515 lines
63 KiB
XML
515 lines
63 KiB
XML
<document ID-DOI="10.37520/fi.2022.017" ID-ISSN="2533-4069" ID-Zenodo-Dep="7522682" checkinTime="1673381873780" checkinUser="felipe" docAuthor="Kvaček, Jiří" docDate="2022" docId="03F987C9115DFFF7FC7967F8E74A8CD8" docLanguage="en" docName="FossilImprint.78.2.425-431.pdf" docOrigin="Fossil Imprint 78 (2)" docSource="http://dx.doi.org/10.37520/fi.2022.017" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.3:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="3" docTitle="Zamites pateri J. KVACEK 2022, sp. nov." docType="treatment" docVersion="3" lastPageNumber="430" masterDocId="FFC0FFB1115FFFF2FFAC676FE50D8E20" masterDocTitle="New Species Of Zamites From The Cenomanian Of The Bohemian Cretaceous Basin" masterLastPageNumber="431" masterPageNumber="425" pageNumber="427" updateTime="1673875764197" updateUser="juliana" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New Species Of Zamites From The Cenomanian Of The Bohemian Cretaceous Basin</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kvaček, Jiří</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Fossil Imprint</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-12-20</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>78</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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<mods:extent unit="page">
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<mods:start>425</mods:start>
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<mods:end>431</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.37520/fi.2022.017</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.37520/fi.2022.017</mods:identifier>
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<mods:identifier type="ISSN">2533-4069</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7522682</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:03F987C9115DFFF7FC7967F8E74A8CD8" httpUri="http://treatment.plazi.org/id/03F987C9115DFFF7FC7967F8E74A8CD8" lastPageId="5" lastPageNumber="430" pageId="2" pageNumber="427">
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<subSubSection box="[981,1344,151,175]" pageId="2" pageNumber="427" type="nomenclature">
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<paragraph blockId="2.[981,1344,151,175]" box="[981,1344,151,175]" pageId="2" pageNumber="427">
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<heading box="[981,1344,151,175]" centered="true" fontSize="8" level="2" pageId="2" pageNumber="427" reason="2">
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<emphasis bold="true" box="[981,1344,151,175]" pageId="2" pageNumber="427">
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<taxonomicName authority="J.KVACEK" authorityName="J. KVACEK" authorityYear="2022" box="[981,1251,151,175]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="2" pageNumber="427" phylum="Tracheophyta" rank="species" species="pateri" status="sp. nov.">
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<emphasis bold="true" box="[981,1139,151,175]" italics="true" pageId="2" pageNumber="427">Zamites pateri</emphasis>
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J.KVAČEK
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</taxonomicName>
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<taxonomicNameLabel box="[1258,1344,151,175]" pageId="2" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[1092,1232,183,207]" pageId="2" pageNumber="427" type="description">
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<paragraph blockId="2.[1092,1232,183,207]" box="[1092,1232,183,207]" pageId="2" pageNumber="427">
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<figureCitation box="[1092,1206,183,207]" captionStart="Text-fig" captionStartId="1.[125,206,1966,1987]" captionTargetBox="[125,1440,151,1950]" captionTargetId="figure-4@1.[-161,1348,77,1084]" captionTargetPageId="1" captionText="Text-fig. 1. Zamites pateri J.KVAČEK sp. nov.; Pecínov locality, holotype, No. NM-F 5185. a: Holotype overview showing fragment of simply pinnate leaf, scale bar 20 mm. b: Abaxial cuticle showing costal and intercostal scale zones and stomata in ill-defined rows, LM micro-photograph, scale bar 100 µm. c: Pinnule detail showing venation pattern, scale bar 5 mm. d: Abaxial cuticle showing costal and intercostal zones, LM micro-photograph, scale bar 500 µm. e: Fragmentary preserved adaxial cuticle showing" figureDoi="http://doi.org/10.5281/zenodo.7522684" httpUri="https://zenodo.org/record/7522684/files/figure.png" pageId="2" pageNumber="427">Text-figs 1</figureCitation>
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,
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<figureCitation box="[1219,1232,183,207]" captionStart="Text-fig" captionStartId="3.[125,206,1879,1900]" captionTargetBox="[125,1440,136,1861]" captionTargetPageId="3" captionText="Text-fig. 2. Zamites pateri J.KVAČEK sp. nov.; Pecínov locality, holotype, No. NM-F 5185. a: Abaxial cuticle showing costal and intercostal zones, SEM micro-photograph, scale bar 100 µm. b: Abaxial cuticle, detail of syndetocheilic stoma, SEM microphotograph, scale bar 10 µm. c: Abaxial cuticle showing transversely oriented stomata, SEM micro-photograph, scale bar 50 µm. d: Abaxial cuticle, detail of syndetocheilic stoma showing ledges of guard cells, SEM micro-photograph, scale bar 10 µm. e: Abaxial cuticle showing costal ordinary cells seen from inside, SEM micro-photograph, scale bar 50 µm. f: External side of abaxial cuticle showing stoma sunken in a stomatal pit surrounded by papillae, SEM micro-photograph, scale bar 10 µm. g: External side of abaxial cuticle showing papillae, SEM micro-photograph, scale bar 100 µm. h: External side of abaxial cuticle showing detail of fused papillae, SEM micro-photograph, scale bar 10 µm." figureDoi="http://doi.org/10.5281/zenodo.7522688" httpUri="https://zenodo.org/record/7522688/files/figure.png" pageId="2" pageNumber="427">2</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection box="[881,1323,230,255]" pageId="2" pageNumber="427" type="type_taxon">
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<paragraph blockId="2.[881,1323,230,255]" box="[881,1323,230,255]" pageId="2" pageNumber="427">
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H o l o t y p e. NM-F 5185,
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<figureCitation box="[1175,1289,231,255]" captionStart="Text-fig" captionStartId="1.[125,206,1966,1987]" captionTargetBox="[125,1440,151,1950]" captionTargetId="figure-4@1.[-161,1348,77,1084]" captionTargetPageId="1" captionText="Text-fig. 1. Zamites pateri J.KVAČEK sp. nov.; Pecínov locality, holotype, No. NM-F 5185. a: Holotype overview showing fragment of simply pinnate leaf, scale bar 20 mm. b: Abaxial cuticle showing costal and intercostal scale zones and stomata in ill-defined rows, LM micro-photograph, scale bar 100 µm. c: Pinnule detail showing venation pattern, scale bar 5 mm. d: Abaxial cuticle showing costal and intercostal zones, LM micro-photograph, scale bar 500 µm. e: Fragmentary preserved adaxial cuticle showing" figureDoi="http://doi.org/10.5281/zenodo.7522684" httpUri="https://zenodo.org/record/7522684/files/figure.png" pageId="2" pageNumber="427">Text-figs 1</figureCitation>
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,
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<figureCitation box="[1303,1323,231,255]" captionStart="Text-fig" captionStartId="3.[125,206,1879,1900]" captionTargetBox="[125,1440,136,1861]" captionTargetPageId="3" captionText="Text-fig. 2. Zamites pateri J.KVAČEK sp. nov.; Pecínov locality, holotype, No. NM-F 5185. a: Abaxial cuticle showing costal and intercostal zones, SEM micro-photograph, scale bar 100 µm. b: Abaxial cuticle, detail of syndetocheilic stoma, SEM microphotograph, scale bar 10 µm. c: Abaxial cuticle showing transversely oriented stomata, SEM micro-photograph, scale bar 50 µm. d: Abaxial cuticle, detail of syndetocheilic stoma showing ledges of guard cells, SEM micro-photograph, scale bar 10 µm. e: Abaxial cuticle showing costal ordinary cells seen from inside, SEM micro-photograph, scale bar 50 µm. f: External side of abaxial cuticle showing stoma sunken in a stomatal pit surrounded by papillae, SEM micro-photograph, scale bar 10 µm. g: External side of abaxial cuticle showing papillae, SEM micro-photograph, scale bar 100 µm. h: External side of abaxial cuticle showing detail of fused papillae, SEM micro-photograph, scale bar 10 µm." figureDoi="http://doi.org/10.5281/zenodo.7522688" httpUri="https://zenodo.org/record/7522688/files/figure.png" pageId="2" pageNumber="427">2.</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="427" type="reference_group">
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<paragraph blockId="2.[843,1479,278,334]" box="[881,1479,278,302]" pageId="2" pageNumber="427">P l a n t F o s s i l N a m e s R e g i s t r y N u m b e r.</paragraph>
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<paragraph blockId="2.[843,1479,278,334]" box="[843,979,310,334]" pageId="2" pageNumber="427">PFN003021.</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="427" type="etymology">
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<paragraph blockId="2.[843,1482,357,446]" pageId="2" pageNumber="427">E t y m o l o g y. The species name is derived from pater, meaning father in Latin. It is devoted to my father and teacher Professor Zlatko Kvaček.</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="427" type="description">
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<paragraph blockId="2.[843,1481,469,526]" pageId="2" pageNumber="427">
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T y p e l o c a l i t y. Pecínov quarry,
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<quantity box="[1306,1381,469,493]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="427" unit="km" value="50.0">50 km</quantity>
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west of
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<collectingRegion box="[843,920,502,526]" country="Czech Republic" name="Praha" pageId="2" pageNumber="427">Prague</collectingRegion>
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, the
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<collectingCountry box="[969,1143,501,525]" name="Czech Republic" pageId="2" pageNumber="427">Czech Republic</collectingCountry>
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.
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</paragraph>
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<paragraph blockId="2.[881,1399,549,573]" box="[881,1399,549,573]" pageId="2" pageNumber="427">T y p e h o r i z o n. Peruc-Korycany Formation.</paragraph>
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<paragraph blockId="2.[881,1283,596,620]" box="[881,1283,596,620]" pageId="2" pageNumber="427">A g e. Cenomanian, Late Cretaceous.</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="427" type="diagnosis">
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<paragraph blockId="2.[843,1483,643,1182]" pageId="2" pageNumber="427">D i a g n o s i s. Leaf simply pinnate, pinnae linear to narrowly lanceolate, entire-margined. Base of pinna slightly contracted, apex unknown. Veins 2–3 per mm, simple or rarely dichotomising. Leaf hypostomatic. Adaxial cuticle very thin, bearing polygonal cells with tightly sinuous anticlinal walls. Mesophyll present, consisting of isodiametric cells. Abaxial cuticle thick, showing distinct marginal band bearing tetragonal ordinary cells. Inner part of leaf consists of costal ordinary cells forming zones typically narrower than intercostal zones, formed by 4–5 rows of syndetocheilic stomata (sometimes ill-defined). Subsidiary cells large, ellipsoidal, thickly cutinised, bearing one or several papillae. Ordinary cells of intercostal zone polygonal in shape with roughly straight anticlinal walls. Ordinary cells of costal zones quadrangular in shape with sinuous anticlinal walls. Epidermal cells of both costal and intercostal zones bearing 3–5 papillae.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="429" pageId="2" pageNumber="427" type="description">
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<paragraph blockId="2.[843,1482,1205,1906]" pageId="2" pageNumber="427">
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D e s c r i p t i o n.The
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<typeStatus box="[1118,1211,1205,1229]" pageId="2" pageNumber="427">holotype</typeStatus>
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corresponds to a fragment of a simply pinnate leaf (
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<figureCitation box="[1117,1245,1238,1262]" captionStart="Text-fig" captionStartId="1.[125,206,1966,1987]" captionTargetBox="[125,1440,151,1950]" captionTargetId="figure-4@1.[-161,1348,77,1084]" captionTargetPageId="1" captionText="Text-fig. 1. Zamites pateri J.KVAČEK sp. nov.; Pecínov locality, holotype, No. NM-F 5185. a: Holotype overview showing fragment of simply pinnate leaf, scale bar 20 mm. b: Abaxial cuticle showing costal and intercostal scale zones and stomata in ill-defined rows, LM micro-photograph, scale bar 100 µm. c: Pinnule detail showing venation pattern, scale bar 5 mm. d: Abaxial cuticle showing costal and intercostal zones, LM micro-photograph, scale bar 500 µm. e: Fragmentary preserved adaxial cuticle showing" figureDoi="http://doi.org/10.5281/zenodo.7522684" httpUri="https://zenodo.org/record/7522684/files/figure.png" pageId="2" pageNumber="427">Text-fig. 1a</figureCitation>
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). Leaves are attached to the upper part of the robust, longitudinally striated rachis,
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<quantity box="[843,935,1302,1326]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="2" pageNumber="427" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">7–9 mm</quantity>
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broad. The
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<typeStatus box="[1076,1169,1302,1326]" pageId="2" pageNumber="427">holotype</typeStatus>
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shows 5 incomplete pinnae arising from a common rachis. The pinnae are attached to the dorsal part of the rachis, where they seem to touch each other. Each pinna is linear to narrowly lanceolate, gradually widening from a contracted base to about
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<quantity box="[1285,1358,1431,1455]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="427" unit="mm" value="10.0">10 mm</quantity>
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wide in the parallel-sided middle region (
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<figureCitation box="[1166,1295,1463,1487]" captionStart="Text-fig" captionStartId="1.[125,206,1966,1987]" captionTargetBox="[125,1440,151,1950]" captionTargetId="figure-4@1.[-161,1348,77,1084]" captionTargetPageId="1" captionText="Text-fig. 1. Zamites pateri J.KVAČEK sp. nov.; Pecínov locality, holotype, No. NM-F 5185. a: Holotype overview showing fragment of simply pinnate leaf, scale bar 20 mm. b: Abaxial cuticle showing costal and intercostal scale zones and stomata in ill-defined rows, LM micro-photograph, scale bar 100 µm. c: Pinnule detail showing venation pattern, scale bar 5 mm. d: Abaxial cuticle showing costal and intercostal zones, LM micro-photograph, scale bar 500 µm. e: Fragmentary preserved adaxial cuticle showing" figureDoi="http://doi.org/10.5281/zenodo.7522684" httpUri="https://zenodo.org/record/7522684/files/figure.png" pageId="2" pageNumber="427">Text-fig. 1a</figureCitation>
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). From the base, parallel, rarely dichotomising veins arise in density 2–3 per
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<quantity box="[843,904,1527,1551]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="427" unit="mm" value="1.0">1 mm</quantity>
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(
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<figureCitation box="[918,1043,1527,1551]" captionStart="Text-fig" captionStartId="1.[125,206,1966,1987]" captionTargetBox="[125,1440,151,1950]" captionTargetId="figure-4@1.[-161,1348,77,1084]" captionTargetPageId="1" captionText="Text-fig. 1. Zamites pateri J.KVAČEK sp. nov.; Pecínov locality, holotype, No. NM-F 5185. a: Holotype overview showing fragment of simply pinnate leaf, scale bar 20 mm. b: Abaxial cuticle showing costal and intercostal scale zones and stomata in ill-defined rows, LM micro-photograph, scale bar 100 µm. c: Pinnule detail showing venation pattern, scale bar 5 mm. d: Abaxial cuticle showing costal and intercostal zones, LM micro-photograph, scale bar 500 µm. e: Fragmentary preserved adaxial cuticle showing" figureDoi="http://doi.org/10.5281/zenodo.7522684" httpUri="https://zenodo.org/record/7522684/files/figure.png" pageId="2" pageNumber="427">Text-fig. 1c</figureCitation>
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).
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</paragraph>
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<paragraph blockId="2.[843,1482,1205,1906]" lastBlockId="4.[129,769,1308,1660]" lastPageId="4" lastPageNumber="429" pageId="2" pageNumber="427">
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The adaxial cuticle is very thin and thus difficult to prepare. It consists of polygonal ordinary cells (25–30 × 30–55 µm). Their anticlinal walls are thin and tightly sinuous (
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<figureCitation box="[850,973,1656,1680]" captionStart="Text-fig" captionStartId="1.[125,206,1966,1987]" captionTargetBox="[125,1440,151,1950]" captionTargetId="figure-4@1.[-161,1348,77,1084]" captionTargetPageId="1" captionText="Text-fig. 1. Zamites pateri J.KVAČEK sp. nov.; Pecínov locality, holotype, No. NM-F 5185. a: Holotype overview showing fragment of simply pinnate leaf, scale bar 20 mm. b: Abaxial cuticle showing costal and intercostal scale zones and stomata in ill-defined rows, LM micro-photograph, scale bar 100 µm. c: Pinnule detail showing venation pattern, scale bar 5 mm. d: Abaxial cuticle showing costal and intercostal zones, LM micro-photograph, scale bar 500 µm. e: Fragmentary preserved adaxial cuticle showing" figureDoi="http://doi.org/10.5281/zenodo.7522684" httpUri="https://zenodo.org/record/7522684/files/figure.png" pageId="2" pageNumber="427">Text-fig. 1e</figureCitation>
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). It is devoid of papillae. The tissue between the adaxial and abaxial cuticle is probably mesophyll, consisting of isodiametric cells (30–40 × 40–50 µm;
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<figureCitation box="[1341,1468,1720,1744]" captionStart="Text-fig" captionStartId="1.[125,206,1966,1987]" captionTargetBox="[125,1440,151,1950]" captionTargetId="figure-4@1.[-161,1348,77,1084]" captionTargetPageId="1" captionText="Text-fig. 1. Zamites pateri J.KVAČEK sp. nov.; Pecínov locality, holotype, No. NM-F 5185. a: Holotype overview showing fragment of simply pinnate leaf, scale bar 20 mm. b: Abaxial cuticle showing costal and intercostal scale zones and stomata in ill-defined rows, LM micro-photograph, scale bar 100 µm. c: Pinnule detail showing venation pattern, scale bar 5 mm. d: Abaxial cuticle showing costal and intercostal zones, LM micro-photograph, scale bar 500 µm. e: Fragmentary preserved adaxial cuticle showing" figureDoi="http://doi.org/10.5281/zenodo.7522684" httpUri="https://zenodo.org/record/7522684/files/figure.png" pageId="2" pageNumber="427">Text-fig. 1f</figureCitation>
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). The abaxial cuticle shows costal and intercostal zones. The costal zones are narrower than the intercostal ones, typically 75–90 µm wide (
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<figureCitation box="[1031,1190,1817,1841]" captionStart="Text-fig" captionStartId="1.[125,206,1966,1987]" captionTargetBox="[125,1440,151,1950]" captionTargetId="figure-4@1.[-161,1348,77,1084]" captionTargetPageId="1" captionText="Text-fig. 1. Zamites pateri J.KVAČEK sp. nov.; Pecínov locality, holotype, No. NM-F 5185. a: Holotype overview showing fragment of simply pinnate leaf, scale bar 20 mm. b: Abaxial cuticle showing costal and intercostal scale zones and stomata in ill-defined rows, LM micro-photograph, scale bar 100 µm. c: Pinnule detail showing venation pattern, scale bar 5 mm. d: Abaxial cuticle showing costal and intercostal zones, LM micro-photograph, scale bar 500 µm. e: Fragmentary preserved adaxial cuticle showing" figureDoi="http://doi.org/10.5281/zenodo.7522684" httpUri="https://zenodo.org/record/7522684/files/figure.png" pageId="2" pageNumber="427">Text-fig. 1b, d</figureCitation>
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). They consist of basically quadrangular ordinary epidermal cells (25–38 × 50–125 µm) with sinuous anticlinal walls (
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<figureCitation box="[1146,1290,1881,1906]" captionStart="Text-fig" captionStartId="3.[125,206,1879,1900]" captionTargetBox="[125,1440,136,1861]" captionTargetPageId="3" captionText="Text-fig. 2. Zamites pateri J.KVAČEK sp. nov.; Pecínov locality, holotype, No. NM-F 5185. a: Abaxial cuticle showing costal and intercostal zones, SEM micro-photograph, scale bar 100 µm. b: Abaxial cuticle, detail of syndetocheilic stoma, SEM microphotograph, scale bar 10 µm. c: Abaxial cuticle showing transversely oriented stomata, SEM micro-photograph, scale bar 50 µm. d: Abaxial cuticle, detail of syndetocheilic stoma showing ledges of guard cells, SEM micro-photograph, scale bar 10 µm. e: Abaxial cuticle showing costal ordinary cells seen from inside, SEM micro-photograph, scale bar 50 µm. f: External side of abaxial cuticle showing stoma sunken in a stomatal pit surrounded by papillae, SEM micro-photograph, scale bar 10 µm. g: External side of abaxial cuticle showing papillae, SEM micro-photograph, scale bar 100 µm. h: External side of abaxial cuticle showing detail of fused papillae, SEM micro-photograph, scale bar 10 µm." figureDoi="http://doi.org/10.5281/zenodo.7522688" httpUri="https://zenodo.org/record/7522688/files/figure.png" pageId="2" pageNumber="427">Text-fig. 2a, e</figureCitation>
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). Intercostal zones are 220–300 µm wide with syndetocheilic stomata (Textfig. 2b–d). Stomata are oriented perpendicularly to the leaf margin, typically arranged in 3–4 ill-defined rows (
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<figureCitation captionStart="Text-fig" captionStartId="1.[125,206,1966,1987]" captionTargetBox="[125,1440,151,1950]" captionTargetId="figure-4@1.[-161,1348,77,1084]" captionTargetPageId="1" captionText="Text-fig. 1. Zamites pateri J.KVAČEK sp. nov.; Pecínov locality, holotype, No. NM-F 5185. a: Holotype overview showing fragment of simply pinnate leaf, scale bar 20 mm. b: Abaxial cuticle showing costal and intercostal scale zones and stomata in ill-defined rows, LM micro-photograph, scale bar 100 µm. c: Pinnule detail showing venation pattern, scale bar 5 mm. d: Abaxial cuticle showing costal and intercostal zones, LM micro-photograph, scale bar 500 µm. e: Fragmentary preserved adaxial cuticle showing" figureDoi="http://doi.org/10.5281/zenodo.7522684" httpUri="https://zenodo.org/record/7522684/files/figure.png" pageId="4" pageNumber="429">Text-figs 1b</figureCitation>
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,
|
||
<figureCitation box="[167,194,1406,1430]" captionStart="Text-fig" captionStartId="3.[125,206,1879,1900]" captionTargetBox="[125,1440,136,1861]" captionTargetPageId="3" captionText="Text-fig. 2. Zamites pateri J.KVAČEK sp. nov.; Pecínov locality, holotype, No. NM-F 5185. a: Abaxial cuticle showing costal and intercostal zones, SEM micro-photograph, scale bar 100 µm. b: Abaxial cuticle, detail of syndetocheilic stoma, SEM microphotograph, scale bar 10 µm. c: Abaxial cuticle showing transversely oriented stomata, SEM micro-photograph, scale bar 50 µm. d: Abaxial cuticle, detail of syndetocheilic stoma showing ledges of guard cells, SEM micro-photograph, scale bar 10 µm. e: Abaxial cuticle showing costal ordinary cells seen from inside, SEM micro-photograph, scale bar 50 µm. f: External side of abaxial cuticle showing stoma sunken in a stomatal pit surrounded by papillae, SEM micro-photograph, scale bar 10 µm. g: External side of abaxial cuticle showing papillae, SEM micro-photograph, scale bar 100 µm. h: External side of abaxial cuticle showing detail of fused papillae, SEM micro-photograph, scale bar 10 µm." figureDoi="http://doi.org/10.5281/zenodo.7522688" httpUri="https://zenodo.org/record/7522688/files/figure.png" pageId="4" pageNumber="429">2a</figureCitation>
|
||
). Externally, they are sunken in pits, surrounded by 2–5 papillae (
|
||
<figureCitation box="[224,347,1439,1463]" captionStart="Text-fig" captionStartId="3.[125,206,1879,1900]" captionTargetBox="[125,1440,136,1861]" captionTargetPageId="3" captionText="Text-fig. 2. Zamites pateri J.KVAČEK sp. nov.; Pecínov locality, holotype, No. NM-F 5185. a: Abaxial cuticle showing costal and intercostal zones, SEM micro-photograph, scale bar 100 µm. b: Abaxial cuticle, detail of syndetocheilic stoma, SEM microphotograph, scale bar 10 µm. c: Abaxial cuticle showing transversely oriented stomata, SEM micro-photograph, scale bar 50 µm. d: Abaxial cuticle, detail of syndetocheilic stoma showing ledges of guard cells, SEM micro-photograph, scale bar 10 µm. e: Abaxial cuticle showing costal ordinary cells seen from inside, SEM micro-photograph, scale bar 50 µm. f: External side of abaxial cuticle showing stoma sunken in a stomatal pit surrounded by papillae, SEM micro-photograph, scale bar 10 µm. g: External side of abaxial cuticle showing papillae, SEM micro-photograph, scale bar 100 µm. h: External side of abaxial cuticle showing detail of fused papillae, SEM micro-photograph, scale bar 10 µm." figureDoi="http://doi.org/10.5281/zenodo.7522688" httpUri="https://zenodo.org/record/7522688/files/figure.png" pageId="4" pageNumber="429">Text-fig. 2f</figureCitation>
|
||
). Their subsidiary cells are 27–35 × 55– 66 µm. Ordinary epidermal cells are isodiametric to elongate (15–25 × 28–40 µm), with straight anticlinal walls (
|
||
<figureCitation captionStart="Text-fig" captionStartId="3.[125,206,1879,1900]" captionTargetBox="[125,1440,136,1861]" captionTargetPageId="3" captionText="Text-fig. 2. Zamites pateri J.KVAČEK sp. nov.; Pecínov locality, holotype, No. NM-F 5185. a: Abaxial cuticle showing costal and intercostal zones, SEM micro-photograph, scale bar 100 µm. b: Abaxial cuticle, detail of syndetocheilic stoma, SEM microphotograph, scale bar 10 µm. c: Abaxial cuticle showing transversely oriented stomata, SEM micro-photograph, scale bar 50 µm. d: Abaxial cuticle, detail of syndetocheilic stoma showing ledges of guard cells, SEM micro-photograph, scale bar 10 µm. e: Abaxial cuticle showing costal ordinary cells seen from inside, SEM micro-photograph, scale bar 50 µm. f: External side of abaxial cuticle showing stoma sunken in a stomatal pit surrounded by papillae, SEM micro-photograph, scale bar 10 µm. g: External side of abaxial cuticle showing papillae, SEM micro-photograph, scale bar 100 µm. h: External side of abaxial cuticle showing detail of fused papillae, SEM micro-photograph, scale bar 10 µm." figureDoi="http://doi.org/10.5281/zenodo.7522688" httpUri="https://zenodo.org/record/7522688/files/figure.png" pageId="4" pageNumber="429">Text-fig. 2c</figureCitation>
|
||
). The external side of the cuticle shows numerous papillae, typically 4–5 per ordinary epidermal cell (
|
||
<figureCitation box="[599,755,1570,1594]" captionStart="Text-fig" captionStartId="3.[125,206,1879,1900]" captionTargetBox="[125,1440,136,1861]" captionTargetPageId="3" captionText="Text-fig. 2. Zamites pateri J.KVAČEK sp. nov.; Pecínov locality, holotype, No. NM-F 5185. a: Abaxial cuticle showing costal and intercostal zones, SEM micro-photograph, scale bar 100 µm. b: Abaxial cuticle, detail of syndetocheilic stoma, SEM microphotograph, scale bar 10 µm. c: Abaxial cuticle showing transversely oriented stomata, SEM micro-photograph, scale bar 50 µm. d: Abaxial cuticle, detail of syndetocheilic stoma showing ledges of guard cells, SEM micro-photograph, scale bar 10 µm. e: Abaxial cuticle showing costal ordinary cells seen from inside, SEM micro-photograph, scale bar 50 µm. f: External side of abaxial cuticle showing stoma sunken in a stomatal pit surrounded by papillae, SEM micro-photograph, scale bar 10 µm. g: External side of abaxial cuticle showing papillae, SEM micro-photograph, scale bar 100 µm. h: External side of abaxial cuticle showing detail of fused papillae, SEM micro-photograph, scale bar 10 µm." figureDoi="http://doi.org/10.5281/zenodo.7522688" httpUri="https://zenodo.org/record/7522688/files/figure.png" pageId="4" pageNumber="429">Text-fig. 2i, h</figureCitation>
|
||
). They are so dense that they are sometimes fused, forming united groups (
|
||
<figureCitation box="[284,410,1635,1659]" captionStart="Text-fig" captionStartId="3.[125,206,1879,1900]" captionTargetBox="[125,1440,136,1861]" captionTargetPageId="3" captionText="Text-fig. 2. Zamites pateri J.KVAČEK sp. nov.; Pecínov locality, holotype, No. NM-F 5185. a: Abaxial cuticle showing costal and intercostal zones, SEM micro-photograph, scale bar 100 µm. b: Abaxial cuticle, detail of syndetocheilic stoma, SEM microphotograph, scale bar 10 µm. c: Abaxial cuticle showing transversely oriented stomata, SEM micro-photograph, scale bar 50 µm. d: Abaxial cuticle, detail of syndetocheilic stoma showing ledges of guard cells, SEM micro-photograph, scale bar 10 µm. e: Abaxial cuticle showing costal ordinary cells seen from inside, SEM micro-photograph, scale bar 50 µm. f: External side of abaxial cuticle showing stoma sunken in a stomatal pit surrounded by papillae, SEM micro-photograph, scale bar 10 µm. g: External side of abaxial cuticle showing papillae, SEM micro-photograph, scale bar 100 µm. h: External side of abaxial cuticle showing detail of fused papillae, SEM micro-photograph, scale bar 10 µm." figureDoi="http://doi.org/10.5281/zenodo.7522688" httpUri="https://zenodo.org/record/7522688/files/figure.png" pageId="4" pageNumber="429">Text-fig. 2h</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<paragraph blockId="2.[147,1463,1995,2075]" pageId="2" pageNumber="427">
|
||
<emphasis bold="true" pageId="2" pageNumber="427">two better-preserved ordinary cells, LM micro-photograph, scale bar 50 µm. f: Abaxial cuticle showing perpendicularly arranged stomata and papillae per ordinary cell, LM micrography, scale bar 50 µm. g: Detail of mesophyll tissue consisting of isodiametric cells, LM micrography, scale bar 50 µm.</emphasis>
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7522688" ID-Zenodo-Dep="7522688" httpUri="https://zenodo.org/record/7522688/files/figure.png" pageId="3" pageNumber="428" startId="3.[125,206,1879,1900]" targetBox="[125,1440,136,1861]" targetPageId="3">
|
||
<paragraph blockId="3.[125,1441,1879,2105]" pageId="3" pageNumber="428">
|
||
<emphasis bold="true" box="[125,492,1879,1900]" pageId="3" pageNumber="428">
|
||
Text-fig. 2.
|
||
<taxonomicName authority="J.KVACEK" authorityName="J. KVACEK" authorityYear="2022" box="[244,492,1879,1900]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="3" pageNumber="428" phylum="Tracheophyta" rank="species" species="pateri" status="sp. nov.">
|
||
<emphasis bold="true" box="[244,388,1879,1900]" italics="true" pageId="3" pageNumber="428">Zamites pateri</emphasis>
|
||
J.
|
||
<subScript attach="left" box="[414,492,1879,1900]" fontSize="6" pageId="3" pageNumber="428">KVAČEK</subScript>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel box="[501,583,1879,1900]" pageId="3" pageNumber="428" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; Pecínov locality, holotype, No. NM-F 5185. a: Abaxial cuticle showing costal and intercostal zones, SEM micro-photograph, scale bar 100 µm. b: Abaxial cuticle, detail of syndetocheilic stoma, SEM microphotograph, scale bar 10 µm. c: Abaxial cuticle showing transversely oriented stomata, SEM micro-photograph, scale bar 50 µm. d: Abaxial cuticle, detail of syndetocheilic stoma showing ledges of guard cells, SEM micro-photograph, scale bar 10 µm. e: Abaxial cuticle showing costal ordinary cells seen from inside, SEM micro-photograph, scale bar 50 µm. f: External side of abaxial cuticle showing stoma sunken in a stomatal pit surrounded by papillae, SEM micro-photograph, scale bar 10 µm. g: External side of abaxial cuticle showing papillae, SEM micro-photograph, scale bar 100 µm. h: External side of abaxial cuticle showing detail of fused papillae, SEM micro-photograph, scale bar 10 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption box="[127,727,152,173]" pageId="4" pageNumber="429" startId="4.[127,183,152,173]" targetBox="[140,1464,220,1225]" targetIsTable="true" targetPageId="4">
|
||
<paragraph blockId="4.[127,727,152,173]" box="[127,727,152,173]" pageId="4" pageNumber="429">
|
||
Table 1. Comparison of
|
||
<taxonomicName box="[372,514,152,173]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="429" phylum="Tracheophyta" rank="species" species="pateri">
|
||
<emphasis bold="true" box="[372,514,152,173]" italics="true" pageId="4" pageNumber="429">Zamites pateri</emphasis>
|
||
</taxonomicName>
|
||
with related species.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="4" pageNumber="429">
|
||
<table box="[140,1464,220,1225]" gridcols="8" gridrows="12" pageId="4" pageNumber="429">
|
||
<tr box="[140,1464,220,316]" gridrow="0" pageId="4" pageNumber="429">
|
||
<th box="[140,304,220,316]" gridcol="0" gridrow="0" pageId="4" pageNumber="429">
|
||
<emphasis bold="true" box="[193,256,259,278]" pageId="4" pageNumber="429">species</emphasis>
|
||
</th>
|
||
<th box="[329,473,220,316]" gridcol="1" gridrow="0" pageId="4" pageNumber="429">
|
||
<emphasis bold="true" box="[331,473,259,278]" pageId="4" pageNumber="429">shape of pinnae</emphasis>
|
||
</th>
|
||
<th box="[496,640,220,316]" gridcol="2" gridrow="0" pageId="4" pageNumber="429">
|
||
<emphasis bold="true" box="[518,619,246,291]" pageId="4" pageNumber="429">orientation of stomata</emphasis>
|
||
</th>
|
||
<th box="[662,807,220,316]" gridcol="3" gridrow="0" pageId="4" pageNumber="429">
|
||
<emphasis bold="true" pageId="4" pageNumber="429">anticlinal walls of abaxial cuticle</emphasis>
|
||
</th>
|
||
<th box="[829,974,220,316]" gridcol="4" gridrow="0" pageId="4" pageNumber="429">
|
||
<emphasis bold="true" pageId="4" pageNumber="429">number of papillae per ordinary cell on abaxial cuticle</emphasis>
|
||
</th>
|
||
<th box="[995,1134,220,316]" gridcol="5" gridrow="0" pageId="4" pageNumber="429">
|
||
<emphasis bold="true" pageId="4" pageNumber="429">cells of adaxial cuticle</emphasis>
|
||
</th>
|
||
<th box="[1180,1288,220,316]" gridcol="6" gridrow="0" pageId="4" pageNumber="429">
|
||
<emphasis bold="true" box="[1180,1288,259,278]" pageId="4" pageNumber="429">hypodermis</emphasis>
|
||
</th>
|
||
<th box="[1327,1464,220,316]" gridcol="7" gridrow="0" pageId="4" pageNumber="429">
|
||
<emphasis bold="true" box="[1370,1429,259,278]" pageId="4" pageNumber="429">source</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr box="[140,1464,333,404]" gridrow="1" pageId="4" pageNumber="429">
|
||
<th box="[140,304,333,404]" gridcol="0" gridrow="1" pageId="4" pageNumber="429">
|
||
<taxonomicName box="[140,267,359,378]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="429" phylum="Tracheophyta" rank="species" species="pateri">
|
||
<emphasis bold="true" box="[140,267,359,378]" italics="true" pageId="4" pageNumber="429">Zamites pateri</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td box="[329,473,333,404]" gridcol="1" gridrow="1" pageId="4" pageNumber="429">linear to narrowly lanceolate</td>
|
||
<td box="[496,640,333,404]" gridcol="2" gridrow="1" pageId="4" pageNumber="429">perpendicular in all cases</td>
|
||
<td box="[662,807,333,404]" gridcol="3" gridrow="1" pageId="4" pageNumber="429">sinuous in all cases</td>
|
||
<td box="[829,974,333,404]" gridcol="4" gridrow="1" pageId="4" pageNumber="429">3–5</td>
|
||
<td box="[995,1134,333,404]" gridcol="5" gridrow="1" pageId="4" pageNumber="429">weakly defined, nearly invisible</td>
|
||
<td box="[1180,1288,333,404]" gridcol="6" gridrow="1" pageId="4" pageNumber="429">present</td>
|
||
<td box="[1327,1464,333,404]" gridcol="7" gridrow="1" pageId="4" pageNumber="429">this paper</td>
|
||
</tr>
|
||
<tr box="[140,1464,421,516]" gridrow="2" pageId="4" pageNumber="429">
|
||
<th box="[140,304,421,516]" gridcol="0" gridrow="2" pageId="4" pageNumber="429">
|
||
<emphasis bold="true" box="[140,270,459,478]" italics="true" pageId="4" pageNumber="429">
|
||
<taxonomicName box="[140,210,459,478]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="429" phylum="Tracheophyta" rank="genus">Zamites</taxonomicName>
|
||
bayeri
|
||
</emphasis>
|
||
</th>
|
||
<td box="[329,473,421,516]" gridcol="1" gridrow="2" pageId="4" pageNumber="429">linear to narrowly lanceolate</td>
|
||
<td box="[496,640,421,516]" gridcol="2" gridrow="2" pageId="4" pageNumber="429">in inner part perpendicular, in marginal parts longitudinal</td>
|
||
<td box="[662,807,421,516]" gridcol="3" gridrow="2" pageId="4" pageNumber="429">straight in intercostal areas</td>
|
||
<td box="[829,974,421,516]" gridcol="4" gridrow="2" pageId="4" pageNumber="429">one to several</td>
|
||
<td box="[995,1134,421,516]" gridcol="5" gridrow="2" pageId="4" pageNumber="429">ordinary cells well-discernible</td>
|
||
<td box="[1180,1288,421,516]" gridcol="6" gridrow="2" pageId="4" pageNumber="429">absent</td>
|
||
<td box="[1327,1464,421,516]" gridcol="7" gridrow="2" pageId="4" pageNumber="429">
|
||
<bibRefCitation author="Knobloch, E. & Kvacek, J." pageId="4" pageNumber="429" pagination="567 - 586" refId="ref3964" refString="Knobloch, E., Kvacek, J. (1997): Bennettitalean and coni- feralean (?) leaves from the Cretaceous (Cenomanian) of the Bohemian Massif, Central Europe. - Cretaceous Research, 18: 567 - 586. https: // doi. org / 10.1006 / cres. 1997.0074" type="journal article" year="1997">Knobloch and Kvaček 1997</bibRefCitation>
|
||
</td>
|
||
</tr>
|
||
<tr box="[140,1464,533,604]" gridrow="3" pageId="4" pageNumber="429">
|
||
<th box="[140,304,533,604]" gridcol="0" gridrow="3" pageId="4" pageNumber="429">
|
||
<emphasis bold="true" box="[140,244,546,591]" italics="true" pageId="4" pageNumber="429">
|
||
<taxonomicName box="[140,210,546,565]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="429" phylum="Tracheophyta" rank="genus">Zamites</taxonomicName>
|
||
carruthersii
|
||
</emphasis>
|
||
</th>
|
||
<td box="[329,473,533,604]" gridcol="1" gridrow="3" pageId="4" pageNumber="429">broadly oval to oblong</td>
|
||
<td box="[496,640,533,604]" gridcol="2" gridrow="3" pageId="4" pageNumber="429">perpendicular in all cases</td>
|
||
<td box="[662,807,533,604]" gridcol="3" gridrow="3" pageId="4" pageNumber="429">sinuous, in some cases straight in intercostal areas</td>
|
||
<td box="[829,974,533,604]" gridcol="4" gridrow="3" pageId="4" pageNumber="429">one to several</td>
|
||
<td box="[995,1134,533,604]" gridcol="5" gridrow="3" pageId="4" pageNumber="429">ordinary cells well-discernible</td>
|
||
<td box="[1180,1288,533,604]" gridcol="6" gridrow="3" pageId="4" pageNumber="429">present</td>
|
||
<td box="[1327,1464,533,604]" gridcol="7" gridrow="3" pageId="4" pageNumber="429">
|
||
<bibRefCitation author="Watson, J. & Sincock, C. A." pageId="4" pageNumber="429" pagination="1 - 228" refId="ref4997" refString="Watson, J., Sincock, C. A. (1992): Bennettitales of the English Wealden. - Monographs, Palaeontographical Society, London, 145 (588): 1 - 228, 23 pls. https: // doi. org / 10.1080 / 25761900.2022.12131771" type="journal article" year="1992">Watson and Sincock 1992</bibRefCitation>
|
||
</td>
|
||
</tr>
|
||
<tr box="[140,1464,621,665]" gridrow="4" pageId="4" pageNumber="429">
|
||
<th box="[140,304,621,665]" gridcol="0" gridrow="4" pageId="4" pageNumber="429">
|
||
<emphasis bold="true" box="[140,304,633,652]" italics="true" pageId="4" pageNumber="429">
|
||
<taxonomicName box="[140,210,633,652]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="429" phylum="Tracheophyta" rank="genus">Zamites</taxonomicName>
|
||
manoniae
|
||
</emphasis>
|
||
</th>
|
||
<td box="[329,473,621,665]" gridcol="1" gridrow="4" pageId="4" pageNumber="429">broadly oval to oblong</td>
|
||
<td box="[496,640,621,665]" gridcol="2" gridrow="4" pageId="4" pageNumber="429">perpendicular in all cases</td>
|
||
<td box="[662,807,621,665]" gridcol="3" gridrow="4" pageId="4" pageNumber="429">slightly and univenly sinuous</td>
|
||
<td box="[829,974,621,665]" gridcol="4" gridrow="4" pageId="4" pageNumber="429">0</td>
|
||
<td box="[995,1134,621,665]" gridcol="5" gridrow="4" pageId="4" pageNumber="429">unknown</td>
|
||
<td box="[1180,1288,621,665]" gridcol="6" gridrow="4" pageId="4" pageNumber="429">absent</td>
|
||
<td box="[1327,1464,621,665]" gridcol="7" gridrow="4" pageId="4" pageNumber="429">
|
||
<bibRefCitation author="Watson, J. & Sincock, C. A." pageId="4" pageNumber="429" pagination="1 - 228" refId="ref4997" refString="Watson, J., Sincock, C. A. (1992): Bennettitales of the English Wealden. - Monographs, Palaeontographical Society, London, 145 (588): 1 - 228, 23 pls. https: // doi. org / 10.1080 / 25761900.2022.12131771" type="journal article" year="1992">Watson and Sincock 1992</bibRefCitation>
|
||
</td>
|
||
</tr>
|
||
<tr box="[140,1464,682,752]" gridrow="5" pageId="4" pageNumber="429">
|
||
<th box="[140,304,682,752]" gridcol="0" gridrow="5" pageId="4" pageNumber="429">
|
||
<emphasis bold="true" box="[140,285,708,727]" italics="true" pageId="4" pageNumber="429">
|
||
<taxonomicName box="[140,210,708,727]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="429" phylum="Tracheophyta" rank="genus">Zamites</taxonomicName>
|
||
dowellii
|
||
</emphasis>
|
||
</th>
|
||
<td box="[329,473,682,752]" gridcol="1" gridrow="5" pageId="4" pageNumber="429">narrowly lanceolate</td>
|
||
<td box="[496,640,682,752]" gridcol="2" gridrow="5" pageId="4" pageNumber="429">perpendicular in all cases</td>
|
||
<td box="[662,807,682,752]" gridcol="3" gridrow="5" pageId="4" pageNumber="429">usually sinuous, occasionally straight</td>
|
||
<td box="[829,974,682,752]" gridcol="4" gridrow="5" pageId="4" pageNumber="429">1–2</td>
|
||
<td box="[995,1134,682,752]" gridcol="5" gridrow="5" pageId="4" pageNumber="429">ordinary cells well-discernible</td>
|
||
<td box="[1180,1288,682,752]" gridcol="6" gridrow="5" pageId="4" pageNumber="429">absent</td>
|
||
<td box="[1327,1464,682,752]" gridcol="7" gridrow="5" pageId="4" pageNumber="429">
|
||
<bibRefCitation author="Watson, J. & Sincock, C. A." pageId="4" pageNumber="429" pagination="1 - 228" refId="ref4997" refString="Watson, J., Sincock, C. A. (1992): Bennettitales of the English Wealden. - Monographs, Palaeontographical Society, London, 145 (588): 1 - 228, 23 pls. https: // doi. org / 10.1080 / 25761900.2022.12131771" type="journal article" year="1992">Watson and Sincock 1992</bibRefCitation>
|
||
</td>
|
||
</tr>
|
||
<tr box="[140,1464,769,840]" gridrow="6" pageId="4" pageNumber="429">
|
||
<th box="[140,304,769,840]" gridcol="0" gridrow="6" pageId="4" pageNumber="429">
|
||
<emphasis bold="true" box="[140,251,782,827]" italics="true" pageId="4" pageNumber="429">
|
||
<taxonomicName box="[140,210,782,801]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="429" phylum="Tracheophyta" rank="genus">Zamites</taxonomicName>
|
||
wendyellisae
|
||
</emphasis>
|
||
</th>
|
||
<td box="[329,473,769,840]" gridcol="1" gridrow="6" pageId="4" pageNumber="429">triangular, narowly lanceolate</td>
|
||
<td box="[496,640,769,840]" gridcol="2" gridrow="6" pageId="4" pageNumber="429">perpendicular or slightly oblique</td>
|
||
<td box="[662,807,769,840]" gridcol="3" gridrow="6" pageId="4" pageNumber="429">strongly sinuous in all cases</td>
|
||
<td box="[829,974,769,840]" gridcol="4" gridrow="6" pageId="4" pageNumber="429">0</td>
|
||
<td box="[995,1134,769,840]" gridcol="5" gridrow="6" pageId="4" pageNumber="429">ordinary cells well-discernible</td>
|
||
<td box="[1180,1288,769,840]" gridcol="6" gridrow="6" pageId="4" pageNumber="429">present</td>
|
||
<td box="[1327,1464,769,840]" gridcol="7" gridrow="6" pageId="4" pageNumber="429">
|
||
<bibRefCitation author="Watson, J. & Sincock, C. A." pageId="4" pageNumber="429" pagination="1 - 228" refId="ref4997" refString="Watson, J., Sincock, C. A. (1992): Bennettitales of the English Wealden. - Monographs, Palaeontographical Society, London, 145 (588): 1 - 228, 23 pls. https: // doi. org / 10.1080 / 25761900.2022.12131771" type="journal article" year="1992">Watson and Sincock 1992</bibRefCitation>
|
||
</td>
|
||
</tr>
|
||
<tr box="[140,1464,857,927]" gridrow="7" pageId="4" pageNumber="429">
|
||
<th box="[140,304,857,927]" gridcol="0" gridrow="7" pageId="4" pageNumber="429">
|
||
<emphasis bold="true" box="[140,270,882,901]" italics="true" pageId="4" pageNumber="429">
|
||
<taxonomicName box="[140,210,882,901]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="429" phylum="Tracheophyta" rank="genus">Zamites</taxonomicName>
|
||
coderi
|
||
</emphasis>
|
||
</th>
|
||
<td box="[329,473,857,927]" gridcol="1" gridrow="7" pageId="4" pageNumber="429">linear to narrowly lanceolate</td>
|
||
<td box="[496,640,857,927]" gridcol="2" gridrow="7" pageId="4" pageNumber="429">perpendicular or slightly oblique</td>
|
||
<td box="[662,807,857,927]" gridcol="3" gridrow="7" pageId="4" pageNumber="429">inuous in all cases</td>
|
||
<td box="[829,974,857,927]" gridcol="4" gridrow="7" pageId="4" pageNumber="429">0</td>
|
||
<td box="[995,1134,857,927]" gridcol="5" gridrow="7" pageId="4" pageNumber="429">ordinary cells well-discernible</td>
|
||
<td box="[1180,1288,857,927]" gridcol="6" gridrow="7" pageId="4" pageNumber="429">absent</td>
|
||
<td box="[1327,1464,857,927]" gridcol="7" gridrow="7" pageId="4" pageNumber="429">
|
||
<bibRefCitation author="Watson, J. & Sincock, C. A." pageId="4" pageNumber="429" pagination="1 - 228" refId="ref4997" refString="Watson, J., Sincock, C. A. (1992): Bennettitales of the English Wealden. - Monographs, Palaeontographical Society, London, 145 (588): 1 - 228, 23 pls. https: // doi. org / 10.1080 / 25761900.2022.12131771" type="journal article" year="1992">Watson and Sincock 1992</bibRefCitation>
|
||
</td>
|
||
</tr>
|
||
<tr box="[140,1464,944,1014]" gridrow="8" pageId="4" pageNumber="429">
|
||
<th box="[140,304,944,1014]" gridcol="0" gridrow="8" pageId="4" pageNumber="429">
|
||
<emphasis bold="true" box="[140,280,969,988]" italics="true" pageId="4" pageNumber="429">
|
||
<taxonomicName box="[140,210,969,988]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="429" phylum="Tracheophyta" rank="genus">Zamites</taxonomicName>
|
||
nicolae
|
||
</emphasis>
|
||
</th>
|
||
<td box="[329,473,944,1014]" gridcol="1" gridrow="8" pageId="4" pageNumber="429">narrowly lanceolate</td>
|
||
<td box="[496,640,944,1014]" gridcol="2" gridrow="8" pageId="4" pageNumber="429">perpendicular in all cases</td>
|
||
<td box="[662,807,944,1014]" gridcol="3" gridrow="8" pageId="4" pageNumber="429">moderately sinuous in all cases</td>
|
||
<td box="[829,974,944,1014]" gridcol="4" gridrow="8" pageId="4" pageNumber="429">1</td>
|
||
<td box="[995,1134,944,1014]" gridcol="5" gridrow="8" pageId="4" pageNumber="429">weakly defined</td>
|
||
<td box="[1180,1288,944,1014]" gridcol="6" gridrow="8" pageId="4" pageNumber="429">absent</td>
|
||
<td box="[1327,1464,944,1014]" gridcol="7" gridrow="8" pageId="4" pageNumber="429">
|
||
<bibRefCitation author="Watson, J. & Sincock, C. A." pageId="4" pageNumber="429" pagination="1 - 228" refId="ref4997" refString="Watson, J., Sincock, C. A. (1992): Bennettitales of the English Wealden. - Monographs, Palaeontographical Society, London, 145 (588): 1 - 228, 23 pls. https: // doi. org / 10.1080 / 25761900.2022.12131771" type="journal article" year="1992">Watson and Sincock 1992</bibRefCitation>
|
||
</td>
|
||
</tr>
|
||
<tr box="[140,1464,1031,1101]" gridrow="9" pageId="4" pageNumber="429">
|
||
<th box="[140,304,1031,1101]" gridcol="0" gridrow="9" pageId="4" pageNumber="429">
|
||
<emphasis bold="true" box="[140,287,1056,1075]" italics="true" pageId="4" pageNumber="429">
|
||
<taxonomicName box="[140,210,1056,1075]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="429" phylum="Tracheophyta" rank="genus">Zamites</taxonomicName>
|
||
tatianae
|
||
</emphasis>
|
||
</th>
|
||
<td box="[329,473,1031,1101]" gridcol="1" gridrow="9" pageId="4" pageNumber="429">linear to narrowly lanceolate</td>
|
||
<td box="[496,640,1031,1101]" gridcol="2" gridrow="9" pageId="4" pageNumber="429">perpendicular in all cases</td>
|
||
<td box="[662,807,1031,1101]" gridcol="3" gridrow="9" pageId="4" pageNumber="429">sinuous in all cases</td>
|
||
<td box="[829,974,1031,1101]" gridcol="4" gridrow="9" pageId="4" pageNumber="429">0</td>
|
||
<td box="[995,1134,1031,1101]" gridcol="5" gridrow="9" pageId="4" pageNumber="429">ordinary cells well-discernible</td>
|
||
<td box="[1180,1288,1031,1101]" gridcol="6" gridrow="9" pageId="4" pageNumber="429">absent</td>
|
||
<td box="[1327,1464,1031,1101]" gridcol="7" gridrow="9" pageId="4" pageNumber="429">
|
||
<bibRefCitation author="Watson, J. & Sincock, C. A." pageId="4" pageNumber="429" pagination="1 - 228" refId="ref4997" refString="Watson, J., Sincock, C. A. (1992): Bennettitales of the English Wealden. - Monographs, Palaeontographical Society, London, 145 (588): 1 - 228, 23 pls. https: // doi. org / 10.1080 / 25761900.2022.12131771" type="journal article" year="1992">Watson and Sincock 1992</bibRefCitation>
|
||
</td>
|
||
</tr>
|
||
<tr box="[140,1464,1118,1163]" gridrow="10" pageId="4" pageNumber="429">
|
||
<th box="[140,304,1118,1163]" gridcol="0" gridrow="10" pageId="4" pageNumber="429">
|
||
<emphasis bold="true" box="[140,255,1118,1163]" italics="true" pageId="4" pageNumber="429">
|
||
<taxonomicName box="[140,210,1118,1137]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="429" phylum="Tracheophyta" rank="genus">Zamites</taxonomicName>
|
||
notokenensis
|
||
</emphasis>
|
||
</th>
|
||
<td box="[329,473,1118,1163]" gridcol="1" gridrow="10" pageId="4" pageNumber="429">oblong</td>
|
||
<td box="[496,640,1118,1163]" gridcol="2" gridrow="10" pageId="4" pageNumber="429">orientation unknown</td>
|
||
<td box="[662,807,1118,1163]" gridcol="3" gridrow="10" pageId="4" pageNumber="429">strongly sinuous in all cases</td>
|
||
<td box="[829,974,1118,1163]" gridcol="4" gridrow="10" pageId="4" pageNumber="429">0</td>
|
||
<td box="[995,1134,1118,1163]" gridcol="5" gridrow="10" pageId="4" pageNumber="429">weakly defined, but discernible</td>
|
||
<td box="[1180,1288,1118,1163]" gridcol="6" gridrow="10" pageId="4" pageNumber="429">absent</td>
|
||
<td box="[1327,1464,1118,1163]" gridcol="7" gridrow="10" pageId="4" pageNumber="429">
|
||
<bibRefCitation author="Watson, J. & Sincock, C. A." pageId="4" pageNumber="429" pagination="1 - 228" refId="ref4997" refString="Watson, J., Sincock, C. A. (1992): Bennettitales of the English Wealden. - Monographs, Palaeontographical Society, London, 145 (588): 1 - 228, 23 pls. https: // doi. org / 10.1080 / 25761900.2022.12131771" type="journal article" year="1992">Watson and Sincock 1992</bibRefCitation>
|
||
</td>
|
||
</tr>
|
||
<tr box="[140,1464,1180,1225]" gridrow="11" pageId="4" pageNumber="429">
|
||
<th box="[140,304,1180,1225]" gridcol="0" gridrow="11" pageId="4" pageNumber="429">
|
||
<emphasis bold="true" box="[140,304,1193,1212]" italics="true" pageId="4" pageNumber="429">
|
||
<taxonomicName box="[140,210,1193,1212]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="429" phylum="Tracheophyta" rank="genus">Zamites</taxonomicName>
|
||
decurrens
|
||
</emphasis>
|
||
</th>
|
||
<td box="[329,473,1180,1225]" gridcol="1" gridrow="11" pageId="4" pageNumber="429">oblanceolate</td>
|
||
<td box="[496,640,1180,1225]" gridcol="2" gridrow="11" pageId="4" pageNumber="429">perpendicular in all cases</td>
|
||
<td box="[662,807,1180,1225]" gridcol="3" gridrow="11" pageId="4" pageNumber="429">straight in all cases</td>
|
||
<td box="[829,974,1180,1225]" gridcol="4" gridrow="11" pageId="4" pageNumber="429">1</td>
|
||
<td box="[995,1134,1180,1225]" gridcol="5" gridrow="11" pageId="4" pageNumber="429">ordinary cells well-discernible</td>
|
||
<td box="[1180,1288,1180,1225]" gridcol="6" gridrow="11" pageId="4" pageNumber="429">absent</td>
|
||
<td box="[1327,1464,1180,1225]" gridcol="7" gridrow="11" pageId="4" pageNumber="429">
|
||
<bibRefCitation author="Menendez, C. A." box="[1327,1464,1193,1212]" pageId="4" pageNumber="429" pagination="3 - 42" refId="ref4506" refString="Menendez, C. A. (1966): Fossil Bennettitales from the Tico flora, Santa Cruz Province, Argentina. - Bulletin of the British Museum (Natural History), 12 (1): 3 - 42. https: // doi. org / 10.5962 / p. 313885" type="journal article" year="1966">Menéndez 1966</bibRefCitation>
|
||
</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<subSubSection lastPageId="5" lastPageNumber="430" pageId="4" pageNumber="429" type="discussion">
|
||
<paragraph blockId="4.[128,768,1683,2100]" lastBlockId="4.[843,1483,1308,2100]" pageId="4" pageNumber="429">
|
||
D i s c u s s i o n.
|
||
<taxonomicName box="[348,501,1684,1707]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="429" phylum="Tracheophyta" rank="species" species="pateri">
|
||
<emphasis box="[348,501,1684,1707]" italics="true" pageId="4" pageNumber="429">Zamites pateri</emphasis>
|
||
</taxonomicName>
|
||
is reminiscent of
|
||
<emphasis italics="true" pageId="4" pageNumber="429">
|
||
<taxonomicName box="[683,767,1684,1707]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="429" phylum="Tracheophyta" rank="genus">Zamites</taxonomicName>
|
||
bayeri
|
||
</emphasis>
|
||
J.KVAČEK, from the same Peruc-Korycany Formation in
|
||
<collectingRegion box="[165,226,1748,1772]" country="Czech Republic" name="Praha" pageId="4" pageNumber="429">Praha</collectingRegion>
|
||
Malá Chuchle (
|
||
<bibRefCitation author="Knobloch, E. & Kvacek, J." box="[424,755,1748,1773]" pageId="4" pageNumber="429" pagination="567 - 586" refId="ref3964" refString="Knobloch, E., Kvacek, J. (1997): Bennettitalean and coni- feralean (?) leaves from the Cretaceous (Cenomanian) of the Bohemian Massif, Central Europe. - Cretaceous Research, 18: 567 - 586. https: // doi. org / 10.1006 / cres. 1997.0074" type="journal article" year="1997">Knobloch and Kvaček 1997</bibRefCitation>
|
||
). They are similar in morphology of the pinnae.
|
||
<taxonomicName class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="429" phylum="Tracheophyta" rank="species" species="pateri">
|
||
<emphasis italics="true" pageId="4" pageNumber="429">Zamites pateri</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<emphasis box="[355,458,1814,1838]" italics="true" pageId="4" pageNumber="429">Z. bayeri</emphasis>
|
||
in having pinnae attached to the dorsal part of the rachis. Further differences are in epidermal structures. Abaxial ordinary epidermal cells in the intercostal zones of the newly described species are nearly straight, bearing 3–5 papillae, whereas
|
||
<emphasis box="[583,687,1945,1969]" italics="true" pageId="4" pageNumber="429">Z. bayeri</emphasis>
|
||
shows ordinary cells more sinuously waved, bearing typically one, rarely several papillae per ordinary epidermal cell. Stomata of
|
||
<emphasis box="[159,257,2043,2067]" italics="true" pageId="4" pageNumber="429">Z. bayeri</emphasis>
|
||
in the marginal parts are occasionally oriented longitudinally to the leaf margin, whereas in
|
||
<taxonomicName box="[620,713,2076,2099]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="429" phylum="Tracheophyta" rank="species" species="pateri">
|
||
<emphasis box="[620,713,2076,2099]" italics="true" pageId="4" pageNumber="429">Z. pateri</emphasis>
|
||
</taxonomicName>
|
||
they are always perpendicularly oriented (
|
||
<tableCitation box="[1247,1317,1308,1332]" captionStart="Table 1" captionStartId="4.[127,183,152,173]" captionTargetId="graphics-441@4.[129,1485,208,1232]" captionTargetPageId="4" captionText="Table 1. Comparison of Zamites pateri with related species." pageId="4" pageNumber="429">Tab. 1</tableCitation>
|
||
). Furthermore,
|
||
<emphasis box="[843,943,1341,1365]" italics="true" pageId="4" pageNumber="429">Z. bayeri</emphasis>
|
||
does not have preserved any mesophyll tissues. Additionally,
|
||
<emphasis box="[995,1096,1375,1399]" italics="true" pageId="4" pageNumber="429">Z. bayeri</emphasis>
|
||
comes from calcareous substrates of Barrandian limestones, whereas
|
||
<taxonomicName box="[1240,1338,1409,1432]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="429" phylum="Tracheophyta" rank="species" species="pateri">
|
||
<emphasis box="[1240,1338,1409,1432]" italics="true" pageId="4" pageNumber="429">Z. pateri</emphasis>
|
||
</taxonomicName>
|
||
comes from silicate substrates of Carboniferous clastic.
|
||
</paragraph>
|
||
<paragraph blockId="4.[843,1483,1308,2100]" pageId="4" pageNumber="429">
|
||
Discussion about other similar impression material, including
|
||
<emphasis box="[957,1169,1508,1532]" italics="true" pageId="4" pageNumber="429">
|
||
<taxonomicName box="[957,1041,1509,1532]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="429" phylum="Tracheophyta" rank="genus">Zamites</taxonomicName>
|
||
bohemicus
|
||
</emphasis>
|
||
VELEN. described from the Peruc-Korycany Formation by earlier authors was published in detail by
|
||
<bibRefCitation author="Knobloch, E. & Kvacek, J." box="[970,1288,1575,1599]" pageId="4" pageNumber="429" pagination="567 - 586" refId="ref3964" refString="Knobloch, E., Kvacek, J. (1997): Bennettitalean and coni- feralean (?) leaves from the Cretaceous (Cenomanian) of the Bohemian Massif, Central Europe. - Cretaceous Research, 18: 567 - 586. https: // doi. org / 10.1006 / cres. 1997.0074" type="journal article" year="1997">Knobloch and Kvaček (1997)</bibRefCitation>
|
||
, and therefore not discussed here.
|
||
</paragraph>
|
||
<paragraph blockId="4.[843,1483,1308,2100]" lastBlockId="5.[106,746,151,760]" lastPageId="5" lastPageNumber="430" pageId="4" pageNumber="429">
|
||
<taxonomicName box="[881,981,1642,1666]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="429" phylum="Tracheophyta" rank="species" species="pateri">
|
||
<emphasis box="[881,981,1642,1666]" italics="true" pageId="4" pageNumber="429">Z. pateri</emphasis>
|
||
</taxonomicName>
|
||
resembles some
|
||
<taxonomicName box="[1188,1365,1642,1666]" pageId="4" pageNumber="429">
|
||
<emphasis box="[1188,1272,1642,1665]" italics="true" pageId="4" pageNumber="429">Zamites</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
described from the English Wealden by
|
||
<bibRefCitation author="Watson, J. & Sincock, C. A." box="[1173,1478,1675,1699]" pageId="4" pageNumber="429" pagination="1 - 228" refId="ref4997" refString="Watson, J., Sincock, C. A. (1992): Bennettitales of the English Wealden. - Monographs, Palaeontographical Society, London, 145 (588): 1 - 228, 23 pls. https: // doi. org / 10.1080 / 25761900.2022.12131771" type="journal article" year="1992">Watson and Sincock (1992)</bibRefCitation>
|
||
. It particularly shares characters with
|
||
<emphasis box="[1252,1370,1709,1733]" italics="true" pageId="4" pageNumber="429">Z. dowellii</emphasis>
|
||
J.WATSON et SINCOCK,
|
||
<emphasis box="[973,1082,1742,1766]" italics="true" pageId="4" pageNumber="429">Z. corderi</emphasis>
|
||
J.WATSON et SINCOCK and
|
||
<emphasis box="[1373,1482,1742,1766]" italics="true" pageId="4" pageNumber="429">Z. nicolae</emphasis>
|
||
J.WATSON et SINCOCK in having long and narrow pinnae.
|
||
<emphasis box="[843,1026,1809,1833]" italics="true" pageId="4" pageNumber="429">Zaimets dowellii</emphasis>
|
||
is similar in possessing papillae on the abaxial cuticle, but differs from the present material in having stomata on the adaxial cuticle, thus being amphistomatic. Additionally, its abaxial cuticle shows stomata with clearly sinuous anticlinal walls.
|
||
<emphasis box="[1143,1324,1942,1966]" italics="true" pageId="4" pageNumber="429">
|
||
<taxonomicName box="[1143,1227,1943,1966]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="429" phylum="Tracheophyta" rank="genus">Zamites</taxonomicName>
|
||
corderi
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName box="[843,938,1976,1999]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="429" phylum="Tracheophyta" rank="species" species="pateri">
|
||
<emphasis box="[843,938,1976,1999]" italics="true" pageId="4" pageNumber="429">Z. pateri</emphasis>
|
||
</taxonomicName>
|
||
in lacking papillae on the abaxial side, while its cuticle shows ordinary epidermal cells with conical trichome bases and well-developed sinuous anticlinal walls.
|
||
<emphasis italics="true" pageId="4" pageNumber="429">
|
||
<taxonomicName box="[1397,1481,2043,2066]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="429" phylum="Tracheophyta" rank="genus">Zamites</taxonomicName>
|
||
nicolae
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName box="[1057,1148,2076,2099]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="4" pageNumber="429" phylum="Tracheophyta" rank="species" species="pateri">
|
||
<emphasis box="[1057,1148,2076,2099]" italics="true" pageId="4" pageNumber="429">Z. pateri</emphasis>
|
||
</taxonomicName>
|
||
in having abaxial ordinary cells with sinuous anticlinal walls, and with only one papilla per ordinary cell.
|
||
<emphasis box="[264,466,184,208]" italics="true" pageId="5" pageNumber="430">
|
||
<taxonomicName box="[264,348,185,208]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="430" phylum="Tracheophyta" rank="genus">Zamites</taxonomicName>
|
||
decurrens
|
||
</emphasis>
|
||
C.A.MENÉNDEZ from the Early Cretaceous of
|
||
<collectingCountry box="[326,432,216,240]" name="Argentina" pageId="5" pageNumber="430">Argentina</collectingCountry>
|
||
(
|
||
<bibRefCitation author="Menendez, C. A." box="[447,623,216,240]" pageId="5" pageNumber="430" pagination="3 - 42" refId="ref4506" refString="Menendez, C. A. (1966): Fossil Bennettitales from the Tico flora, Santa Cruz Province, Argentina. - Bulletin of the British Museum (Natural History), 12 (1): 3 - 42. https: // doi. org / 10.5962 / p. 313885" type="journal article" year="1966">Menéndez 1966</bibRefCitation>
|
||
) resembles
|
||
<taxonomicName box="[106,197,250,273]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="430" phylum="Tracheophyta" rank="species" species="pateri">
|
||
<emphasis box="[106,197,250,273]" italics="true" pageId="5" pageNumber="430">Z. pateri</emphasis>
|
||
</taxonomicName>
|
||
in cuticle, having stomata perpendicularly oriented, forming rows and belts. It differs from
|
||
<taxonomicName box="[539,634,282,305]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="430" phylum="Tracheophyta" rank="species" species="pateri">
|
||
<emphasis box="[539,634,282,305]" italics="true" pageId="5" pageNumber="430">Z. pateri</emphasis>
|
||
</taxonomicName>
|
||
in having spatulate pinnae. Its adaxial cuticle is thicker, consisting of uniform elongated cells. Its abaxial cuticle shows less dense stomata, forming regular rows (
|
||
<tableCitation box="[443,510,379,403]" captionStart="Table 1" captionStartId="4.[127,183,152,173]" captionTargetId="graphics-441@4.[129,1485,208,1232]" captionTargetPageId="4" captionText="Table 1. Comparison of Zamites pateri with related species." pageId="5" pageNumber="430">Tab. 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="5.[106,746,151,760]" pageId="5" pageNumber="430">
|
||
Other Early Cretaceous species of
|
||
<taxonomicName authority="from" authorityName="FROM" box="[489,626,411,435]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="425" phylum="Tracheophyta" rank="genus">
|
||
<emphasis box="[489,572,412,435]" italics="true" pageId="5" pageNumber="430">Zamites</emphasis>
|
||
from
|
||
</taxonomicName>
|
||
the English Wealden show more profound differences from
|
||
<taxonomicName box="[617,708,445,468]" class="Cycadopsida" genus="Zamites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cycadales" pageId="5" pageNumber="430" phylum="Tracheophyta" rank="species" species="pateri">
|
||
<emphasis box="[617,708,445,468]" italics="true" pageId="5" pageNumber="430">Z. pateri</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis italics="true" pageId="5" pageNumber="430">Z. carruthersii</emphasis>
|
||
SEWARD and
|
||
<emphasis box="[377,511,477,500]" italics="true" pageId="5" pageNumber="430">Z. manoniae</emphasis>
|
||
J.WATSON et SINCOCK differ in having narrowly to broadly oval pinnae with rounded bases and bluntly rounded apex;
|
||
<emphasis box="[468,634,541,565]" italics="true" pageId="5" pageNumber="430">Z. wendyellisae</emphasis>
|
||
J.WATSON et SINCOCK differs in having pinnae triangular in outline;
|
||
<emphasis italics="true" pageId="5" pageNumber="430">Z. notokenensis</emphasis>
|
||
J.WATSON et SINCOCK and
|
||
<emphasis box="[522,637,607,630]" italics="true" pageId="5" pageNumber="430">Z. tatianae</emphasis>
|
||
J.WATSON et SINCOCK differ in having smooth or tuberate but not papillate external side of abaxial cuticle (
|
||
<bibRefCitation author="Watson, J. & Sincock, C. A." box="[451,734,671,695]" pageId="5" pageNumber="430" pagination="1 - 228" refId="ref4997" refString="Watson, J., Sincock, C. A. (1992): Bennettitales of the English Wealden. - Monographs, Palaeontographical Society, London, 145 (588): 1 - 228, 23 pls. https: // doi. org / 10.1080 / 25761900.2022.12131771" type="journal article" year="1992">Watson and Sincock 1992</bibRefCitation>
|
||
). Further differential characters in epidermal micromorphology of the discussed species are shown in
|
||
<tableCitation box="[499,577,736,760]" captionStart="Table 1" captionStartId="4.[127,183,152,173]" captionTargetId="graphics-441@4.[129,1485,208,1232]" captionTargetPageId="4" captionText="Table 1. Comparison of Zamites pateri with related species." pageId="5" pageNumber="430">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |