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<document id="AF9ECAA9668F6FDAA5EDDFD5558B8EA0" ID="10.11646/zootaxa.4238.3.1SLASH10400" ID-CLB-Dataset="36345" ID-DOI="10.11646/zootaxa.4238.3.1" ID-GBIF-Dataset="030203e2-e6a1-4b59-bfa0-0689bf817b08" ID-ISSN="1175-5326" ID-Zenodo-Dep="345756" ID-ZooBank="D123950C-DF2B-45EE-9966-813339A83491" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1488787785636" checkinUser="plazi" docAuthor="Krikken, J. &amp; Huijbregts, J." docDate="2017" docId="03E5BB4DE839FFF2AAE4FBE4FA47FCCF" docLanguage="en" docName="zootaxa.4238.3.1SLASH10400.pdf" docOrigin="Zootaxa 4238 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onthophagus begoniophilus Krikken &amp; Huijbregts, new species" docType="treatment" docVersion="7" lastPageNumber="333" masterDocId="FFDCC335E824FFD2AA73FF98FFDAFFDA" masterDocTitle="Sulawesi Onthophagus: seven new species in select groups (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="334" masterPageNumber="301" pageNumber="330" updateTime="1698437609924" updateUser="ExternalLinkService">
<mods:mods id="247E7D6D7354D2503CB9E1E341F422E1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="90D2EAF043BEF956D44BEE5D84088B3D">Sulawesi Onthophagus: seven new species in select groups (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:name id="466C0893A27ACE1E0037B5CAD61DD71C" type="personal">
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<mods:namePart id="CE2A88C128E267193A936000F224C6AC">Krikken, J.</mods:namePart>
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<mods:namePart id="0A5A9BB88336C9D3E27692A86515833B">Huijbregts, J.</mods:namePart>
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<mods:title id="F47B3D4DD7B1E4A2AC38ED80D2E65325">Zootaxa</mods:title>
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<mods:date id="BE1F3275BFC24C76E94EC1C22C5DAFB2">2017</mods:date>
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<mods:classification id="AB0E160657EFAE9B4838E3E3A28CC29E">journal article</mods:classification>
<mods:identifier id="4935F063D5B0AD5C6B67D865949E5F05" type="CLB-Dataset">36345</mods:identifier>
<mods:identifier id="3CE85435EF3539EC01586C377B5F4D17" type="DOI">10.11646/zootaxa.4238.3.1</mods:identifier>
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<treatment id="03E5BB4DE839FFF2AAE4FBE4FA47FCCF" ID-DOI="http://doi.org/10.5281/zenodo.6030205" ID-GBIF-Taxon="127719077" ID-Zenodo-Dep="6030205" LSID="urn:lsid:plazi:treatment:03E5BB4DE839FFF2AAE4FBE4FA47FCCF" httpUri="http://treatment.plazi.org/id/03E5BB4DE839FFF2AAE4FBE4FA47FCCF" lastPageId="32" lastPageNumber="333" pageId="29" pageNumber="330">
<subSubSection id="C35659D0E839FFCFAAE4FBE4FE44FB6D" pageId="29" pageNumber="330" type="nomenclature">
<paragraph id="8BF30A5BE839FFCFAAE4FBE4FC71FB4C" blockId="29.[151,939,1148,1207]" box="[151,939,1148,1174]" pageId="29" pageNumber="330">
<heading id="D0BBBD37E839FFCFAAE4FBE4FC71FB4C" bold="true" box="[151,939,1148,1174]" fontSize="11" level="1" pageId="29" pageNumber="330" reason="1">
<emphasis id="B938D649E839FFCFAAE4FBE4FC71FB4C" bold="true" box="[151,939,1148,1174]" pageId="29" pageNumber="330">
<taxonomicName id="4C4C71D8E839FFCFAAE4FBE4FCD4FB4C" ID-CoL="7XMFG" authority="Krikken &amp; Huijbregts" authorityName="Krikken &amp; Huijbregts" box="[151,782,1148,1174]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="begoniophilus" status="sp. nov.">
<emphasis id="B938D649E839FFCFAAE4FBE4FE34FB4C" bold="true" box="[151,494,1148,1174]" italics="true" pageId="29" pageNumber="330">Onthophagus begoniophilus</emphasis>
Krikken &amp; Huijbregts
</taxonomicName>
,
<taxonomicNameLabel id="A20B6B32E839FFCFA968FBE4FC71FB4C" box="[795,939,1148,1174]" pageId="29" pageNumber="330" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF30A5BE839FFCFAAE4FB06FE44FB6D" blockId="29.[151,939,1148,1207]" box="[151,414,1182,1207]" pageId="29" pageNumber="330">
(
<figureCitation id="137716DEE839FFCFAAECFB06FF34FB6D" box="[159,238,1182,1207]" captionStart="FIGURES 9 – 12" captionStartId="6.[151,264,1319,1342]" captionTargetBox="[172,1423,274,1297]" captionTargetId="figure@6.[151,1436,193,1298]" captionTargetPageId="6" captionText="FIGURES 9 – 12. Sulawesi Onthophagus species, oblique habitus. 9, O. begoniophilus, paratype male from Ambang. 10, O. annulopunctatus, paratype male from Mogogonipa. 11, O. hollowayi, holotype male from Dumoga. 12, O. seseba, holotype male from Batui (pygidium up). Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345758/files/figure.png" pageId="29" pageNumber="330">Figs. 9</figureCitation>
,
<figureCitation id="137716DEE839FFCFAA89FB07FE99FB6D" box="[250,323,1183,1207]" captionStart="FIGURES 19 – 24" captionStartId="8.[151,264,1879,1902]" captionTargetBox="[171,1436,193,1838]" captionTargetId="figure@8.[151,1436,193,1858]" captionTargetPageId="8" captionText="FIGURES 19 – 24. Sulawesi Onthophagus species, dorsal, oblique (20 only) forebody. 19, O. sangirensis, male from Manado. 20, O. holosericus, female from Kendari, oblique. 21, O. holosericus, male from Marena. 22, O. holosericus, female from Marena. 23, O. begoniophilus, holotype male major from Ambang. 24, O. begoniophilus, paratype male medium from Ambang. Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345760/files/figure.png" pageId="29" pageNumber="330">23–24</figureCitation>
,
<figureCitation id="137716DEE839FFCFAB3DFB07FE4CFB6D" box="[334,406,1183,1207]" captionStart="FIGURES 68 – 81" captionStartId="30.[151,264,1760,1783]" captionTargetBox="[253,1344,205,1735]" captionTargetId="figure@30.[239,1344,191,1735]" captionTargetPageId="30" captionText="FIGURES 68 – 81. Sulawesi Onthophagus body parts. 68, O. seseba, head full-face, holotype male from Batui. 69, O. seseba, left elytron dorsal, holotype male from Batui. 70, O. seseba, protibial upper side, holotype male from Batui. 71, O. seseba, metatibial upperside, holotype male from Batui. 72, O. seseba, aedeagus lateral, holotype male from Batui. 73, O. seseba, head full-face, paratype female. 72, O. begoniophilus, aedeagus lateral, holotype from Ambang. 73, O. begoniophilus, head fullface, holotype from Ambang. 74, O. begoniophilus, head full-face, holotype from Ambang. 75, O. begoniophilus, left elytron dorsal, holotype from Ambang. 76, O. begoniophilus, head full-face, paratype male. 77, O. begoniophilus, left elytron caudal, paratype male. 78, O. begoniophilus, protibial upper side, paratype male. 79, O. begoniophilus, metatibial upperside, paratype male. 80, O. begoniophilus, aedeagus lateral, paratype male. 81, O. begoniophilus, head full-face, paratype female. Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345766/files/figure.png" pageId="29" pageNumber="330">72–81</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35659D0E839FFCFAAE4FB7EFDAAF815" pageId="29" pageNumber="330" type="materials_examined">
<paragraph id="8BF30A5BE839FFCFAAE4FB7EFF0FFA9C" blockId="29.[151,1437,1254,1999]" pageId="29" pageNumber="330">
<materialsCitation id="3B240006E839FFCFAAE4FB7EFF0FFA9C" ID-GBIF-Occurrence="1438340107" collectingDate="1985-04-09" collectingDateMax="1985-04-13" collectingDateMin="1985-04-09" collectionCode="RMNH" collectorName="Mt Ambang NR &amp; J. Huijbregts" country="Indonesia" elevation="1120" location="Kotamobagu" pageId="29" pageNumber="330" specimenCount="1" stateProvince="Sulawesi" typeStatus="holotype">
<emphasis id="B938D649E839FFCFAAE4FB7EFE52FB25" bold="true" box="[151,392,1254,1279]" pageId="29" pageNumber="330">Material examined.</emphasis>
<typeStatus id="54F7B4F9E839FFCFABE5FB7EFE24FB25" box="[406,510,1254,1279]" pageId="29" pageNumber="330" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="ED5D929EE839FFCFA829FB7FFD68FB25" box="[602,690,1255,1279]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="29" pageNumber="330">RMNH</collectionCode>
) from
<collectingCountry id="F35B4ACBE839FFCFA97FFB7EFCA7FB25" box="[780,893,1254,1279]" name="Indonesia" pageId="29" pageNumber="330">Indonesia</collectingCountry>
: N
<collectingRegion id="4988C4B9E839FFCFA9C1FB7EFBC1FB25" box="[946,1051,1254,1279]" country="Indonesia" name="Sulawesi Utara" pageId="29" pageNumber="330">Sulawesi</collectingRegion>
:
<collectorName id="26B96F8DE839FFCFAE5DFB7FFB2CFB24" box="[1070,1270,1254,1279]" pageId="29" pageNumber="330">Mt Ambang NR</collectorName>
,
<quantity id="4CB4A7BEE839FFCFAF7BFB7FFA8DFB25" box="[1288,1367,1254,1279]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="29" pageNumber="330" unit="km" value="20.0">20 km</quantity>
E of
<location id="8E935C80E839FFCFAAE4FA92FEF1FAF9" LSID="urn:lsid:plazi:treatment:03E5BB4DE839FFF2AAE4FBE4FA47FCCF:8E935C80E839FFCFAAE4FA92FEF1FAF9" box="[151,299,1290,1315]" country="Indonesia" name="Kotamobagu" pageId="29" pageNumber="330" stateProvince="Sulawesi">Kotamobagu</location>
, alt.
<elevation id="0061ED68E839FFCFAB10FA93FE61FAF8" box="[355,443,1291,1315]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="29" pageNumber="330" unit="m" value="1120.0">
<quantity id="4CB4A7BEE839FFCFAB10FA93FE61FAF8" box="[355,443,1291,1315]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="29" pageNumber="330" unit="m" value="1120.0">1120 m</quantity>
</elevation>
,
<collectingDate id="EFB6D573E839FFCFABBBFA92FDBCFAF9" box="[456,614,1290,1315]" pageId="29" pageNumber="330" value="1985-04-09" valueMax="1985-04-13" valueMin="1985-04-09">
<date id="FFF22C9BE839FFCFABBBFA92FDBCFAF9" box="[456,614,1290,1315]" pageId="29" pageNumber="330" value="1985-04-09" valueMax="1985-04-13" valueMin="1985-04-09">09-13.iv.1985</date>
</collectingDate>
,
<collectorName id="26B96F8DE839FFCFA800FA93FCDEFAF9" box="[627,772,1290,1315]" pageId="29" pageNumber="330">J. Huijbregts</collectorName>
HH443A; multistratal evergreen forest,
<collectingMethod id="520D724CE839FFCFAEA1FA92FF0BFA9C" pageId="29" pageNumber="330">human excrement traps</collectingMethod>
.
</materialsCitation>
</paragraph>
<paragraph id="8BF30A5BE839FFCFAAB4FACAFD5CFAB1" blockId="29.[151,1437,1254,1999]" box="[199,646,1362,1387]" pageId="29" pageNumber="330">
The
<specimenCount id="9D4AC1D2E839FFCFAA8AFACBFEA3FAB1" box="[249,377,1363,1387]" pageId="29" pageNumber="330" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
are summarized below:
</paragraph>
<paragraph id="8BF30A5BE839FFCFAAE4FAEFFCADFA55" blockId="29.[151,1437,1254,1999]" box="[151,887,1399,1423]" pageId="29" pageNumber="330">
<collectingRegion id="4988C4B9E839FFCFAAE4FAEFFE57FA55" box="[151,397,1399,1423]" country="Indonesia" name="Sulawesi Utara" pageId="29" pageNumber="330">
<collectionCode id="ED5D929EE839FFCFAAE4FAEFFE57FA55" box="[151,397,1399,1423]" pageId="29" pageNumber="330">NORTH SULAWESI</collectionCode>
</collectingRegion>
:
<collectionCode id="ED5D929EE839FFCFABE4FAEFFCADFA55" box="[407,887,1399,1423]" pageId="29" pageNumber="330">MOUNT AMBANG NATURE RESERVE</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE839FFCFAAE4FA03FDEAFA0C" blockId="29.[151,1437,1254,1999]" pageId="29" pageNumber="330">
- Kotamobagu (
<quantity id="4CB4A7BEE839FFCFAB1FFA03FE63FA69" box="[364,441,1434,1459]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="29" pageNumber="330" unit="km" value="20.0">20 km</quantity>
E of),
<quantity id="4CB4A7BEE839FFCFA865FA03FDABFA68" box="[534,625,1435,1459]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="29" pageNumber="330" unit="m" value="1120.0">1120 m</quantity>
,
<date id="FFF22C9BE839FFCFA8F1FA02FCC2FA69" box="[642,792,1434,1459]" pageId="29" pageNumber="330" value="1985-09-09" valueMax="1985-09-13" valueMin="1985-09-09">9–13.ix.1985</date>
, J. Huijbregts, hh443a, multistratal evergreen forest, 2 human excrement traps,
<quantity id="4CB4A7BEE839FFCFABD9FA27FE0FFA0D" box="[426,469,1470,1495]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="29" pageNumber="330" unit="in" value="2.0">2 in</quantity>
<collectionCode id="ED5D929EE839FFCFABAFFA27FDEAFA0C" box="[476,560,1471,1494]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="29" pageNumber="330">RMNH</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE839FFCFAAE4FA7BFEC7F9C4" blockId="29.[151,1437,1254,1999]" pageId="29" pageNumber="330">
- Kotamobagu, (
<quantity id="4CB4A7BEE839FFCFAB1EFA7BFE6FFA21" box="[365,437,1506,1531]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="29" pageNumber="330" unit="km" value="20.0">20 km</quantity>
E of),
<quantity id="4CB4A7BEE839FFCFA872FA7BFD8DFA20" box="[513,599,1507,1531]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="29" pageNumber="330" unit="m" value="1200.0">1200 m</quantity>
,
<date id="FFF22C9BE839FFCFA810FA7AFD31FA21" box="[611,747,1506,1531]" pageId="29" pageNumber="330" value="1985-11-05" valueMax="1985-11-08" valueMin="1985-11-05">5–8.xi.1985</date>
, J. Krikken, pw12b, multistratal evergreen forest, 2 fish traps,
<quantity id="4CB4A7BEE839FFCFAAE4F99FFF18F9C5" box="[151,194,1542,1567]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="29" pageNumber="330" unit="in" value="1.0">1 in</quantity>
<collectionCode id="ED5D929EE839FFCFAABAF99FFEC7F9C4" box="[201,285,1543,1566]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="29" pageNumber="330">RMNH</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE839FFCFAAB4F9B2FDAAF815" blockId="29.[151,1437,1254,1999]" pageId="29" pageNumber="330">
<emphasis id="B938D649E839FFCFAAB4F9B2FE60F999" bold="true" box="[199,442,1578,1603]" pageId="29" pageNumber="330">Diagnostic remarks.</emphasis>
For tentative group diagnosis, see
<bibRefCitation id="EFDD77AAE839FFCFA923F9B2FB74F999" author="Krikken" box="[848,1198,1578,1603]" pageId="29" pageNumber="330" refString="Krikken, J. &amp; Huijbregts, J. (2011 a) New Southeast Asian Onthophagus species in the deflexicollis group (Coleoptera: Scarabaeidae: Scarabaeinae). Tijdschrift voor Entomologie, 154, 203 - 213." type="journal article" year="2011" yearSuffix="a">Krikken &amp; Huijbregts (2011a)</bibRefCitation>
. This new species is characterized by the combination of the three long projections on its clypeus (in major males) and the matt, asperate elytral microsculpture with associated more-or-less claviform stubbles. In this size class (body length roughly
<quantity id="4CB4A7BEE839FFCFAA8EF90EFEBEF975" box="[253,356,1686,1711]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="29" pageNumber="330" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
) some other Oriental-Papuasian
<taxonomicName id="4C4C71D8E839FFCFA887F90EFC51F974" box="[756,907,1686,1710]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="330" phylum="Arthropoda" rank="genus">
<emphasis id="B938D649E839FFCFA887F90EFC51F974" box="[756,907,1686,1710]" italics="true" pageId="29" pageNumber="330">Onthophagus</emphasis>
</taxonomicName>
with a T-shaped projection or an elongate anteromedian clypeal plate have recently been placed in an operational group with
<taxonomicName id="4C4C71D8E839FFCFAE01F923FE8EF92D" authority="Lansberge, 1883" authorityName="Lansberge" authorityYear="1883" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="deflexicollis">
<emphasis id="B938D649E839FFCFAE01F923FA41F909" box="[1138,1435,1723,1747]" italics="true" pageId="29" pageNumber="330">Onthophagus deflexicollis</emphasis>
<bibRefCitation id="EFDD77AAE839FFCFAAE4F946FE8EF92D" author="Lansberge" box="[151,340,1758,1783]" pageId="29" pageNumber="330" refString="Lansberge, J. W. van (1883) Revision des Onthophagus de l'archipel indo-neerlandaise, avec description des especes nouvelles. Notes Leyden Museum, 5, 41 - 82." type="journal article" year="1883">Lansberge, 1883</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFDD77AAE839FFCFAB16F946FD69F92D" author="Krikken" box="[357,691,1758,1783]" pageId="29" pageNumber="330" refString="Krikken, J. &amp; Huijbregts, J. (2011 a) New Southeast Asian Onthophagus species in the deflexicollis group (Coleoptera: Scarabaeidae: Scarabaeinae). Tijdschrift voor Entomologie, 154, 203 - 213." type="journal article" year="2011" yearSuffix="a">Krikken &amp; Huijbregts 2011a</bibRefCitation>
, see also group
<quantity id="4CB4A7BEE839FFCFA901F947FC75F92D" box="[882,943,1758,1783]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="29" pageNumber="330" unit="in" value="14.0">14 in</quantity>
<bibRefCitation id="EFDD77AAE839FFCFA9CAF947FB5BF92D" author="Boucomont" box="[953,1153,1759,1783]" pageId="29" pageNumber="330" refString="Boucomont, A. (1914) Les Coprophages de l'Archipel malais (Coleopt.). Annales de la Societe Entomologique de France, 73, 238 - 350." type="journal article" year="1914">Boucomont 1914</bibRefCitation>
).
<taxonomicName id="4C4C71D8E839FFCFAEE4F946FE2DF8C1" authority="Huijbregts &amp; Krikken, 2012" authorityName="Huijbregts &amp; Krikken" authorityYear="2012" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="alfuricus">
<emphasis id="B938D649E839FFCFAEE4F946FA46F92C" box="[1175,1436,1758,1782]" italics="true" pageId="29" pageNumber="330">Onthophagus alfuricus</emphasis>
Huijbregts &amp; Krikken, 2012
</taxonomicName>
, which has conspicuous anterolateral clypeal lobes similar to those in
<taxonomicName id="4C4C71D8E839FFCFAFF2F89BFEECF8E4" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="begoniophilus">
<emphasis id="B938D649E839FFCFAFF2F89BFEECF8E4" italics="true" pageId="29" pageNumber="330">O. begoniophilus</emphasis>
</taxonomicName>
, was tentatively added to this operational group, and occurs in the nearby Moluccas—this species, contrary to our present novelty, is smooth and shiny. A more distant “giant” Oriental congener (up to approximately
<quantity id="4CB4A7BEE839FFCFAB30F8F7FE4FF85C" box="[323,405,1903,1927]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="29" pageNumber="330" unit="mm" value="13.0">13 mm</quantity>
long) with similarly expanded clypeal lobes is
<taxonomicName id="4C4C71D8E839FFCFA9B1F8F6FEFDF871" authority="Arrow, 1910" authorityName="Arrow" authorityYear="1910" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="egregius" subGenus="Parascatonomus">
<emphasis id="B938D649E839FFCFA9B1F8F6FB83F85C" box="[962,1113,1902,1926]" italics="true" pageId="29" pageNumber="330">Onthophagus</emphasis>
(
<emphasis id="B938D649E839FFCFAE1FF8F7FAF2F85C" box="[1132,1320,1903,1926]" italics="true" pageId="29" pageNumber="330">Parascatonomus</emphasis>
)
<emphasis id="B938D649E839FFCFAF4FF8F7FA46F85C" box="[1340,1436,1903,1926]" italics="true" pageId="29" pageNumber="330">egregius</emphasis>
Arrow, 1910
</taxonomicName>
, ranging from
<collectingRegion id="4988C4B9E839FFCFABBEF80BFE25F870" box="[461,511,1939,1962]" country="Indonesia" name="Jawa Timur" pageId="29" pageNumber="330">Java</collectingRegion>
and Borneo to Indochina, but this species is certainly not closely related to any of the various smaller forms mentioned here.
</paragraph>
</subSubSection>
<caption id="DF335AD3E83AFFCCAAE4F978FEABF836" httpUri="https://zenodo.org/record/345766/files/figure.png" pageId="30" pageNumber="331" targetBox="[253,1344,205,1735]" targetPageId="30">
<paragraph id="8BF30A5BE83AFFCCAAE4F978FEABF836" blockId="30.[151,1436,1760,2028]" pageId="30" pageNumber="331">
<emphasis id="B938D649E83AFFCCAAE4F978FE8EF92C" bold="true" box="[151,340,1760,1783]" pageId="30" pageNumber="331">FIGURES 68–81.</emphasis>
Sulawesi
<taxonomicName id="4C4C71D8E83AFFCCABCEF979FD9FF92D" box="[445,581,1761,1783]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="331" phylum="Arthropoda" rank="genus">
<emphasis id="B938D649E83AFFCCABCEF979FD9FF92D" box="[445,581,1761,1783]" italics="true" pageId="30" pageNumber="331">Onthophagus</emphasis>
</taxonomicName>
body parts. 68,
<taxonomicName id="4C4C71D8E83AFFCCA89FF97AFC8AF92D" box="[748,848,1761,1783]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="331" phylum="Arthropoda" rank="species" species="seseba">
<emphasis id="B938D649E83AFFCCA89FF97AFC8AF92D" box="[748,848,1761,1783]" italics="true" pageId="30" pageNumber="331">O. seseba</emphasis>
</taxonomicName>
, head full-face, holotype male from Batui. 69,
<taxonomicName id="4C4C71D8E83AFFCCAF41F97AFA4CF92D" box="[1330,1430,1761,1783]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="331" phylum="Arthropoda" rank="species" species="seseba">
<emphasis id="B938D649E83AFFCCAF41F97AFA4CF92D" box="[1330,1430,1761,1783]" italics="true" pageId="30" pageNumber="331">O. seseba</emphasis>
</taxonomicName>
, left elytron dorsal, holotype male from Batui. 70,
<taxonomicName id="4C4C71D8E83AFFCCA8D1F898FCDCF8CF" box="[674,774,1791,1813]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="331" phylum="Arthropoda" rank="species" species="seseba">
<emphasis id="B938D649E83AFFCCA8D1F898FCDCF8CF" box="[674,774,1791,1813]" italics="true" pageId="30" pageNumber="331">O. seseba</emphasis>
</taxonomicName>
, protibial upper side, holotype male from Batui. 71,
<taxonomicName id="4C4C71D8E83AFFCCAF42F898FA4CF8CF" box="[1329,1430,1791,1813]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="331" phylum="Arthropoda" rank="species" species="seseba">
<emphasis id="B938D649E83AFFCCAF42F898FA4CF8CF" box="[1329,1430,1791,1813]" italics="true" pageId="30" pageNumber="331">O. seseba</emphasis>
</taxonomicName>
, metatibial upperside, holotype male from Batui. 72,
<taxonomicName id="4C4C71D8E83AFFCCA8DBF887FCD1F8EE" box="[680,779,1822,1844]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="331" phylum="Arthropoda" rank="species" species="seseba">
<emphasis id="B938D649E83AFFCCA8DBF887FCD1F8EE" box="[680,779,1822,1844]" italics="true" pageId="30" pageNumber="331">O. seseba</emphasis>
</taxonomicName>
, aedeagus lateral, holotype male from Batui. 73,
<taxonomicName id="4C4C71D8E83AFFCCAE8EF887FA85F8EE" box="[1277,1375,1822,1844]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="331" phylum="Arthropoda" rank="species" species="seseba">
<emphasis id="B938D649E83AFFCCAE8EF887FA85F8EE" box="[1277,1375,1822,1844]" italics="true" pageId="30" pageNumber="331">O. seseba</emphasis>
</taxonomicName>
, head full-face, paratype female. 72,
<taxonomicName id="4C4C71D8E83AFFCCABABF8A6FD53F889" box="[472,649,1853,1875]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="331" phylum="Arthropoda" rank="species" species="begoniophilus">
<emphasis id="B938D649E83AFFCCABABF8A6FD53F889" box="[472,649,1853,1875]" italics="true" pageId="30" pageNumber="331">O. begoniophilus</emphasis>
</taxonomicName>
, aedeagus lateral, holotype from Ambang. 73,
<taxonomicName id="4C4C71D8E83AFFCCAE04F8A6FAF2F889" box="[1143,1320,1853,1875]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="331" phylum="Arthropoda" rank="species" species="begoniophilus">
<emphasis id="B938D649E83AFFCCAE04F8A6FAF2F889" box="[1143,1320,1853,1875]" italics="true" pageId="30" pageNumber="331">O. begoniophilus</emphasis>
</taxonomicName>
, head fullface, holotype from Ambang. 74,
<taxonomicName id="4C4C71D8E83AFFCCAB80F8C4FD78F8A8" box="[499,674,1884,1906]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="331" phylum="Arthropoda" rank="species" species="begoniophilus">
<emphasis id="B938D649E83AFFCCAB80F8C4FD78F8A8" box="[499,674,1884,1906]" italics="true" pageId="30" pageNumber="331">O. begoniophilus</emphasis>
</taxonomicName>
, head full-face, holotype from Ambang. 75,
<taxonomicName id="4C4C71D8E83AFFCCAE1DF8C4FAC7F8A8" box="[1134,1309,1884,1906]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="331" phylum="Arthropoda" rank="species" species="begoniophilus">
<emphasis id="B938D649E83AFFCCAE1DF8C4FAC7F8A8" box="[1134,1309,1884,1906]" italics="true" pageId="30" pageNumber="331">O. begoniophilus</emphasis>
</taxonomicName>
, left elytron dorsal, holotype from Ambang. 76,
<taxonomicName id="4C4C71D8E83AFFCCA876F8E3FD6EF84A" box="[517,692,1914,1936]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="331" phylum="Arthropoda" rank="species" species="begoniophilus">
<emphasis id="B938D649E83AFFCCA876F8E3FD6EF84A" box="[517,692,1914,1936]" italics="true" pageId="30" pageNumber="331">O. begoniophilus</emphasis>
</taxonomicName>
, head full-face, paratype male. 77,
<taxonomicName id="4C4C71D8E83AFFCCAE6CF8E3FB14F84A" box="[1055,1230,1914,1936]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="331" phylum="Arthropoda" rank="species" species="begoniophilus">
<emphasis id="B938D649E83AFFCCAE6CF8E3FB14F84A" box="[1055,1230,1914,1936]" italics="true" pageId="30" pageNumber="331">O. begoniophilus</emphasis>
</taxonomicName>
, left elytron caudal, paratype male. 78,
<taxonomicName id="4C4C71D8E83AFFCCAB29F802FDD3F875" box="[346,521,1945,1967]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="331" phylum="Arthropoda" rank="species" species="begoniophilus">
<emphasis id="B938D649E83AFFCCAB29F802FDD3F875" box="[346,521,1945,1967]" italics="true" pageId="30" pageNumber="331">O. begoniophilus</emphasis>
</taxonomicName>
, protibial upper side, paratype male. 79,
<taxonomicName id="4C4C71D8E83AFFCCA9DFF802FB81F875" box="[940,1115,1945,1967]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="331" phylum="Arthropoda" rank="species" species="begoniophilus">
<emphasis id="B938D649E83AFFCCA9DFF802FB81F875" box="[940,1115,1945,1967]" italics="true" pageId="30" pageNumber="331">O. begoniophilus</emphasis>
</taxonomicName>
, metatibial upperside, paratype male. 80,
<taxonomicName id="4C4C71D8E83AFFCCAA8FF820FE70F817" box="[252,426,1975,1997]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="331" phylum="Arthropoda" rank="species" species="begoniophilus">
<emphasis id="B938D649E83AFFCCAA8FF820FE70F817" box="[252,426,1975,1997]" italics="true" pageId="30" pageNumber="331">O. begoniophilus</emphasis>
</taxonomicName>
, aedeagus lateral, paratype male. 81,
<taxonomicName id="4C4C71D8E83AFFCCA959F820FC02F817" box="[810,984,1975,1997]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="331" phylum="Arthropoda" rank="species" species="begoniophilus">
<emphasis id="B938D649E83AFFCCA959F820FC02F817" box="[810,984,1975,1997]" italics="true" pageId="30" pageNumber="331">O. begoniophilus</emphasis>
</taxonomicName>
, head full-face, paratype female. Scale lines approximately 1 mm.
</paragraph>
</caption>
<subSubSection id="C35659D0E83BFFF2AAB4FF0FFEB8FD2A" lastPageId="32" lastPageNumber="333" pageId="31" pageNumber="332" type="description">
<paragraph id="8BF30A5BE83BFFCDAAB4FF0FFDCDFF22" blockId="31.[151,1437,151,789]" pageId="31" pageNumber="332">
<emphasis id="B938D649E83BFFCDAAB4FF0FFE83FF6A" bold="true" box="[199,345,151,176]" pageId="31" pageNumber="332">Description.</emphasis>
<typeStatus id="54F7B4F9E83BFFCDAB1BFF0FFE09FF6A" box="[360,467,151,176]" pageId="31" pageNumber="332" type="holotype">Holotype</typeStatus>
, male. Body length approximately
<quantity id="4CB4A7BEE83BFFCDA9FAFF00FC31FF75" box="[905,1003,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="31" pageNumber="332" unit="mm" value="4.8">4.8 mm</quantity>
. Black-brown, largely matt, due to microreticulation (
<date id="FFF22C9BE83BFFCDAB1AFF24FE4CFF0E" box="[361,406,188,212]" pageId="31" pageNumber="332" value="1980-10">x80</date>
) and other microsculpture. Pilosity abundant, mainly pale yellow, nearly all body bristles characteristically claviform, short.
</paragraph>
<paragraph id="8BF30A5BE83BFFCDAAB4FE9CFAF7FE76" blockId="31.[151,1437,151,789]" pageId="31" pageNumber="332">Clypeal apex produced into long, slender, nearly T-shaped, reflexed horn; clypeus on either side produced into apically angulate, anteriorly directed, reflexed lobe; genal border obtusely angular halfway; clypeogenal transition (in this major male) virtually continuous; frontovertex without any protrusions; clypeal surface anteromedially impunctate, shiny; remainder of head densely to crowdedly (sub)ocellate-punctate, matt. Eyes with approximately 6 facet rows across widest point; ratio interocular distance / single maximum eye width approximately 9.</paragraph>
<paragraph id="8BF30A5BE83BFFCDAAB4FE20FF12FDBA" blockId="31.[151,1437,151,789]" pageId="31" pageNumber="332">Pronotum moderately, evenly convex, discal midline not impressed; anterior section of lateral border slightly, widely round; anterolateral margin slightly reflexed, angle rounded off, approximately 90°, posterior section slightly sinuate; pronotal base medially virtually immarginate; pronotal derm with crowded large, elongate-elliptic, superficial, (sub)ocellate, punctures (many bristle-bearing); punctures mostly separated by 0.5 their diameter or less.</paragraph>
<paragraph id="8BF30A5BE83BFFCDAAB4FDF4FAD5FCCF" blockId="31.[151,1437,151,789]" pageId="31" pageNumber="332">
Elytra brown, lighter (rufous) on basal parts; striae discally well defined, with distinct punctures, mainly separated by 2–4 times their diameter, crenulate on interstriae; interstriae 1–6 slightly generally slightly convex on disc, distinctly raised on steeper distal and lateral declivities (particularly on humeral area); punctures of lateral striae deeply impressed; all interstriae with irregular rows of bristle-bearing microgranules (laterally strongly asperate); interstrial derm irregularly microreticulate (
<date id="FFF22C9BE83BFFCDA887FD64FCF8FCCE" box="[756,802,764,788]" pageId="31" pageNumber="332" value="1980-10">x80</date>
); slight intrahumeral impression noticeable.
</paragraph>
<caption id="DF335AD3E83BFFCDAAE4F8C9FDCFF819" httpUri="https://zenodo.org/record/345767/files/figure.png" pageId="31" pageNumber="332" targetBox="[206,1436,841,1851]" targetPageId="31">
<paragraph id="8BF30A5BE83BFFCDAAE4F8C9FDCFF819" blockId="31.[151,1436,1873,1987]" pageId="31" pageNumber="332">
<emphasis id="B938D649E83BFFCDAAE4F8C9FEFEF8BC" bold="true" box="[151,292,1873,1895]" pageId="31" pageNumber="332">FIGURE 82.</emphasis>
Map of Sulawesi and nearby islands with indication of collecting regions mentioned in Material examined sections. In North-South arrangement, as follows: Sangihe Islands, Tangkoko Nature Reserve, Manado Region, Mount Ambang Nature Reserve, Dumoga-Bone National Park, Palu Region, Lore-Lindu National Park, Banggai District, Sula Islands, Kendari District, and Kolaka District.
</paragraph>
</caption>
<paragraph id="8BF30A5BE804FFF2AAB4FF0FFCEDFEC7" blockId="32.[151,1437,151,789]" pageId="32" pageNumber="333">Antennal club yellow. Metaventrite with vague discal midline impression; most of derm with relatively large, bristle-bearing, ocellate punctation. Abdominal ventrites brown, each with transverse row of bristle-bearing, ocellate punctures. Pygidium slightly convex, matt, brown; virtually crowded with large, superficial, bristlebearing, (sub)ocellate punctures; base with transverse ridge.</paragraph>
<paragraph id="8BF30A5BE804FFF2AAB4FEB0FE2CFE77" blockId="32.[151,1437,151,789]" pageId="32" pageNumber="333">
Legs brown. Protibia with 3+(1) larger denticles, spur elongate-acuminate (apex worn in
<typeStatus id="54F7B4F9E804FFF2AED4FEBFFAD4FE9A" box="[1191,1294,295,320]" pageId="32" pageNumber="333" type="holotype">holotype</typeStatus>
). Mesotibiae and metatibiae slender, spurs unmodified. Femoral undersides with numerous bristle-bearing punctures. Metarsomere 1 robust, almost as long as metatarsomeres 2–4 combined; approximate proportions of spur and tarsomeres 1–5: 10//
<date id="FFF22C9BE804FFF2AB0DFE0CFE1CFE77" box="[382,454,404,429]" pageId="32" pageNumber="333" value="2015-04-07">15/7/4</date>
/3/5.
</paragraph>
<paragraph id="8BF30A5BE804FFF2AAB4FE2FFDCEFE0A" blockId="32.[151,1437,151,789]" box="[199,532,439,464]" pageId="32" pageNumber="333">
Aedeagus,
<figureCitation id="137716DEE804FFF2AB30FE2FFE48FE0A" box="[323,402,439,464]" captionStart="FIGURES 68 – 81" captionStartId="30.[151,264,1760,1783]" captionTargetBox="[253,1344,205,1735]" captionTargetId="figure@30.[239,1344,191,1735]" captionTargetPageId="30" captionText="FIGURES 68 – 81. Sulawesi Onthophagus body parts. 68, O. seseba, head full-face, holotype male from Batui. 69, O. seseba, left elytron dorsal, holotype male from Batui. 70, O. seseba, protibial upper side, holotype male from Batui. 71, O. seseba, metatibial upperside, holotype male from Batui. 72, O. seseba, aedeagus lateral, holotype male from Batui. 73, O. seseba, head full-face, paratype female. 72, O. begoniophilus, aedeagus lateral, holotype from Ambang. 73, O. begoniophilus, head fullface, holotype from Ambang. 74, O. begoniophilus, head full-face, holotype from Ambang. 75, O. begoniophilus, left elytron dorsal, holotype from Ambang. 76, O. begoniophilus, head full-face, paratype male. 77, O. begoniophilus, left elytron caudal, paratype male. 78, O. begoniophilus, protibial upper side, paratype male. 79, O. begoniophilus, metatibial upperside, paratype male. 80, O. begoniophilus, aedeagus lateral, paratype male. 81, O. begoniophilus, head full-face, paratype female. Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345766/files/figure.png" pageId="32" pageNumber="333">Fig. 80</figureCitation>
(
<typeStatus id="54F7B4F9E804FFF2ABD2FE20FDD2FE0A" box="[417,520,440,464]" pageId="32" pageNumber="333" type="paratype">paratype</typeStatus>
).
</paragraph>
<paragraph id="8BF30A5BE804FFF2AAB4FE45FD66FDC2" blockId="32.[151,1437,151,789]" pageId="32" pageNumber="333">Measurements in mm. Maximum width of head 1.4. Median length of pronotum 1.3, maximum width 2.3. Sutural length of elytra 1.9, maximum width 2.5.</paragraph>
<paragraph id="8BF30A5BE804FFF2AAB4FDBCFBEDFD7D" blockId="32.[151,1437,151,789]" pageId="32" pageNumber="333">
<emphasis id="B938D649E804FFF2AAB4FDBCFDBBFDE7" bold="true" box="[199,609,548,573]" pageId="32" pageNumber="333">Variation and sexual dimorphism.</emphasis>
Body length roughly
<quantity id="4CB4A7BEE804FFF2A928FDBCFC66FDE6" box="[859,956,548,573]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="32" pageNumber="333" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
. Shape of male head very polymorphic. A minor male (length
<quantity id="4CB4A7BEE804FFF2ABFAFDD0FE3CFDBA" box="[393,486,584,608]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="32" pageNumber="333" unit="mm" value="3.9">3.9 mm</quantity>
) has the clypeal projections poorly developed (less elongate), the apicomedian projection being simply elongate-lobate (tip simply truncate, not T-shaped); the male
<typeStatus id="54F7B4F9E804FFF2AE1EFDF5FB17FD5F" box="[1133,1229,621,645]" pageId="32" pageNumber="333" type="paratype">paratype</typeStatus>
illustrated here is intermediate. Their clypeogenal edge is at most shallowly interrupted at the suture.
</paragraph>
<paragraph id="8BF30A5BE804FFF2AAB4FD2CFEB8FD2A" blockId="32.[151,1437,151,789]" pageId="32" pageNumber="333">
<emphasis id="B938D649E804FFF2AAB4FD2CFEC6FD16" box="[199,284,692,716]" italics="true" pageId="32" pageNumber="333">Female</emphasis>
has clypeal border anteromedially simply shortly bidentate-sinuate; marginal surface of clypeus more rugulate-punctate.
</paragraph>
</subSubSection>
<subSubSection id="C35659D0E804FFF2AAB4FD64FA47FCCF" box="[199,1437,764,789]" pageId="32" pageNumber="333" type="etymology">
<paragraph id="8BF30A5BE804FFF2AAB4FD64FA47FCCF" blockId="32.[151,1437,151,789]" box="[199,1437,764,789]" pageId="32" pageNumber="333">
<emphasis id="B938D649E804FFF2AAB4FD64FE94FCCF" bold="true" box="[199,334,764,789]" pageId="32" pageNumber="333">Etymology.</emphasis>
Named after the begonia flowers on the floor of the upland forest in which this species was found.
</paragraph>
</subSubSection>
</treatment>
</document>
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