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<document id="E8468B6250F68D77567CE4D4D0157993" ID-CLB-Dataset="300009" ID-DOI="10.32225/bkpmnh.2021.50_55" ID-GBIF-Dataset="70224806-ca38-47ca-bd8a-63da3ac532f9" ID-ISSN="2189-6720" ID-Zenodo-Dep="12774639" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1721260925551" checkinUser="felipe" docAuthor="Watanabe, Kyohei" docDate="2021" docId="03F987931B00A751FF2E6539FC157C60" docLanguage="en" docName="BullKanagawaPrefectMus.50.55-136.pdf" docOrigin="Bulletin of the Kanagawa Prefectural Museum (Natural Science) 50" docSource="https://doi.org/10.32225/bkpmnh.2021.50_55" docStyle="DocumentStyle:020B98DC1285EBBD02620724ECB44EDE.1:BullKanagawaPrefectMus.2013-.journal_article.english" docStyleId="020B98DC1285EBBD02620724ECB44EDE" docStyleName="BullKanagawaPrefectMus.2013-.journal_article.english" docStyleVersion="1" docTitle="Nipponaetes haeussleri" docType="treatment" docVersion="9" lastPageNumber="124" masterDocId="FFC0FFEB1B45A714FFCA6633FFD77447" masterDocTitle="Taxonomic and Zoogeographic Study of the Japanese Phygadeuontinae (Hymenoptera, Ichneumonidae), with Descriptions of 17 New Species" masterLastPageNumber="136" masterPageNumber="55" pageNumber="124" updateTime="1721329445643" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="AA28CF2F58015DF0FBA4D6E14CDDE3BF">Taxonomic and Zoogeographic Study of the Japanese Phygadeuontinae (Hymenoptera, Ichneumonidae), with Descriptions of 17 New Species</mods:title>
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<mods:namePart id="1431076BAC03C2439C8729E4D4A0DB87">Watanabe, Kyohei</mods:namePart>
<mods:affiliation id="4BA2A1F9676777780A21C2740C0DCC7A">Kanagawa Prefectural Museum of Natural History, 499 Iryuda, Odawara, Kanagawa 250 – 0031, Japan</mods:affiliation>
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<mods:title id="3568431C30E97A7F74A118F175F57904">Bulletin of the Kanagawa Prefectural Museum (Natural Science)</mods:title>
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<mods:date id="96A659FFFB25181A88AF8ED75C7E3ACB">2021</mods:date>
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<mods:number id="4B9DAC922E3E05A35AF90256E3F91636">2021-03-30</mods:number>
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<treatment id="03F987931B00A751FF2E6539FC157C60" ID-DOI="http://doi.org/10.5281/zenodo.12773558" ID-GBIF-Taxon="235610758" ID-Zenodo-Dep="12773558" LSID="urn:lsid:plazi:treatment:03F987931B00A751FF2E6539FC157C60" httpUri="http://treatment.plazi.org/id/03F987931B00A751FF2E6539FC157C60" lastPageNumber="124" pageId="69" pageNumber="124">
<subSubSection id="C34A650E1B00A751FF2E6539FD437765" box="[228,660,778,802]" pageId="69" pageNumber="124" type="nomenclature">
<paragraph id="8BEF36851B00A751FF2E6539FD437765" blockId="69.[228,660,778,880]" box="[228,660,778,802]" pageId="69" pageNumber="124">
<taxonomicName id="4C504D061B00A751FF2E6539FD437765" ID-CoL="47GWS" authority="(Uchida, 1933)" baseAuthorityName="Uchida" baseAuthorityYear="1933" box="[228,660,778,802]" class="Insecta" family="Ichneumonidae" genus="Nipponaetes" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="124" phylum="Arthropoda" rank="species" species="haeussleri">
<emphasis id="B924EA971B00A751FF2E6539FE3E7765" bold="true" box="[228,489,778,802]" italics="true" pageId="69" pageNumber="124">Nipponaetes haeussleri</emphasis>
(Uchida, 1933)
</taxonomicName>
</paragraph>
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<subSubSection id="C34A650E1B00A751FED36502FD88770E" box="[281,607,817,841]" pageId="69" pageNumber="124" type="vernacular_names">
<paragraph id="8BEF36851B00A751FED36502FD88770E" blockId="69.[228,660,778,880]" box="[281,607,817,841]" pageId="69" pageNumber="124">(SJN: Kobu-togari-himebachi)</paragraph>
</subSubSection>
<subSubSection id="C34A650E1B00A751FEFF656BFD947737" box="[309,579,856,880]" pageId="69" pageNumber="124" type="description">
<paragraph id="8BEF36851B00A751FEFF656BFD947737" blockId="69.[228,660,778,880]" box="[309,579,856,880]" pageId="69" pageNumber="124">
(
<figureCitation id="136B2A001B00A751FEF6656BFE137737" box="[316,452,856,880]" captionStart="Fig" captionStartId="69.[151,184,661,683]" captionTargetBox="[151,1398,151,630]" captionTargetId="figure-565@69.[151,1398,151,630]" captionTargetPageId="69" captionText="Fig. 57. Nipponaetes haeussleri (Uchida, 1933), KPM-NK 81832, female from Japan ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, fore wing; E, T I to T III, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774769" httpUri="https://zenodo.org/record/12774769/files/figure.png" pageId="69" pageNumber="124">Figs 57A–D</figureCitation>
,
<figureCitation id="136B2A001B00A751FE04656BFE2C7737" box="[462,507,856,880]" captionStart="Fig" captionStartId="72.[190,223,1065,1087]" captionTargetBox="[195,1428,161,998]" captionTargetId="figure-227@72.[189,1436,149,1010]" captionTargetPageId="72" captionText="Fig. 62. Bathythrix kuwanae Viereck, 1912 (A, KPM-NK 81718), Ba. margaretae Sawoniewicz, 1980 (B, KPM-NK 55017), Ba. thomsoni (Kerrich, 1942) (C, KPM-NK 81675), Diaglyptidea conformis (Gmelin, 1790) (D, KPM-NK 81645), Nipponaetes haeussleri (Uchida, 1933) (E, KPM-NK 81832), Acrolyta japonica sp. nov. (F, N, KPM-NK 54996, holotype), Acr. rufocincta (Gravenhorst, 1829) (G, KPM-NK 81732), Surculus japonicus sp. nov. (H, KPM-NK 81861, holotype), Isadelphus nigrus sp. nov. (I, KPM-NK 82003, holotype), Lochetica japonica sp. nov. (J, KPM-NK 81989, holotype), Megacara similis Sheng, 1999 (K, KPM-NK 81837), Gnotus chionops (Gravenhorst, 1829) (L, KPM-NK 82100), Gn. striatus (Uchida, 1930) (M, KPM-NK 82060), and Acr. flavicoxis Sheng &amp; Sun, 2014 (O, KPM-NK 81837), females from Japan ― A–C, head, frontal view; D, E, head, lateral view; F–K, anterior margin of clypeus; L, M, mandible; N, O, basal part of antenna, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12774781" httpUri="https://zenodo.org/record/12774781/files/figure.png" pageId="69" pageNumber="124">62E</figureCitation>
,
<figureCitation id="136B2A001B00A751FDCD656BFDEB7737" box="[519,572,856,880]" captionStart="Fig" captionStartId="74.[191,224,1111,1133]" captionTargetBox="[193,1432,157,1080]" captionTargetId="figure-134@74.[189,1438,151,1085]" captionTargetPageId="74" captionText="Fig.65.Acrolyta japonica sp. nov. (A, KPM-NK 54996, holotype), Diaglyptidea conformis (Gmelin,1790) (B, KPM-NK 81650), Micraris ryukyuensis sp. nov. (C, KPM-NK 81873, holotype), Bathythrix kuwanae Viereck, 1912 (D, KPM-NK 81718), Bentyra ryukyuana sp. nov. (E, KPM-NK 81887, holotype), Chirotica matsukemushii (Matsumura, 1926) (F, KPM-NK 81906), Paraphylax politus sp. nov. (G, KPM-NK 55079, holotype), Pa. transstriatus sp. nov. (H, KPM-NK 55080, holotype), Pa. yakushimensis sp. nov. (I, KPM-NK 55081, holotype), Pa. yambarensis sp. nov. (J, KPM-NK 55085, holotype), Ethelurgus sodalis kumatai Kusigemati, 1983 (K, KPM- NK 81938), Aclastus etorofuensis (Uchida, 1936) (L, KPM-NK 81976), Hemiteles kuro sp. nov. (M, KPM-NK 55024, holotype), H. yamatonis sp. nov. (O, KPM-NK 55033, paratype), Isadelphus nigrus sp. nov. (P, KPM-NK 82003, holotype), Lochetica japonica sp. nov. (Q, KPM-NK 81989, holotype), Mastrus molestae (Uchida, 1933) (R, KPM-NK 81987), Ma.takadai Momoi, 1970 (R, KPM-NK 81979), Gnotus chionops (Gravenhorst, 1829) (L, KPM-NK 82104), Gn. striatus (Uchida, 1930) (M, KPM-NK 82061), Orthizema semanotae Sheng &amp; Sun, 2014 (V, KPM-NK 82047), Nipponaetes haeussleri (Uchida, 1933) (W, KPM-NK 81832), and Surculus japonicus sp. nov. (X, KPM-NK 81861, holotype), females from Japan ― A-W, carinae around area superomedia of propodeum, dorsal view; X, T I, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12774787" httpUri="https://zenodo.org/record/12774787/files/figure.png" pageId="69" pageNumber="124">65W</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34A650E1B00A751FF5D6595FD0E77F9" box="[151,729,934,958]" pageId="69" pageNumber="124" type="reference_group">
<paragraph id="8BEF36851B00A751FF5D6595FD0E77F9" blockId="69.[151,729,934,958]" box="[151,729,934,958]" pageId="69" pageNumber="124">
<treatmentCitationGroup id="AB4011AB1B00A751FF5D6595FD0E77F9" box="[151,729,934,958]" pageId="69" pageNumber="124">
<treatmentCitation id="0AF110941B00A751FF5D6595FD0277F9" author="Uchida, T." box="[151,725,934,958]" page="159" pageId="69" pageNumber="124" year="1933">
<taxonomicName id="4C504D061B00A751FF5D6595FD0277F9" authority="Uchida, 1933 b: 159" authorityName="Uchida" authorityPageNumber="159" authorityYear="1933" box="[151,725,934,958]" class="Insecta" family="Ichneumonidae" genus="Hemiteles" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="124" phylum="Arthropoda" rank="species" species="haeussleri" subGenus="Nipponaetes">
<emphasis id="B924EA971B00A751FF5D6595FF2A77F9" box="[151,253,934,958]" italics="true" pageId="69" pageNumber="124">Hemiteles</emphasis>
(
<emphasis id="B924EA971B00A751FEC16594FE5D77F9" box="[267,394,935,958]" italics="true" pageId="69" pageNumber="124">Nipponaetes</emphasis>
)
<emphasis id="B924EA971B00A751FE536595FDD577F9" box="[409,514,934,958]" italics="true" pageId="69" pageNumber="124">haeussleri</emphasis>
<bibRefCitation id="EFC14B741B00A751FDC26595FD0277F9" author="Uchida, T." box="[520,725,934,958]" pageId="69" pageNumber="124" pagination="153 - 164" refId="ref59004" refString="Uchida, T., 1933 b. Uber die Schmarotzerhymenopteren von Grapholitha molesta Busck. in Japan. Insecta matsumurana, 7: 153 - 164." type="journal article" year="1933">Uchida, 1933b: 159</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34A650E1B00A751FF6565C0FBAC7198" pageId="69" pageNumber="124" type="description">
<paragraph id="8BEF36851B00A751FF6565C0FE077075" blockId="69.[151,738,1011,2087]" pageId="69" pageNumber="124">
<emphasis id="B924EA971B00A751FF6565C0FEFA704C" bold="true" box="[175,301,1011,1035]" pageId="69" pageNumber="124">Description</emphasis>
. Female (n=3). Body length
<quantity id="4CA89B601B00A751FD9465C7FD09704C" box="[606,734,1012,1036]" metricMagnitude="-3" metricUnit="m" metricValue="3.3499999999999996" metricValueMax="3.4" metricValueMin="3.3" pageId="69" pageNumber="124" unit="mm" value="3.3499999999999996" valueMax="3.4" valueMin="3.3">3.3–3.4 mm</quantity>
. Body covered with silver setae.
</paragraph>
<paragraph id="8BEF36851B00A751FF656271FDAF71D6" blockId="69.[151,738,1011,2087]" pageId="69" pageNumber="124">
Head. Clypeus 1.8 times as wide as maximum length, its anterior margin truncate (
<figureCitation id="136B2A001B00A751FE00625BFDFB70C7" box="[458,556,1128,1152]" captionStart="Fig" captionStartId="69.[151,184,661,683]" captionTargetBox="[151,1398,151,630]" captionTargetId="figure-565@69.[151,1398,151,630]" captionTargetPageId="69" captionText="Fig. 57. Nipponaetes haeussleri (Uchida, 1933), KPM-NK 81832, female from Japan ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, fore wing; E, T I to T III, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774769" httpUri="https://zenodo.org/record/12774769/files/figure.png" pageId="69" pageNumber="124">Fig. 57B</figureCitation>
), its median part slightly concave. Face polished, densely punctate, with a median strong convexity (
<figureCitation id="136B2A001B00A751FE7D6285FDEE7089" box="[439,569,1206,1230]" captionStart="Fig" captionStartId="69.[151,184,661,683]" captionTargetBox="[151,1398,151,630]" captionTargetId="figure-565@69.[151,1398,151,630]" captionTargetPageId="69" captionText="Fig. 57. Nipponaetes haeussleri (Uchida, 1933), KPM-NK 81832, female from Japan ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, fore wing; E, T I to T III, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774769" httpUri="https://zenodo.org/record/12774769/files/figure.png" pageId="69" pageNumber="124">Figs 57A, B</figureCitation>
,
<figureCitation id="136B2A001B00A751FD8C6285FDA57089" box="[582,626,1206,1230]" captionStart="Fig" captionStartId="72.[190,223,1065,1087]" captionTargetBox="[195,1428,161,998]" captionTargetId="figure-227@72.[189,1436,149,1010]" captionTargetPageId="72" captionText="Fig. 62. Bathythrix kuwanae Viereck, 1912 (A, KPM-NK 81718), Ba. margaretae Sawoniewicz, 1980 (B, KPM-NK 55017), Ba. thomsoni (Kerrich, 1942) (C, KPM-NK 81675), Diaglyptidea conformis (Gmelin, 1790) (D, KPM-NK 81645), Nipponaetes haeussleri (Uchida, 1933) (E, KPM-NK 81832), Acrolyta japonica sp. nov. (F, N, KPM-NK 54996, holotype), Acr. rufocincta (Gravenhorst, 1829) (G, KPM-NK 81732), Surculus japonicus sp. nov. (H, KPM-NK 81861, holotype), Isadelphus nigrus sp. nov. (I, KPM-NK 82003, holotype), Lochetica japonica sp. nov. (J, KPM-NK 81989, holotype), Megacara similis Sheng, 1999 (K, KPM-NK 81837), Gnotus chionops (Gravenhorst, 1829) (L, KPM-NK 82100), Gn. striatus (Uchida, 1930) (M, KPM-NK 82060), and Acr. flavicoxis Sheng &amp; Sun, 2014 (O, KPM-NK 81837), females from Japan ― A–C, head, frontal view; D, E, head, lateral view; F–K, anterior margin of clypeus; L, M, mandible; N, O, basal part of antenna, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12774781" httpUri="https://zenodo.org/record/12774781/files/figure.png" pageId="69" pageNumber="124">62E</figureCitation>
). MSL 1.3 times as long as BWM. Frons, vertex, and gena matt. OOL shorter than POL. Occipital carina complete, joined with hypostomal carina distant from mandibular base. Upper tooth of mandible distinctly longer than lower tooth. Base of mandible flat.Antenna with 20 flagellomeres.
</paragraph>
<paragraph id="8BEF36851B00A751FF656393FE6073CC" blockId="69.[151,738,1011,2087]" pageId="69" pageNumber="124">
Mesosoma. Lateral part of pronotum largely granulate, with epomia. Mesoscutum granulate and polished, with distinct notaulus. Scutellum polished, coriaceous, lateral carina present except for apex. Mesopleuron polished, granulate except for smooth speculum. Metapleuron granulate, with a complete juxtacoxal carina. Propodeal carinae complete and strong (
<figureCitation id="136B2A001B00A751FE2B60B9FD8772E6" box="[481,592,1673,1698]" captionStart="Fig" captionStartId="74.[191,224,1111,1133]" captionTargetBox="[193,1432,157,1080]" captionTargetId="figure-134@74.[189,1438,151,1085]" captionTargetPageId="74" captionText="Fig.65.Acrolyta japonica sp. nov. (A, KPM-NK 54996, holotype), Diaglyptidea conformis (Gmelin,1790) (B, KPM-NK 81650), Micraris ryukyuensis sp. nov. (C, KPM-NK 81873, holotype), Bathythrix kuwanae Viereck, 1912 (D, KPM-NK 81718), Bentyra ryukyuana sp. nov. (E, KPM-NK 81887, holotype), Chirotica matsukemushii (Matsumura, 1926) (F, KPM-NK 81906), Paraphylax politus sp. nov. (G, KPM-NK 55079, holotype), Pa. transstriatus sp. nov. (H, KPM-NK 55080, holotype), Pa. yakushimensis sp. nov. (I, KPM-NK 55081, holotype), Pa. yambarensis sp. nov. (J, KPM-NK 55085, holotype), Ethelurgus sodalis kumatai Kusigemati, 1983 (K, KPM- NK 81938), Aclastus etorofuensis (Uchida, 1936) (L, KPM-NK 81976), Hemiteles kuro sp. nov. (M, KPM-NK 55024, holotype), H. yamatonis sp. nov. (O, KPM-NK 55033, paratype), Isadelphus nigrus sp. nov. (P, KPM-NK 82003, holotype), Lochetica japonica sp. nov. (Q, KPM-NK 81989, holotype), Mastrus molestae (Uchida, 1933) (R, KPM-NK 81987), Ma.takadai Momoi, 1970 (R, KPM-NK 81979), Gnotus chionops (Gravenhorst, 1829) (L, KPM-NK 82104), Gn. striatus (Uchida, 1930) (M, KPM-NK 82061), Orthizema semanotae Sheng &amp; Sun, 2014 (V, KPM-NK 82047), Nipponaetes haeussleri (Uchida, 1933) (W, KPM-NK 81832), and Surculus japonicus sp. nov. (X, KPM-NK 81861, holotype), females from Japan ― A-W, carinae around area superomedia of propodeum, dorsal view; X, T I, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12774787" httpUri="https://zenodo.org/record/12774787/files/figure.png" pageId="69" pageNumber="124">Fig. 65W</figureCitation>
). Propodeum polished and irregularly or transversally rugulose. Length of fore wing
<quantity id="4CA89B601B00A751FED760EBFE4D72A8" box="[285,410,1752,1776]" metricMagnitude="-3" metricUnit="m" metricValue="3.0999999999999996" metricValueMax="3.3" metricValueMin="2.9" pageId="69" pageNumber="124" unit="mm" value="3.0999999999999996" valueMax="3.3" valueMin="2.9">2.9–3.3 mm</quantity>
. Fore wing without areolet (
<figureCitation id="136B2A001B00A751FD7360E4FF107351" captionStart="Fig" captionStartId="69.[151,184,661,683]" captionTargetBox="[151,1398,151,630]" captionTargetId="figure-565@69.[151,1398,151,630]" captionTargetPageId="69" captionText="Fig. 57. Nipponaetes haeussleri (Uchida, 1933), KPM-NK 81832, female from Japan ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, fore wing; E, T I to T III, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774769" httpUri="https://zenodo.org/record/12774769/files/figure.png" pageId="69" pageNumber="124">Fig. 57D</figureCitation>
). Nervellus subvertical, intercepted near the middle (
<figureCitation id="136B2A001B00A751FF556116FED5737A" box="[159,258,1829,1853]" captionStart="Fig" captionStartId="69.[151,184,661,683]" captionTargetBox="[151,1398,151,630]" captionTargetId="figure-565@69.[151,1398,151,630]" captionTargetPageId="69" captionText="Fig. 57. Nipponaetes haeussleri (Uchida, 1933), KPM-NK 81832, female from Japan ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, fore wing; E, T I to T III, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774769" httpUri="https://zenodo.org/record/12774769/files/figure.png" pageId="69" pageNumber="124">Fig. 57A</figureCitation>
). Hind femur 5.0 times as long as maximum depth in lateral view. Hind TS I: II: III: IV: V = 2.0: 0.9: 0.6: 0.4: 0.5. Tarsal claws simple.
</paragraph>
<paragraph id="8BEF36851B00A751FF6561A8FB4F77A2" blockId="69.[151,738,1011,2087]" lastBlockId="69.[813,1399,778,2087]" pageId="69" pageNumber="124">
Metasoma. T I 1.4–1.5 times as long as maximum width, longitudinally striated medially and laterally. Median dorsal carina of T I present except for posterior part. Dorsolateral carina of T I complete. T II granulate, sometimes covered with longitudinal striae anteriorly and oblique striae posteriorly except for smooth area on posterior part (
<figureCitation id="136B2A001B00A751FC75656BFBEB7737" box="[959,1084,856,880]" captionStart="Fig" captionStartId="69.[151,184,661,683]" captionTargetBox="[151,1398,151,630]" captionTargetId="figure-565@69.[151,1398,151,630]" captionTargetPageId="69" captionText="Fig. 57. Nipponaetes haeussleri (Uchida, 1933), KPM-NK 81832, female from Japan ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, fore wing; E, T I to T III, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774769" httpUri="https://zenodo.org/record/12774769/files/figure.png" pageId="69" pageNumber="124">Figs 57C, E</figureCitation>
). T III to T V sparsely punctate except for weak, oblique striae on antero-lateral area of T III. Ovipositor sheath 0.75 times as long as hind tibia. Ovipositor straight, without a nodus.
</paragraph>
<paragraph id="8BEF36851B00A751FC8F65C7FC1071FF" blockId="69.[813,1399,778,2087]" pageId="69" pageNumber="124">
Coloration (
<figureCitation id="136B2A001B00A751FC0B65C7FB92704B" box="[961,1093,1012,1036]" captionStart="Fig" captionStartId="69.[151,184,661,683]" captionTargetBox="[151,1398,151,630]" captionTargetId="figure-565@69.[151,1398,151,630]" captionTargetPageId="69" captionText="Fig. 57. Nipponaetes haeussleri (Uchida, 1933), KPM-NK 81832, female from Japan ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, fore wing; E, T I to T III, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774769" httpUri="https://zenodo.org/record/12774769/files/figure.png" pageId="69" pageNumber="124">Figs 57A–D</figureCitation>
). Body (excluding wings and legs) black to blackish brown. Ventral part of scape and pedicel and base of FL I reddish brown. Mandible yellow except for darkened apex. A small spot of malar space, palpi, and tegula yellow. T II more or less tinged with reddish brown. Membranous part of metasomal sternites and ovipositor yellowish brown. Wings hyaline. Veins and pterostigma yellowish brown to brown. Legs reddish brown to yellowish brown. Apex of hind femur and base and apical parts of hind tibia blackish brown. Apical part of T I and T III to T VII sometimes tinged with dark reddish brown.
</paragraph>
<paragraph id="8BEF36851B00A751FC8F63F4FBAC7198" blockId="69.[813,1399,778,2087]" box="[837,1147,1479,1503]" pageId="69" pageNumber="124">Male. Not studied in this study.</paragraph>
</subSubSection>
<subSubSection id="C34A650E1B00A751FC8F63DDFA8672E6" pageId="69" pageNumber="124" type="materials_examined">
<paragraph id="8BEF36851B00A751FC8F63DDFA8672E6" blockId="69.[813,1399,778,2087]" pageId="69" pageNumber="124">
<emphasis id="B924EA971B00A751FC8F63DDFBFB7241" bold="true" box="[837,1068,1518,1542]" pageId="69" pageNumber="124">Materials examined</emphasis>
.
<materialsCitation id="3B383CD81B00A751FBF163DDFCAE7213" ID-GBIF-Occurrence="4906826371" collectingDate="2011-04-29" collectionCode="KPM-NK" collectorName="M. Ito" country="Japan" location="Kanmaki" municipality="Takatsuki City" pageId="69" pageNumber="124" specimenCode="KPM-NK 81832" specimenCount="1" specimenCount-female="1" stateProvince="Osaka Pref.">
<collectingCountry id="F34776151B00A751FBF163DDFB467241" box="[1083,1169,1518,1542]" name="Japan" pageId="69" pageNumber="124">JAPAN</collectingCountry>
:
<specimenCode id="DBF69EFE1B00A751FB5763DDFA817241" box="[1181,1366,1518,1542]" collectionCode="KPM-NK" pageId="69" pageNumber="124">KPM-NK 81832</specimenCode>
,
<specimenCount id="9D56FD0C1B00A751FAA963DDFAA57242" box="[1379,1394,1518,1541]" count="1" pageId="69" pageNumber="124" type="female">F</specimenCount>
,
<collectingRegion id="4994F8671B00A751FCE76026FC7D726A" box="[813,938,1557,1581]" country="Japan" name="Osaka" pageId="69" pageNumber="124">Osaka Pref.</collectingRegion>
,
<collectingMunicipality id="6B8BACFF1B00A751FC7F6026FB99726A" box="[949,1102,1557,1581]" pageId="69" pageNumber="124">Takatsuki City</collectingMunicipality>
,
<location id="8E8F605E1B00A751FB906026FB6B726A" LSID="urn:lsid:plazi:treatment:03F987931B00A751FF2E6539FC157C60:8E8F605E1B00A751FB906026FB6B726A" box="[1114,1212,1557,1581]" country="Japan" municipality="Takatsuki City" name="Kanmaki" pageId="69" pageNumber="124" stateProvince="Osaka Pref.">Kanmaki</location>
,
<date id="FFEE10451B00A751FB026026FA9B726A" box="[1224,1356,1557,1581]" pageId="69" pageNumber="124" value="2011-04-29">
<collectingDate id="EFAAE9AD1B00A751FB026026FA9B726A" box="[1224,1356,1557,1581]" pageId="69" pageNumber="124" value="2011-04-29">29. IV. 2011</collectingDate>
</date>
,
<collectorName id="26A553531B00A751FA936026FC9D7214" pageId="69" pageNumber="124">M. Ito</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B383CD81B00A751FC4F600FFAA0723C" ID-GBIF-Occurrence="4906826369" collectingDate="1964-07-12" collectionCode="KU" collectorName="K. Yasumatsu &amp; T. Nishida" country="Japan" determinerName="Momoi" location="Yamashiro" municipality="Miyazaki" pageId="69" pageNumber="124" specimenCount="1" stateProvince="Miyazaki Pref.">
<collectionCode id="ED41AE401B00A751FC4F600FFC7A7214" box="[901,941,1596,1619]" country="USA" name="Biodiversity Institute, University of Kansas" pageId="69" pageNumber="124" type="University or college">KU</collectionCode>
,
<specimenCount id="9D56FD0C1B00A751FC73600FFC1F7214" box="[953,968,1596,1619]" pageId="69" pageNumber="124" type="female">F</specimenCount>
(det.
<determinerName id="7E33DA411B00A751FBCF600FFB827213" box="[1029,1109,1596,1620]" pageId="69" pageNumber="124">Momoi</determinerName>
),
<collectingRegion id="4994F8671B00A751FBA2600FFAD27213" box="[1128,1285,1596,1620]" country="Japan" name="Miyazaki" pageId="69" pageNumber="124">Miyazaki Pref.</collectingRegion>
,
<collectingMunicipality id="6B8BACFF1B00A751FADA600FFAA47213" box="[1296,1395,1596,1620]" pageId="69" pageNumber="124">Miyazaki</collectingMunicipality>
,
<location id="8E8F605E1B00A751FCE76050FC4C723C" LSID="urn:lsid:plazi:treatment:03F987931B00A751FF2E6539FC157C60:8E8F605E1B00A751FCE76050FC4C723C" box="[813,923,1635,1659]" country="Japan" municipality="Miyazaki" name="Yamashiro" pageId="69" pageNumber="124" stateProvince="Miyazaki Pref.">Yamashiro</location>
,
<date id="FFEE10451B00A751FC6E6050FBF9723C" box="[932,1070,1635,1659]" pageId="69" pageNumber="124" value="1964-07-12">
<collectingDate id="EFAAE9AD1B00A751FC6E6050FBF9723C" box="[932,1070,1635,1659]" pageId="69" pageNumber="124" value="1964-07-12">12. VII. 1964</collectingDate>
</date>
,
<collectorName id="26A553531B00A751FBF26050FB14723D" box="[1080,1219,1635,1658]" pageId="69" pageNumber="124">K. Yasumatsu</collectorName>
&amp;
<collectorName id="26A553531B00A751FB296050FA9B723C" box="[1251,1356,1635,1659]" pageId="69" pageNumber="124">T. Nishida</collectorName>
leg.
</materialsCitation>
<materialsCitation id="3B383CD81B00A751FCE760B9FA9972E6" ID-GBIF-Occurrence="4906826433" box="[813,1358,1674,1698]" collectionCode="SEHU" collectorName="Keikido &amp; Suigen" country="South Korea" location="Keikido" municipality="Suigen" pageId="69" pageNumber="124" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F34776151B00A751FCE760B9FC5E72E5" box="[813,905,1674,1698]" name="South Korea" pageId="69" pageNumber="124">KOREA</collectingCountry>
:
<collectionCode id="ED41AE401B00A751FC5B60B9FC0172E5" box="[913,982,1674,1698]" collectionName="SEHU" pageId="69" pageNumber="124">SEHU</collectionCode>
,
<specimenCount id="9D56FD0C1B00A751FC1560B9FC3972E6" box="[991,1006,1674,1697]" pageId="69" pageNumber="124" type="female">F</specimenCount>
(
<typeStatus id="54EB88271B00A751FC3060B9FB8172E5" box="[1018,1110,1674,1698]" pageId="69" pageNumber="124" type="holotype">holotype</typeStatus>
),
<location id="8E8F605E1B00A751FBAC60B9FB6E72E5" LSID="urn:lsid:plazi:treatment:03F987931B00A751FF2E6539FC157C60:8E8F605E1B00A751FBAC60B9FB6E72E5" box="[1126,1209,1674,1698]" country="South Korea" municipality="Suigen" name="Keikido" pageId="69" pageNumber="124">Keikido</location>
,
<collectingMunicipality id="6B8BACFF1B00A751FB0860B9FADD72E5" box="[1218,1290,1674,1698]" pageId="69" pageNumber="124">Suigen</collectingMunicipality>
, 30. X
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A650E1B00A751FC8F6083FAA172A8" pageId="69" pageNumber="124" type="distribution">
<paragraph id="8BEF36851B00A751FC8F6083FAA172A8" blockId="69.[813,1399,778,2087]" pageId="69" pageNumber="124">
<emphasis id="B924EA971B00A751FC8F6083FC10728F" bold="true" box="[837,967,1712,1736]" pageId="69" pageNumber="124">Distribution</emphasis>
.
<collectingCountry id="F34776151B00A751FC1F6082FBD8728F" box="[981,1039,1713,1736]" name="Japan" pageId="69" pageNumber="124">Japan</collectingCountry>
(Honshu*, Kyushu, Amamioshima Is., and Okinawajima Is.);
<collectingCountry id="F34776151B00A751FBF860EBFBA572A8" box="[1074,1138,1752,1775]" name="South Korea" pageId="69" pageNumber="124">Korea</collectingCountry>
, the
<collectingCountry id="F34776151B00A751FB5560E4FAC672A8" box="[1183,1297,1751,1775]" name="Philippines" pageId="69" pageNumber="124">Philippines</collectingCountry>
, and
<collectingCountry id="F34776151B00A751FA8960E4FAA172A8" box="[1347,1398,1751,1775]" name="India" pageId="69" pageNumber="124">India</collectingCountry>
</paragraph>
</subSubSection>
<subSubSection id="C34A650E1B00A751FC8F60CDFA8A739E" pageId="69" pageNumber="124" type="biology_ecology">
<paragraph id="8BEF36851B00A751FC8F60CDFA8A739E" blockId="69.[813,1399,778,2087]" pageId="69" pageNumber="124">
<emphasis id="B924EA971B00A751FC8F60CDFC6F7351" bold="true" box="[837,952,1790,1814]" pageId="69" pageNumber="124">Bionomics</emphasis>
. Host record:
<taxonomicName id="4C504D061B00A751FB9860CDFCB8737A" authority="(Busck, 1916)" baseAuthorityName="Busck" baseAuthorityYear="1916" class="Insecta" family="Tortricidae" genus="Grapholita" kingdom="Animalia" order="Lepidoptera" pageId="69" pageNumber="124" phylum="Arthropoda" rank="species" species="molesta">
<emphasis id="B924EA971B00A751FB9860CDFAC87351" box="[1106,1311,1790,1814]" italics="true" pageId="69" pageNumber="124">Grapholita molesta</emphasis>
(Busck, 1916)
</taxonomicName>
(
<taxonomicName id="4C504D061B00A751FC486116FBC3737A" authorityName="Linnaeus" authorityYear="1758" box="[898,1044,1829,1853]" class="Insecta" kingdom="Animalia" order="Lepidoptera" pageId="69" pageNumber="124" phylum="Arthropoda" rank="order">Lepidoptera</taxonomicName>
,
<taxonomicName id="4C504D061B00A751FBE96116FB72737A" box="[1059,1189,1829,1853]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="69" pageNumber="124" phylum="Arthropoda" rank="family">Tortricidae</taxonomicName>
) (
<bibRefCitation id="EFC14B741B00A751FB756116FAA7737A" author="Uchida, T." box="[1215,1392,1829,1853]" pageId="69" pageNumber="124" pagination="153 - 164" refId="ref59004" refString="Uchida, T., 1933 b. Uber die Schmarotzerhymenopteren von Grapholitha molesta Busck. in Japan. Insecta matsumurana, 7: 153 - 164." type="journal article" year="1933">Uchida, 1933b</bibRefCitation>
;
<collectingCountry id="F34776151B00A751FCE7617FFCBB7323" box="[813,876,1868,1892]" name="Japan" pageId="69" pageNumber="124">Japan</collectingCountry>
);
<taxonomicName id="4C504D061B00A751FCB5617EFB047323" authority="(Haliday, 1834)" baseAuthorityName="Haliday" baseAuthorityYear="1834" box="[895,1235,1868,1892]" class="Insecta" family="Braconidae" genus="Cotesia" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="124" phylum="Arthropoda" rank="species" species="ruficrus">
<emphasis id="B924EA971B00A751FCB5617EFBF07323" box="[895,1063,1868,1892]" italics="true" pageId="69" pageNumber="124">Cotesia ruficrus</emphasis>
(Haliday, 1834)
</taxonomicName>
(
<taxonomicName id="4C504D061B00A751FB28617FFAA47323" box="[1250,1395,1868,1892]" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="124" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
,
<taxonomicName id="4C504D061B00A751FCE76140FC7D73CC" box="[813,938,1907,1931]" class="Insecta" family="Braconidae" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="124" phylum="Arthropoda" rank="family">Braconidae</taxonomicName>
) (
<bibRefCitation id="EFC14B741B00A751FC086140FBFB73CC" author="He, J. H." box="[962,1068,1907,1931]" pageId="69" pageNumber="124" pagination="77 - 110" refId="ref56371" refString="He, J. H., 1984. A checklist of Ichneumon-flies parasitic on rice pests from China (Hymenoptera: Ichneumonidae). Acta Agriculturae Universitatis Zhejiangensis, 10 (1): 77 - 110. [In Chinese with English summary]" type="journal article" year="1984">He, 1984</bibRefCitation>
;
<collectingCountry id="F34776151B00A751FBF06140FBA873CC" box="[1082,1151,1907,1931]" name="China" pageId="69" pageNumber="124">China</collectingCountry>
); Egg sac of Araneida (
<taxonomicName id="4C504D061B00A751FCF961A9FC5D73F5" box="[819,906,1946,1970]" class="Pilidiophora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="69" pageNumber="124" phylum="Nemertea" rank="order">Araneae</taxonomicName>
) (
<bibRefCitation id="EFC14B741B00A751FC5461A9FB8073F5" author="Broad, G. R. &amp; N. M. Laurenne &amp; D. L. J. Quicke" box="[926,1111,1946,1970]" pageId="69" pageNumber="124" pagination="651 - 655" refId="ref55318" refString="Broad, G. R., N. M. Laurenne &amp; D. L. J. Quicke, 2004. The genus Nipponaetes (Hymenoptera: Ichneumonidae: Cryptinae) in Costa Rica, with a reassessment of the generic limits. European Journal of Entomology, 101: 651 - 655." type="journal article" year="2004">
Broad
<emphasis id="B924EA971B00A751FC2961A8FBC273F5" box="[995,1045,1946,1970]" italics="true" pageId="69" pageNumber="124">et al.</emphasis>
, 2004
</bibRefCitation>
;
<collectingCountry id="F34776151B00A751FBAA61A9FB4073F5" box="[1120,1175,1946,1970]" name="India" pageId="69" pageNumber="124">India</collectingCountry>
). The above specimen deposited in KU is collected by sweeping in paddy field.
</paragraph>
</subSubSection>
<subSubSection id="C34A650E1B00A751FC8F61DBFC157C60" pageId="69" pageNumber="124" type="discussion">
<paragraph id="8BEF36851B00A751FC8F61DBFC157C60" blockId="69.[813,1399,778,2087]" pageId="69" pageNumber="124">
<emphasis id="B924EA971B00A751FC8F61DBFC667C47" bold="true" box="[837,945,2024,2048]" pageId="69" pageNumber="124">Remarks</emphasis>
. This is the first record of this species from Honshu.
</paragraph>
</subSubSection>
</treatment>
</document>
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