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<document id="D9063E4579F28A5789A7986A94D818C3" ID-CLB-Dataset="20138" ID-DOI="10.5852/ejt.2022.806.1697" ID-GBIF-Dataset="28f53295-9155-4b84-ab43-45e852842817" ID-ISSN="2118-9773" ID-Zenodo-Dep="6373945" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1647883015262" checkinUser="felipe" docAuthor="Dang, Minh Quan, Nguyen, Manh Ha, Hoang, Nghia Son, Nguyen, Thi Mai Huong, Truong, Ba Vuong &amp; Dang, Van Son" docDate="2022" docId="0387F954292CFF86FE6A172AFE0DCCCC" docLanguage="en" docName="ejt-806_19-31.pdf" docOrigin="European Journal of Taxonomy 806" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Lasianthus sonlangensis V. S. Dang, Vuong &amp; Quan 2022, sp. nov." docType="treatment" docVersion="6" lastPageNumber="29" masterDocId="FFBE812C292AFF8CFFDD1542FF27C806" masterDocTitle="Two new species of Lasianthus Jack (Rubiaceae) from southern Vietnam" masterLastPageNumber="31" masterPageNumber="19" pageNumber="25" updateTime="1699319045260" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="40F55D1FE35BD930A8E14F52EEDD73B9">Two new species of Lasianthus Jack (Rubiaceae) from southern Vietnam</mods:title>
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<mods:namePart id="F33DC7579956C9BF189BB8B2DFA40779">Dang, Minh Quan</mods:namePart>
<mods:affiliation id="B3051AF9DA93E5AB382CC3CC4449A0F2">School of Education, Can Tho University, 3 / 2 Street, Ninh Kieu District, Can Tho City, Vietnam. Pedo-Geography and Soil Resources, Institute of Geography (IG), Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam. Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam. Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan.</mods:affiliation>
<mods:nameIdentifier id="3094CC0715107A4D3D4A9B369429080A" type="email">dmquan@ctu.edu.vn</mods:nameIdentifier>
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<mods:namePart id="C893D7CFD30B70436B3E559E9CC8DD55">Nguyen, Manh Ha</mods:namePart>
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<mods:namePart id="F79BC758BF5721B6C50804A6C790EAF2">Hoang, Nghia Son</mods:namePart>
<mods:nameIdentifier id="C8F31A66FAC13F291C0BFA14821F2311" type="email">hoangnghiason@yahoo.com&amp;tho1010@gmail.com</mods:nameIdentifier>
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<mods:namePart id="9CA1CA59FC21B37C1D0D549FBF054BB0">Nguyen, Thi Mai Huong</mods:namePart>
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<mods:namePart id="486E17C13611C5BC4BEA45B9C2608444">Truong, Ba Vuong</mods:namePart>
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<mods:namePart id="AD5C84E8FE8CF8F338653223048A45E7">Dang, Van Son</mods:namePart>
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<mods:title id="18EA86731E7F7C37212F711F87DB9BDD">European Journal of Taxonomy</mods:title>
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<mods:date id="0B6510D4AEA279B02BCCA191F78402D7">2022</mods:date>
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<treatment id="0387F954292CFF86FE6A172AFE0DCCCC" ID-DOI="http://doi.org/10.5281/zenodo.6379962" ID-GBIF-Taxon="194208392" ID-Zenodo-Dep="6379962" LSID="urn:lsid:plazi:treatment:0387F954292CFF86FE6A172AFE0DCCCC" httpUri="http://treatment.plazi.org/id/0387F954292CFF86FE6A172AFE0DCCCC" lastPageId="10" lastPageNumber="29" pageId="6" pageNumber="25">
<subSubSection id="C3341BC9292CFF8AFE6A172AFCA7CACA" pageId="6" pageNumber="25" type="distribution">
<subSubSection id="C3341BC9292CFF8AFE6A172AFB5BCA85" box="[439,1148,616,643]" pageId="6" pageNumber="25" type="nomenclature">
<paragraph id="8B914842292CFF8AFE6A172AFB5BCA85" blockId="6.[439,1148,616,716]" box="[439,1148,616,643]" pageId="6" pageNumber="25">
<taxonomicName id="4C2E33C1292CFF8AFE6A172AFB3CCA84" authority="V.S.Dang, Vuong &amp; Quan" authorityName="V. S. Dang, Vuong &amp; Quan" authorityYear="2022" box="[439,1051,616,643]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="25" phylum="Tracheophyta" rank="species" species="sonlangensis" status="sp. nov.">
<emphasis id="B95A9450292CFF8AFE6A172AFDC5CA85" bold="true" box="[439,738,616,643]" italics="true" pageId="6" pageNumber="25">Lasianthus sonlangensis</emphasis>
V.S.Dang, Vuong &amp; Quan
</taxonomicName>
<taxonomicNameLabel id="A269292B292CFF8AFBFF172BFB5BCA85" box="[1058,1148,617,643]" pageId="6" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C3341BC9292CFF8AFD9A17CFFCA7CACA" pageId="6" pageNumber="25" type="description">
<paragraph id="8B914842292CFF8AFD9A17CFFCCACAAE" blockId="6.[439,1148,616,716]" box="[583,1005,653,680]" pageId="6" pageNumber="25">urn:lsid:ipni.org:names:77295608-1</paragraph>
<paragraph id="8B914842292CFF8AFD6917F0FCA7CACA" blockId="6.[439,1148,616,716]" box="[692,896,689,716]" pageId="6" pageNumber="25">
<figureCitation id="131554C7292CFF8AFD6917F0FC3ACACA" box="[692,797,689,716]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="7.[189,232,1845,1871]" captionStartId-1="8.[189,232,1892,1918]" captionTargetBox-0="[189,1399,265,1806]" captionTargetBox-1="[232,1346,282,1842]" captionTargetId-0="figure-11@7.[189,1399,265,1806]" captionTargetId-1="figure-16@8.[189,1398,262,1857]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Fig. 4. Lasianthus sonlangensis V.S.Dang, Vuong &amp; Quan sp. nov. A. Branch. B. Stipule and flower bud. C. Abaxial leafs surfaces. DF. Close-up of infructescence. GJ. Close-up of flowers. Photos by Ba-Vuong Truong on 24 Apr. 2021." captionText-1="Fig. 5. Lasianthus sonlangensis V.S.Dang, Vuong &amp; Quan sp. nov. A. Branch. B. Abaxial and adaxial leafs surfaces. C. Stipule and flower bud. D. Infructescence. E. Flower view from above. F. Flower in side view. G. Flower in vertical section. H. Stigma. I. Stigma apical portion. Line drawings from the holotype (Dang 465) by Ba-Vuong Truong." figureDoi-0="http://doi.org/10.5281/zenodo.6373953" figureDoi-1="http://doi.org/10.5281/zenodo.6373955" httpUri-0="https://zenodo.org/record/6373953/files/figure.png" httpUri-1="https://zenodo.org/record/6373955/files/figure.png" pageId="6" pageNumber="25">Figs 45</figureCitation>
;
<tableCitation id="C6AC7DF9292CFF8AFCF417F0FCA7CACA" box="[809,896,690,716]" captionStart="Table 2" captionStartId="10.[189,258,265,291]" captionTargetPageId="10" captionText="Table 2. Comparison of Lasianthus sonlangensis V.S.Dang, Vuong &amp; Quan sp. nov. and L. caeruleus Pitard. * = from Pitard 1924; ** = from Dang 411, VNM." httpUri="http://table.plazi.org/id/DF5118CA2920FF86FF60144BFC46C941" pageId="6" pageNumber="25" tableUuid="DF5118CA2920FF86FF60144BFC46C941">Table 2</tableCitation>
</paragraph>
</subSubSection>
</subSubSection>
<subSubSection id="C3341BC9292CFF8AFF6017B6FBBACBD4" pageId="6" pageNumber="25" type="diagnosis">
<paragraph id="8B914842292CFF8AFF6017B6FE11CB08" blockId="6.[189,310,756,782]" box="[189,310,756,782]" pageId="6" pageNumber="25">
<heading id="D0D9FF2E292CFF8AFF6017B6FE11CB08" bold="true" box="[189,310,756,782]" fontSize="11" level="3" pageId="6" pageNumber="25" reason="3">
<emphasis id="B95A9450292CFF8AFF6017B6FE11CB08" bold="true" box="[189,310,756,782]" pageId="6" pageNumber="25">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B914842292CFF8AFF601667FBBACBD4" blockId="6.[189,1399,805,978]" pageId="6" pageNumber="25">
<taxonomicName id="4C2E33C1292CFF8AFF601667FEC3CB39" authority="Dang &amp; Nguyen &amp; Hoang &amp; Nguyen &amp; Truong &amp; Dang, 2022" authorityName="Dang &amp; Nguyen &amp; Hoang &amp; Nguyen &amp; Truong &amp; Dang" authorityYear="2022" box="[189,484,805,831]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="25" phylum="Tracheophyta" rank="species" species="sonlangensis" status="sp. nov.">
<emphasis id="B95A9450292CFF8AFF601667FEC3CB39" box="[189,484,805,831]" italics="true" pageId="6" pageNumber="25">Lasianthus sonlangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269292B292CFF8AFE301667FD6CCB39" box="[493,587,805,831]" pageId="6" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
is morphologically similar to
<taxonomicName id="4C2E33C1292CFF8AFC631667FB84CB39" authority="Pitard" authorityName="Pitard" box="[958,1187,805,831]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="25" phylum="Tracheophyta" rank="species" species="caeruleus">
<emphasis id="B95A9450292CFF8AFC631667FB75CB39" box="[958,1106,805,831]" italics="true" pageId="6" pageNumber="25">L. caeruleus</emphasis>
Pitard
</taxonomicName>
(
<figureCitation id="131554C7292CFF8AFB681667FBD9CB46" box="[1205,1278,805,832]" captionStart="Fig" captionStartId="9.[189,233,1858,1884]" captionTargetBox="[189,1398,265,1820]" captionTargetId="figure-11@9.[189,1398,265,1820]" captionTargetPageId="9" captionText="Fig. 6. Lasianthus caeruleus Pitard. A. Flowering branchs. B. Stipule. C. Abaxial leaf surfaces. DE. Close-up of calyces, flower buds and young fruits. F. Close-up of inflorescences and flowers. Photos by Van-Son Dang on 26 Jul. 2019." figureDoi="http://doi.org/10.5281/zenodo.6373957" httpUri="https://zenodo.org/record/6373957/files/figure.png" pageId="6" pageNumber="25">Fig. 6</figureCitation>
) which is endemic to
<collectingCountry id="F33908D2292CFF8AFE91160BFE96CB65" box="[332,433,841,867]" name="Vietnam" pageId="6" pageNumber="25">Vietnam</collectingCountry>
in the shape of leaves and inflorescences, but differs from the latter by having secondary veins 57 pairs (vs 45 pairs), petioles
<quantity id="4CD6E5A7292CFF8AFCD3162CFC57CB8F" box="[782,880,878,905]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="6" pageNumber="25" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
long (vs
<quantity id="4CD6E5A7292CFF8AFC3D162CFB64CB8F" box="[992,1091,878,905]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="6" pageNumber="25" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
long), bracts present
<quantity id="4CD6E5A7292CFF8AFA9E162CFE23CBAB" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="6" pageNumber="25" unit="mm" value="0.8" valueMax="1.0" valueMin="0.6">0.6 1 mm</quantity>
long (vs absent), calyx tube
<quantity id="4CD6E5A7292CFF8AFDB016D1FDCFCBAB" box="[621,744,915,942]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="6" pageNumber="25" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.52 mm</quantity>
long (vs ca
<quantity id="4CD6E5A7292CFF8AFC5816D1FCEBCBAB" box="[901,972,915,941]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="25" unit="mm" value="1.0">1 mm</quantity>
long), calyx lobes
<quantity id="4CD6E5A7292CFF8AFB6616D1FA11CBAB" box="[1211,1334,915,941]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="6" pageNumber="25" unit="mm" value="0.85" valueMax="1.0" valueMin="0.7">0.71 mm</quantity>
long (vs ca
<quantity id="4CD6E5A7292CFF8AFED116F5FE41CBD4" box="[268,358,951,978]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="25" unit="mm" value="0.5">0.5 mm</quantity>
long) and corolla lobes 4 (vs 5). For a further comparison see
<tableCitation id="C6AC7DF9292CFF8AFB9D16F5FBBDCBD4" box="[1088,1178,951,978]" captionStart="Table 2" captionStartId="10.[189,258,265,291]" captionTargetPageId="10" captionText="Table 2. Comparison of Lasianthus sonlangensis V.S.Dang, Vuong &amp; Quan sp. nov. and L. caeruleus Pitard. * = from Pitard 1924; ** = from Dang 411, VNM." httpUri="http://table.plazi.org/id/DF5118CA2920FF86FF60144BFC46C941" pageId="6" pageNumber="25" tableUuid="DF5118CA2920FF86FF60144BFC46C941">Table 2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3341BC9292CFF8AFF6016B8FA10CC42" box="[189,1335,1018,1095]" pageId="6" pageNumber="25" type="etymology">
<paragraph id="8B914842292CFF8AFF6016B8FE63CC12" blockId="6.[189,324,1018,1044]" box="[189,324,1018,1044]" pageId="6" pageNumber="25">
<heading id="D0D9FF2E292CFF8AFF6016B8FE63CC12" bold="true" box="[189,324,1018,1044]" fontSize="11" level="3" pageId="6" pageNumber="25" reason="3">
<emphasis id="B95A9450292CFF8AFF6016B8FE63CC12" bold="true" box="[189,324,1018,1044]" pageId="6" pageNumber="25">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B914842292CFF8AFF601168FA10CC42" blockId="6.[189,1335,1066,1095]" box="[189,1335,1066,1095]" pageId="6" pageNumber="25">The specific epithet is derived from commune ʻSƠn Langʼ where the new species was discovered.</paragraph>
</subSubSection>
<subSubSection id="C3341BC9292CFF8AFF60112FFEEFCCBE" pageId="6" pageNumber="25" type="vernacular_names">
<paragraph id="8B914842292CFF8AFF60112FFEB3CC81" blockId="6.[189,404,1133,1159]" box="[189,404,1133,1159]" pageId="6" pageNumber="25">
<heading id="D0D9FF2E292CFF8AFF60112FFEB3CC81" bold="true" box="[189,404,1133,1159]" fontSize="11" level="3" pageId="6" pageNumber="25" reason="3">
<emphasis id="B95A9450292CFF8AFF60112FFEB3CC81" bold="true" box="[189,404,1133,1159]" pageId="6" pageNumber="25">Vernacular name</emphasis>
</heading>
</paragraph>
<paragraph id="8B914842292CFF8AFF6011DFFEEFCCBE" blockId="6.[189,456,1181,1210]" box="[189,456,1181,1210]" pageId="6" pageNumber="25">Xú hƯƠng ʻSƠn Langʼ.</paragraph>
</subSubSection>
<subSubSection id="C3341BC9292CFF8AFF6011A2FE72CD9E" pageId="6" pageNumber="25" type="materials_examined">
<paragraph id="8B914842292CFF8AFF6011A2FE4BCCFC" blockId="6.[189,364,1248,1274]" box="[189,364,1248,1274]" pageId="6" pageNumber="25">
<heading id="D0D9FF2E292CFF8AFF6011A2FE4BCCFC" bold="true" box="[189,364,1248,1274]" fontSize="11" level="3" pageId="6" pageNumber="25" reason="3">
<emphasis id="B95A9450292CFF8AFF6011A2FE4BCCFC" bold="true" box="[189,364,1248,1274]" pageId="6" pageNumber="25">
<typeStatus id="5495F6E0292CFF8AFF6011A2FFDDCCFC" box="[189,250,1248,1274]" pageId="6" pageNumber="25">Type</typeStatus>
material
</emphasis>
</heading>
</paragraph>
<paragraph id="8B914842292CFF8AFF601052FE72CD9E" blockId="6.[189,1399,1296,1432]" pageId="6" pageNumber="25">
<materialsCitation id="3B46421F292CFF8AFF601052FA20CD72" ID-GBIF-Occurrence="3714053306" collectedFrom="primary evergreen forests" collectingDate="2021-04-24" collectionCode="VNM" collectorName="Van-Son Dang &amp; Ba Vuong Truong &amp; Minh Quan Dang &amp; Nghia Son Hoang Dang" country="Vietnam" county="K'Bang District" elevation="1000" latitude="14.487522" location="Kon Chu Rang Nature Reserve" longLatPrecision="1" longitude="108.536415" municipality="Son Lang Commune" pageId="6" pageNumber="25" specimenCount="1" stateProvince="Gia Lai Province" typeStatus="holotype">
<collectingCountry id="F33908D2292CFF8AFF601052FE63CD2C" box="[189,324,1296,1322]" name="Vietnam" pageId="6" pageNumber="25">VIETNAM</collectingCountry>
<collectingRegion id="49EA86A0292CFF8AFE801052FD0ECD2C" box="[349,553,1296,1322]" country="Vietnam" name="Gia Lai" pageId="6" pageNumber="25">Gia Lai Province</collectingRegion>
,
<collectingCounty id="62F030CE292CFF8AFDE81052FDD3CD2C" box="[565,756,1296,1322]" pageId="6" pageNumber="25">KBang District</collectingCounty>
,
<collectingMunicipality id="6BF5D238292CFF8AFCDD1052FCD1CD2C" box="[768,1014,1296,1323]" pageId="6" pageNumber="25">Son Lang Commune</collectingMunicipality>
,
<location id="8EF11E99292CFF8AFBDF1053FA55CD2D" LSID="urn:lsid:plazi:treatment:0387F954292CFF86FE6A172AFE0DCCCC:8EF11E99292CFF8AFBDF1053FA55CD2D" box="[1026,1394,1296,1323]" country="Vietnam" county="K'Bang District" latitude="14.487522" longLatPrecision="1" longitude="108.536415" municipality="Son Lang Commune" name="Kon Chu Rang Nature Reserve" pageId="6" pageNumber="25" stateProvince="Gia Lai Province">Kon Chu Rang Nature Reserve</location>
,
<collectedFrom id="38320BBF292CFF8AFF601077FECFCD49" box="[189,488,1333,1360]" pageId="6" pageNumber="25">primary evergreen forests</collectedFrom>
;
<geoCoordinate id="EE1A2E85292CFF8AFE2F1071FD8ECD56" box="[498,681,1331,1360]" degrees="14" direction="north" minutes="29" orientation="latitude" pageId="6" pageNumber="25" precision="1" seconds="15.08" value="14.487522">14°2915.08″ N</geoCoordinate>
,
<geoCoordinate id="EE1A2E85292CFF8AFD6E1071FC52CD56" box="[691,885,1331,1360]" degrees="108" direction="east" minutes="32" orientation="longitude" pageId="6" pageNumber="25" precision="1" seconds="11.10" value="108.536415">108°3211.10″ E</geoCoordinate>
; at
<elevation id="0003AF71292CFF8AFC471077FB4FCD49" box="[922,1128,1333,1360]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="6" pageNumber="25" unit="m" value="1000.0" valueMax="1100.0" valueMin="900.0">
<quantity id="4CD6E5A7292CFF8AFC471077FB0DCD56" box="[922,1066,1333,1360]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="6" pageNumber="25" unit="m" value="1000.0" valueMax="1100.0" valueMin="900.0">9001100 m</quantity>
a.s.l.
</elevation>
;
<date id="FF906E82292CFF8AFBA91077FA2BCD49" box="[1140,1292,1333,1359]" pageId="6" pageNumber="25" value="2021-04-24">
<collectingDate id="EFD4976A292CFF8AFBA91077FA2BCD49" box="[1140,1292,1333,1359]" pageId="6" pageNumber="25" value="2021-04-24">24 Apr. 2021</collectingDate>
</date>
;
<emphasis id="B95A9450292CFF8AFAC81077FB0BCD72" italics="true" pageId="6" pageNumber="25">
<collectorName id="26DB2D94292CFF8AFAC81077FE26CD75" pageId="6" pageNumber="25">Van-Son Dang</collectorName>
,
<collectorName id="26DB2D94292CFF8AFED01018FEF9CD75" box="[269,478,1370,1395]" pageId="6" pageNumber="25">Ba Vuong Truong</collectorName>
,
<collectorName id="26DB2D94292CFF8AFE371018FD9FCD75" box="[490,696,1370,1396]" pageId="6" pageNumber="25">Minh Quan Dang</collectorName>
&amp;
<collectorName id="26DB2D94292CFF8AFD031018FCDFCD75" box="[734,1016,1370,1396]" pageId="6" pageNumber="25">Nghia Son Hoang Dang</collectorName>
465
</emphasis>
;
<typeStatus id="5495F6E0292CFF8AFBE61018FB81CD72" box="[1083,1190,1370,1396]" pageId="6" pageNumber="25" type="holotype">holotype</typeStatus>
:
<collectionCode id="ED3FD087292CFF8AFB6C1018FBD8CD72" box="[1201,1279,1370,1396]" country="Viet Nam" httpUri="http://grbio.org/cool/tj07-dt2h" name="Institute of Tropical Biology" pageId="6" pageNumber="25" type="Herbarium">VNM</collectionCode>
!;
</materialsCitation>
<materialsCitation id="3B46421F292CFF8AFAD31018FE6ACD9E" ID-GBIF-Occurrence="3714053302" collectedFrom="primary evergreen forests" collectingDate="2021-04-24" collectionCode="HN, VNM" collectorName="Van-Son Dang &amp; Ba Vuong Truong &amp; Minh Quan Dang &amp; Nghia Son Hoang Dang" country="Vietnam" county="K'Bang District" elevation="1000" latitude="14.487522" location="Kon Chu Rang Nature Reserve" longLatPrecision="1" longitude="108.536415" municipality="Son Lang Commune" pageId="6" pageNumber="25" specimenCount="1" stateProvince="Gia Lai Province" typeStatus="isotype">
<typeStatus id="5495F6E0292CFF8AFAD31018FA54CD72" box="[1294,1395,1370,1396]" pageId="6" pageNumber="25" type="isotype">isotypes</typeStatus>
:
<collectionCode id="ED3FD087292CFF8AFF60103CFFD7CD9E" box="[189,240,1406,1432]" pageId="6" pageNumber="25">HN</collectionCode>
!,
<collectionCode id="ED3FD087292CFF8AFF22103CFE6ACD9E" box="[255,333,1406,1432]" pageId="6" pageNumber="25">VNM</collectionCode>
</materialsCitation>
!.
</paragraph>
</subSubSection>
<subSubSection id="C3341BC9292CFF8AFF601083FE93CFEF" pageId="6" pageNumber="25" type="description">
<paragraph id="8B914842292CFF8AFF601083FE69CDDD" blockId="6.[189,334,1473,1499]" box="[189,334,1473,1499]" pageId="6" pageNumber="25">
<heading id="D0D9FF2E292CFF8AFF601083FE69CDDD" bold="true" box="[189,334,1473,1499]" fontSize="11" level="3" pageId="6" pageNumber="25" reason="3">
<emphasis id="B95A9450292CFF8AFF601083FE69CDDD" bold="true" box="[189,334,1473,1499]" pageId="6" pageNumber="25">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B914842292CFF8AFF6010B3FE93CFEF" blockId="6.[188,1400,1521,2025]" pageId="6" pageNumber="25">
Shrubs, evergreen, up to
<quantity id="4CD6E5A7292CFF8AFE0110B3FD2FCE0A" box="[476,520,1521,1548]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="25" unit="m" value="1.0">1 m</quantity>
tall; branches and branchlets slender,
<quantity id="4CD6E5A7292CFF8AFC6210B3FB12CE0A" box="[959,1077,1521,1548]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="6" pageNumber="25" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">11.5 mm</quantity>
diam., glabrous, internodes
<quantity id="4CD6E5A7292CFF8AFF611354FE63CE37" box="[188,324,1558,1585]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="6" pageNumber="25" unit="cm" value="2.0" valueMax="2.5" valueMin="1.5">1.52.5 cm</quantity>
long. Leaves opposite, blades oblong to oblong-laceolate, 57 ×
<quantity id="4CD6E5A7292CFF8AFBB71354FBF8CE37" box="[1130,1247,1558,1585]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="6" pageNumber="25" unit="cm" value="1.25" valueMax="1.5" valueMin="1.0">11.5 cm</quantity>
, coriaceous, yellowish brown when dry, glabrous on both sides, apex acute to acuminate, aristate
<quantity id="4CD6E5A7292CFF8AFB661379FA15CE53" box="[1211,1330,1595,1621]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="6" pageNumber="25" unit="mm" value="0.85" valueMax="1.0" valueMin="0.7">0.71 mm</quantity>
long, glabrous, base acute, margin entire; midrib slightly prominent adaxially, prominent abaxially; secondary veins 57 pairs, ascending at an angle of 6075° from midrib, curved to the margin, flat adaxially, slighly prominent abaxially; tertiary veins subparallel to subreticulate; petioles
<quantity id="4CD6E5A7292CFF8AFC2013EBFB7ACEC5" box="[1021,1117,1705,1731]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="6" pageNumber="25" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
long, grabrous. Stipules triangular,
<quantity id="4CD6E5A7292CFF8AFEE0138CFE90CEEE" box="[317,439,1742,1768]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="6" pageNumber="25" unit="mm" value="0.9" valueMax="1.0" valueMin="0.8">0.81 mm</quantity>
long, persistent, glabrous except barbate margin. Inflorescences cymose, sessile, 19-flowered; bracts small,
<quantity id="4CD6E5A7292CFF8AFDCF13B0FDABCF0B" box="[530,652,1778,1805]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="6" pageNumber="25" unit="mm" value="0.8" valueMax="1.0" valueMin="0.6">0.61 mm</quantity>
long, glabrous except barbate margin. Flowers sessile; calyx campanulate,
<quantity id="4CD6E5A7292CFF8AFEBC1255FEE3CF37" box="[353,452,1815,1841]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="6" pageNumber="25" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
long, white to whitish purple, calyx tube
<quantity id="4CD6E5A7292CFF8AFC6A1255FB08CF37" box="[951,1071,1815,1842]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="6" pageNumber="25" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.52 mm</quantity>
long, glabrous, calyx lobes 4, linear-triangular,
<quantity id="4CD6E5A7292CFF8AFE771279FD05CF50" box="[426,546,1851,1878]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="6" pageNumber="25" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.51 mm</quantity>
long, glabrous except barbate margin; corolla tube cylindrical,
<quantity id="4CD6E5A7292CFF8AFACB127EFA5FCF50" box="[1302,1400,1852,1878]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="6" pageNumber="25" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">89 mm</quantity>
long, white, corolla tube
<quantity id="4CD6E5A7292CFF8AFE321222FD74CF7D" box="[495,595,1888,1915]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="6" pageNumber="25" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">67 mm</quantity>
long, puberulous outside in upper part, corolla lobes 4, triangular,
<quantity id="4CD6E5A7292CFF8AFF6212C7FE10CF99" box="[191,311,1925,1952]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="6" pageNumber="25" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.52 mm</quantity>
long, puberulous outside, floccose inside in the throat; stamens 4, filaments short, anthers oblong,
<quantity id="4CD6E5A7292CFF8AFEC012E8FEB3CFC2" box="[285,404,1962,1988]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="6" pageNumber="25" unit="mm" value="1.1" valueMax="1.2" valueMin="1.0">11.2 mm</quantity>
long; ovary with 45 locules, style linear,
<quantity id="4CD6E5A7292CFF8AFC5A12E8FCCECFC2" box="[903,1001,1962,1988]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="6" pageNumber="25" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">78 mm</quantity>
long, stigma lobes 4,
<quantity id="4CD6E5A7292CFF8AFB3712E8FA50CFC2" box="[1258,1399,1962,1988]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="6" pageNumber="25" unit="mm" value="0.55" valueMax="0.7" valueMin="0.4">0.40.7 mm</quantity>
long. Fruits not seen.
</paragraph>
</subSubSection>
<caption id="DF5118CA292DFF8BFF601277FD79CF93" ID-DOI="http://doi.org/10.5281/zenodo.6373953" ID-Zenodo-Dep="6373953" httpUri="https://zenodo.org/record/6373953/files/figure.png" pageId="7" pageNumber="26" startId="7.[189,232,1845,1871]" targetBox="[189,1399,265,1806]" targetPageId="7">
<paragraph id="8B914842292DFF8BFF601277FD79CF93" blockId="7.[189,1399,1845,1942]" pageId="7" pageNumber="26">
<emphasis id="B95A9450292DFF8BFF601277FE2ACF49" bold="true" box="[189,269,1845,1871]" pageId="7" pageNumber="26">Fig. 4.</emphasis>
<taxonomicName id="4C2E33C1292DFF8BFECB1277FC5ACF49" authority="V.S.Dang, Vuong &amp; Quan" authorityName="V. S. Dang, Vuong &amp; Quan" authorityYear="2022" box="[278,893,1845,1871]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="7" pageNumber="26" phylum="Tracheophyta" rank="species" species="sonlangensis" status="sp. nov.">
<emphasis id="B95A9450292DFF8BFECB1277FD1DCF49" box="[278,570,1845,1871]" italics="true" pageId="7" pageNumber="26">Lasianthus sonlangensis</emphasis>
V.S.Dang, Vuong &amp; Quan
</taxonomicName>
<taxonomicNameLabel id="A269292B292DFF8BFC5A1277FCC3CF49" box="[903,996,1845,1871]" pageId="7" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B95A9450292DFF8BFC311277FB27CF49" bold="true" box="[1004,1024,1845,1871]" pageId="7" pageNumber="26">A</emphasis>
. Branch.
<emphasis id="B95A9450292DFF8BFBA51277FBADCF49" bold="true" box="[1144,1162,1845,1871]" pageId="7" pageNumber="26">B</emphasis>
. Stipule and flower bud.
<emphasis id="B95A9450292DFF8BFF24121AFE29CF74" bold="true" box="[249,270,1880,1906]" pageId="7" pageNumber="26">C</emphasis>
. Abaxial leafs surfaces.
<emphasis id="B95A9450292DFF8BFDEE121AFD40CF74" bold="true" box="[563,615,1880,1906]" pageId="7" pageNumber="26">DF</emphasis>
. Close-up of infructescence.
<emphasis id="B95A9450292DFF8BFC18121AFCD0CF74" bold="true" box="[965,1015,1880,1906]" pageId="7" pageNumber="26">GJ</emphasis>
. Close-up of flowers. Photos by Ba-Vuong Truong on 24 Apr. 2021.
</paragraph>
</caption>
<caption id="DF5118CA2922FF84FF601226FD9CCFEE" ID-DOI="http://doi.org/10.5281/zenodo.6373955" ID-Zenodo-Dep="6373955" httpUri="https://zenodo.org/record/6373955/files/figure.png" pageId="8" pageNumber="27" startId="8.[189,232,1892,1918]" targetBox="[232,1346,282,1842]" targetPageId="8">
<paragraph id="8B9148422922FF84FF601226FD9CCFEE" blockId="8.[189,1399,1892,2024]" pageId="8" pageNumber="27">
<emphasis id="B95A94502922FF84FF601226FE2BCF78" bold="true" box="[189,268,1892,1918]" pageId="8" pageNumber="27">Fig. 5.</emphasis>
<taxonomicName id="4C2E33C12922FF84FEC91226FC53CF78" authority="V.S.Dang, Vuong &amp; Quan" authorityName="V. S. Dang, Vuong &amp; Quan" authorityYear="2022" box="[276,884,1892,1919]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="8" pageNumber="27" phylum="Tracheophyta" rank="species" species="sonlangensis" status="sp. nov.">
<emphasis id="B95A94502922FF84FEC91226FD12CF78" box="[276,565,1892,1918]" italics="true" pageId="8" pageNumber="27">Lasianthus sonlangensis</emphasis>
V.S.Dang, Vuong &amp; Quan
</taxonomicName>
<taxonomicNameLabel id="A269292B2922FF84FCA11227FCF0CF79" box="[892,983,1893,1919]" pageId="8" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B95A94502922FF84FC021226FCD4CF78" bold="true" box="[991,1011,1892,1918]" pageId="8" pageNumber="27">A</emphasis>
. Branch.
<emphasis id="B95A94502922FF84FBB41226FB5CCF78" bold="true" box="[1129,1147,1892,1918]" pageId="8" pageNumber="27">B</emphasis>
. Abaxial and adaxial leafs surfaces.
<emphasis id="B95A94502922FF84FEB012C5FEA6CFA7" bold="true" box="[365,385,1927,1953]" pageId="8" pageNumber="27">C</emphasis>
. Stipule and flower bud.
<emphasis id="B95A94502922FF84FD7012C5FDE5CFA7" bold="true" box="[685,706,1927,1953]" pageId="8" pageNumber="27">D</emphasis>
. Infructescence.
<emphasis id="B95A94502922FF84FC5012C5FC87CFA7" bold="true" box="[909,928,1927,1953]" pageId="8" pageNumber="27">E</emphasis>
. Flower view from above.
<emphasis id="B95A94502922FF84FB3E12C5FBD2CFA7" bold="true" box="[1251,1269,1927,1953]" pageId="8" pageNumber="27">F</emphasis>
. Flower in side view.
<emphasis id="B95A94502922FF84FE9D12E8FE72CFC2" bold="true" box="[320,341,1962,1988]" pageId="8" pageNumber="27">G</emphasis>
. Flower in vertical section.
<emphasis id="B95A94502922FF84FD7412E9FDE7CFC3" bold="true" box="[681,704,1963,1989]" pageId="8" pageNumber="27">H</emphasis>
. Stigma.
<emphasis id="B95A94502922FF84FCEA12E9FC66CFC3" bold="true" box="[823,833,1963,1989]" pageId="8" pageNumber="27">I</emphasis>
. Stigma apical portion. Line drawings from the holotype (
<emphasis id="B95A94502922FF84FEE8128CFE8ECFE1" box="[309,425,1997,2023]" italics="true" pageId="8" pageNumber="27">Dang 465</emphasis>
) by Ba-Vuong Truong.
</paragraph>
</caption>
<caption id="DF5118CA2923FF85FF601200FD8FCFA5" ID-DOI="http://doi.org/10.5281/zenodo.6373957" ID-Zenodo-Dep="6373957" httpUri="https://zenodo.org/record/6373957/files/figure.png" pageId="9" pageNumber="28" startId="9.[189,233,1858,1884]" targetBox="[189,1398,265,1820]" targetPageId="9">
<paragraph id="8B9148422923FF85FF601200FD8FCFA5" blockId="9.[189,1399,1858,1955]" pageId="9" pageNumber="28">
<emphasis id="B95A94502923FF85FF601200FE36CF5A" bold="true" box="[189,273,1858,1884]" pageId="9" pageNumber="28">Fig. 6.</emphasis>
<taxonomicName id="4C2E33C12923FF85FEC01200FC16CF5A" authority="Pitard. A. Flowering" box="[285,817,1858,1885]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="9" pageNumber="28" phylum="Tracheophyta" rank="species" species="caeruleus">
<emphasis id="B95A94502923FF85FEC01200FD03CF5A" box="[285,548,1858,1884]" italics="true" pageId="9" pageNumber="28">Lasianthus caeruleus</emphasis>
Pitard.
<emphasis id="B95A94502923FF85FD561200FD87CF5A" bold="true" box="[651,672,1858,1884]" pageId="9" pageNumber="28">A</emphasis>
. Flowering
</taxonomicName>
branchs.
<emphasis id="B95A94502923FF85FC721200FCE4CF5A" bold="true" box="[943,963,1858,1884]" pageId="9" pageNumber="28">B</emphasis>
. Stipule.
<emphasis id="B95A94502923FF85FBE21200FB73CF5A" bold="true" box="[1087,1108,1858,1884]" pageId="9" pageNumber="28">C</emphasis>
. Abaxial leaf surfaces.
<emphasis id="B95A94502923FF85FF601227FFD1CF79" bold="true" box="[189,246,1893,1919]" pageId="9" pageNumber="28">DE</emphasis>
. Close-up of calyces, flower buds and young fruits.
<emphasis id="B95A94502923FF85FCAB1227FCAFCF79" bold="true" box="[886,904,1893,1919]" pageId="9" pageNumber="28">F</emphasis>
. Close-up of inflorescences and flowers. Photos by Van-Son Dang on 26 Jul. 2019.
</paragraph>
</caption>
<caption id="DF5118CA2920FF86FF60144BFC46C941" ID-Table-UUID="DF5118CA2920FF86FF60144BFC46C941" httpUri="http://table.plazi.org/id/DF5118CA2920FF86FF60144BFC46C941" pageId="10" pageNumber="29" startId="10.[189,258,265,291]" targetBox="[197,1256,366,712]" targetIsTable="true" targetPageId="10">
<paragraph id="8B9148422920FF86FF60144BFC46C941" blockId="10.[189,1398,265,327]" pageId="10" pageNumber="29">
<emphasis id="B95A94502920FF86FF60144BFE06C925" bold="true" box="[189,289,265,291]" pageId="10" pageNumber="29">Table 2.</emphasis>
Comparison of
<taxonomicName id="4C2E33C12920FF86FE38144BFB64C925" authority="V.S.Dang, Vuong &amp; Quan" authorityName="V. S. Dang, Vuong &amp; Quan" authorityYear="2022" box="[485,1091,265,292]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="10" pageNumber="29" phylum="Tracheophyta" rank="species" species="sonlangensis" status="sp. nov.">
<emphasis id="B95A94502920FF86FE38144BFC20C925" box="[485,775,265,291]" italics="true" pageId="10" pageNumber="29">Lasianthus sonlangensis</emphasis>
V.S.Dang, Vuong &amp; Quan
</taxonomicName>
<taxonomicNameLabel id="A269292B2920FF86FB911448FB8FC922" box="[1100,1192,266,292]" pageId="10" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C2E33C12920FF86FB381448FE2CC940" authority="Pitard." authorityName="Pitard." class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="10" pageNumber="29" phylum="Tracheophyta" rank="species" species="caeruleus">
<emphasis id="B95A94502920FF86FB381448FA51C925" box="[1253,1398,265,291]" italics="true" pageId="10" pageNumber="29">L. caeruleus</emphasis>
Pitard.
</taxonomicName>
* = from
<bibRefCitation id="EFBF35B32920FF86FE5D146EFD2CC940" author="Pitard J." box="[384,523,300,326]" pageId="10" pageNumber="29" pagination="395" refId="ref5138" refString="Pitard J. 1924. Rubiacees. In: Lecomte P. H., Humbert H. &amp; Gagnepain F. (eds) Flore generale de l'Indo- Chine 3: 395. Masson, Paris." type="journal article" year="1924">Pitard 1924</bibRefCitation>
; ** = from
<emphasis id="B95A94502920FF86FD49146FFC20C940" box="[660,775,300,326]" italics="true" pageId="10" pageNumber="29">Dang 411</emphasis>
, VNM.
</paragraph>
</caption>
<paragraph id="8B9148422920FF86FF18142CFBCFCAC1" pageId="10" pageNumber="29">
<table id="F92EBAE229200073FF18142CFBCFCACE" box="[197,1256,366,712]" gridcols="3" gridrows="10" pageId="10" pageNumber="29">
<tr id="351E4A0029200073FF18142CFBCFC98F" box="[197,1256,366,393]" gridrow="0" pageId="10" pageNumber="29">
<th id="76CF237C29200073FF18142CFEB7C98F" box="[197,400,366,393]" gridcol="0" gridrow="0" pageId="10" pageNumber="29">
<emphasis id="B95A94502920FF86FF18142CFE76C98E" bold="true" box="[197,337,366,392]" pageId="10" pageNumber="29">Characters</emphasis>
</th>
<th id="76CF237C29200073FE38142CFC52C98F" box="[485,885,366,393]" gridcol="1" gridrow="0" pageId="10" pageNumber="29">
<emphasis id="B95A94502920FF86FE38142CFC52C98E" bold="true" box="[485,885,366,393]" pageId="10" pageNumber="29">
<taxonomicName id="4C2E33C12920FF86FE38142CFC28C98F" authority="Dang &amp; Nguyen &amp; Hoang &amp; Nguyen &amp; Truong &amp; Dang, 2022" authorityName="Dang &amp; Nguyen &amp; Hoang &amp; Nguyen &amp; Truong &amp; Dang" authorityYear="2022" box="[485,783,366,393]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="10" pageNumber="29" phylum="Tracheophyta" rank="species" species="sonlangensis" status="sp. nov.">
<emphasis id="B95A94502920FF86FE38142CFC28C98F" bold="true" box="[485,783,366,393]" italics="true" pageId="10" pageNumber="29">Lasianthus sonlangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269292B2920FF86FCCA142CFC52C98E" box="[791,885,366,392]" pageId="10" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="76CF237C29200073FC70142CFBCFC98F" box="[941,1256,366,393]" gridcol="2" gridrow="0" pageId="10" pageNumber="29">
<emphasis id="B95A94502920FF86FC70142CFB97C98F" bold="true" box="[941,1200,366,393]" italics="true" pageId="10" pageNumber="29">Lasianthus caeruleus</emphasis>
</th>
</tr>
<tr id="351E4A0029200073FF1814D3FBCFC9AA" box="[197,1256,401,428]" gridrow="1" pageId="10" pageNumber="29">
<th id="76CF237C29200073FF1814D3FEB7C9AA" box="[197,400,401,428]" gridcol="0" gridrow="1" pageId="10" pageNumber="29">
<emphasis id="B95A94502920FF86FF1814D3FE72C9AD" bold="true" box="[197,341,401,427]" pageId="10" pageNumber="29">Leaf blades</emphasis>
</th>
<td id="76CF237C29200073FE3814D3FC52C9AA" box="[485,885,401,428]" gridcol="1" gridrow="1" pageId="10" pageNumber="29">57 × 11.5 cm</td>
<td id="76CF237C29200073FC7014D3FBCFC9AA" box="[941,1256,401,428]" gridcol="2" gridrow="1" pageId="10" pageNumber="29">4.57 × 0.81.5 cm *</td>
</tr>
<tr id="351E4A0029200073FF1814F6FBCFC9D6" box="[197,1256,436,464]" gridrow="2" pageId="10" pageNumber="29">
<th id="76CF237C29200073FF1814F6FEB7C9D6" box="[197,400,436,464]" gridcol="0" gridrow="2" pageId="10" pageNumber="29">
<emphasis id="B95A94502920FF86FF1814F6FEB7C9C8" bold="true" box="[197,400,436,462]" pageId="10" pageNumber="29">Secondary veins</emphasis>
</th>
<td id="76CF237C29200073FE3814F6FC52C9D6" box="[485,885,436,464]" gridcol="1" gridrow="2" pageId="10" pageNumber="29">57 pairs</td>
<td id="76CF237C29200073FC7014F6FBCFC9D6" box="[941,1256,436,464]" gridcol="2" gridrow="2" pageId="10" pageNumber="29">45 pairs *</td>
</tr>
<tr id="351E4A0029200073FF18149AFBCFC9F5" box="[197,1256,472,499]" gridrow="3" pageId="10" pageNumber="29">
<th id="76CF237C29200073FF18149AFEB7C9F5" box="[197,400,472,499]" gridcol="0" gridrow="3" pageId="10" pageNumber="29">
<emphasis id="B95A94502920FF86FF18149AFE02C9F4" bold="true" box="[197,293,472,498]" pageId="10" pageNumber="29">Petioles</emphasis>
</th>
<td id="76CF237C29200073FE38149AFC52C9F5" box="[485,885,472,499]" gridcol="1" gridrow="3" pageId="10" pageNumber="29">23 mm long</td>
<td id="76CF237C29200073FC70149AFBCFC9F5" box="[941,1256,472,499]" gridcol="2" gridrow="3" pageId="10" pageNumber="29">45 mm long *</td>
</tr>
<tr id="351E4A0029200073FF1814B9FBCFCA10" box="[197,1256,507,534]" gridrow="4" pageId="10" pageNumber="29">
<th id="76CF237C29200073FF1814B9FEB7CA10" box="[197,400,507,534]" gridcol="0" gridrow="4" pageId="10" pageNumber="29">
<emphasis id="B95A94502920FF86FF1814B9FE0FCA13" bold="true" box="[197,296,507,533]" pageId="10" pageNumber="29">Stipules</emphasis>
</th>
<td id="76CF237C29200073FE3814B9FC52CA10" box="[485,885,507,534]" gridcol="1" gridrow="4" pageId="10" pageNumber="29">persistent, 0.81 mm long</td>
<td id="76CF237C29200073FC7014B9FBCFCA10" box="[941,1256,507,534]" gridcol="2" gridrow="4" pageId="10" pageNumber="29">often deciduous *</td>
</tr>
<tr id="351E4A0029200073FF18175DFBCFCA3C" box="[197,1256,543,570]" gridrow="5" pageId="10" pageNumber="29">
<th id="76CF237C29200073FF18175DFEB7CA3C" box="[197,400,543,570]" gridcol="0" gridrow="5" pageId="10" pageNumber="29">
<emphasis id="B95A94502920FF86FF18175DFE31CA3F" bold="true" box="[197,278,543,569]" pageId="10" pageNumber="29">Bracts</emphasis>
</th>
<td id="76CF237C29200073FE38175DFC52CA3C" box="[485,885,543,570]" gridcol="1" gridrow="5" pageId="10" pageNumber="29">0.61 mm long</td>
<td id="76CF237C29200073FC70175DFBCFCA3C" box="[941,1256,543,570]" gridcol="2" gridrow="5" pageId="10" pageNumber="29">absent*</td>
</tr>
<tr id="351E4A0029200073FF181700FBCFCA58" box="[197,1256,578,606]" gridrow="6" pageId="10" pageNumber="29">
<th id="76CF237C29200073FF181700FEB7CA58" box="[197,400,578,606]" gridcol="0" gridrow="6" pageId="10" pageNumber="29">
<emphasis id="B95A94502920FF86FF181700FE6BCA5B" bold="true" box="[197,332,578,605]" pageId="10" pageNumber="29">Calyx tube</emphasis>
</th>
<td id="76CF237C29200073FE381700FC52CA58" box="[485,885,578,606]" gridcol="1" gridrow="6" pageId="10" pageNumber="29">1.52 mm long</td>
<td id="76CF237C29200073FC701700FBCFCA58" box="[941,1256,578,606]" gridcol="2" gridrow="6" pageId="10" pageNumber="29">ca 1 mm long *</td>
</tr>
<tr id="351E4A0029200073FF181724FBCFCA87" box="[197,1256,614,641]" gridrow="7" pageId="10" pageNumber="29">
<th id="76CF237C29200073FF181724FEB7CA87" box="[197,400,614,641]" gridcol="0" gridrow="7" pageId="10" pageNumber="29">
<emphasis id="B95A94502920FF86FF181724FE72CA86" bold="true" box="[197,341,614,640]" pageId="10" pageNumber="29">Calyx lobes</emphasis>
</th>
<td id="76CF237C29200073FE381724FC52CA87" box="[485,885,614,641]" gridcol="1" gridrow="7" pageId="10" pageNumber="29">0.71 mm long, 4-lobed</td>
<td id="76CF237C29200073FC701724FBCFCA87" box="[941,1256,614,641]" gridcol="2" gridrow="7" pageId="10" pageNumber="29">ca 0.5 mm long, 5-lobed *</td>
</tr>
<tr id="351E4A0029200073FF1817CBFBCFCAA3" box="[197,1256,649,677]" gridrow="8" pageId="10" pageNumber="29">
<th id="76CF237C29200073FF1817CBFEB7CAA3" box="[197,400,649,677]" gridcol="0" gridrow="8" pageId="10" pageNumber="29">
<emphasis id="B95A94502920FF86FF1817CBFE46CAA2" bold="true" box="[197,353,649,676]" pageId="10" pageNumber="29">Corolla tube</emphasis>
</th>
<td id="76CF237C29200073FE3817CBFC52CAA3" box="[485,885,649,677]" gridcol="1" gridrow="8" pageId="10" pageNumber="29">67 mm long</td>
<td id="76CF237C29200073FC7017CBFBCFCAA3" box="[941,1256,649,677]" gridcol="2" gridrow="8" pageId="10" pageNumber="29">45 mm long **</td>
</tr>
<tr id="351E4A0029200073FF1817EFFBCFCACE" box="[197,1256,685,712]" gridrow="9" pageId="10" pageNumber="29">
<th id="76CF237C29200073FF1817EFFEB7CACE" box="[197,400,685,712]" gridcol="0" gridrow="9" pageId="10" pageNumber="29">
<emphasis id="B95A94502920FF86FF1817EFFE4DCAC1" bold="true" box="[197,362,685,711]" pageId="10" pageNumber="29">Corolla lobes</emphasis>
</th>
<td id="76CF237C29200073FE3817EFFC52CACE" box="[485,885,685,712]" gridcol="1" gridrow="9" pageId="10" pageNumber="29">1.52 mm long, 4-lobed</td>
<td id="76CF237C29200073FC7017EFFBCFCACE" box="[941,1256,685,712]" gridcol="2" gridrow="9" pageId="10" pageNumber="29">1.52 mm long, 5-lobed **</td>
</tr>
</table>
</paragraph>
<subSubSection id="C3341BC92920FF86FF601641FCCFCBF2" pageId="10" pageNumber="29" type="distribution">
<paragraph id="8B9148422920FF86FF601641FD5ACB1B" blockId="10.[189,637,771,797]" box="[189,637,771,797]" pageId="10" pageNumber="29">
<heading id="D0D9FF2E2920FF86FF601641FD5ACB1B" bold="true" box="[189,637,771,797]" fontSize="11" level="3" pageId="10" pageNumber="29" reason="3">
<emphasis id="B95A94502920FF86FF601641FD5ACB1B" bold="true" box="[189,637,771,797]" pageId="10" pageNumber="29">Distribution, habitat and phenology</emphasis>
</heading>
</paragraph>
<paragraph id="8B9148422920FF86FF601672FCCFCBF2" blockId="10.[189,1400,816,1013]" pageId="10" pageNumber="29">
<taxonomicName id="4C2E33C12920FF86FF601672FEF9CB4C" authority="Dang &amp; Nguyen &amp; Hoang &amp; Nguyen &amp; Truong &amp; Dang, 2022" authorityName="Dang &amp; Nguyen &amp; Hoang &amp; Nguyen &amp; Truong &amp; Dang" authorityYear="2022" box="[189,478,816,842]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="10" pageNumber="29" phylum="Tracheophyta" rank="species" species="sonlangensis" status="sp. nov.">
<emphasis id="B95A94502920FF86FF601672FEF9CB4C" box="[189,478,816,842]" italics="true" pageId="10" pageNumber="29">Lasianthus sonlangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269292B2920FF86FE3B1673FD66CB4D" box="[486,577,817,843]" pageId="10" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
was discovered in Kon Chu Rang Nature Reserve,
<collectingRegion id="49EA86A02920FF86FB781672FA55CB4C" box="[1189,1394,816,842]" country="Vietnam" name="Gia Lai" pageId="10" pageNumber="29">Gia Lai Province</collectingRegion>
, southern
<collectingCountry id="F33908D22920FF86FEF31610FEB0CB6A" box="[302,407,850,876]" name="Vietnam" pageId="10" pageNumber="29">Vietnam</collectingCountry>
. It occurs in the understory of primary evergreen forest together with
<taxonomicName id="4C2E33C12920FF86FB281610FEE9CB89" authority="(Roxb.) Merr." authorityName="Merr." baseAuthorityName="Roxb." class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="10" pageNumber="29" phylum="Tracheophyta" rank="species" species="hirsutus">
<emphasis id="B95A94502920FF86FB281610FE3ACB88" italics="true" pageId="10" pageNumber="29">Lasianthus hirsutus</emphasis>
(Roxb.) Merr.
</taxonomicName>
(
<taxonomicName id="4C2E33C12920FF86FE031636FD84CB89" authority="Juss." authorityName="Juss." box="[478,675,884,911]" class="Magnoliopsida" family="Rubiaceae" kingdom="Plantae" order="Gentianales" pageId="10" pageNumber="29" phylum="Tracheophyta" rank="family">Rubiaceae Juss.</taxonomicName>
),
<taxonomicName id="4C2E33C12920FF86FD611636FB77CB88" authority="E. Walker" authorityName="E. Walker" box="[700,1104,884,910]" class="Magnoliopsida" family="Primulaceae" genus="Ardisia" kingdom="Plantae" order="Ericales" pageId="10" pageNumber="29" phylum="Tracheophyta" rank="species" species="perpendicularis">
<emphasis id="B95A94502920FF86FD611636FCFDCB88" box="[700,986,884,910]" italics="true" pageId="10" pageNumber="29">Ardisia perpendicularis</emphasis>
E.Walker
</taxonomicName>
(
<taxonomicName id="4C2E33C12920FF86FBBC1636FE31CBB7" authority="Batsch ex Borkh." authorityName="Borkh." class="Magnoliopsida" family="Primulaceae" kingdom="Plantae" order="Ericales" pageId="10" pageNumber="29" phylum="Tracheophyta" rank="family">Primulaceae Batsch ex Borkh.</taxonomicName>
),
<taxonomicName id="4C2E33C12920FF86FEF116D4FD93CBB6" authority="Blume" authorityName="Blume" box="[300,692,918,944]" class="Magnoliopsida" family="Sapindaceae" genus="Mischocarpus" kingdom="Plantae" order="Sapindales" pageId="10" pageNumber="29" phylum="Tracheophyta" rank="species" species="sundaicus">
<emphasis id="B95A94502920FF86FEF116D4FD7DCBB6" box="[300,602,918,944]" italics="true" pageId="10" pageNumber="29">Mischocarpus sundaicus</emphasis>
Blume
</taxonomicName>
(
<taxonomicName id="4C2E33C12920FF86FD1B16D4FC80CBB7" authority="Juss." authorityName="Juss." box="[710,935,918,945]" class="Magnoliopsida" family="Sapindaceae" kingdom="Plantae" order="Sapindales" pageId="10" pageNumber="29" phylum="Tracheophyta" rank="family">Sapindaceae Juss.</taxonomicName>
),
<taxonomicName id="4C2E33C12920FF86FC6216D5FA51CBB6" authority="Tardieu" authorityName="Tardieu" box="[959,1398,918,944]" class="Magnoliopsida" family="Malvaceae" genus="Pterospermum" kingdom="Plantae" order="Malvales" pageId="10" pageNumber="29" phylum="Tracheophyta" rank="species" species="mucronatum">
<emphasis id="B95A94502920FF86FC6216D5FA36CBB6" box="[959,1297,919,944]" italics="true" pageId="10" pageNumber="29">Pterospermum mucronatum</emphasis>
Tardieu
</taxonomicName>
(
<taxonomicName id="4C2E33C12920FF86FF1A16FAFEA8CBD5" authority="Juss." authorityName="Juss." box="[199,399,952,979]" class="Magnoliopsida" family="Malvaceae" kingdom="Plantae" order="Malvales" pageId="10" pageNumber="29" phylum="Tracheophyta" rank="family">Malvaceae Juss.</taxonomicName>
) and
<taxonomicName id="4C2E33C12920FF86FE0816FAFD9ACBD4" authorityName="I.M.Turner" authorityYear="2009" baseAuthorityName="Hook.f. &amp; Thomson" box="[469,701,952,978]" class="Magnoliopsida" family="Annonaceae" genus="Trivalvaria" kingdom="Plantae" order="Magnoliales" pageId="10" pageNumber="29" phylum="Tracheophyta" rank="species" species="costata">
<emphasis id="B95A94502920FF86FE0816FAFD9ACBD4" box="[469,701,952,978]" italics="true" pageId="10" pageNumber="29">Trivalvaria costata</emphasis>
</taxonomicName>
(Hook.f. &amp; Thomson) I.M.Turner (
<taxonomicName id="4C2E33C12920FF86FBB016FBFA6ECBD5" authority="Juss." authorityName="Juss." box="[1133,1353,953,979]" class="Magnoliopsida" family="Annonaceae" kingdom="Plantae" order="Magnoliales" pageId="10" pageNumber="29" phylum="Tracheophyta" rank="family">Annonaceae Juss.</taxonomicName>
), at
<quantity id="4CD6E5A72920FF86FF621699FE76CBF3" box="[191,337,987,1013]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="10" pageNumber="29" unit="m" value="1000.0" valueMax="1100.0" valueMin="900.0">9001100 m</quantity>
elevation. Flowering specimens were collected in April.
</paragraph>
</subSubSection>
<subSubSection id="C3341BC92920FF86FF601158FE0DCCCC" pageId="10" pageNumber="29" type="description">
<paragraph id="8B9148422920FF86FF601158FEFCCC33" blockId="10.[189,475,1050,1077]" box="[189,475,1050,1077]" pageId="10" pageNumber="29">
<heading id="D0D9FF2E2920FF86FF601158FEFCCC33" bold="true" box="[189,475,1050,1077]" fontSize="11" level="3" pageId="10" pageNumber="29" reason="3">
<emphasis id="B95A94502920FF86FF601158FEFCCC33" bold="true" box="[189,475,1050,1077]" pageId="10" pageNumber="29">Population and threats</emphasis>
</heading>
</paragraph>
<paragraph id="8B9148422920FF86FF60110AFE0DCCCC" blockId="10.[187,1400,1096,1226]" pageId="10" pageNumber="29">
<taxonomicName id="4C2E33C12920FF86FF60110AFEC3CC64" authority="Dang &amp; Nguyen &amp; Hoang &amp; Nguyen &amp; Truong &amp; Dang, 2022" authorityName="Dang &amp; Nguyen &amp; Hoang &amp; Nguyen &amp; Truong &amp; Dang" authorityYear="2022" box="[189,484,1096,1122]" class="Magnoliopsida" family="Rubiaceae" genus="Lasianthus" kingdom="Plantae" order="Gentianales" pageId="10" pageNumber="29" phylum="Tracheophyta" rank="species" species="sonlangensis" status="sp. nov.">
<emphasis id="B95A94502920FF86FF60110AFEC3CC64" box="[189,484,1096,1122]" italics="true" pageId="10" pageNumber="29">Lasianthus sonlangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A269292B2920FF86FE30110BFD6CCC65" box="[493,587,1097,1123]" pageId="10" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
is currently known from only one locality with a small population of less than 10 individuals. The forest habitat where we found this species is frequently disturbed by anthropogenic factors, such as spontaneous tourism activities and collection of non-timber forest products (NTFPs).
</paragraph>
</subSubSection>
</treatment>
</document>