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<mods:title id="1DF29DB2ED96CC10C9C1E6C24FD55A87">A study of the scale insect genera Puto Signoret (Hemiptera: Sternorrhyncha: Coccoidea: Putoidae) and Ceroputo Šulc (Pseudococcidae) with a comparison to Phenacoccus Cockerell (Pseudococcidae)</mods:title>
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<subSubSection id="C34A6561FFA9FF9CFF259BC0E010A5A8" box="[151,324,1493,1519]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8BEF36EAFFA9FF9CFF259BC0E010A5A8" blockId="13.[151,324,1493,1519]" box="[151,324,1493,1519]" pageId="13" pageNumber="14">
<heading id="D0A78186FFA9FF9CFF259BC0E010A5A8" bold="true" box="[151,324,1493,1519]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<taxonomicName id="4C504D69FFA9FF9CFF259BC0E010A5A8" authority="Sulc" authorityName="Sulc" box="[151,324,1493,1519]" class="Insecta" family="Pseudococcidae" genus="Ceroputo" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAF8FFA9FF9CFF259BC0E010A5A8" bold="true" box="[151,324,1493,1519]" pageId="13" pageNumber="14">
<emphasis id="B924EAF8FFA9FF9CFF259BC0E053A5A8" bold="true" box="[151,263,1493,1519]" italics="true" pageId="13" pageNumber="14">Ceroputo</emphasis>
Šulc
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34A6561FFA9FF9FFF259809E434A15B" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="reference_group">
<paragraph id="8BEF36EAFFA9FF9CFF259809E0C6A628" blockId="13.[151,1436,1563,1647]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="AB4011C4FFA9FF9CFF259809E0C6A628" pageId="13" pageNumber="14">
<taxonomicName id="4C504D69FFA9FF9CFF259809E0D7A676" authority="Sulc, 1898: 1" authorityName="Sulc" authorityPageNumber="1" authorityYear="1898" box="[151,387,1563,1585]" class="Insecta" family="Pseudococcidae" genus="Ceroputo" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAF8FFA9FF9CFF259809E1A2A676" box="[151,246,1564,1585]" italics="true" pageId="13" pageNumber="14">Ceroputo</emphasis>
<treatmentCitation id="0AF110FBFFA9FF9CFF4E980EE0D7A676" author="Sulc" box="[252,387,1563,1585]" page="1" pageId="13" pageNumber="14" year="1898">
<bibRefCitation id="EFC14B1BFFA9FF9CFF4E980EE03FA676" author="Sulc" box="[252,363,1563,1585]" pageId="13" pageNumber="14" refString="Sulc, K. (1898) Studie o coccidech. - II. Sitzungsberichte der K. Bohmisch Gesellschaft der Wissenschaften (1897) 66, 1 - 19." type="journal article" year="1898">Šulc, 1898</bibRefCitation>
: 1
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="54EB8848FFA9FF9CFE3D9808E095A675" box="[399,449,1565,1586]" pageId="13" pageNumber="14">Type</typeStatus>
species:
<taxonomicName id="4C504D69FFA9FF9CFDAC9809E248A676" authority="Sulc" authorityName="Sulc" box="[542,796,1563,1586]" class="Insecta" family="Pseudococcidae" genus="Ceroputo" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pilosellae">
<emphasis id="B924EAF8FFA9FF9CFDAC9809E3B1A675" box="[542,741,1564,1586]" italics="true" pageId="13" pageNumber="14">Ceroputo pilosellae</emphasis>
Šulc
</taxonomicName>
, by original designation.
<taxonomicName id="4C504D69FFA9FF9CFF25982EE096A617" authority="Kanda, 1959: 239" authorityName="Kanda" authorityPageNumber="239" authorityYear="1959" box="[151,450,1594,1616]" class="Insecta" family="Pseudococcidae" genus="Leococcus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAF8FFA9FF9CFF25982EE056A617" box="[151,258,1595,1616]" italics="true" pageId="13" pageNumber="14">Leococcus</emphasis>
<treatmentCitation id="0AF110FBFFA9FF9CFEBB982FE096A617" author="Kanda" box="[265,450,1594,1616]" page="239" pageId="13" pageNumber="14" year="1959">
<bibRefCitation id="EFC14B1BFFA9FF9CFEBB982FE0DAA617" author="Kanda" box="[265,398,1594,1616]" pageId="13" pageNumber="14" refString="Kanda, S. (1959) Two new genera and two new species of the Pseudococcidae (Homoptera). Kontyu, 27, 239 - 243." type="journal article" year="1959">Kanda, 1959</bibRefCitation>
: 239
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="54EB8848FFA9FF9CFE7E982EE0AAA617" box="[460,510,1595,1616]" pageId="13" pageNumber="14">Type</typeStatus>
species:
<taxonomicName id="4C504D69FFA9FF9CFDEE982EE2C6A617" authority="Kanda" authorityName="Kanda" box="[604,914,1594,1616]" class="Insecta" family="Pseudococcidae" genus="Leococcus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="erigeroneus">
<emphasis id="B924EAF8FFA9FF9CFDEE982EE212A617" box="[604,838,1595,1616]" italics="true" pageId="13" pageNumber="14">Leococcus erigeroneus</emphasis>
Kanda
</taxonomicName>
, by monotypy and original designation. Synonymy by
<treatmentCitation id="0AF110FBFFA9FF9CFF54984FE0DAA628" author="Tang" box="[230,398,1626,1647]" page="388" pageId="13" pageNumber="14" year="1992">
<bibRefCitation id="EFC14B1BFFA9FF9CFF54984FE00FA628" author="Tang" box="[230,347,1626,1647]" pageId="13" pageNumber="14" refString="Tang, F. D. (1992) The Pseudococcidae of China (Homoptera: Coccinea of Insects). Beijing, Chinese Agricultural Science Technology Press, 767 pp." type="book" year="1992">Tang, 1992</bibRefCitation>
: 388
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEF36EAFFA9FF9FFF259888E434A15B" blockId="13.[151,1436,1692,2004]" lastBlockId="14.[151,1437,151,680]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<taxonomicName id="4C504D69FFA9FF9CFF259888E054A6F3" box="[151,256,1693,1716]" class="Insecta" family="Pseudococcidae" genus="Ceroputo" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAF8FFA9FF9CFF259888E054A6F3" box="[151,256,1693,1716]" italics="true" pageId="13" pageNumber="14">Ceroputo</emphasis>
</taxonomicName>
was synonymised with
<taxonomicName id="4C504D69FFA9FF9CFDA09888E313A6F3" ID-CoL="75LQ" box="[530,583,1693,1716]" class="Insecta" family="Pseudococcidae" genus="Puto" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAF8FFA9FF9CFDA09888E313A6F3" box="[530,583,1693,1716]" italics="true" pageId="13" pageNumber="14">Puto</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFC14B1BFFA9FF9CFDC19889E253A6F3" author="Ferris" box="[627,775,1692,1716]" pageId="13" pageNumber="14" refString="Ferris, G. F. (1918) The California species of mealy bugs. Leland Stanford Junior University Publications, University Series, Palo Alto, 78 pp." type="book" year="1918">Ferris (1918)</bibRefCitation>
[not by
<bibRefCitation id="EFC14B1BFFA9FF9CFCDB9889E2A8A6F3" author="Danzig" box="[873,1020,1692,1716]" pageId="13" pageNumber="14" refString="Danzig, E. M. (1980) Scale insects of the Far East of the USSR (Homoptera, Coccinea) SSSR [Keys to the fauna of the USSR]. (in Russian) Nauka, Leningrad, 368 pp." type="book" year="1980">Danzig 1980</bibRefCitation>
, as suggested by
<bibRefCitation id="EFC14B1BFFA9FF9CFB769888E43FA6F3" author="Ben-Dov" box="[1220,1387,1692,1716]" pageId="13" pageNumber="14" refString="Ben-Dov, Y. (1994) A Systematic Catalogue of the Mealybugs of the World (Insecta: Homoptera: Coccoidea: Pseudococcidae and Putoidae) with Data on Geographical Distribution, Host Plants, Biology and Economic Importance. Intercept Ltd, Andover, U. K., 686 pp." type="book" year="1994">Ben-Dov 1994</bibRefCitation>
and ScaleNet], but treated as a valid genus by
<bibRefCitation id="EFC14B1BFFA9FF9CFD3498D5E241A690" author="Tang" box="[646,789,1727,1752]" pageId="13" pageNumber="14" refString="Tang, F. D. (1992) The Pseudococcidae of China (Homoptera: Coccinea of Insects). Beijing, Chinese Agricultural Science Technology Press, 767 pp." type="book" year="1992">Tang (1992)</bibRefCitation>
.
<bibRefCitation id="EFC14B1BFFA9FF9CFC9198AAE556A690" box="[803,1026,1727,1752]" pageId="13" pageNumber="14" refString="Hardy, N. B., Gullan, P. J. &amp; Hodgson, C. J. (2008) A classification of mealybugs (Hemiptera: Coccoidea) based on integrated molecular and morphological data. Systematic Entomolog y, 33, 51 - 71." type="journal article">
Hardy
<emphasis id="B924EAF8FFA9FF9CFCC198D4E2F3A69F" box="[883,935,1728,1752]" italics="true" pageId="13" pageNumber="14">et al</emphasis>
. (2008)
</bibRefCitation>
treated
<taxonomicName id="4C504D69FFA9FF9CFBD098D5E59FA690" box="[1122,1227,1728,1751]" class="Insecta" family="Pseudococcidae" genus="Ceroputo" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAF8FFA9FF9CFBD098D5E59FA690" box="[1122,1227,1728,1751]" italics="true" pageId="13" pageNumber="14">Ceroputo</emphasis>
</taxonomicName>
as a genus of the mealybug subfamily Phenacoccinae, which they redefined. We concur with
<bibRefCitation id="EFC14B1BFFA9FF9CFC4498F0E5D5A6BB" author="Tang" box="[1014,1153,1764,1789]" pageId="13" pageNumber="14" refString="Tang, F. D. (1992) The Pseudococcidae of China (Homoptera: Coccinea of Insects). Beijing, Chinese Agricultural Science Technology Press, 767 pp." type="book" year="1992">Tang (1992)</bibRefCitation>
and
<bibRefCitation id="EFC14B1BFFA9FF9CFB0898F1E4CCA6BB" box="[1210,1432,1764,1788]" pageId="13" pageNumber="14" refString="Hardy, N. B., Gullan, P. J. &amp; Hodgson, C. J. (2008) A classification of mealybugs (Hemiptera: Coccoidea) based on integrated molecular and morphological data. Systematic Entomolog y, 33, 51 - 71." type="journal article">
Hardy
<emphasis id="B924EAF8FFA9FF9CFABA98F0E46FA6BB" box="[1288,1339,1764,1788]" italics="true" pageId="13" pageNumber="14">et al</emphasis>
. (2008)
</bibRefCitation>
. The molecular phylogenetic studies by M. Bora Kaydan (pers. comm.) also have confirmed the placement of
<taxonomicName id="4C504D69FFA9FF9CFA30991DE050A703" class="Insecta" family="Pseudococcidae" genus="Ceroputo" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pilosellae">
<emphasis id="B924EAF8FFA9FF9CFA30991DE050A703" italics="true" pageId="13" pageNumber="14">C. pilosellae</emphasis>
</taxonomicName>
in the
<taxonomicName id="4C504D69FFA9FF9CFEE19939E352A703" ID-CoL="F8D" box="[339,518,1836,1860]" class="Insecta" family="Pseudococcidae" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="family">Pseudococcidae</taxonomicName>
near some species of
<taxonomicName id="4C504D69FFA9FF9CFD4D9939E2C7A703" ID-CoL="6N5N" box="[767,915,1836,1860]" class="Insecta" family="Pseudococcidae" genus="Phenacoccus" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAF8FFA9FF9CFD4D9939E2C7A703" box="[767,915,1836,1860]" italics="true" pageId="13" pageNumber="14">Phenacoccus</emphasis>
</taxonomicName>
. Here we diagnose
<taxonomicName id="4C504D69FFA9FF9CFBC19938E58FA703" box="[1139,1243,1837,1860]" class="Insecta" family="Pseudococcidae" genus="Ceroputo" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAF8FFA9FF9CFBC19938E58FA703" box="[1139,1243,1837,1860]" italics="true" pageId="13" pageNumber="14">Ceroputo</emphasis>
</taxonomicName>
based largely on the adult female and transfer several species to it from
<taxonomicName id="4C504D69FFA9FF9CFCB09944E263A72F" ID-CoL="75LQ" box="[770,823,1873,1896]" class="Insecta" family="Pseudococcidae" genus="Puto" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAF8FFA9FF9CFCB09944E263A72F" box="[770,823,1873,1896]" italics="true" pageId="13" pageNumber="14">Puto</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFC14B1BFFA9FF9CFCF7995AE29AA720" author="Afifi" box="[837,974,1871,1895]" pageId="13" pageNumber="14" refString="Afifi, S. A. (1968) Morphology and taxonomy of the adult males of Pseudococcidae and Eriococcidae (Homoptera: Coccoidea). Bulletin of the British Museum (Natural History) Entomology, Supplement, 13, 1 - 210." type="journal article" year="1968">Afifi (1968)</bibRefCitation>
described the adult male of
<taxonomicName id="4C504D69FFA9FF9CFABC9945E4C8A72F" box="[1294,1436,1872,1896]" class="Insecta" family="Pseudococcidae" genus="Ceroputo" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pilosellae">
<emphasis id="B924EAF8FFA9FF9CFABC9945E4C8A72F" box="[1294,1436,1872,1896]" italics="true" pageId="13" pageNumber="14">C. pilosellae</emphasis>
</taxonomicName>
(based on specimens from
<taxonomicName id="4C504D69FFA9FF9CFE709960E339A7CB" ID-CoL="6JK5R" box="[450,621,1909,1932]" class="Magnoliopsida" family="Rosaceae" genus="Fragaria" kingdom="Plantae" order="Rosales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="species" species="vesca">
<emphasis id="B924EAF8FFA9FF9CFE709960E339A7CB" box="[450,621,1909,1932]" italics="true" pageId="13" pageNumber="14">Fragaria vesca</emphasis>
</taxonomicName>
from Belgrade, in the former
<collectingCountry id="F347767AFFA9FF9CFC0E9961E56CA7CB" box="[956,1080,1908,1932]" name="Serbia" pageId="13" pageNumber="14">Yugoslavia</collectingCountry>
now
<collectingCountry id="F347767AFFA9FF9CFBC49961E5EBA7CB" box="[1142,1215,1908,1932]" name="Serbia" pageId="13" pageNumber="14">Serbia</collectingCountry>
) as having a pair of dorsal and a pair of ventral simple eyes and a pair of lateral ocelli. It also has two pairs of lateral filaments on the abdomen, features that are consistent with this species being a true pseudococcid. We also recognise
<taxonomicName id="4C504D69FFA9FF9FFA9499A8E1B4A0E8" authority="Kanda" authorityName="Kanda" class="Insecta" family="Pseudococcidae" genus="Leococcus" higherTaxonomySource="CoL" kingdom="Animalia" lastPageId="14" lastPageNumber="15" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAF8FFA9FF9CFA9499A8E4C8A793" box="[1318,1436,1981,2004]" italics="true" pageId="13" pageNumber="14">Leococcus</emphasis>
Kanda
</taxonomicName>
as a junior synonym of
<taxonomicName id="4C504D69FFAAFF9FFE489E8DE337A0E8" box="[506,611,152,175]" class="Insecta" family="Pseudococcidae" genus="Ceroputo" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAF8FFAAFF9FFE489E8DE337A0E8" box="[506,611,152,175]" italics="true" pageId="14" pageNumber="15">Ceroputo</emphasis>
</taxonomicName>
, in agreement with
<bibRefCitation id="EFC14B1BFFAAFF9FFCF89E8DE28CA0E8" author="Tang" box="[842,984,151,176]" pageId="14" pageNumber="15" refString="Tang, F. D. (1992) The Pseudococcidae of China (Homoptera: Coccinea of Insects). Beijing, Chinese Agricultural Science Technology Press, 767 pp." type="book" year="1992">Tang (1992)</bibRefCitation>
. Previously
<taxonomicName id="4C504D69FFAAFF9FFBD49E8CE588A0F7" box="[1126,1244,153,176]" class="Insecta" family="Pseudococcidae" genus="Leococcus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAF8FFAAFF9FFBD49E8CE588A0F7" box="[1126,1244,153,176]" italics="true" pageId="14" pageNumber="15">Leococcus</emphasis>
</taxonomicName>
was treated as a junior synonym of
<taxonomicName id="4C504D69FFAAFF9FFEC39EA8E0F2A093" ID-CoL="75LQ" box="[369,422,189,212]" class="Insecta" family="Pseudococcidae" genus="Puto" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAF8FFAAFF9FFEC39EA8E0F2A093" box="[369,422,189,212]" italics="true" pageId="14" pageNumber="15">Puto</emphasis>
</taxonomicName>
, a synonymy suggested by
<bibRefCitation id="EFC14B1BFFAAFF9FFD539EA9E2D4A093" author="Kawai" box="[737,896,188,212]" pageId="14" pageNumber="15" refString="Kawai, S. (1980) Scale Insects of Japan in Colors. Tokyo, Japan, National Agricultural Education Association, 455 pp." type="book" year="1980">Kawai (1980)</bibRefCitation>
.
<bibRefCitation id="EFC14B1BFFAAFF9FFC3F9EA9E57DA093" author="Kanda" box="[909,1065,188,212]" pageId="14" pageNumber="15" refString="Kanda, S. (1959) Two new genera and two new species of the Pseudococcidae (Homoptera). Kontyu, 27, 239 - 243." type="journal article" year="1959">Kanda (1959)</bibRefCitation>
stated how his genus
<taxonomicName id="4C504D69FFAAFF9FFA949EA8E4C8A093" box="[1318,1436,189,212]" class="Insecta" family="Pseudococcidae" genus="Leococcus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAF8FFAAFF9FFA949EA8E4C8A093" box="[1318,1436,189,212]" italics="true" pageId="14" pageNumber="15">Leococcus</emphasis>
</taxonomicName>
differed from
<taxonomicName id="4C504D69FFAAFF9FFE819EF4E03CA0BF" ID-CoL="75LQ" box="[307,360,225,248]" class="Insecta" family="Pseudococcidae" genus="Puto" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAF8FFAAFF9FFE819EF4E03CA0BF" box="[307,360,225,248]" italics="true" pageId="14" pageNumber="15">Puto</emphasis>
</taxonomicName>
and
<taxonomicName id="4C504D69FFAAFF9FFE2C9EF5E353A0B0" box="[414,519,224,247]" class="Insecta" family="Pseudococcidae" genus="Ceroputo" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAF8FFAAFF9FFE2C9EF5E353A0B0" box="[414,519,224,247]" italics="true" pageId="14" pageNumber="15">Ceroputo</emphasis>
</taxonomicName>
but the features that he listed are all shared with the
<typeStatus id="54EB8848FFAAFF9FFBE69EF5E5D0A0BF" box="[1108,1156,224,248]" pageId="14" pageNumber="15">type</typeStatus>
species of
<taxonomicName id="4C504D69FFAAFF9FFAB39EF5E43EA0B0" box="[1281,1386,224,247]" class="Insecta" family="Pseudococcidae" genus="Ceroputo" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAF8FFAAFF9FFAB39EF5E43EA0B0" box="[1281,1386,224,247]" italics="true" pageId="14" pageNumber="15">Ceroputo</emphasis>
</taxonomicName>
. We recognise a total of six species in
<taxonomicName id="4C504D69FFAAFF9FFDA29F10E32DA15B" box="[528,633,261,284]" class="Insecta" family="Pseudococcidae" genus="Ceroputo" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAF8FFAAFF9FFDA29F10E32DA15B" box="[528,633,261,284]" italics="true" pageId="14" pageNumber="15">Ceroputo</emphasis>
</taxonomicName>
and discuss each of these species following the generic diagnosis.
</paragraph>
</subSubSection>
<subSubSection id="C34A6561FFAAFF9FFF759F32E4C8A2E0" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="8BEF36EAFFAAFF9FFF759F32E4C8A2E0" blockId="14.[151,1437,151,680]" pageId="14" pageNumber="15">
<emphasis id="B924EAF8FFAAFF9FFF759F32E0FDA107" bold="true" box="[199,425,295,320]" pageId="14" pageNumber="15">Diagnosis features.</emphasis>
<emphasis id="B924EAF8FFAAFF9FFE029F3DE36BA107" box="[432,575,296,320]" italics="true" pageId="14" pageNumber="15">Adult female</emphasis>
: eye height as great as or greater than length of first antennal segment (usually very much smaller in typical
<taxonomicName id="4C504D69FFAAFF9FFDA19F59E3F3A123" box="[531,679,332,356]" class="Insecta" family="Pseudococcidae" genus="Phenacoccus" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAF8FFAAFF9FFDA19F59E3F3A123" box="[531,679,332,356]" italics="true" pageId="14" pageNumber="15">Phenacoccus</emphasis>
</taxonomicName>
, except
<taxonomicName id="4C504D69FFAAFF9FFCB49F59E2FDA123" box="[774,937,332,356]" class="Insecta" family="Pseudococcidae" genus="Phenacoccus" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="solenopsis">
<emphasis id="B924EAF8FFAAFF9FFCB49F59E2FDA123" box="[774,937,332,356]" italics="true" pageId="14" pageNumber="15">
<collectingCountry id="F347767AFFAAFF9FFCB49F59E27CA123" box="[774,808,332,356]" name="Philippines" pageId="14" pageNumber="15">Ph</collectingCountry>
. solenopsis
</emphasis>
</taxonomicName>
); antennae with 89 segments; 02 (usually 0) antennal intersegmental sensilla present between each of segments IIIIV, IVV and VIVII; 2 campaniform sensilla on each surface of each trochanter; claw lacking basal spurs; claw digitules capitate but tarsal digitules either capitate or not; more than 17 pairs of cerarii on sclerotised plates; long tubular ducts present or absent on frons (anterior to mouthparts); trilocular pores on venter usually smaller than those on dorsum, and pores in cerarii at most only slightly larger than those on rest of dorsum; quinquelocular pores sometimes present.
<emphasis id="B924EAF8FFAAFF9FFAA99C14E1BFA27B" italics="true" pageId="14" pageNumber="15">First-instar nymph:</emphasis>
6-segmented antennae; multilocular pores with&gt;5 loculi absent.
<emphasis id="B924EAF8FFAAFF9FFC649C31E45CA27B" box="[982,1288,548,572]" italics="true" pageId="14" pageNumber="15">
Adult male of
<taxonomicName id="4C504D69FFAAFF9FFBCB9C30E45CA27B" box="[1145,1288,548,572]" class="Insecta" family="Pseudococcidae" genus="Ceroputo" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pilosellae">C. pilosellae</taxonomicName>
</emphasis>
(adult males unknown in other species): head with one pair of dorsal and one pair of ventral simple eyes plus a pair of lateral ocelli; with 2 pairs of lateral filaments on posterior abdominal segments (each filament from glandular pouch on each side of abdominal segments VII and VIII); penial sheath 1-segmented, apex of aedeagus simple (
<bibRefCitation id="EFC14B1BFFAAFF9FFABF9C9AE4DBA2E0" author="Afifi" box="[1293,1423,655,679]" pageId="14" pageNumber="15" refString="Afifi, S. A. (1968) Morphology and taxonomy of the adult males of Pseudococcidae and Eriococcidae (Homoptera: Coccoidea). Bulletin of the British Museum (Natural History) Entomology, Supplement, 13, 1 - 210." type="journal article" year="1968">Afifi, 1968</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>