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<document id="F54A5FB9AC5BCF015C41830E9DEBBB13" ID-DOI="10.11646/zootaxa.5535.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="14023694" ID-ZooBank="42A7715C-0CE3-4CE7-BF67-BDB5D5B9F12B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1730455184296" checkinUser="plazi" docAuthor="Enushchenko, Ilya V., Makarov, Kirill V. &amp; Shavrin, Alexey V." docDate="2024" docId="039F21082243684FFF30F935DE28CE31" docLanguage="en" docName="zootaxa.5535.1.1.pdf" docOrigin="Zootaxa 5535 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5535.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Gyrophaena zhagaensis Pace 2003" docType="treatment" docVersion="2" lastPageNumber="64" masterDocId="FFA65970227D6870FFA7FF92DD71CC28" masterDocTitle="Review of the subtribe Gyrophaenina Kraatz, 1856 (Coleoptera: Staphylinidae: Aleocharinae: Homalotini) of Kunashir Island, with notes on species of Far Eastern Russia" masterLastPageNumber="81" masterPageNumber="1" pageNumber="63" updateTime="1730455996750" updateUser="ExternalLinkService">
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<mods:title id="A430B0DD146336B32EB7249214AAF34A">Review of the subtribe Gyrophaenina Kraatz, 1856 (Coleoptera: Staphylinidae: Aleocharinae: Homalotini) of Kunashir Island, with notes on species of Far Eastern Russia</mods:title>
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<mods:namePart id="25A8A87ED2C74BFBF871724871C69069">Enushchenko, Ilya V.</mods:namePart>
<mods:nameIdentifier id="645F4E9A4E65B771593740A6E1CDEC92" type="ORCID">0000-0002-8166-6079</mods:nameIdentifier>
<mods:affiliation id="B3D967917E7CA10A827D00581477A973">Testing Laboratory of the Baikalskiy branch of the All-Russian Plant Quarantine Center, Krasnoyarskaya str. 77, Irkutsk, 664023, Russia. deschampsia @ yandex. ru; https: // orcid. org / 0000 - 0002 - 8166 - 6079</mods:affiliation>
<mods:nameIdentifier id="AC5BC49DAC2CE2532049A2DFF795686F" type="email">deschampsia@yandex.ru</mods:nameIdentifier>
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<mods:name id="3219226D631D05040805FA6473ADEF20" type="personal">
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<mods:namePart id="8439B0B4EBF3120C5C8CC6D03282CACA">Makarov, Kirill V.</mods:namePart>
<mods:nameIdentifier id="C31EB844C74A94E3282A4C72D9E3AB79" type="ORCID">0000-0002-9184-7869</mods:nameIdentifier>
<mods:affiliation id="BFB080580BFD4EC452FAAB797E6B5D50">Moscow State Pedagogical University, Malaya Pirogovskaya str., 1 / 1, 119435, Moscow kvmac @ inbox. ru; http: // orcid. org / 0000 - 0002 - 9184 - 7869</mods:affiliation>
<mods:nameIdentifier id="2B7CDB35900F76FCA2FA6BA94D382971" type="email">kvmac@inbox.ru</mods:nameIdentifier>
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<mods:roleTerm id="0C0F7B3874DCA294B7C77EE7CFDBC2F6">Author</mods:roleTerm>
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<mods:namePart id="A403918153EF53DBE34D43B44D260C49">Shavrin, Alexey V.</mods:namePart>
<mods:nameIdentifier id="A3B82C7ADCE2C9E856B16DF63585798C" type="ORCID">0000-0001-6487-4530</mods:nameIdentifier>
<mods:affiliation id="221EF84F717433D4528F9980B8A4CE21">Institute of Life Sciences and Technologies, Daugavpils University, Vienibas 13, Daugavpils, LV- 5401, Latvia. ashavrin @ hotmail. com; https: // orcid. org / 0000 - 0001 - 6487 - 4530 * Corresponding author</mods:affiliation>
<mods:nameIdentifier id="3E105FCFE45942E624C451030C092EE9" type="email">ashavrin@hotmail.com</mods:nameIdentifier>
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<mods:title id="B2C9AF934F0ED99160FCE5331608F805">Zootaxa</mods:title>
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<mods:date id="77BB775F99BE85657EF33481770D38A4">2024</mods:date>
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<subSubSection id="C32CC3952243684EFF30F935DF80CAE9" box="[151,753,1703,1730]" pageId="62" pageNumber="63" type="nomenclature">
<paragraph id="8B89901E2243684EFF30F935DF80CAE9" blockId="62.[151,753,1703,1765]" box="[151,753,1703,1730]" pageId="62" pageNumber="63">
<emphasis id="B9424C0C2243684EFF30F935DF80CAE9" bold="true" box="[151,753,1703,1730]" pageId="62" pageNumber="63">
3.2.17.
<taxonomicName id="4C36EB9D2243684EFF48F935DF80CAE9" ID-CoL="6L8TR" authority="PaCe, 2003" authorityName="Pace" authorityYear="2003" box="[239,753,1703,1730]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="zhagaensis">
<emphasis id="B9424C0C2243684EFF48F935DCF4CAEA" bold="true" box="[239,389,1703,1730]" italics="true" pageId="62" pageNumber="63">Gyrophaena</emphasis>
(s.str.)
<emphasis id="B9424C0C2243684EFE46F935DF14CAEA" bold="true" box="[481,613,1703,1730]" italics="true" pageId="62" pageNumber="63">zhagaensis</emphasis>
<bibRefCitation id="EFA7EDEF2243684EFDCBF935DF80CAE9" author="Pace, R." box="[620,753,1703,1729]" pageId="62" pageNumber="63" pagination="621 - 660" refId="ref52519" refString="Pace, R. (2003) Gyrophaenini della Cina (Coleoptera, Staphylinidae). Revue Suisse de Zoologie, 110 (3), 621 - 660. https: // doi. org / 10.5962 / bhl. part. 80203" type="journal article" year="2003">PaCe, 2003</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
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<subSubSection id="C32CC3952243684EFF30F959DC0ACACD" box="[151,379,1739,1765]" pageId="62" pageNumber="63" type="description">
<paragraph id="8B89901E2243684EFF30F959DC0ACACD" blockId="62.[151,753,1703,1765]" box="[151,379,1739,1765]" pageId="62" pageNumber="63">
(
<figureCitation id="130D8C9B2243684EFF38F959DC4CCACD" box="[159,317,1739,1765]" captionStart="FIGURES 257–264" captionStartId="62.[151,264,1533,1557]" captionTargetBox="[158,1431,158,1511]" captionTargetId="figure-21@62.[151,1435,151,1517]" captionTargetPageId="62" captionText="FIGURES 257–264. Gyrophaena (s.str.) williamsi (Figs 257–259) and G. (s.str.) zhagaensis (Figs 260–264): 257, 260—habitus, 258, 261—aedeagus, lateral view, 262—paramere, 259, 263—male abdominal tergite VIII, 264—male abdominal sternite VIII. Scale bars: 1.0 mm (Figs 257, 260), 0.1 mm (Figs 258–259, 261–264)." figureDoi="http://doi.org/10.5281/zenodo.14023772" httpUri="https://zenodo.org/record/14023772/files/figure.png" pageId="62" pageNumber="63">Figs 260–264</figureCitation>
,
<figureCitation id="130D8C9B2243684EFEEFF959DC05CACD" box="[328,372,1739,1765]" captionStart="FIGURES 305–311" captionStartId="70.[151,264,1784,1808]" captionTargetBox="[151,1436,181,1760]" captionTargetId="figure-21@70.[151,1436,181,1760]" captionTargetPageId="70" captionText="FIGURES 305–311. Distribution of Gyrophaena in Eastern Siberia and Far Eastern Russia: 305—G. (s.str.) pseudonana, 306—G. (s.str.) pseudonitidula, 307—G. (s.str.) pulchella, 308—G. (s.str.) triquetra, 309—G. (s.str.) williamsi, 310—G. (s.str.) zhagaensis, 311—G. (Phaenogyra) strictula." figureDoi="http://doi.org/10.5281/zenodo.14023790" httpUri="https://zenodo.org/record/14023790/files/figure.png" pageId="62" pageNumber="63">310</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32CC3952243684EFF30F882DF09CB6E" pageId="62" pageNumber="63" type="reference_group">
<paragraph id="8B89901E2243684EFF30F882DF95CB00" blockId="62.[151,740,1808,1862]" box="[151,740,1808,1832]" pageId="62" pageNumber="63">
<treatmentCitationGroup id="AB26B7302243684EFF30F882DF95CB00" box="[151,740,1808,1832]" pageId="62" pageNumber="63">
<treatmentCitation id="0A97B60F2243684EFF30F882DF47CB00" author="Pace, R." box="[151,566,1808,1832]" page="643" pageId="62" pageNumber="63" year="2003">
<taxonomicName id="4C36EB9D2243684EFF30F882DF47CB00" ID-CoL="6L8TR" authority="Pace, 2003: 643" authorityName="Pace" authorityPageNumber="643" authorityYear="2003" box="[151,566,1808,1832]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="zhagaensis">
<emphasis id="B9424C0C2243684EFF30F882DCFBCB00" box="[151,394,1808,1832]" italics="true" pageId="62" pageNumber="63">Gyrophaena zhagaensis</emphasis>
<bibRefCitation id="EFA7EDEF2243684EFE36F882DF47CB00" author="Pace, R." box="[401,566,1808,1832]" pageId="62" pageNumber="63" pagination="621 - 660" refId="ref52519" refString="Pace, R. (2003) Gyrophaenini della Cina (Coleoptera, Staphylinidae). Revue Suisse de Zoologie, 110 (3), 621 - 660. https: // doi. org / 10.5962 / bhl. part. 80203" type="journal article" year="2003">Pace, 2003: 643</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0A97B60F2243684EFDE6F882DF91CB00" author="Pace, R." box="[577,736,1808,1832]" httpUri="http://treatment.plazi.org/id/B301E003882EFFAAFF7DFD04FCFBFC30" page="129" pageId="62" pageNumber="63" year="2010">
<bibRefCitation id="EFA7EDEF2243684EFDE6F882DF91CB00" author="Pace, R." box="[577,736,1808,1832]" pageId="62" pageNumber="63" pagination="125 - 193" refId="ref52707" refString="Pace, R. (2010) Biodiversita delle Aleocharinae della Cina: Gyrophaenini (Coleoptera, Staphylinidae). Beitrage zur Entomologie, 60 (1), 125 - 193. https: // doi. org / 10.21248 / contrib. entomol. 60.1.125 - 193" type="journal article" year="2010">Pace 2010: 129</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
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<treatmentCitationGroup id="AB26B7302243684EFF30F8BCDF09CB6E" box="[151,632,1838,1862]" pageId="62" pageNumber="63">
<treatmentCitation id="0A97B60F2243684EFF30F8BCDF64CB6E" author="Pace, R." box="[151,533,1838,1862]" page="117" pageId="62" pageNumber="63" year="2007">
<taxonomicName id="4C36EB9D2243684EFF30F8BCDF64CB6E" ID-CoL="6KX6G" authority="Pace, 2007: 117" authorityName="Pace" authorityPageNumber="117" authorityYear="2007" box="[151,533,1838,1862]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="peniculi" status="syn. nov.">
<emphasis id="B9424C0C2243684EFF30F8BCDC1CCB6E" box="[151,365,1838,1862]" italics="true" pageId="62" pageNumber="63">Gyrophaena peniculi</emphasis>
<bibRefCitation id="EFA7EDEF2243684EFED4F8BDDF64CB6E" author="Pace, R." box="[371,533,1838,1862]" pageId="62" pageNumber="63" pagination="103 - 129" refId="ref52664" refString="Pace, R. (2007) Le specie dei generi Gyrophaena Mann. e Brachida Muls. &amp; Rey di Taiwan (Coleoptera, Staphylinidae). Bollettino del Museo Civico di Storia di Verona, 31, 103 - 129." type="journal article" year="2007">Pace, 2007: 117</bibRefCitation>
</taxonomicName>
</treatmentCitation>
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<emphasis id="B9424C0C2243684EFDBCF8BCDF31CB6E" bold="true" box="[539,576,1838,1862]" pageId="62" pageNumber="63">syn</emphasis>
.
<emphasis id="B9424C0C2243684EFDEBF8BCDF09CB6E" bold="true" box="[588,632,1838,1862]" pageId="62" pageNumber="63">nov.</emphasis>
</taxonomicNameLabel>
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</paragraph>
</subSubSection>
<subSubSection id="C32CC3952243684EFF30F8E6DF49CBD3" pageId="62" pageNumber="63" type="materials_examined">
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<emphasis id="B9424C0C2243684EFF30F8E6DC0BCBA6" bold="true" box="[151,378,1908,1934]" pageId="62" pageNumber="63">Material examined</emphasis>
.
<materialsCitation id="3B5E9A432243684EFE2FF8E6DDB6CB9B" collectingDate="1999-08-07" collectingDateMax="1999-08-09" collectingDateMin="1999-08-07" collectionCode="V" location="Siniy Mts." municipality="Spasskiy District" pageId="62" pageNumber="63" specimenCount="1" specimenCount-male="1" stateProvince="Primorskiy">
<emphasis id="B9424C0C2243684EFE2FF8E6DF0ACBA6" bold="true" box="[392,635,1908,1934]" pageId="62" pageNumber="63">
<collectingRegion id="49F25EFC2243684EFE2FF8E6DF0ACBA6" box="[392,635,1908,1934]" country="Russia" name="Primorskiy" pageId="62" pageNumber="63">MARITIME PROVINCE</collectingRegion>
</emphasis>
:
<specimenCount id="9D305B972243684EFD2CF8E7DFC9CBA7" box="[651,696,1909,1935]" count="1" pageId="62" pageNumber="63" type="male">1 ♂</specimenCount>
:
<collectingMunicipality id="6BED0A642243684EFD63F8E6DEF7CBA6" box="[708,902,1908,1934]" pageId="62" pageNumber="63">Spasskiy District</collectingMunicipality>
,
<location id="8EE9C6C52243684EFC36F8E6D978CBA7" LSID="urn:lsid:plazi:treatment:039F21082243684FFF30F935DE28CE31:8EE9C6C52243684EFC36F8E6D978CBA7" box="[913,1033,1908,1935]" municipality="Spasskiy District" name="Siniy Mts." pageId="62" pageNumber="63" stateProvince="Primorskiy">Siniy Mts.</location>
,
<locationDeviation id="9A3B893C2243684EFBB2F8E7D99FCBA6" box="[1045,1262,1908,1935]" location="Evseyevka." pageId="62" pageNumber="63">
<quantity id="4CCE3DFB2243684EFBB2F8E7D93FCBA6" box="[1045,1102,1908,1935]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="62" pageNumber="63" unit="km" value="4.0">4 km</quantity>
E Evseyevka.
</locationDeviation>
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<collectingDate id="EFCC4F362243684EFB52F8E7D8E9CBA7" box="[1269,1432,1909,1935]" pageId="62" pageNumber="63" value="1999-08-07" valueMax="1999-08-09" valueMin="1999-08-07">7–9.VIII.1999</collectingDate>
</date>
. A.
<collectionCode id="ED2708DB2243684EFF16F80BDDB6CB9B" box="[177,199,1945,1971]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="62" pageNumber="63" type="Museum">V</collectionCode>
</materialsCitation>
.
<collectorName id="26C3F5C82243684EFF71F80ADC5CCB9A" box="[214,301,1944,1970]" pageId="62" pageNumber="63">Shavrin</collectorName>
leg. (
<collectionCode id="ED2708DB2243684EFED2F80BDCB9CB9B" box="[373,456,1945,1971]" pageId="62" pageNumber="63">MSPU</collectionCode>
);
<materialsCitation id="3B5E9A432243684EFE78F80BDF00CBFF" collectingDate="1986-07-21" collectionCode="V" county="Khasanskiy District" location="Cheremshan" municipality="Cheryomukhovaya River" pageId="62" pageNumber="63" specimenCount="1" specimenCount-male="1" stateProvince="MARITIME PROVINCE">
<specimenCount id="9D305B972243684EFE78F80BDF60CB9B" box="[479,529,1945,1971]" count="1" pageId="62" pageNumber="63" type="male">1 ♂</specimenCount>
:
<collectingCounty id="62E8E8922243684EFD86F80ADE77CB9A" box="[545,774,1944,1970]" pageId="62" pageNumber="63">Khasanskiy District</collectingCounty>
,
<collectingMunicipality id="6BED0A642243684EFCB1F80AD947CB9A" box="[790,1078,1944,1970]" pageId="62" pageNumber="63">Cheryomukhovaya River</collectingMunicipality>
,
<quantity id="4CCE3DFB2243684EFBE1F80AD9E0CB9A" box="[1094,1169,1944,1971]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="62" pageNumber="63" unit="km" value="15.0">15 km</quantity>
down of
<location id="8EE9C6C52243684EFAAEF80AD8E9CB9A" LSID="urn:lsid:plazi:treatment:039F21082243684FFF30F935DE28CE31:8EE9C6C52243684EFAAEF80AD8E9CB9A" box="[1289,1432,1944,1970]" county="Khasanskiy District" municipality="Cheryomukhovaya River" name="Cheremshan" pageId="62" pageNumber="63" stateProvince="MARITIME PROVINCE">Cheremshan</location>
,
<location id="8EE9C6C52243684EFF30F82EDC63CBFE" LSID="urn:lsid:plazi:treatment:039F21082243684FFF30F935DE28CE31:8EE9C6C52243684EFF30F82EDC63CBFE" box="[151,274,1980,2006]" county="Khasanskiy District" municipality="Cheryomukhovaya River" name="Kedrovaya" pageId="62" pageNumber="63" stateProvince="MARITIME PROVINCE">Kedrovaya</location>
Pad`
<location id="8EE9C6C52243684EFEF9F82FDCC4CBFF" LSID="urn:lsid:plazi:treatment:039F21082243684FFF30F935DE28CE31:8EE9C6C52243684EFEF9F82FDCC4CBFF" box="[350,437,1981,2007]" county="Khasanskiy District" municipality="Cheryomukhovaya River" name="Stream." pageId="62" pageNumber="63" stateProvince="MARITIME PROVINCE">Stream.</location>
<date id="FF88B6DE2243684EFE66F82EDF3CCBFE" box="[449,589,1980,2006]" pageId="62" pageNumber="63" value="1986-07-21">
<collectingDate id="EFCC4F362243684EFE66F82EDF3CCBFE" box="[449,589,1980,2006]" pageId="62" pageNumber="63" value="1986-07-21">21.VII.1986</collectingDate>
</date>
.
<collectionCode id="ED2708DB2243684EFDFAF82FDF00CBFF" box="[605,625,1981,2007]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="62" pageNumber="63" type="Museum">V</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B5E9A432243684EFD26F82EDE43CBFF" box="[641,818,1980,2007]" collectionCode="V" location="Zherikhin" pageId="62" pageNumber="63" specimenCount="1" stateProvince="MARITIME PROVINCE">
<location id="8EE9C6C52243684EFD26F82EDF9ECBFE" LSID="urn:lsid:plazi:treatment:039F21082243684FFF30F935DE28CE31:8EE9C6C52243684EFD26F82EDF9ECBFE" box="[641,751,1980,2006]" name="Zherikhin" pageId="62" pageNumber="63" stateProvince="MARITIME PROVINCE">Zherikhin</location>
&amp;
<collectionCode id="ED2708DB2243684EFCB9F82FDE43CBFF" box="[798,818,1981,2007]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="62" pageNumber="63" type="Museum">V</collectionCode>
</materialsCitation>
. Grachev leg. (cIE);
<materialsCitation id="3B5E9A432243684EFB96F82FDF42CBD3" collectingDate="2009-09-18" collectorName="S. A. Shabalin" location="Lazo" municipality="Lazovskiy District" pageId="62" pageNumber="63" specimenCount="1" specimenCount-male="1" stateProvince="MARITIME PROVINCE">
<specimenCount id="9D305B972243684EFB96F82FD912CBFF" box="[1073,1123,1981,2007]" count="1" pageId="62" pageNumber="63" type="male">1 ♂</specimenCount>
:
<collectingMunicipality id="6BED0A642243684EFBD2F82ED83CCBFE" box="[1141,1357,1980,2006]" pageId="62" pageNumber="63">Lazovskiy District</collectingMunicipality>
,
<location id="8EE9C6C52243684EFAFAF82FD8E9CBFF" LSID="urn:lsid:plazi:treatment:039F21082243684FFF30F935DE28CE31:8EE9C6C52243684EFAFAF82FD8E9CBFF" box="[1373,1432,1981,2007]" municipality="Lazovskiy District" name="Lazo" pageId="62" pageNumber="63" stateProvince="MARITIME PROVINCE">Lazo</location>
,
<date id="FF88B6DE2243684EFF30F873DC6BCBD3" box="[151,282,2017,2043]" pageId="62" pageNumber="63" value="2009-09-18">
<collectingDate id="EFCC4F362243684EFF30F873DC6BCBD3" box="[151,282,2017,2043]" pageId="62" pageNumber="63" value="2009-09-18">18.IX.2009</collectingDate>
</date>
.
<collectorName id="26C3F5C82243684EFE82F873DCB1CBD2" box="[293,448,2016,2043]" pageId="62" pageNumber="63">S.A. Shabalin</collectorName>
leg. (cIE)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32CC3952242684FFF60FF04D972CCFD" pageId="63" pageNumber="64" type="distribution">
<paragraph id="8B89901E2242684FFF60FF04D972CCFD" blockId="63.[151,1437,150,537]" pageId="63" pageNumber="64">
<emphasis id="B9424C0C2242684FFF60FF04DC10CC98" bold="true" box="[199,353,150,176]" pageId="63" pageNumber="64">Distribution.</emphasis>
The eastern Palaearctic
<taxonomicName id="4C36EB9D2242684FFD2CFF05DE83CC99" authorityName="Pace" authorityYear="2003" box="[651,1010,151,177]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="zhagaensis">
<emphasis id="B9424C0C2242684FFD2CFF05DE67CC99" box="[651,790,151,177]" italics="true" pageId="63" pageNumber="64">Gyrophaena</emphasis>
(s.str.)
<emphasis id="B9424C0C2242684FFCD0FF05DE83CC99" box="[887,1010,151,177]" italics="true" pageId="63" pageNumber="64">zhagaensis</emphasis>
</taxonomicName>
is known from Far Eastern
<collectingCountry id="F321D08E2242684FFAF5FF05D8ECCC99" box="[1362,1437,151,177]" name="Russia" pageId="63" pageNumber="64">Russia</collectingCountry>
(
<collectingRegion id="49F25EFC2242684FFF38FF29DC05CCFD" box="[159,372,187,213]" country="Russia" name="Primorskiy" pageId="63" pageNumber="64">Maritime Province</collectingRegion>
(
<figureCitation id="130D8C9B2242684FFE24FF29DC95CCFD" box="[387,484,187,213]" captionStart="FIGURES 305–311" captionStartId="70.[151,264,1784,1808]" captionTargetBox="[151,1436,181,1760]" captionTargetId="figure-21@70.[151,1436,181,1760]" captionTargetPageId="70" captionText="FIGURES 305–311. Distribution of Gyrophaena in Eastern Siberia and Far Eastern Russia: 305—G. (s.str.) pseudonana, 306—G. (s.str.) pseudonitidula, 307—G. (s.str.) pulchella, 308—G. (s.str.) triquetra, 309—G. (s.str.) williamsi, 310—G. (s.str.) zhagaensis, 311—G. (Phaenogyra) strictula." figureDoi="http://doi.org/10.5281/zenodo.14023790" httpUri="https://zenodo.org/record/14023790/files/figure.png" pageId="63" pageNumber="64">Fig. 310</figureCitation>
)),
<collectingCountry id="F321D08E2242684FFDA5FF29DF34CCFD" box="[514,581,187,213]" name="China" pageId="63" pageNumber="64">China</collectingCountry>
(
<collectingRegion id="49F25EFC2242684FFDF4FF29DFEDCCFD" box="[595,668,187,213]" country="China" name="Hubei" pageId="63" pageNumber="64">Hubei</collectingRegion>
,
<collectingRegion id="49F25EFC2242684FFD00FF29DE75CCFD" box="[679,772,187,213]" country="China" name="Shaanxi" pageId="63" pageNumber="64">Shaanxi</collectingRegion>
,
<collectingRegion id="49F25EFC2242684FFCA8FF29DE1ACCFD" box="[783,875,187,213]" country="China" name="Sichuan" pageId="63" pageNumber="64">Sichuan</collectingRegion>
) and
<collectingCountry id="F321D08E2242684FFC0EFF29DE8ECCFD" box="[937,1023,187,213]" name="Taiwan" pageId="63" pageNumber="64">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C32CC3952242684FFF60FF4DDE28CE31" pageId="63" pageNumber="64" type="discussion">
<paragraph id="8B89901E2242684FFF60FF4DDC4ACD85" blockId="63.[151,1437,150,537]" pageId="63" pageNumber="64">
<emphasis id="B9424C0C2242684FFF60FF4DDC45CCD1" bold="true" box="[199,308,223,249]" pageId="63" pageNumber="64">Remarks</emphasis>
.
<taxonomicName id="4C36EB9D2242684FFEE2FF4DDFAACCD1" authority="Pace, 2003" authorityName="Pace" authorityYear="2003" box="[325,731,223,249]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="zhagaensis">
<emphasis id="B9424C0C2242684FFEE2FF4DDF25CCD1" box="[325,596,223,249]" italics="true" pageId="63" pageNumber="64">Gyrophaena zhagaensis</emphasis>
<bibRefCitation id="EFA7EDEF2242684FFDF9FF4DDFAACCD1" author="Pace, R." box="[606,731,223,249]" pageId="63" pageNumber="64" pagination="621 - 660" refId="ref52519" refString="Pace, R. (2003) Gyrophaenini della Cina (Coleoptera, Staphylinidae). Revue Suisse de Zoologie, 110 (3), 621 - 660. https: // doi. org / 10.5962 / bhl. part. 80203" type="journal article" year="2003">Pace, 2003</bibRefCitation>
</taxonomicName>
was originally described from
<collectingCountry id="F321D08E2242684FFBE1FF4DD9F8CCD1" box="[1094,1161,223,249]" name="China" pageId="63" pageNumber="64">China</collectingCountry>
(
<collectingRegion id="49F25EFC2242684FFB3DFF4DD988CCD1" box="[1178,1273,223,249]" country="China" name="Sichuan" pageId="63" pageNumber="64">Sichuan</collectingRegion>
,
<collectingRegion id="49F25EFC2242684FFAA0FF4DD812CCD1" box="[1287,1379,223,249]" country="China" name="Shaanxi" pageId="63" pageNumber="64">Shaanxi</collectingRegion>
) and was compared with the Nepalese
<taxonomicName id="4C36EB9D2242684FFDACFE91DE66CD35" authority="Pace, 1989" authorityName="Pace" authorityYear="1989" box="[523,791,259,285]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="narendra">
<emphasis id="B9424C0C2242684FFDACFE91DFE2CD35" box="[523,659,259,285]" italics="true" pageId="63" pageNumber="64">G. narendra</emphasis>
<bibRefCitation id="EFA7EDEF2242684FFD3EFE91DE66CD35" author="Pace, R." box="[665,791,259,285]" pageId="63" pageNumber="64" pagination="483 - 539" refId="ref52378" refString="Pace, R. (1989) Aleocharinae nepalesi del Museo du Ginevra: Parte I (Coleoptera, Staphylinidae) (104 º contributo alla conoscenza delle Aleocharinae). Revue suisse de Zoologie, 96, 483 - 539. https: // doi. org / 10.5962 / bhl. part. 82045" type="journal article" year="1989">Pace, 1989</bibRefCitation>
</taxonomicName>
. Later,
<bibRefCitation id="EFA7EDEF2242684FFCCFFE91DE9FCD35" author="Pace, R." box="[872,1006,259,285]" pageId="63" pageNumber="64" pagination="125 - 193" refId="ref52707" refString="Pace, R. (2010) Biodiversita delle Aleocharinae della Cina: Gyrophaenini (Coleoptera, Staphylinidae). Beitrage zur Entomologie, 60 (1), 125 - 193. https: // doi. org / 10.21248 / contrib. entomol. 60.1.125 - 193" type="journal article" year="2010">Pace (2010)</bibRefCitation>
recorded
<taxonomicName id="4C36EB9D2242684FFBFBFE91D989CD35" authorityName="Pace" authorityYear="2003" box="[1116,1272,259,285]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="zhagaensis">
<emphasis id="B9424C0C2242684FFBFBFE91D989CD35" box="[1116,1272,259,285]" italics="true" pageId="63" pageNumber="64">G. zhagaensis</emphasis>
</taxonomicName>
from
<collectingRegion id="49F25EFC2242684FFA9CFE91D8E9CD35" box="[1339,1432,259,285]" country="China" name="Sichuan" pageId="63" pageNumber="64">Sichuan</collectingRegion>
,
<collectingRegion id="49F25EFC2242684FFF30FEB5DDAACD69" box="[151,219,295,321]" country="China" name="Hubei" pageId="63" pageNumber="64">Hubei</collectingRegion>
and
<collectingRegion id="49F25EFC2242684FFEB0FEB5DC04CD69" box="[279,373,295,321]" country="China" name="Shaanxi" pageId="63" pageNumber="64">Shaanxi</collectingRegion>
.
<taxonomicName id="4C36EB9D2242684FFE26FEB5DF1ECD69" authorityName="Pace" authorityYear="2007" box="[385,623,295,321]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="peniculi">
<emphasis id="B9424C0C2242684FFE26FEB5DF1ECD69" box="[385,623,295,321]" italics="true" pageId="63" pageNumber="64">Gyrophaena peniculi</emphasis>
</taxonomicName>
was originally described from
<collectingCountry id="F321D08E2242684FFC7FFEB5D95FCD69" box="[984,1070,295,321]" name="Taiwan" pageId="63" pageNumber="64">Taiwan</collectingCountry>
. The shape of the aedeagus and the male abdominal tergite VIII from Maritime Province (see above) are similar to these in
<taxonomicName id="4C36EB9D2242684FFB1DFED9D828CD4D" authorityName="Pace" authorityYear="2003" box="[1210,1369,331,357]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="zhagaensis">
<emphasis id="B9424C0C2242684FFB1DFED9D828CD4D" box="[1210,1369,331,357]" italics="true" pageId="63" pageNumber="64">G. zhagaensis</emphasis>
</taxonomicName>
(
<figureCitation id="130D8C9B2242684FFACDFED9DDACCDA1" captionStart="FIGURES 77–87" captionStartId="24.[151,264,1846,1870]" captionTargetBox="[157,1430,533,1817]" captionTargetId="figure-165@24.[151,1436,525,1823]" captionTargetPageId="24" captionText="FIGURES 77–87. Gyrophaena (s.str.) melanoceta: 77—habitus (holotype), 78–79—aedeagus, lateral view (holotype), 80— aedeagus, ventral view, 81—paramere, 82–83—male abdominal tergite VIII, 84—male abdominal sternite VIII, 85—female abdominal tergite VIII, 86—female abdominal sternite VIII, 87—spermatheca. Scale bars: 1.0 mm (Fig. 77), 0.1 mm (Figs 78–87)." figureDoi="http://doi.org/10.5281/zenodo.14023722" httpUri="https://zenodo.org/record/14023722/files/figure.png" pageId="63" pageNumber="64">
Figs
<quantity id="4CCE3DFB2242684FFF30FEFDDDACCDA1" box="[151,221,367,393]" metricMagnitude="0" metricUnit="m" metricValue="1.9939" metricValueMax="2.032" metricValueMin="1.9558" pageId="63" pageNumber="64" unit="in" value="78.5" valueMax="80.0" valueMin="77.0">77–80</quantity>
</figureCitation>
in
<bibRefCitation id="EFA7EDEF2242684FFEA5FEFDDC09CDA1" author="Pace, R." box="[258,376,367,393]" pageId="63" pageNumber="64" pagination="621 - 660" refId="ref52519" refString="Pace, R. (2003) Gyrophaenini della Cina (Coleoptera, Staphylinidae). Revue Suisse de Zoologie, 110 (3), 621 - 660. https: // doi. org / 10.5962 / bhl. part. 80203" type="journal article" year="2003">Pace 2003</bibRefCitation>
) and
<taxonomicName id="4C36EB9D2242684FFE10FEFDDF45CDA1" authorityName="Pace" authorityYear="2007" box="[439,564,367,393]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="peniculi">
<emphasis id="B9424C0C2242684FFE10FEFDDF45CDA1" box="[439,564,367,393]" italics="true" pageId="63" pageNumber="64">G. peniculi</emphasis>
</taxonomicName>
(
<figureCitation id="130D8C9B2242684FFDE4FEFDDFB3CDA1" box="[579,706,367,393]" captionStart="FIGURES 77–87" captionStartId="24.[151,264,1846,1870]" captionTargetBox="[157,1430,533,1817]" captionTargetId="figure-165@24.[151,1436,525,1823]" captionTargetPageId="24" captionText="FIGURES 77–87. Gyrophaena (s.str.) melanoceta: 77—habitus (holotype), 78–79—aedeagus, lateral view (holotype), 80— aedeagus, ventral view, 81—paramere, 82–83—male abdominal tergite VIII, 84—male abdominal sternite VIII, 85—female abdominal tergite VIII, 86—female abdominal sternite VIII, 87—spermatheca. Scale bars: 1.0 mm (Fig. 77), 0.1 mm (Figs 78–87)." figureDoi="http://doi.org/10.5281/zenodo.14023722" httpUri="https://zenodo.org/record/14023722/files/figure.png" pageId="63" pageNumber="64">
Figs
<quantity id="4CCE3DFB2242684FFDDBFEFDDFB3CDA1" box="[636,706,367,393]" metricMagnitude="0" metricUnit="m" metricValue="2.0446999999999997" metricValueMax="2.0827999999999998" metricValueMin="2.0065999999999997" pageId="63" pageNumber="64" unit="in" value="80.5" valueMax="82.0" valueMin="79.0">79–82</quantity>
</figureCitation>
in
<bibRefCitation id="EFA7EDEF2242684FFD40FEFDDE11CDA1" author="Pace, R." box="[743,864,367,393]" pageId="63" pageNumber="64" pagination="103 - 129" refId="ref52664" refString="Pace, R. (2007) Le specie dei generi Gyrophaena Mann. e Brachida Muls. &amp; Rey di Taiwan (Coleoptera, Staphylinidae). Bollettino del Museo Civico di Storia di Verona, 31, 103 - 129." type="journal article" year="2007">Pace 2007</bibRefCitation>
). Thus, the latter species is here synonymized with
<taxonomicName id="4C36EB9D2242684FFF30FE01DC45CD85" authorityName="Pace" authorityYear="2003" box="[151,308,403,429]" class="Insecta" family="Staphylinidae" genus="Gyrophaena" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="zhagaensis">
<emphasis id="B9424C0C2242684FFF30FE01DDDACD85" box="[151,171,403,429]" italics="true" pageId="63" pageNumber="64">G</emphasis>
.
<emphasis id="B9424C0C2242684FFF1EFE01DC45CD85" box="[185,308,403,429]" italics="true" pageId="63" pageNumber="64">zhagaensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B89901E2242684FFF60FE25DFCCCDDD" blockId="63.[151,1437,150,537]" pageId="63" pageNumber="64">
Habitus as in
<figureCitation id="130D8C9B2242684FFEF8FE25DCB1CDF9" box="[351,448,439,465]" captionStart="FIGURES 257–264" captionStartId="62.[151,264,1533,1557]" captionTargetBox="[158,1431,158,1511]" captionTargetId="figure-21@62.[151,1435,151,1517]" captionTargetPageId="62" captionText="FIGURES 257–264. Gyrophaena (s.str.) williamsi (Figs 257–259) and G. (s.str.) zhagaensis (Figs 260–264): 257, 260—habitus, 258, 261—aedeagus, lateral view, 262—paramere, 259, 263—male abdominal tergite VIII, 264—male abdominal sternite VIII. Scale bars: 1.0 mm (Figs 257, 260), 0.1 mm (Figs 258–259, 261–264)." figureDoi="http://doi.org/10.5281/zenodo.14023772" httpUri="https://zenodo.org/record/14023772/files/figure.png" pageId="63" pageNumber="64">Fig. 260</figureCitation>
. Aedeagus as in
<figureCitation id="130D8C9B2242684FFDD1FE25DFA7CDF9" box="[630,726,439,465]" captionStart="FIGURES 257–264" captionStartId="62.[151,264,1533,1557]" captionTargetBox="[158,1431,158,1511]" captionTargetId="figure-21@62.[151,1435,151,1517]" captionTargetPageId="62" captionText="FIGURES 257–264. Gyrophaena (s.str.) williamsi (Figs 257–259) and G. (s.str.) zhagaensis (Figs 260–264): 257, 260—habitus, 258, 261—aedeagus, lateral view, 262—paramere, 259, 263—male abdominal tergite VIII, 264—male abdominal sternite VIII. Scale bars: 1.0 mm (Figs 257, 260), 0.1 mm (Figs 258–259, 261–264)." figureDoi="http://doi.org/10.5281/zenodo.14023772" httpUri="https://zenodo.org/record/14023772/files/figure.png" pageId="63" pageNumber="64">Fig. 261</figureCitation>
. Paramere as in
<figureCitation id="130D8C9B2242684FFC2CFE25DE9DCDF9" box="[907,1004,439,465]" captionStart="FIGURES 257–264" captionStartId="62.[151,264,1533,1557]" captionTargetBox="[158,1431,158,1511]" captionTargetId="figure-21@62.[151,1435,151,1517]" captionTargetPageId="62" captionText="FIGURES 257–264. Gyrophaena (s.str.) williamsi (Figs 257–259) and G. (s.str.) zhagaensis (Figs 260–264): 257, 260—habitus, 258, 261—aedeagus, lateral view, 262—paramere, 259, 263—male abdominal tergite VIII, 264—male abdominal sternite VIII. Scale bars: 1.0 mm (Figs 257, 260), 0.1 mm (Figs 258–259, 261–264)." figureDoi="http://doi.org/10.5281/zenodo.14023772" httpUri="https://zenodo.org/record/14023772/files/figure.png" pageId="63" pageNumber="64">Fig. 262</figureCitation>
. Male abdominal tergite VIII as in
<figureCitation id="130D8C9B2242684FFAD7FE25DDB5CDDD" captionStart="FIGURES 257–264" captionStartId="62.[151,264,1533,1557]" captionTargetBox="[158,1431,158,1511]" captionTargetId="figure-21@62.[151,1435,151,1517]" captionTargetPageId="62" captionText="FIGURES 257–264. Gyrophaena (s.str.) williamsi (Figs 257–259) and G. (s.str.) zhagaensis (Figs 260–264): 257, 260—habitus, 258, 261—aedeagus, lateral view, 262—paramere, 259, 263—male abdominal tergite VIII, 264—male abdominal sternite VIII. Scale bars: 1.0 mm (Figs 257, 260), 0.1 mm (Figs 258–259, 261–264)." figureDoi="http://doi.org/10.5281/zenodo.14023772" httpUri="https://zenodo.org/record/14023772/files/figure.png" pageId="63" pageNumber="64">Fig. 263</figureCitation>
. Male abdominal sternite VIII as in
<figureCitation id="130D8C9B2242684FFDFFFE49DFC8CDDD" box="[600,697,475,501]" captionStart="FIGURES 257–264" captionStartId="62.[151,264,1533,1557]" captionTargetBox="[158,1431,158,1511]" captionTargetId="figure-21@62.[151,1435,151,1517]" captionTargetPageId="62" captionText="FIGURES 257–264. Gyrophaena (s.str.) williamsi (Figs 257–259) and G. (s.str.) zhagaensis (Figs 260–264): 257, 260—habitus, 258, 261—aedeagus, lateral view, 262—paramere, 259, 263—male abdominal tergite VIII, 264—male abdominal sternite VIII. Scale bars: 1.0 mm (Figs 257, 260), 0.1 mm (Figs 258–259, 261–264)." figureDoi="http://doi.org/10.5281/zenodo.14023772" httpUri="https://zenodo.org/record/14023772/files/figure.png" pageId="63" pageNumber="64">Fig. 264</figureCitation>
.
</paragraph>
<paragraph id="8B89901E2242684FFF60FE6DDE28CE31" blockId="63.[151,1437,150,537]" box="[199,857,511,537]" pageId="63" pageNumber="64">
The species is here recorded from
<collectingCountry id="F321D08E2242684FFDEEFE6DDFE5CE31" box="[585,660,511,537]" name="Russia" pageId="63" pageNumber="64">Russia</collectingCountry>
for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>
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