treatments-xml/data/F9/3F/F7/F93FF74CE97CFFD90ACE09A8DA71FE58.xml
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<document id="0D689231BF8BF91675C1B3C5C995B036" ID-CLB-Dataset="148093" ID-DOI="10.11646/zootaxa.5188.3.5" ID-GBIF-Dataset="8b16cbb8-a846-4ee3-8692-2c677f338147" ID-ISSN="1175-5326" ID-Zenodo-Dep="7091693" ID-ZooBank="5F0BCAF8-8397-418C-8882-8865B2086530" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1663565831141" checkinUser="plazi" docAuthor="Lyubarsky, Georgy Yu. &amp; Bukejs, Andris" docDate="2022" docId="F93FF74CE97CFFD90ACE09A8DA71FE58" docLanguage="en" docName="zootaxa.5188.3.5.pdf" docOrigin="Zootaxa 5188 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Atomaria (Anchicera) perkovskyi Lyubarsky &amp; Bukejs 2022, sp. nov." docType="treatment" docVersion="6" lastPageNumber="288" masterDocId="05068F34E97FFFDC0A590A60D945FFAD" masterDocTitle="TwonewfossilspeciesofthegenusAtomariaStephens (Coleoptera: Cryptophagidae) from Eocene European amber with a key to species described from fossil resins" masterLastPageNumber="289" masterPageNumber="283" pageNumber="286" updateTime="1699382941167" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9355F860E232A659C45B69B756DF2145">TwonewfossilspeciesofthegenusAtomariaStephens (Coleoptera: Cryptophagidae) from Eocene European amber with a key to species described from fossil resins</mods:title>
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<mods:namePart id="8B3140293AFD1C1F31EC3E5D82FA364E">Lyubarsky, Georgy Yu.</mods:namePart>
<mods:affiliation id="839518DB71CC8AD657885C8864023B40">Zoological Museum, Moscow State University, Bolshaya Nikitskaya Str. 2, Moscow 103009, Russia.</mods:affiliation>
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<mods:namePart id="83C2632B320F1B44239BF99693B52E52">Bukejs, Andris</mods:namePart>
<mods:nameIdentifier id="C4ED4C9C5177E0222696FF46AC2F4B14" type="ORCID">0000-0001-7165-3023</mods:nameIdentifier>
<mods:affiliation id="28B8BCBB3D033CCE0E4CBB308BA31FDD">Institute of Life Sciences and Technologies, Daugavpils University, Vienîbas 13, Daugavpils LV- 5401, Latvia. carabidae @ inbox. lv; https: // orcid. org / 0000 - 0001 - 7165 - 3023</mods:affiliation>
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<mods:title id="990150C8AB6353B4D4A8A0FFB68EB8F2">Zootaxa</mods:title>
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<treatment id="F93FF74CE97CFFD90ACE09A8DA71FE58" ID-DOI="http://doi.org/10.5281/zenodo.7091691" ID-GBIF-Taxon="201101895" ID-Zenodo-Dep="7091691" LSID="urn:lsid:plazi:treatment:F93FF74CE97CFFD90ACE09A8DA71FE58" httpUri="http://treatment.plazi.org/id/F93FF74CE97CFFD90ACE09A8DA71FE58" lastPageId="5" lastPageNumber="288" pageId="3" pageNumber="286">
<subSubSection id="398C15D1E97CFFDF0ACE09A8DBCAFC4F" box="[151,655,967,994]" pageId="3" pageNumber="286" type="nomenclature">
<paragraph id="7129465AE97CFFDF0ACE09A8DBCAFC4F" blockId="3.[151,655,967,1030]" box="[151,655,967,994]" pageId="3" pageNumber="286">
<heading id="2A61F136E97CFFDF0ACE09A8DBCAFC4F" bold="true" box="[151,655,967,994]" fontSize="11" level="1" pageId="3" pageNumber="286" reason="1">
<emphasis id="43E29A48E97CFFDF0ACE09A8DBCAFC4F" bold="true" box="[151,655,967,994]" pageId="3" pageNumber="286">
<taxonomicName id="B6963DD9E97CFFDF0ACE09A8DB6FFC4F" authority="Lyubarsky &amp; Bukejs, 2022" authorityName="Lyubarsky &amp; Bukejs" authorityYear="2022" box="[151,554,967,994]" class="Insecta" family="Cryptophagidae" genus="Atomaria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="286" phylum="Arthropoda" rank="species" species="perkovskyi" status="sp. nov." subGenus="Anchicera">
<emphasis id="43E29A48E97CFFDF0ACE09A8DB6FFC4F" bold="true" box="[151,554,967,994]" italics="true" pageId="3" pageNumber="286">Atomaria (Anchicera) perkovskyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58D12733E97CFFDF086809A8DBCAFC4F" box="[561,655,968,994]" pageId="3" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="398C15D1E97CFFDF0ACE098CD84EFBAB" box="[151,267,1004,1030]" pageId="3" pageNumber="286" type="description">
<paragraph id="7129465AE97CFFDF0ACE098CD84EFBAB" blockId="3.[151,655,967,1030]" box="[151,267,1004,1030]" pageId="3" pageNumber="286">
(
<figureCitation id="E9AD5ADFE97CFFDF0AC6098CD846FBAB" box="[159,259,1004,1030]" captionStart="FIGURES 34" captionStartId="4.[151,264,1946,1970]" captionTargetBox="[153,1435,453,1922]" captionTargetId="figure-139@4.[151,1435,453,1922]" captionTargetPageId="4" captionText="FIGURES 34. Atomaria (Anchicera) perkovskyi sp. nov., No 6832 [MAIG]: 3—habitus, dorsal view; 4—habitus, ventrolateral view. Scale bar = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.7091697" httpUri="https://zenodo.org/record/7091697/files/figure.png" pageId="3" pageNumber="286">Figs 34</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="398C15D1E97CFFDF0ACE0E54DB95FA8B" pageId="3" pageNumber="286" type="materials_examined">
<paragraph id="7129465AE97CFFDF0ACE0E54D851FB73" blockId="3.[151,1437,1076,2039]" pageId="3" pageNumber="286">
<emphasis id="43E29A48E97CFFDF0ACE0E54D802FBE3" bold="true" box="[151,327,1076,1102]" pageId="3" pageNumber="286">Type material.</emphasis>
<typeStatus id="AE2DF8F8E97CFFDF0B160E54D8F9FBE3" box="[335,444,1076,1102]" pageId="3" pageNumber="286" type="holotype">Holotype</typeStatus>
: collection number No 6832 [
<collectionCode id="1787DE9FE97CFFDF09560E54DA1AFBE3" box="[783,863,1076,1102]" pageId="3" pageNumber="286">MAIG</collectionCode>
] (ex coll. Jonas Damzen JDC-10149
<collectionCode id="1787DE9FE97CFFDF0F5E0E54DC5FFBE3" box="[1287,1306,1076,1102]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="286">R</collectionCode>
); adult, sex unkown. A complete beetle with partially exposed metathoracic wings is included in a transparent, yellow amber piece with dimensions of 38×
<quantity id="B66EEBBFE97CFFDF0B820E1CDB6CFB3A" box="[475,553,1148,1175]" metricMagnitude="-2" metricUnit="m" metricValue="5.2" pageId="3" pageNumber="286" unit="mm" value="52.0">52 mm</quantity>
and a maximum thickness of
<quantity id="B66EEBBFE97CFFDF09370E1CDAFBFB3A" box="[878,958,1148,1175]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="286" unit="mm" value="15.0">15 mm</quantity>
; preserved without supplementary fixation. Ventral side of specimen completely obscured by milky opacity. Syninclusions: four small Nematocera (Diptera) specimens.
</paragraph>
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<materialsCitation id="C1FE4C07E97CFFDF0A9E0E88DAF2FAAF" ID-GBIF-Occurrence="3912658302" box="[199,951,1256,1283]" location="Late Eocene" pageId="3" pageNumber="286" specimenCount="1" stateProvince="Rivne" typeStatus="holotype">
<emphasis id="43E29A48E97CFFDF0A9E0E88D82AFAAF" bold="true" box="[199,367,1256,1282]" pageId="3" pageNumber="286">
<typeStatus id="AE2DF8F8E97CFFDF0A9E0E88D847FAAF" box="[199,258,1256,1282]" pageId="3" pageNumber="286">Type</typeStatus>
stratum.
</emphasis>
<collectingRegion id="B35288B8E97CFFDF0B2F0E89D884FAAE" box="[374,449,1257,1283]" country="Ukraine" name="Rivne" pageId="3" pageNumber="286">Rovno</collectingRegion>
amber,
<location id="74491081E97CFFDF08430E89DBE0FAAE" LSID="urn:lsid:plazi:treatment:F93FF74CE97CFFD90ACE09A8DA71FE58:74491081E97CFFDF08430E89DBE0FAAE" box="[538,677,1257,1283]" name="Late Eocene" pageId="3" pageNumber="286" stateProvince="Rivne">Late Eocene</location>
(
<bibRefCitation id="15073BABE97CFFDF08ED0E88DAEAFAAF" author="Perkovsky, E. E. &amp; Rasnitsyn, A. P. &amp; Vlaskin, A. P. &amp; Taraschuk, M. V." box="[692,943,1256,1283]" pageId="3" pageNumber="286" pagination="229 - 245" refId="ref4885" refString="Perkovsky, E. E., Rasnitsyn, A. P., Vlaskin, A. P. &amp; Taraschuk, M. V. (2007) A comparative analysis of the Baltic and Rovno amber arthropod faunas: representative samples. African Invertebrates, 48, 229 - 245. [https: // hdl. handle. net / 10520 / EJC 84578]" type="journal article" year="2007">
Perkovsky
<emphasis id="43E29A48E97CFFDF096A0E89DA21FAAF" box="[819,868,1256,1282]" italics="true" pageId="3" pageNumber="286">et al</emphasis>
. 2007
</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="7129465AE97CFFDF0A9E0F6CDB95FA8B" blockId="3.[151,1437,1076,2039]" box="[199,720,1292,1318]" pageId="3" pageNumber="286">
<materialsCitation id="C1FE4C07E97CFFDF0A9E0F6CDB88FA8B" ID-GBIF-Occurrence="3912658301" box="[199,717,1292,1318]" country="Ukraine" location="Rivne Oblast" pageId="3" pageNumber="286" specimenCount="1" stateProvince="Rivne" typeStatus="holotype">
<emphasis id="43E29A48E97CFFDF0A9E0F6CD823FA8B" bold="true" box="[199,358,1292,1318]" pageId="3" pageNumber="286">
<typeStatus id="AE2DF8F8E97CFFDF0A9E0F6CD847FA8B" box="[199,258,1292,1318]" pageId="3" pageNumber="286">Type</typeStatus>
locality.
</emphasis>
<collectingRegion id="B35288B8E97CFFDF0B350F6CD8BAFA8B" box="[364,511,1292,1318]" country="Ukraine" name="Rivne" pageId="3" pageNumber="286">Rivne Oblast</collectingRegion>
(region),
<collectingCountry id="098106CAE97CFFDF08360F6CDB88FA8B" box="[623,717,1292,1318]" name="Ukraine" pageId="3" pageNumber="286">Ukraine</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="398C15D1E97CFFD90A9E0F50D896FF78" lastPageId="5" lastPageNumber="288" pageId="3" pageNumber="286" type="description">
<paragraph id="7129465AE97CFFDF0A9E0F50DACCFA77" blockId="3.[151,1437,1076,2039]" pageId="3" pageNumber="286">
<emphasis id="43E29A48E97CFFDF0A9E0F50D81CFAE7" bold="true" box="[199,345,1328,1354]" pageId="3" pageNumber="286">Description.</emphasis>
Measurements: body length (from anterior margin of head to elytral apex along midline) about
<quantity id="B66EEBBFE97CFFDF0ACE0F34D9BBFAC2" box="[151,254,1364,1391]" metricMagnitude="-3" metricUnit="m" metricValue="1.19" pageId="3" pageNumber="286" unit="mm" value="1.19">1.19 mm</quantity>
, body maximum width across both elytron
<quantity id="B66EEBBFE97CFFDF08B10F34DA15FAC2" box="[744,848,1364,1391]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="286" unit="mm" value="0.5">0.50 mm</quantity>
; head length
<quantity id="B66EEBBFE97CFFDF09BF0F34DD08FAC2" box="[998,1101,1364,1391]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="286" unit="mm" value="0.16">0.16 mm</quantity>
, head maximum width across eyes
<quantity id="B66EEBBFE97CFFDF0A8D0F18D87AFA3E" box="[212,319,1400,1427]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="286" unit="mm" value="0.27">0.27 mm</quantity>
; pronotum length
<quantity id="B66EEBBFE97CFFDF084F0F18DBC5FA3E" box="[534,640,1400,1427]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="286" unit="mm" value="0.3">0.30 mm</quantity>
, pronotum maximum width
<quantity id="B66EEBBFE97CFFDF09970F18DD7CFA3E" box="[974,1081,1400,1427]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="3" pageNumber="286" unit="mm" value="0.38">0.38 mm</quantity>
; elytra length
<quantity id="B66EEBBFE97CFFDF0EBA0F18DC08FA3E" box="[1251,1357,1400,1427]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="3" pageNumber="286" unit="mm" value="0.73">0.73 mm</quantity>
, elytra maximum width
<quantity id="B66EEBBFE97CFFDF0B0C0FFCD8F9FA1A" box="[341,444,1436,1463]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="286" unit="mm" value="0.5">0.50 mm</quantity>
. Body broadly elongate, moderately arched, slightly convex; integument uniformly darkbrown (as preserved); sparsely covered with semierect pubescence.
</paragraph>
<paragraph id="7129465AE97CFFDF0A9E0F84DD4FF8CA" blockId="3.[151,1437,1076,2039]" pageId="3" pageNumber="286">Head transverse, about 1.7× as wide as long; coarsely punctate, distance between punctures equal to 0.5-1.0× diameter of one puncture, punctation denser laterally. Compound eyes hemispherical, convex, rather large, with large facets. Antennae inserted close to eyes, under lateral margin of forehead; antennal insertions widely separated; distance between antennal insertions larger than distance between antennae and аnterior margin of eye. Antennae 11-segmented with 3-segmented loose club, slender, short, reaching about to posterior one-third of pronotal length; antennomere 1 subcylindrical, elongate, 1.6× as long as wide; antennomere 2 conical, elongate, 1.5× as long as wide, slightly shorter than antennomere 1; antennomere 3 conical, short, nearly as long as wide, narrower and distinctly shorter than antennomere 2; antennomere 4 nearly as long as wide; antennomere 5 conical, nearly as long as wide, 1.1× as long as wide; antennomeres 68 trapezoidal, as long as wide; antennomeres 910 trapezoidal, strongly transverse, 1.5× as wide as long; antennomere 11 widely oval, with rounded apex, nearly as long as wide; relative length ratios of antennomeres 111 equal to 16:12:7:7:8:5:5:6:8:10:16.</paragraph>
<paragraph id="7129465AE97CFFD80A9E0D11DD90FF78" blockId="3.[151,1437,1076,2039]" lastBlockId="4.[151,1437,151,429]" lastPageId="4" lastPageNumber="287" pageId="3" pageNumber="286">Pronotum transverse, 1.2× as wide as long, slightly narrowed posteriad and anteriad, with maximum width in middle of its length; with slightly convex disc; with finely margined posterior and lateral sides; lateral side borders visible from above only in basal half. Pronotal punctation moderately coarse and dense, distance between punctures equal to 1.5-2.0× diameter of one puncture; puncture diameter slightly smaller than eye facet diameter. Posterior and anterior angles obtuse. Anterior edge weakly sinuate, without excision; posterior edge strongly bisinuate, lobed, with shallow depression and basal pits; lateral edges widely rounded, without callosity and teeth.</paragraph>
<paragraph id="7129465AE97BFFD80A9E0ABFDC66FF54" blockId="4.[151,1437,151,429]" box="[199,1315,223,249]" pageId="4" pageNumber="287">Scutellar shield small, oval, strongly transverse, 2.0× as wide as long, covered with fine punctation.</paragraph>
<paragraph id="7129465AE97BFFD80A9E0B63DB9CFE00" blockId="4.[151,1437,151,429]" pageId="4" pageNumber="287">Elytra short-oval, moderately convex, with maximum width about in anterior one-third of its length, 1.5× as long as wide and 2.4 times as long as pronotum length; completely covering the abdomen; lateral sides widely rounded; humeri rounded, slightly prominent. Elytral punctation irregular, moderately coarse and dense; (nearly as pronotal punctation), distance between punctures equal to 1.5-2.0× diameter of one puncture. Epipleura incomplete. Macropterous, metathoracic wings fully developed.</paragraph>
<caption id="25E916D2E97BFFD80ACE0DFAD8B6F878" ID-DOI="http://doi.org/10.5281/zenodo.7091697" ID-Zenodo-Dep="7091697" httpUri="https://zenodo.org/record/7091697/files/figure.png" pageId="4" pageNumber="287" startId="4.[151,264,1946,1970]" targetBox="[153,1435,453,1922]" targetPageId="4">
<paragraph id="7129465AE97BFFD80ACE0DFAD8B6F878" blockId="4.[151,1436,1946,2006]" pageId="4" pageNumber="287">
<emphasis id="43E29A48E97BFFD80ACE0DFAD878F81F" bold="true" box="[151,317,1946,1970]" pageId="4" pageNumber="287">FIGURES 34.</emphasis>
<taxonomicName id="B6963DD9E97BFFD80B1F0DFADBD9F81F" authority="Lyubarsky &amp; Bukejs, 2022" authorityName="Lyubarsky &amp; Bukejs" authorityYear="2022" box="[326,668,1946,1970]" class="Insecta" family="Cryptophagidae" genus="Atomaria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="287" phylum="Arthropoda" rank="species" species="perkovskyi" status="sp. nov." subGenus="Anchicera">
<emphasis id="43E29A48E97BFFD80B1F0DFADBD9F81F" box="[326,668,1946,1970]" italics="true" pageId="4" pageNumber="287">Atomaria (Anchicera) perkovskyi</emphasis>
</taxonomicName>
<emphasis id="43E29A48E97BFFD808FC0DFADBBDF81F" bold="true" box="[677,760,1946,1970]" pageId="4" pageNumber="287">
<taxonomicNameLabel id="58D12733E97BFFD808FC0DFADBBDF81F" box="[677,760,1946,1970]" pageId="4" pageNumber="287" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, No 6832 [MAIG]: 3—habitus, dorsal view; 4—habitus, ventrolateral view. Scale bar = 0.25 mm.
</paragraph>
</caption>
<paragraph id="7129465AE97AFFD90A9E0AF7D896FF78" blockId="5.[151,1437,151,501]" pageId="5" pageNumber="288">Legs slender, long, relatively similar in shape, finely and densely punctate. Femora elongate-oval, flattened. Pretarsal claws simple, thin.</paragraph>
</subSubSection>
<subSubSection id="398C15D1E97AFFD90A9E0ABFDBF9FEB0" pageId="5" pageNumber="288" type="etymology">
<paragraph id="7129465AE97AFFD90A9E0ABFDBF9FEB0" blockId="5.[151,1437,151,501]" pageId="5" pageNumber="288">
<emphasis id="43E29A48E97AFFD90A9E0ABFD80BFF54" bold="true" box="[199,334,223,249]" pageId="5" pageNumber="288">Etymology.</emphasis>
The species is named in honor of our dear colleague Dr. Evgeny E. Perkovsky (
<collectingRegion id="B35288B8E97AFFD90EB50ABFDC60FF54" box="[1260,1317,223,249]" country="Ukraine" name="Kyiv" pageId="5" pageNumber="288">Kiev</collectingRegion>
,
<collectingCountry id="098106CAE97AFFD90F6B0ABFDCD4FF54" box="[1330,1425,223,249]" name="Ukraine" pageId="5" pageNumber="288">Ukraine</collectingCountry>
), leading researcher of inclusions in
<collectingRegion id="B35288B8E97AFFD908470B63DB2CFEB0" box="[542,617,259,285]" country="Ukraine" name="Rivne" pageId="5" pageNumber="288">Rovno</collectingRegion>
amber.
</paragraph>
</subSubSection>
<subSubSection id="398C15D1E97AFFD90A9E0B46DA71FE58" pageId="5" pageNumber="288" type="diagnosis">
<paragraph id="7129465AE97AFFD90A9E0B46DA71FE58" blockId="5.[151,1437,151,501]" pageId="5" pageNumber="288">
<emphasis id="43E29A48E97AFFD90A9E0B46D889FEED" bold="true" box="[199,460,294,320]" pageId="5" pageNumber="288">Differential diagnosis.</emphasis>
<taxonomicName id="B6963DD9E97AFFD90B890B47DBF2FEEC" authority="Lyubarsky &amp; Bukejs, 2022" authorityName="Lyubarsky &amp; Bukejs" authorityYear="2022" box="[464,695,295,321]" class="Insecta" family="Cryptophagidae" genus="Atomaria" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="288" phylum="Arthropoda" rank="species" species="perkovskyi" status="sp. nov.">
<emphasis id="43E29A48E97AFFD90B890B47DBF2FEEC" box="[464,695,295,321]" italics="true" pageId="5" pageNumber="288">Atomaria perkovskyi</emphasis>
</taxonomicName>
<emphasis id="43E29A48E97AFFD908E20B47DA56FEEC" bold="true" box="[699,787,295,321]" pageId="5" pageNumber="288">
<taxonomicNameLabel id="58D12733E97AFFD908E20B47DA56FEEC" box="[699,787,295,321]" pageId="5" pageNumber="288" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from related species by a combination of characters: antennomere 3 short, nearly as long as wide, distinctly shorter than antennomere 2; antennomeres 9 and 10 strongly transverse; antennomere 5 conical, nearly as long as wide, 1.1× as long as wide; pronotum narrowed toward base; lateral side borders of pronotum visible from above only in basal half of its length; and pronotum maximum width in middle of its length. A new species can distinguished from all other extinct Eocene
<taxonomicName id="B6963DD9E97AFFD90F6B0BD7DCD9FE7C" authorityName="Stephens" authorityYear="1829" box="[1330,1436,439,465]" class="Insecta" family="Cryptophagidae" genus="Atomaria" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="288" phylum="Arthropoda" rank="genus">
<emphasis id="43E29A48E97AFFD90F6B0BD7DCD9FE7C" box="[1330,1436,439,465]" italics="true" pageId="5" pageNumber="288">Atomaria</emphasis>
</taxonomicName>
representatives using the identification key provided below.
</paragraph>
</subSubSection>
</treatment>
</document>