364 lines
44 KiB
XML
364 lines
44 KiB
XML
<document id="30C979E3598487640E2A0629EA20B5F8" ID-DOI="10.5281/zenodo.173478" ID-GBIF-Dataset="1e858e0a-a143-49df-bd29-aa89c25c41e8" ID-ISSN="11755326" ID-Zenodo-Dep="173478" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459775041540" checkinUser="plazi" docAuthor="Duellman, William E., Lehr, Edgar & Venegas, Pablo J." docDate="2006" docId="03A50A52FFFBFF82FE80E748FC50F90D" docLanguage="en" docName="zt01285p064.pdf" docOrigin="Zootaxa 1285" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Eleutherodactylus corrugatus Duellman, Lehr & Venegas, 2006, new species" docType="treatment" docVersion="7" lastPageNumber="57" masterDocId="FF9C722AFFFAFF84FF88E04EFFB0FFDB" masterDocTitle="Two new species of Eleutherodactylus (Anura: Leptodactylidae) from the Andes of northern Peru" masterLastPageNumber="64" masterPageNumber="51" pageNumber="52" updateTime="1698217738710" updateUser="plazi">
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<mods:mods id="BD2D886EA72D0D496B289E7854440CC3" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4EA8FCBBC2BAD07AA129617A8C0343B3">
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<mods:title id="D5E9C476D5064AF354C19ECBBEBAFA5F">Two new species of Eleutherodactylus (Anura: Leptodactylidae) from the Andes of northern Peru</mods:title>
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<mods:name id="CCD75D869782EAF3291D222456CBEC45" type="personal">
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<mods:role id="8896BFE164D4243FE3910EAF9A200810">
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<mods:roleTerm id="5F37DE16E0B3FC2DAE5AF6AE0B753AB3">Author</mods:roleTerm>
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<mods:namePart id="4B6620C999DC6A1A308A46DBF5E90015">Duellman, William E.</mods:namePart>
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<mods:roleTerm id="E4D6698E2E0F15AEB5AADFFAD2724B24">Author</mods:roleTerm>
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<mods:namePart id="852EC93F2B3A74FFB848B90775F5879E">Lehr, Edgar</mods:namePart>
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</mods:name>
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<mods:name id="9C368C9FF84909A5357731FD56E5EF1A" type="personal">
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<mods:roleTerm id="A60F0DF0A39B0C355ABD49B6004F16CD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="33CCE0D27581422A1F3388FBF1EB4FC5">Venegas, Pablo J.</mods:namePart>
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<mods:typeOfResource id="2EAF604818EA671FBB21EDE3AE61491D">text</mods:typeOfResource>
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<mods:titleInfo id="F61CE97873958F41AD0D8A01E89ACA10">
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<mods:title id="17DC10243145A43BCF2121B13D79FA48">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="1C79F5DEF09709F17FD4CF4D5FEE60CA">
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<mods:date id="8706924C2CE26C9D81B3C3083590BCD1">2006</mods:date>
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<mods:detail id="1A47CD241F949F3B48EAC7385D9ED4B0" type="volume">
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<mods:number id="F8A7E1140C75ACCDEEC408259EB85ADD">1285</mods:number>
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<mods:classification id="EFBC4B52B08908568CDD5F04CCF66F39">journal article</mods:classification>
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<mods:identifier id="7EFAB61EFDE2A40394945B4802F21CB3" type="DOI">10.5281/zenodo.173478</mods:identifier>
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<mods:identifier id="09B6980F1B52AAC527154233B0FCF4CB" type="GBIF-Dataset">1e858e0a-a143-49df-bd29-aa89c25c41e8</mods:identifier>
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<treatment id="03A50A52FFFBFF82FE80E748FC50F90D" ID-DOI="http://doi.org/10.5281/zenodo.6263393" ID-GBIF-Taxon="119347814" ID-Zenodo-Dep="6263393" LSID="urn:lsid:plazi:treatment:03A50A52FFFBFF82FE80E748FC50F90D" httpUri="http://treatment.plazi.org/id/03A50A52FFFBFF82FE80E748FC50F90D" lastPageId="6" lastPageNumber="57" pageId="1" pageNumber="52">
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<subSubSection id="C316E8CFFFFBFF85FE80E748FCB6F8FB" box="[264,774,1798,1824]" pageId="1" pageNumber="52" type="nomenclature">
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<paragraph id="8BB3BB44FFFBFF85FE80E748FCB6F8FB" blockId="1.[264,774,1798,1824]" box="[264,774,1798,1824]" pageId="1" pageNumber="52">
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<heading id="D0FB0C28FFFBFF85FE80E748FCB6F8FB" bold="true" box="[264,774,1798,1824]" fontSize="11" level="1" pageId="1" pageNumber="52" reason="1">
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<emphasis id="B9786756FFFBFF85FE80E748FCB6F8FB" bold="true" box="[264,774,1798,1824]" pageId="1" pageNumber="52">
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<taxonomicName id="4C0CC0C7FFFBFF85FE80E748FDDFF8FB" box="[264,623,1798,1824]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="52" phylum="Chordata" rank="species" species="corrugatus" status="sp. nov.">
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<emphasis id="B9786756FFFBFF85FE80E748FDDFF8FB" bold="true" box="[264,623,1798,1824]" italics="true" pageId="1" pageNumber="52">Eleutherodactylus corrugatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24BDA2DFFFBFF85FDFEE748FCB6F8FB" box="[630,774,1798,1824]" pageId="1" pageNumber="52" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C316E8CFFFFBFF86FE80E71FFEC2FD95" lastPageId="2" lastPageNumber="53" pageId="1" pageNumber="52" type="materials_examined">
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<paragraph id="8BB3BB44FFFBFF86FE80E71FFD55FE85" blockId="1.[264,1322,1873,1899]" lastBlockId="2.[264,1325,284,590]" lastPageId="2" lastPageNumber="53" pageId="1" pageNumber="52">
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<emphasis id="B9786756FFFBFF85FE80E71FFECDF8B1" box="[264,381,1873,1898]" italics="true" pageId="1" pageNumber="52">
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<typeStatus id="54B705E6FFFBFF85FE80E71FFEC6F8B1" box="[264,374,1873,1898]" pageId="1" pageNumber="52" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<collectionCode id="ED1D2381FFFBFF85FE0DE71FFE44F8B0" box="[389,500,1873,1899]" pageId="1" pageNumber="52">MHNSM</collectionCode>
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28063, an adult female, from Ullilen, Provincia de Mariscal Cáceres, Departamento de San Martín,
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<collectingCountry id="F31BFBD4FFF8FF86FDE5E152FD13FEED" box="[621,675,284,310]" name="Peru" pageId="2" pageNumber="53">Peru</collectingCountry>
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(
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<geoCoordinate id="EE38DD83FFF8FF86FD3BE152FCF3FEED" box="[691,835,284,310]" direction="south" orientation="latitude" pageId="2" pageNumber="53" precision="15" value="-6.846945">06°50'49" S</geoCoordinate>
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,
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<geoCoordinate id="EE38DD83FFF8FF86FCD9E152FC5AFEED" box="[849,1002,284,310]" direction="west" orientation="longitude" pageId="2" pageNumber="53" precision="15" value="-77.69444">77°41'40" W</geoCoordinate>
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,
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<quantity id="4CF416A1FFF8FF86FC7EE152FBE0FEED" box="[1014,1104,284,310]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="53" unit="m" value="3000.0">3000 m</quantity>
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a.s.l.) obtained on
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<date id="FFB29D84FFF8FF86FE80E10AFE5BFE85" box="[264,491,324,350]" pageId="2" pageNumber="53" value="2005-11-30">30 November 2005</date>
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by Pablo J. Venegas.
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</paragraph>
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<paragraph id="8BB3BB44FFF8FF86FEB0E122FEC2FD95" blockId="2.[264,1325,284,590]" pageId="2" pageNumber="53">
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<emphasis id="B9786756FFF8FF86FEB0E122FE09FE5E" box="[312,441,364,389]" italics="true" pageId="2" pageNumber="53">
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<typeStatus id="54B705E6FFF8FF86FEB0E122FE02FE5E" box="[312,434,364,389]" pageId="2" pageNumber="53" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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<collectionCode id="ED1D2381FFF8FF86FE49E122FD80FE5D" box="[449,560,364,390]" pageId="2" pageNumber="53">MHNSM</collectionCode>
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28062, 28064–67, two males, three females, and two juveniles, from Quintecocha (
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<geoCoordinate id="EE38DD83FFF8FF86FE64E1DAFDC9FE75" box="[492,633,404,430]" direction="south" orientation="latitude" pageId="2" pageNumber="53" precision="15" value="-6.858333">06º51'30" S</geoCoordinate>
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,
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<geoCoordinate id="EE38DD83FFF8FF86FD0EE1DAFCACFE75" box="[646,796,404,430]" direction="west" orientation="longitude" pageId="2" pageNumber="53" precision="15" value="-77.7">77º42' 00"W</geoCoordinate>
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,
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<quantity id="4CF416A1FFF8FF86FCAFE1DAFC30FE75" box="[807,896,404,430]" metricMagnitude="3" metricUnit="m" metricValue="3.13" pageId="2" pageNumber="53" unit="m" value="3130.0">3130 m</quantity>
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a.s.l.) obtained on
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<date id="FFB29D84FFF8FF86FBE8E1DAFEF3FE0D" pageId="2" pageNumber="53" value="2005-11-20" valueMax="2005-11-24" valueMin="2005-11-20">20–24 November 2005</date>
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by Pablo J. Venegas, an
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<collectionCode id="ED1D2381FFF8FF86FDF8E1F2FD6FFE0D" box="[624,735,444,470]" pageId="2" pageNumber="53">MHNSM</collectionCode>
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24444, 28068–69, one female and two juveniles, from Laguna El Plomo, (
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<geoCoordinate id="EE38DD83FFF8FF86FDB9E1AAFD76FE25" box="[561,710,484,510]" direction="south" orientation="latitude" pageId="2" pageNumber="53" precision="1" value="-6.8508887">6º51'03.2" S</geoCoordinate>
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,
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<geoCoordinate id="EE38DD83FFF8FF86FD5CE1AAFCDAFE25" box="[724,874,484,510]" direction="west" orientation="longitude" pageId="2" pageNumber="53" precision="15" value="-77.71667">77º43'00" W</geoCoordinate>
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,
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<quantity id="4CF416A1FFF8FF86FCFFE1AAFC61FE25" box="[887,977,484,510]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="2" pageNumber="53" unit="m" value="3300.0">3300 m</quantity>
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a.s.l.), Provincia de Mariscal Cáceres, Departamento de San Martín,
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<collectingCountry id="F31BFBD4FFF8FF86FD56E242FCA7FDFD" box="[734,791,524,550]" name="Peru" pageId="2" pageNumber="53">Peru</collectingCountry>
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||
, obtained on
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<date id="FFB29D84FFF8FF86FC31E242FB2FFDFD" box="[953,1183,524,550]" pageId="2" pageNumber="53" value="2005-11-28">28 November 2005</date>
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by Pablo J. Venegas.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C316E8CFFFF8FF87FE80E2CAFE2CFE25" lastPageId="3" lastPageNumber="54" pageId="2" pageNumber="53" type="diagnosis">
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||
<paragraph id="8BB3BB44FFF8FF86FE80E2CAFECFFD46" blockId="2.[264,1324,644,1870]" box="[264,383,644,669]" pageId="2" pageNumber="53">
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||
<emphasis id="B9786756FFF8FF86FE80E2CAFECFFD46" box="[264,383,644,669]" italics="true" pageId="2" pageNumber="53">Diagnosis</emphasis>
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||
</paragraph>
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<paragraph id="8BB3BB44FFF8FF86FEB0E2E2FAB8FAC5" blockId="2.[264,1324,644,1870]" pageId="2" pageNumber="53">
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A member of the
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<taxonomicName id="4C0CC0C7FFF8FF86FD8DE2E2FB91FD1E" box="[517,1057,684,710]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="53" phylum="Chordata" rank="species" species="orestes" subGenus="Eleutherodactylus">
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<emphasis id="B9786756FFF8FF86FD8DE2E2FD69FD1E" box="[517,729,684,709]" italics="true" pageId="2" pageNumber="53">Eleutherodactylus</emphasis>
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||
(
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<emphasis id="B9786756FFF8FF86FD62E2E2FC0EFD1E" box="[746,958,684,709]" italics="true" pageId="2" pageNumber="53">Eleutherodactylus</emphasis>
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)
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<emphasis id="B9786756FFF8FF86FC47E2E2FB91FD1E" box="[975,1057,684,709]" italics="true" pageId="2" pageNumber="53">orestes</emphasis>
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</taxonomicName>
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Group having (1) skin on dorsum shagreen to finely tuberculate with irregular longitudinal ridges; skin on venter areolate; discoidal fold not evident; dorsolateral folds absent; (2) tympanic membrane smooth; tympanic annulus distinct, round, its length slightly more than half that of eye; (3) snout short, rounded in dorsal view and profile, bearing low terminal tubercle; lips rounded; (4) upper eyelid bearing prominent conical tubercle, 60–75% IOD; cranial crests absent; (5) dentigerous processes of vomers absent; (6) males having vocal slits and nonspinous nuptial pads; (7) Finger I shorter than II; discs on outer fingers expanded, elliptical, half again width of digit proximal to pad; (8) fingers bearing broad lateral fringes; (9) ulnar tubercles small, elongate; (10) heel bearing conical tubercle; outer edge of tarsus with low, diffuse tubercle; inner tarsal fold barely evident distally; (11) inner metatarsal tubercle broadly ovoid, about
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<date id="FFB29D84FFF8FF86FCBCE42AFCE2FBA6" box="[820,850,1122,1150]" pageId="2" pageNumber="53">6x</date>
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elongate outer metatarsal tubercle; supernumerary plantar tubercles present; (12) toes bearing broad lateral fringes; Toe V slightly longer than Toe III; discs slightly smaller than those on fingers; (13) dorsum reddish tan with brown markings; venter creamy orange with brown reticulations and flecks; groin pale orange; (14) SVL
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<quantity id="4CF416A1FFF8FF86FD3AE54AFCECFAC5" box="[690,860,1284,1310]" metricMagnitude="-2" metricUnit="m" metricValue="1.9649999999999999" metricValueMax="1.98" metricValueMin="1.95" pageId="2" pageNumber="53" unit="mm" value="19.65" valueMax="19.8" valueMin="19.5">19.5–19.8 mm</quantity>
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in males,
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<quantity id="4CF416A1FFF8FF86FC5DE54AFB30FAC5" box="[981,1152,1284,1310]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" metricValueMax="2.62" metricValueMin="2.58" pageId="2" pageNumber="53" unit="mm" value="26.0" valueMax="26.2" valueMin="25.8">25.8–26.2 mm</quantity>
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in females.
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</paragraph>
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<paragraph id="8BB3BB44FFF8FF87FEB0E562FE2CFE25" blockId="2.[264,1324,644,1870]" lastBlockId="3.[264,1323,284,510]" lastPageId="3" lastPageNumber="54" pageId="2" pageNumber="53">
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Among the 10 species assigned to the
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<taxonomicName id="4C0CC0C7FFF8FF86FC8AE562FB81FA9E" box="[770,1073,1324,1349]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="53" phylum="Chordata" rank="species" species="orestes">
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||
<emphasis id="B9786756FFF8FF86FC8AE562FB81FA9E" box="[770,1073,1324,1349]" italics="true" pageId="2" pageNumber="53">Eleutherodactylus orestes</emphasis>
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||
</taxonomicName>
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||
Group (
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||
<bibRefCitation id="EF9DC6B5FFF8FF86FB1DE562FE57FAB5" author="Duellman" pageId="2" pageNumber="63" refString="Duellman, W. E. & Pramuk, J. B. (1999) Frogs of the genus Eleutherodactylus (Anura: Leptodactylidae) in the Andes of northern Peru. Scientific Papers, Natural History Museum University of Kansas, 13, 1 - 78." type="journal article" year="1999">Duellman & and Pramuk 1999</bibRefCitation>
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||
; this paper),
|
||
<taxonomicName id="4C0CC0C7FFF8FF86FD1AE51AFC8AFAB6" box="[658,826,1364,1389]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="53" phylum="Chordata" rank="species" species="corrugatus">
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||
<emphasis id="B9786756FFF8FF86FD1AE51AFC8AFAB6" box="[658,826,1364,1389]" italics="true" pageId="2" pageNumber="53">E. corrugatus</emphasis>
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</taxonomicName>
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is unique in having prominent, conical tubercles on the upper eyelids and heels, and by lacking contrasting markings in the groin. Four other species of
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<taxonomicName id="4C0CC0C7FFF8FF86FD8EE5EAFD6AFA66" box="[518,730,1444,1469]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="53" phylum="Chordata" rank="genus">
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<emphasis id="B9786756FFF8FF86FD8EE5EAFD6AFA66" box="[518,730,1444,1469]" italics="true" pageId="2" pageNumber="53">Eleutherodactylus</emphasis>
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</taxonomicName>
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in northern
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<collectingCountry id="F31BFBD4FFF8FF86FCE2E5EAFC10FA65" box="[874,928,1444,1470]" name="Peru" pageId="2" pageNumber="53">Peru</collectingCountry>
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have prominent conical tubercles on the upper eyelids and heels; in all of these, Toe V is much longer than Toe III. Three of these species (
|
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<taxonomicName id="4C0CC0C7FFF8FF86FE3CE5BAFDCBF9D5" authority="Jimenez" authorityName="Jimenez" box="[436,635,1524,1550]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="53" phylum="Chordata" rank="species" species="galdi">
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<emphasis id="B9786756FFF8FF86FE3CE5BAFDA2F9D6" box="[436,530,1524,1549]" italics="true" pageId="2" pageNumber="53">E. galdi</emphasis>
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Jiménez
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</taxonomicName>
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de la Espada,
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<taxonomicName id="4C0CC0C7FFF8FF86FCA2E5BAFB61F9D5" authority="Duellman and Pramuk" authorityName="Duellman and Pramuk" box="[810,1233,1524,1550]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="53" phylum="Chordata" rank="species" species="muscosus">
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<emphasis id="B9786756FFF8FF86FCA2E5BAFC0CF9D6" box="[810,956,1524,1549]" italics="true" pageId="2" pageNumber="53">E. muscosus</emphasis>
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||
Duellman and Pramuk
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</taxonomicName>
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, and
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<taxonomicName id="4C0CC0C7FFF8FF86FA99E5BAFDF3F9ED" authority="Duellman and Wild" authorityName="Duellman and Wild" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="53" phylum="Chordata" rank="species" species="wiensi">
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<emphasis id="B9786756FFF8FF86FA99E5BAFEE3F9EE" italics="true" pageId="2" pageNumber="53">E. wiensi</emphasis>
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Duellman and Wild
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</taxonomicName>
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) are larger frogs (males to
|
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<quantity id="4CF416A1FFF8FF86FC09E652FC64F9ED" box="[897,980,1564,1590]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="2" pageNumber="53" unit="mm" value="33.0">33 mm</quantity>
|
||
and females to
|
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<quantity id="4CF416A1FFF8FF86FB06E652FB51F9ED" box="[1166,1249,1564,1590]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="2" pageNumber="53" unit="mm" value="36.0">36 mm</quantity>
|
||
SVL) that have large, truncate discs. Furthermore,
|
||
<taxonomicName id="4C0CC0C7FFF8FF86FC9CE60AFCC1F986" box="[788,881,1604,1629]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="53" phylum="Chordata" rank="species" species="galdi">
|
||
<emphasis id="B9786756FFF8FF86FC9CE60AFCC1F986" box="[788,881,1604,1629]" italics="true" pageId="2" pageNumber="53">E. galdi</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C0CC0C7FFF8FF86FB85E60AFB1EF986" box="[1037,1198,1604,1629]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="53" phylum="Chordata" rank="species" species="corrugatus">
|
||
<emphasis id="B9786756FFF8FF86FB85E60AFB1EF986" box="[1037,1198,1604,1629]" italics="true" pageId="2" pageNumber="53">E. corrugatus</emphasis>
|
||
</taxonomicName>
|
||
by having an acuminate snout, conical tubercles along the other edges of the forearm and foot, and cranial crests. In
|
||
<emphasis id="B9786756FFF8FF86FE5DE6DAFDDFF976" box="[469,623,1684,1709]" italics="true" pageId="2" pageNumber="53">
|
||
<taxonomicName id="4C0CC0C7FFF8FF86FE5DE6DAFDDBF976" box="[469,619,1684,1709]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="53" phylum="Chordata" rank="species" species="muscosus">E. muscosus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
there are two subconical tubercles on the posterior part of the upper eyelid, and the belly and ventral surfaces of the hind limbs are dark brown with creamywhite spots.
|
||
<taxonomicName id="4C0CC0C7FFF8FF86FD89E6AAFC9AF926" box="[513,810,1764,1789]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="53" phylum="Chordata" rank="species" species="wiensi">
|
||
<emphasis id="B9786756FFF8FF86FD89E6AAFC9AF926" box="[513,810,1764,1789]" italics="true" pageId="2" pageNumber="53">Eleutherodactylus wiensi</emphasis>
|
||
</taxonomicName>
|
||
lacks a tympanic membrane, and only the ventral part of the tympanic annulus is visible. The fourth species,
|
||
<taxonomicName id="4C0CC0C7FFF8FF86FB95E742FA95F8FD" authority="Lynch" authorityName="Lynch" box="[1053,1317,1804,1830]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="53" phylum="Chordata" rank="species" species="colodactylus">
|
||
<emphasis id="B9786756FFF8FF86FB95E742FB62F8FE" box="[1053,1234,1804,1829]" italics="true" pageId="2" pageNumber="53">E. colodactylus</emphasis>
|
||
Lynch
|
||
</taxonomicName>
|
||
, has short, stocky fingers and a yellow spot in the groin; it lacks vocal slits, a tympanic membrane, and tympanic annulus. The only other species in northern
|
||
<collectingCountry id="F31BFBD4FFF9FF87FBEFE152FB2DFEED" box="[1127,1181,284,310]" name="Peru" pageId="3" pageNumber="54">Peru</collectingCountry>
|
||
that bear a tubercle on the snout are
|
||
<taxonomicName id="4C0CC0C7FFF9FF87FDB4E10AFC5AFE85" authority="Duellman and Pramuk" authorityName="Duellman and Pramuk" box="[572,1002,324,350]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="54" phylum="Chordata" rank="species" species="anemerus">
|
||
<emphasis id="B9786756FFF9FF87FDB4E10AFD61FE86" box="[572,721,324,349]" italics="true" pageId="3" pageNumber="54">E. anemerus</emphasis>
|
||
Duellman and Pramuk
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C0CC0C7FFF9FF87FBA4E10AFA9AFE85" authority="Lynch." authorityName="Lynch." box="[1068,1322,324,350]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="54" phylum="Chordata" rank="species" species="proserpens">
|
||
<emphasis id="B9786756FFF9FF87FBA4E10AFB60FE86" box="[1068,1232,324,349]" italics="true" pageId="3" pageNumber="54">E. proserpens</emphasis>
|
||
Lynch.
|
||
</taxonomicName>
|
||
<taxonomicName id="4C0CC0C7FFF9FF87FE80E122FDEAFE5E" box="[264,602,364,389]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="54" phylum="Chordata" rank="species" species="anemerus">
|
||
<emphasis id="B9786756FFF9FF87FE80E122FDEAFE5E" box="[264,602,364,389]" italics="true" pageId="3" pageNumber="54">Eleutherodactylus anemerus</emphasis>
|
||
</taxonomicName>
|
||
lacks tubercles on the upper eyelids and heels, and has a uniform orangered dorsum;
|
||
<taxonomicName id="4C0CC0C7FFF9FF87FDD3E1DAFD4CFE76" box="[603,764,404,429]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="54" phylum="Chordata" rank="species" species="proserpens">
|
||
<emphasis id="B9786756FFF9FF87FDD3E1DAFD4CFE76" box="[603,764,404,429]" italics="true" pageId="3" pageNumber="54">E. proserpens</emphasis>
|
||
</taxonomicName>
|
||
has an acuminate snout, prominent dentigerous processes of the vomers, and low, round tubercles on the upper eyelids, and lacks tubercles on the heels.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF73EBCCFFF9FF87FE80E628FC6AF97B" httpUri="https://zenodo.org/record/173479/files/figure.png" pageId="3" pageNumber="54" targetBox="[359,1225,568,1606]" targetPageId="3">
|
||
<paragraph id="8BB3BB44FFF9FF87FE80E628FC6AF97B" blockId="3.[264,1322,1638,1696]" pageId="3" pageNumber="54">
|
||
<emphasis id="B9786756FFF9FF87FE80E628FE3DF9A5" bold="true" box="[264,397,1638,1662]" pageId="3" pageNumber="54">FIGURE 1.</emphasis>
|
||
Holotype of
|
||
<emphasis id="B9786756FFF9FF87FDACE628FCDAF9A6" box="[548,874,1638,1661]" italics="true" pageId="3" pageNumber="54">
|
||
<taxonomicName id="4C0CC0C7FFF9FF87FDACE628FCD6F9A6" box="[548,870,1638,1661]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="54" phylum="Chordata" rank="species" species="corrugatus">Eleutherodactylus corrugatus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
MHNSM 28063. Lateral (A) and dorsal (B) views of head. C. Palmar view of hand. D. Plantar view of foot.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C316E8CFFFF9FF82FE80E69AFEC3FBF5" lastPageId="6" lastPageNumber="57" pageId="3" pageNumber="54" type="description">
|
||
<paragraph id="8BB3BB44FFF9FF87FE80E69AFDF8F936" blockId="3.[264,1325,1748,1893]" box="[264,584,1748,1773]" pageId="3" pageNumber="54">
|
||
<emphasis id="B9786756FFF9FF87FE80E69AFDF8F936" box="[264,584,1748,1773]" italics="true" pageId="3" pageNumber="54">
|
||
Description of the
|
||
<typeStatus id="54B705E6FFF9FF87FE6BE69AFDF8F936" box="[483,584,1748,1773]" pageId="3" pageNumber="54" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB3BB44FFF9FF80FEB0E6B5FE61FD45" blockId="3.[264,1325,1748,1893]" lastBlockId="4.[264,1325,284,1870]" lastPageId="4" lastPageNumber="55" pageId="3" pageNumber="54">
|
||
Adult female with robust body; head narrow, not as wide as body, wider than long; head width 38.9% of SVL; head length 35.9% of SVL, 92.2% of head width; snout short with small terminal tubercle barely evident in lateral view; snout rounded in dorsal view, bluntly rounded in profile (
|
||
<figureCitation id="1337A7C1FFFEFF80FDDEE152FD16FEED" box="[598,678,284,310]" captionStart="FIGURE 1" captionStartId="3.[264,367,1638,1662]" captionTargetBox="[359,1225,568,1606]" captionTargetId="figure@3.[345,1241,549,1611]" captionTargetPageId="3" captionText="FIGURE 1. Holotype of Eleutherodactylus corrugatus, MHNSM 28063. Lateral (A) and dorsal (B) views of head. C. Palmar view of hand. D. Plantar view of foot." httpUri="https://zenodo.org/record/173479/files/figure.png" pageId="4" pageNumber="55">Figs. 1</figureCitation>
|
||
A and B); diameter of eye slightly greater than eyenostril distance; nostrils distinctly protuberant laterally; canthus rostralis curved in dorsal view, nearly angular in section; loreal region concave; lips rounded; upper eyelid bearing prominent, conical tubercle; width of upper eyelid 74% of IOD; tympanic annulus round, its posterodorsal part obscured by laterally projecting supratympanic fold; diameter of tympanum 55.6% of diameter of eye, separated from eye by distance slightly less than diameter of tympanum; two subconical postrictal tubercles. Choanae small, ovoid, not concealed by palatal shelf of maxillae; vomers lacking visible dentigerous processes; tongue cordiform, as wide as long, shallowly notched posteriorly, free behind for about half of its length.
|
||
</paragraph>
|
||
<paragraph id="8BB3BB44FFFEFF80FEB0E2E2FC7DFC55" blockId="4.[264,1325,284,1870]" pageId="4" pageNumber="55">Skin on dorsum shagreen, becoming finely tuberculate laterally, with low irregular paravertebral and scapular ridges, longitudinally sinusoidal ridge on upper flank, and low ridges and subconical tubercles on limbs; skin on flanks finely tuberculate above, tubercles becoming larger ventrally; skin on throat, chest, belly, and ventral surfaces of thighs coarsely areolate; other ventral surfaces smooth; discoidal and thoracic folds not evident; cloacal sheath short; ornamentation in cloacal region absent.</paragraph>
|
||
<paragraph id="8BB3BB44FFFEFF80FEB0E3D2FC11FB7D" blockId="4.[264,1325,284,1870]" pageId="4" pageNumber="55">
|
||
Single, low, elongate ulnar tubercle on each forearm; palmar tubercle elevated, bifid, about twice as large as elliptical thenar tubercle; subarticular tubercles prominent, round, and subconical in section; supernumerary tubercles low, round, about half size of subarticular tubercles; fingers short, bearing broad lateral fringes (
|
||
<figureCitation id="1337A7C1FFFEFF80FBCEE45AFB20FBF5" box="[1094,1168,1044,1070]" captionStart="FIGURE 1" captionStartId="3.[264,367,1638,1662]" captionTargetBox="[359,1225,568,1606]" captionTargetId="figure@3.[345,1241,549,1611]" captionTargetPageId="3" captionText="FIGURE 1. Holotype of Eleutherodactylus corrugatus, MHNSM 28063. Lateral (A) and dorsal (B) views of head. C. Palmar view of hand. D. Plantar view of foot." httpUri="https://zenodo.org/record/173479/files/figure.png" pageId="4" pageNumber="55">Fig. 1</figureCitation>
|
||
C); Finger I shorter than Finger II; disc on Finger I slightly expanded; those on Fingers II–IV expanded, elliptical, half again as wide as digit proximal to disc; ventral digital pads well defined by circumferential grooves, weakest on Finger I.
|
||
</paragraph>
|
||
<paragraph id="8BB3BB44FFFEFF80FEB0E4FAFAA5FA3D" blockId="4.[264,1325,284,1870]" pageId="4" pageNumber="55">
|
||
<collectingCountry id="F31BFBD4FFFEFF80FEB0E4FAFEC3FB15" box="[312,371,1204,1230]" name="India" pageId="4" pageNumber="55">Hind</collectingCountry>
|
||
limbs moderately robust; tibia length 45.0% of SVL; foot length 45.4% of SVL; heel bearing distinct conical tubercle; outer edge of tarsus with three, low, subconical tubercles; inner tarsal fold barely evident distally; inner metatarsal tubercle elevated, ovoid, about
|
||
<date id="FFB29D84FFFEFF80FE2EE562FE74FA9E" box="[422,452,1322,1350]" pageId="4" pageNumber="55">6x</date>
|
||
elongate outer metatarsal tubercle; toes unwebbed, bearing broad lateral fringes (
|
||
<figureCitation id="1337A7C1FFFEFF80FEE3E51AFE1EFAB5" box="[363,430,1364,1390]" captionStart="FIGURE 1" captionStartId="3.[264,367,1638,1662]" captionTargetBox="[359,1225,568,1606]" captionTargetId="figure@3.[345,1241,549,1611]" captionTargetPageId="3" captionText="FIGURE 1. Holotype of Eleutherodactylus corrugatus, MHNSM 28063. Lateral (A) and dorsal (B) views of head. C. Palmar view of hand. D. Plantar view of foot." httpUri="https://zenodo.org/record/173479/files/figure.png" pageId="4" pageNumber="55">Fig. 1</figureCitation>
|
||
D); discs on toes about as large as those on fingers; all toes having ventral pads well defined by circumferential grooves; relative lengths of toes 1<2<3<5<4; Toe V slightly longer than Toe III; disc on Toe III not reaching distal subarticular tubercle on Toe IV; disc on Toe V extending to proximal border of distal subarticular tubercle on Toe IV.
|
||
</paragraph>
|
||
<paragraph id="8BB3BB44FFFEFF81FEB0E5BAFAA5FE5D" blockId="4.[264,1325,284,1870]" lastBlockId="5.[264,1323,284,590]" lastPageId="5" lastPageNumber="56" pageId="4" pageNumber="55">
|
||
Coloration of
|
||
<typeStatus id="54B705E6FFFEFF80FE6AE5BAFDF9F9D5" box="[482,585,1524,1550]" pageId="4" pageNumber="55" type="holotype">holotype</typeStatus>
|
||
in preservative: Dorsum grayish brown with brownishblack markings consisting of interocular bar with broad extension posteromedially to occiput, shallow Wshaped mark in scapular region, small diffuse spots on head and body, three faint diagonal marks on each flank, diffuse canthal stripe, two irregular labial bars on each upper lip, welldefined crescentshaped mark below supratympanic fold, one elongate diagonal bar and one transverse bar on each thigh, and irregular transverse bars on limbs (4 on shank, 3 on tarsus, 3 on forearm). Broad creamytan transverse mark on top of head anterior to interocular bar and with posterolateral corners on supraocular tubercles; groin pale tan; posterior surfaces of thighs dark brown with bold, cream mottling. Throat, chest, belly, and ventral surfaces of limbs creamy tan with brown flecks, most dense on margin of chin; ventral surfaces of outer two fingers and toes brown; ventral surfaces of other digits yellowish tan; dorsal surfaces of Fingers I and II and Toes I–III cream with brown spots.
|
||
</paragraph>
|
||
<paragraph id="8BB3BB44FFFFFF81FEB0E1DAFD60FD95" blockId="5.[264,1323,284,590]" pageId="5" pageNumber="56">
|
||
Coloration of
|
||
<typeStatus id="54B705E6FFFFFF81FE65E1DAFDE4FE75" box="[493,596,404,430]" pageId="5" pageNumber="56" type="holotype">holotype</typeStatus>
|
||
in life: Dorsum reddish tan with dark brown markings, including labial bars, interocular bar, diagonal bars on flanks, transverse bars on limbs, and with broad cream band across top of head and upper eyelids (
|
||
<figureCitation id="1337A7C1FFFFFF81FB8DE1AAFBE1FE25" box="[1029,1105,484,510]" captionStart="FIGURE 2" captionStartId="5.[264,368,1214,1238]" captionTargetBox="[409,1177,649,1187]" captionTargetId="figure@5.[409,1177,649,1187]" captionTargetPageId="5" captionText="FIGURE 2. Holotype of Eleutherodactylus corrugatus in life, MHNSM 28063, 26.2 mm SVL. Photo by P. Venegas." httpUri="https://zenodo.org/record/173480/files/figure.png" pageId="5" pageNumber="56">Fig. 2</figureCitation>
|
||
); venter dull pale orange with brown flecks and reticulations; iris reddish tan dorsally, grayish bronze below narrow median black triangular marks.
|
||
</paragraph>
|
||
<caption id="DF73EBCCFFFFFF81FE80E4F0FE58FB23" httpUri="https://zenodo.org/record/173480/files/figure.png" pageId="5" pageNumber="56" targetBox="[409,1177,649,1187]" targetPageId="5">
|
||
<paragraph id="8BB3BB44FFFFFF81FE80E4F0FE58FB23" blockId="5.[264,1323,1214,1272]" pageId="5" pageNumber="56">
|
||
<emphasis id="B9786756FFFFFF81FE80E4F0FE3EFB0D" bold="true" box="[264,398,1214,1238]" pageId="5" pageNumber="56">FIGURE 2.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="4C0CC0C7FFFFFF81FDADE4F0FCD5FB0E" box="[549,869,1214,1237]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="56" phylum="Chordata" rank="species" species="corrugatus">
|
||
<emphasis id="B9786756FFFFFF81FDADE4F0FCD5FB0E" box="[549,869,1214,1237]" italics="true" pageId="5" pageNumber="56">Eleutherodactylus corrugatus</emphasis>
|
||
</taxonomicName>
|
||
in life, MHNSM 28063, 26.2 mm SVL. Photo by P. Venegas.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB3BB44FFFFFF81FEB0E565FC87FA4E" blockId="5.[264,1322,1323,1429]" pageId="5" pageNumber="56">
|
||
Measurements of
|
||
<typeStatus id="54B705E6FFFFFF81FD82E565FDC1FA9E" box="[522,625,1323,1349]" pageId="5" pageNumber="56" type="holotype">holotype</typeStatus>
|
||
in mm: SVL 26.2, tibia length, 11.8, foot length 11.6, head length 9.4, head width 10.2, IOD 2.7, width of upper eyelid 2.0; eye–nostril distance 2.3; diameter of eye 2.7, diameter of tympanum 1.5.
|
||
</paragraph>
|
||
<paragraph id="8BB3BB44FFFFFF81FE80E582FEC1FA3E" blockId="5.[264,1325,1484,1869]" box="[264,369,1484,1509]" pageId="5" pageNumber="56">
|
||
<emphasis id="B9786756FFFFFF81FE80E582FEC1FA3E" box="[264,369,1484,1509]" italics="true" pageId="5" pageNumber="56">Va r i a t i o n</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB3BB44FFFFFF82FEB0E5BDFE3DFEED" blockId="5.[264,1325,1484,1869]" lastBlockId="6.[264,1325,284,1070]" lastPageId="6" lastPageNumber="57" pageId="5" pageNumber="56">
|
||
Conical tubercles are present on the eyelids and heels of all specimens, but those on the eyelid are not as well developed in juveniles as in adults. Also, a tubercle on the snout is not evident in the two juveniles (MHNSM 23066, 23068), one male (MHNSM 23069), and one female (MHNSM 23065). The degree of tuberculation and development of dermal ridges on the dorsum and flanks in two females (MHNSM 23062, 23065) and one male (MHNSM 23067) is equivalent to that in the female
|
||
<typeStatus id="54B705E6FFFFFF81FCF7E6F5FC59F90E" box="[895,1001,1723,1749]" pageId="5" pageNumber="56" type="holotype">holotype</typeStatus>
|
||
, whereas the dorsal part of the flank is shagreen in two females (MHNSM 23064, 24444) and one male (MHNSM 23069). The skin on the dorsum is simply shagreen in the two juveniles. Males have vocal slits and weakly spinous, unpigmented nuptial pads on the dorsomedial surface of the base of Finger I.
|
||
</paragraph>
|
||
<paragraph id="8BB3BB44FFFCFF82FEB0E10AFC0CFCCD" blockId="6.[264,1325,284,1070]" pageId="6" pageNumber="57">Males are smaller than females and have proportionately longer legs, narrower IODs, and smaller tympani. The ranges and means (in parentheses) of measurements (in mm) of five females are followed by the ranges of two males: SVL 24.3–26.2 (25.7), 19.5–19.8; tibia length 11.0–11.9 (11.5), 9.4–9.5; foot length 11.6–12.0 (11.8), 9.6–9.7; head length 8.9–9.8 (9.3), 6.5–6.8; head width 9.3–10.5 (9.9), 7.2; IOD 2.6–2.9 (2.8), 2.2–2.4; width of upper eyelid 1.8–2.2 (2.0), 1.5–1.6; eye–nostril distance 1.9–2.4 (2.2), 1.8–1.9; diameter of eye 2.4–3.0 (2.7), 2.2–2.3; diameter of tympanum 1.5–1.7 (1.6). The ranges of proportions (as percentages) in five females is followed by those of two males: tibia/SVL 43.5–46.1, 48.0–48.2; foot/SVL 44.8–47.7, 47.5–47.7; head length/SVL 34.0–39.1, 32.8–34.9; head width/SVL 36.0–40.2, 36.4–36.9; upper eyelid/IOD 69.2–75.9, 66.7–68.1; eye–nostril distance/eye 79.2–85.2, 81.8–82.6; tympanum/eye 55.6–66.7, 47.8–50.0. Two juveniles have SVLs of 13.0 and 16.0 mm.</paragraph>
|
||
<paragraph id="8BB3BB44FFFCFF82FEB0E36AFEC3FBF5" blockId="6.[264,1325,284,1070]" pageId="6" pageNumber="57">
|
||
In preservative, only one specimen (a juvenile, MHNSM 23066) resembles the
|
||
<typeStatus id="54B705E6FFFCFF82FE80E302FEDEFCBD" box="[264,366,844,870]" pageId="6" pageNumber="57" type="holotype">holotype</typeStatus>
|
||
in having a broad, pale transverse band on the head; this juvenile also has a brown venter with a median cream line. The other juvenile (MHNSM 23068) has dense brown pigment on the throat and belly. The dorsum in one male (MHNSM 23069) is tan with dark brown middorsal and dorsolateral stripes. In all specimens, the groin is pale tan to cream, and the posterior surfaces of the thighs are brown with varying amounts of cream mottling.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C316E8CFFFFCFF82FE80E42AFA9CF9ED" pageId="6" pageNumber="57" type="distribution">
|
||
<paragraph id="8BB3BB44FFFCFF82FE80E42AFD9DFBA6" blockId="6.[264,1324,1124,1590]" box="[264,557,1124,1149]" pageId="6" pageNumber="57">
|
||
<emphasis id="B9786756FFFCFF82FE80E42AFD9DFBA6" box="[264,557,1124,1149]" italics="true" pageId="6" pageNumber="57">Distribution and ecology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB3BB44FFFCFF82FEB0E4C2FA9CF9ED" blockId="6.[264,1324,1124,1590]" pageId="6" pageNumber="57">
|
||
<taxonomicName id="4C0CC0C7FFFCFF82FEB0E4C2FD11FB7E" box="[312,673,1164,1189]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="57" phylum="Chordata" rank="species" species="corrugatus">
|
||
<emphasis id="B9786756FFFCFF82FEB0E4C2FD11FB7E" box="[312,673,1164,1189]" italics="true" pageId="6" pageNumber="57">Eleutherodactylus corrugatus</emphasis>
|
||
</taxonomicName>
|
||
is known from three localities at elevations of
|
||
<quantity id="4CF416A1FFFCFF82FE80E4FAFE03FB15" box="[264,435,1204,1230]" metricMagnitude="3" metricUnit="m" metricValue="3.15" metricValueMax="3.3" metricValueMin="3.0" pageId="6" pageNumber="57" unit="m" value="3150.0" valueMax="3300.0" valueMin="3000.0">3000–3300 m</quantity>
|
||
in the northern part of the Cordillera Central in northern
|
||
<collectingCountry id="F31BFBD4FFFCFF82FB3FE4FAFB41FB15" box="[1207,1265,1204,1230]" name="Peru" pageId="6" pageNumber="57">Peru</collectingCountry>
|
||
. All individuals were found on leaves of low bushes,
|
||
<quantity id="4CF416A1FFFCFF82FCE8E492FC57FB2D" box="[864,999,1244,1270]" metricMagnitude="-1" metricUnit="m" metricValue="9.0" metricValueMax="15.0" metricValueMin="3.0" pageId="6" pageNumber="57" unit="cm" value="90.0" valueMax="150.0" valueMin="30.0">30–150 cm</quantity>
|
||
above ground at night. At Ullilen and Quintecocha the frogs were found in very humid montane forest, and at Laguna El Plomo individuals were in patches of elfin forest. Sympatric species at Ullilen include
|
||
<taxonomicName id="4C0CC0C7FFFCFF82FEE0E51AFE42FAB6" box="[360,498,1364,1389]" class="Amphibia" family="Dendrobatidae" genus="Colostethus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="57" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9786756FFFCFF82FEE0E51AFE42FAB6" box="[360,498,1364,1389]" italics="true" pageId="6" pageNumber="57">Colostethus</emphasis>
|
||
</taxonomicName>
|
||
sp.,
|
||
<taxonomicName id="4C0CC0C7FFFCFF82FDA6E51AFCB2FAB6" box="[558,770,1364,1389]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="57" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9786756FFFCFF82FDA6E51AFCB2FAB6" box="[558,770,1364,1389]" italics="true" pageId="6" pageNumber="57">Eleutherodactylus</emphasis>
|
||
</taxonomicName>
|
||
wagteri Venegas, and
|
||
<taxonomicName id="4C0CC0C7FFFCFF82FB9CE51AFDD4FA4D" authority="Duellman and Venegas. The" authorityName="Duellman and Venegas. The" class="Amphibia" family="Hemiphractidae" genus="Gastrotheca" kingdom="Animalia" order="Anura" pageId="6" pageNumber="57" phylum="Chordata" rank="species" species="ossilaginis">
|
||
<emphasis id="B9786756FFFCFF82FB9CE51AFA9AFAB6" box="[1044,1322,1364,1389]" italics="true" pageId="6" pageNumber="57">Gastrotheca ossilaginis</emphasis>
|
||
Duellman and Venegas. The
|
||
</taxonomicName>
|
||
other species found at Laguna de Quintecocha are
|
||
<taxonomicName id="4C0CC0C7FFFCFF82FB7BE532FD6EFA65" authority="Duellman and Schulte" authorityName="Duellman and Schulte" class="Amphibia" family="Bufonidae" genus="Bufo" kingdom="Animalia" order="Anura" pageId="6" pageNumber="57" phylum="Chordata" rank="species" species="arborescandens">
|
||
<emphasis id="B9786756FFFCFF82FB7BE532FE72FA66" italics="true" pageId="6" pageNumber="57">Bufo arborescandens</emphasis>
|
||
Duellman and Schulte
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0CC0C7FFFCFF82FD79E5EAFA96FA65" authority="Duellman and Schulte" authorityName="Duellman and Schulte" box="[753,1318,1444,1470]" class="Amphibia" family="Centrolenidae" genus="Centrolene" kingdom="Animalia" order="Anura" pageId="6" pageNumber="57" phylum="Chordata" rank="species" species="lemniscatus">
|
||
<emphasis id="B9786756FFFCFF82FD79E5EAFBB8FA66" box="[753,1032,1444,1469]" italics="true" pageId="6" pageNumber="57">Centrolene lemniscatus</emphasis>
|
||
Duellman and Schulte
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B9786756FFFCFF82FE80E582FD3AFA3E" box="[264,650,1484,1509]" italics="true" pageId="6" pageNumber="57">
|
||
<taxonomicName id="4C0CC0C7FFFCFF82FE80E582FE6FFA3E" box="[264,479,1484,1509]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="57" phylum="Chordata" rank="genus">Eleutherodactylus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0CC0C7FFFCFF82FE65E582FD3AFA3E" authority="Lynch" authorityName="Lynch" box="[493,650,1484,1509]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="57" phylum="Chordata" rank="species" species="bromeliaceus">bromeliaceus</taxonomicName>
|
||
</emphasis>
|
||
Lynch,
|
||
<taxonomicName id="4C0CC0C7FFFCFF82FD67E582FB2EFA3D" authority="Duellman and Pramuk" authorityName="Duellman and Pramuk" box="[751,1182,1484,1510]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="57" phylum="Chordata" rank="species" species="rufioculis">
|
||
<emphasis id="B9786756FFFCFF82FD67E582FC33FA3E" box="[751,899,1484,1509]" italics="true" pageId="6" pageNumber="57">E. rufioculis</emphasis>
|
||
Duellman and Pramuk
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0CC0C7FFFCFF82FB24E582FECFF9D5" authority="Duellman" authorityName="Duellman" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="57" phylum="Chordata" rank="species" species="schultei">
|
||
<emphasis id="B9786756FFFCFF82FB24E582FA9BFA3E" box="[1196,1323,1484,1509]" italics="true" pageId="6" pageNumber="57">E. schultei</emphasis>
|
||
Duellman
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0CC0C7FFFCFF82FE1EE5BAFDC5F9D6" box="[406,629,1524,1549]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="57" phylum="Chordata" rank="species" species="stictoboubonus" status="sp. nov.">
|
||
<emphasis id="B9786756FFFCFF82FE1EE5BAFDC5F9D6" box="[406,629,1524,1549]" italics="true" pageId="6" pageNumber="57">E. stictoboubonus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24BDA2DFFFCFF82FD0FE5BAFC92F9D5" box="[647,802,1524,1550]" pageId="6" pageNumber="57" rank="species">new species</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C0CC0C7FFFCFF82FCB2E5BAFBEAF9D6" box="[826,1114,1524,1549]" class="Amphibia" family="Hemiphractidae" genus="Gastrotheca" kingdom="Animalia" order="Anura" pageId="6" pageNumber="57" phylum="Chordata" rank="species" species="ossilaginis">
|
||
<emphasis id="B9786756FFFCFF82FCB2E5BAFBEAF9D6" box="[826,1114,1524,1549]" italics="true" pageId="6" pageNumber="57">Gastrotheca ossilaginis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0CC0C7FFFCFF82FB23E5BAFE6BF9ED" authority="Wiens" authorityName="Wiens" class="Amphibia" family="Brachycephalidae" genus="Telmatbius" kingdom="Animalia" order="Anura" pageId="6" pageNumber="57" phylum="Chordata" rank="species" species="atahualpai">
|
||
<emphasis id="B9786756FFFCFF82FB23E5BAFE37F9EE" italics="true" pageId="6" pageNumber="57">Telmatbius atahualpai</emphasis>
|
||
Wiens
|
||
</taxonomicName>
|
||
, whereas the latter was the only other species found at Laguna El Plomo.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C316E8CFFFFCFF82FE80E622FC50F90D" pageId="6" pageNumber="57" type="etymology">
|
||
<paragraph id="8BB3BB44FFFCFF82FE80E622FE35F95E" blockId="6.[264,1323,1644,1750]" box="[264,389,1644,1669]" pageId="6" pageNumber="57">
|
||
<emphasis id="B9786756FFFCFF82FE80E622FE35F95E" box="[264,389,1644,1669]" italics="true" pageId="6" pageNumber="57">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB3BB44FFFCFF82FEB0E6DAFC50F90D" blockId="6.[264,1323,1644,1750]" pageId="6" pageNumber="57">The specific epithet is a Latin adjective meaning wrinkled or ridged; the name alludes to the texture of the skin on the dorsal surfaces of this species.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |