treatments-xml/data/03/C9/00/03C9000DFF99FFAB0BC1CEED6828F896.xml
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<document id="451399935EEEBC69EC58BA08FEBA737B" ID-CLB-Dataset="32218" ID-DOI="10.11646/zootaxa.4317.2.5" ID-GBIF-Dataset="795485ed-5f1b-46d8-8ce6-1216e579b222" ID-ISSN="1175-5326" ID-Zenodo-Dep="884300" ID-ZooBank="440236D4-7C5C-4409-8Ab3-B5F752A7F326" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1504526935299" checkinUser="plazi" docAuthor="Caviedes-Solís, Itzue W." docDate="2017" docId="03C9000DFF99FFAB0BC1CEED6828F896" docLanguage="en" docName="zootaxa.4317.2.5.pdf" docOrigin="Zootaxa 4317 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ptychohyla zoque Caviedes-Solís 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="286" masterDocId="FFF07875FF98FFAC0B56CB276A33FF9B" masterDocTitle="A new species of treefrog of the genus Ptychohyla (Anura: Hylidae) from Southern Mexico" masterLastPageNumber="290" masterPageNumber="279" pageNumber="280" updateTime="1698467826383" updateUser="ExternalLinkService">
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<mods:title id="20A37AA46D02E4357DF538521748DCAA">A new species of treefrog of the genus Ptychohyla (Anura: Hylidae) from Southern Mexico</mods:title>
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<mods:namePart id="D1C04861DD7A78F71B60B1A963713238">Caviedes-Solís, Itzue W.</mods:namePart>
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<mods:title id="3EDDC3A53E58172358F9D6BF50FD90CB">Zootaxa</mods:title>
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<mods:date id="BF5824E96B49DBF37C1CFBA85751476E">2017</mods:date>
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<mods:number id="EDFE986776A0842BCE645110D9FED3AF">2017-04-04</mods:number>
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<treatment id="03C9000DFF99FFAB0BC1CEED6828F896" ID-DOI="http://doi.org/10.5281/zenodo.5998815" ID-GBIF-Taxon="133169960" ID-Zenodo-Dep="5998815" LSID="urn:lsid:plazi:treatment:03C9000DFF99FFAB0BC1CEED6828F896" httpUri="http://treatment.plazi.org/id/03C9000DFF99FFAB0BC1CEED6828F896" lastPageId="7" lastPageNumber="286" pageId="1" pageNumber="280">
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<heading id="D0970677FF99FFAD0BC1CEED6BE3FA7E" bold="true" box="[151,464,1482,1509]" fontSize="11" level="1" pageId="1" pageNumber="280" reason="1">
<emphasis id="B9146D09FF99FFAD0BC1CEED6BE3FA7E" bold="true" box="[151,464,1482,1509]" pageId="1" pageNumber="280">
<taxonomicName id="4C60CA98FF99FFAD0BC1CEED6B58FA7F" ID-CoL="78NC8" authority="Caviedes-Solís, 2017" authorityName="Caviedes-Solís" authorityYear="2017" box="[151,363,1482,1508]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="1" pageNumber="280" phylum="Chordata" rank="species" species="zoque" status="sp. nov.">
<emphasis id="B9146D09FF99FFAD0BC1CEED6B58FA7F" bold="true" box="[151,363,1482,1508]" italics="true" pageId="1" pageNumber="280">Ptychohyla zoque</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A227D072FF99FFAD0A25CEEC6BE3FA7E" box="[371,464,1483,1509]" pageId="1" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDFB11BFF99FFAD0BC1CEC96B1CF9B1" blockId="1.[151,592,1482,1578]" pageId="1" pageNumber="280">
Zoque Treefrog, Rana arborícola
<taxonomicName id="4C60CA98FF99FFAD095BCEC96863F99E" authorityName="Caviedes-Solís" authorityYear="2017" box="[525,592,1518,1541]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="1" pageNumber="280" phylum="Chordata" rank="species" species="zoque">zoque</taxonomicName>
(
<figureCitation id="135BAD9EFF99FFAD0BC8CD366B16F9B1" box="[158,293,1553,1578]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1057,1079]" captionStartId-1="3.[151,250,1949,1971]" captionTargetBox-0="[151,1436,197,1035]" captionTargetBox-1="[389,1202,1042,1916]" captionTargetId-0="figure@2.[151,1436,193,1035]" captionTargetId-1="figure@3.[367,1219,1021,1928]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Specimens in life of Ptychohyla zoque. A: Male holotype (MZFC 24004, 36.4 mm SVL); B: Female paratype (UTA-A 59271, 55.65 SVL); C: male paratype (CARIE 1135); D: Female and male paratypes in amplexus (CARIE 1215 and 1216)." captionText-1="FIGURE 2. Ventral aspect of hand and foot of Ptychohyla zoque, holotype (MZFC 24004). Scale bar = 3 mm." httpUri-0="https://zenodo.org/record/884302/files/figure.png" httpUri-1="https://zenodo.org/record/884304/files/figure.png" pageId="1" pageNumber="280">Figures 12</figureCitation>
)
</paragraph>
</subSubSection>
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<paragraph id="8BDFB11BFF99FFAD0BC1CD7E6E74F9F4" blockId="1.[151,1095,1625,1647]" box="[151,1095,1625,1647]" pageId="1" pageNumber="280">
<taxonomicName id="4C60CA98FF99FFAD0BC1CD7E6B3AF9F4" authority="Caviedes-Solís, 2017" authorityName="Caviedes-Solís" authorityYear="2017" box="[151,265,1625,1647]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="1" pageNumber="280" phylum="Chordata" rank="genus" status="sp. nov.">
<emphasis id="B9146D09FF99FFAD0BC1CD7E6B3AF9F4" box="[151,265,1625,1647]" italics="true" pageId="1" pageNumber="280">Ptychohyla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A227D072FF99FFAD0A59CD7D6B6DF9F4" box="[271,350,1626,1647]" pageId="1" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B9146D09FF99FFAD0A0BCD7E6B45F9F5" bold="true" box="[349,374,1625,1646]" pageId="1" pageNumber="280"></emphasis>
<bibRefCitation id="EFF1CCEAFF99FFAD0A20CD7E6E71F9F4" box="[374,1090,1625,1647]" pageId="1" pageNumber="280" refString="Aguilar-Lopez, J. L., Pineda, E., Luria-Manzano, R. &amp; Canseco-Marquez, L. (2016) Species Diversity, Distribution, and Conservation Status in a Mesoamerican Region: Amphibians of the Uxpanapa-Chimalapas Region, Mexico. Tropical Conservation Science, 9 (4), 1 - 16. http: // doi. org / 10.1177 / 1940082916670003" type="journal article">Aguilar-López, Pineda, Luría-Manzano, and Canseco-Márquez (2016)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37AE290FF99FFAF0BC1CDBB683EFF63" lastPageId="3" lastPageNumber="282" pageId="1" pageNumber="280" type="materials_examined">
<paragraph id="8BDFB11BFF99FFAD0BC1CDBB68C4F966" blockId="1.[151,1436,1692,2005]" pageId="1" pageNumber="280">
<materialsCitation id="3B08BB46FF99FFAD0BC1CDBB68C4F966" ID-GBIF-Occurrence="1606511269" collectingDate="2008-07-06" country="Mexico" elevation="76" latitude="17.180666" location="Moral" longLatPrecision="13" longitude="-94.5855" municipality="Uxpanapa" pageId="1" pageNumber="280" specimenCount="1" stateProvince="Veracruz" typeStatus="holotype">
<emphasis id="B9146D09FF99FFAD0BC1CDBB6B38F92E" bold="true" box="[151,267,1692,1717]" pageId="1" pageNumber="280">
<typeStatus id="54DB0FB9FF99FFAD0BC1CDBB6B35F92E" box="[151,262,1692,1717]" pageId="1" pageNumber="280" type="holotype">Holotype</typeStatus>
.
</emphasis>
UNAM-MZFC 24004 (original field number RLM 062), an adult male (
<figureCitation id="135BAD9EFF99FFAD0F6ECDBB6E45F92F" box="[1080,1142,1692,1717]" captionStart="FIGURE 1" captionStartId="2.[151,250,1057,1079]" captionTargetBox="[151,1436,197,1035]" captionTargetId="figure@2.[151,1436,193,1035]" captionTargetPageId="2" captionText="FIGURE 1. Specimens in life of Ptychohyla zoque. A: Male holotype (MZFC 24004, 36.4 mm SVL); B: Female paratype (UTA-A 59271, 55.65 SVL); C: male paratype (CARIE 1135); D: Female and male paratypes in amplexus (CARIE 1215 and 1216)." httpUri="https://zenodo.org/record/884302/files/figure.png" pageId="1" pageNumber="280">Fig. 1</figureCitation>
A) from
<locationDeviation id="9A6DA839FF99FFAD0F8ACDBB6FAFF92F" box="[1244,1436,1692,1717]" pageId="1" pageNumber="280">
<quantity id="4C981CFEFF99FFAD0F8ACDBB6F19F92E" box="[1244,1322,1692,1717]" metricMagnitude="3" metricUnit="m" metricValue="6.5" pageId="1" pageNumber="280" unit="km" value="6.5">6.5 km</quantity>
SSE Paso
</locationDeviation>
del
<location id="8EBFE7C0FF99FFAD0B97CD986B3BF943" LSID="urn:lsid:plazi:treatment:03C9000DFF99FFAB0BC1CEED6828F896:8EBFE7C0FF99FFAD0B97CD986B3BF943" box="[193,264,1727,1752]" country="Mexico" latitude="17.180666" longLatPrecision="13" longitude="-94.5855" municipality="Uxpanapa" name="Moral" pageId="1" pageNumber="280" stateProvince="Veracruz">Moral</location>
,
<collectingMunicipality id="6BBB2B61FF99FFAD0A42CDE76BBAF94C" box="[276,393,1728,1751]" pageId="1" pageNumber="280">Uxpanapa</collectingMunicipality>
,
<collectingRegion id="49A47FF9FF99FFAD0AC2CDE76BC8F94C" box="[404,507,1728,1751]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="1" pageNumber="280">Veracruz</collectingRegion>
,
<collectingCountry id="F377F18BFF99FFAD0950CD98686FF943" box="[518,604,1727,1752]" name="Mexico" pageId="1" pageNumber="280">Mexico</collectingCountry>
(
<geoCoordinate id="EE54D7DCFF99FFAD093ACDE768CDF94C" box="[620,766,1728,1752]" direction="north" orientation="latitude" pageId="1" pageNumber="280" precision="9" value="17.180666">17° 10.84' N</geoCoordinate>
,
<geoCoordinate id="EE54D7DCFF99FFAD085FCDE76993F94C" box="[777,928,1727,1752]" direction="west" orientation="longitude" pageId="1" pageNumber="280" precision="9" value="-94.5855">94° 35.13' W</geoCoordinate>
;
<elevation id="004D5628FF99FFAD08FACD9869D5F94C" box="[940,998,1727,1752]" metricMagnitude="1" metricUnit="m" metricValue="7.6" pageId="1" pageNumber="280" unit="m" value="76.0">
<quantity id="4C981CFEFF99FFAD08FACD9869D5F94C" box="[940,998,1727,1752]" metricMagnitude="1" metricUnit="m" metricValue="7.6" pageId="1" pageNumber="280" unit="m" value="76.0">76 m</quantity>
</elevation>
elevation) collected on
<collectingDate id="EF9A6E33FF99FFAD0FA3CD986F4AF943" box="[1269,1401,1727,1752]" pageId="1" pageNumber="280" value="2008-07-06">
<date id="FFDE97DBFF99FFAD0FA3CD986F4AF943" box="[1269,1401,1727,1752]" pageId="1" pageNumber="280" value="2008-07-06">6 July 2008</date>
</collectingDate>
by Ricardo Luría-Manzano and José Luis Aguilar-López.
</materialsCitation>
</paragraph>
<paragraph id="8BDFB11BFF99FFAD0B91CC206E58F84E" blockId="1.[151,1436,1692,2005]" pageId="1" pageNumber="280">
<materialsCitation id="3B08BB46FF99FFAD0B91CC206E34F8DF" ID-GBIF-Occurrence="1606511277" collectingDate="1993-05-09" county="Eight" elevation="600" latitude="17.054167" location="Chimalapas region" longLatPrecision="13" longitude="-94.65383" municipality="Chalchijapa" pageId="1" pageNumber="280" specimenCount="1" stateProvince="Oaxaca" typeStatus="paratype">
<emphasis id="B9146D09FF99FFAD0B91CC206B75F8BB" bold="true" box="[199,326,1799,1824]" pageId="1" pageNumber="280">
<typeStatus id="54DB0FB9FF99FFAD0B91CC206B72F8BB" box="[199,321,1799,1824]" pageId="1" pageNumber="280" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCounty id="62BEC997FF99FFAD0A07CC206BBEF8BB" box="[337,397,1799,1824]" pageId="1" pageNumber="280">Eight</collectingCounty>
specimens. UNAM-MZFC 1 8667 (adult female) from
<collectingMunicipality id="6BBB2B61FF99FFAD0F4CCC206EACF8BB" box="[1050,1183,1799,1824]" pageId="1" pageNumber="280">Chalchijapa</collectingMunicipality>
(
<location id="8EBFE7C0FF99FFAD0FE4CC206FA3F8BB" LSID="urn:lsid:plazi:treatment:03C9000DFF99FFAB0BC1CEED6828F896:8EBFE7C0FF99FFAD0FE4CC206FA3F8BB" box="[1202,1424,1799,1824]" county="Eight" latitude="17.054167" longLatPrecision="13" longitude="-94.65383" municipality="Chalchijapa" name="Chimalapas region" pageId="1" pageNumber="280" stateProvince="Oaxaca">Chimalapas region</location>
),
<collectingRegion id="49A47FF9FF99FFAD0BC1CC0B6AD8F8DF" box="[151,235,1836,1860]" country="Mexico" name="Oaxaca" pageId="1" pageNumber="280">Oaxaca</collectingRegion>
(
<geoCoordinate id="EE54D7DCFF99FFAD0BACCC0B6B44F8DE" box="[250,375,1836,1861]" direction="north" orientation="latitude" pageId="1" pageNumber="280" precision="9" value="17.054167">17° 3.25N</geoCoordinate>
,
<geoCoordinate id="EE54D7DCFF99FFAD0AD4CC0B6829F8DF" box="[386,538,1836,1860]" direction="west" orientation="longitude" pageId="1" pageNumber="280" precision="9" value="-94.65383">94° 39.23 W</geoCoordinate>
;
<elevation id="004D5628FF99FFAD0972CC0B6858F8DF" box="[548,619,1836,1861]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="280" unit="m" value="600.0">
<quantity id="4C981CFEFF99FFAD0972CC0B6858F8DF" box="[548,619,1836,1861]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="280" unit="m" value="600.0">600 m</quantity>
</elevation>
elevation), collected on
<collectingDate id="EF9A6E33FF99FFAD0828CC0B6E34F8DF" box="[894,1031,1836,1861]" pageId="1" pageNumber="280" value="1993-05-09">
<date id="FFDE97DBFF99FFAD0828CC0B6E34F8DF" box="[894,1031,1836,1861]" pageId="1" pageNumber="280" value="1993-05-09">9 May 1993</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B08BB46FF99FFAD0F44CC0B6B94F817" ID-GBIF-Occurrence="1606510444" collectingDate="2009-03-05" collectingDateMax="2009-03-06" collectingDateMin="2009-03-05" collectionCode="UTA" elevation="230" latitude="17.18" location="Moral" longLatPrecision="92" longitude="-94.589836" municipality="Paso" pageId="1" pageNumber="280" specimenCode="MZFC 24005-06" specimenCount="1" stateProvince="UTA" typeStatus="paratype">
three females
<specimenCode id="DBC61960FF99FFAD0FF9CC0B6F5EF8DE" box="[1199,1389,1836,1861]" collectionCode="MZFC" country="Mexico" httpUri="http://grbio.org/cool/fg24-x4zf" name="Museo de Zoologia &amp;quot;Alfonso L. Herrera&amp;quot;" pageId="1" pageNumber="280">MZFC 24005-06</specimenCode>
and
<collectionCode id="ED7129DEFF99FFAD0BC1CC776AFDF8F3" box="[151,206,1872,1896]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34873" name="University of Texas at Arlington" pageId="1" pageNumber="280">
<collectingRegion id="49A47FF9FF99FFAD0BC1CC776AFDF8F3" box="[151,206,1872,1896]" country="United States of America" name="Utah" pageId="1" pageNumber="280">UTA</collectingRegion>
</collectionCode>
<accessionNumber id="94332CF8FF99FFAD0B83CC776B04F8F3" box="[213,311,1871,1896]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A59271" pageId="1" pageNumber="280" type="EnaNcbi">A 59271</accessionNumber>
(
<figureCitation id="135BAD9EFF99FFAD0A10CC686BB6F8F3" box="[326,389,1871,1896]" captionStart="FIGURE 1" captionStartId="2.[151,250,1057,1079]" captionTargetBox="[151,1436,197,1035]" captionTargetId="figure@2.[151,1436,193,1035]" captionTargetPageId="2" captionText="FIGURE 1. Specimens in life of Ptychohyla zoque. A: Male holotype (MZFC 24004, 36.4 mm SVL); B: Female paratype (UTA-A 59271, 55.65 SVL); C: male paratype (CARIE 1135); D: Female and male paratypes in amplexus (CARIE 1215 and 1216)." httpUri="https://zenodo.org/record/884302/files/figure.png" pageId="1" pageNumber="280">Fig. 1</figureCitation>
B) from
<collectingMunicipality id="6BBB2B61FF99FFAD0ABFCC77682DF8FC" box="[489,542,1872,1895]" pageId="1" pageNumber="280">Paso</collectingMunicipality>
del
<location id="8EBFE7C0FF99FFAD0918CC6868A6F8F3" LSID="urn:lsid:plazi:treatment:03C9000DFF99FFAB0BC1CEED6828F896:8EBFE7C0FF99FFAD0918CC6868A6F8F3" box="[590,661,1871,1896]" latitude="17.18" longLatPrecision="92" longitude="-94.589836" municipality="Paso" name="Moral" pageId="1" pageNumber="280" stateProvince="UTA">Moral</location>
,
<collectingRegion id="49A47FF9FF99FFAD09F7CC776937F8FC" box="[673,772,1872,1895]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="1" pageNumber="280">Veracruz</collectingRegion>
(
<geoCoordinate id="EE54D7DCFF99FFAD0842CC7769AAF8FC" box="[788,921,1872,1896]" direction="north" orientation="latitude" pageId="1" pageNumber="280" precision="92" value="17.18">17° 10.8 N</geoCoordinate>
,
<geoCoordinate id="EE54D7DCFF99FFAD08F2CC776E0FF8FC" box="[932,1084,1871,1896]" direction="west" orientation="longitude" pageId="1" pageNumber="280" precision="9" value="-94.589836">94° 35.39 W</geoCoordinate>
;
<elevation id="004D5628FF99FFAD0F1ECC776EBCF8FC" box="[1096,1167,1872,1896]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="280" unit="m" value="230.0">
<quantity id="4C981CFEFF99FFAD0F1ECC776EBCF8FC" box="[1096,1167,1872,1896]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="1" pageNumber="280" unit="m" value="230.0">230 m</quantity>
</elevation>
elevation), collected
<collectingDate id="EF9A6E33FF99FFAD0ED6CC686B06F817" pageId="1" pageNumber="280" value="2009-03-05" valueMax="2009-03-06" valueMin="2009-03-05">
<date id="FFDE97DBFF99FFAD0ED6CC686B06F817" pageId="1" pageNumber="280" value="2009-03-05" valueMax="2009-03-06" valueMin="2009-03-05">5 6 March 2009</date>
</collectingDate>
(
<figureCitation id="135BAD9EFF99FFAD0A10CC536BB5F817" box="[326,390,1908,1933]" captionStart="FIGURE 1" captionStartId="2.[151,250,1057,1079]" captionTargetBox="[151,1436,197,1035]" captionTargetId="figure@2.[151,1436,193,1035]" captionTargetPageId="2" captionText="FIGURE 1. Specimens in life of Ptychohyla zoque. A: Male holotype (MZFC 24004, 36.4 mm SVL); B: Female paratype (UTA-A 59271, 55.65 SVL); C: male paratype (CARIE 1135); D: Female and male paratypes in amplexus (CARIE 1215 and 1216)." httpUri="https://zenodo.org/record/884302/files/figure.png" pageId="1" pageNumber="280">Fig. 1</figureCitation>
B)
</materialsCitation>
;
<materialsCitation id="3B08BB46FF99FFAD0AE2CC536F0FF817" ID-GBIF-Occurrence="1606510464" box="[436,1340,1908,1933]" collectingDate="2013-04-10" collectingDateMax="2009-03-06" collectingDateMin="2009-03-05" collectionCode="CARIE" elevation="230" latitude="17.18" location="Moral" longLatPrecision="92" longitude="-94.589836" municipality="Paso" pageId="1" pageNumber="280" specimenCode="CARIE 1135" specimenCount="1" stateProvince="UTA" typeStatus="holotype">
<specimenCode id="DBC61960FF99FFAD0AE2CC536874F816" box="[436,583,1908,1933]" collectionCode="CARIE" pageId="1" pageNumber="280">CARIE 1135</specimenCode>
(adult male,
<figureCitation id="135BAD9EFF99FFAD098BCC536928F817" box="[733,795,1908,1933]" captionStart="FIGURE 1" captionStartId="2.[151,250,1057,1079]" captionTargetBox="[151,1436,197,1035]" captionTargetId="figure@2.[151,1436,193,1035]" captionTargetPageId="2" captionText="FIGURE 1. Specimens in life of Ptychohyla zoque. A: Male holotype (MZFC 24004, 36.4 mm SVL); B: Female paratype (UTA-A 59271, 55.65 SVL); C: male paratype (CARIE 1135); D: Female and male paratypes in amplexus (CARIE 1215 and 1216)." httpUri="https://zenodo.org/record/884302/files/figure.png" pageId="1" pageNumber="280">Fig. 1</figureCitation>
C) same locality as the
<typeStatus id="54DB0FB9FF99FFAD0F7FCC536EBDF816" box="[1065,1166,1908,1933]" pageId="1" pageNumber="280" type="holotype">holotype</typeStatus>
,
<collectingDate id="EF9A6E33FF99FFAD0FCCCC536F0FF817" box="[1178,1340,1908,1933]" pageId="1" pageNumber="280" value="2013-04-10">
<date id="FFDE97DBFF99FFAD0FCCCC536F0FF817" box="[1178,1340,1908,1933]" pageId="1" pageNumber="280" value="2013-04-10">10 April 2013</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="3B08BB46FF99FFAD0E1ECC536E58F84E" ID-GBIF-Occurrence="1606510471" collectingDate="2013-05-16" collectingDateMax="2013-06-12" collectingDateMin="2013-05-16" collectionCode="CARIE" elevation="246" latitude="17.189" location="Uxpanapa" longLatPrecision="88" longitude="-94.486664" municipality="Arroyo Zarco" pageId="1" pageNumber="280" specimenCode="CARIE 1138, CARIE 1215-16" specimenCount="1" stateProvince="Veracruz" typeStatus="holotype">
<specimenCode id="DBC61960FF99FFAD0E1ECC536AFDF82B" collectionCode="CARIE" pageId="1" pageNumber="280">CARIE 1138</specimenCode>
(adult male)
<collectingDate id="EF9A6E33FF99FFAD0A35CCBF6BC4F82B" box="[355,503,1944,1969]" pageId="1" pageNumber="280" value="2013-05-16">
<date id="FFDE97DBFF99FFAD0A35CCBF6BC4F82B" box="[355,503,1944,1969]" pageId="1" pageNumber="280" value="2013-05-16">16 May 2013</date>
</collectingDate>
and
<specimenCode id="DBC61960FF99FFAD097BCCBF68DAF82A" box="[557,745,1944,1969]" collectionCode="CARIE" pageId="1" pageNumber="280">
CARIE
<date id="FFDE97DBFF99FFAD09DECCBF68DAF82A" box="[648,745,1944,1969]" pageId="1" pageNumber="280" value="1215" valueMax="1216">121516</date>
</specimenCode>
(one adult male and one adult female,
<figureCitation id="135BAD9EFF99FFAD0FCACCBF6EEBF82B" box="[1180,1240,1944,1969]" captionStart="FIGURE 1" captionStartId="2.[151,250,1057,1079]" captionTargetBox="[151,1436,197,1035]" captionTargetId="figure@2.[151,1436,193,1035]" captionTargetPageId="2" captionText="FIGURE 1. Specimens in life of Ptychohyla zoque. A: Male holotype (MZFC 24004, 36.4 mm SVL); B: Female paratype (UTA-A 59271, 55.65 SVL); C: male paratype (CARIE 1135); D: Female and male paratypes in amplexus (CARIE 1215 and 1216)." httpUri="https://zenodo.org/record/884302/files/figure.png" pageId="1" pageNumber="280">Fig. 1</figureCitation>
D)
<collectingDate id="EF9A6E33FF99FFAD0E57CCBF6FABF82B" box="[1281,1432,1944,1968]" pageId="1" pageNumber="280" value="2013-06-12">
<date id="FFDE97DBFF99FFAD0E57CCBF6FABF82B" box="[1281,1432,1944,1968]" pageId="1" pageNumber="280" value="2013-06-12">12 June 2013</date>
</collectingDate>
, from
<collectingMunicipality id="6BBB2B61FF99FFAD0B82CC9B6B43F84F" box="[212,368,1980,2004]" pageId="1" pageNumber="280">Arroyo Zarco</collectingMunicipality>
,
<location id="8EBFE7C0FF99FFAD0A2DCC9A6BC3F84F" LSID="urn:lsid:plazi:treatment:03C9000DFF99FFAB0BC1CEED6828F896:8EBFE7C0FF99FFAD0A2DCC9A6BC3F84F" box="[379,496,1981,2004]" latitude="17.189" longLatPrecision="88" longitude="-94.486664" municipality="Arroyo Zarco" name="Uxpanapa" pageId="1" pageNumber="280" stateProvince="Veracruz">Uxpanapa</location>
,
<collectingRegion id="49A47FF9FF99FFAD0AADCC9A686CF84F" box="[507,607,1981,2004]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="1" pageNumber="280">Veracruz</collectingRegion>
(
<geoCoordinate id="EE54D7DCFF99FFAD0939CC9B6931F84F" box="[623,770,1980,2004]" direction="north" orientation="latitude" pageId="1" pageNumber="280" precision="9" value="17.189">17° 11.34 N</geoCoordinate>
,
<geoCoordinate id="EE54D7DCFF99FFAD0858CC9B69A9F84F" box="[782,922,1980,2004]" direction="west" orientation="longitude" pageId="1" pageNumber="280" precision="92" value="-94.486664">94° 29.2 W</geoCoordinate>
;
<elevation id="004D5628FF99FFAD08F3CC9B69DFF84F" box="[933,1004,1980,2005]" metricMagnitude="2" metricUnit="m" metricValue="2.46" pageId="1" pageNumber="280" unit="m" value="246.0">
<quantity id="4C981CFEFF99FFAD08F3CC9B69DFF84F" box="[933,1004,1980,2005]" metricMagnitude="2" metricUnit="m" metricValue="2.46" pageId="1" pageNumber="280" unit="m" value="246.0">246 m</quantity>
</elevation>
elevation).
</materialsCitation>
</paragraph>
<caption id="DF1FE193FF9AFFAE0BC1CF066AEAFBEF" httpUri="https://zenodo.org/record/884302/files/figure.png" pageId="2" pageNumber="281" targetBox="[151,1436,197,1035]" targetPageId="2">
<paragraph id="8BDFB11BFF9AFFAE0BC1CF066AEAFBEF" blockId="2.[151,1436,1057,1140]" pageId="2" pageNumber="281">
<emphasis id="B9146D09FF9AFFAE0BC1CF066B26FBAD" bold="true" box="[151,277,1057,1079]" pageId="2" pageNumber="281">FIGURE 1.</emphasis>
Specimens in life of
<taxonomicName id="4C60CA98FF9AFFAE0AAFCF06689CFBAC" authorityName="Caviedes-Solís" authorityYear="2017" box="[505,687,1057,1079]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="2" pageNumber="281" phylum="Chordata" rank="species" species="zoque">
<emphasis id="B9146D09FF9AFFAE0AAFCF06689CFBAC" box="[505,687,1057,1079]" italics="true" pageId="2" pageNumber="281">Ptychohyla zoque</emphasis>
</taxonomicName>
. A: Male holotype (MZFC 24004, 36.4 mm SVL); B: Female paratype (UTA-A 59271, 55.65 SVL); C: male paratype (CARIE 1135); D: Female and male paratypes in amplexus (CARIE 1215 and 1216).
</paragraph>
</caption>
<caption id="DF1FE193FF9AFFAE0BC1CFBC6B28FB49" pageId="2" pageNumber="281">
<paragraph id="8BDFB11BFF9AFFAE0BC1CFBC6B28FB49" blockId="2.[151,1436,1178,1234]" pageId="2" pageNumber="281">
<emphasis id="B9146D09FF9AFFAE0BC1CFBC6B3DFB29" bold="true" box="[151,270,1179,1202]" pageId="2" pageNumber="281">TABLE 1.</emphasis>
Morphometric variation in type-series of
<taxonomicName id="4C60CA98FF9AFFAE09B3CFBD6995FB29" authorityName="Caviedes-Solís" authorityYear="2017" box="[741,934,1178,1202]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="2" pageNumber="281" phylum="Chordata" rank="species" species="zoque">
<emphasis id="B9146D09FF9AFFAE09B3CFBD6995FB29" box="[741,934,1178,1202]" italics="true" pageId="2" pageNumber="281">Ptychohyla zoque</emphasis>
</taxonomicName>
: range (in millimeters) followed by mean in parentheses.
</paragraph>
</caption>
<paragraph id="8BDFB11BFF9AFFAE0BC9CFC86E7CFA9F" blockId="2.[159,1186,1262,1540]" box="[159,1103,1262,1284]" pageId="2" pageNumber="281">Measurement Males Females</paragraph>
<paragraph id="8BDFB11BFF9AFFAE091ACE2B6E0BFAB9" blockId="2.[159,1186,1262,1540]" box="[588,1080,1292,1315]" pageId="2" pageNumber="281">
(
<emphasis id="B9146D09FF9AFFAE0903CE296852FAB8" box="[597,609,1294,1315]" italics="true" pageId="2" pageNumber="281">n</emphasis>
= 4) (
<emphasis id="B9146D09FF9AFFAE0F54CE296E3DFAB8" box="[1026,1038,1294,1315]" italics="true" pageId="2" pageNumber="281">n</emphasis>
= 5)
</paragraph>
<paragraph id="8BDFB11BFF9AFFAF0BC9CE11683EFF63" blockId="2.[159,1186,1262,1540]" lastBlockId="3.[151,1437,151,968]" lastPageId="3" lastPageNumber="282" pageId="2" pageNumber="281">
SVL 35.036.4 (35.8) 46.257.8 (54.3) Tibia length 20.322.0 (20.9) 28.935.6 (32.9) Foot length 14.216.7 (15.4) 20.926.3 (24.2) Head length 12.012.9 (12.5) 17.220.7 (19.3) Head width 12.113.5 (12.9) 17.421.7 (20.3) Diameter of eye 4.04.8 (4.4) 5.25.6 (5.5) Diameter of tympanum 1.92.3 (2.1) 2.43.0 (2.7)
<emphasis id="B9146D09FF9AFFAE0B91CD646B72F9C7" bold="true" box="[199,321,1603,1628]" pageId="2" pageNumber="281">Diagnosis.</emphasis>
<taxonomicName id="4C60CA98FF9AFFAE0A11CD63683EF9C0" authorityName="Caviedes-Solís" authorityYear="2017" box="[327,525,1604,1628]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="2" pageNumber="281" phylum="Chordata" rank="species" species="zoque">
<emphasis id="B9146D09FF9AFFAE0A11CD63683EF9C0" box="[327,525,1604,1628]" italics="true" pageId="2" pageNumber="281">Ptychohyla zoque</emphasis>
</taxonomicName>
is placed into the genus
<taxonomicName id="4C60CA98FF9AFFAE0876CD6369ADF9C7" authorityName="Caviedes-Solís" authorityYear="2017" box="[800,926,1604,1628]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="2" pageNumber="281" phylum="Chordata" rank="genus">
<emphasis id="B9146D09FF9AFFAE0876CD6369ADF9C7" box="[800,926,1604,1628]" italics="true" pageId="2" pageNumber="281">Ptychohyla</emphasis>
</taxonomicName>
because of characteristics described below. It has a green dorsal coloration with brown blotches and numerous irregular white spots, and the iris is pale pink with brown reticulations. There is a marked sexual size dimorphism, with males attaining
<quantity id="4C981CFEFF9AFFAE0F1ACDAC6E83F938" box="[1100,1200,1675,1700]" metricMagnitude="-2" metricUnit="m" metricValue="3.6399999999999997" pageId="2" pageNumber="281" unit="mm" value="36.4">36.4 mm</quantity>
in SVL, and females
<quantity id="4C981CFEFF9AFFAE0BC1CD886B30F95C" box="[151,259,1711,1736]" metricMagnitude="-2" metricUnit="m" metricValue="5.779999999999999" pageId="2" pageNumber="281" unit="mm" value="57.8">57.8 mm</quantity>
. Breeding males have small nonspinous nuptial excrescences, and lack glands on the venter (either ventrolateral or chest).
<taxonomicName id="4C60CA98FF9AFFAE0ACFCDF3686DF970" authorityName="Caviedes-Solís" authorityYear="2017" box="[409,606,1748,1772]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="2" pageNumber="281" phylum="Chordata" rank="species" species="zoque">
<emphasis id="B9146D09FF9AFFAE0ACFCDF3686DF970" box="[409,606,1748,1772]" italics="true" pageId="2" pageNumber="281">Ptychohyla zoque</emphasis>
</taxonomicName>
may be distinguished from all other species in the genus
<taxonomicName id="4C60CA98FF9AFFAE0F8DCDF36F6AF977" authorityName="Caviedes-Solís" authorityYear="2017" box="[1243,1369,1748,1772]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="2" pageNumber="281" phylum="Chordata" rank="genus">
<emphasis id="B9146D09FF9AFFAE0F8DCDF36F6AF977" box="[1243,1369,1748,1772]" italics="true" pageId="2" pageNumber="281">Ptychohyla</emphasis>
</taxonomicName>
by its green dorsal coloration and iris pale pink. Although it superficially resembles
<taxonomicName id="4C60CA98FF9AFFAE08A8CDDF6EAAF894" box="[1022,1177,1783,1807]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="2" pageNumber="281" phylum="Chordata" rank="species" species="acrochorda">
<emphasis id="B9146D09FF9AFFAE08A8CDDF6EAAF894" box="[1022,1177,1783,1807]" italics="true" pageId="2" pageNumber="281">P. acrochorda</emphasis>
</taxonomicName>
,
<taxonomicName id="4C60CA98FF9AFFAE0FF1CDDF6F78F894" baseAuthorityName="Taylor" baseAuthorityYear="1937" box="[1191,1355,1783,1807]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="2" pageNumber="281" phylum="Chordata" rank="species" species="erythromma">
<emphasis id="B9146D09FF9AFFAE0FF1CDDF6F78F894" box="[1191,1355,1783,1807]" italics="true" pageId="2" pageNumber="281">P. erythromma</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C60CA98FF9AFFAE0ED1CDDF6B02F8A8" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="2" pageNumber="281" phylum="Chordata" rank="species" species="sanctaecrucis">
<emphasis id="B9146D09FF9AFFAE0ED1CDDF6B02F8A8" italics="true" pageId="2" pageNumber="281">P. sanctaecrucis</emphasis>
</taxonomicName>
in regard to dorsal coloration,
<taxonomicName id="4C60CA98FF9AFFAE09C9CC3B68CCF8A8" authorityName="Caviedes-Solís" authorityYear="2017" box="[671,767,1820,1843]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="2" pageNumber="281" phylum="Chordata" rank="species" species="zoque">
<emphasis id="B9146D09FF9AFFAE09C9CC3B68CCF8A8" box="[671,767,1820,1843]" italics="true" pageId="2" pageNumber="281">P. zoque</emphasis>
</taxonomicName>
may be distinguished by its iris of pale pink with brown reticulations (whitish or silver in
<taxonomicName id="4C60CA98FF9AFFAE095FCC676896F8CC" box="[521,677,1855,1879]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="2" pageNumber="281" phylum="Chordata" rank="species" species="acrochorda">
<emphasis id="B9146D09FF9AFFAE095FCC676896F8CC" box="[521,677,1855,1879]" italics="true" pageId="2" pageNumber="281">P. acrochorda</emphasis>
</taxonomicName>
, red in
<taxonomicName id="4C60CA98FF9AFFAE09AFCC6769AFF8CC" baseAuthorityName="Taylor" baseAuthorityYear="1937" box="[761,924,1855,1879]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="2" pageNumber="281" phylum="Chordata" rank="species" species="erythromma">
<emphasis id="B9146D09FF9AFFAE09AFCC6769AFF8CC" box="[761,924,1855,1879]" italics="true" pageId="2" pageNumber="281">P. erythromma</emphasis>
</taxonomicName>
, and pale tan in
<taxonomicName id="4C60CA98FF9AFFAE0F04CC676F34F8CC" box="[1106,1287,1856,1879]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="2" pageNumber="281" phylum="Chordata" rank="species" species="sanctaecrucis">
<emphasis id="B9146D09FF9AFFAE0F04CC676F34F8CC" box="[1106,1287,1856,1879]" italics="true" pageId="2" pageNumber="281">P. sanctaecrucis</emphasis>
</taxonomicName>
).
<taxonomicName id="4C60CA98FF9AFFAE0E48CC186AEBF8E0" authorityName="Caviedes-Solís" authorityYear="2017" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="2" pageNumber="281" phylum="Chordata" rank="species" species="zoque">
<emphasis id="B9146D09FF9AFFAE0E48CC186AEBF8E0" italics="true" pageId="2" pageNumber="281">Ptychohyla zoque</emphasis>
</taxonomicName>
can also be distinguished by the presence of small nonspinous nuptial excrescences (dark nuptial excrescences composed of spines in
<taxonomicName id="4C60CA98FF9AFFAE0979CCAF68F8F804" box="[559,715,1927,1951]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="2" pageNumber="281" phylum="Chordata" rank="species" species="acrochorda">
<emphasis id="B9146D09FF9AFFAE0979CCAF68F8F804" box="[559,715,1927,1951]" italics="true" pageId="2" pageNumber="281">P. acrochorda</emphasis>
</taxonomicName>
and
<taxonomicName id="4C60CA98FF9AFFAE0857CCAF6985F804" box="[769,950,1928,1951]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="2" pageNumber="281" phylum="Chordata" rank="species" species="sanctaecrucis">
<emphasis id="B9146D09FF9AFFAE0857CCAF6985F804" box="[769,950,1928,1951]" italics="true" pageId="2" pageNumber="281">P. sanctaecrucis</emphasis>
</taxonomicName>
, unknown in
<taxonomicName id="4C60CA98FF9AFFAE0F19CCAF6EC1F804" baseAuthorityName="Taylor" baseAuthorityYear="1937" box="[1103,1266,1927,1951]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="2" pageNumber="281" phylum="Chordata" rank="species" species="erythromma">
<emphasis id="B9146D09FF9AFFAE0F19CCAF6EC1F804" box="[1103,1266,1927,1951]" italics="true" pageId="2" pageNumber="281">P. erythromma</emphasis>
</taxonomicName>
). Additionally, the new species is distinguished from
<taxonomicName id="4C60CA98FF9AFFAE090ACC8B68CDF85F" box="[604,766,1964,1988]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="2" pageNumber="281" phylum="Chordata" rank="species" species="acrochorda">
<emphasis id="B9146D09FF9AFFAE090ACC8B68CDF85F" box="[604,766,1964,1988]" italics="true" pageId="2" pageNumber="281">P. acrochorda</emphasis>
</taxonomicName>
and
<taxonomicName id="4C60CA98FF9AFFAE086BCC8B69C4F858" box="[829,1015,1964,1987]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="2" pageNumber="281" phylum="Chordata" rank="species" species="sanctaecrucis">
<emphasis id="B9146D09FF9AFFAE086BCC8B69C4F858" box="[829,1015,1964,1987]" italics="true" pageId="2" pageNumber="281">P. sanctaecrucis</emphasis>
</taxonomicName>
by the absence of a chest gland in breeding males (present in the latter two species).
<taxonomicName id="4C60CA98FF9AFFAE09B4CCE8699FF87C" authorityName="Caviedes-Solís" authorityYear="2017" box="[738,940,1999,2023]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="2" pageNumber="281" phylum="Chordata" rank="species" species="zoque">
<emphasis id="B9146D09FF9AFFAE09B4CCE8699FF87C" box="[738,940,1999,2023]" italics="true" pageId="2" pageNumber="281">Ptychohyla zoque</emphasis>
</taxonomicName>
and
<taxonomicName id="4C60CA98FF9AFFAE08BECCE86F36F87C" baseAuthorityName="Duellman" baseAuthorityYear="1961" box="[1000,1285,1999,2023]" class="Amphibia" family="Hylidae" genus="Duellmanohyla" kingdom="Animalia" order="Anura" pageId="2" pageNumber="281" phylum="Chordata" rank="species" species="ignicolor">
<emphasis id="B9146D09FF9AFFAE08BECCE86F36F87C" box="[1000,1285,1999,2023]" italics="true" pageId="2" pageNumber="281">Duellmanohyla ignicolor</emphasis>
</taxonomicName>
have similar dorsal color patterns of brown blotches and numerous irregular white spots, and both share a pinkish iris. However,
<taxonomicName id="4C60CA98FF9BFFAF0BC1CB9A6AC6FF4F" authorityName="Caviedes-Solís" authorityYear="2017" box="[151,245,189,212]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="282" phylum="Chordata" rank="species" species="zoque">
<emphasis id="B9146D09FF9BFFAF0BC1CB9A6AC6FF4F" box="[151,245,189,212]" italics="true" pageId="3" pageNumber="282">P. zoque</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C60CA98FF9BFFAF097DCB9A6885FF4F" baseAuthorityName="Duellman" baseAuthorityYear="1961" box="[555,694,188,212]" class="Amphibia" family="Hylidae" genus="Duellmanohyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="282" phylum="Chordata" rank="species" species="ignicolor">
<emphasis id="B9146D09FF9BFFAF097DCB9A6885FF4F" box="[555,694,188,212]" italics="true" pageId="3" pageNumber="282">D. ignicolor</emphasis>
</taxonomicName>
by having nuptial excrescences and lacking noticeably thickened and extended ventrolateral gland.
</paragraph>
</subSubSection>
<subSubSection id="C37AE290FF9BFFAA0B91CA236FAEFBB5" lastPageId="6" lastPageNumber="285" pageId="3" pageNumber="282" type="description">
<paragraph id="8BDFB11BFF9BFFAF0B91CA2369FDFC53" blockId="3.[151,1437,151,968]" pageId="3" pageNumber="282">
<emphasis id="B9146D09FF9BFFAF0B91CA236822FE86" bold="true" box="[199,529,260,285]" pageId="3" pageNumber="282">
Description of the
<typeStatus id="54DB0FB9FF9BFFAF0AF5CA23683FFE86" box="[419,524,260,285]" pageId="3" pageNumber="282" type="holotype">holotype</typeStatus>
.
</emphasis>
An adult male, SVL
<quantity id="4C981CFEFF9BFFAF0857CA23695BFE87" box="[769,872,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="3.6399999999999997" pageId="3" pageNumber="282" unit="mm" value="36.4">36.4 mm</quantity>
, tibia length
<quantity id="4C981CFEFF9BFFAF08AFCA236E52FE87" box="[1017,1121,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="2.09" pageId="3" pageNumber="282" unit="mm" value="20.9">20.9 mm</quantity>
; tibia length/SVL 0.57; foot length
<quantity id="4C981CFEFF9BFFAF0BB3CA006B7DFEA4" box="[229,334,295,320]" metricMagnitude="-2" metricUnit="m" metricValue="1.54" pageId="3" pageNumber="282" unit="mm" value="15.4">15.4 mm</quantity>
; foot length/SVL 0.42; head length
<quantity id="4C981CFEFF9BFFAF09BDCA0F6967FEA4" box="[747,852,296,320]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="3" pageNumber="282" unit="mm" value="12.7">12.7 mm</quantity>
; head length/SVL 0.34; head width
<quantity id="4C981CFEFF9BFFAF0FA4CA0F6F69FEA4" box="[1266,1370,296,320]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="3" pageNumber="282" unit="mm" value="12.8">12.8 mm</quantity>
; head width/SVL 0.35; eye diameter
<quantity id="4C981CFEFF9BFFAF0AADCA6B6864FEFF" box="[507,599,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="3" pageNumber="282" unit="mm" value="4.8">4.8 mm</quantity>
; tympanum diameter
<quantity id="4C981CFEFF9BFFAF0807CA6B699EFEFF" box="[849,941,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="282" unit="mm" value="1.9">1.9 mm</quantity>
; tympanum/eye 0.39. Snout in lateral view truncate, slightly rounded in dorsal view; canthus distinct and slightly angular; loreal region concave; lips moderately thick, not flared; nostrils protuberant; internarial distance
<quantity id="4C981CFEFF9BFFAF08F7CAB369C9FE37" box="[929,1018,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="282" unit="mm" value="2.2">2.2 mm</quantity>
; top of head flat; interorbital distance
<quantity id="4C981CFEFF9BFFAF0BC1CA9F6AC0FE54" box="[151,243,440,464]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="3" pageNumber="282" unit="mm" value="4.3">4.3 mm</quantity>
. Supratympanic fold well developed, extending from posterior corner of eye above tympanum to point above insertion of forearm, covering upper edge of tympanum; tympanum distinct, rounded. Forearm moderately robust; ulnar tubercles unequally elevated. Prepollex ossified, large and blunt. Nuptial excrescences small and nonspinous, located on prepollical protrusion, inner and outer surface of Finger I, and dorsal surface of webbing between Fingers I and II; subarticular tubercles rounded, distal one on Fingers III and IV bifid; discs on fingers large, disc of the third finger slightly smaller than diameter of the tympanum; webbing formula
<date id="FFDE97DBFF9BFFAF0F93C94B6ED8FD1E" box="[1221,1259,620,645]" pageId="3" pageNumber="282">
<emphasis id="B9146D09FF9BFFAF0F93C94B6EE8FD1E" bold="true" box="[1221,1243,620,645]" pageId="3" pageNumber="282">II</emphasis>
1
</date>
½—
<date id="FFDE97DBFF9BFFAF0E74C94B6F65FD1E" box="[1314,1366,620,645]" pageId="3" pageNumber="282">
3
<emphasis id="B9146D09FF9BFFAF0E63C94B6F65FD1E" bold="true" box="[1333,1366,620,645]" pageId="3" pageNumber="282">III</emphasis>
</date>
2½—
<date id="FFDE97DBFF9BFFAF0BC1C9B76AE2FD33" box="[151,209,655,680]" pageId="3" pageNumber="282">
2
<emphasis id="B9146D09FF9BFFAF0BE4C9A86AE2FD33" bold="true" box="[178,209,655,680]" pageId="3" pageNumber="282">IV</emphasis>
</date>
(
<figureCitation id="135BAD9EFF9BFFAF0BB1C9A86B00FD33" box="[231,307,655,680]" captionStart="FIGURE 2" captionStartId="3.[151,250,1949,1971]" captionTargetBox="[389,1202,1042,1916]" captionTargetId="figure@3.[367,1219,1021,1928]" captionTargetPageId="3" captionText="FIGURE 2. Ventral aspect of hand and foot of Ptychohyla zoque, holotype (MZFC 24004). Scale bar = 3 mm." httpUri="https://zenodo.org/record/884304/files/figure.png" pageId="3" pageNumber="282">Fig. 2</figureCitation>
). Heels overlapping when hind limbs adpressed at right angle to body; tibiotarsal articulation extending slightly beyond nostril; row of tubercles forming dermal ridge on the tarsus, extending from heel to disc of fifth toe; heel tubercle present; subarticular tubercles rounded; length of toes from shortest to longest 1-2-3-5-4; webbing formula
<emphasis id="B9146D09FF9BFFAF0A36C9DB6B58FC8E" bold="true" box="[352,363,764,789]" pageId="3" pageNumber="282">I</emphasis>
1—
<date id="FFDE97DBFF9BFFAF0ACBC9DB6BF9FC8E" box="[413,458,764,789]" pageId="3" pageNumber="282">
2
<emphasis id="B9146D09FF9BFFAF0AE2C9DB6BF9FC8E" bold="true" box="[436,458,764,789]" pageId="3" pageNumber="282">II</emphasis>
</date>
1½—
<date id="FFDE97DBFF9BFFAF0947C9DB687BFC8E" box="[529,584,764,789]" pageId="3" pageNumber="282">
2
<emphasis id="B9146D09FF9BFFAF0971C9DB687BFC8E" bold="true" box="[551,584,764,789]" pageId="3" pageNumber="282">III</emphasis>
</date>
1—
<date id="FFDE97DBFF9BFFAF092CC9DB6883FC8E" box="[634,688,764,789]" pageId="3" pageNumber="282">
2
<emphasis id="B9146D09FF9BFFAF09C7C9DB6883FC8E" bold="true" box="[657,688,764,789]" pageId="3" pageNumber="282">IV</emphasis>
</date>
2 —
<date id="FFDE97DBFF9BFFAF09B9C9DB6925FC8E" box="[751,790,764,789]" pageId="3" pageNumber="282">
1
<emphasis id="B9146D09FF9BFFAF0857C9DB6925FC8E" bold="true" box="[769,790,764,789]" pageId="3" pageNumber="282">V</emphasis>
</date>
(
<figureCitation id="135BAD9EFF9BFFAF0873C9DB695FFC8F" box="[805,876,764,789]" captionStart="FIGURE 2" captionStartId="3.[151,250,1949,1971]" captionTargetBox="[389,1202,1042,1916]" captionTargetId="figure@3.[367,1219,1021,1928]" captionTargetPageId="3" captionText="FIGURE 2. Ventral aspect of hand and foot of Ptychohyla zoque, holotype (MZFC 24004). Scale bar = 3 mm." httpUri="https://zenodo.org/record/884304/files/figure.png" pageId="3" pageNumber="282">Fig. 2</figureCitation>
). Toes moderately long; discs of toes about same size as those on fingers (
<figureCitation id="135BAD9EFF9BFFAF0A96C8386839FCA3" box="[448,522,799,824]" captionStart="FIGURE 2" captionStartId="3.[151,250,1949,1971]" captionTargetBox="[389,1202,1042,1916]" captionTargetId="figure@3.[367,1219,1021,1928]" captionTargetPageId="3" captionText="FIGURE 2. Ventral aspect of hand and foot of Ptychohyla zoque, holotype (MZFC 24004). Scale bar = 3 mm." httpUri="https://zenodo.org/record/884304/files/figure.png" pageId="3" pageNumber="282">Fig. 2</figureCitation>
). Cloacal opening directed posteroventrally at midlevel of thighs with several curved folds below cloacal opening; cloacal sheath short; a few white tubercles below and lateral to vent. Skin on dorsum smooth; that of throat, chest, belly and ventral surface of thighs granular. Ventrolateral glands absent. Tongue cordiform with posterior notch; vomerine teeth five on right side, six on left, situated on small, elliptical elevations between ovoid internal nares. Vocal slits present on both sides.
</paragraph>
<caption id="DF1FE193FF9BFFAF0BC1CCBA6ECEF828" box="[151,1277,1949,1971]" httpUri="https://zenodo.org/record/884304/files/figure.png" pageId="3" pageNumber="282" targetBox="[389,1202,1042,1916]" targetPageId="3">
<paragraph id="8BDFB11BFF9BFFAF0BC1CCBA6ECEF828" blockId="3.[151,1277,1949,1971]" box="[151,1277,1949,1971]" pageId="3" pageNumber="282">
<emphasis id="B9146D09FF9BFFAF0BC1CCBA6B2BF829" bold="true" box="[151,280,1949,1971]" pageId="3" pageNumber="282">FIGURE 2.</emphasis>
Ventral aspect of hand and foot of
<taxonomicName id="4C60CA98FF9BFFAF092ACCBA6903F828" authorityName="Caviedes-Solís" authorityYear="2017" box="[636,816,1949,1971]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="3" pageNumber="282" phylum="Chordata" rank="species" species="zoque">
<emphasis id="B9146D09FF9BFFAF092ACCBA6903F828" box="[636,816,1949,1971]" italics="true" pageId="3" pageNumber="282">Ptychohyla zoque</emphasis>
</taxonomicName>
, holotype (MZFC 24004). Scale bar = 3 mm.
</paragraph>
</caption>
<paragraph id="8BDFB11BFF9CFFA80B21CA9D6868FE73" blockId="4.[119,1702,441,533]" pageId="4" pageNumber="283">
<emphasis id="B9146D09FF9CFFA80B21CA9D6AF4FE55" bold="true" box="[119,199,442,462]" pageId="4" pageNumber="283">Table 2.</emphasis>
Selected characteristics of all species of the genus
<taxonomicName id="4C60CA98FF9CFFA809F4CA9E6939FE55" authorityName="Caviedes-Solís" authorityYear="2017" box="[674,778,441,462]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="4" pageNumber="283" phylum="Chordata" rank="genus">
<emphasis id="B9146D09FF9CFFA809F4CA9E6939FE55" box="[674,778,441,462]" italics="true" pageId="4" pageNumber="283">Ptychohyla</emphasis>
</taxonomicName>
. Data of
<taxonomicName id="4C60CA98FF9CFFA80836CA9E69D5FE55" box="[864,998,441,462]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="4" pageNumber="283" phylum="Chordata" rank="species" species="acrochorda">
<emphasis id="B9146D09FF9CFFA80836CA9E69D5FE55" box="[864,998,441,462]" italics="true" pageId="4" pageNumber="283">P. acrochorda</emphasis>
</taxonomicName>
were obtained from
<bibRefCitation id="EFF1CCEAFF9CFFA80FF1CA9D6FE2FE55" author="Campbell" box="[1191,1489,442,462]" pageId="4" pageNumber="283" refString="Campbell, J. A. &amp; Duellman, W. E. (2000) New Species of Stream-breeding Hylid Frogs from the Northern Versant of the Highlands of Oaxaca, Mexico. S cientific Papers Natural History Museum The University of Kansas, 16, 1 - 28." type="journal article" year="2000">Campbell and Duellman (2000)</bibRefCitation>
, and the data of other species were obtained from
<bibRefCitation id="EFF1CCEAFF9CFFA80A2DCAF36827FE73" author="Duellman" box="[379,532,468,488]" pageId="4" pageNumber="283" refString="Duellman, W. E. (1963) A review of the Middle American tree frogs of the genus Ptychohyla. University of Kansas Publications, Museum of Natural History, 15 (7), 297 - 349. http: // doi. org / 10.5962 / bhl. part. 7290" type="journal article" year="1963">Duellman (1963</bibRefCitation>
,
<bibRefCitation id="EFF1CCEAFF9CFFA80948CAF36863FE73" author="Duellman" box="[542,592,468,488]" pageId="4" pageNumber="283" refString="Duellman, W. E. (2001) Hylid Frogs of Middle America. 2 volumes. Society for the Study of Amphibians and Reptiles. 1 - 1158 pp." type="journal article" year="2001">2001</bibRefCitation>
).
</paragraph>
<paragraph id="8BDFB11BFF9CFFA80AFECAC86CA4FA5A" blockId="4.[119,1702,441,533]" lastBlockId="4.[138,1744,1457,1650]" pageId="4" pageNumber="283">
<emphasis id="B9146D09FF9CFFA80AFECAC86C95FD8E" italics="true" pageId="4" pageNumber="283">
P. P. P. P. P. P.
<taxonomicName id="4C60CA98FF9CFFA80D0ACAC8688FFD8E" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="4" pageNumber="283" phylum="Chordata" rank="subSpecies" species="acrochorda" subSpecies="dendrophasma">P. acrochorda dendrophasma</taxonomicName>
<taxonomicName id="4C60CA98FF9CFFA8084BC923694BFD8E" baseAuthorityName="Taylor" baseAuthorityYear="1937" box="[797,888,516,533]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="4" pageNumber="283" phylum="Chordata" rank="species" species="erythromma">erythromma</taxonomicName>
<taxonomicName id="4C60CA98FF9CFFA808BDC9236E76FD8E" baseAuthorityName="Kellogg" baseAuthorityYear="1928" box="[1003,1093,516,533]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="4" pageNumber="283" phylum="Chordata" rank="species" species="euthysanota">euthysanota</taxonomicName>
<taxonomicName id="4C60CA98FF9CFFA80FEDC9236F21FD8E" box="[1211,1298,516,533]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="4" pageNumber="283" phylum="Chordata" rank="species" species="hypomykter">hypomykter</taxonomicName>
legleri
<taxonomicName id="4C60CA98FF9CFFA80D74C9236C95FD8E" baseAuthorityName="Ahl" baseAuthorityYear="1934" box="[1570,1702,516,533]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="4" pageNumber="283" phylum="Chordata" rank="species" species="leonhardschultzei">leonhardschultzei</taxonomicName>
</emphasis>
<emphasis id="B9146D09FF9CFFA80BDCCE966B3EFA5A" bold="true" box="[138,269,1457,1473]" pageId="4" pageNumber="283">Distribution and</emphasis>
Atlantic slopes of Sierra Northwestern edge of Pacific slopes of the Pacific slopes from West central Pacific slopes of the Pacific slopes
</paragraph>
<paragraph id="8BDFB11BFF9CFFA80BDCC9166CB7FA1A" pageId="4" pageNumber="283">
<table id="F96043BBFF9C00530BDCC9166C88FA0D" box="[138,1723,561,1430]" gridcols="8" gridrows="11" pageId="4" pageNumber="283">
<tr id="3550B359FF9C00530BDCC9166C88FDCD" box="[138,1723,561,598]" gridrow="0" pageId="4" pageNumber="283">
<th id="7681DA25FF9C00530BDCC9166B1CFDCD" box="[138,303,561,598]" gridcol="0" gridrow="0" pageId="4" pageNumber="283">
<emphasis id="B9146D09FF9CFFA80BDCC9166AD5FDCD" bold="true" pageId="4" pageNumber="283">Maximum SVL in males (mm)</emphasis>
</th>
<th id="7681DA25FF9C00530A1FC9166824FDCD" box="[329,535,561,598]" gridcol="1" gridrow="0" pageId="4" pageNumber="283">36.6</th>
<th id="7681DA25FF9C00530979C91668EFFDCD" box="[559,732,561,598]" gridcol="2" gridrow="0" pageId="4" pageNumber="283">84.1</th>
<th id="7681DA25FF9C005309A3C9166992FDCD" box="[757,929,561,598]" gridcol="3" gridrow="0" pageId="4" pageNumber="283">----</th>
<th id="7681DA25FF9C005308EAC9166E47FDCD" box="[956,1140,561,598]" gridcol="4" gridrow="0" pageId="4" pageNumber="283">37.3</th>
<th id="7681DA25FF9C00530FC7C9166F08FDCD" box="[1169,1339,561,598]" gridcol="5" gridrow="0" pageId="4" pageNumber="283">35.4</th>
<th id="7681DA25FF9C00530E00C9166FC2FDCD" box="[1366,1521,561,598]" gridcol="6" gridrow="0" pageId="4" pageNumber="283">36.7</th>
<th id="7681DA25FF9C00530D5BC9166C88FDCD" box="[1549,1723,561,598]" gridcol="7" gridrow="0" pageId="4" pageNumber="283">35.1</th>
</tr>
<tr id="3550B359FF9C00530BDCC9576C88FD0D" box="[138,1723,624,662]" gridrow="1" pageId="4" pageNumber="283">
<th id="7681DA25FF9C00530BDCC9576B1CFD0D" box="[138,303,624,662]" gridcol="0" gridrow="1" pageId="4" pageNumber="283">
<emphasis id="B9146D09FF9CFFA80BDCC9566AC6FD0D" bold="true" pageId="4" pageNumber="283">Maximum SVL in females (mm)</emphasis>
</th>
<td id="7681DA25FF9C00530A1FC9576824FD0D" box="[329,535,624,662]" gridcol="1" gridrow="1" pageId="4" pageNumber="283">57.6</td>
<td id="7681DA25FF9C00530979C95768EFFD0D" box="[559,732,624,662]" gridcol="2" gridrow="1" pageId="4" pageNumber="283">Unknown</td>
<td id="7681DA25FF9C005309A3C9576992FD0D" box="[757,929,624,662]" gridcol="3" gridrow="1" pageId="4" pageNumber="283">26.2</td>
<td id="7681DA25FF9C005308EAC9576E47FD0D" box="[956,1140,624,662]" gridcol="4" gridrow="1" pageId="4" pageNumber="283">53.3</td>
<td id="7681DA25FF9C00530FC7C9576F08FD0D" box="[1169,1339,624,662]" gridcol="5" gridrow="1" pageId="4" pageNumber="283">39.6</td>
<td id="7681DA25FF9C00530E00C9576FC2FD0D" box="[1366,1521,624,662]" gridcol="6" gridrow="1" pageId="4" pageNumber="283">37</td>
<td id="7681DA25FF9C00530D5BC9576C88FD0D" box="[1549,1723,624,662]" gridcol="7" gridrow="1" pageId="4" pageNumber="283">43.4</td>
</tr>
<tr id="3550B359FF9C00530BDCC9976C88FD70" box="[138,1723,688,747]" gridrow="2" pageId="4" pageNumber="283">
<th id="7681DA25FF9C00530BDCC9976B1CFD70" box="[138,303,688,747]" gridcol="0" gridrow="2" pageId="4" pageNumber="283">
<emphasis id="B9146D09FF9CFFA80BDCC9966AC9FD5A" bold="true" box="[138,250,689,705]" pageId="4" pageNumber="283">Iris coloration</emphasis>
</th>
<td id="7681DA25FF9C00530A1FC9976824FD70" box="[329,535,688,747]" gridcol="1" gridrow="2" pageId="4" pageNumber="283">Whitish or silver with reddish brown reticulations</td>
<td id="7681DA25FF9C00530979C99768EFFD70" box="[559,732,688,747]" gridcol="2" gridrow="2" pageId="4" pageNumber="283">Gold with black reticulations</td>
<td id="7681DA25FF9C005309A3C9976992FD70" box="[757,929,688,747]" gridcol="3" gridrow="2" pageId="4" pageNumber="283">Red</td>
<td id="7681DA25FF9C005308EAC9976E47FD70" box="[956,1140,688,747]" gridcol="4" gridrow="2" pageId="4" pageNumber="283">Reddish-bronze</td>
<td id="7681DA25FF9C00530FC7C9976F08FD70" box="[1169,1339,688,747]" gridcol="5" gridrow="2" pageId="4" pageNumber="283">Dull grayish bronze</td>
<td id="7681DA25FF9C00530E00C9976FC2FD70" box="[1366,1521,688,747]" gridcol="6" gridrow="2" pageId="4" pageNumber="283">Red</td>
<td id="7681DA25FF9C00530D5BC9976C88FD70" box="[1549,1723,688,747]" gridcol="7" gridrow="2" pageId="4" pageNumber="283">Reddish bronze with fine black flecks or reticulations</td>
</tr>
<tr id="3550B359FF9C00530BDCC8216C88FC8D" box="[138,1723,774,790]" gridrow="3" pageId="4" pageNumber="283">
<th id="7681DA25FF9C00530BDCC8216B1CFC8D" box="[138,303,774,790]" gridcol="0" gridrow="3" pageId="4" pageNumber="283">
<emphasis id="B9146D09FF9CFFA80BDCC8216B1DFC8D" bold="true" box="[138,302,774,790]" pageId="4" pageNumber="283">Ventrolateral glands</emphasis>
</th>
<td id="7681DA25FF9C00530A1FC8216824FC8D" box="[329,535,774,790]" gridcol="1" gridrow="3" pageId="4" pageNumber="283">Absent</td>
<td id="7681DA25FF9C00530979C82168EFFC8D" box="[559,732,774,790]" gridcol="2" gridrow="3" pageId="4" pageNumber="283">Absent</td>
<td id="7681DA25FF9C005309A3C8216992FC8D" box="[757,929,774,790]" gridcol="3" gridrow="3" pageId="4" pageNumber="283">---------</td>
<td id="7681DA25FF9C005308EAC8216E47FC8D" box="[956,1140,774,790]" gridcol="4" gridrow="3" pageId="4" pageNumber="283">Well developed</td>
<td id="7681DA25FF9C00530FC7C8216F08FC8D" box="[1169,1339,774,790]" gridcol="5" gridrow="3" pageId="4" pageNumber="283">Well developed</td>
<td id="7681DA25FF9C00530E00C8216FC2FC8D" box="[1366,1521,774,790]" gridcol="6" gridrow="3" pageId="4" pageNumber="283">Well developed</td>
<td id="7681DA25FF9C00530D5BC8216C88FC8D" box="[1549,1723,774,790]" gridcol="7" gridrow="3" pageId="4" pageNumber="283">Well developed</td>
</tr>
<tr id="3550B359FF9C00530BDCC8176C88FCDA" box="[138,1723,816,833]" gridrow="4" pageId="4" pageNumber="283">
<th id="7681DA25FF9C00530BDCC8176B1CFCDA" box="[138,303,816,833]" gridcol="0" gridrow="4" pageId="4" pageNumber="283">
<emphasis id="B9146D09FF9CFFA80BDCC8166ADBFCDA" bold="true" box="[138,232,817,833]" pageId="4" pageNumber="283">Chest gland</emphasis>
</th>
<td id="7681DA25FF9C00530A1FC8176824FCDA" box="[329,535,816,833]" gridcol="1" gridrow="4" pageId="4" pageNumber="283">Present</td>
<td id="7681DA25FF9C00530979C81768EFFCDA" box="[559,732,816,833]" gridcol="2" gridrow="4" pageId="4" pageNumber="283">Absent</td>
<td id="7681DA25FF9C005309A3C8176992FCDA" box="[757,929,816,833]" gridcol="3" gridrow="4" pageId="4" pageNumber="283">Absent</td>
<td id="7681DA25FF9C005308EAC8176E47FCDA" box="[956,1140,816,833]" gridcol="4" gridrow="4" pageId="4" pageNumber="283">Absent</td>
<td id="7681DA25FF9C00530FC7C8176F08FCDA" box="[1169,1339,816,833]" gridcol="5" gridrow="4" pageId="4" pageNumber="283">Absent</td>
<td id="7681DA25FF9C00530E00C8176FC2FCDA" box="[1366,1521,816,833]" gridcol="6" gridrow="4" pageId="4" pageNumber="283">Absent</td>
<td id="7681DA25FF9C00530D5BC8176C88FCDA" box="[1549,1723,816,833]" gridcol="7" gridrow="4" pageId="4" pageNumber="283">Absent</td>
</tr>
<tr id="3550B359FF9C00530BDCC8576C88FC30" box="[138,1723,880,939]" gridrow="5" pageId="4" pageNumber="283">
<th id="7681DA25FF9C00530BDCC8576B1CFC30" box="[138,303,880,939]" gridcol="0" gridrow="5" pageId="4" pageNumber="283">
<emphasis id="B9146D09FF9CFFA80BDCC8566B1CFC1A" bold="true" box="[138,303,881,897]" pageId="4" pageNumber="283">Nuptial excrescences</emphasis>
</th>
<td id="7681DA25FF9C00530A1FC8576824FC30" box="[329,535,880,939]" gridcol="1" gridrow="5" pageId="4" pageNumber="283">Dark keratinized nuptial excrescence of several hundred tiny spines</td>
<td id="7681DA25FF9C00530979C85768EFFC30" box="[559,732,880,939]" gridcol="2" gridrow="5" pageId="4" pageNumber="283">Unknown</td>
<td id="7681DA25FF9C005309A3C8576992FC30" box="[757,929,880,939]" gridcol="3" gridrow="5" pageId="4" pageNumber="283">---------</td>
<td id="7681DA25FF9C005308EAC8576E47FC30" box="[956,1140,880,939]" gridcol="4" gridrow="5" pageId="4" pageNumber="283">Cluster of about 44 to 143 small spines</td>
<td id="7681DA25FF9C00530FC7C8576F08FC30" box="[1169,1339,880,939]" gridcol="5" gridrow="5" pageId="4" pageNumber="283">35 to 66 sharply pointed spines</td>
<td id="7681DA25FF9C00530E00C8576FC2FC30" box="[1366,1521,880,939]" gridcol="6" gridrow="5" pageId="4" pageNumber="283">Enlarged</td>
<td id="7681DA25FF9C00530D5BC8576C88FC30" box="[1549,1723,880,939]" gridcol="7" gridrow="5" pageId="4" pageNumber="283">Cluster of 24 to 80 spines</td>
</tr>
<tr id="3550B359FF9C00530BDCC8E16C88FB8D" box="[138,1723,966,1046]" gridrow="6" pageId="4" pageNumber="283">
<th id="7681DA25FF9C00530BDCC8E16B1CFB8D" box="[138,303,966,1046]" gridcol="0" gridrow="6" pageId="4" pageNumber="283">
<emphasis id="B9146D09FF9CFFA80BDCC8E16B20FC4D" bold="true" box="[138,275,966,982]" pageId="4" pageNumber="283">Dorsal coloration</emphasis>
</th>
<td id="7681DA25FF9C00530A1FC8E16824FB8D" box="[329,535,966,1046]" gridcol="1" gridrow="6" pageId="4" pageNumber="283">Lime-green with a small amount of indistinct pale brown mottling</td>
<td id="7681DA25FF9C00530979C8E168EFFB8D" box="[559,732,966,1046]" gridcol="2" gridrow="6" pageId="4" pageNumber="283">Reddish brown with indistinct dark brown smudging of mottlings</td>
<td id="7681DA25FF9C005309A3C8E16992FB8D" box="[757,929,966,1046]" gridcol="3" gridrow="6" pageId="4" pageNumber="283">Olive green to gray with irregular darker green blotches</td>
<td id="7681DA25FF9C005308EAC8E16E47FB8D" box="[956,1140,966,1046]" gridcol="4" gridrow="6" pageId="4" pageNumber="283">Reddish brown with small indistinct darker brown markings</td>
<td id="7681DA25FF9C00530FC7C8E16F08FB8D" box="[1169,1339,966,1046]" gridcol="5" gridrow="6" pageId="4" pageNumber="283">Yellowish tan with small irregular brown markings</td>
<td id="7681DA25FF9C00530E00C8E16FC2FB8D" box="[1366,1521,966,1046]" gridcol="6" gridrow="6" pageId="4" pageNumber="283">Reddish brown or dull olive-green</td>
<td id="7681DA25FF9C00530D5BC8E16C88FB8D" box="[1549,1723,966,1046]" gridcol="7" gridrow="6" pageId="4" pageNumber="283">Pale tan, yellowishbrown, or pale reddishbrown with irregular, darker brown blotches</td>
</tr>
<tr id="3550B359FF9C00530BDCCF166C88FBCD" box="[138,1723,1073,1110]" gridrow="7" pageId="4" pageNumber="283">
<th id="7681DA25FF9C00530BDCCF166B1CFBCD" box="[138,303,1073,1110]" gridcol="0" gridrow="7" pageId="4" pageNumber="283">
<emphasis id="B9146D09FF9CFFA80BDCCF166ADCFBDA" bold="true" box="[138,239,1073,1089]" pageId="4" pageNumber="283">Labial stripe</emphasis>
</th>
<td id="7681DA25FF9C00530A1FCF166824FBCD" box="[329,535,1073,1110]" gridcol="1" gridrow="7" pageId="4" pageNumber="283">White fine on upper lip</td>
<td id="7681DA25FF9C00530979CF1668EFFBCD" box="[559,732,1073,1110]" gridcol="2" gridrow="7" pageId="4" pageNumber="283">Absent</td>
<td id="7681DA25FF9C005309A3CF166992FBCD" box="[757,929,1073,1110]" gridcol="3" gridrow="7" pageId="4" pageNumber="283">Narrow discontinuous with line</td>
<td id="7681DA25FF9C005308EACF166E47FBCD" box="[956,1140,1073,1110]" gridcol="4" gridrow="7" pageId="4" pageNumber="283">White line along the edge of the upper lip</td>
<td id="7681DA25FF9C00530FC7CF166F08FBCD" box="[1169,1339,1073,1110]" gridcol="5" gridrow="7" pageId="4" pageNumber="283">Fine on upper lip</td>
<td id="7681DA25FF9C00530E00CF166FC2FBCD" box="[1366,1521,1073,1110]" gridcol="6" gridrow="7" pageId="4" pageNumber="283">White stripe on upper lip</td>
<td id="7681DA25FF9C00530D5BCF166C88FBCD" box="[1549,1723,1073,1110]" gridcol="7" gridrow="7" pageId="4" pageNumber="283">Absent</td>
</tr>
<tr id="3550B359FF9C00530BDCCF566C88FB5A" box="[138,1723,1137,1217]" gridrow="8" pageId="4" pageNumber="283">
<th id="7681DA25FF9C00530BDCCF566B1CFB5A" box="[138,303,1137,1217]" gridcol="0" gridrow="8" pageId="4" pageNumber="283">
<emphasis id="B9146D09FF9CFFA80BDCCF566AF3FB1A" bold="true" box="[138,192,1137,1153]" pageId="4" pageNumber="283">Flanks</emphasis>
</th>
<td id="7681DA25FF9C00530A1FCF566824FB5A" box="[329,535,1137,1217]" gridcol="1" gridrow="8" pageId="4" pageNumber="283">Reticulate pattern</td>
<td id="7681DA25FF9C00530979CF5668EFFB5A" box="[559,732,1137,1217]" gridcol="2" gridrow="8" pageId="4" pageNumber="283">Whitish with brown markings</td>
<td id="7681DA25FF9C005309A3CF566992FB5A" box="[757,929,1137,1217]" gridcol="3" gridrow="8" pageId="4" pageNumber="283">Large white spots more or less confluent, bordered by irregular blackish spots</td>
<td id="7681DA25FF9C005308EACF566E47FB5A" box="[956,1140,1137,1217]" gridcol="4" gridrow="8" pageId="4" pageNumber="283">White lateral stripe</td>
<td id="7681DA25FF9C00530FC7CF566F08FB5A" box="[1169,1339,1137,1217]" gridcol="5" gridrow="8" pageId="4" pageNumber="283">Marked by dark brown spots</td>
<td id="7681DA25FF9C00530E00CF566FC2FB5A" box="[1366,1521,1137,1217]" gridcol="6" gridrow="8" pageId="4" pageNumber="283">Creamy yellow stripe extend the length of the flank</td>
<td id="7681DA25FF9C00530D5BCF566C88FB5A" box="[1549,1723,1137,1217]" gridcol="7" gridrow="8" pageId="4" pageNumber="283">Creamy tan with irregular brown spots or mottling</td>
</tr>
<tr id="3550B359FF9C00530BDCCFFC6C88FAB0" box="[138,1723,1243,1323]" gridrow="9" pageId="4" pageNumber="283">
<th id="7681DA25FF9C00530BDCCFFC6B1CFAB0" box="[138,303,1243,1323]" gridcol="0" gridrow="9" pageId="4" pageNumber="283">
<emphasis id="B9146D09FF9CFFA80BDCCFFB6AD2FB77" bold="true" box="[138,225,1244,1260]" pageId="4" pageNumber="283">Tarsal fold</emphasis>
</th>
<td id="7681DA25FF9C00530A1FCFFC6824FAB0" box="[329,535,1243,1323]" gridcol="1" gridrow="9" pageId="4" pageNumber="283">Narrow, low, ex- tending full length of tarsus</td>
<td id="7681DA25FF9C00530979CFFC68EFFAB0" box="[559,732,1243,1323]" gridcol="2" gridrow="9" pageId="4" pageNumber="283">Dermal fold extends from the heel along the ventrolateral edge of the tarsus</td>
<td id="7681DA25FF9C005309A3CFFC6992FAB0" box="[757,929,1243,1323]" gridcol="3" gridrow="9" pageId="4" pageNumber="283">Fain present</td>
<td id="7681DA25FF9C005308EACFFC6E47FAB0" box="[956,1140,1243,1323]" gridcol="4" gridrow="9" pageId="4" pageNumber="283">A low rounded extends the full length</td>
<td id="7681DA25FF9C00530FC7CFFC6F08FAB0" box="[1169,1339,1243,1323]" gridcol="5" gridrow="9" pageId="4" pageNumber="283">Distinct, low and rounded extend the full length of tarsus</td>
<td id="7681DA25FF9C00530E00CFFC6FC2FAB0" box="[1366,1521,1243,1323]" gridcol="6" gridrow="9" pageId="4" pageNumber="283">Low on the distal half of the tarsus</td>
<td id="7681DA25FF9C00530D5BCFFC6C88FAB0" box="[1549,1723,1243,1323]" gridcol="7" gridrow="9" pageId="4" pageNumber="283">Row of tubercles forms a indistinct fold</td>
</tr>
<tr id="3550B359FF9C00530BDCCE616C88FA0D" box="[138,1723,1350,1430]" gridrow="10" pageId="4" pageNumber="283">
<th id="7681DA25FF9C00530BDCCE616B1CFA0D" box="[138,303,1350,1430]" gridcol="0" gridrow="10" pageId="4" pageNumber="283">
<emphasis id="B9146D09FF9CFFA80BDCCE606AE3FACC" bold="true" box="[138,208,1351,1367]" pageId="4" pageNumber="283">Forearm</emphasis>
</th>
<td id="7681DA25FF9C00530A1FCE616824FA0D" box="[329,535,1350,1430]" gridcol="1" gridrow="10" pageId="4" pageNumber="283">Distinctly raised dermal ridge punctuated with enlarged tubercles</td>
<td id="7681DA25FF9C00530979CE6168EFFA0D" box="[559,732,1350,1430]" gridcol="2" gridrow="10" pageId="4" pageNumber="283">Poorly developed dermal fold</td>
<td id="7681DA25FF9C005309A3CE616992FA0D" box="[757,929,1350,1430]" gridcol="3" gridrow="10" pageId="4" pageNumber="283">Inconspicuous dermal fold, but not enlarged tubercles</td>
<td id="7681DA25FF9C005308EACE616E47FA0D" box="[956,1140,1350,1430]" gridcol="4" gridrow="10" pageId="4" pageNumber="283">A row of small tubercles</td>
<td id="7681DA25FF9C00530FC7CE616F08FA0D" box="[1169,1339,1350,1430]" gridcol="5" gridrow="10" pageId="4" pageNumber="283">Row of low tubercles</td>
<td id="7681DA25FF9C00530E00CE616FC2FA0D" box="[1366,1521,1350,1430]" gridcol="6" gridrow="10" pageId="4" pageNumber="283">Row of tubercles fused in some specimens into a low, thick dermal fold</td>
<td id="7681DA25FF9C00530D5BCE616C88FA0D" box="[1549,1723,1350,1430]" gridcol="7" gridrow="10" pageId="4" pageNumber="283">Low and rounded extends the full length of tarsus</td>
</tr>
</table>
</paragraph>
<paragraph id="8BDFB11BFF9CFFA80BDCCEE06E65F9B7" blockId="4.[138,1744,1457,1650]" pageId="4" pageNumber="283">
<materialsCitation id="3B08BB46FF9CFFA80BDCCEE06E65F9B7" ID-GBIF-Occurrence="1606510479" collectorName="Juarez &amp; Sierra &amp; Sierra Madre &amp; Sur &amp; de Talamanca" country="Mexico" county="eastern Costa Rica Sur" elevation="747" location="Sierra de Talamanca" municipality="central Nicaragua" pageId="4" pageNumber="283" specimenCount="1" stateProvince="Oaxaca, Mexico">
<emphasis id="B9146D09FF9CFFA80BDCCEE06AE2FA4C" bold="true" box="[138,209,1479,1495]" pageId="4" pageNumber="283">elevation</emphasis>
<collectorName id="2695D4CDFF9CFFA80A1BCEE16B4DFA4D" box="[333,382,1478,1494]" pageId="4" pageNumber="283">Juarez</collectorName>
,
<collectingRegion id="49A47FF9FF9CFFA80AD0CEE16833FA4D" box="[390,512,1478,1494]" isEnumeration="true" pageId="4" pageNumber="283">Oaxaca, México</collectingRegion>
at the
<collectorName id="2695D4CDFF9CFFA80933CEE168A2FA4D" box="[613,657,1478,1494]" pageId="4" pageNumber="283">Sierra</collectorName>
de los
<collectorName id="2695D4CDFF9CFFA809ABCEE1696DFA4D" box="[765,862,1478,1494]" pageId="4" pageNumber="283">Sierra Madre</collectorName>
del
<collectorName id="2695D4CDFF9CFFA80828CEE169ABFA4D" box="[894,920,1478,1494]" pageId="4" pageNumber="283">Sur</collectorName>
southeastern
<collectingRegion id="49A47FF9FF9CFFA80F7ACEE16ED8FA4D" box="[1068,1259,1478,1494]" isEnumeration="true" pageId="4" pageNumber="283">
<collectingRegion id="49A47FF9FF9CFFA80F7ACEE16E56FA4D" box="[1068,1125,1478,1494]" country="Mexico" name="Mexico" pageId="4" pageNumber="283">
<collectingCountry id="F377F18BFF9CFFA80F7ACEE16E56FA4D" box="[1068,1125,1478,1494]" name="Mexico" pageId="4" pageNumber="283">México</collectingCountry>
</collectingRegion>
<collectingRegion id="49A47FF9FF9CFFA80FCDCEE16ED8FA4D" box="[1179,1259,1478,1494]" country="Guatemala" name="Guatemala" pageId="4" pageNumber="283">
<collectingCountry id="F377F18BFF9CFFA80FCDCEE16ED8FA4D" box="[1179,1259,1478,1494]" name="Guatemala" pageId="4" pageNumber="283">Guatemala</collectingCountry>
</collectingRegion>
</collectingRegion>
to north-
<location id="8EBFE7C0FF9CFFA80E03CEE16FC3FA4D" LSID="urn:lsid:plazi:treatment:03C9000DFF99FFAB0BC1CEED6828F896:8EBFE7C0FF9CFFA80E03CEE16FC3FA4D" box="[1365,1520,1478,1494]" country="Mexico" county="eastern Costa Rica Sur" municipality="central Nicaragua" name="Sierra de Talamanca" pageId="4" pageNumber="283" stateProvince="Oaxaca, Mexico">
Sierra
<collectorName id="2695D4CDFF9CFFA80ED0CEE16FC3FA4D" box="[1414,1520,1478,1494]" pageId="4" pageNumber="283">de Talamanca</collectorName>
</location>
, of the
<collectorName id="2695D4CDFF9CFFA80D68CEE16CADFA4D" box="[1598,1694,1478,1494]" pageId="4" pageNumber="283">Sierra Madre</collectorName>
del
<elevation id="004D5628FF9CFFA80A27CEFB6BF5FA77" box="[369,454,1499,1516]" metricMagnitude="2" metricUnit="m" metricValue="7.47" metricValueMax="9.0" metricValueMin="5.9399999999999995" pageId="4" pageNumber="283" unit="m" value="747.0" valueMax="900.0" valueMin="594.0">
<quantity id="4C981CFEFF9CFFA80A27CEFB6BF5FA77" box="[369,454,1499,1516]" metricMagnitude="2" metricUnit="m" metricValue="7.47" metricValueMax="9.0" metricValueMin="5.9399999999999995" pageId="4" pageNumber="283" unit="m" value="747.0" valueMax="900.0" valueMin="594.0">
594
<emphasis id="B9146D09FF9CFFA80ADBCEFC6BA5FA77" box="[397,406,1499,1516]" italics="true" pageId="4" pageNumber="283"></emphasis>
900 m
</quantity>
</elevation>
elev. Cuchumatanes, of
<collectingRegion id="49A47FF9FF9CFFA80843CEFB696BFA77" box="[789,856,1500,1516]" country="Mexico" name="Guerrero" pageId="4" pageNumber="283">Guerrero</collectingRegion>
at 700
<emphasis id="B9146D09FF9CFFA808DDCEFC69A7FA77" box="[907,916,1499,1516]" italics="true" pageId="4" pageNumber="283"></emphasis>
(
<collectingRegion id="49A47FF9FF9CFFA8089ACEFB6E3AFA77" box="[972,1033,1500,1516]" country="Mexico" name="Chiapas" pageId="4" pageNumber="283">Chiapas</collectingRegion>
and extreme
<collectingMunicipality id="6BBB2B61FF9CFFA80FCDCEFB6F2DFA77" box="[1179,1310,1500,1516]" pageId="4" pageNumber="283">central Nicaragua</collectingMunicipality>
at
<collectingCounty id="62BEC997FF9CFFA80E09CEFB6C09FA77" box="[1375,1594,1500,1516]" pageId="4" pageNumber="283">eastern Costa Rica Sur</collectingCounty>
in
<collectingRegion id="49A47FF9FF9CFFA80D04CEFB6CBAFA77" box="[1618,1673,1500,1516]" country="Mexico" name="Oaxaca" pageId="4" pageNumber="283">Oaxaca</collectingRegion>
and Huehetenango,
<elevation id="004D5628FF9CFFA80849CED6697DF99A" box="[799,846,1521,1537]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="4" pageNumber="283" unit="m" value="950.0">
<quantity id="4C981CFEFF9CFFA80849CED6697DF99A" box="[799,846,1521,1537]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="4" pageNumber="283" unit="m" value="950.0">950 m</quantity>
</elevation>
elev.
<collectingRegion id="49A47FF9FF9CFFA80898CED66E34F99A" box="[974,1031,1521,1537]" country="Mexico" name="Oaxaca" pageId="4" pageNumber="283">Oaxaca</collectingRegion>
) and pacific
<elevation id="004D5628FF9CFFA80FF5CED66F32F99A" box="[1187,1281,1521,1538]" metricMagnitude="3" metricUnit="m" metricValue="1.205" metricValueMax="2.07" metricValueMin="0.33999999999999997" pageId="4" pageNumber="283" unit="m" value="1205.0" valueMax="2070.0" valueMin="340.0">
<quantity id="4C981CFEFF9CFFA80FF5CED66F32F99A" box="[1187,1281,1521,1538]" metricMagnitude="3" metricUnit="m" metricValue="1.205" metricValueMax="2.07" metricValueMin="0.33999999999999997" pageId="4" pageNumber="283" unit="m" value="1205.0" valueMax="2070.0" valueMin="340.0">
340
<emphasis id="B9146D09FF9CFFA80FE9CED66EFBF999" box="[1215,1224,1521,1538]" italics="true" pageId="4" pageNumber="283"></emphasis>
2070 m
</quantity>
</elevation>
elev. and western
<collectingRegion id="49A47FF9FF9CFFA80EE3CED66CADF99A" box="[1461,1694,1521,1537]" isEnumeration="true" pageId="4" pageNumber="283">
<collectingRegion id="49A47FF9FF9CFFA80EE3CED66FC3F99A" box="[1461,1520,1521,1537]" country="Panama" name="Panama" pageId="4" pageNumber="283">Panamá</collectingRegion>
<collectingRegion id="49A47FF9FF9CFFA80D4FCED66C6DF99A" box="[1561,1630,1521,1537]" country="Mexico" name="Guerrero" pageId="4" pageNumber="283">Guerrero</collectingRegion>
, Mexico
</collectingRegion>
at
<collectingCountry id="F377F18BFF9CFFA80960CD2168B5F98D" box="[566,646,1542,1558]" name="Guatemala" pageId="4" pageNumber="283">Guatemala</collectingCountry>
at
<elevation id="004D5628FF9CFFA809CBCD2168E5F98D" box="[669,726,1542,1558]" metricMagnitude="3" metricUnit="m" metricValue="1.27" pageId="4" pageNumber="283" unit="m" value="1270.0">
<quantity id="4C981CFEFF9CFFA809CBCD2168E5F98D" box="[669,726,1542,1558]" metricMagnitude="3" metricUnit="m" metricValue="1.27" pageId="4" pageNumber="283" unit="m" value="1270.0">1270 m</quantity>
</elevation>
slopes of
<collectingCountry id="F377F18BFF9CFFA80F47CD216E50F98D" box="[1041,1123,1542,1558]" name="Guatemala" pageId="4" pageNumber="283">Guatemala</collectingCountry>
, at
<elevation id="004D5628FF9CFFA80E3CCD216FFBF98D" box="[1386,1480,1542,1559]" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="1.6" metricValueMin="0.7" pageId="4" pageNumber="283" unit="m" value="1150.0" valueMax="1600.0" valueMin="700.0">
<quantity id="4C981CFEFF9CFFA80E3CCD216FFBF98D" box="[1386,1480,1542,1559]" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="1.6" metricValueMin="0.7" pageId="4" pageNumber="283" unit="m" value="1150.0" valueMax="1600.0" valueMin="700.0">
700
<emphasis id="B9146D09FF9CFFA80ED0CD216FBCF98C" box="[1414,1423,1542,1559]" italics="true" pageId="4" pageNumber="283"></emphasis>
1600 m
</quantity>
</elevation>
elev.
<elevation id="004D5628FF9CFFA80D77CD216C4CF98D" box="[1569,1663,1542,1559]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="2.0" metricValueMin="0.7" pageId="4" pageNumber="283" unit="m" value="1350.0" valueMax="2000.0" valueMin="700.0">
<quantity id="4C981CFEFF9CFFA80D77CD216C4CF98D" box="[1569,1663,1542,1559]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="2.0" metricValueMin="0.7" pageId="4" pageNumber="283" unit="m" value="1350.0" valueMax="2000.0" valueMin="700.0">
700
<emphasis id="B9146D09FF9CFFA80D6BCD216C75F98C" box="[1597,1606,1542,1559]" italics="true" pageId="4" pageNumber="283"></emphasis>
2000 m
</quantity>
</elevation>
elev. elev.
<collectingCountry id="F377F18BFF9CFFA8088CCD3B6E11F9B7" box="[986,1058,1564,1580]" name="Honduras" pageId="4" pageNumber="283">Honduras</collectingCountry>
and
<collectingCountry id="F377F18BFF9CFFA80F10CD3B6E65F9B7" box="[1094,1110,1564,1580]" name="El Salvador" pageId="4" pageNumber="283">El</collectingCountry>
</materialsCitation>
</paragraph>
<paragraph id="8BDFB11BFF9CFFA808EBCD166E40F9DA" blockId="4.[138,1744,1457,1650]" box="[957,1139,1585,1602]" pageId="4" pageNumber="283">
Salvador at
<quantity id="4C981CFEFF9CFFA80F40CD166E40F9DA" box="[1046,1139,1585,1602]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="12.0" metricValueMin="6.0" pageId="4" pageNumber="283" unit="m" value="900.0" valueMax="1200.0" valueMin="600.0">
600
<emphasis id="B9146D09FF9CFFA80F64CD166E08F9D9" box="[1074,1083,1585,1602]" italics="true" pageId="4" pageNumber="283"></emphasis>
1200 m
</quantity>
</paragraph>
<paragraph id="8BDFB11BFF9CFFA80F50CD616E18F9CD" blockId="4.[138,1744,1457,1650]" box="[1030,1067,1606,1622]" pageId="4" pageNumber="283">elev.</paragraph>
<paragraph id="8BDFB11BFF9CFFAA0E97CD7A6892FE01" blockId="4.[138,1744,1457,1650]" lastBlockId="6.[151,1437,151,1070]" lastPageId="6" lastPageNumber="285" pageId="4" pageNumber="283">
<emphasis id="B9146D09FF9CFFA80E97CD7A6CE3F9E9" box="[1473,1744,1629,1650]" italics="true" pageId="4" pageNumber="283">…continued on the next page</emphasis>
In life, dorsum of head, body, forelimbs, and hind limbs green with brown blotches and numerous irregular white spots concentrated on posterior part of dorsum; well-defined white stripe along upper jaw; flanks mottled with irregular white spots on pale brown background; narrow white stripe on ventrolateral edge of forearm and hand, and on ventrolateral edge of tarsus and foot; brown stripe parallel to white stripe on forearm, extending onto Finger IV; similar brown stripe on tarsus. White stripe above cloacal opening; brown patch with a few small white spots below cloacal opening. Belly, chin, chest, and ventral surfaces of limbs white. Ventral surface of Finger IV, and webbing between Fingers III and IV, dark brown; and all ventral surface of foot, including webbing, dark brown. Iris pale pink with brown reticulations.
</paragraph>
<paragraph id="8BDFB11BFF9DFFA90B2BCA9F6CFFFA61" pageId="5" pageNumber="284">
<table id="F96043BBFF9D00530B2BCA9F6CC4F9E0" box="[125,1783,440,1659]" gridcols="8" gridrows="12" pageId="5" pageNumber="284">
<tr id="3550B359FF9D00530B2BCA9F6CC4FE62" box="[125,1783,440,505]" gridrow="0" pageId="5" pageNumber="284">
<th id="7681DA25FF9D00530B2BCA9F6B00FE62" box="[125,307,440,505]" gridcol="0" gridrow="0" pageId="5" pageNumber="284">
<emphasis id="B9146D09FF9DFFA90B2BCA9F6AFEFE57" bold="true" box="[125,205,440,460]" pageId="5" pageNumber="284">Table 2.</emphasis>
Continued
</th>
<th id="7681DA25FF9D00530A69CA9F6BC9FE62" box="[319,506,440,505]" gridcol="1" gridrow="0" pageId="5" pageNumber="284">
<taxonomicName id="4C60CA98FF9DFFA90AC2CAF46BE8FE63" baseAuthorityName="Tanner" baseAuthorityYear="1957" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="284" phylum="Chordata" rank="species" species="macrotympanum">
<emphasis id="B9146D09FF9DFFA90AC2CAF46BE8FE63" italics="true" pageId="5" pageNumber="284">P. macrotympanum</emphasis>
</taxonomicName>
</th>
<th id="7681DA25FF9D00530942CA9F68FCFE62" box="[532,719,440,505]" gridcol="2" gridrow="0" pageId="5" pageNumber="284">
<taxonomicName id="4C60CA98FF9DFFA9093CCAF468A3FE62" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="284" phylum="Chordata" rank="species" species="panchoi">
<emphasis id="B9146D09FF9DFFA9093CCAF468A3FE62" italics="true" pageId="5" pageNumber="284">P. panchoi</emphasis>
</taxonomicName>
</th>
<th id="7681DA25FF9D005309BBCA9F6980FE62" box="[749,947,440,505]" gridcol="3" gridrow="0" pageId="5" pageNumber="284">
<taxonomicName id="4C60CA98FF9DFFA9081ECAF469B0FE62" baseAuthorityName="Mertens" baseAuthorityYear="1952" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="284" phylum="Chordata" rank="species" species="salvadorensis">
<emphasis id="B9146D09FF9DFFA9081ECAF469B0FE62" italics="true" pageId="5" pageNumber="284">P. salvadorensis</emphasis>
</taxonomicName>
</th>
<th id="7681DA25FF9D00530886CA9F6E5FFE62" box="[976,1132,440,505]" gridcol="4" gridrow="0" pageId="5" pageNumber="284">
<taxonomicName id="4C60CA98FF9DFFA90F40CAF46E62FE63" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="284" phylum="Chordata" rank="species" species="sanctaecrucis">
<emphasis id="B9146D09FF9DFFA90F40CAF46E62FE63" italics="true" pageId="5" pageNumber="284">P. sanctaecrucis</emphasis>
</taxonomicName>
</th>
<th id="7681DA25FF9D00530FD0CA9F6F03FE62" box="[1158,1328,440,505]" gridcol="5" gridrow="0" pageId="5" pageNumber="284">
<taxonomicName id="4C60CA98FF9DFFA90F85CAF46F37FE62" baseAuthorityName="Schmidt" baseAuthorityYear="1936" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="284" phylum="Chordata" rank="species" species="spinipollex">
<emphasis id="B9146D09FF9DFFA90F85CAF46F37FE62" italics="true" pageId="5" pageNumber="284">P. spinipollex</emphasis>
</taxonomicName>
</th>
<th id="7681DA25FF9D00530E1CCA9F6C22FE62" box="[1354,1553,440,505]" gridcol="6" gridrow="0" pageId="5" pageNumber="284">
<taxonomicName id="4C60CA98FF9DFFA90EF0CAF46FE3FE62" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="284" phylum="Chordata" rank="species" species="zophodes">
<emphasis id="B9146D09FF9DFFA90EF0CAF46FE3FE62" italics="true" pageId="5" pageNumber="284">P. zophodes</emphasis>
</taxonomicName>
</th>
<th id="7681DA25FF9D00530D7DCA9F6CC4FE62" box="[1579,1783,440,505]" gridcol="7" gridrow="0" pageId="5" pageNumber="284">
<taxonomicName id="4C60CA98FF9DFFA90DDCCAF46C9BFE62" authorityName="Caviedes-Solís" authorityYear="2017" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="5" pageNumber="284" phylum="Chordata" rank="species" species="zoque">
<emphasis id="B9146D09FF9DFFA90DDCCAF46C9BFE62" bold="true" italics="true" pageId="5" pageNumber="284">P. zoque</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="3550B359FF9D00530B2BC9336CC4FDA1" box="[125,1783,532,570]" gridrow="1" pageId="5" pageNumber="284">
<th id="7681DA25FF9D00530B2BC9336B00FDA1" box="[125,307,532,570]" gridcol="0" gridrow="1" pageId="5" pageNumber="284">
<emphasis id="B9146D09FF9DFFA90B2BC9336AEAFDA1" bold="true" pageId="5" pageNumber="284">Maximum SVL in males (mm)</emphasis>
</th>
<td id="7681DA25FF9D00530A69C9336BC9FDA1" box="[319,506,532,570]" gridcol="1" gridrow="1" pageId="5" pageNumber="284">38.8</td>
<td id="7681DA25FF9D00530942C93368FCFDA1" box="[532,719,532,570]" gridcol="2" gridrow="1" pageId="5" pageNumber="284">34.2</td>
<td id="7681DA25FF9D005309BBC9336980FDA1" box="[749,947,532,570]" gridcol="3" gridrow="1" pageId="5" pageNumber="284">34.9</td>
<td id="7681DA25FF9D00530886C9336E5FFDA1" box="[976,1132,532,570]" gridcol="4" gridrow="1" pageId="5" pageNumber="284">32.5</td>
<td id="7681DA25FF9D00530FD0C9336F03FDA1" box="[1158,1328,532,570]" gridcol="5" gridrow="1" pageId="5" pageNumber="284">39.1</td>
<td id="7681DA25FF9D00530E1CC9336C22FDA1" box="[1354,1553,532,570]" gridcol="6" gridrow="1" pageId="5" pageNumber="284">37.4</td>
<td id="7681DA25FF9D00530D7DC9336CC4FDA1" box="[1579,1783,532,570]" gridcol="7" gridrow="1" pageId="5" pageNumber="284">36.4</td>
</tr>
<tr id="3550B359FF9D00530B2BC9736CC4FDE1" box="[125,1783,596,634]" gridrow="2" pageId="5" pageNumber="284">
<th id="7681DA25FF9D00530B2BC9736B00FDE1" box="[125,307,596,634]" gridcol="0" gridrow="2" pageId="5" pageNumber="284">
<emphasis id="B9146D09FF9DFFA90B2BC9736ADBFDE1" bold="true" pageId="5" pageNumber="284">Maximum SVL in females (mm)</emphasis>
</th>
<td id="7681DA25FF9D00530A69C9736BC9FDE1" box="[319,506,596,634]" gridcol="1" gridrow="2" pageId="5" pageNumber="284">44.8</td>
<td id="7681DA25FF9D00530942C97368FCFDE1" box="[532,719,596,634]" gridcol="2" gridrow="2" pageId="5" pageNumber="284">37.3</td>
<td id="7681DA25FF9D005309BBC9736980FDE1" box="[749,947,596,634]" gridcol="3" gridrow="2" pageId="5" pageNumber="284">41.8</td>
<td id="7681DA25FF9D00530886C9736E5FFDE1" box="[976,1132,596,634]" gridcol="4" gridrow="2" pageId="5" pageNumber="284">50.7</td>
<td id="7681DA25FF9D00530FD0C9736F03FDE1" box="[1158,1328,596,634]" gridcol="5" gridrow="2" pageId="5" pageNumber="284">46.1</td>
<td id="7681DA25FF9D00530E1CC9736C22FDE1" box="[1354,1553,596,634]" gridcol="6" gridrow="2" pageId="5" pageNumber="284">43.6</td>
<td id="7681DA25FF9D00530D7DC9736CC4FDE1" box="[1579,1783,596,634]" gridcol="7" gridrow="2" pageId="5" pageNumber="284">57.8</td>
</tr>
<tr id="3550B359FF9D00530B2BC9B36CC4FD54" box="[125,1783,660,719]" gridrow="3" pageId="5" pageNumber="284">
<th id="7681DA25FF9D00530B2BC9B36B00FD54" box="[125,307,660,719]" gridcol="0" gridrow="3" pageId="5" pageNumber="284">
<emphasis id="B9146D09FF9DFFA90B2BC9B36ADEFD3F" bold="true" box="[125,237,660,676]" pageId="5" pageNumber="284">Iris coloration</emphasis>
</th>
<td id="7681DA25FF9D00530A69C9B36BC9FD54" box="[319,506,660,719]" gridcol="1" gridrow="3" pageId="5" pageNumber="284">Redish-bronze</td>
<td id="7681DA25FF9D00530942C9B368FCFD54" box="[532,719,660,719]" gridcol="2" gridrow="3" pageId="5" pageNumber="284">Deep reddish orange</td>
<td id="7681DA25FF9D005309BBC9B36980FD54" box="[749,947,660,719]" gridcol="3" gridrow="3" pageId="5" pageNumber="284">Copper color with fine black flecks</td>
<td id="7681DA25FF9D00530886C9B36E5FFD54" box="[976,1132,660,719]" gridcol="4" gridrow="3" pageId="5" pageNumber="284">Pale tan, almost white with fine brown markings</td>
<td id="7681DA25FF9D00530FD0C9B36F03FD54" box="[1158,1328,660,719]" gridcol="5" gridrow="3" pageId="5" pageNumber="284">Cooper in male, and chocolate in female</td>
<td id="7681DA25FF9D00530E1CC9B36C22FD54" box="[1354,1553,660,719]" gridcol="6" gridrow="3" pageId="5" pageNumber="284">Reddish brown to cooper with fine black reticulations</td>
<td id="7681DA25FF9D00530D7DC9B36CC4FD54" box="[1579,1783,660,719]" gridcol="7" gridrow="3" pageId="5" pageNumber="284">Pale pink with brown reticulations</td>
</tr>
<tr id="3550B359FF9D00530B2BC9CD6CC4FD61" box="[125,1783,746,762]" gridrow="4" pageId="5" pageNumber="284">
<th id="7681DA25FF9D00530B2BC9CD6B00FD61" box="[125,307,746,762]" gridcol="0" gridrow="4" pageId="5" pageNumber="284">
<emphasis id="B9146D09FF9DFFA90B2BC9CD6B13FD61" bold="true" box="[125,288,746,762]" pageId="5" pageNumber="284">Ventrolateral glands</emphasis>
</th>
<td id="7681DA25FF9D00530A69C9CD6BC9FD61" box="[319,506,746,762]" gridcol="1" gridrow="4" pageId="5" pageNumber="284">Well developed</td>
<td id="7681DA25FF9D00530942C9CD68FCFD61" box="[532,719,746,762]" gridcol="2" gridrow="4" pageId="5" pageNumber="284">Barely evident</td>
<td id="7681DA25FF9D005309BBC9CD6980FD61" box="[749,947,746,762]" gridcol="3" gridrow="4" pageId="5" pageNumber="284">Absent</td>
<td id="7681DA25FF9D00530886C9CD6E5FFD61" box="[976,1132,746,762]" gridcol="4" gridrow="4" pageId="5" pageNumber="284">Absent</td>
<td id="7681DA25FF9D00530FD0C9CD6F03FD61" box="[1158,1328,746,762]" gridcol="5" gridrow="4" pageId="5" pageNumber="284">Absent</td>
<td id="7681DA25FF9D00530E1CC9CD6C22FD61" box="[1354,1553,746,762]" gridcol="6" gridrow="4" pageId="5" pageNumber="284">Well developed</td>
<td id="7681DA25FF9D00530D7DC9CD6CC4FD61" box="[1579,1783,746,762]" gridcol="7" gridrow="4" pageId="5" pageNumber="284">Absent</td>
</tr>
<tr id="3550B359FF9D00530B2BC80D6CC4FCA1" box="[125,1783,810,826]" gridrow="5" pageId="5" pageNumber="284">
<th id="7681DA25FF9D00530B2BC80D6B00FCA1" box="[125,307,810,826]" gridcol="0" gridrow="5" pageId="5" pageNumber="284">
<emphasis id="B9146D09FF9DFFA90B2BC80D6AE8FCA1" bold="true" box="[125,219,810,826]" pageId="5" pageNumber="284">Chest gland</emphasis>
</th>
<td id="7681DA25FF9D00530A69C80D6BC9FCA1" box="[319,506,810,826]" gridcol="1" gridrow="5" pageId="5" pageNumber="284">Absent</td>
<td id="7681DA25FF9D00530942C80D68FCFCA1" box="[532,719,810,826]" gridcol="2" gridrow="5" pageId="5" pageNumber="284">Absent</td>
<td id="7681DA25FF9D005309BBC80D6980FCA1" box="[749,947,810,826]" gridcol="3" gridrow="5" pageId="5" pageNumber="284">Absent</td>
<td id="7681DA25FF9D00530886C80D6E5FFCA1" box="[976,1132,810,826]" gridcol="4" gridrow="5" pageId="5" pageNumber="284">Present</td>
<td id="7681DA25FF9D00530FD0C80D6F03FCA1" box="[1158,1328,810,826]" gridcol="5" gridrow="5" pageId="5" pageNumber="284">Absent</td>
<td id="7681DA25FF9D00530E1CC80D6C22FCA1" box="[1354,1553,810,826]" gridcol="6" gridrow="5" pageId="5" pageNumber="284">Absent</td>
<td id="7681DA25FF9D00530D7DC80D6CC4FCA1" box="[1579,1783,810,826]" gridcol="7" gridrow="5" pageId="5" pageNumber="284">Absent</td>
</tr>
<tr id="3550B359FF9D00530B2BC8736CC4FC14" box="[125,1783,852,911]" gridrow="6" pageId="5" pageNumber="284">
<th id="7681DA25FF9D00530B2BC8736B00FC14" box="[125,307,852,911]" gridcol="0" gridrow="6" pageId="5" pageNumber="284">
<emphasis id="B9146D09FF9DFFA90B2BC8736B12FCFF" bold="true" box="[125,289,852,868]" pageId="5" pageNumber="284">Nuptial excrescences</emphasis>
</th>
<td id="7681DA25FF9D00530A69C8736BC9FC14" box="[319,506,852,911]" gridcol="1" gridrow="6" pageId="5" pageNumber="284">Small horn spines (94- 100)</td>
<td id="7681DA25FF9D00530942C87368FCFC14" box="[532,719,852,911]" gridcol="2" gridrow="6" pageId="5" pageNumber="284">Moderately large, pointed, keratinous spines</td>
<td id="7681DA25FF9D005309BBC8736980FC14" box="[749,947,852,911]" gridcol="3" gridrow="6" pageId="5" pageNumber="284">Large, pointed, keratinous spines</td>
<td id="7681DA25FF9D00530886C8736E5FFC14" box="[976,1132,852,911]" gridcol="4" gridrow="6" pageId="5" pageNumber="284">A patch of minutes pointed, keratinous spines</td>
<td id="7681DA25FF9D00530FD0C8736F03FC14" box="[1158,1328,852,911]" gridcol="5" gridrow="6" pageId="5" pageNumber="284">Large, pointed, keratinous spines</td>
<td id="7681DA25FF9D00530E1CC8736C22FC14" box="[1354,1553,852,911]" gridcol="6" gridrow="6" pageId="5" pageNumber="284">More than 50 keratinized spines</td>
<td id="7681DA25FF9D00530D7DC8736CC4FC14" box="[1579,1783,852,911]" gridcol="7" gridrow="6" pageId="5" pageNumber="284">Small, nonspinous</td>
</tr>
<tr id="3550B359FF9D00530B2BC8986CC4FBA1" box="[125,1783,959,1082]" gridrow="7" pageId="5" pageNumber="284">
<th id="7681DA25FF9D00530B2BC8986B00FBA1" box="[125,307,959,1082]" gridcol="0" gridrow="7" pageId="5" pageNumber="284">
<emphasis id="B9146D09FF9DFFA90B2BC8986B35FC54" bold="true" box="[125,262,959,975]" pageId="5" pageNumber="284">Dorsal coloration</emphasis>
</th>
<td id="7681DA25FF9D00530A69C8986BC9FBA1" box="[319,506,959,1082]" gridcol="1" gridrow="7" pageId="5" pageNumber="284">Pale tan with dark flecks</td>
<td id="7681DA25FF9D00530942C89868FCFBA1" box="[532,719,959,1082]" gridcol="2" gridrow="7" pageId="5" pageNumber="284">Grayish tan to dark brown with darker gray or darker brown mottling on dorsum</td>
<td id="7681DA25FF9D005309BBC8986980FBA1" box="[749,947,959,1082]" gridcol="3" gridrow="7" pageId="5" pageNumber="284">Mottled green (male) and brown (female)</td>
<td id="7681DA25FF9D00530886C8986E5FFBA1" box="[976,1132,959,1082]" gridcol="4" gridrow="7" pageId="5" pageNumber="284">Lime green with scattered pale green dots or olive-green dots to brown with a lichenous pattern of dark brown</td>
<td id="7681DA25FF9D00530FD0C8986F03FBA1" box="[1158,1328,959,1082]" gridcol="5" gridrow="7" pageId="5" pageNumber="284">Mottled shades of olive green in males and brown dorsum with darker brown mottling with or without white flecks</td>
<td id="7681DA25FF9D00530E1CC8986C22FBA1" box="[1354,1553,959,1082]" gridcol="6" gridrow="7" pageId="5" pageNumber="284">Dark brown to dull greenish brown with or without irregular dark brown to black mottling</td>
<td id="7681DA25FF9D00530D7DC8986CC4FBA1" box="[1579,1783,959,1082]" gridcol="7" gridrow="7" pageId="5" pageNumber="284">Green with brown blotches and numerous irregular white spots concentrated on posterior part of dorsum</td>
</tr>
<tr id="3550B359FF9D00530B2BCF736CC4FBE1" box="[125,1783,1108,1146]" gridrow="8" pageId="5" pageNumber="284">
<th id="7681DA25FF9D00530B2BCF736B00FBE1" box="[125,307,1108,1146]" gridcol="0" gridrow="8" pageId="5" pageNumber="284">
<emphasis id="B9146D09FF9DFFA90B2BCF726AD1FBFE" bold="true" box="[125,226,1109,1125]" pageId="5" pageNumber="284">Labial stripe</emphasis>
</th>
<td id="7681DA25FF9D00530A69CF736BC9FBE1" box="[319,506,1108,1146]" gridcol="1" gridrow="8" pageId="5" pageNumber="284">A thin white line extends the length of upper lipe</td>
<td id="7681DA25FF9D00530942CF7368FCFBE1" box="[532,719,1108,1146]" gridcol="2" gridrow="8" pageId="5" pageNumber="284">Present</td>
<td id="7681DA25FF9D005309BBCF736980FBE1" box="[749,947,1108,1146]" gridcol="3" gridrow="8" pageId="5" pageNumber="284">Narrow white stripe on upper lipe</td>
<td id="7681DA25FF9D00530886CF736E5FFBE1" box="[976,1132,1108,1146]" gridcol="4" gridrow="8" pageId="5" pageNumber="284">Thin pale</td>
<td id="7681DA25FF9D00530FD0CF736F03FBE1" box="[1158,1328,1108,1146]" gridcol="5" gridrow="8" pageId="5" pageNumber="284">Absent</td>
<td id="7681DA25FF9D00530E1CCF736C22FBE1" box="[1354,1553,1108,1146]" gridcol="6" gridrow="8" pageId="5" pageNumber="284">------</td>
<td id="7681DA25FF9D00530D7DCF736CC4FBE1" box="[1579,1783,1108,1146]" gridcol="7" gridrow="8" pageId="5" pageNumber="284">White fine line on upper lipe</td>
</tr>
<tr id="3550B359FF9D00530B2BCFB36CC4FB7E" box="[125,1783,1172,1253]" gridrow="9" pageId="5" pageNumber="284">
<th id="7681DA25FF9D00530B2BCFB36B00FB7E" box="[125,307,1172,1253]" gridcol="0" gridrow="9" pageId="5" pageNumber="284">
<emphasis id="B9146D09FF9DFFA90B2BCFB26A80FB3E" bold="true" box="[125,179,1173,1189]" pageId="5" pageNumber="284">Flanks</emphasis>
</th>
<td id="7681DA25FF9D00530A69CFB36BC9FB7E" box="[319,506,1172,1253]" gridcol="1" gridrow="9" pageId="5" pageNumber="284">No lateral stripe</td>
<td id="7681DA25FF9D00530942CFB368FCFB7E" box="[532,719,1172,1253]" gridcol="2" gridrow="9" pageId="5" pageNumber="284">White ventrolateral stripe</td>
<td id="7681DA25FF9D005309BBCFB36980FB7E" box="[749,947,1172,1253]" gridcol="3" gridrow="9" pageId="5" pageNumber="284">Mottled silvery white and dark brown</td>
<td id="7681DA25FF9D00530886CFB36E5FFB7E" box="[976,1132,1172,1253]" gridcol="4" gridrow="9" pageId="5" pageNumber="284">Purplish band with flecks</td>
<td id="7681DA25FF9D00530FD0CFB36F03FB7E" box="[1158,1328,1172,1253]" gridcol="5" gridrow="9" pageId="5" pageNumber="284">A stripe with diffuse edges along the flank is cream anteriorly and yellow posteriorly</td>
<td id="7681DA25FF9D00530E1CCFB36C22FB7E" box="[1354,1553,1172,1253]" gridcol="6" gridrow="9" pageId="5" pageNumber="284">White with black mottling or brown</td>
<td id="7681DA25FF9D00530D7DCFB36CC4FB7E" box="[1579,1783,1172,1253]" gridcol="7" gridrow="9" pageId="5" pageNumber="284">Mottled with irregular white spots on pale brown background</td>
</tr>
<tr id="3550B359FF9D00530B2BCFD86CC4FAA1" box="[125,1783,1279,1338]" gridrow="10" pageId="5" pageNumber="284">
<th id="7681DA25FF9D00530B2BCFD86B00FAA1" box="[125,307,1279,1338]" gridcol="0" gridrow="10" pageId="5" pageNumber="284">
<emphasis id="B9146D09FF9DFFA90B2BCFD86AE7FA94" bold="true" box="[125,212,1279,1295]" pageId="5" pageNumber="284">Tarsal fold</emphasis>
</th>
<td id="7681DA25FF9D00530A69CFD86BC9FAA1" box="[319,506,1279,1338]" gridcol="1" gridrow="10" pageId="5" pageNumber="284">Weak on distal two thirds of tarsus</td>
<td id="7681DA25FF9D00530942CFD868FCFAA1" box="[532,719,1279,1338]" gridcol="2" gridrow="10" pageId="5" pageNumber="284">Distict, no row of tubercles</td>
<td id="7681DA25FF9D005309BBCFD86980FAA1" box="[749,947,1279,1338]" gridcol="3" gridrow="10" pageId="5" pageNumber="284">Weak present distally on tarsus</td>
<td id="7681DA25FF9D00530886CFD86E5FFAA1" box="[976,1132,1279,1338]" gridcol="4" gridrow="10" pageId="5" pageNumber="284">Weakly developed</td>
<td id="7681DA25FF9D00530FD0CFD86F03FAA1" box="[1158,1328,1279,1338]" gridcol="5" gridrow="10" pageId="5" pageNumber="284">Weak</td>
<td id="7681DA25FF9D00530E1CCFD86C22FAA1" box="[1354,1553,1279,1338]" gridcol="6" gridrow="10" pageId="5" pageNumber="284">Wide, low and extends about three-fourths of the length of the tarsum</td>
<td id="7681DA25FF9D00530D7DCFD86CC4FAA1" box="[1579,1783,1279,1338]" gridcol="7" gridrow="10" pageId="5" pageNumber="284">Row of tubercles forming a dermal ridge</td>
</tr>
<tr id="3550B359FF9D00530B2BCE726CC4F9E0" box="[125,1783,1365,1659]" gridrow="11" pageId="5" pageNumber="284">
<th id="7681DA25FF9D00530B2BCE726B00F9E0" box="[125,307,1365,1659]" gridcol="0" gridrow="11" pageId="5" pageNumber="284">
<emphasis id="B9146D09FF9DFFA90B2BCE726AF7FA7E" bold="true" pageId="5" pageNumber="284">Forearm Distribution and elevation</emphasis>
</th>
<td id="7681DA25FF9D00530A69CE726BC9F9E0" box="[319,506,1365,1659]" gridcol="1" gridrow="11" pageId="5" pageNumber="284">
Dermal fold, but no row of tubercles Northern slopes of the Chiapan highlands in Mexico and the Sierra de los Cuchumatanes in Guatemala. Also in the upper Grijalva Basin in Chiapas, Mexico and Guatemala at 700
<emphasis id="B9146D09FF9DFFA90A91CD736BE3F9FE" box="[455,464,1620,1637]" italics="true" pageId="5" pageNumber="284"></emphasis>
1700 m elev.
</td>
<td id="7681DA25FF9D00530942CE7268FCF9E0" box="[532,719,1365,1659]" gridcol="2" gridrow="11" pageId="5" pageNumber="284">
A row of elevated tubercles Northern slopes of the Sierra de las Minas and Montañas del Mico in eastern Guatemala at 100
<emphasis id="B9146D09FF9DFFA90906CD33686AF9BE" box="[592,601,1556,1573]" italics="true" pageId="5" pageNumber="284"></emphasis>
895 m elev.
</td>
<td id="7681DA25FF9D005309BBCE726980F9E0" box="[749,947,1365,1659]" gridcol="3" gridrow="11" pageId="5" pageNumber="284">
A row of low tubercles fused into a nearly continuous fold Pacific versant from northwestern El Salvador to south central Honduras and southeastern Guatemala. Also along the Atlantic versant near the Continental divide in southwestern Honduras at 1440
<emphasis id="B9146D09FF9DFFA90878CD4D6904F9E0" box="[814,823,1642,1659]" italics="true" pageId="5" pageNumber="284"></emphasis>
2050 m elev.
</td>
<td id="7681DA25FF9D00530886CE726E5FF9E0" box="[976,1132,1365,1659]" gridcol="4" gridrow="11" pageId="5" pageNumber="284">
Low of elevated tubercles forming a distinct ridge Sierra Santa Cruz in eastern Guatemala at 366
<emphasis id="B9146D09FF9DFFA908A1CECD6E33FA60" box="[1015,1024,1514,1531]" italics="true" pageId="5" pageNumber="284"></emphasis>
1150 m elev.
</td>
<td id="7681DA25FF9D00530FD0CE726F03F9E0" box="[1158,1328,1365,1659]" gridcol="5" gridrow="11" pageId="5" pageNumber="284">
Row of elevated tubercles along the ventrolateral edge of the forearm forms a distinct ridges Cordillera Nombre de Dios in north-central Honduras at 160
<emphasis id="B9146D09FF9DFFA90E57CECD6F39FA60" box="[1281,1290,1514,1531]" italics="true" pageId="5" pageNumber="284"></emphasis>
1580 m elev.
</td>
<td id="7681DA25FF9D00530E1CCE726C22F9E0" box="[1354,1553,1365,1659]" gridcol="6" gridrow="11" pageId="5" pageNumber="284">
A distinct dermal ridge punctuated with enlarged tubercles on the ventrolateral edge Atlantic slopes of northern Oaxaca and southern Puebla at 400
<emphasis id="B9146D09FF9DFFA90E96CECD6FFAFA60" box="[1472,1481,1514,1531]" italics="true" pageId="5" pageNumber="284"></emphasis>
1500 m elev.
</td>
<td id="7681DA25FF9D00530D7DCE726CC4F9E0" box="[1579,1783,1365,1659]" gridcol="7" gridrow="11" pageId="5" pageNumber="284">
Ulnar tubercles unequally elevated Southern Mexico in Oaxaca and Veracruz at 76
<emphasis id="B9146D09FF9DFFA90D3DCECD6C47FA60" box="[1643,1652,1514,1531]" italics="true" pageId="5" pageNumber="284"></emphasis>
600 m elev.
</td>
</tr>
</table>
</paragraph>
<paragraph id="8BDFB11BFF9EFFAA0B91CA856967FD3E" blockId="6.[151,1437,151,1070]" pageId="6" pageNumber="285">
<emphasis id="B9146D09FF9EFFAA0B91CA856B0FFE20" bold="true" box="[199,316,418,443]" pageId="6" pageNumber="285">Variation.</emphasis>
Marked sexual size dimorphism is evident, with a ratio of sexual dimorphism of 1.52, and females attaining the greater SVL (
<tableCitation id="C6E284A0FF9EFFAA0A8ECAE36802FE47" box="[472,561,452,477]" captionStart="TABLE 1" captionStartId="2.[151,239,1179,1202]" captionText="TABLE 1. Morphometric variation in type-series of Ptychohyla zoque: range (in millimeters) followed by mean in parentheses." pageId="6" pageNumber="285">Table 1</tableCitation>
). Variation in body measurements among four males and five females is summarized in
<tableCitation id="C6E284A0FF9EFFAA0A13CAC26BA9FE65" box="[325,410,485,510]" captionStart="TABLE 1" captionStartId="2.[151,239,1179,1202]" captionText="TABLE 1. Morphometric variation in type-series of Ptychohyla zoque: range (in millimeters) followed by mean in parentheses." pageId="6" pageNumber="285">Table 1</tableCitation>
. Two males (CARIE 1138, 1216) and three females (MZFC, 18667, 24005; CARIE 1215) lack the scattered white spots on dorsum; additionally, one male (CARIE 1135) and two females (MZFC 24006; CARIE 1215) have a concentration of black spots along the edge of the lower lip, and UTA
<accessionNumber id="94332CF8FF9EFFAA0FA3C90F6F68FDDB" box="[1269,1371,551,576]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A59271" pageId="6" pageNumber="285" type="EnaNcbi">A 59271</accessionNumber>
have concentrated brown spots on the chin and chest that are absent in the other specimens. Variation in vomerine teeth is as follows: 5/7 (MZFC 18667); 5/4 (MZFC 24005); 6/5 (MZFC 24006); 6/6 (UTA-A 59271); 4/5 (CARIE 1135); 5/7 (CARIE 1138); 4/5 (CARIE 1215) and 4/5 (CARIE 1216).
</paragraph>
<paragraph id="8BDFB11BFF9EFFAA0B91C98A6FAEFBB5" blockId="6.[151,1437,151,1070]" pageId="6" pageNumber="285">
<emphasis id="B9146D09FF9EFFAA0B91C98A6B87FD5D" bold="true" box="[199,436,685,710]" pageId="6" pageNumber="285">Advertisement call.</emphasis>
The description of the advertisement call is based on one male (SVL =
<quantity id="4C981CFEFF9EFFAA0E7FC98A6FA7FD5D" box="[1321,1428,685,710]" metricMagnitude="-2" metricUnit="m" metricValue="3.61" pageId="6" pageNumber="285" unit="mm" value="36.1">36.1 mm</quantity>
) recorded at 0 140 h on
<date id="FFDE97DBFF9EFFAA0AC7C9F56828FD71" box="[401,539,722,746]" pageId="6" pageNumber="285" value="2014-05-04">4 May 2014</date>
, when it was calling on a leaf at
<quantity id="4C981CFEFF9EFFAA08D1C9F569F8FD72" box="[903,971,722,746]" metricMagnitude="-1" metricUnit="m" metricValue="9.2" pageId="6" pageNumber="285" unit="cm" value="92.0">92 cm</quantity>
above ground, over a quiet pool next to a stream, in the locality of Arroyo Zarco, Uxpanapa,
<collectingRegion id="49A47FF9FF9EFFAA09B6C9D16977FC96" box="[736,836,758,781]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="6" pageNumber="285">Veracruz</collectingRegion>
(
<geoCoordinate id="EE54D7DCFF9EFFAA0803C9D169D8FC96" box="[853,1003,757,782]" direction="north" orientation="latitude" pageId="6" pageNumber="285" precision="9" value="17.191">17° 11.46 N</geoCoordinate>
,
<geoCoordinate id="EE54D7DCFF9EFFAA08A1C9D16EA7FC96" box="[1015,1172,758,782]" direction="west" orientation="longitude" pageId="6" pageNumber="285" precision="9" value="-94.488335">94° 29.30 W</geoCoordinate>
;
<quantity id="4C981CFEFF9EFFAA0FF7C9D26EDAFC96" box="[1185,1257,757,782]" metricMagnitude="2" metricUnit="m" metricValue="2.16" pageId="6" pageNumber="285" unit="m" value="216.0">216 m</quantity>
elevation). The advertisement call of
<taxonomicName id="4C60CA98FF9EFFAA0ADFC83D6BD5FCAA" authorityName="Caviedes-Solís" authorityYear="2017" box="[393,486,794,817]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="285" phylum="Chordata" rank="species" species="zoque">
<emphasis id="B9146D09FF9EFFAA0ADFC83D6BD5FCAA" box="[393,486,794,817]" italics="true" pageId="6" pageNumber="285">P. zoque</emphasis>
</taxonomicName>
is very soft and difficult to hear because of noise produced by other species of frogs (mainly
<taxonomicName id="4C60CA98FF9EFFAA0BA0C81A6BECFCCE" baseAuthorityName="Duellman and Hoyt" baseAuthorityYear="1961" box="[246,479,829,853]" class="Amphibia" family="Hylidae" genus="Exerodonta" kingdom="Animalia" order="Anura" pageId="6" pageNumber="285" phylum="Chordata" rank="species" species="bivocata">
<emphasis id="B9146D09FF9EFFAA0BA0C81A6BECFCCE" box="[246,479,829,853]" italics="true" pageId="6" pageNumber="285">Exerodonta bivocata</emphasis>
</taxonomicName>
), insects, and the stream. It consists of two notes with different structure (
<figureCitation id="135BAD9EFF9EFFAA0E7FC81A6F5DFCCD" box="[1321,1390,829,854]" captionStart="FIGURE 3" captionStartId="6.[151,250,1668,1690]" captionTargetBox="[163,1420,1125,1647]" captionTargetId="figure@6.[163,1420,1125,1647]" captionTargetPageId="6" captionText="FIGURE 3. Advertisement call of Ptychohyla zoque, recorded in the locality of Arroyo Zarco, municipality of Uxpanapa, Veracruz. Oscillogram (A), and audio spectrogram (B) of two calls composed by two notes with different structure." httpUri="https://zenodo.org/record/884306/files/figure.png" pageId="6" pageNumber="285">Fig. 3</figureCitation>
); in some cases the first note is immediately followed by the second, but in some instances the notes seem to be emitted independently. The first is a short, pulsed note (
<emphasis id="B9146D09FF9EFFAA09FCC8A1688BFC06" box="[682,696,902,925]" italics="true" pageId="6" pageNumber="285">n</emphasis>
= 9), with a mean number of pulses of 9.6 ± 4 (range 314), mean pulse duration 0.008 ±
<geoCoordinate id="EE54D7DCFF9EFFAA0AC8C88D6BC2FC5A" box="[414,497,938,962]" direction="south" orientation="latitude" pageId="6" pageNumber="285" precision="55" value="-0.003">0.003 s</geoCoordinate>
(range 0.0050.011), mean interpulse interval 0.015 ±.
<geoCoordinate id="EE54D7DCFF9EFFAA0F3EC88D6E9BFC5A" box="[1128,1192,938,962]" direction="south" orientation="latitude" pageId="6" pageNumber="285" precision="15" value="-0.0019444445">0 0 7 s</geoCoordinate>
(range 0.0050.027), mean dominant frequency 2516.9 ± 103.5 Hz (range 2325.62670.1), and mean note duration 0.249 ±
<geoCoordinate id="EE54D7DCFF9EFFAA0E1EC8EA6FAFFC7E" box="[1352,1436,973,998]" direction="south" orientation="latitude" pageId="6" pageNumber="285" precision="55" value="-0.136">0.136 s</geoCoordinate>
(range 0.0430.395). The second note is shorter than the first, and is not pulsed (n= 8), it has a mean dominant frequency 2024.1 ± 220.8 Hz (range 1808.82497.8), and a mean note duration 0.01 ±
<geoCoordinate id="EE54D7DCFF9EFFAA0F0DCF316E9FFBB6" box="[1115,1196,1046,1070]" direction="south" orientation="latitude" pageId="6" pageNumber="285" precision="55" value="-0.003">0.003 s</geoCoordinate>
(range 0.0050.013).
</paragraph>
</subSubSection>
<caption id="DF1FE193FF9EFFAA0BC1CDA36F14F922" httpUri="https://zenodo.org/record/884306/files/figure.png" pageId="6" pageNumber="285" targetBox="[163,1420,1125,1647]" targetPageId="6">
<paragraph id="8BDFB11BFF9EFFAA0BC1CDA36F14F922" blockId="6.[151,1436,1668,1721]" pageId="6" pageNumber="285">
<emphasis id="B9146D09FF9EFFAA0BC1CDA36B2EF902" bold="true" box="[151,285,1668,1690]" pageId="6" pageNumber="285">FIGURE 3.</emphasis>
Advertisement call of
<taxonomicName id="4C60CA98FF9EFFAA0959CDA368F6F901" authorityName="Caviedes-Solís" authorityYear="2017" box="[527,709,1668,1690]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="285" phylum="Chordata" rank="species" species="zoque">
<emphasis id="B9146D09FF9EFFAA0959CDA368F6F901" box="[527,709,1668,1690]" italics="true" pageId="6" pageNumber="285">Ptychohyla zoque</emphasis>
</taxonomicName>
, recorded in the locality of Arroyo Zarco, municipality of Uxpanapa, Veracruz. Oscillogram (A), and audio spectrogram (B) of two calls composed by two notes with different structure.
</paragraph>
</caption>
<subSubSection id="C37AE290FF9EFFAB0B91CDC76B07F9E7" lastPageId="7" lastPageNumber="286" pageId="6" pageNumber="285" type="distribution">
<paragraph id="8BDFB11BFF9EFFAB0B91CDC769CDFEFE" blockId="6.[151,1436,1760,2037]" lastBlockId="7.[151,1436,151,357]" lastPageId="7" lastPageNumber="286" pageId="6" pageNumber="285">
<emphasis id="B9146D09FF9EFFAA0B91CDC76BC9F962" bold="true" box="[199,506,1760,1785]" pageId="6" pageNumber="285">Distribution and ecology.</emphasis>
<taxonomicName id="4C60CA98FF9EFFAA0953CDC768FCF963" authorityName="Caviedes-Solís" authorityYear="2017" box="[517,719,1760,1784]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="6" pageNumber="285" phylum="Chordata" rank="species" species="zoque">
<emphasis id="B9146D09FF9EFFAA0953CDC768FCF963" box="[517,719,1760,1784]" italics="true" pageId="6" pageNumber="285">Ptychohyla zoque</emphasis>
</taxonomicName>
occurs in evergreen tropical forest, and is known from three localities in southern
<collectingRegion id="49A47FF9FF9EFFAA0AD0CC236BEFF886" box="[390,476,1796,1821]" country="Mexico" name="Mexico" pageId="6" pageNumber="285">
<collectingCountry id="F377F18BFF9EFFAA0AD0CC236BEFF886" box="[390,476,1796,1821]" name="Mexico" pageId="6" pageNumber="285">Mexico</collectingCountry>
</collectingRegion>
in the Selva Zoque, two in southern
<collectingRegion id="49A47FF9FF9EFFAA082ECC2269E8F887" box="[888,987,1797,1820]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="6" pageNumber="285">Veracruz</collectingRegion>
(Paso del Moral and Arroyo Zarco) and one in northeastern
<collectingRegion id="49A47FF9FF9EFFAA0A2ACC0F6BE3F8DB" box="[380,464,1832,1856]" country="Mexico" name="Oaxaca" pageId="6" pageNumber="285">Oaxaca</collectingRegion>
(Chalchijapa) (
<figureCitation id="135BAD9EFF9EFFAA09D5CC0F68F8F8DB" box="[643,715,1832,1857]" captionStart="FIGURE 4" captionStartId="7.[151,250,1214,1236]" captionTargetBox="[151,1436,409,1193]" captionTargetId="figure@7.[151,1436,409,1193]" captionTargetPageId="7" captionText="FIGURE 4. Geographic distribution of the species of Ptychohyla in Mexico." httpUri="https://zenodo.org/record/884308/files/figure.png" pageId="6" pageNumber="285">Fig. 4</figureCitation>
). Known elevational range is
<quantity id="4C981CFEFF9EFFAA0F70CC0F6EAAF8DB" box="[1062,1177,1832,1857]" metricMagnitude="2" metricUnit="m" metricValue="3.38" metricValueMax="6.0" metricValueMin="0.76" pageId="6" pageNumber="285" unit="m" value="338.0" valueMax="600.0" valueMin="76.0">76600 m</quantity>
asl. Seven of the nine specimens that comprise the
<typeStatus id="54DB0FB9FF9EFFAA0ABCCC6A6829F8FE" box="[490,538,1869,1893]" pageId="6" pageNumber="285">type</typeStatus>
series plus 20 additional specimens not collected (
<specimenCount id="9D667A92FF9EFFAA0F25CC6B6EE1F8FE" box="[1139,1234,1868,1893]" pageId="6" pageNumber="285" type="adult">18 adult</specimenCount>
males, one adult female, and one juvenile) were observed at night, on branches or leaves at
<quantity id="4C981CFEFF9EFFAA08B0CC576E57F813" box="[998,1124,1904,1928]" metricMagnitude="0" metricUnit="m" metricValue="1.24" metricValueMax="2.28" metricValueMin="0.2" pageId="6" pageNumber="285" unit="cm" value="124.0" valueMax="228.0" valueMin="20.0">20228 cm</quantity>
above ground, and close to streams. Exceptions from this pattern were the female from Chalchijapa (MZFC 18667), which was sleeping on a leaf at 0 830 h near a stream, a male from Paso del Moral (CARIE 1135), which was found on the ground at 1504 h near a stream, and a juvenile of undetermined sex (not collected), which was on a leaf at 1730 h, approximately
<quantity id="4C981CFEFF9EFFAB0ED6CCFA6A9EFF34" lastPageId="7" lastPageNumber="286" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="285" unit="m" value="30.0">30 m</quantity>
away from the nearest stream. Twenty-five of the 29 frogs were found in the dry season (March-May), whereas only four were observed in the beginning of the rainy season (June-July). All males had well developed nuptial excrescences regardless of month collected. An amplexing pair (CARIE
<date id="FFDE97DBFF9FFFAB0887CBF86E48FF63" box="[977,1147,223,248]" pageId="7" pageNumber="286" value="1215" valueMax="1216">1215 and 1216</date>
) was collected in Arroyo Zarco, and was held overnight in a plastic bag, where
<specimenCount id="9D667A92FF9FFFAB0844CA23694AFE86" box="[786,889,260,285]" pageId="7" pageNumber="286" type="egg">341 eggs</specimenCount>
were deposited a few hours later. Although all males appeared to be in breeding condition, only two observed on
<date id="FFDE97DBFF9FFFAB082ECA0F6EADFEDB" box="[888,1182,295,320]" pageId="7" pageNumber="286" value="2014-03-11" valueMax="2014-05-04" valueMin="2014-03-11">
11 March and 4 May
<quantity id="4C981CFEFF9FFFAB0F30CA0F6EADFEDB" box="[1126,1182,296,320]" metricMagnitude="1" metricUnit="m" metricValue="5.11556" pageId="7" pageNumber="286" unit="in" value="2014.0">2014</quantity>
</date>
in Arroyo Zarco, were calling from perches on leaves 228 and
<quantity id="4C981CFEFF9FFFAB0902CA6B68AAFEFF" box="[596,665,332,356]" metricMagnitude="-1" metricUnit="m" metricValue="9.2" pageId="7" pageNumber="286" unit="cm" value="92.0">92 cm</quantity>
above the ground, respectively.
</paragraph>
<caption id="DF1FE193FF9FFFAB0BC1CF99699AFB4F" box="[151,937,1214,1236]" httpUri="https://zenodo.org/record/884308/files/figure.png" pageId="7" pageNumber="286" targetBox="[151,1436,409,1193]" targetPageId="7">
<paragraph id="8BDFB11BFF9FFFAB0BC1CF99699AFB4F" blockId="7.[151,937,1214,1236]" box="[151,937,1214,1236]" pageId="7" pageNumber="286">
<emphasis id="B9146D09FF9FFFAB0BC1CF996B2BFB4F" bold="true" box="[151,280,1214,1236]" pageId="7" pageNumber="286">FIGURE 4.</emphasis>
Geographic distribution of the species of
<taxonomicName id="4C60CA98FF9FFFAB0994CF996907FB4F" authorityName="Caviedes-Solís" authorityYear="2017" box="[706,820,1214,1236]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="7" pageNumber="286" phylum="Chordata" rank="genus">
<emphasis id="B9146D09FF9FFFAB0994CF996907FB4F" box="[706,820,1214,1236]" italics="true" pageId="7" pageNumber="286">Ptychohyla</emphasis>
</taxonomicName>
in Mexico.
</paragraph>
</caption>
<paragraph id="8BDFB11BFF9FFFAB0B91CFDB6B07F9E7" blockId="7.[151,1436,1276,1805]" pageId="7" pageNumber="286">
<taxonomicName id="4C60CA98FF9FFFAB0B91CFDB6BA2FA8F" authorityName="Caviedes-Solís" authorityYear="2017" box="[199,401,1276,1300]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="7" pageNumber="286" phylum="Chordata" rank="species" species="zoque">
<emphasis id="B9146D09FF9FFFAB0B91CFDB6BA2FA8F" box="[199,401,1276,1300]" italics="true" pageId="7" pageNumber="286">Ptychohyla zoque</emphasis>
</taxonomicName>
is sympatric with other species of amphibians including
<taxonomicName id="4C60CA98FF9FFFAB0F62CFDB6F13FA8F" baseAuthorityName="Duméril" baseAuthorityYear="1853" box="[1076,1312,1276,1300]" class="Amphibia" family="Hylidae" genus="Agalychnis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="286" phylum="Chordata" rank="species" species="moreletii">
<emphasis id="B9146D09FF9FFFAB0F62CFDB6F13FA8F" box="[1076,1312,1276,1300]" italics="true" pageId="7" pageNumber="286">Agalychnis moreletii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C60CA98FF9FFFAB0E64CFDB6ADFFAAC" baseAuthorityName="Steindachner" baseAuthorityYear="1864" class="Amphibia" family="Hylidae" genus="Anotheca" kingdom="Animalia" order="Anura" pageId="7" pageNumber="286" phylum="Chordata" rank="species" species="spinosa">
<emphasis id="B9146D09FF9FFFAB0E64CFDB6ADFFAAC" italics="true" pageId="7" pageNumber="286">Anotheca spinosa</emphasis>
</taxonomicName>
,
<emphasis id="B9146D09FF9FFFAB0A5CCE076ECEFAC7" italics="true" pageId="7" pageNumber="286">
<taxonomicName id="4C60CA98FF9FFFAB0A5CCE07682CFAAC" baseAuthorityName="Firschein and Smith" baseAuthorityYear="1957" box="[266,543,1312,1336]" class="Amphibia" family="Bufonidae" genus="Incilius" kingdom="Animalia" order="Anura" pageId="7" pageNumber="286" phylum="Chordata" rank="species" species="macrocristatus">Incilius macrocristatus</taxonomicName>
,
<taxonomicName id="4C60CA98FF9FFFAB096CCE06698DFAA3" authority=", Hyalinobatrachium" authorityName="Hyalinobatrachium" box="[570,958,1312,1336]" class="Amphibia" family="Bufonidae" genus="Incilius" kingdom="Animalia" order="Anura" pageId="7" pageNumber="286" phylum="Chordata" rank="species" species="valliceps">I. valliceps, Hyalinobatrachium</taxonomicName>
fleischmanni,
<taxonomicName id="4C60CA98FF9FFFAB0FD5CE076F5BFAA3" baseAuthorityName="Boulenger" baseAuthorityYear="1898" box="[1155,1384,1312,1336]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="7" pageNumber="286" phylum="Chordata" rank="species" species="alfredi">Craugastor alfredi</taxonomicName>
,
<taxonomicName id="4C60CA98FF9FFFAB0ED5CE076B04FAC7" baseAuthorityName="Peters" baseAuthorityYear="1870" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="7" pageNumber="286" phylum="Chordata" rank="species" species="berkenbuschii">C. berkenbuschii</taxonomicName>
,
<taxonomicName id="4C60CA98FF9FFFAB0A10CE636847FAC7" authority=", Eleutherodactylus" authorityName="Eleutherodactylus" box="[326,628,1348,1372]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="7" pageNumber="286" phylum="Chordata" rank="species" species="loki">C. loki, Eleutherodactylus</taxonomicName>
leprus,
<taxonomicName id="4C60CA98FF9FFFAB098ECE636E33FAC7" baseAuthorityName="Duellman" baseAuthorityYear="1961" box="[728,1024,1348,1372]" class="Amphibia" family="Hylidae" genus="Duellmanohyla" kingdom="Animalia" order="Anura" pageId="7" pageNumber="286" phylum="Chordata" rank="species" species="chamulae">Duellmanohyla chamulae</taxonomicName>
,
<taxonomicName id="4C60CA98FF9FFFAB0F59CE636ECEFAC7" baseAuthorityName="Duellman and Hoyt" baseAuthorityYear="1961" box="[1039,1277,1348,1372]" class="Amphibia" family="Hylidae" genus="Exerodonta" kingdom="Animalia" order="Anura" pageId="7" pageNumber="286" phylum="Chordata" rank="species" species="bivocata">Exerodonta bivocata</taxonomicName>
</emphasis>
,
<taxonomicName id="4C60CA98FF9FFFAB0E46CE636AE2FA1B" baseAuthorityName="Cope" baseAuthorityYear="1866" class="Amphibia" family="Microhylidae" genus="Hypopachus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="286" phylum="Chordata" rank="species" species="ustus">
<emphasis id="B9146D09FF9FFFAB0E46CE636AE2FA1B" italics="true" pageId="7" pageNumber="286">Hypopachus ustus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C60CA98FF9FFFAB0BBECE4F6BF2FA1B" box="[232,449,1384,1408]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="7" pageNumber="286" phylum="Chordata" rank="species" species="horribilis">
<emphasis id="B9146D09FF9FFFAB0BBECE4F6BF2FA1B" box="[232,449,1384,1408]" italics="true" pageId="7" pageNumber="286">Rhinella horribilis</emphasis>
</taxonomicName>
,
<emphasis id="B9146D09FF9FFFAB0A8ECE4F6BE8FA3F" italics="true" pageId="7" pageNumber="286">
<taxonomicName id="4C60CA98FF9FFFAB0A8ECE4F6894FA1B" baseAuthorityName="Duméril and Bibron" baseAuthorityYear="1841" box="[472,679,1384,1408]" class="Amphibia" family="Hylidae" genus="Smilisca" kingdom="Animalia" order="Anura" pageId="7" pageNumber="286" phylum="Chordata" rank="species" species="baudinii">Smilisca baudinii</taxonomicName>
,
<taxonomicName id="4C60CA98FF9FFFAB09EACE4F69DDFA1B" authority=", Lithobates" authorityName="Lithobates" box="[700,1006,1384,1408]" class="Amphibia" family="Hylidae" genus="Smilisca" kingdom="Animalia" order="Anura" pageId="7" pageNumber="286" phylum="Chordata" rank="species" species="cyanosticta">S. cyanosticta, Lithobates</taxonomicName>
vaillanti, Bolitoglossa alberchi,
<taxonomicName id="4C60CA98FF9FFFAB0ED2CE4E6B32FA3F" pageId="7" pageNumber="286" rank="species" species="rufescens">B. rufescens</taxonomicName>
and
<taxonomicName id="4C60CA98FF9FFFAB0A6ACEAA6BE4FA3F" box="[316,471,1421,1444]" pageId="7" pageNumber="286" rank="species" species="veracrucis">B. veracrucis</taxonomicName>
.
</emphasis>
The new species is relatively common in the evergreen tropical forests, being more abundant than other members of the family
<taxonomicName id="4C60CA98FF9FFFAB09C6CE8868DAFA53" authorityName="Rafinesque" authorityYear="1815" box="[656,745,1455,1480]" class="Amphibia" family="Hylidae" kingdom="Animalia" order="Anura" pageId="7" pageNumber="286" phylum="Chordata" rank="family">Hylidae</taxonomicName>
such as
<taxonomicName id="4C60CA98FF9FFFAB0818CE976E42FA53" baseAuthorityName="Duellman" baseAuthorityYear="1961" box="[846,1137,1456,1480]" class="Amphibia" family="Hylidae" genus="Duellmanohyla" kingdom="Animalia" order="Anura" pageId="7" pageNumber="286" phylum="Chordata" rank="species" species="chamulae">
<emphasis id="B9146D09FF9FFFAB0818CE976E42FA53" box="[846,1137,1456,1480]" italics="true" pageId="7" pageNumber="286">Duellmanohyla chamulae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C60CA98FF9FFFAB0FD7CE976F59FA53" baseAuthorityName="Duméril" baseAuthorityYear="1853" box="[1153,1386,1456,1480]" class="Amphibia" family="Hylidae" genus="Agalychnis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="286" phylum="Chordata" rank="species" species="moreletii">
<emphasis id="B9146D09FF9FFFAB0FD7CE976F59FA53" box="[1153,1386,1456,1480]" italics="true" pageId="7" pageNumber="286">Agalychnis moreletii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C60CA98FF9FFFAB0BC1CEF36B50FA77" baseAuthorityName="Steindachner" baseAuthorityYear="1864" box="[151,355,1492,1516]" class="Amphibia" family="Hylidae" genus="Anotheca" kingdom="Animalia" order="Anura" pageId="7" pageNumber="286" phylum="Chordata" rank="species" species="spinosa">
<emphasis id="B9146D09FF9FFFAB0BC1CEF36B50FA77" box="[151,355,1492,1516]" italics="true" pageId="7" pageNumber="286">Anotheca spinosa</emphasis>
</taxonomicName>
, but less abundant than
<taxonomicName id="4C60CA98FF9FFFAB09D8CEF3694FFA77" baseAuthorityName="Duellman and Hoyt" baseAuthorityYear="1961" box="[654,892,1492,1516]" class="Amphibia" family="Hylidae" genus="Exerodonta" kingdom="Animalia" order="Anura" pageId="7" pageNumber="286" phylum="Chordata" rank="species" species="bivocata">
<emphasis id="B9146D09FF9FFFAB09D8CEF3694FFA77" box="[654,892,1492,1516]" italics="true" pageId="7" pageNumber="286">Exerodonta bivocata</emphasis>
</taxonomicName>
and anurans in other families represented by
<taxonomicName id="4C60CA98FF9FFFAB0BC1CEDF6B8FF98B" baseAuthorityName="Peters" baseAuthorityYear="1870" box="[151,444,1528,1552]" class="Amphibia" family="Craugastoridae" genus="Craugastor" kingdom="Animalia" order="Anura" pageId="7" pageNumber="286" phylum="Chordata" rank="species" species="berkenbuschii">
<emphasis id="B9146D09FF9FFFAB0BC1CEDF6B8FF98B" box="[151,444,1528,1552]" italics="true" pageId="7" pageNumber="286">Craugastor berkenbuschii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C60CA98FF9FFFAB0AA5CEDF68C7F994" baseAuthorityName="Firschein and Smith" baseAuthorityYear="1957" box="[499,756,1528,1552]" class="Amphibia" family="Bufonidae" genus="Incilius" kingdom="Animalia" order="Anura" pageId="7" pageNumber="286" phylum="Chordata" rank="species" species="macrocristatus">
<emphasis id="B9146D09FF9FFFAB0AA5CEDF68C7F994" box="[499,756,1528,1552]" italics="true" pageId="7" pageNumber="286">Incilius macrocristatus</emphasis>
</taxonomicName>
. This species appears to be sensitive to anthropogenic forest degradation, since it has been observed only in undisturbed evergreen tropical forest sites (
<figureCitation id="135BAD9EFF9FFFAB0FD9CD3B6EE1F9AE" box="[1167,1234,1564,1589]" captionStart="FIGURE 5" captionStartId="8.[151,250,1272,1294]" captionTargetBox="[161,1413,196,1245]" captionTargetId="figure@8.[161,1413,196,1245]" captionTargetPageId="8" captionText="FIGURE 5. General habitat at Arroyo Zarco (A); Section of a stream at the same locality (B); modified habitat near the type locality (C, D)." httpUri="https://zenodo.org/record/884310/files/figure.png" pageId="7" pageNumber="286">Fig. 5</figureCitation>
A and B), but not in altered environments (
<figureCitation id="135BAD9EFF9FFFAB0AF9CD186BD8F9C3" box="[431,491,1599,1624]" captionStart="FIGURE 5" captionStartId="8.[151,250,1272,1294]" captionTargetBox="[161,1413,196,1245]" captionTargetId="figure@8.[161,1413,196,1245]" captionTargetPageId="8" captionText="FIGURE 5. General habitat at Arroyo Zarco (A); Section of a stream at the same locality (B); modified habitat near the type locality (C, D)." httpUri="https://zenodo.org/record/884310/files/figure.png" pageId="7" pageNumber="286">Fig 5</figureCitation>
C and D) such as cattle pastures, rubber plantations and secondary forests (Aguilar-
<bibRefCitation id="EFF1CCEAFF9FFFAB0BC1CD436B1BF9E7" author="Lopez" box="[151,296,1636,1660]" pageId="7" pageNumber="286" refString="Aguilar-Lopez, J. L. (2010) Transformacion del habitat y diversidad de anfibios en un paisaje tropical del sur de Mexico. Master of Science Thesis. Instituto de Ecologia A. C. Xalapa, Mexico, 60 pp." type="book" year="2010">López, 2010</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37AE290FF9FFFAB0B91CDA06828F896" pageId="7" pageNumber="286" type="etymology">
<paragraph id="8BDFB11BFF9FFFAB0B91CDA06828F896" blockId="7.[151,1436,1276,1805]" pageId="7" pageNumber="286">
<emphasis id="B9146D09FF9FFFAB0B91CDA06B7DF904" bold="true" box="[199,334,1671,1696]" pageId="7" pageNumber="286">Etymology.</emphasis>
The species epithet,
<taxonomicName id="4C60CA98FF9FFFAB0919CDAE68A3F93B" authorityName="Caviedes-Solís" authorityYear="2017" box="[591,656,1673,1696]" class="Amphibia" family="Hylidae" genus="Ptychohyla" kingdom="Animalia" order="Anura" pageId="7" pageNumber="286" phylum="Chordata" rank="species" species="zoque">
<emphasis id="B9146D09FF9FFFAB0919CDAE68A3F93B" box="[591,656,1673,1696]" italics="true" pageId="7" pageNumber="286">zoque</emphasis>
</taxonomicName>
, is an indeclinable word and refers to the Selva Zoque, a region comprised of tropical forest and covering the areas about Uxpanapa (
<collectingRegion id="49A47FF9FF9FFFAB08C1CD8B6E32F95F" box="[919,1025,1708,1732]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="7" pageNumber="286">Veracruz</collectingRegion>
), Chimalapas (
<collectingRegion id="49A47FF9FF9FFFAB0FF3CD8B6ECFF95F" box="[1189,1276,1708,1732]" country="Mexico" name="Oaxaca" pageId="7" pageNumber="286">Oaxaca</collectingRegion>
) and El Ocote (
<collectingRegion id="49A47FF9FF9FFFAB0BC9CDE86B33F973" box="[159,256,1743,1768]" country="Mexico" name="Chiapas" pageId="7" pageNumber="286">Chiapas</collectingRegion>
). This region includes part of the historical range of Zoque culture, an ethnic group that has inhabited this region since pre-Columbian times.
</paragraph>
</subSubSection>
</treatment>
</document>