250 lines
30 KiB
XML
250 lines
30 KiB
XML
<document id="D03DCA5E0B5CDA5A89B86C68377E38F2" ID-DOI="10.1080/00222933.2017.1324050" ID-ISSN="1464-5262" ID-Zenodo-Dep="5181888" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628698566541" checkinUser="felipe" docAuthor="Schmitter-Soto, Juan J." docDate="2017" docId="03806F39C965FFD3FE58FAEED230FAE2" docLanguage="en" docName="JNatHist.51.23-24.1331-1424.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 51 (21 - 24)" docSource="http://dx.doi.org/10.1080/00222933.2017.1324050" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Astyanax cubilhuitz Schmitter-Soto 2017, sp. nov." docType="treatment" docVersion="4" lastPageNumber="1382" masterDocId="FFB91741C957FFE7FFBEFFB5D316FFA7" masterDocTitle="A revision of Astyanax (Characiformes: Characidae) in Central and North America, with the description of nine new species" masterLastPageNumber="1424" masterPageNumber="1331" pageNumber="1380" updateTime="1699050015946" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="521DB456B9B59CC2A43C114DE0CF2F2A">A revision of Astyanax (Characiformes: Characidae) in Central and North America, with the description of nine new species</mods:title>
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<mods:name id="D0CE7163B97214A9CCC2EAB9CBDF7784" type="personal">
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<mods:namePart id="5D1D48CA38A1159583C9A616FDC1978F">Schmitter-Soto, Juan J.</mods:namePart>
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<mods:title id="3713C13849069C6D121DFC9AFE05643F">Journal of Natural History</mods:title>
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<mods:part id="89DDD8095CB5FDD313988AF32AB53C42">
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<mods:date id="E685DCD646FE29BD34FBCCF7C7F69B5E">2017</mods:date>
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<mods:detail id="B73E416C6ED9E9C7D54201574016BE3D" type="series">
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<mods:title id="8A474419A5414179F3F8C89DE083A8E1">J. Nat. Hist.</mods:title>
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<mods:detail id="3DC6425ADC55F83FD44728236A09EDAE" type="pubDate">
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<mods:number id="0A0BA8B7C77013AE538AF59914F7464B">2017-07-03</mods:number>
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<mods:number id="F299978A6F4F76050F89595D5D32AD3D">51</mods:number>
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<mods:number id="54AE3F65AB7DB9572B5C9EC4D838E223">21 - 24</mods:number>
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<mods:url id="4A5924E243C5C45330D16FB02D5FF707">http://dx.doi.org/10.1080/00222933.2017.1324050</mods:url>
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<mods:identifier id="2D3A45DCA21A25F452AA96B9439BFC9A" type="DOI">10.1080/00222933.2017.1324050</mods:identifier>
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<mods:identifier id="95E298CC315FD3BFD1E26D231C9840D3" type="ISSN">1464-5262</mods:identifier>
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<treatment id="03806F39C965FFD3FE58FAEED230FAE2" ID-DOI="http://doi.org/10.5281/zenodo.5191154" ID-Zenodo-Dep="5191154" LSID="urn:lsid:plazi:treatment:03806F39C965FFD3FE58FAEED230FAE2" httpUri="http://treatment.plazi.org/id/03806F39C965FFD3FE58FAEED230FAE2" lastPageId="52" lastPageNumber="1382" pageId="50" pageNumber="1380">
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<subSubSection id="C3338DA4C965FFD5FE58FAEED028FAD2" box="[486,830,1371,1397]" pageId="50" pageNumber="1380" type="nomenclature">
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<paragraph id="8B96DE2FC965FFD5FE58FAEED028FAD2" blockId="50.[486,830,1371,1432]" box="[486,830,1371,1397]" pageId="50" pageNumber="1380">
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<heading id="D0DE6943C965FFD5FE58FAEED028FAD2" box="[486,830,1371,1397]" centered="true" fontSize="11" level="2" pageId="50" pageNumber="1380" reason="5">
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<taxonomicName id="4C29A5ACC965FFD5FE58FAEED1CEFAD2" authority="Schmitter-Soto, 2017" authorityName="Schmitter-Soto" authorityYear="2017" box="[486,728,1371,1397]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="50" pageNumber="1380" phylum="Chordata" rank="species" species="cubilhuitz" status="sp. nov.">
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<emphasis id="B95D023DC965FFD5FE58FAEED1CEFAD2" bold="true" box="[486,728,1371,1397]" italics="true" pageId="50" pageNumber="1380">Astyanax cubilhuitz</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26EBF46C965FFD5FD5FFAE9D028FAD2" box="[737,830,1372,1397]" pageId="50" pageNumber="1380" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3338DA4C965FFD5FD88FACAD192FA46" pageId="50" pageNumber="1380" type="description">
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<paragraph id="8B96DE2FC965FFD5FD88FACAD1FBFA3F" blockId="50.[486,830,1371,1432]" box="[566,749,1407,1432]" pageId="50" pageNumber="1380">
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(
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<figureCitation id="1312C2AAC965FFD5FD80FACAD1ACFA3F" box="[574,698,1407,1432]" captionStart="Figure 24" captionStartId="37.[160,225,1645,1668]" captionTargetBox="[163,1151,941,1611]" captionTargetId="figure-183@37.[163,1153,941,1614]" captionTargetPageId="37" captionText="Figure 24. Map of northern Central America, showing distribution localities of Astyanax baileyi, A. belizianus, A. brevimanus, A. cubilhuitz sp. nov., A. dorioni and A. petenensis. A larger symbol denotes the type locality." figureDoi="http://doi.org/10.5281/zenodo.5181937" httpUri="https://zenodo.org/record/5181937/files/figure.png" pageId="50" pageNumber="1380">Figures 24</figureCitation>
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,
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<figureCitation id="1312C2AAC965FFD5FD77FACAD1F0FA3F" box="[713,742,1407,1432]" captionStart="Figure 32" captionStartId="52.[160,225,581,604]" captionTargetBox="[163,1153,142,550]" captionTargetId="figure-181@52.[163,1153,142,550]" captionTargetPageId="52" captionText="Figure 32. Astyanax cubilhuitz sp. nov., holotype. UMMZ 250290." figureDoi="http://doi.org/10.5281/zenodo.5181951" httpUri="https://zenodo.org/record/5181951/files/figure.png" pageId="50" pageNumber="1380">32</figureCitation>
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)
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</paragraph>
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<paragraph id="8B96DE2FC965FFD5FF1EFA7CD192FA46" blockId="50.[160,644,1481,1505]" box="[160,644,1481,1505]" pageId="50" pageNumber="1380">
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<taxonomicName id="4C29A5ACC965FFD5FF1EFA7CD215FA46" authorityName="Baird and Girard" authorityYear="1854" box="[160,259,1481,1505]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="50" pageNumber="1380" phylum="Chordata" rank="genus">
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<emphasis id="B95D023DC965FFD5FF1EFA7CD215FA46" box="[160,259,1481,1505]" italics="true" pageId="50" pageNumber="1380">Astyanax</emphasis>
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</taxonomicName>
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‘Cubilhuitz’,
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<bibRefCitation id="EFB8A3DEC965FFD5FE29FA7CD168FA46" author="Schmitter-Soto JJ" box="[407,638,1481,1505]" pageId="50" pageNumber="1380" pagination="101 - 130" refId="ref49172" refString="Schmitter-Soto JJ. 2016. A phylogeny of Astyanax (Characiformes: Characidae) in Central and North America. Zootaxa. 4109: 101 - 130." type="journal article" year="2016">Schmitter-Soto 2016</bibRefCitation>
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.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3338DA4C965FFD4FF1EF9B8D792FF6E" lastPageId="51" lastPageNumber="1381" pageId="50" pageNumber="1380" type="diagnosis">
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<paragraph id="8B96DE2FC965FFD5FF1EF9B8D202F981" blockId="50.[160,1156,1549,1713]" box="[160,276,1549,1574]" pageId="50" pageNumber="1380">
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<emphasis id="B95D023DC965FFD5FF1EF9B8D202F981" bold="true" box="[160,276,1549,1574]" italics="true" pageId="50" pageNumber="1380">Diagnosis</emphasis>
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</paragraph>
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<paragraph id="8B96DE2FC965FFD4FF1EF984D792FF6E" blockId="50.[160,1156,1549,1713]" lastBlockId="51.[160,1156,142,201]" lastPageId="51" lastPageNumber="1381" pageId="50" pageNumber="1380">
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Diagnosed from other
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<taxonomicName id="4C29A5ACC965FFD5FE06F984D16AF9EE" box="[440,636,1585,1609]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="50" pageNumber="1380" phylum="Chordata" rank="species" species="undetermined">
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<emphasis id="B95D023DC965FFD5FE06F984D10DF9EE" box="[440,539,1585,1609]" italics="true" pageId="50" pageNumber="1380">Astyanax</emphasis>
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species
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</taxonomicName>
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in north-eastern
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<collectingCountry id="F33E9EBFC965FFD5FCE6F984D0C7F9EE" box="[856,977,1585,1609]" name="Guatemala" pageId="50" pageNumber="1380">Guatemala</collectingCountry>
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and
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<collectingCountry id="F33E9EBFC965FFD5FBA5F984D74AF9EE" box="[1051,1116,1585,1609]" name="Belize" pageId="50" pageNumber="1380">Belize</collectingCountry>
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as follows: body depth, 35% SL or less (35% or more in
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<taxonomicName id="4C29A5ACC965FFD5FC9EF9E6D0D1F9CC" authority="Schmitter-Soto, 2017" authorityName="Schmitter-Soto" authorityYear="2017" box="[800,967,1619,1643]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="50" pageNumber="1380" phylum="Chordata" rank="species" species="bacalarensis" status="sp. nov.">
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<emphasis id="B95D023DC965FFD5FC9EF9E6D0D1F9CC" box="[800,967,1619,1643]" italics="true" pageId="50" pageNumber="1380">A. bacalarensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26EBF46C965FFD5FC6CF9E6D73BF9CC" box="[978,1069,1619,1643]" pageId="50" pageNumber="1380" rank="species">sp. nov.</taxonomicNameLabel>
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and
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<taxonomicName id="4C29A5ACC965FFD5FBD0F9E6D21AF929" baseAuthorityName="Bocourt" baseAuthorityYear="1868" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="50" pageNumber="1380" phylum="Chordata" rank="species" species="belizianus">
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<emphasis id="B95D023DC965FFD5FBD0F9E6D21AF929" italics="true" pageId="50" pageNumber="1380">A. belizianus</emphasis>
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</taxonomicName>
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); eye larger, diameter 35% HL (vs 32% or less); mean predorsal length, 52% SL (49% in
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<taxonomicName id="4C29A5ACC965FFD5FF40F92CD248F916" authority="Schmitter-Soto, 2017" authorityName="Schmitter-Soto" authorityYear="2017" box="[254,350,1689,1713]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="50" pageNumber="1380" phylum="Chordata" rank="species" species="macal" status="sp. nov.">
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<emphasis id="B95D023DC965FFD5FF40F92CD248F916" box="[254,350,1689,1713]" italics="true" pageId="50" pageNumber="1380">A. macal</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26EBF46C965FFD5FED9F92CD2A9F916" box="[359,447,1689,1713]" pageId="50" pageNumber="1380" rank="species">sp. nov.</taxonomicNameLabel>
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and
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<taxonomicName id="4C29A5ACC965FFD5FE44F92CD19EF916" baseAuthorityName="Gunther" baseAuthorityYear="1864" box="[506,648,1689,1713]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="50" pageNumber="1380" phylum="Chordata" rank="species" species="petenensis">
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<emphasis id="B95D023DC965FFD5FE44F92CD19EF916" box="[506,648,1689,1713]" italics="true" pageId="50" pageNumber="1380">A. petenensis</emphasis>
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</taxonomicName>
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); mean anal-base length, 26% SL (vs 28% or more); longest anal-fin ray, mean 17% SL (vs 14%); upper pharyngeal plate, oval (crescent-shaped, in
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<taxonomicName id="4C29A5ACC964FFD4FE35FF04D105FF6E" baseAuthorityName="Bocourt" baseAuthorityYear="1868" box="[395,531,177,201]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="51" pageNumber="1381" phylum="Chordata" rank="species" species="belizianus">
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<emphasis id="B95D023DC964FFD4FE35FF04D105FF6E" box="[395,531,177,201]" italics="true" pageId="51" pageNumber="1381">A. belizianus</emphasis>
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</taxonomicName>
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); humeral spot, rectangular (vs indistinct or P-shaped).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3338DA4C964FFD4FF1EFF40D1A8F93B" pageId="51" pageNumber="1381" type="description">
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<paragraph id="8B96DE2FC964FFD4FF1EFF40D230FEA9" blockId="51.[160,1157,245,1692]" box="[160,294,245,270]" pageId="51" pageNumber="1381">
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<emphasis id="B95D023DC964FFD4FF1EFF40D230FEA9" bold="true" box="[160,294,245,270]" italics="true" pageId="51" pageNumber="1381">Description</emphasis>
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</paragraph>
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<paragraph id="8B96DE2FC964FFD4FF1EFEACD793FE96" blockId="51.[160,1157,245,1692]" box="[160,1157,281,305]" pageId="51" pageNumber="1381">
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A species of
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<taxonomicName id="4C29A5ACC964FFD4FE92FEACD299FE96" authorityName="Baird and Girard" authorityYear="1854" box="[300,399,281,305]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="51" pageNumber="1381" phylum="Chordata" rank="genus">
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<emphasis id="B95D023DC964FFD4FE92FEACD299FE96" box="[300,399,281,305]" italics="true" pageId="51" pageNumber="1381">Astyanax</emphasis>
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</taxonomicName>
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, subgenus
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<taxonomicName id="4C29A5ACC964FFD4FDB1FEACD164FE96" authorityName="Baird and Girard" authorityYear="1854" box="[527,626,281,305]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="51" pageNumber="1381" phylum="Chordata" rank="genus">
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<emphasis id="B95D023DC964FFD4FDB1FEACD164FE96" box="[527,626,281,305]" italics="true" pageId="51" pageNumber="1381">Astyanax</emphasis>
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</taxonomicName>
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(i.e. with a complete predorsal series of scales).
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</paragraph>
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<paragraph id="8B96DE2FC964FFD4FF7EFE89D219FE3E" blockId="51.[160,1157,245,1692]" pageId="51" pageNumber="1381">Head profile straight; snout round. Lips even; mouth terminal. Pectoral fins always reach pelvic fin origin; anal and dorsal fins may overlap. Lobes of caudal fin, subequal.</paragraph>
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<paragraph id="8B96DE2FC964FFD4FF7EFE11D270FD09" blockId="51.[160,1157,245,1692]" pageId="51" pageNumber="1381">
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D. 9; A. 21–26, modally 23; pect. 12. Procurrent unsegmented dorsal rays on caudal fin, 10 or more. Gill rakers on first arch, 21–23, modally 22; on lower limb, 12–14, modally 13. Scales on lateral line, 34–38, mean 36.5; predorsal scales, 12–13; scale rows from lateral line to base of first dorsal-fin ray, 6.5–8, modally 7; to base of pelvic fin, modally 6, up to 6.5; to base of pectoral fin, 3.5–4; circumpeduncular scales, 13–16, mean 15. A single scale row on anal fin base, rather short. Nuptial tubercles not seen. Total vertebrae 32–33, 18–19 caudal. Detailed frequencies are given in
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<tableCitation id="C6ABEB94C964FFD4FEB4FD23D276FD09" box="[266,352,662,686]" captionStart="Table 3" captionStartId="16.[160,216,142,165]" captionText="Table 3. Meristics of Central and North American species in the genus Astyanax. Numbers are" pageId="51" pageNumber="1381">Table 3</tableCitation>
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.
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</paragraph>
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<paragraph id="8B96DE2FC964FFD4FF7EFD0CD0B2FCB1" blockId="51.[160,1157,245,1692]" pageId="51" pageNumber="1381">
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Largest examined specimen, 53.0 mm SL. Body slender, 32–35% SL. Head length, 25–28% SL; orbital diameter somewhat large, 33–37% HL; interorbital distance, 7.2–9.1% SL, mean 7.9% SL (further morphometric data are given in
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<tableCitation id="C6ABEB94C964FFD4FCFAFD4BD08EFCB1" box="[836,920,766,790]" captionStart="Table 4" captionStartId="19.[160,216,137,160]" captionText="Table 4. Selected morphometric proportions of Central and North American species in the genus Astyanax (abbreviated by first letters of specific epithet). Specimens measured are those marked ‘dig’ in Appendix. All measurements given as percentage of standard length. Data are means; whole range in parentheses." pageId="51" pageNumber="1381">Table 4</tableCitation>
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).
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</paragraph>
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<paragraph id="8B96DE2FC964FFD4FF7EFC94D1AFF9F1" blockId="51.[160,1157,245,1692]" pageId="51" pageNumber="1381">Anterior fontanel long, straight-sided, blunt-tipped. Supraoccipital process in dorsal view, short, wide-based; slightly concave in lateral view. Vomer rostrally slightly concave. Dentigerous arm of premaxilla, longer; 0–4 teeth. Highest tooth on dentary, first or third; posterior teeth, abruptly smaller. Dorsal edge of longer articular arm, straight. Maxillary, with a convex anteroinferior edge; 1–3 teeth. Quadrate, dorsal process expanded. Metapterygoid, rostral arm longer than ventral, 2 dorsorostral projections. Infraorbital II, triangular with an angled base; infraorbital III, inferoposteriorly semicircular; infraorbital IV, squarish with a projection; a wide contact between infraorbitals II and III. Urohyal rostral end turned up, blunt, its ventrorostral edge convex, its ventral apex about equidistant between the rostral and caudal ends; ceratohyal foramen oval; rostral vertices of ceratohyal round, the ventral side straight. Epibranchial III, insertion of uncinate process angled, the distal segment of the main body straight. Upper pharyngeal bones, oval. Dorsal side of hyomandibular, convex. Opercular dorsal edge, convex; sides of dorsal half, parallel; posterior edge, dorsally concave, ventrally straight-convex; ventral tip, rounded. Interopercle, posterior edge, straight-convex, with a spine. Preopercle, anterodorsal edge, with a median convexity; ventral rim, straight; posteroventral edge, curved; 2 divergent canals at angle. Five predorsal bony elements, distally expanded; rostral edge of first pterygiophore angled. Coracoid with 1 interdigitation in suture to cleithrum, a concave caudal edge, a single posteroinferior spine. Caudad process of postcleithrum, digitiform. Proximal edge of pelvic bone, convex. Postanal element, short. Dorsal tip of scaphium, truncate; caudal tip, sharp; dorsoposterior edge, slightly concave. Neural spines under dorsal fin, straight. Sixth caudal vertebra from tail, with a haemal spine displaced caudad. Rostral edge of largest hypuric plate, pointed. Epuric plate on last neural spine, distally spiny.</paragraph>
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<paragraph id="8B96DE2FC964FFD4FF7EF9D4D1A8F93B" blockId="51.[160,1157,245,1692]" pageId="51" pageNumber="1381">Humeral spot, approximately rectangular. Pigment on anal fin, uniformly sparse. Caudal spot, both on peduncle and on fin rays.</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF568EA7C963FFD3FF1EFDF0D03BFDFB" ID-DOI="http://doi.org/10.5281/zenodo.5181951" ID-Zenodo-Dep="5181951" box="[160,813,581,604]" httpUri="https://zenodo.org/record/5181951/files/figure.png" pageId="52" pageNumber="1382" startId="52.[160,225,581,604]" targetBox="[163,1153,142,550]" targetPageId="52">
|
||
<paragraph id="8B96DE2FC963FFD3FF1EFDF0D03BFDFB" blockId="52.[160,813,581,604]" box="[160,813,581,604]" pageId="52" pageNumber="1382">
|
||
<emphasis id="B95D023DC963FFD3FF1EFDF0D21FFDFB" bold="true" box="[160,265,581,604]" pageId="52" pageNumber="1382">Figure 32.</emphasis>
|
||
<taxonomicName id="4C29A5ACC963FFD3FEAFFDF0D2DAFDFB" authority="Schmitter-Soto, 2017" authorityName="Schmitter-Soto" authorityYear="2017" box="[273,460,581,604]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="52" pageNumber="1382" phylum="Chordata" rank="species" species="cubilhuitz" status="sp. nov.">
|
||
<emphasis id="B95D023DC963FFD3FEAFFDF0D2DAFDFB" box="[273,460,581,604]" italics="true" pageId="52" pageNumber="1382">Astyanax cubilhuitz</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26EBF46C963FFD3FE6BFDF0D130FDFB" box="[469,550,581,604]" pageId="52" pageNumber="1382" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype. UMMZ 250290.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3338DA4C963FFD3FF1EFD2CD109FC9A" pageId="52" pageNumber="1382" type="materials_examined">
|
||
<paragraph id="8B96DE2FC963FFD3FF1EFD2CD2E2FD15" blockId="52.[160,1156,665,829]" box="[160,500,665,690]" pageId="52" pageNumber="1382">
|
||
<emphasis id="B95D023DC963FFD3FF1EFD2CD2E2FD15" bold="true" box="[160,500,665,690]" italics="true" pageId="52" pageNumber="1382">Type material and depositor</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B96DE2FC963FFD3FF1EFD08D109FC9A" blockId="52.[160,1156,665,829]" pageId="52" pageNumber="1382">
|
||
<materialsCitation id="3B41D472C963FFD3FF1EFD08D1BCFCBD" collectionCode="R" collectorName="Icbolay' & D. E. Rosen & Bailey" country="Guatemala" elevation="300" location="Rio Dolores" municipality="Cubilhuitz" pageId="52" pageNumber="1382" specimenCode="UMMZ 250290" specimenCount="1" stateProvince="Alta Verapaz" typeStatus="holotype">
|
||
<typeStatus id="5492608DC963FFD3FF1EFD08D211FD72" box="[160,263,701,725]" pageId="52" pageNumber="1382" type="holotype">Holotype</typeStatus>
|
||
<specimenCode id="DB8F7654C963FFD3FEAEFD08D2AFFD72" box="[272,441,701,725]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="52" pageNumber="1382" type="University or college">UMMZ 250290</specimenCode>
|
||
, 53.0 mm SL, ‘spring origin of short stream with underground drainage to
|
||
<location id="8EF688F4C963FFD3FE92FD6AD2A7FD50" LSID="urn:lsid:plazi:treatment:03806F39C965FFD3FE58FAEED230FAE2:8EF688F4C963FFD3FE92FD6AD2A7FD50" box="[300,433,735,759]" country="Guatemala" municipality="Cubilhuitz" name="Rio Dolores" pageId="52" pageNumber="1382" stateProvince="Alta Verapaz">Río Dolores</location>
|
||
,
|
||
<collectingMunicipality id="6BF24455C963FFD3FE03FD6AD127FD50" box="[445,561,735,759]" pageId="52" pageNumber="1382">Cubilhuitz</collectingMunicipality>
|
||
,
|
||
<elevation id="0004391CC963FFD3FD83FD6AD1E1FD50" box="[573,759,735,759]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="52" pageNumber="1382" unit="m" value="300.0">
|
||
elevation
|
||
<quantity id="4CD173CAC963FFD3FD13FD6AD1E1FD50" box="[685,759,735,759]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="52" pageNumber="1382" unit="m" value="300.0">300 m</quantity>
|
||
</elevation>
|
||
,
|
||
<collectorName id="26DCBBF9C963FFD3FCBAFD6AD04FFD50" box="[772,857,735,759]" pageId="52" pageNumber="1382">Icbolay’</collectorName>
|
||
,
|
||
<collectingRegion id="49ED10CDC963FFD3FCD6FD6AD0EFFD50" box="[872,1017,735,759]" country="Guatemala" name="Alta Verapaz" pageId="52" pageNumber="1382">Alta Verapaz</collectingRegion>
|
||
,
|
||
<collectingCountry id="F33E9EBFC963FFD3FBBAFD6AD796FD50" box="[1028,1152,735,759]" name="Guatemala" pageId="52" pageNumber="1382">Guatemala</collectingCountry>
|
||
, coll.
|
||
<collectorName id="26DCBBF9C963FFD3FF6BFCB7D25DFCBD" box="[213,331,770,794]" pageId="52" pageNumber="1382">D.E. Rosen</collectorName>
|
||
and
|
||
<collectionCode id="ED3846EAC963FFD3FE3BFCB7D280FCBD" box="[389,406,770,794]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="52" pageNumber="1382">R</collectionCode>
|
||
.
|
||
<collectorName id="26DCBBF9C963FFD3FE1FFCB7D2F3FCBD" box="[417,485,770,794]" pageId="52" pageNumber="1382">Bailey</collectorName>
|
||
, 1968 (
|
||
<figureCitation id="1312C2AAC963FFD3FD89FCB7D1B7FCBD" box="[567,673,770,794]" captionStart="Figure 32" captionStartId="52.[160,225,581,604]" captionTargetBox="[163,1153,142,550]" captionTargetId="figure-181@52.[163,1153,142,550]" captionTargetPageId="52" captionText="Figure 32. Astyanax cubilhuitz sp. nov., holotype. UMMZ 250290." figureDoi="http://doi.org/10.5281/zenodo.5181951" httpUri="https://zenodo.org/record/5181951/files/figure.png" pageId="52" pageNumber="1382">Figure 32</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<typeStatus id="5492608DC963FFD3FD09FCB7D031FCBD" box="[695,807,770,794]" pageId="52" pageNumber="1382" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<collectionCode id="ED3846EAC963FFD3FC8FFCB7D06AFCBD" box="[817,892,770,794]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="52" pageNumber="1382" type="University or college">UMMZ</collectionCode>
|
||
188007,
|
||
<specimenCount id="9D2F15A6C963FFD3FC5AFCB7D796FCBD" box="[996,1152,770,794]" pageId="52" pageNumber="1382" type="generic">55 specimens</specimenCount>
|
||
, same collection data as
|
||
<typeStatus id="5492608DC963FFD3FE0AFC90D10DFC9A" box="[436,539,805,829]" pageId="52" pageNumber="1382" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3338DA4C963FFD3FF1EFCDCD1E3FC02" box="[160,757,873,933]" pageId="52" pageNumber="1382" type="etymology">
|
||
<paragraph id="8B96DE2FC963FFD3FF1EFCDCD20BFC25" blockId="52.[160,757,873,933]" box="[160,285,873,898]" pageId="52" pageNumber="1382">
|
||
<emphasis id="B95D023DC963FFD3FF1EFCDCD20BFC25" bold="true" box="[160,285,873,898]" italics="true" pageId="52" pageNumber="1382">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B96DE2FC963FFD3FF1EFC38D1E3FC02" blockId="52.[160,757,873,933]" box="[160,757,909,933]" pageId="52" pageNumber="1382">
|
||
Named after the
|
||
<typeStatus id="5492608DC963FFD3FEDAFC38D280FC02" box="[356,406,909,933]" pageId="52" pageNumber="1382">type</typeStatus>
|
||
locality. A noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3338DA4C963FFD3FF1EFC64D17CFB30" pageId="52" pageNumber="1382" type="distribution">
|
||
<paragraph id="8B96DE2FC963FFD3FF1EFC64D23AFC4D" blockId="52.[160,1156,977,1071]" box="[160,300,977,1002]" pageId="52" pageNumber="1382">
|
||
<emphasis id="B95D023DC963FFD3FF1EFC64D23AFC4D" bold="true" box="[160,300,977,1002]" italics="true" pageId="52" pageNumber="1382">Distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B96DE2FC963FFD3FF1EFC40D2B4FB88" blockId="52.[160,1156,977,1071]" pageId="52" pageNumber="1382">
|
||
Río Dolores, a stream near Cubilhuitz isolated by a subterranean segment,
|
||
<collectingRegion id="49ED10CDC963FFD3FC4EFC40D796FBAA" box="[1008,1152,1013,1037]" country="Guatemala" name="Alta Verapaz" pageId="52" pageNumber="1382">Alta Verapaz</collectingRegion>
|
||
,
|
||
<collectingCountry id="F33E9EBFC963FFD3FF1EFBA2D20FFB88" box="[160,281,1047,1071]" name="Guatemala" pageId="52" pageNumber="1382">Guatemala</collectingCountry>
|
||
(
|
||
<figureCitation id="1312C2AAC963FFD3FE94FBA2D282FB88" box="[298,404,1047,1071]" captionStart="Figure 24" captionStartId="37.[160,225,1645,1668]" captionTargetBox="[163,1151,941,1611]" captionTargetId="figure-183@37.[163,1153,941,1614]" captionTargetPageId="37" captionText="Figure 24. Map of northern Central America, showing distribution localities of Astyanax baileyi, A. belizianus, A. brevimanus, A. cubilhuitz sp. nov., A. dorioni and A. petenensis. A larger symbol denotes the type locality." figureDoi="http://doi.org/10.5281/zenodo.5181937" httpUri="https://zenodo.org/record/5181937/files/figure.png" pageId="52" pageNumber="1382">Figure 24</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B96DE2FC963FFD3FF1EFBE9D2C0FBD2" blockId="52.[160,618,1116,1175]" box="[160,470,1116,1141]" pageId="52" pageNumber="1382">
|
||
<emphasis id="B95D023DC963FFD3FF1EFBE9D2C0FBD2" bold="true" box="[160,470,1116,1141]" italics="true" pageId="52" pageNumber="1382">Proposed common names</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B96DE2FC963FFD3FF1EFBCAD17CFB30" blockId="52.[160,618,1116,1175]" box="[160,618,1151,1175]" pageId="52" pageNumber="1382">Cubilhuitz tetra, sardinita del Cubilhuitz.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3338DA4C963FFD3FF1EFB71D230FAE2" pageId="52" pageNumber="1382" type="discussion">
|
||
<paragraph id="8B96DE2FC963FFD3FF1EFB71D210FB7A" blockId="52.[160,1155,1220,1349]" box="[160,262,1220,1245]" pageId="52" pageNumber="1382">
|
||
<emphasis id="B95D023DC963FFD3FF1EFB71D210FB7A" bold="true" box="[160,262,1220,1245]" italics="true" pageId="52" pageNumber="1382">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B96DE2FC963FFD3FF1EFB52D230FAE2" blockId="52.[160,1155,1220,1349]" pageId="52" pageNumber="1382">
|
||
Although probably not syntopic with
|
||
<taxonomicName id="4C29A5ACC963FFD3FD83FB52D1A9FB58" authority="Schmitter-Soto, 2017" authorityName="Schmitter-Soto" authorityYear="2017" box="[573,703,1255,1279]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="52" pageNumber="1382" phylum="Chordata" rank="species" species="cubilhuitz" status="sp. nov.">
|
||
<emphasis id="B95D023DC963FFD3FD83FB52D1A9FB58" box="[573,703,1255,1279]" italics="true" pageId="52" pageNumber="1382">A. cubilhuitz</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26EBF46C963FFD3FD78FB52D00AFB58" box="[710,796,1255,1279]" pageId="52" pageNumber="1382" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, several fish species are endemic to this karstic region, such as poeciliids in genera
|
||
<taxonomicName id="4C29A5ACC963FFD3FD54FABFD066FA85" authorityName="Heckel" authorityYear="1848" box="[746,880,1290,1314]" class="Actinopterygii" family="Poeciliidae" genus="Xiphophorus" kingdom="Animalia" order="Cyprinodontiformes" pageId="52" pageNumber="1382" phylum="Chordata" rank="genus">
|
||
<emphasis id="B95D023DC963FFD3FD54FABFD066FA85" box="[746,880,1290,1314]" italics="true" pageId="52" pageNumber="1382">Xiphophorus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C29A5ACC963FFD3FC0EFABFD738FA85" authorityName="Agassiz" authorityYear="1853" box="[944,1070,1290,1314]" class="Actinopterygii" family="Poeciliidae" genus="Heterandria" kingdom="Animalia" order="Cyprinodontiformes" pageId="52" pageNumber="1382" phylum="Chordata" rank="genus">
|
||
<emphasis id="B95D023DC963FFD3FC0EFABFD738FA85" box="[944,1070,1290,1314]" italics="true" pageId="52" pageNumber="1382">Heterandria</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFB8A3DEC963FFD3FB81FABFD3C0FAE2" author="Rosen DE" pageId="52" pageNumber="1382" pagination="159 - 188" refId="ref48999" refString="Rosen DE. 1978. Vicariant patterns and historical explanation in biogeography. Syst Biol. 27: 159 - 188." type="journal article" year="1978">Rosen 1978</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFB8A3DEC963FFD3FF5AFA98D20CFAE2" author="Rosen DE" box="[228,282,1325,1349]" pageId="52" pageNumber="1382" pagination="267 - 376" refId="ref49021" refString="Rosen DE. 1979. Fishes from the uplands and intermontane basins of Guatemala: revisionary studies and comparative geography. Bull Am Mus Nat Hist. 162: 267 - 376." type="journal article" year="1979">1979</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |