</emphasis>
.
<emphasis id="EB8DEAFEFF90FFA58979062017DA2CA9" box="[576,704,638,659]" italics="true" pageId="3" pageNumber="1424">breviusculus</emphasis>
. The latter species differs from
<taxonomicName id="1EF94D6FFF90FFA58F35062014052C8A" authorityName="Skalický" authorityYear="2019" class="Insecta" family="Heteroceridae" genus="Augyles" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1424" phylum="Arthropoda" rank="species" species="davranogloui">
<emphasis id="EB8DEAFEFF90FFA58F35062014052C8A" italics="true" pageId="3" pageNumber="1424">A. davranogloui</emphasis>
</taxonomicName>
by its ground colour and elytral pattern (reddish brown to black, dark specimens with an exceedingly diffuse, pale apical spot) and in the morphology of male genitalia (see Figs. 80 and
<quantity id="1E019B09FF90FFA58A9E068814FD2CD6" box="[423,487,726,748]" metricMagnitude="0" metricUnit="m" metricValue="3.175" pageId="3" pageNumber="1424" unit="in" value="125.0">125 in</quantity>
<bibRefCitation id="BD684B1DFF90FFA58AD4068817DE2CD6" author="CHARPENTIER R" box="[493,708,726,748]" pageId="3" pageNumber="1424" pagination="214 - 343" refId="ref4565" refString="CHARPENTIER R. (1965): A monograph of the family Heteroceridae (Coleoptera) of the Ethiopian Region. - South Afr. Anim. Life 11: 214 - 343." type="journal article" year="1965">CHARPENTIER (1965)</bibRefCitation>
and Figs
<quantity id="1E019B09FF90FFA5881E0688166A2CD6" box="[807,880,726,748]" metricMagnitude="-1" metricUnit="m" metricValue="2.921" metricValueMax="3.556" metricValueMin="2.286" pageId="3" pageNumber="1424" unit="in" value="11.5" valueMax="14.0" valueMin="9.0">9-14 in</quantity>
this paper).
</paragraph>
<paragraph id="D94636ECFF90FFA58BAE06A214542D76" blockId="3.[151,1058,763,844]" pageId="3" pageNumber="1424">E t y m o l o g y: The new species is named in honour of Leonidas-Romanos Davranoglou, an excellent Greek entomologist who specializes in hemipteran morphology and taxonomy.</paragraph>
</subSubSection>
</treatment>
</document>
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