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<document id="13B1F44330BAE04AE26EA5FAAF89910B" ID="10.11646/zootaxa.4238.3.1SLASH10400" ID-CLB-Dataset="36345" ID-DOI="10.11646/zootaxa.4238.3.1" ID-GBIF-Dataset="030203e2-e6a1-4b59-bfa0-0689bf817b08" ID-ISSN="1175-5326" ID-Zenodo-Dep="345756" ID-ZooBank="D123950C-DF2B-45EE-9966-813339A83491" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1488787785636" checkinUser="plazi" docAuthor="Krikken, J. &amp; Huijbregts, J." docDate="2017" docId="03E5BB4DE83CFFCBAAE4F8B2FAEBF9F7" docLanguage="en" docName="zootaxa.4238.3.1SLASH10400.pdf" docOrigin="Zootaxa 4238 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onthophagus hollowayi Krikken &amp; Huijbregts, new species" docType="treatment" docVersion="7" lastPageNumber="326" masterDocId="FFDCC335E824FFD2AA73FF98FFDAFFDA" masterDocTitle="Sulawesi Onthophagus: seven new species in select groups (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="334" masterPageNumber="301" pageNumber="325" updateTime="1698437609924" updateUser="ExternalLinkService">
<mods:mods id="4ABB5E7B55A9EA14EA59828F26E45C5C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="80C5245BF200AE4222D3AF159A7AF1A1">
<mods:title id="823BDAFC6A7383EF91DA3DB7900E65C4">Sulawesi Onthophagus: seven new species in select groups (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
</mods:titleInfo>
<mods:name id="BF36CE3D74BA0B41C1509DC609E72A64" type="personal">
<mods:role id="5978D6239C8CEB7BAA2885A4C3FFF8E6">
<mods:roleTerm id="26C03845361B877B452560D17DA92D54">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7C6F21DAEE3C6AAA31ABD18A8C8D2D77">Krikken, J.</mods:namePart>
</mods:name>
<mods:name id="BA294328B2286D37D18D516ED12EAE9C" type="personal">
<mods:role id="CD69A4662735CFB880D73FCF474AC1C7">
<mods:roleTerm id="743D035EF0ECB78DA0F417A296C30B43">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D990E84B9258869606B11A253BE76FDF">Huijbregts, J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="41C627CACD778220FB0276BC1846CF34">text</mods:typeOfResource>
<mods:relatedItem id="E28AB6E902281E23668FDB9B7F6E17C3" type="host">
<mods:titleInfo id="AD0C8D8E33FE0B1FF05C663A1D341EE5">
<mods:title id="C61CD2D10CD7D642D0241DA665682BB9">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A3D5B612681C84A4D5317C30971D1193">
<mods:date id="4CCC26C2B10213BBAACA1115BFE00BAD">2017</mods:date>
<mods:detail id="00EC1447A43F325FB387EE1295814905" type="volume">
<mods:number id="A886FE67D404D6F5ED1668046F553CA9">4238</mods:number>
</mods:detail>
<mods:detail id="C2296619A32E909D2E51AD06220C53F3" type="issue">
<mods:number id="2C4D1DC04AFB9282B9A8D5B80382A95D">3</mods:number>
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<mods:extent id="13FD8F12E4C0819B8E406294E632A7F8" unit="page">
<mods:start id="85E5CC455F65FE811A4889786E1658BB">301</mods:start>
<mods:end id="9B8AA938BEA9BE32F16452DEBB3B4018">334</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:classification id="028A28B70069C5D4F23E4956084847AC">journal article</mods:classification>
<mods:identifier id="A83041168A7806D9D685694185AA2CDE" type="CLB-Dataset">36345</mods:identifier>
<mods:identifier id="A7A75AAD11C02A2480F811E96C5252D4" type="DOI">10.11646/zootaxa.4238.3.1</mods:identifier>
<mods:identifier id="35ADF20781B0D1F37D49F469FD07A30B" type="GBIF-Dataset">030203e2-e6a1-4b59-bfa0-0689bf817b08</mods:identifier>
<mods:identifier id="288F76CA8941FBC30DEA6003E3D35D6E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E7B9D7A219087477D418D6E1411255D7" type="Zenodo-Dep">345756</mods:identifier>
<mods:identifier id="02422AB44C19825B96B789546C112235" type="ZooBank">D123950C-DF2B-45EE-9966-813339A83491</mods:identifier>
</mods:mods>
<treatment id="03E5BB4DE83CFFCBAAE4F8B2FAEBF9F7" ID-DOI="http://doi.org/10.5281/zenodo.6030199" ID-GBIF-Taxon="127719075" ID-Zenodo-Dep="6030199" LSID="urn:lsid:plazi:treatment:03E5BB4DE83CFFCBAAE4F8B2FAEBF9F7" httpUri="http://treatment.plazi.org/id/03E5BB4DE83CFFCBAAE4F8B2FAEBF9F7" lastPageId="25" lastPageNumber="326" pageId="24" pageNumber="325">
<subSubSection id="C35659D0E83CFFCAAAE4F8B2FE5BF8BC" pageId="24" pageNumber="325" type="nomenclature">
<paragraph id="8BF30A5BE83CFFCAAAE4F8B2FCAFF89F" blockId="24.[151,885,1834,1894]" box="[151,885,1834,1861]" pageId="24" pageNumber="325">
<heading id="D0BBBD37E83CFFCAAAE4F8B2FCAFF89F" bold="true" box="[151,885,1834,1861]" fontSize="11" level="1" pageId="24" pageNumber="325" reason="1">
<emphasis id="B938D649E83CFFCAAAE4F8B2FCAFF89F" bold="true" box="[151,885,1834,1861]" pageId="24" pageNumber="325">
<taxonomicName id="4C4C71D8E83CFFCAAAE4F8B2FD02F89F" ID-CoL="7XN44" authority="Krikken &amp; Huijbregts" authorityName="Krikken &amp; Huijbregts" box="[151,728,1834,1861]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="hollowayi" status="sp. nov.">
<emphasis id="B938D649E83CFFCAAAE4F8B2FE62F89E" bold="true" box="[151,440,1834,1860]" italics="true" pageId="24" pageNumber="325">Onthophagus hollowayi</emphasis>
Krikken &amp; Huijbregts
</taxonomicName>
,
<taxonomicNameLabel id="A20B6B32E83CFFCAA896F8B3FCAFF89F" box="[741,885,1835,1861]" pageId="24" pageNumber="325" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF30A5BE83CFFCAAAE4F8D5FE5BF8BC" blockId="24.[151,885,1834,1894]" box="[151,385,1869,1894]" pageId="24" pageNumber="325">
(
<figureCitation id="137716DEE83CFFCAAAECF8D5FF21F8BC" box="[159,251,1869,1894]" captionStart="FIGURES 9 – 12" captionStartId="6.[151,264,1319,1342]" captionTargetBox="[172,1423,274,1297]" captionTargetId="figure@6.[151,1436,193,1298]" captionTargetPageId="6" captionText="FIGURES 9 – 12. Sulawesi Onthophagus species, oblique habitus. 9, O. begoniophilus, paratype male from Ambang. 10, O. annulopunctatus, paratype male from Mogogonipa. 11, O. hollowayi, holotype male from Dumoga. 12, O. seseba, holotype male from Batui (pygidium up). Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345758/files/figure.png" pageId="24" pageNumber="325">Figs. 11</figureCitation>
,
<figureCitation id="137716DEE83CFFCAAB74F8D6FEFFF8BC" box="[263,293,1870,1894]" captionStart="FIGURES 25 – 30" captionStartId="9.[151,264,1925,1948]" captionTargetBox="[170,1436,198,1875]" captionTargetId="figure@9.[151,1436,193,1904]" captionTargetPageId="9" captionText="FIGURES 25 – 30. Sulawesi Onthophagus species, dorsal forebody. 25, O. sopu, holotype male from Sopu. 26, O. sopu, paratype male from Sopu. 27, O. manguliensis, male from Mangoli. 28, O. annulopunctatus, paratype male from Mogogonipa. 29, O. hollowayi, holotype male from Dumoga. 30, O. seseba, holotype male from Batui. Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345761/files/figure.png" pageId="24" pageNumber="325">29</figureCitation>
,
<figureCitation id="137716DEE83CFFCAAB42F8D5FE94F8BC" box="[305,334,1869,1894]" captionStart="FIGURES 56 – 66" captionStartId="27.[151,264,1705,1728]" captionTargetBox="[182,1432,265,1662]" captionTargetId="figure@27.[151,1436,265,1684]" captionTargetPageId="27" captionText="FIGURES 56 – 66. Sulawesi Onthophagus body parts. 56, O. annulopunctatus, head full-face, paratype male from Mogogonipa. 57, O. annulopunctatus, left elytron dorsal, paratype male Sanggona. 58, O. annulopunctatus, protibial upper side, paratype male from Mogogonipa. 59, O. annulopunctatus, metatibial upperside, paratype male from Mogogonipa. 60, O. annulopunctatus, aedeagus lateral, paratype male Toraut. 61, O. annulopunctatus, head full-face, paratype female from Marena. 62, O. hollowayi, head full-face, holotype male from Dumoga. 63, O. hollowayi, left elytron dorsal, holotype male from Dumoga. 64, O. hollowayi, protibial upper side, holotype male from Dumoga. 65, O. hollowayi, metatibial upperside, holotype male from Dumoga. 66, O. hollowayi, aedeagus lateral, holotype male from Dumoga. 67, O. hollowayi, head full-face, paratype female from Malibagu. Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345765/files/figure.png" pageId="24" pageNumber="325">62</figureCitation>
–67)
</paragraph>
</subSubSection>
<subSubSection id="C35659D0E83CFFCBAAE4F80DFA46FE5D" lastPageId="25" lastPageNumber="326" pageId="24" pageNumber="325" type="materials_examined">
<paragraph id="8BF30A5BE83CFFCAAAE4F80DFD5CF82C" blockId="24.[151,1436,1941,2038]" pageId="24" pageNumber="325">
<materialsCitation id="3B240006E83CFFCAAAE4F80DFD5CF82C" ID-GBIF-Occurrence="1438340119" collectingDate="1985-02-24" collectingDateMax="1985-02-26" collectingDateMin="1985-02-24" collectionCode="BMNH" collectorName="Plot A" country="Indonesia" elevation="200" location="North Sulawesi" pageId="24" pageNumber="325" specimenCount="5" stateProvince="North Sulawesi" typeStatus="holotype">
<emphasis id="B938D649E83CFFCAAAE4F80DFE5BF874" bold="true" box="[151,385,1941,1966]" pageId="24" pageNumber="325">Material examined.</emphasis>
<typeStatus id="54F7B4F9E83CFFCAABFBF80DFE2AF874" box="[392,496,1941,1966]" pageId="24" pageNumber="325" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="ED5D929EE83CFFCAA84EF80EFD4EF874" box="[573,660,1942,1966]" pageId="24" pageNumber="325">BMNH</collectionCode>
) from
<collectingCountry id="F35B4ACBE83CFFCAA892F80DFC88F874" box="[737,850,1941,1966]" name="Indonesia" pageId="24" pageNumber="325">Indonesia</collectingCountry>
:
<collectingRegion id="4988C4B9E83CFFCAA92EF80DFBCAF874" box="[861,1040,1941,1966]" country="Indonesia" name="Sulawesi Utara" pageId="24" pageNumber="325">North Sulawesi</collectingRegion>
: Dumoga Bone National Park:
<collectorName id="26B96F8DE83CFFCAAF1CF80DFF74F808" pageId="24" pageNumber="325">Plot A</collectorName>
,
<collectingDate id="EFB6D573E83CFFCAAACAF821FE8EF808" box="[185,340,1977,2002]" pageId="24" pageNumber="325" value="1985-02-24" valueMax="1985-02-26" valueMin="1985-02-24">
<date id="FFF22C9BE83CFFCAAACAF821FE8EF808" box="[185,340,1977,2002]" pageId="24" pageNumber="325" value="1985-02-24" valueMax="1985-02-26" valueMin="1985-02-24">24–26.ii.1985</date>
</collectingDate>
Project Wallace;
<collectingMethod id="520D724CE83CFFCAA869F821FCC3F80B" box="[538,793,1977,2002]" pageId="24" pageNumber="325">flight interception trap</collectingMethod>
, lowland forest, approximately
<elevation id="0061ED68E83CFFCAAE0FF822FB1FF80B" box="[1148,1221,1978,2002]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="325" unit="m" value="200.0">
<quantity id="4CB4A7BEE83CFFCAAE0FF822FB1FF80B" box="[1148,1221,1978,2002]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="325" unit="m" value="200.0">200 m</quantity>
</elevation>
, B.M. 1985-10. The
<specimenCount id="9D4AC1D2E83CFFCAAA8AF845FEA3F82C" box="[249,377,2013,2038]" pageId="24" pageNumber="325" type="generic" typeStatus="paratypes">5 paratypes</specimenCount>
are summarized below:
</materialsCitation>
</paragraph>
<paragraph id="8BF30A5BE83DFFCBAAE4FF00FC91FF6A" blockId="25.[151,1437,152,1581]" box="[151,843,152,176]" pageId="25" pageNumber="326">
<collectingRegion id="4988C4B9E83DFFCBAAE4FF00FE57FF6A" box="[151,397,152,176]" country="Indonesia" name="Sulawesi Utara" pageId="25" pageNumber="326">
<collectionCode id="ED5D929EE83DFFCBAAE4FF00FE57FF6A" box="[151,397,152,176]" pageId="25" pageNumber="326">NORTH SULAWESI</collectionCode>
</collectingRegion>
:
<collectionCode id="ED5D929EE83DFFCBABE4FF00FC91FF6A" box="[407,843,152,176]" pageId="25" pageNumber="326">DUMOGA BONE NATIONAL PARK</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE83DFFCBAAE4FF25FF36FF2D" blockId="25.[151,1437,152,1581]" pageId="25" pageNumber="326">
- Malibagu Rd
<quantity id="4CB4A7BEE83DFFCBAB14FF24FE72FF0E" box="[359,424,188,212]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="25" pageNumber="326" unit="m" value="110.0">110m</quantity>
,
<date id="FFF22C9BE83DFFCBABC5FF24FD8CFF0F" box="[438,598,188,213]" pageId="25" pageNumber="326" value="1985-11-14" valueMax="1985-11-19" valueMin="1985-11-14">14-19.xi.1985</date>
, J. Krikken, pw38b, multistratal evergreen forest edge, 2 fish traps,
<quantity id="4CB4A7BEE83DFFCBAF1DFF24FA46FF0F" box="[1390,1436,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="25" pageNumber="326" unit="in" value="1.0">1 in</quantity>
<collectionCode id="ED5D929EE83DFFCBAAE4FF78FF36FF2D" box="[151,236,224,247]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="25" pageNumber="326">RMNH</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE83DFFCBAAE4FE9DFB4FFEC6" blockId="25.[151,1437,152,1581]" box="[151,1173,260,285]" pageId="25" pageNumber="326">
- Plot A,
<quantity id="4CB4A7BEE83DFFCBAB6EFE9CFEBCFEC6" box="[285,358,260,284]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="25" pageNumber="326" unit="m" value="200.0">200 m</quantity>
,
<date id="FFF22C9BE83DFFCBAB01FE9CFDCDFEC7" box="[370,535,260,285]" pageId="25" pageNumber="326" value="1985-03-13" valueMax="1985-03-20" valueMin="1985-03-13">13–20.iii.1985</date>
, lowland rain forest, flight interception trap,
<quantity id="4CB4A7BEE83DFFCBAE7CFE9CFBE0FEC7" box="[1039,1082,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="25" pageNumber="326" unit="in" value="1.0">1 in</quantity>
<collectionCode id="ED5D929EE83DFFCBAE32FE9DFB4FFEC6" box="[1089,1173,261,284]" pageId="25" pageNumber="326">BMNH</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE83DFFCBAAE4FEB0FB62FEE5" blockId="25.[151,1437,152,1581]" box="[151,1208,295,320]" pageId="25" pageNumber="326">
- Plot A,
<quantity id="4CB4A7BEE83DFFCBAB6EFEB0FEBCFEE5" box="[285,358,296,320]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="25" pageNumber="326" unit="m" value="200.0">200 m</quantity>
,
<date id="FFF22C9BE83DFFCBAB01FEBFFDE1FE9A" box="[370,571,295,320]" pageId="25" pageNumber="326" value="1985-11-30" valueMax="1985-12-06" valueMin="1985-11-30">30.xi–06.xii.1985</date>
, lowland rain forest, flight interception trap,
<quantity id="4CB4A7BEE83DFFCBAE41FEB0FB87FE9A" box="[1074,1117,295,320]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="25" pageNumber="326" unit="in" value="1.0">1 in</quantity>
<collectionCode id="ED5D929EE83DFFCBAE17FEB0FB62FEE5" box="[1124,1208,296,319]" pageId="25" pageNumber="326">BMNH</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE83DFFCBAAE4FED5FB5CFEBE" blockId="25.[151,1437,152,1581]" box="[151,1158,332,357]" pageId="25" pageNumber="326">
- Plot D,
<quantity id="4CB4A7BEE83DFFCBAB6EFED4FEBCFEBE" box="[285,358,332,356]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="25" pageNumber="326" unit="m" value="400.0">400 m</quantity>
,
<date id="FFF22C9BE83DFFCBAB01FED4FDCAFEBF" box="[370,528,332,357]" pageId="25" pageNumber="326" value="1985-02-06" valueMax="1985-02-13" valueMin="1985-02-06">06–13.ii.1985</date>
, lowland rainforest, flight interception trap,
<quantity id="4CB4A7BEE83DFFCBAE73FED4FBF1FEBF" box="[1024,1067,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="25" pageNumber="326" unit="in" value="1.0">1 in</quantity>
<collectionCode id="ED5D929EE83DFFCBAE41FED5FB5CFEBE" box="[1074,1158,333,356]" pageId="25" pageNumber="326">BMNH</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE83DFFCBAAE4FEE8FA46FE5D" blockId="25.[151,1437,152,1581]" box="[151,1436,367,392]" pageId="25" pageNumber="326">
- Toraut R
<quantity id="4CB4A7BEE83DFFCBAB5FFEE8FE54FE5D" box="[300,398,367,392]" metricMagnitude="3" metricUnit="m" metricValue="3.245" pageId="25" pageNumber="326" unit="m" value="3245.0">3, 245 m</quantity>
,
<date id="FFF22C9BE83DFFCBABE9FEF7FDEEFE52" box="[410,564,367,392]" pageId="25" pageNumber="326" value="1985-05-27" valueMax="1985-05-31" valueMin="1985-05-27">27–31.v.1985</date>
, J. Huijbregts, hh343, multistratal evergreen forest, 4 fungi traps,
<quantity id="4CB4A7BEE83DFFCBAF65FEE8FA9BFE52" box="[1302,1345,367,392]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="25" pageNumber="326" unit="in" value="1.0">1 in</quantity>
<collectionCode id="ED5D929EE83DFFCBAF3BFEE8FA46FE5D" box="[1352,1436,368,391]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="25" pageNumber="326">RMNH</collectionCode>
</paragraph>
</subSubSection>
<subSubSection id="C35659D0E83DFFCBAAB4FE0CFCE6FA3F" pageId="25" pageNumber="326" type="description">
<paragraph id="8BF30A5BE83DFFCBAAB4FE0CFC65FE0A" blockId="25.[151,1437,152,1581]" pageId="25" pageNumber="326">
<emphasis id="B938D649E83DFFCBAAB4FE0CFE83FE77" bold="true" box="[199,345,404,429]" pageId="25" pageNumber="326">Description.</emphasis>
<typeStatus id="54F7B4F9E83DFFCBAB13FE0CFE11FE77" box="[352,459,404,429]" pageId="25" pageNumber="326" type="holotype">Holotype</typeStatus>
, male. Body length approximately
<quantity id="4CB4A7BEE83DFFCBA925FE0CFC75FE76" box="[854,943,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="25" pageNumber="326" unit="mm" value="4.8">4.8 mm</quantity>
. Colour black-brown, almost entirely shiny. Pilosity pale yellow, dorsal side virtually glabrous (with microstubbles).
</paragraph>
<paragraph id="8BF30A5BE83DFFCBAAB4FE44FB44FD5E" blockId="25.[151,1437,152,1581]" pageId="25" pageNumber="326">Clypeal apex medially produced into short, reflexed lobe, apex truncate; clypeogenal transition nearly continuous at boder; clypeofrontal ridge arcuate, obsolescent; clypeal surface finely, distinctly punctate, except for somewhat smooth anterior part; rest of cephalic surface abundantly, evenly punctate; punctures separated by 1–3 times their diameter. Vertex slightly elevated, somewhat callose medially. Eyes with approximately 8 facet rows across widest point; ratio interocular distance / single maximum eye width approximately 8.</paragraph>
<paragraph id="8BF30A5BE83DFFCBAAB4FD08FC12FCE2" blockId="25.[151,1437,152,1581]" pageId="25" pageNumber="326">
Pronotum moderately convex, with anteromedian pair of slight approximated callosities (more distinct in profile); discal midline not impressed; anterior section of lateral border virtually straight; anterolateral angle weakly round, approximately 85°; sides in dorsal view more strongly round halfway their length; posterior section of lateral border distinctly sinuate, posterolateral angles obtuse; base finely marginate (
<date id="FFF22C9BE83DFFCBAE02FD64FB45FCCF" box="[1137,1183,764,789]" pageId="25" pageNumber="326" value="1960-10">x60</date>
); entire pronotal derm with abundant punctation; punctures simple, separated by 1–4 diameters.
</paragraph>
<paragraph id="8BF30A5BE83DFFCBAAB4FCDCFCDEFC7F" blockId="25.[151,1437,152,1581]" pageId="25" pageNumber="326">Elytral striae discally well defined, with distinct punctures, mainly separated by 3–5 times their diameter, weakly crenulate on interstriae; interstriae slightly convex, with numerous fine, subserially arranged punctures; elytral margins (including epipleuron) narrowly brown.</paragraph>
<paragraph id="8BF30A5BE83DFFCBAAB4FC37FDC9FB82" blockId="25.[151,1437,152,1581]" pageId="25" pageNumber="326">Antennal club light yellow. Metasternum with vague midline impression, disc with scattered micropunctation, anteromedian lobe abundantly punctate-setose; lateral wings with numerous seta-bearing, ocellate punctures. Abdominal ventrites brown, each with transverse row of seta-bearing, ocellate punctures. Pygidial disc also brown with abundant, distinct seta-bearing punctures; length of setae up to approximately 2 times punctural diameter; base finely marginate (ridge low).</paragraph>
<paragraph id="8BF30A5BE83DFFCBAAB4FBFDFB35FB32" blockId="25.[151,1437,152,1581]" pageId="25" pageNumber="326">
Legs brown. Protibia with 3+(1) larger denticles, spur elongate-acuminate. Mesotibial spurs unmodified; metatibial spur long, distally feebly sinuate, apex truncate. Profemoral underside moderately punctate-setose, mesofemoral and metafemoral undersides with sparse, seta-bearing punctures. Metatarsomere 1 about as longs as metatarsomere 2–4 combined; approximate proportions of spur and tarsomeres 1–5: 13//
<date id="FFF22C9BE83DFFCBAE04FB57FB65FB32" box="[1143,1215,1231,1256]" pageId="25" pageNumber="326" value="2011-04-06">11/6/4</date>
/3/6.
</paragraph>
<paragraph id="8BF30A5BE83DFFCBAAB4FB6CFDCCFAD7" blockId="25.[151,1437,152,1581]" box="[199,534,1268,1293]" pageId="25" pageNumber="326">
Aedeagus,
<figureCitation id="137716DEE83DFFCBAB30FB6CFE48FAD7" box="[323,402,1268,1293]" captionStart="FIGURES 56 – 66" captionStartId="27.[151,264,1705,1728]" captionTargetBox="[182,1432,265,1662]" captionTargetId="figure@27.[151,1436,265,1684]" captionTargetPageId="27" captionText="FIGURES 56 – 66. Sulawesi Onthophagus body parts. 56, O. annulopunctatus, head full-face, paratype male from Mogogonipa. 57, O. annulopunctatus, left elytron dorsal, paratype male Sanggona. 58, O. annulopunctatus, protibial upper side, paratype male from Mogogonipa. 59, O. annulopunctatus, metatibial upperside, paratype male from Mogogonipa. 60, O. annulopunctatus, aedeagus lateral, paratype male Toraut. 61, O. annulopunctatus, head full-face, paratype female from Marena. 62, O. hollowayi, head full-face, holotype male from Dumoga. 63, O. hollowayi, left elytron dorsal, holotype male from Dumoga. 64, O. hollowayi, protibial upper side, holotype male from Dumoga. 65, O. hollowayi, metatibial upperside, holotype male from Dumoga. 66, O. hollowayi, aedeagus lateral, holotype male from Dumoga. 67, O. hollowayi, head full-face, paratype female from Malibagu. Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345765/files/figure.png" pageId="25" pageNumber="326">Fig. 66</figureCitation>
(
<typeStatus id="54F7B4F9E83DFFCBABD1FB6CFDD0FAD7" box="[418,522,1268,1293]" pageId="25" pageNumber="326" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="8BF30A5BE83DFFCBAAB4FA81FD66FA8E" blockId="25.[151,1437,152,1581]" pageId="25" pageNumber="326">Measurements in mm. Maximum width of head 1.5. Median length of pronotum 1.7, maximum width 2.5. Sutural length of elytra 2.0, maximum width 2.7.</paragraph>
<paragraph id="8BF30A5BE83DFFCBAAB4FAC7FCA6FAA2" blockId="25.[151,1437,152,1581]" box="[199,892,1375,1400]" pageId="25" pageNumber="326">
<emphasis id="B938D649E83DFFCBAAB4FAC7FD84FAA2" bold="true" box="[199,606,1375,1400]" pageId="25" pageNumber="326">Variation and sexual dimorphism.</emphasis>
Body length
<quantity id="4CB4A7BEE83DFFCBA886FAF8FCA3FAA2" box="[757,889,1376,1400]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="4.8" metricValueMin="4.4" pageId="25" pageNumber="326" unit="mm" value="4.6" valueMax="4.8" valueMin="4.4">4.4–4.8mm</quantity>
.
</paragraph>
<paragraph id="8BF30A5BE83DFFCBAAB4FA1CFCE6FA3F" blockId="25.[151,1437,152,1581]" pageId="25" pageNumber="326">
<emphasis id="B938D649E83DFFCBAAB4FA1CFEC6FA46" box="[199,284,1412,1436]" italics="true" pageId="25" pageNumber="326">Female</emphasis>
with same labels as
<typeStatus id="54F7B4F9E83DFFCBA871FA1CFDBEFA47" box="[514,612,1412,1437]" pageId="25" pageNumber="326" type="holotype">holotype</typeStatus>
has clypeal margin hardly reflexed, evenly round; clypeal derm rugulate, clypeofrontal ridge arcuate, distinct; medially between eyes with transverse callosity. Pronotal surface anteriorly evenly round, without medially impressed anterior declivity.
</paragraph>
</subSubSection>
<subSubSection id="C35659D0E83DFFCBAAB4FA77FAEBF9F7" pageId="25" pageNumber="326" type="etymology">
<paragraph id="8BF30A5BE83DFFCBAAB4FA77FAEBF9F7" blockId="25.[151,1437,152,1581]" pageId="25" pageNumber="326">
<emphasis id="B938D649E83DFFCBAAB4FA77FE94F9D2" bold="true" box="[199,334,1519,1544]" pageId="25" pageNumber="326">Etymology.</emphasis>
Named after our colleague Jeremy D. Holloway, who played a role in the organization of Project Wallace 1985 and strongly stimulated our interest in Indo-Australian biogeography with his publications.
</paragraph>
</subSubSection>
</treatment>
</document>
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