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<document id="4E6B7F627DC087C8F850346AB014012A" ID-CLB-Dataset="55542" ID-DOI="10.11646/zootaxa.5134.4.4" ID-GBIF-Dataset="074aea58-8df7-44e0-a546-1663ade77a3f" ID-ISSN="1175-5326" ID-Zenodo-Dep="6543169" ID-ZooBank="1CDBA232-1CBB-44F7-895F-11A069652242" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1652342946175" checkinUser="plazi" docAuthor="Okajima, Shuji &amp; Masumoto, Masami" docDate="2022" docId="03E4B36E8F2BFFF2FF0141B6FEC383A5" docLanguage="en" docName="zootaxa.5134.4.4.pdf" docOrigin="Zootaxa 5134 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nazonothrips toshifumii Okajima &amp; Masumoto 2022, sp. n." docType="treatment" docVersion="7" lastPageNumber="565" masterDocId="FFDDCB168F28FFF6FF964338FF83816E" masterDocTitle="Enigmatic male dimorphism in the Phlaeothripinae (Thysanoptera, Phlaeothripidae), with description of a new genus and species" masterLastPageNumber="568" masterPageNumber="561" pageNumber="564" updateTime="1699330271474" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E8D1F16F449C033BD93B8C5460951DE2">Enigmatic male dimorphism in the Phlaeothripinae (Thysanoptera, Phlaeothripidae), with description of a new genus and species</mods:title>
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<mods:namePart id="B116AB81EED9734280B28E0782252058">Okajima, Shuji</mods:namePart>
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<mods:affiliation id="BC8BBE54B69A363E236FDA66AD3B2A82">Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. &amp; 7 okajimas 2 @ gmail. com; https: // orcid. org / 0000 - 0001 - 7249 - 671 X</mods:affiliation>
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<mods:namePart id="B4621417DB918BB53766F806E0FF82E6">Masumoto, Masami</mods:namePart>
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<mods:affiliation id="EEE695B7E32ED8C9D53228D416A81C85">Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan. &amp; masumotoms @ gmail. com; https: // orcid. org / 0000 - 0001 - 9049 - 2448</mods:affiliation>
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<treatment id="03E4B36E8F2BFFF2FF0141B6FEC383A5" ID-DOI="http://doi.org/10.5281/zenodo.6543104" ID-GBIF-Taxon="195304239" ID-Zenodo-Dep="6543104" LSID="urn:lsid:plazi:treatment:03E4B36E8F2BFFF2FF0141B6FEC383A5" httpUri="http://treatment.plazi.org/id/03E4B36E8F2BFFF2FF0141B6FEC383A5" lastPageId="4" lastPageNumber="565" pageId="3" pageNumber="564">
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<heading id="D0BAB5148F2BFFF5FF0141B6FD8483C7" bold="true" box="[151,519,654,681]" fontSize="11" level="1" pageId="3" pageNumber="564" reason="1">
<emphasis id="B939DE6A8F2BFFF5FF0141B6FD8483C7" bold="true" box="[151,519,654,681]" pageId="3" pageNumber="564">
<taxonomicName id="4C4D79FB8F2BFFF5FF0141B6FE3D83C7" authority="Okajima &amp; Masumoto, 2022" authorityName="Okajima &amp; Masumoto" authorityYear="2022" box="[151,446,654,681]" class="Insecta" family="Phlaeothripidae" genus="Nazonothrips" kingdom="Animalia" order="Thysanoptera" pageId="3" pageNumber="564" phylum="Arthropoda" rank="species" species="toshifumii" status="sp. nov.">
<emphasis id="B939DE6A8F2BFFF5FF0141B6FE3D83C7" bold="true" box="[151,446,654,681]" italics="true" pageId="3" pageNumber="564">Nazonothrips toshifumii</emphasis>
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<taxonomicNameLabel id="A20A63118F2BFFF5FE5341B7FD8483C7" box="[453,519,655,681]" pageId="3" pageNumber="564" rank="species">sp. n.</taxonomicNameLabel>
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<subSubSection id="C35751F38F2BFFF2FF01418BFC5883EC" lastPageId="4" lastPageNumber="565" pageId="3" pageNumber="564" type="description">
<paragraph id="8BF202788F2BFFF5FF01418BFE9A83A3" blockId="3.[151,519,654,717]" box="[151,281,691,717]" pageId="3" pageNumber="564">
(
<figureCitation id="13761EFD8F2BFFF5FF09418BFE9183A3" box="[159,274,691,717]" captionStart-0="FIGURES 213" captionStart-1="FIGURES 1420" captionStart-2="FIGURES 2124" captionStartId-0="5.[151,264,1914,1938]" captionStartId-1="6.[151,264,1610,1634]" captionStartId-2="7.[151,264,1936,1960]" captionTargetBox-0="[209,1378,201,1890]" captionTargetBox-1="[153,1434,181,1586]" captionTargetBox-2="[202,1384,181,1912]" captionTargetId-0="figure-17@5.[209,1378,201,1890]" captionTargetId-1="figure-24@6.[153,1434,181,1586]" captionTargetId-2="figure-16@7.[202,1384,181,1912]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURES 213. Nazonothrips toshifumii gen. et sp. n., female, 23. (2) Macroptera; (3) Aptera. Gynecoid male aptera, 45. (4) Large individual; (5) Small individual.Allometric male aptera, 67. (6) Large individual; (7) Small individual. Female macroptera, 813. (8) Head &amp; pronotum; (9) Antenna; (10) Mesonotumpelta; (11) Pro-metasterna; (12) Tergite V; (13) Tergite IX &amp; tube." captionText-1="FIGURES 1420. Nazonothrips toshifumii gen. et sp. n., headpelta, 1416. (14) Female aptera; (15) Gynecoid male aptera, large individual; (16) Gynecoid male aptera, small individual. Pro-metasterna, 1718. (17) Female aptera; (18) Gynecoid male aptera. Fore leg, 1920. (19) Female aptera; (20) Gynecoid male aptera." captionText-2="FIGURES 2124. Nazonothrips toshifumii gen. et sp. n., allometric male aptera. (21) Headpelta, small individual; (22) Head pronotum, large individual; (23) Pro-metasterna; (24) Fore leg." figureDoi-0="http://doi.org/10.5281/zenodo.6543176" figureDoi-1="http://doi.org/10.5281/zenodo.6543182" figureDoi-2="http://doi.org/10.5281/zenodo.6543190" httpUri-0="https://zenodo.org/record/6543176/files/figure.png" httpUri-1="https://zenodo.org/record/6543182/files/figure.png" httpUri-2="https://zenodo.org/record/6543190/files/figure.png" pageId="3" pageNumber="564">Figs 224</figureCitation>
)
</paragraph>
<paragraph id="8BF202788F2BFFF5FF0141CFFD388467" blockId="3.[151,1437,759,2009]" pageId="3" pageNumber="564">
<emphasis id="B939DE6A8F2BFFF5FF0141CFFEFA827E" box="[151,377,759,785]" italics="true" pageId="3" pageNumber="564">Female macroptera</emphasis>
(
<figureCitation id="13761EFD8F2BFFF5FE1C41CFFD9D827F" box="[394,542,759,785]" captionStart="FIGURES 213" captionStartId="5.[151,264,1914,1938]" captionTargetBox="[209,1378,201,1890]" captionTargetId="figure-17@5.[209,1378,201,1890]" captionTargetPageId="5" captionText="FIGURES 213. Nazonothrips toshifumii gen. et sp. n., female, 23. (2) Macroptera; (3) Aptera. Gynecoid male aptera, 45. (4) Large individual; (5) Small individual.Allometric male aptera, 67. (6) Large individual; (7) Small individual. Female macroptera, 813. (8) Head &amp; pronotum; (9) Antenna; (10) Mesonotumpelta; (11) Pro-metasterna; (12) Tergite V; (13) Tergite IX &amp; tube." figureDoi="http://doi.org/10.5281/zenodo.6543176" httpUri="https://zenodo.org/record/6543176/files/figure.png" pageId="3" pageNumber="564">Figs 2, 813</figureCitation>
). Body brown; abdominal segment IX and tube yellowish. Antennae yellowish brown, somewhat paler than head, segment II slightly paler than other segments. Fore femora yellowish, with bases brownish; mid and hind femora brown; tibiae yellowish, mid and hind tibiae shaded with brown. Fore wings, remaining part, weakly shaded. Prominent setae pale. Head wider than long, dorsal surface smooth; cheeks short and weekly narrowed toward base. Postocular setae situated on cheeks just behind eyes, short and nearly pointed; all of other cephalic setae minute. Eyes large and bulged, about 0.7 times as long as head, smaller on ventral surface. Ocelli largely developed; anterior ocellus 13µm in diameter, 20µm apart from posterior pair; posterior ocelli recessed into eyes, 2022µm in diameter, 15µm apart from each other. Antennal segment III a little smaller than segment IV; segment VIII about 3.0 times as long as wide; segments III and IV with two (1 + 1) and four (2 + 2) sense-cones, respectively. Pronotum shield comparatively weak, a little shorter than head, much wider than long, margins weakly eroded; anteroangular and posterosngulsr setae pointed. Ferna eroded marginally. Mesoeusternum largely eroded anteromedially, this eroded area is inverted triangular shape. Mesonotum smooth; metanotum transverse, without reticulation, with a pair of median week setae. Fore tarsal tooth moderately developed. Abdominal tergite IX S1 and S2 setae sharply pointed, S2 longer than S1. Tube about 2/3 as long as head, and 1.15 times as long as wide.
</paragraph>
<paragraph id="8BF202788F2BFFF5FF51462BFB9D84D3" blockId="3.[151,1437,759,2009]" pageId="3" pageNumber="564">
<emphasis id="B939DE6A8F2BFFF5FF51462BFEF58443" bold="true" box="[199,374,1299,1325]" pageId="3" pageNumber="564">Measurements</emphasis>
(
<typeStatus id="54F6BCDA8F2BFFF5FE11462BFE688443" box="[391,491,1299,1325]" pageId="3" pageNumber="564" type="holotype">holotype</typeStatus>
female macroptera in microns). Body length about 1300 (distended). Head length 110, width 125; postocular setae less than 30. Pronotum length 95, width 193; anteroangular setae 4245; posteroangular setae 5557; epimeral setae 4244; coxal setae about 30. Antenna total length 310, segments IVIII length (width) as follows: 35 (27), 35 (29), 36 (32), 40 (33), 40 (30), 37 (23), 37 (22), 38 (12). Setae on tergite IX: S1 7376, S2 90100. Tube length 75, maximum width 65; terminal setae about 40.
</paragraph>
<paragraph id="8BF202788F2BFFF5FF5146FFFCD28747" blockId="3.[151,1437,759,2009]" pageId="3" pageNumber="564">
<emphasis id="B939DE6A8F2BFFF5FF5146FFFEEF848E" box="[199,364,1479,1505]" italics="true" pageId="3" pageNumber="564">Female aptera</emphasis>
(
<figureCitation id="13761EFD8F2BFFF5FEEB46FFFE45848F" box="[381,454,1479,1505]" captionStart="FIGURES 213" captionStartId="5.[151,264,1914,1938]" captionTargetBox="[209,1378,201,1890]" captionTargetId="figure-17@5.[209,1378,201,1890]" captionTargetPageId="5" captionText="FIGURES 213. Nazonothrips toshifumii gen. et sp. n., female, 23. (2) Macroptera; (3) Aptera. Gynecoid male aptera, 45. (4) Large individual; (5) Small individual.Allometric male aptera, 67. (6) Large individual; (7) Small individual. Female macroptera, 813. (8) Head &amp; pronotum; (9) Antenna; (10) Mesonotumpelta; (11) Pro-metasterna; (12) Tergite V; (13) Tergite IX &amp; tube." figureDoi="http://doi.org/10.5281/zenodo.6543176" httpUri="https://zenodo.org/record/6543176/files/figure.png" pageId="3" pageNumber="564">Figs 3</figureCitation>
,
<figureCitation id="13761EFD8F2BFFF5FE4246FFFDC4848F" box="[468,583,1479,1505]" captionStart="FIGURES 1420" captionStartId="6.[151,264,1610,1634]" captionTargetBox="[153,1434,181,1586]" captionTargetId="figure-24@6.[153,1434,181,1586]" captionTargetPageId="6" captionText="FIGURES 1420. Nazonothrips toshifumii gen. et sp. n., headpelta, 1416. (14) Female aptera; (15) Gynecoid male aptera, large individual; (16) Gynecoid male aptera, small individual. Pro-metasterna, 1718. (17) Female aptera; (18) Gynecoid male aptera. Fore leg, 1920. (19) Female aptera; (20) Gynecoid male aptera." figureDoi="http://doi.org/10.5281/zenodo.6543182" httpUri="https://zenodo.org/record/6543182/files/figure.png" pageId="3" pageNumber="564">14, 17, 19</figureCitation>
). Colour similar to macropterous female, but head and antennae somewhat paler, brownish yellow. Head small and square, almost as long as wide. Eyes with about 10 facets. Pronotum almost as long as head. Mesothorax transverse. Pelta strongly eroded.
</paragraph>
<paragraph id="8BF202788F2BFFF5FF51450BFE4D87B3" blockId="3.[151,1437,759,2009]" pageId="3" pageNumber="564">
<emphasis id="B939DE6A8F2BFFF5FF51450BFEF58723" bold="true" box="[199,374,1587,1613]" pageId="3" pageNumber="564">Measurements</emphasis>
(
<typeStatus id="54F6BCDA8F2BFFF5FE13450BFE648723" box="[389,487,1587,1613]" pageId="3" pageNumber="564" type="paratype">paratype</typeStatus>
female aptera in microns). Body length 1220 (distended). Head length 86, width 87: postocular setae?. Pronotum length 85, width 160; anteroangular setae 4550; posteroangular setae 5055; epimeral setae 45; coxal setae 3035. Antennal segments I-VIII length (width) as follows: 33 (27), 37 (27), 27 (22), 30 (28), 33 (26), 32 (23), 31 (20), 38 (12). Setae on tergite IX: S1 75-80, S2 about 90. Tube length 75, maximum width 65; terminal setae about 50.
</paragraph>
<paragraph id="8BF202788F2BFFF5FF5145DFFCDA8603" blockId="3.[151,1437,759,2009]" pageId="3" pageNumber="564">
<emphasis id="B939DE6A8F2BFFF5FF5145DFFE50866E" box="[199,467,1767,1793]" italics="true" pageId="3" pageNumber="564">Allometric male aptera</emphasis>
(
<figureCitation id="13761EFD8F2BFFF5FE7045DFFDD6866F" box="[486,597,1767,1793]" captionStart="FIGURES 213" captionStartId="5.[151,264,1914,1938]" captionTargetBox="[209,1378,201,1890]" captionTargetId="figure-17@5.[209,1378,201,1890]" captionTargetPageId="5" captionText="FIGURES 213. Nazonothrips toshifumii gen. et sp. n., female, 23. (2) Macroptera; (3) Aptera. Gynecoid male aptera, 45. (4) Large individual; (5) Small individual.Allometric male aptera, 67. (6) Large individual; (7) Small individual. Female macroptera, 813. (8) Head &amp; pronotum; (9) Antenna; (10) Mesonotumpelta; (11) Pro-metasterna; (12) Tergite V; (13) Tergite IX &amp; tube." figureDoi="http://doi.org/10.5281/zenodo.6543176" httpUri="https://zenodo.org/record/6543176/files/figure.png" pageId="3" pageNumber="564">Figs. 67</figureCitation>
,
<figureCitation id="13761EFD8F2BFFF5FDF345DFFD2C866F" box="[613,687,1767,1793]" captionStart="FIGURES 2124" captionStartId="7.[151,264,1936,1960]" captionTargetBox="[202,1384,181,1912]" captionTargetId="figure-16@7.[202,1384,181,1912]" captionTargetPageId="7" captionText="FIGURES 2124. Nazonothrips toshifumii gen. et sp. n., allometric male aptera. (21) Headpelta, small individual; (22) Head pronotum, large individual; (23) Pro-metasterna; (24) Fore leg." figureDoi="http://doi.org/10.5281/zenodo.6543190" httpUri="https://zenodo.org/record/6543190/files/figure.png" pageId="3" pageNumber="564">2124</figureCitation>
).
<emphasis id="B939DE6A8F2BFFF5FD5045DEFBEF866F" bold="true" box="[710,1132,1766,1793]" italics="true" pageId="3" pageNumber="564">Prothorax always largely developed</emphasis>
. Head similar to apterous female in small individuals, but longer than wide in large individuals. Prothorax larger than pterothorax, at least wider than metathorax. Fore legs largely developed in large individuals, with stout fore tarsal tooth. Colour similar to apterous female, often somewhat paler abdominal segments.
</paragraph>
<paragraph id="8BF202788F2BFFF2FF51444FFACA81BB" blockId="3.[151,1437,759,2009]" lastBlockId="4.[151,1437,150,715]" lastPageId="4" lastPageNumber="565" pageId="3" pageNumber="564">
<emphasis id="B939DE6A8F2BFFF5FF51444FFEF586FF" bold="true" box="[199,374,1911,1937]" pageId="3" pageNumber="564">Measurements</emphasis>
(
<typeStatus id="54F6BCDA8F2BFFF5FE10444FFE6B86FF" box="[390,488,1911,1937]" pageId="3" pageNumber="564" type="paratype">paratype</typeStatus>
small-large apterous males in microns). Body length 10001300 (distended). Head length 80125, width 8390: postocular setae less than 3050. Pronotum length 140232, width 160-260; anteroangular setae 65105; posteroangular setae 50-90; epimeral setae 50-90; coxal setae 45-80. Antenna total length 250-305, segments IVIII length as follows: 3542, 3040, 2533, 2737, 3236, 3240, 3037, 3441. Setae on tergite IX: S1 70100, S2 75100. Tube length 6580, maximum width 5460; terminal setae about 3530.
</paragraph>
<paragraph id="8BF202788F2CFFF2FF5143E7FB2E8073" blockId="4.[151,1437,150,715]" pageId="4" pageNumber="565">
<emphasis id="B939DE6A8F2CFFF2FF5143E7FE3E8196" box="[199,445,223,249]" italics="true" pageId="4" pageNumber="565">Gynecoid male aptera</emphasis>
(
<figureCitation id="13761EFD8F2CFFF2FE5D43E7FDB68197" box="[459,565,222,249]" captionStart="FIGURES 213" captionStartId="5.[151,264,1914,1938]" captionTargetBox="[209,1378,201,1890]" captionTargetId="figure-17@5.[209,1378,201,1890]" captionTargetPageId="5" captionText="FIGURES 213. Nazonothrips toshifumii gen. et sp. n., female, 23. (2) Macroptera; (3) Aptera. Gynecoid male aptera, 45. (4) Large individual; (5) Small individual.Allometric male aptera, 67. (6) Large individual; (7) Small individual. Female macroptera, 813. (8) Head &amp; pronotum; (9) Antenna; (10) Mesonotumpelta; (11) Pro-metasterna; (12) Tergite V; (13) Tergite IX &amp; tube." figureDoi="http://doi.org/10.5281/zenodo.6543176" httpUri="https://zenodo.org/record/6543176/files/figure.png" pageId="4" pageNumber="565">Figs. 45</figureCitation>
,
<figureCitation id="13761EFD8F2CFFF2FDD643E6FD588197" box="[576,731,222,249]" captionStart="FIGURES 1420" captionStartId="6.[151,264,1610,1634]" captionTargetBox="[153,1434,181,1586]" captionTargetId="figure-24@6.[153,1434,181,1586]" captionTargetPageId="6" captionText="FIGURES 1420. Nazonothrips toshifumii gen. et sp. n., headpelta, 1416. (14) Female aptera; (15) Gynecoid male aptera, large individual; (16) Gynecoid male aptera, small individual. Pro-metasterna, 1718. (17) Female aptera; (18) Gynecoid male aptera. Fore leg, 1920. (19) Female aptera; (20) Gynecoid male aptera." figureDoi="http://doi.org/10.5281/zenodo.6543182" httpUri="https://zenodo.org/record/6543182/files/figure.png" pageId="4" pageNumber="565">1516, 18, 20</figureCitation>
).
<emphasis id="B939DE6A8F2CFFF2FD7B43E6FAC18197" bold="true" box="[749,1346,222,249]" italics="true" pageId="4" pageNumber="565">Prothorax always moderately developed as in female</emphasis>
. Colour and structure similar to female. Prothorax smaller than pterothorax, at least about same sized.
</paragraph>
<paragraph id="8BF202788F2CFFF2FF51421EFB6B80BE" blockId="4.[151,1437,150,715]" pageId="4" pageNumber="565">
<emphasis id="B939DE6A8F2CFFF2FF51421EFEF5802E" bold="true" box="[199,374,294,320]" pageId="4" pageNumber="565">Measurements</emphasis>
(
<typeStatus id="54F6BCDA8F2CFFF2FE1E421EFE69802E" box="[392,490,294,320]" pageId="4" pageNumber="565" type="paratype">paratype</typeStatus>
small-large apterous males in microns). Body length 9801280 (distended). Head length 8295, width 7580: postocular setae less than 3060. Pronotum length 120142, width 155183; anteroangular setae 6090; posteroangular setae 50100; epimeral setae 4058; coxal setae 4045. Antenna total length 250260, segments IVIII length as follows: 3337, 3035, 2527, 2728, 3032, 3033, 3030, 3533. Setae on tergite IX: S1 7595, S2 7790. Tube length 7075, maximum width 5560; terminal setae 2835.
</paragraph>
<paragraph id="8BF202788F2CFFF2FF5142E1FC138354" blockId="4.[151,1437,150,715]" pageId="4" pageNumber="565">
<emphasis id="B939DE6A8F2CFFF2FF5142E1FE26809D" bold="true" box="[199,421,473,499]" pageId="4" pageNumber="565">Specimens studied</emphasis>
.
<materialsCitation id="3B2508258F2CFFF2FE2342E1FE8C8379" ID-GBIF-Occurrence="3766757304" collectingDate="1993-10-10" collectionCode="T" country="Taiwan" location="Taitung-hsien" municipality="Chihpen" pageId="4" pageNumber="565" specimenCount="1" typeStatus="holotype">
<typeStatus id="54F6BCDA8F2CFFF2FE2342E1FD9E809D" box="[437,541,473,499]" pageId="4" pageNumber="565" type="holotype">Holotype</typeStatus>
macropterous female,
<emphasis id="B939DE6A8F2CFFF2FCB242E1FCFF809D" bold="true" box="[804,892,473,499]" pageId="4" pageNumber="565">
<collectingCountry id="F35A42E88F2CFFF2FCB242E1FCFF809D" box="[804,892,473,499]" name="Taiwan" pageId="4" pageNumber="565">Taiwan</collectingCountry>
</emphasis>
,
<location id="8E9254A38F2CFFF2FC1D42E1FBAB809D" LSID="urn:lsid:plazi:treatment:03E4B36E8F2BFFF2FF0141B6FEC383A5:8E9254A38F2CFFF2FC1D42E1FBAB809D" box="[907,1064,473,499]" country="Taiwan" municipality="Chihpen" name="Taitung-hsien" pageId="4" pageNumber="565">Taitung-hsien</location>
,
<collectingMunicipality id="6B9698028F2CFFF2FBA342E1FB14809D" box="[1077,1175,473,499]" pageId="4" pageNumber="565">Chihpen</collectingMunicipality>
, on fungus,
<date id="FFF324B88F2CFFF2FAB342E1FA1B809D" box="[1317,1432,473,499]" pageId="4" pageNumber="565" value="1993-10-10">
<collectingDate id="EFB7DD508F2CFFF2FAB342E1FA1B809D" box="[1317,1432,473,499]" pageId="4" pageNumber="565" value="1993-10-10">10.x.1993</collectingDate>
</date>
,
<collectionCode id="ED5C9ABD8F2CFFF2FF0142C5FF2A8379" box="[151,169,509,535]" name="Tavera, Department of Geology and Geophysics" pageId="4" pageNumber="565">T</collectionCode>
. Nonaka
</materialsCitation>
.
<materialsCitation id="3B2508258F2CFFF2FE8C42C5FCBB8379" ID-GBIF-Occurrence="3766757301" box="[282,824,509,536]" collectingDate="1993-10-10" collectionCode="T" country="Taiwan" location="Taitung-hsien" municipality="Chihpen" pageId="4" pageNumber="565" specimenCount="45" typeStatus="paratype">
<typeStatus id="54F6BCDA8F2CFFF2FE8C42C5FE048379" box="[282,391,509,535]" pageId="4" pageNumber="565" type="paratype">Paratypes</typeStatus>
<specimenCount id="9D4BC9F18F2CFFF2FE1842C5FDA58379" box="[398,550,509,536]" pageId="4" pageNumber="565" type="generic">45 specimens</specimenCount>
collected with holotype
</materialsCitation>
:
<specimenCount id="9D4BC9F18F2CFFF2FCD542C5FC278379" box="[835,932,509,535]" pageId="4" pageNumber="565" type="female">1 female</specimenCount>
(mac.),
<materialsCitation id="3B2508258F2CFFF2FB9442C5FC0F8354" ID-GBIF-Occurrence="3766757303" collectingDate="1993-10-10" collectionCode="T" country="Taiwan" location="Taitung-hsien" municipality="Chihpen" pageId="4" pageNumber="565" specimenCount="30" specimenCount-female="30" typeStatus="paratype">
<specimenCount id="9D4BC9F18F2CFFF2FB9442C5FBFF8379" box="[1026,1148,509,535]" pageId="4" pageNumber="565" type="female">30 females</specimenCount>
(apt.), 8 allometric males (apt.) and 6 gynecoid males (apt.), collected together with holotype
</materialsCitation>
.
</paragraph>
<paragraph id="8BF202788F2CFFF2FF51417CFC5883EC" blockId="4.[151,1437,150,715]" pageId="4" pageNumber="565">
Non-paratypic material:
<emphasis id="B939DE6A8F2CFFF2FE4A417CFDD88330" bold="true" box="[476,603,580,606]" pageId="4" pageNumber="565">
<collectingCountry id="F35A42E88F2CFFF2FE4A417CFDD48330" box="[476,599,580,606]" name="Singapore" pageId="4" pageNumber="565">Singapore</collectingCountry>
,
</emphasis>
Macritchie Park, on dead branches,
<specimenCount id="9D4BC9F18F2CFFF2FC6E417CFBDA8330" box="[1016,1113,580,606]" pageId="4" pageNumber="565" type="female">1 female</specimenCount>
(mac.),
<date id="FFF324B88F2CFFF2FB2E417CFAB98330" box="[1208,1338,580,606]" pageId="4" pageNumber="565" value="1976-07-22">22.vii.1976</date>
, S. Okajima;
<materialsCitation id="3B2508258F2CFFF2FF4E4151FC5483EC" ID-GBIF-Occurrence="3766757302" box="[216,983,616,643]" collectingDate="1990-08-07" collectionCode="T" collectorName="Nonaka &amp; S. Okajima" pageId="4" pageNumber="565" specimenCount="1" specimenCount-female="1">
same locality,
<specimenCount id="9D4BC9F18F2CFFF2FEEE4150FE5A83EC" box="[376,473,616,642]" pageId="4" pageNumber="565" type="female">1 female</specimenCount>
(mac.),
<date id="FFF324B88F2CFFF2FDA14150FD3083EC" box="[567,691,616,642]" pageId="4" pageNumber="565" value="1990-08-07">
<collectingDate id="EFB7DD508F2CFFF2FDA14150FD3083EC" box="[567,691,616,642]" pageId="4" pageNumber="565" value="1990-08-07">7.viii.1990</collectingDate>
</date>
,
<collectionCode id="ED5C9ABD8F2CFFF2FD284151FD5383ED" box="[702,720,617,643]" name="Tavera, Department of Geology and Geophysics" pageId="4" pageNumber="565">T</collectionCode>
.
<collectorName id="26B867AE8F2CFFF2FD4D4150FCB183EC" box="[731,818,616,642]" pageId="4" pageNumber="565">Nonaka</collectorName>
&amp;
<collectorName id="26B867AE8F2CFFF2FCC04150FC5483EC" box="[854,983,616,642]" pageId="4" pageNumber="565">S. Okajima</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35751F38F2CFFF2FF5141B4FEC383A5" pageId="4" pageNumber="565" type="discussion">
<paragraph id="8BF202788F2CFFF2FF5141B4FEC383A5" blockId="4.[151,1437,150,715]" pageId="4" pageNumber="565">
<emphasis id="B939DE6A8F2CFFF2FF5141B4FECC83C8" bold="true" box="[199,335,652,678]" pageId="4" pageNumber="565">Comments.</emphasis>
The two macropterous females listed above as non-paratypic specimens are not distinguishable from the
<typeStatus id="54F6BCDA8F2CFFF2FF684189FEBE83A5" box="[254,317,689,715]" pageId="4" pageNumber="565">types</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>