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<document id="FA167C5AFA4D7820697A4B6B5EBDDC35" ID-DOI="10.5281/zenodo.273733" ID-GBIF-Dataset="2f8785c9-b5f9-453d-be7c-a673d94c540c" ID-ISSN="1175-5326" ID-Zenodo-Dep="273733" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459834519907" checkinUser="plazi" docAuthor="Viquez, Carlos &amp; De, Luis F." docDate="2007" docId="03F987867A563369FF3BFDBABF35F86F" docLanguage="en" docName="zt01463p045.pdf" docOrigin="Zootaxa 1463" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Mastigoproctus ayalai Viquez &amp; De, 2007, sp. n." docType="treatment" docVersion="9" lastPageNumber="42" masterDocId="FFC0FFFE7A57336AFFACFF85BA47FFF9" masterDocTitle="A new species of Mastigoproctus Pocock, 1894 (Thelyphonida: Thelyphonidae) from Venezuela" masterLastPageNumber="45" masterPageNumber="39" pageNumber="40" updateTime="1698223301027" updateUser="plazi">
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<mods:title id="EF8535CF3753B3D0B05D968844CAAF36">A new species of Mastigoproctus Pocock, 1894 (Thelyphonida: Thelyphonidae) from Venezuela</mods:title>
</mods:titleInfo>
<mods:name id="4F75FA207F8E3937FD35F59E4ABE4E2E" type="personal">
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<mods:namePart id="08C874CA0BC38A6A45A1B6C3D6F31C0A">Viquez, Carlos</mods:namePart>
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<mods:namePart id="CCDFC97CA3E3AA62759C54182DAF5337">De, Luis F.</mods:namePart>
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<mods:typeOfResource id="487D13C0F5E01175F06F1D3CAC318814">text</mods:typeOfResource>
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<mods:title id="B24E7173CECBC299A0FD2E4A5245195C">Zootaxa</mods:title>
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<mods:date id="DD2388408E531DF11E6813997CDD5644">2007</mods:date>
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<mods:number id="9BAA5BEBE3C88A087B14C86AEA99E212">1463</mods:number>
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<treatment id="03F987867A563369FF3BFDBABF35F86F" ID-DOI="http://doi.org/10.5281/zenodo.5689417" ID-GBIF-Taxon="119358527" ID-Zenodo-Dep="5689417" LSID="urn:lsid:plazi:treatment:03F987867A563369FF3BFDBABF35F86F" httpUri="http://treatment.plazi.org/id/03F987867A563369FF3BFDBABF35F86F" lastPageId="3" lastPageNumber="42" pageId="1" pageNumber="40">
<subSubSection id="C34A651B7A56336BFF3BFDBAB82EFD85" pageId="1" pageNumber="40" type="nomenclature">
<paragraph id="8BEF36907A56336BFF3BFDBABBABFDA0" blockId="1.[151,617,575,636]" box="[151,492,575,601]" pageId="1" pageNumber="40">
<heading id="D0A781FC7A56336BFF3BFDBABBABFDA0" bold="true" box="[151,492,575,601]" fontSize="11" level="1" pageId="1" pageNumber="40" reason="1">
<emphasis id="B924EA827A56336BFF3BFDBABBABFDA0" bold="true" box="[151,492,575,601]" pageId="1" pageNumber="40">
<taxonomicName id="4C504D137A56336BFF3BFDBABBE4FDA0" ID-CoL="3YD43" box="[151,419,575,601]" class="Arachnida" family="Thelyphonidae" genus="Mastigoproctus" kingdom="Animalia" order="Uropygi" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="ayalai" status="sp. nov.">
<emphasis id="B924EA827A56336BFF3BFDBABBE4FDA0" bold="true" box="[151,419,575,601]" italics="true" pageId="1" pageNumber="40">Mastigoproctus ayalai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757F97A56336BFE06FDBABBABFDA0" box="[426,492,575,601]" pageId="1" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF36907A56336BFF3BFDE7B82EFD85" blockId="1.[151,617,575,636]" box="[151,617,610,636]" pageId="1" pageNumber="40">
(
<figureCitation id="136B2A157A56336BFF33FDE7BAB6FD85" box="[159,241,610,636]" captionStart="FIGURE 1" captionStartId="2.[151,255,1616,1640]" captionTargetBox="[272,1299,181,1579]" captionTargetId="figure@2.[264,1323,179,1590]" captionTargetPageId="2" captionText="FIGURE 1. Mastigoproctus ayalai sp. n. Male holotype. A, carapace; B, opisthosoma, dorsal aspect; C D, pedipalp: C, dorsal aspect; D, ventral aspect; E, sternites II IV; F, pigidium, lateral aspect." httpUri="https://zenodo.org/record/273734/files/figure.png" pageId="1" pageNumber="40">Figs. 1</figureCitation>
AF, 2 AF, 3 AB, 4,
<tableCitation id="C6D2032B7A56336BFDA4FDE7B81AFD85" box="[520,605,610,636]" captionStart="TABLE 1" captionStartId="5.[151,239,963,987]" captionTargetBox="[159,1261,1020,1640]" captionTargetPageId="5" captionText="TABLE 1. Measurements (mm) of Mastigoproctus ayalai sp. n. W, width; H, height; L, length." httpUri="http://table.plazi.org/id/DF2F66187A52336FFF3BFC46BED5FC22" pageId="1" pageNumber="40" tableUuid="DF2F66187A52336FFF3BFC46BED5FC22">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34A651B7A56336BFF3BFD37BEEEFCBD" pageId="1" pageNumber="40" type="materials_examined">
<paragraph id="8BEF36907A56336BFF3BFD37BEEEFCBD" blockId="1.[151,1437,690,2036]" pageId="1" pageNumber="40">
<typeStatus id="54EB88327A56336BFF3BFD37BA95FD35" box="[151,210,690,716]" pageId="1" pageNumber="40">
<emphasis id="B924EA827A56336BFF3BFD37BA95FD35" bold="true" box="[151,210,690,716]" pageId="1" pageNumber="40">Type</emphasis>
</typeStatus>
. Male
<typeStatus id="54EB88327A56336BFE81FD37BBD4FD35" box="[301,403,690,716]" pageId="1" pageNumber="40" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED41AE557A56336BFE0AFD37BBB0FD35" LSID="urn:lsid:biocol.org:col:33834" box="[422,503,690,716]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="1" pageNumber="40">MIZA</collectionCode>
), Quebrada Trapichote (
<quantity id="4CA89B757A56336BFCB7FD37B92AFD35" box="[795,877,690,716]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="40" unit="m" value="400.0">400 m</quantity>
), La
<taxonomicName id="4C504D137A56336BFC03FD37BE2BFD35" authority="Pijiguaos" authorityName="Pijiguaos" box="[943,1132,690,716]" class="Magnoliopsida" family="Convolvulaceae" genus="Mina" higherTaxonomySource="CoL" kingdom="Plantae" order="Solanales" pageId="1" pageNumber="40" phylum="Tracheophyta" rank="genus">Mina, Pijiguaos</taxonomicName>
(
<geoCoordinate id="EE6450577A56336BFBD3FD37BEAEFD35" box="[1151,1257,690,716]" direction="north" orientation="latitude" pageId="1" pageNumber="40" precision="925" value="6.5666666">6º 34 N</geoCoordinate>
<geoCoordinate id="EE6450577A56336BFAA1FD37BFC9FD35" box="[1293,1422,690,716]" direction="west" orientation="longitude" pageId="1" pageNumber="40" precision="925" value="-66.75">66º 45 W</geoCoordinate>
), estado Bolívar,
<collectingCountry id="F34776007A56336BFEE1FD5FBB8FFD0D" box="[333,456,730,756]" name="Venezuela" pageId="1" pageNumber="40">Venezuela</collectingCountry>
,
<date id="FFEE10507A56336BFE78FD5FB809FD0D" box="[468,590,730,756]" pageId="1" pageNumber="40" value="1992-06">June 1992</date>
, J. M. Ayala.
<typeStatus id="54EB88327A56336BFD41FD5FB923FD0A" box="[749,868,730,755]" pageId="1" pageNumber="40" type="paratype">
<emphasis id="B924EA827A56336BFD41FD5FB923FD0A" box="[749,868,730,755]" italics="true" pageId="1" pageNumber="40">Paratypes</emphasis>
</typeStatus>
: One adult male (IES) and one immature female (
<collectionCode id="ED41AE557A56336BFF33FC87BAA9FCE5" box="[159,238,770,796]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="1" pageNumber="40">INBio</collectionCode>
),
<specimenCount id="9D56FD197A56336BFEAFFC87BB30FCE5" box="[259,375,770,796]" pageId="1" pageNumber="40" type="female">8 females</specimenCount>
,
<specimenCount id="9D56FD197A56336BFE2FFC87BB9AFCE5" box="[387,477,770,796]" pageId="1" pageNumber="40" type="male">6 males</specimenCount>
(
<collectionCode id="ED41AE557A56336BFE41FC87B801FCE5" box="[493,582,770,796]" pageId="1" pageNumber="40">CMGS</collectionCode>
),
<specimenCount id="9D56FD197A56336BFDF6FC87B883FCE5" box="[602,708,770,796]" pageId="1" pageNumber="40" type="female">1 female</specimenCount>
,
<specimenCount id="9D56FD197A56336BFD63FC87B959FCE5" box="[719,798,770,796]" pageId="1" pageNumber="40" type="male">1 male</specimenCount>
(
<collectionCode id="ED41AE557A56336BFC82FC87B9C5FCE5" box="[814,898,770,796]" pageId="1" pageNumber="40">CJMA</collectionCode>
), same data as
<typeStatus id="54EB88327A56336BFB9EFC87BEDAFCE5" box="[1074,1181,770,796]" pageId="1" pageNumber="40" type="holotype">holotype</typeStatus>
. One female (
<collectionCode id="ED41AE557A56336BFAECFC87BFC8FCE5" box="[1344,1423,770,796]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="1" pageNumber="40">INBio</collectionCode>
), Los Pijiguaos (
<geoCoordinate id="EE6450577A56336BFEEBFCAFBBEDFCBD" box="[327,426,810,836]" direction="north" orientation="latitude" pageId="1" pageNumber="40" precision="925" value="6.5666666">6º 34 N</geoCoordinate>
<geoCoordinate id="EE6450577A56336BFE64FCAFB804FCBD" box="[456,579,810,836]" direction="west" orientation="longitude" pageId="1" pageNumber="40" precision="925" value="-66.75">66º 45 W</geoCoordinate>
), estado Bolívar,
<collectingCountry id="F34776007A56336BFCA3FCAFB9CDFCBD" box="[783,906,810,836]" name="Venezuela" pageId="1" pageNumber="40">Venezuela</collectingCountry>
,
<date id="FFEE10507A56336BFC3AFCAFBE57FCBD" box="[918,1040,810,836]" pageId="1" pageNumber="40" value="1991-06">June 1991</date>
, M. Ayala L.
</paragraph>
</subSubSection>
<subSubSection id="C34A651B7A56336BFF6BFCD7B8B6FC95" box="[199,753,850,876]" pageId="1" pageNumber="40" type="distribution">
<paragraph id="8BEF36907A56336BFF6BFCD7B8B6FC95" blockId="1.[151,1437,690,2036]" box="[199,753,850,876]" pageId="1" pageNumber="40">
<emphasis id="B924EA827A56336BFF6BFCD7BB27FC95" bold="true" box="[199,352,850,876]" pageId="1" pageNumber="40">Distribution</emphasis>
. Bolivar state,
<collectingCountry id="F34776007A56336BFDBEFCD7B8CCFC95" box="[530,651,850,876]" name="Venezuela" pageId="1" pageNumber="40">Venezuela</collectingCountry>
(
<figureCitation id="136B2A157A56336BFD36FCD7B8A4FC95" box="[666,739,850,876]" captionStart="FIGURE 4" captionStartId="4.[151,255,1915,1939]" captionText="FIGURE 4. Geographical distribution of Mastigoproctus ayalai sp. n." pageId="1" pageNumber="40">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34A651B7A56336BFF6BFCFFBE9AFC6D" box="[199,1245,890,916]" pageId="1" pageNumber="40" type="etymology">
<paragraph id="8BEF36907A56336BFF6BFCFFBE9AFC6D" blockId="1.[151,1437,690,2036]" box="[199,1245,890,916]" pageId="1" pageNumber="40">
<emphasis id="B924EA827A56336BFF6BFCFFBB0BFC6D" bold="true" box="[199,332,890,916]" pageId="1" pageNumber="40">Etymology</emphasis>
. This species is named in honor of J. M. Ayala L., collector of the
<typeStatus id="54EB88327A56336BFBF5FCFFBECCFC6D" box="[1113,1163,890,916]" pageId="1" pageNumber="40">type</typeStatus>
series.
</paragraph>
</subSubSection>
<subSubSection id="C34A651B7A56336BFF6BFC27B827FAB5" pageId="1" pageNumber="40" type="diagnosis">
<paragraph id="8BEF36907A56336BFF6BFC27B827FAB5" blockId="1.[151,1437,690,2036]" pageId="1" pageNumber="40">
<emphasis id="B924EA827A56336BFF6BFC27BB07FC45" bold="true" box="[199,320,930,956]" pageId="1" pageNumber="40">Diagnosis</emphasis>
. Total length
<quantity id="4CA89B757A56336BFE49FC27B82FFC45" box="[485,616,930,956]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="1" pageNumber="40" unit="mm" value="45.0" valueMax="50.0" valueMin="40.0">4050 mm</quantity>
; dark brown. Male: sternite II (genital plate) with a pair of anterolateral depressions; sternite III with a central, sub-triangular field, covered with short, strong bristles; IIIIV undivided. Pedipalps: trochanter moderately granulose, the remainder smooth; tibia higher than wide, not forming a conspicuous angle with respect to its longitudinal axis; scarce reddish hairs on the fingers; small ommatoids, ovoid in shape. Female: seminal receptacle tubular, J-shaped and distally swollen, with basal duct forming an angle of 45º with respect to the longitudinal axis of the opisthosoma. This species may be separated from
<taxonomicName id="4C504D137A56336BFE89FB16B86EFB55" authority="Hirst" authorityName="Hirst" box="[293,553,1170,1196]" class="Arachnida" family="Thelyphonidae" genus="Mastigoproctus" kingdom="Animalia" order="Uropygi" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="formidabilis">
<emphasis id="B924EA827A56336BFE89FB16BBA6FB55" box="[293,481,1171,1196]" italics="true" pageId="1" pageNumber="40">M. formidabilis</emphasis>
Hirst
</taxonomicName>
(
<figureCitation id="136B2A157A56336BFD93FB17B8CBFB55" box="[575,652,1170,1196]" captionStart="FIGURE 5" captionStartId="5.[151,255,825,849]" captionTargetBox="[264,1314,200,797]" captionTargetId="figure@5.[264,1323,194,801]" captionTargetPageId="5" captionText="FIGURE 5. Mastigoproctus formidabilis. Male and female sintype. A B, male pedipalp; A, female genitalia, dorsal aspect (arrow shows the seminal receptacle); B, Mastigoproctus colombianus. C D, male pedipalp; C, female genitalia, dorsal aspect (arrow shows the seminal receptacle); D." httpUri="https://zenodo.org/record/273737/files/figure.png" pageId="1" pageNumber="40">Fig. 5</figureCitation>
), by having more attenuate male pedipalp, with tibia ventrally unarmed (in
<taxonomicName id="4C504D137A56336BFE80FB3EBBA6FB2D" box="[300,481,1211,1236]" class="Arachnida" family="Thelyphonidae" genus="Mastigoproctus" kingdom="Animalia" order="Uropygi" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="formidabilis">
<emphasis id="B924EA827A56336BFE80FB3EBBA6FB2D" box="[300,481,1211,1236]" italics="true" pageId="1" pageNumber="40">M. formidabilis</emphasis>
</taxonomicName>
the tibia has a large ventral tooth); also, female seminal receptacles are very short in
<taxonomicName id="4C504D137A56336BFF18FB66BB2DFB05" box="[180,362,1251,1276]" class="Arachnida" family="Thelyphonidae" genus="Mastigoproctus" kingdom="Animalia" order="Uropygi" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="formidabilis">
<emphasis id="B924EA827A56336BFF18FB66BB2DFB05" box="[180,362,1251,1276]" italics="true" pageId="1" pageNumber="40">M. formidabilis</emphasis>
</taxonomicName>
, resembling those of
<taxonomicName id="4C504D137A56336BFDCAFB66B9FAFB05" authority="Valerio" authorityName="Valerio" box="[614,957,1250,1276]" class="Arachnida" family="Thelyphonidae" genus="Valeriophonus" kingdom="Animalia" order="Uropygi" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="nara">
<emphasis id="B924EA827A56336BFDCAFB66B909FB05" box="[614,846,1251,1276]" italics="true" pageId="1" pageNumber="40">Valeriophonus nara</emphasis>
(Valerio)
</taxonomicName>
from
<collectingCountry id="F34776007A56336BFBA8FB67BEC0FB05" box="[1028,1159,1250,1276]" name="Costa Rica" pageId="1" pageNumber="40">Costa Rica</collectingCountry>
. The Colombian
<taxonomicName id="4C504D137A56336BFAE3FB66B80DFADD" authority="Mello-Leitao" authorityName="Mello-Leitao" class="Arachnida" family="Thelyphonidae" genus="Mastigoproctus" kingdom="Animalia" order="Uropygi" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="colombianus">
<emphasis id="B924EA827A56336BFAE3FB66BBEFFADD" italics="true" pageId="1" pageNumber="40">Mastigoproctus colombianus</emphasis>
Mello-Leitão
</taxonomicName>
has shorter and digitiform seminal receptacles, as well as stouter, armed and densely punctate pedipalp (
<figureCitation id="136B2A157A56336BFDA4FAB7B817FAB5" box="[520,592,1330,1356]" captionStart="FIGURE 5" captionStartId="5.[151,255,825,849]" captionTargetBox="[264,1314,200,797]" captionTargetId="figure@5.[264,1323,194,801]" captionTargetPageId="5" captionText="FIGURE 5. Mastigoproctus formidabilis. Male and female sintype. A B, male pedipalp; A, female genitalia, dorsal aspect (arrow shows the seminal receptacle); B, Mastigoproctus colombianus. C D, male pedipalp; C, female genitalia, dorsal aspect (arrow shows the seminal receptacle); D." httpUri="https://zenodo.org/record/273737/files/figure.png" pageId="1" pageNumber="40">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34A651B7A563368FF6BFADFBF78F809" lastPageId="2" lastPageNumber="41" pageId="1" pageNumber="40" type="description">
<paragraph id="8BEF36907A56336BFF6BFADFB80CFA15" blockId="1.[151,1437,690,2036]" pageId="1" pageNumber="40">
<emphasis id="B924EA827A56336BFF6BFADFBB3DFA8D" bold="true" box="[199,378,1370,1396]" pageId="1" pageNumber="40">
Male
<typeStatus id="54EB88327A56336BFEA2FADFBB3DFA8D" box="[270,378,1370,1396]" pageId="1" pageNumber="40" type="holotype">holotype</typeStatus>
</emphasis>
. Reddish brown in color, paler on leg I. Carapace with interocular triangle mostly rugose; the remaining part of the carapace is densely granulose; median eyes separated by 1.4 times one ocular diameter, and 1.0 mm from the anterior margin; posterior fovea is wide and moderately deep; epistoma present. Carapace 1.7 times longer than wide.
</paragraph>
<paragraph id="8BEF36907A56336BFF6BFA7FB94CF9C5" blockId="1.[151,1437,690,2036]" pageId="1" pageNumber="40">Tergites with abundant fine granules; IIIII divided by a median longitudinal suture. Segment X laterally divided; XII with one pair of small ovoid ommatoids.</paragraph>
<paragraph id="8BEF36907A56336BFF6BF9CFBFC9F94D" blockId="1.[151,1437,690,2036]" pageId="1" pageNumber="40">
Sternite II (genital plate) with one pair of anterolateral depressions, posterior margin of the sternal plate slightly convex; III two times longer than IV, with a central sub-triangular field, rather elevated and covered with abundant short rigid bristles; IIIV with feeble rugosity. Genitalia broad, twice as wide as long (
<figureCitation id="136B2A157A56336BFA9BF91FBF39F94D" box="[1335,1406,1690,1716]" captionStart="FIGURE 3" captionStartId="4.[151,255,1054,1078]" captionTargetBox="[449,1139,203,1026]" captionTargetId="figure@4.[442,1145,194,1030]" captionTargetPageId="4" captionText="FIGURE 3. Mastigoproctus ayalai sp. n. Genitalia. A, male, ventral aspect, B, female, dorsal aspect (arrow shows the seminal receptacle)." httpUri="https://zenodo.org/record/273736/files/figure.png" pageId="1" pageNumber="40">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8BEF36907A56336BFF6BF947B93EF85D" blockId="1.[151,1437,690,2036]" pageId="1" pageNumber="40">Pedipalps elongate, slightly pilose. Coxa externally granulose, ventrally smooth, with scarce and fine punctuation; apophysis with one distal tooth. Trochanter moderately granulose, with two ventral teeth and five acute dorsal internal ones. Femur sub-cylindrical, smooth, 1.5 times longer than patella, with one small dorsal internal tooth and one strong ventral internal tooth. Patella sub-cylindrical, smooth, apophysis long, without denticles. Tibia (manus) very long, sub-cylindrical and smooth, 1.2 times higher than wide, without internal ventrodistal tooth; movable finger 0.5 times longer than femur.</paragraph>
<paragraph id="8BEF36907A56336BFF6BF837BEA6F80D" blockId="1.[151,1437,690,2036]" pageId="1" pageNumber="40">Legs. Tarsal segments of first leg unmodified. Coxa, trochanter and femur of IIIV externally granulose. Tibiae of IIIIV with one ventral distal spur. Basitarsus of IIIV with two ventral distal spurs.</paragraph>
<caption id="DF2F66187A553368FF3BF9D5B996F973" httpUri="https://zenodo.org/record/273734/files/figure.png" pageId="2" pageNumber="41" targetBox="[272,1299,181,1579]" targetPageId="2">
<paragraph id="8BEF36907A553368FF3BF9D5B996F973" blockId="2.[151,1436,1616,1674]" pageId="2" pageNumber="41">
<emphasis id="B924EA827A553368FF3BF9D5BB5EF991" bold="true" box="[151,281,1616,1640]" pageId="2" pageNumber="41">FIGURE 1.</emphasis>
<taxonomicName id="4C504D137A553368FE8CF9D5B857F99E" box="[288,528,1616,1639]" class="Arachnida" family="Thelyphonidae" genus="Mastigoproctus" kingdom="Animalia" order="Uropygi" pageId="2" pageNumber="41" phylum="Arthropoda" rank="species" species="ayalai" status="sp. nov.">
<emphasis id="B924EA827A553368FE8CF9D5B857F99E" box="[288,528,1616,1639]" italics="true" pageId="2" pageNumber="41">Mastigoproctus ayalai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757F97A553368FDBAF9D5B815F991" box="[534,594,1616,1640]" pageId="2" pageNumber="41" rank="species">
<emphasis id="B924EA827A553368FDBAF9D5B815F991" bold="true" box="[534,594,1616,1640]" pageId="2" pageNumber="41">sp. n.</emphasis>
</taxonomicNameLabel>
Male holotype. A, carapace; B, opisthosoma, dorsal aspect; CD, pedipalp: C, dorsal aspect; D, ventral aspect; E, sternites IIIV; F, pigidium, lateral aspect.
</paragraph>
</caption>
<paragraph id="8BEF36907A553368FF6BF938B880F881" blockId="2.[151,1436,1725,2032]" pageId="2" pageNumber="41">
<emphasis id="B924EA827A553368FF6BF938BB65F92E" bold="true" box="[199,290,1725,1751]" pageId="2" pageNumber="41">Female</emphasis>
. The female differs from the male by having shorter and more robust pedipalps, tibia unmodified, tibial apophysis serrate and shorter; both patella and tibia with one ventral distal tooth. Sternite II (genital plate) smooth, without lateral depressions, with the posterior margin medially and weakly bilobulate; III slight longer than IV, without pilose field; IV with the anterior margin straight. Seminal receptacle tubular, J-shaped, distally swollen and backward directed (
<figureCitation id="136B2A157A553368FDDCF8DBB8F0F881" box="[624,695,1886,1912]" captionStart="FIGURE 3" captionStartId="4.[151,255,1054,1078]" captionTargetBox="[449,1139,203,1026]" captionTargetId="figure@4.[442,1145,194,1030]" captionTargetPageId="4" captionText="FIGURE 3. Mastigoproctus ayalai sp. n. Genitalia. A, male, ventral aspect, B, female, dorsal aspect (arrow shows the seminal receptacle)." httpUri="https://zenodo.org/record/273736/files/figure.png" pageId="2" pageNumber="41">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8BEF36907A553368FF6BF800BF78F809" blockId="2.[151,1436,1725,2032]" pageId="2" pageNumber="41">
<emphasis id="B924EA827A553368FF6BF800BBC0F866" bold="true" box="[199,391,1925,1951]" pageId="2" pageNumber="41">Natural history</emphasis>
. Most of the adult specimens were found under fallen logs, in tropical rain forest. Some of the vinegarroons also came from crevices on walls along the sides of semidry creeks, where they constructed small tunnels each ending in a more wide excavation (J. M. Ayala, personal communication).
</paragraph>
</subSubSection>
<caption id="DF2F66187A543369FF3BF9EAB9B1F950" httpUri="https://zenodo.org/record/273735/files/figure.png" pageId="3" pageNumber="42" targetBox="[268,1318,216,1620]" targetPageId="3">
<paragraph id="8BEF36907A543369FF3BF9EAB9B1F950" blockId="3.[151,1437,1647,1705]" pageId="3" pageNumber="42">
<emphasis id="B924EA827A543369FF3BF9EABB5DF97E" bold="true" box="[151,282,1647,1671]" pageId="3" pageNumber="42">FIGURE 2.</emphasis>
<taxonomicName id="4C504D137A543369FE8DF9EAB856F97F" box="[289,529,1647,1670]" class="Arachnida" family="Thelyphonidae" genus="Mastigoproctus" kingdom="Animalia" order="Uropygi" pageId="3" pageNumber="42" phylum="Arthropoda" rank="species" species="ayalai" status="sp. nov.">
<emphasis id="B924EA827A543369FE8DF9EAB856F97F" box="[289,529,1647,1670]" italics="true" pageId="3" pageNumber="42">Mastigoproctus ayalai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757F97A543369FDB5F9EAB812F97E" box="[537,597,1647,1671]" pageId="3" pageNumber="42" rank="species">
<emphasis id="B924EA827A543369FDB5F9EAB812F97E" bold="true" box="[537,597,1647,1671]" pageId="3" pageNumber="42">sp. n.</emphasis>
</taxonomicNameLabel>
Female paratype. A, carapace; B, opisthosoma, dorsal aspect; CD, pedipalp: C, dorsal aspect; D, ventral aspect; E, sternites IIIII; F, pigidium, lateral aspect..
</paragraph>
</caption>
<subSubSection id="C34A651B7A543369FF6BF959BF35F86F" pageId="3" pageNumber="42" type="discussion">
<paragraph id="8BEF36907A543369FF6BF959BF35F86F" blockId="3.[151,1436,1756,1942]" pageId="3" pageNumber="42">
<emphasis id="B924EA827A543369FF6BF959BB09F90F" bold="true" box="[199,334,1756,1782]" pageId="3" pageNumber="42">Comments</emphasis>
. According with recent changes in the taxonomy of Central American whip scorpions (
<bibRefCitation id="EFC14B617A543369FAEBF959BB21F8E7" author="Viquez" pageId="3" pageNumber="45" refString="Viquez, C. and Armas, L. F. de (2005). Dos nuevos generos de vinagrillos de America Central y las Antillas (Arachnida: Thelyphonida). Boletin de la Sociedad Entomologica Aragonesa, 37, 95 - 98." type="journal article" year="2005">Viquez and Armas, 2005</bibRefCitation>
,
<bibRefCitation id="EFC14B617A543369FED9F881BBF4F8E7" author="Viquez" box="[373,435,1796,1822]" pageId="3" pageNumber="45" refString="Viquez, C. &amp; Armas, L. F. de (2006). Un nuevo genero y dos nuevas especies de vinagrillos centroamericanos (Arachnida: Thelyphonida). Boletin de la Sociedad Entomologica Aragonesa, 38, 37 - 41." type="journal article" year="2006">2006</bibRefCitation>
; Viquez &amp; Armas in press), the genus
<taxonomicName id="4C504D137A543369FC2BF880BE79F8E7" box="[903,1086,1797,1822]" class="Arachnida" family="Thelyphonidae" genus="Mastigoproctus" kingdom="Animalia" order="Uropygi" pageId="3" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="B924EA827A543369FC2BF880BE79F8E7" box="[903,1086,1797,1822]" italics="true" pageId="3" pageNumber="42">Mastigoproctus</emphasis>
</taxonomicName>
is now restricted to southern
<collectingCountry id="F34776007A543369FF3BF8A9BAADF8BF" box="[151,234,1836,1862]" name="United States of America" pageId="3" pageNumber="42">U.S.A.</collectingCountry>
,
<collectingCountry id="F34776007A543369FF54F8A9BB13F8BF" box="[248,340,1836,1862]" name="Mexico" pageId="3" pageNumber="42">Mexico</collectingCountry>
, Greater Antilles (
<collectingCountry id="F34776007A543369FD9CF8A9B835F8BF" box="[560,626,1836,1862]" name="Cuba" pageId="3" pageNumber="42">Cuba</collectingCountry>
, and Hispaniola), and South
<collectingCountry id="F34776007A543369FC61F8A9BE71F8BF" box="[973,1078,1836,1862]" name="United States of America" pageId="3" pageNumber="42">America</collectingCountry>
. It is now apparent for us that South American whip scorpions ascribe to this genus are an heterogeneous assemblage of species and for that reason they are needed of a profound revision in order of clarifying its taxonomic status and relationships.
</paragraph>
</subSubSection>
</treatment>
</document>