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<document id="C873E93AB7B330D5669DCE8262E9F8E5" ID-DOI="10.24199/j.mmv.2003.60.21" ID-ISSN="1447-2554" ID-Zenodo-Dep="12207731" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718987894808" checkinUser="felipe" docAuthor="Mesibov, Robert" docDate="2003" docId="AB398E709D794C13FCABF91F6EC54BF3" docLanguage="en" docName="MemMusVictoria.60.2.197-206.pdf" docOrigin="Memoirs of Museum Victoria 60 (2)" docSource="https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-60-issue-2-2003/pages-207-219/" docStyle="DocumentStyle:5B3B5C052A93BFEEE334AD6097A94E45.2:MemMusVictoria.2003-.journal_article.type1" docStyleId="5B3B5C052A93BFEEE334AD6097A94E45" docStyleName="MemMusVictoria.2003-.journal_article.type1" docStyleVersion="2" docTitle="Dasystigma margaretae Mesibov 2003, comb. nov." docType="treatment" docVersion="2" lastPageNumber="204" masterDocId="5700F6089D7F4C14FF9AFFD06A754D78" masterDocTitle="A new genus of Tasmanian millipedes (Diplopoda: Polydesmida: Dalodesmidae) with unusual spiracles and a mosaic distribution" masterLastPageNumber="206" masterPageNumber="197" pageNumber="203" updateTime="1718988523422" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="07FEEA9E3F98A66AA57775976FB01C3E">A new genus of Tasmanian millipedes (Diplopoda: Polydesmida: Dalodesmidae) with unusual spiracles and a mosaic distribution</mods:title>
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<mods:namePart id="380E68D0760A0A4837CDF1043FE60FD3">Mesibov, Robert</mods:namePart>
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<mods:title id="311B3C9E3FA409E6204BC1AAED9D4A95">Memoirs of Museum Victoria</mods:title>
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<treatment id="AB398E709D794C13FCABF91F6EC54BF3" LSID="urn:lsid:plazi:treatment:AB398E709D794C13FCABF91F6EC54BF3" httpUri="http://treatment.plazi.org/id/AB398E709D794C13FCABF91F6EC54BF3" lastPageId="7" lastPageNumber="204" pageId="6" pageNumber="203">
<subSubSection id="6B8A6CED9D794C12FCABF91F6F4D4B9D" box="[817,1336,1743,1765]" pageId="6" pageNumber="203" type="nomenclature">
<paragraph id="232F3F669D794C12FCABF91F6F4D4B9D" blockId="6.[817,1336,1743,1765]" box="[817,1336,1743,1765]" pageId="6" pageNumber="203">
<heading id="7867880A9D794C12FCABF91F6F4D4B9D" box="[817,1336,1743,1765]" fontSize="9" level="2" pageId="6" pageNumber="203" reason="5">
<taxonomicName id="E49044E59D794C12FCABF91F6EB54B9D" ID-CoL="6CF25" authority="(Jeekel, 1984)" authorityName="Mesibov" authorityYear="2003" baseAuthorityName="Jeekel" baseAuthorityYear="1984" box="[817,1216,1743,1765]" class="Diplopoda" family="Dalodesmidae" genus="Dasystigma" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="203" phylum="Arthropoda" rank="species" species="margaretae" status="comb. nov.">
<emphasis id="11E4E3749D794C12FCABF91F6E5C4B9D" bold="true" box="[817,1065,1743,1765]" italics="true" pageId="6" pageNumber="203">Dasystigma margaretae</emphasis>
(
<bibRefCitation id="470142979D794C12FBADF91F6ECC4B9D" author="Jeekel, C. A. W." box="[1079,1209,1743,1765]" pageId="6" pageNumber="203" pagination="85 - 102" refId="ref6748" refString="Jeekel, C. A. W. 1984. Millipedes from Australia, 7: The identity of the genus Lissodesmus Chamberlin, with the description of four new species from Tasmania (Diplopoda, Polydesmida, Dalodesmidae). Papers and Proceedings of the Royal Society of Tasmania 118: 85 - 102." type="journal article" year="1984">Jeekel, 1984</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="0AD75E0F9D794C12FB5DF91F6F4D4B9D" box="[1223,1336,1743,1765]" pageId="6" pageNumber="203" rank="species">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6B8A6CED9D794C12FCABF92B6F7A4A69" box="[817,1295,1787,1809]" pageId="6" pageNumber="203" type="description">
<paragraph id="232F3F669D794C12FCABF92B6F7A4A69" blockId="6.[817,1295,1787,1809]" box="[817,1295,1787,1809]" pageId="6" pageNumber="203">
<figureCitation id="BBAB23E39D794C12FCABF92B69E44A69" box="[817,913,1787,1809]" captionStart="Figure 1" captionStartId="1.[131,190,1834,1854]" captionTargetBox="[129,772,1182,1816]" captionTargetId="figure-519@1.[129,772,1182,1816]" captionTargetPageId="1" captionText="Figure 1. Whole-animal view of Dasystigma margaretae (Jeekel, 1984) comb. nov. A, Rocka Rivulet male, QVM 23:24949; B, Tiger Creek male, QVM 23:24950. Scale bar = 10 mm." pageId="6" pageNumber="203">Figures 1</figureCitation>
,
<figureCitation id="BBAB23E39D794C12FC04F92B699F4A69" box="[926,1002,1787,1809]" captionStart="Figure 3" captionStartId="2.[139,198,602,622]" captionTargetBox="[329,1300,224,582]" captionTargetId="figure-411@2.[307,1327,220,584]" captionTargetPageId="2" captionText="Figure 3. Dorsal views of right margin of tergite 14 of Dasystigma females. A, D. bonhami sp. nov., Sand R., QVM 23:40811; B, D. huonense sp. nov., Edwards Rd, QVM 23:15269; C, D. margaretae (Jeekel, 1984) comb. nov., Fingal Tier, QVM 23:15230; D, D. margaretae (Jeekel, 1984) comb. nov., Tarraleah, QVM 23:15252; E, D. tyleri sp. nov., Trackham Creek, QVM 23:15260. PCS = posterior corner seta." pageId="6" pageNumber="203">3C, 3D</figureCitation>
,
<figureCitation id="BBAB23E39D794C12FC6FF92B6E714A69" box="[1013,1028,1787,1809]" captionStart="Figure 4" captionStartId="2.[820,881,1788,1808]" captionTargetBox="[943,1334,909,1769]" captionTargetId="figure-485@2.[890,1353,902,1770]" captionTargetPageId="2" captionText="Figure 4. Mesal view of right gonopod of male holotype of Dasystigma margaretae (Jeekel, 1984) comb. nov., redrawn from Jeekel (1984). f = femoral process, pf = prefemoral process, s = solenomerite, tt = tibiotarsus, u = uncus. Dashed line marks path of prostatic groove." pageId="6" pageNumber="203">4</figureCitation>
,
<figureCitation id="BBAB23E39D794C12FB95F92B6E2E4A69" box="[1039,1115,1787,1809]" captionStart="Figure 5" captionStartId="3.[131,190,1759,1779]" captionTargetBox="[204,1376,221,1717]" captionTargetId="graphics-7@3.[204,1443,173,1211]" captionTargetPageId="3" captionText="Figure 5. Spiracles on midbody segment of A, Dasystigma bonhami sp. nov., Sandspit R. male, QVM 23:15219; B, D. huonense sp. nov., Huon R. (Arve Rd) male paratype, QVM 23:40805; C, D. margaretae (Jeekel, 1984) comb. nov., Tooms Lake male, QVM 23:15214; D, D. margaretae (Jeekel, 1984), Dromedary Creek male, QVM 23:15212; E, D. tyleri sp. nov., White Spur male, QVM 23:15190. Scale bar in all cases = 0.25 mm; anterior to right for AD, anterior to left for E." pageId="6" pageNumber="203">5C, 5D</figureCitation>
,
<figureCitation id="BBAB23E39D794C12FBFCF92B6EF44A69" box="[1126,1153,1787,1809]" captionStart="Figure 10" captionStartId="5.[131,190,723,743]" captionTargetBox="[130,1449,221,705]" captionTargetId="figure-651@5.[574,1006,220,704]" captionTargetPageId="5" captionText="Figure 10. Dasystigma margaretae (Jeekel, 1984) comb. nov. Gonopods in situ, scale bar = 0.5 mm in all cases. A, Tooms Lake male, QVM 23:15214; B, Lake Augusta male paratype, (DPI-NT) 19A17; C, Coles Creek male (“Derwent form”), QVM 23:24932." pageId="6" pageNumber="203">10</figureCitation>
,
<figureCitation id="BBAB23E39D794C12FB17F92B6ED24A69" box="[1165,1191,1787,1809]" captionStart="Figure 11" captionStartId="6.[139,200,942,962]" captionTargetBox="[191,706,225,922]" captionTargetId="figure-554@6.[153,740,220,924]" captionTargetPageId="6" captionText="Figure 11. Dasystigma margaretae (Jeekel, 1984) comb. nov. Approximately mesal views of left gonopod telopodite of (left) Lake Augusta male paratype (DPI-NT) 19A17 and (right) Tarraleah male (“Derwent form”), QVM 23:15203. Setation not shown." pageId="6" pageNumber="203">11</figureCitation>
,
<figureCitation id="BBAB23E39D794C12FB29F92B6EB94A69" box="[1203,1228,1787,1809]" captionStart="Figure 14" captionStartId="8.[820,879,1736,1756]" captionTargetBox="[821,1466,913,1697]" captionTargetId="figure-704@8.[813,1471,903,1704]" captionTargetPageId="8" captionText="Figure 14. Known localities (to Jun 2002). Dasystigma bonhami sp. nov., s. D. huonense sp. nov., l. D. margaretae (Jeekel, 1984) comb. nov., n. D.margaretae (Jeekel, 1984) comb. nov. “Derwent form”, nn. D. tyleri sp. nov., +.Unidentified Dasystigma species, •. Polygons have been drawn through outermost known localities of each of the four named species to more clearly delineate the species ranges. Arrow marks Little Quoin (see text)." pageId="6" pageNumber="203">14</figureCitation>
(map)
</paragraph>
</subSubSection>
<subSubSection id="6B8A6CED9D794C12FCD6F8F56EBA4A41" box="[844,1231,1829,1849]" pageId="6" pageNumber="203" type="reference_group">
<paragraph id="232F3F669D794C12FCD6F8F56EBA4A41" blockId="6.[844,1231,1829,1849]" box="[844,1231,1829,1849]" pageId="6" pageNumber="203">
<treatmentCitationGroup id="038018489D794C12FCD6F8F56EBA4A41" box="[844,1231,1829,1849]" pageId="6" pageNumber="203">
<treatmentCitation id="A23119779D794C12FCD6F8F56EBF4A41" author="Jeekel, C. A. W." box="[844,1226,1829,1849]" page="99" pageId="6" pageNumber="203" year="1984">
<taxonomicName id="E49044E59D794C12FCD6F8F56EBF4A41" ID-CoL="6QL3R" authority="Jeekel, 1984: 99" authorityName="Jeekel" authorityPageNumber="99" authorityYear="1984" box="[844,1226,1829,1849]" class="Diplopoda" family="Dalodesmidae" genus="Lissodesmus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="203" phylum="Arthropoda" rank="species" species="margaretae">
<emphasis id="11E4E3749D794C12FCD6F8F56E584A41" box="[844,1069,1829,1849]" italics="true" pageId="6" pageNumber="203">Lissodesmus margaretae</emphasis>
<bibRefCitation id="470142979D794C12FBA9F8F56EBF4A41" author="Jeekel, C. A. W." box="[1075,1226,1829,1849]" pageId="6" pageNumber="203" pagination="85 - 102" refId="ref6748" refString="Jeekel, C. A. W. 1984. Millipedes from Australia, 7: The identity of the genus Lissodesmus Chamberlin, with the description of four new species from Tasmania (Diplopoda, Polydesmida, Dalodesmidae). Papers and Proceedings of the Royal Society of Tasmania 118: 85 - 102." type="journal article" year="1984">Jeekel, 1984: 99</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="6B8A6CED9D794C13FCABF89D6EC54BF3" lastPageId="7" lastPageNumber="204" pageId="6" pageNumber="203" type="materials_examined">
<paragraph id="232F3F669D794C13FCABF89D6B674F4C" blockId="6.[817,1458,1869,1916]" lastBlockId="7.[131,772,219,830]" lastPageId="7" lastPageNumber="204" pageId="6" pageNumber="203">
<emphasis id="11E4E3749D794C12FCABF89D69914A19" box="[817,996,1869,1889]" italics="true" pageId="6" pageNumber="203">Material examined</emphasis>
.
<materialsCitation id="93F8353B9D794C13FC6DF89D6B7B4F4C" collectingDate="1979-04-25" collectorName="When I &amp; The &amp; After &amp; McCartney &amp; New Town Laboratories &amp; Water &amp; Environment" country="Australia" county="New Town" lastPageId="7" lastPageNumber="204" location="Department of Agriculture" municipality="Tasmanian Department of Primary Industries" pageId="6" pageNumber="203" specimenCount="3" stateProvince="Tasmania" typeStatus="holotype">
<typeStatus id="FC2B81C49D794C12FC6DF89D6E3E4A19" box="[1015,1099,1869,1889]" pageId="6" pageNumber="203" type="holotype">Holotype</typeStatus>
and
<typeStatus id="FC2B81C49D794C12FB1DF89D6E944A19" box="[1159,1249,1869,1889]" pageId="6" pageNumber="203" type="paratype">paratypes</typeStatus>
.
<collectingCountry id="5B877FF69D794C12FB75F89D6F314A19" box="[1263,1348,1869,1889]" name="Australia" pageId="6" pageNumber="203">Australia</collectingCountry>
,
<collectingRegion id="E154F1849D794C12FACFF89D6FDA4A19" box="[1365,1455,1869,1889]" country="Australia" name="Tasmania" pageId="6" pageNumber="203">Tasmania</collectingRegion>
. “Lake Augusta,
<date id="572E19A69D794C12FC43F8B86E344A04" box="[985,1089,1896,1916]" pageId="6" pageNumber="203" value="1979-04-25">
<collectingDate id="476AE04E9D794C12FC43F8B86E344A04" box="[985,1089,1896,1916]" pageId="6" pageNumber="203" value="1979-04-25">25.IV.1979</collectingDate>
</date>
[on cushion plant]
<date id="572E19A69D794C12FA83F8B86FDB4A04" box="[1305,1454,1896,1916]" pageId="6" pageNumber="203" value="1979-04-25">
<collectingDate id="476AE04E9D794C12FA83F8B86FDB4A04" box="[1305,1454,1896,1916]" pageId="6" pageNumber="203" value="1979-04-25">25 April 1979</collectingDate>
</date>
, Tasmanian Department of Agriculture 19A17,
<typeStatus id="FC2B81C49D784C13FDC9FF3068D04D8C" box="[595,677,224,244]" pageId="7" pageNumber="204" type="holotype">holotype</typeStatus>
, 6, 1 + O (fragm.),
<specimenCount id="3596F4EF9D784C13FF43FF2A6B7E4C76" box="[217,267,250,270]" count="2" pageId="7" pageNumber="204" type="generic">2 juv.</specimenCount>
O + (19 somites),
<specimenCount id="3596F4EF9D784C13FE01FF2A6BBB4C76" box="[411,462,250,270]" count="1" pageId="7" pageNumber="204" type="generic">1 juv.</specimenCount>
O + (18 somites)
<typeStatus id="FC2B81C49D784C13FDCDFF2A68C54C76" box="[599,688,250,270]" pageId="7" pageNumber="204" type="paratype">paratypes</typeStatus>
” (
<bibRefCitation id="470142979D784C13FD59FF2A6AA24C51" author="Jeekel, C. A. W." pageId="7" pageNumber="204" pagination="85 - 102" refId="ref6748" refString="Jeekel, C. A. W. 1984. Millipedes from Australia, 7: The identity of the genus Lissodesmus Chamberlin, with the description of four new species from Tasmania (Diplopoda, Polydesmida, Dalodesmidae). Papers and Proceedings of the Royal Society of Tasmania 118: 85 - 102." type="journal article" year="1984">Jeekel, 1984: 99</bibRefCitation>
).
<collectorName id="8E655AB09D784C13FF7DFEC56B594C51" box="[231,300,277,297]" pageId="7" pageNumber="204">When I</collectorName>
inspected the type-containing vial in 2001, I found two cotton-plugged genitalia tubes and (under a cotton pad) a number of body fragments.
<collectorName id="8E655AB09D784C13FEA4FE9A6B144C26" box="[318,353,330,350]" pageId="7" pageNumber="204">The</collectorName>
genitalia tubes contained the male
<typeStatus id="FC2B81C49D784C13FD2EFE9A69764C26" box="[692,771,330,350]" pageId="7" pageNumber="204" type="holotype">holotype</typeStatus>
and the fragmented mature female
<typeStatus id="FC2B81C49D784C13FE5AFEB568654C01" box="[448,528,357,377]" pageId="7" pageNumber="204" type="paratype">paratype</typeStatus>
, respectively, both in good condition.
<collectorName id="8E655AB09D784C13FF79FE506B664CEC" box="[227,275,384,404]" pageId="7" pageNumber="204">After</collectorName>
removing a
<typeStatus id="FC2B81C49D784C13FE1CFE506BA14CEC" box="[390,468,384,404]" pageId="7" pageNumber="204" type="paratype">paratype</typeStatus>
male segment 7 for SEM examination, I placed the remaining body fragments in a small cotton-plugged glass tube.
<collectorName id="8E655AB09D784C13FEA3FE656B294CB1" box="[313,348,437,457]" pageId="7" pageNumber="204">The</collectorName>
three tubes and all accompanying labels were then sealed in an alcohol-filled
<collectorName id="8E655AB09D784C13FE3EFE00687F4C9C" box="[420,522,464,484]" pageId="7" pageNumber="204">McCartney</collectorName>
vial for continuing storage at the
<collectorName id="8E655AB09D784C13FF53FE3A6BC14C86" box="[201,436,490,510]" pageId="7" pageNumber="204">
<collectingCounty id="CA4E47EA9D784C13FF53FE3A6B414C86" box="[201,308,490,510]" pageId="7" pageNumber="204">New Town</collectingCounty>
Laboratories
</collectorName>
of the
<collectingMunicipality id="C34BA51C9D784C13FD93FE3A6B5A4F61" pageId="7" pageNumber="204">Tasmanian Department of Primary Industries</collectingMunicipality>
,
<collectorName id="8E655AB09D784C13FEA2FDD56B1B4F61" box="[312,366,517,537]" pageId="7" pageNumber="204">Water</collectorName>
and
<collectorName id="8E655AB09D784C13FE01FDD568644F61" box="[411,529,517,537]" pageId="7" pageNumber="204">Environment</collectorName>
(formerly the
<location id="264F69BD9D784C13FD02FDD56B7D4F4C" LSID="urn:lsid:plazi:treatment:AB398E709D794C13FCABF91F6EC54BF3:264F69BD9D784C13FD02FDD56B7D4F4C" country="Australia" county="New Town" municipality="Tasmanian Department of Primary Industries" name="Department of Agriculture" pageId="7" pageNumber="204" stateProvince="Tasmania">Department of Agriculture</location>
)
</materialsCitation>
.
</paragraph>
<paragraph id="232F3F669D784C13FF04FDEA6B774E46" blockId="7.[131,772,219,830]" pageId="7" pageNumber="204">
Other material.
<materialsCitation id="93F8353B9D784C13FEACFDEA6A8B4E46" collectorName="Blackman R. &amp; Florentine R. &amp; Little Florentine R. &amp; Pinnacles Creek, R" county="Tooms White Gum Reserve" location="Anglers Creek" municipality="Tooms Lake" pageId="7" pageNumber="204" specimenCount="235" specimenCount-female="66" specimenCount-juvenile="72" specimenCount-male="97">
<specimenCount id="3596F4EF9D784C13FEACFDEA6BFB4F36" box="[310,398,570,590]" count="97" pageId="7" pageNumber="204" type="male">97 males</specimenCount>
,
<specimenCount id="3596F4EF9D784C13FE06FDEA68764F36" box="[412,515,570,590]" count="66" pageId="7" pageNumber="204" type="female">66 females</specimenCount>
and
<specimenCount id="3596F4EF9D784C13FDA3FDEA68D94F36" box="[569,684,570,590]" count="72" pageId="7" pageNumber="204" type="juvenile">72 juveniles</specimenCount>
from 52 unique localities including Alberts Marsh,
<location id="264F69BD9D784C13FD89FD8568EE4F11" LSID="urn:lsid:plazi:treatment:AB398E709D794C13FCABF91F6EC54BF3:264F69BD9D784C13FD89FD8568EE4F11" box="[531,667,597,617]" county="Tooms White Gum Reserve" municipality="Tooms Lake" name="Anglers Creek" pageId="7" pageNumber="204">Anglers Creek</location>
,
<collectorName id="8E655AB09D784C13FD3DFD856AEF4FFC" pageId="7" pageNumber="204">Blackman R.</collectorName>
,
<location id="264F69BD9D784C13FF33FDA06B324FFC" LSID="urn:lsid:plazi:treatment:AB398E709D794C13FCABF91F6EC54BF3:264F69BD9D784C13FF33FDA06B324FFC" box="[169,327,624,644]" county="Tooms White Gum Reserve" municipality="Tooms Lake" name="Blue Tier Creek" pageId="7" pageNumber="204">Blue Tier Creek</location>
, Boyer,
<location id="264F69BD9D784C13FE07FDA068464FFC" LSID="urn:lsid:plazi:treatment:AB398E709D794C13FCABF91F6EC54BF3:264F69BD9D784C13FE07FDA068464FFC" box="[413,563,624,644]" county="Tooms White Gum Reserve" municipality="Tooms Lake" name="Brumbys Creek" pageId="7" pageNumber="204">Brumbys Creek</location>
, Butlers Gorge, Coal Marsh,
<location id="264F69BD9D784C13FF50FD5A6B494FE6" LSID="urn:lsid:plazi:treatment:AB398E709D794C13FCABF91F6EC54BF3:264F69BD9D784C13FF50FD5A6B494FE6" box="[202,316,650,670]" county="Tooms White Gum Reserve" municipality="Tooms Lake" name="Coles Creek" pageId="7" pageNumber="204">Coles Creek</location>
,
<location id="264F69BD9D784C13FEDDFD5A6B984FE6" LSID="urn:lsid:plazi:treatment:AB398E709D794C13FCABF91F6EC54BF3:264F69BD9D784C13FEDDFD5A6B984FE6" box="[327,493,650,670]" county="Tooms White Gum Reserve" municipality="Tooms Lake" name="Dromedary Creek" pageId="7" pageNumber="204">Dromedary Creek</location>
, Fingal Tier,
<location id="264F69BD9D784C13FDF3FD5A688A4FE6" LSID="urn:lsid:plazi:treatment:AB398E709D794C13FCABF91F6EC54BF3:264F69BD9D784C13FDF3FD5A688A4FE6" box="[617,767,650,670]" county="Tooms White Gum Reserve" municipality="Tooms Lake" name="Flexmore Creek" pageId="7" pageNumber="204">Flexmore Creek</location>
,
<collectorName id="8E655AB09D784C13FF19FD756A884FC1" box="[131,253,677,697]" pageId="7" pageNumber="204">Florentine R.</collectorName>
,
<location id="264F69BD9D784C13FE9FFD756B194FC1" LSID="urn:lsid:plazi:treatment:AB398E709D794C13FCABF91F6EC54BF3:264F69BD9D784C13FE9FFD756B194FC1" box="[261,364,677,697]" county="Tooms White Gum Reserve" municipality="Tooms Lake" name="Gulf Creek" pageId="7" pageNumber="204">Gulf Creek</location>
,
<location id="264F69BD9D784C13FEEFFD756B974FC1" LSID="urn:lsid:plazi:treatment:AB398E709D794C13FCABF91F6EC54BF3:264F69BD9D784C13FEEFFD756B974FC1" box="[373,482,677,697]" county="Tooms White Gum Reserve" municipality="Tooms Lake" name="Halls Creek" pageId="7" pageNumber="204">Halls Creek</location>
,
<location id="264F69BD9D784C13FE71FD7568124FC1" LSID="urn:lsid:plazi:treatment:AB398E709D794C13FCABF91F6EC54BF3:264F69BD9D784C13FE71FD7568124FC1" box="[491,615,677,697]" county="Tooms White Gum Reserve" municipality="Tooms Lake" name="Lake Dobson" pageId="7" pageNumber="204">Lake Dobson</location>
, Liawenee,
<collectorName id="8E655AB09D784C13FD48FD756A8B4FAC" pageId="7" pageNumber="204">Little Florentine R.</collectorName>
, Lookout Hill,
<location id="264F69BD9D784C13FE10FD106B924FAC" LSID="urn:lsid:plazi:treatment:AB398E709D794C13FCABF91F6EC54BF3:264F69BD9D784C13FE10FD106B924FAC" box="[394,487,704,724]" county="Tooms White Gum Reserve" municipality="Tooms Lake" name="Lost Falls" pageId="7" pageNumber="204">Lost Falls</location>
, Meehan Range,
<location id="264F69BD9D784C13FD1CFD106ACE4F96" LSID="urn:lsid:plazi:treatment:AB398E709D794C13FCABF91F6EC54BF3:264F69BD9D784C13FD1CFD106ACE4F96" county="Tooms White Gum Reserve" municipality="Tooms Lake" name="Mossy Marsh Creek" pageId="7" pageNumber="204">Mossy Marsh Creek</location>
,
<location id="264F69BD9D784C13FF52FD0A6BF14F96" LSID="urn:lsid:plazi:treatment:AB398E709D794C13FCABF91F6EC54BF3:264F69BD9D784C13FF52FD0A6BF14F96" box="[200,388,730,750]" county="Tooms White Gum Reserve" municipality="Tooms Lake" name="Mt Mismanagement" pageId="7" pageNumber="204">Mt Mismanagement</location>
, Native Tier, Old Mans Head,
<collectorName id="8E655AB09D784C13FD36FD0A6A954E71" pageId="7" pageNumber="204">
<location id="264F69BD9D784C13FD36FD0A6AC94E71" LSID="urn:lsid:plazi:treatment:AB398E709D794C13FCABF91F6EC54BF3:264F69BD9D784C13FD36FD0A6AC94E71" county="Tooms White Gum Reserve" municipality="Tooms Lake" name="Pinnacles Creek" pageId="7" pageNumber="204">Pinnacles Creek</location>
, R
</collectorName>
. Dee, Rocka Rivulet, Sassafras Hill, St Pauls Dome, Tarraleah,
<location id="264F69BD9D784C13FF7FFCC06B214E5C" LSID="urn:lsid:plazi:treatment:AB398E709D794C13FCABF91F6EC54BF3:264F69BD9D784C13FF7FFCC06B214E5C" box="[229,340,784,804]" county="Tooms White Gum Reserve" municipality="Tooms Lake" name="Tiger Creek" pageId="7" pageNumber="204">Tiger Creek</location>
,
<collectingMunicipality id="C34BA51C9D784C13FEC4FCC06BA74E5C" box="[350,466,784,804]" pageId="7" pageNumber="204">Tooms Lake</collectingMunicipality>
,
<collectingCounty id="CA4E47EA9D784C13FE46FCC068AE4E5C" box="[476,731,784,804]" pageId="7" pageNumber="204">Tooms White Gum Reserve</collectingCounty>
and Yangena Hill
</materialsCitation>
.
</paragraph>
<paragraph id="232F3F669D784C13FF19FC85684A4E85" blockId="7.[131,771,853,1021]" pageId="7" pageNumber="204">
<emphasis id="11E4E3749D784C13FF19FC856A844E13" box="[131,241,853,875]" italics="true" pageId="7" pageNumber="204">Diagnosis.</emphasis>
Differs from
<taxonomicName id="E49044E59D784C13FEE7FC856B834E13" authorityName="Mesibov" authorityYear="2003" box="[381,502,853,875]" class="Diplopoda" family="Dalodesmidae" genus="Dasystigma" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="204" phylum="Arthropoda" rank="species" species="bonhami">
<emphasis id="11E4E3749D784C13FEE7FC856B834E13" box="[381,502,853,875]" italics="true" pageId="7" pageNumber="204">D. bonhami</emphasis>
</taxonomicName>
in having a broader, more flexed and more prominently toothed femoral process; from
<taxonomicName id="E49044E59D784C13FF19FC5F6B704EDD" authorityName="Mesibov" authorityYear="2003" box="[131,261,911,933]" class="Diplopoda" family="Dalodesmidae" genus="Dasystigma" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="204" phylum="Arthropoda" rank="species" species="huonense">
<emphasis id="11E4E3749D784C13FF19FC5F6B704EDD" box="[131,261,911,933]" italics="true" pageId="7" pageNumber="204">D. huonense</emphasis>
</taxonomicName>
and
<taxonomicName id="E49044E59D784C13FEA1FC5F6BE74EDD" authorityName="Mesibov" authorityYear="2003" box="[315,402,911,933]" class="Diplopoda" family="Dalodesmidae" genus="Dasystigma" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="204" phylum="Arthropoda" rank="species" species="tyleri">
<emphasis id="11E4E3749D784C13FEA1FC5F6BE74EDD" box="[315,402,911,933]" italics="true" pageId="7" pageNumber="204">D. tyleri</emphasis>
</taxonomicName>
in having the posterior spiracle on diplosegments located above the anterior leg; from
<taxonomicName id="E49044E59D784C13FD09FC7D689D4EBB" authorityName="Mesibov" authorityYear="2003" box="[659,744,941,963]" class="Diplopoda" family="Dalodesmidae" genus="Dasystigma" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="204" phylum="Arthropoda" rank="species" species="tyleri">
<emphasis id="11E4E3749D784C13FD09FC7D689D4EBB" box="[659,744,941,963]" italics="true" pageId="7" pageNumber="204">D. tyleri</emphasis>
</taxonomicName>
in having much “hairier” spiracles, spiracles also being larger than those in
<taxonomicName id="E49044E59D784C13FF40FC376B5A4E85" authorityName="Mesibov" authorityYear="2003" box="[218,303,999,1021]" class="Diplopoda" family="Dalodesmidae" genus="Dasystigma" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="204" phylum="Arthropoda" rank="species" species="tyleri">
<emphasis id="11E4E3749D784C13FF40FC376B5A4E85" box="[218,303,999,1021]" italics="true" pageId="7" pageNumber="204">D. tyleri</emphasis>
</taxonomicName>
(or in other dalodesmids).
</paragraph>
<paragraph id="232F3F669D784C13FF19FBC368B84899" blockId="7.[131,772,1043,1505]" pageId="7" pageNumber="204">
<emphasis id="11E4E3749D784C13FF19FBC36B774951" box="[131,258,1043,1065]" italics="true" pageId="7" pageNumber="204">Description.</emphasis>
As for the genus except in the following details. Both spiracles on diplosegments (
<figureCitation id="BBAB23E39D784C13FE6CFBE1683F493F" box="[502,586,1073,1095]" captionStart="Figure 5" captionStartId="3.[131,190,1759,1779]" captionTargetBox="[204,1376,221,1717]" captionTargetId="graphics-7@3.[204,1443,173,1211]" captionTargetPageId="3" captionText="Figure 5. Spiracles on midbody segment of A, Dasystigma bonhami sp. nov., Sandspit R. male, QVM 23:15219; B, D. huonense sp. nov., Huon R. (Arve Rd) male paratype, QVM 23:40805; C, D. margaretae (Jeekel, 1984) comb. nov., Tooms Lake male, QVM 23:15214; D, D. margaretae (Jeekel, 1984), Dromedary Creek male, QVM 23:15212; E, D. tyleri sp. nov., White Spur male, QVM 23:15190. Scale bar in all cases = 0.25 mm; anterior to right for AD, anterior to left for E." pageId="7" pageNumber="204">Fig. 5C</figureCitation>
) positioned over anterior leg, hair-like structures emergent from spiracles and apparent at low magnification, spiracles unusually large for a dalodesmid. Paranotal margin slightly convex; posterior corner variably projected (
<figureCitation id="BBAB23E39D784C13FEC4FB766B9049C4" box="[350,485,1190,1212]" captionStart="Figure 3" captionStartId="2.[139,198,602,622]" captionTargetBox="[329,1300,224,582]" captionTargetId="figure-411@2.[307,1327,220,584]" captionTargetPageId="2" captionText="Figure 3. Dorsal views of right margin of tergite 14 of Dasystigma females. A, D. bonhami sp. nov., Sand R., QVM 23:40811; B, D. huonense sp. nov., Edwards Rd, QVM 23:15269; C, D. margaretae (Jeekel, 1984) comb. nov., Fingal Tier, QVM 23:15230; D, D. margaretae (Jeekel, 1984) comb. nov., Tarraleah, QVM 23:15252; E, D. tyleri sp. nov., Trackham Creek, QVM 23:15260. PCS = posterior corner seta." pageId="7" pageNumber="204">Figs 3C, D</figureCitation>
; see
<emphasis id="11E4E3749D784C13FDB1FB7668F049C4" box="[555,645,1190,1212]" italics="true" pageId="7" pageNumber="204">Remarks</emphasis>
). Gonopod telopodite (
<figureCitation id="BBAB23E39D784C13FF63FB136B3F49A1" box="[249,330,1219,1241]" captionStart="Figure 10" captionStartId="5.[131,190,723,743]" captionTargetBox="[130,1449,221,705]" captionTargetId="figure-651@5.[574,1006,220,704]" captionTargetPageId="5" captionText="Figure 10. Dasystigma margaretae (Jeekel, 1984) comb. nov. Gonopods in situ, scale bar = 0.5 mm in all cases. A, Tooms Lake male, QVM 23:15214; B, Lake Augusta male paratype, (DPI-NT) 19A17; C, Coles Creek male (“Derwent form”), QVM 23:24932." pageId="7" pageNumber="204">Figs 10</figureCitation>
,
<figureCitation id="BBAB23E39D784C13FEC2FB136B0449A1" box="[344,369,1219,1241]" captionStart="Figure 11" captionStartId="6.[139,200,942,962]" captionTargetBox="[191,706,225,922]" captionTargetId="figure-554@6.[153,740,220,924]" captionTargetPageId="6" captionText="Figure 11. Dasystigma margaretae (Jeekel, 1984) comb. nov. Approximately mesal views of left gonopod telopodite of (left) Lake Augusta male paratype (DPI-NT) 19A17 and (right) Tarraleah male (“Derwent form”), QVM 23:15203. Setation not shown." pageId="7" pageNumber="204">11</figureCitation>
) with prefemoral process bent mesad, sharply narrowing distad, apex curving caudad, apical edge slightly serrulate, a single tooth on mesal edge of process at about three-quarters of process length. Femoral process curving caudad and mesad and terminating just proximal to level of tooth on mesal edge of prefemoral process. Femoral process massive, divided into a broadly cuneate anterior portion and a posterior spike; distal and posterior margin of cuneate portion with numerous heavy teeth, spike crossing below cuneate portion in manner of a thumb bent slightly towards palm.
</paragraph>
<caption id="77EF6FEE9D784C13FCB1FBFF6E0B4900" pageId="7" pageNumber="204" startId="7.[811,870,1071,1091]" targetBox="[891,1379,224,1050]" targetPageId="7" targetType="figure">
<paragraph id="232F3F669D784C13FCB1FBFF6E0B4900" blockId="7.[811,1449,1071,1144]" pageId="7" pageNumber="204">
Figure 13.
<taxonomicName id="E49044E59D784C13FC0AFBFF6E47493B" authorityName="Mesibov" authorityYear="2003" box="[912,1074,1071,1091]" class="Diplopoda" family="Dalodesmidae" genus="Dasystigma" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="204" phylum="Arthropoda" rank="species" species="tyleri" status="sp. nov.">
<emphasis id="11E4E3749D784C13FC0AFBFF6E47493B" box="[912,1074,1071,1091]" italics="true" pageId="7" pageNumber="204">Dasystigma tyleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0AD75E0F9D784C13FBA3FBFF6EF4493B" box="[1081,1153,1071,1091]" pageId="7" pageNumber="204" rank="species">sp. nov.</taxonomicNameLabel>
Approximately lateral (left) and mesal (right) views of left gonopod telopodite of Franklin R. male, QVM 23:15189. Setation not shown.
</paragraph>
</caption>
<paragraph id="232F3F669D784C13FF19FA276B504B80" blockId="7.[131,771,1527,1784]" pageId="7" pageNumber="204">
<emphasis id="11E4E3749D784C13FF19FA276BCE4B75" box="[131,443,1527,1549]" italics="true" pageId="7" pageNumber="204">Distribution and macrohabitat</emphasis>
. Common in dry and wet eucalypt forest and in subalpine woodland over c. 12 000 km
<quantity id="E46892839D784C13FD45F9C469764B53" box="[735,771,1556,1579]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="7" pageNumber="204" unit="in" value="2.0">
<superScript id="D4E5922E9D784C13FD45F9C468904B67" attach="left" box="[735,741,1556,1567]" fontSize="5" pageId="7" pageNumber="204">2</superScript>
in
</quantity>
eastern and central
<collectingRegion id="E154F1849D784C13FED6F9E26BD84B30" box="[332,429,1586,1608]" country="Australia" name="Tasmania" pageId="7" pageNumber="204">Tasmania</collectingRegion>
(
<figureCitation id="BBAB23E39D784C13FE26F9E2687C4B30" box="[444,521,1586,1608]" captionStart="Figure 14" captionStartId="8.[820,879,1736,1756]" captionTargetBox="[821,1466,913,1697]" captionTargetId="figure-704@8.[813,1471,903,1704]" captionTargetPageId="8" captionText="Figure 14. Known localities (to Jun 2002). Dasystigma bonhami sp. nov., s. D. huonense sp. nov., l. D. margaretae (Jeekel, 1984) comb. nov., n. D.margaretae (Jeekel, 1984) comb. nov. “Derwent form”, nn. D. tyleri sp. nov., +.Unidentified Dasystigma species, •. Polygons have been drawn through outermost known localities of each of the four named species to more clearly delineate the species ranges. Arrow marks Little Quoin (see text)." pageId="7" pageNumber="204">Fig. 14</figureCitation>
), from the south side of the Fingal Valley south to the north side of the Little Swanport R. valley, and from near the east coast west to the Little Florentine R.; so far known at altitudes c.
<quantity id="E46892839D784C13FDCAF95A68A64BD8" box="[592,723,1674,1696]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="11.0" metricValueMin="1.0" pageId="7" pageNumber="204" unit="m" value="600.0" valueMax="1100.0" valueMin="100.0">1001100 m</quantity>
elevation. Locally abundant in places in the Eastern Tiers, on the eastern fringe of the Central Plateau and in wet forests in the Derwent valley.
</paragraph>
<paragraph id="232F3F669D784C13FF19F8DE6EC54BF3" blockId="7.[131,771,1806,1916]" lastBlockId="7.[809,1449,1184,1675]" pageId="7" pageNumber="204">
<emphasis id="11E4E3749D784C13FF19F8DE6A964A5C" box="[131,227,1806,1828]" italics="true" pageId="7" pageNumber="204">Remarks.</emphasis>
<bibRefCitation id="470142979D784C13FF74F8DE6B094A5C" author="Jeekel, C. A. W." box="[238,380,1806,1828]" pageId="7" pageNumber="204" pagination="85 - 102" refId="ref6748" refString="Jeekel, C. A. W. 1984. Millipedes from Australia, 7: The identity of the genus Lissodesmus Chamberlin, with the description of four new species from Tasmania (Diplopoda, Polydesmida, Dalodesmidae). Papers and Proceedings of the Royal Society of Tasmania 118: 85 - 102." type="journal article" year="1984">Jeekel (1984)</bibRefCitation>
gave a complete description of this species from the
<typeStatus id="FC2B81C49D784C13FEA4F8FB6B1F4A39" box="[318,362,1835,1857]" pageId="7" pageNumber="204">type</typeStatus>
locality, Lake Augusta on Tasmanias Central Plateau; for the sake of consistency I have included my own summary. This taxon is the most variable within
<taxonomicName id="E49044E59D784C13FCB3FB7069D449CE" authorityName="Mesibov" authorityYear="2003" box="[809,929,1184,1206]" class="Diplopoda" family="Dalodesmidae" genus="Dasystigma" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="204" phylum="Arthropoda" rank="genus">
<emphasis id="11E4E3749D784C13FCB3FB7069D449CE" box="[809,929,1184,1206]" italics="true" pageId="7" pageNumber="204">Dasystigma</emphasis>
</taxonomicName>
. The prefemoral process is straighter and narrower in the eastern portion of the range (
<figureCitation id="BBAB23E39D784C13FB0DFB6D6E8049AB" box="[1175,1269,1213,1235]" captionStart="Figure 10" captionStartId="5.[131,190,723,743]" captionTargetBox="[130,1449,221,705]" captionTargetId="figure-651@5.[574,1006,220,704]" captionTargetPageId="5" captionText="Figure 10. Dasystigma margaretae (Jeekel, 1984) comb. nov. Gonopods in situ, scale bar = 0.5 mm in all cases. A, Tooms Lake male, QVM 23:15214; B, Lake Augusta male paratype, (DPI-NT) 19A17; C, Coles Creek male (“Derwent form”), QVM 23:24932." pageId="7" pageNumber="204">Fig. 10A</figureCitation>
) than in the west (
<figureCitation id="BBAB23E39D784C13FCAAFB0B69F94989" box="[816,908,1243,1265]" captionStart="Figure 10" captionStartId="5.[131,190,723,743]" captionTargetBox="[130,1449,221,705]" captionTargetId="figure-651@5.[574,1006,220,704]" captionTargetPageId="5" captionText="Figure 10. Dasystigma margaretae (Jeekel, 1984) comb. nov. Gonopods in situ, scale bar = 0.5 mm in all cases. A, Tooms Lake male, QVM 23:15214; B, Lake Augusta male paratype, (DPI-NT) 19A17; C, Coles Creek male (“Derwent form”), QVM 23:24932." pageId="7" pageNumber="204">Fig. 10B</figureCitation>
) and the femoral process less bent mesad. A distinctive variant, here called the “Derwent form,” is restricted to the valley of the Derwent R. (
<figureCitation id="BBAB23E39D784C13FBD1FAC56EEF4853" box="[1099,1178,1301,1323]" captionStart="Figure 14" captionStartId="8.[820,879,1736,1756]" captionTargetBox="[821,1466,913,1697]" captionTargetId="figure-704@8.[813,1471,903,1704]" captionTargetPageId="8" captionText="Figure 14. Known localities (to Jun 2002). Dasystigma bonhami sp. nov., s. D. huonense sp. nov., l. D. margaretae (Jeekel, 1984) comb. nov., n. D.margaretae (Jeekel, 1984) comb. nov. “Derwent form”, nn. D. tyleri sp. nov., +.Unidentified Dasystigma species, •. Polygons have been drawn through outermost known localities of each of the four named species to more clearly delineate the species ranges. Arrow marks Little Quoin (see text)." pageId="7" pageNumber="204">Fig. 14</figureCitation>
), where it replaces more typical
<taxonomicName id="E49044E59D784C13FCEFFAE36E7D4831" authorityName="Jeekel" authorityYear="1984" box="[885,1032,1331,1353]" class="Diplopoda" family="Dalodesmidae" genus="Lissodesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="204" phylum="Arthropoda" rank="species" species="margaretae">
<emphasis id="11E4E3749D784C13FCEFFAE36E7D4831" box="[885,1032,1331,1353]" italics="true" pageId="7" pageNumber="204">D. margaretae</emphasis>
</taxonomicName>
. In the “Derwent form” the spiracles are somewhat less “hairy” (
<figureCitation id="BBAB23E39D784C13FBBCFA806E0F481E" box="[1062,1146,1360,1382]" captionStart="Figure 5" captionStartId="3.[131,190,1759,1779]" captionTargetBox="[204,1376,221,1717]" captionTargetId="graphics-7@3.[204,1443,173,1211]" captionTargetPageId="3" captionText="Figure 5. Spiracles on midbody segment of A, Dasystigma bonhami sp. nov., Sandspit R. male, QVM 23:15219; B, D. huonense sp. nov., Huon R. (Arve Rd) male paratype, QVM 23:40805; C, D. margaretae (Jeekel, 1984) comb. nov., Tooms Lake male, QVM 23:15214; D, D. margaretae (Jeekel, 1984), Dromedary Creek male, QVM 23:15212; E, D. tyleri sp. nov., White Spur male, QVM 23:15190. Scale bar in all cases = 0.25 mm; anterior to right for AD, anterior to left for E." pageId="7" pageNumber="204">Fig. 5D</figureCitation>
) and the prefemoral process somewhat broader (
<figureCitation id="BBAB23E39D784C13FC75FABD6E3B48FB" box="[1007,1102,1389,1411]" captionStart="Figure 10" captionStartId="5.[131,190,723,743]" captionTargetBox="[130,1449,221,705]" captionTargetId="figure-651@5.[574,1006,220,704]" captionTargetPageId="5" captionText="Figure 10. Dasystigma margaretae (Jeekel, 1984) comb. nov. Gonopods in situ, scale bar = 0.5 mm in all cases. A, Tooms Lake male, QVM 23:15214; B, Lake Augusta male paratype, (DPI-NT) 19A17; C, Coles Creek male (“Derwent form”), QVM 23:24932." pageId="7" pageNumber="204">Figs 10C</figureCitation>
,
<figureCitation id="BBAB23E39D784C13FBC2FABD6E0648FB" box="[1112,1139,1389,1411]" captionStart="Figure 11" captionStartId="6.[139,200,942,962]" captionTargetBox="[191,706,225,922]" captionTargetId="figure-554@6.[153,740,220,924]" captionTargetPageId="6" captionText="Figure 11. Dasystigma margaretae (Jeekel, 1984) comb. nov. Approximately mesal views of left gonopod telopodite of (left) Lake Augusta male paratype (DPI-NT) 19A17 and (right) Tarraleah male (“Derwent form”), QVM 23:15203. Setation not shown." pageId="7" pageNumber="204">11</figureCitation>
). The most striking difference, however, is in the form of the paranota: the margins have moved ventrad relative to those in typical
<taxonomicName id="E49044E59D784C13FB44FA786F2348C6" authorityName="Mesibov" authorityYear="2003" box="[1246,1366,1448,1470]" class="Diplopoda" family="Dalodesmidae" genus="Dasystigma" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="204" phylum="Arthropoda" rank="genus">
<emphasis id="11E4E3749D784C13FB44FA786F2348C6" box="[1246,1366,1448,1470]" italics="true" pageId="7" pageNumber="204">Dasystigma</emphasis>
</taxonomicName>
and the posterior corners are very strongly projected caudad and laterad (
<figureCitation id="BBAB23E39D784C13FCE4FA3369A54881" box="[894,976,1507,1529]" captionStart="Figure 3" captionStartId="2.[139,198,602,622]" captionTargetBox="[329,1300,224,582]" captionTargetId="figure-411@2.[307,1327,220,584]" captionTargetPageId="2" captionText="Figure 3. Dorsal views of right margin of tergite 14 of Dasystigma females. A, D. bonhami sp. nov., Sand R., QVM 23:40811; B, D. huonense sp. nov., Edwards Rd, QVM 23:15269; C, D. margaretae (Jeekel, 1984) comb. nov., Fingal Tier, QVM 23:15230; D, D. margaretae (Jeekel, 1984) comb. nov., Tarraleah, QVM 23:15252; E, D. tyleri sp. nov., Trackham Creek, QVM 23:15260. PCS = posterior corner seta." pageId="7" pageNumber="204">Fig. 3D</figureCitation>
). In future, genetic data may justify the erection of a new species for this geographically and morphologically distinctive variant. I am reluctant at this time to name the “Derwent form” formally, as it differs less from more typical
<taxonomicName id="E49044E59D784C13FCE3F9886E674B16" authorityName="Jeekel" authorityYear="1984" box="[889,1042,1624,1646]" class="Diplopoda" family="Dalodesmidae" genus="Lissodesmus" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="204" phylum="Arthropoda" rank="species" species="margaretae">
<emphasis id="11E4E3749D784C13FCE3F9886E674B16" box="[889,1042,1624,1646]" italics="true" pageId="7" pageNumber="204">D. margaretae</emphasis>
</taxonomicName>
in gonopod details than do the three forms recognised here as new species.
</paragraph>
</subSubSection>
</treatment>
</document>