treatments-xml/data/8A/28/87/8A288798FFE0E71F7EC3FF61FCC8C141.xml
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<document ID-DOI="http://dx.doi.org/10.11646/zootaxa.3862.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="4930959" ID-ZooBank="3FA375FE-E4E0-4509-BE02-EE5E786B07C6" approvalRequired="20" approvalRequired_for_treatments="20" checkinTime="1623452360688" checkinUser="felipe" docAuthor="Köhler, Gunther, Pérez, Raúl Gómez Trejo, Petersen, Claus Bo P., Méndez, Fausto R. &amp; Cruz, De La" docDate="2014" docId="8A288798FFE0E71F7EC3FF61FCC8C141" docLanguage="en" docName="zt03862p210.pdf.imf" docOrigin="Zootaxa 3862 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.10:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="10" docTitle="Anolis nietoi Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz 2014, sp. nov." docType="treatment" docVersion="7" lastPageNumber="39" masterDocId="7611FFE0FFC0E7397E54FFF6FFD3C631" masterDocTitle="A revision of the Mexican Anolis (Reptilia, Squamata, Dactyloidae) from the Pacific versant west of the Isthmus de Tehuantepec in the states of Oaxaca, Guerrero, and Puebla, with the description of six new species" masterLastPageNumber="210" masterPageNumber="1" pageNumber="33" updateTime="1673621398369" updateUser="juliana">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the Mexican Anolis (Reptilia, Squamata, Dactyloidae) from the Pacific versant west of the Isthmus de Tehuantepec in the states of Oaxaca, Guerrero, and Puebla, with the description of six new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Köhler, Gunther</mods:namePart>
<mods:nameIdentifier type="email">gkoehler@senckenberg.de</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pérez, Raúl Gómez Trejo</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Petersen, Claus Bo P.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Méndez, Fausto R.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cruz, De La</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="pubDate">
<mods:number>2014-09-19</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>3862</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>210</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.11646/zootaxa.3862.1.1</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">4930959</mods:identifier>
<mods:identifier type="ZooBank">3FA375FE-E4E0-4509-BE02-EE5E786B07C6</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5583550" ID-Zenodo-Dep="5583550" LSID="urn:lsid:plazi:treatment:8A288798FFE0E71F7EC3FF61FCC8C141" httpUri="http://treatment.plazi.org/id/8A288798FFE0E71F7EC3FF61FCC8C141" lastPageId="38" lastPageNumber="39" pageId="32" pageNumber="33">
<subSubSection box="[151,406,151,178]" pageId="32" pageNumber="33" type="nomenclature">
<paragraph blockId="32.[151,406,151,211]" box="[151,406,151,178]" pageId="32" pageNumber="33">
<heading bold="true" box="[151,406,151,178]" fontSize="11" level="2" pageId="32" pageNumber="33" reason="4">
<emphasis bold="true" box="[151,406,151,178]" pageId="32" pageNumber="33">
<taxonomicName authority="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz, 2014" authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[151,305,151,177]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="nietoi" status="sp. nov.">
<emphasis bold="true" box="[151,305,151,177]" italics="true" pageId="32" pageNumber="33">Anolis nietoi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[312,406,152,178]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,283,186,211]" pageId="32" pageNumber="33" type="description">
<paragraph blockId="32.[151,406,151,211]" box="[151,283,186,211]" pageId="32" pageNumber="33">
<figureCitation box="[151,283,186,211]" captionStart-0="FIGURE 20" captionStart-1="FIGURE 21" captionStart-2="FIGURE 22" captionStart-3="FIGURE 23" captionStart-4="FIGURE 24" captionStart-5="FIGURE 25" captionStartId-0="32.[151,250,1968,1990]" captionStartId-1="33.[151,250,1709,1731]" captionStartId-2="34.[151,250,1537,1559]" captionStartId-3="35.[151,250,2000,2022]" captionStartId-4="36.[151,250,1945,1967]" captionStartId-5="37.[151,250,1498,1520]" captionTargetBox-0="[157,1436,658,1943]" captionTargetBox-1="[151,1436,823,1687]" captionTargetBox-2="[153,1436,199,1506]" captionTargetBox-3="[151,1436,651,1957]" captionTargetBox-4="[151,1436,179,1926]" captionTargetBox-5="[151,1426,155,1461]" captionTargetId-0="figure-185@32.[157,1442,658,1943]" captionTargetId-1="figure-386@33.[151,1436,823,1687]" captionTargetId-2="figure-246@34.[153,1438,199,1506]" captionTargetId-3="figure-259@35.[151,1436,651,1957]" captionTargetId-4="figure-71@36.[151,1436,177,1926]" captionTargetId-5="figure-282@37.[141,1426,155,1461]" captionTargetPageId-0="32" captionTargetPageId-1="33" captionTargetPageId-2="34" captionTargetPageId-3="35" captionTargetPageId-4="36" captionTargetPageId-5="37" captionText-0="FIGURE 20. Dewlaps of Anolis nietoi in life. (a) SMF 96406; (b) SMF 96403; (c) SMF 96402; (d) SMF 96407. (ac) are males, (d) is a female." captionText-1="FIGURE 21. Anolis nietoi in life. (a) SMF 96406; (b) IBH 27051; (c) SMF 96398; (d) SMF 96403. (ab) are males, (cd) are females." captionText-2="FIGURE 22. Holotype of Anolis nietoi (SMF 96404): (a) dorsal view; (b) lateral view; (c) ventral view; (d) lateral view of head; (e) dorsal view of head; (f) ventral view of head. Scale bars equal 5.0 mm in (ac) and 1.0 mm in (df), respectively." captionText-3="FIGURE 23. Anolis nietoi: head scalation in (a,c,e) SMF 96406; (b,d,f) SMF 96398. Scale bars equal 1.0 mm." captionText-4="FIGURE 24. Anolis nietoi: nasal region in (a) SMF 96406; (b) SMF 96398; superciliary region in (c) SMF 96403; (d) SMF 96406; chin region in (e) SMF 96403; (f) SMF 96406; cloacal region in (g) SMF 96395; (h) SMF 96390. Scale bars equal 1.0 mm." captionText-5="FIGURE 25. Anolis nietoi: dorsal scales in (a) IBH 26573; (b) SMF 96213; flank scales in (c) SMF 96406; (d) SMF 96398; ventral scales in (e) SMF 96406; (f) SMF 96398. Scale bars equal 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4931003" figureDoi-1="http://doi.org/10.5281/zenodo.4931005" figureDoi-2="http://doi.org/10.5281/zenodo.4931007" figureDoi-3="http://doi.org/10.5281/zenodo.4931009" figureDoi-4="http://doi.org/10.5281/zenodo.4931011" figureDoi-5="http://doi.org/10.5281/zenodo.4931013" httpUri-0="https://zenodo.org/record/4931003/files/figure.png" httpUri-1="https://zenodo.org/record/4931005/files/figure.png" httpUri-2="https://zenodo.org/record/4931007/files/figure.png" httpUri-3="https://zenodo.org/record/4931009/files/figure.png" httpUri-4="https://zenodo.org/record/4931011/files/figure.png" httpUri-5="https://zenodo.org/record/4931013/files/figure.png" pageId="32" pageNumber="33">Figs. 2025</figureCitation>
</paragraph>
</subSubSection>
<subSubSection box="[151,440,258,280]" pageId="32" pageNumber="33" type="reference_group">
<paragraph blockId="32.[151,440,258,280]" box="[151,440,258,280]" pageId="32" pageNumber="33">
<taxonomicName authority=": Ahl 1934" authorityName=": Ahl" authorityYear="1934" box="[151,440,258,280]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="nebuloides">
<emphasis box="[151,332,258,280]" italics="true" pageId="32" pageNumber="33">Anolis nebuloides</emphasis>
:
<bibRefCitation author="Ahl, E." box="[345,440,258,280]" pageId="32" pageNumber="33" pagination="184 - 186" refId="ref87687" refString="Ahl, E. (1934) Uber eine Sammlung von Reptilien und Amphibien aus Mexico. Zoologischer Anzeiger, Leipzig, 106 (7 / 8), 184 - 186." type="journal article" year="1934">Ahl 1934</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="32" pageNumber="33" type="materials_examined">
<paragraph blockId="32.[151,1437,325,602]" pageId="32" pageNumber="33">
<materialsCitation accessionNumber="SMF 96404" collectingDate="2013-01-12" collectionCode="SMF" country="Mexico" elevation="1185" latitude="17.06753" location="Cascada Iliatenco" longLatPrecision="1" longitude="-98.77796" pageId="32" pageNumber="33" specimenCode="WGS84, GK-4260" specimenCount="1" stateProvince="Guerrero" typeStatus="holotype">
<emphasis bold="true" box="[151,260,325,350]" pageId="32" pageNumber="33">
<typeStatus box="[151,260,325,350]" pageId="32" pageNumber="33">Holotype</typeStatus>
</emphasis>
.
<accessionNumber box="[279,421,325,350]" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF96404" pageId="32" pageNumber="33">
<collectionCode box="[279,335,325,350]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="32" pageNumber="33">SMF</collectionCode>
96404
</accessionNumber>
, an adult male from
<location LSID="urn:lsid:plazi:treatment:8A288798FFE0E71F7EC3FF61FCC8C141:075E6055FFE0E7197CF3FEB3FCA0C76F" box="[679,883,325,350]" country="Mexico" latitude="17.06753" longLatPrecision="1" longitude="-98.77796" name="Cascada Iliatenco" pageId="32" pageNumber="33" stateProvince="Guerrero">Cascada Iliatenco</location>
(
<geoCoordinate box="[904,1044,325,350]" degrees="17.06753" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="1" value="17.06753">17.06753°N</geoCoordinate>
,
<geoCoordinate box="[1061,1204,325,350]" degrees="98.77796" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="1" value="-98.77796">98.77796°W</geoCoordinate>
,
<specimenCode box="[1221,1315,325,350]" collectionCode="WGS" pageId="32" pageNumber="33">WGS84</specimenCode>
),
<quantity box="[1340,1431,325,350]" metricMagnitude="3" metricUnit="m" metricValue="1.185" pageId="32" pageNumber="33" unit="m" value="1185.0">
<elevation box="[1340,1431,325,350]" metricMagnitude="3" metricUnit="m" metricValue="1.185" pageId="32" pageNumber="33" unit="m" value="1185.0">1185 m</elevation>
</quantity>
,
<collectingRegion box="[151,372,361,386]" country="Mexico" name="Guerrero" pageId="32" pageNumber="33">Estado de Guerrero</collectingRegion>
,
<collectingCountry box="[382,471,361,386]" name="Mexico" pageId="32" pageNumber="33">Mexico</collectingCountry>
; collected
<date box="[591,776,361,386]" pageId="32" pageNumber="33" value="2013-01-12">
<collectingDate box="[591,776,361,386]" pageId="32" pageNumber="33" value="2013-01-12">12 January 2013</collectingDate>
</date>
by Raúl Gómez Trejo Pérez. Field tag number
<specimenCode box="[1307,1417,361,386]" collectionCode="GK-" pageId="32" pageNumber="33">GK-4260</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph blockId="32.[151,1437,325,602]" pageId="32" pageNumber="33">
<materialsCitation accessionNumber="SMF 96403, SMF 96407" box="[199,1278,397,422]" collectingDate="2013-01-12" collectingDateMax="2013-01-13" collectingDateMin="2013-01-12" collectionCode="IBH, SMF" collectorName="All &amp; Raul Gomez Trejo Perez" country="Mexico" elevation="1120" latitude="17.05023" location="All" longLatPrecision="1" longitude="-98.68941" pageId="32" pageNumber="33" specimenCode="GK-4252-53, 4256, 96405-07, SMF 96397-99" specimenCount="3" stateProvince="Guerrero" typeStatus="paratype">
<emphasis bold="true" box="[199,319,397,422]" pageId="32" pageNumber="33">
<typeStatus box="[199,319,397,422]" pageId="32" pageNumber="33">Paratypes</typeStatus>
</emphasis>
.
<location LSID="urn:lsid:plazi:treatment:8A288798FFE0E71F7EC3FF61FCC8C141:075E6055FFE0E7197F06FE7BFEA5C797" box="[338,374,397,422]" country="Mexico" latitude="17.05023" longLatPrecision="1" longitude="-98.68941" name="All" pageId="32" pageNumber="33" stateProvince="Guerrero">All</location>
from
<collectingRegion box="[453,685,397,422]" country="Mexico" name="Guerrero" pageId="32" pageNumber="33">Estado de Guerrero</collectingRegion>
,
<collectingCountry box="[701,791,397,422]" name="Mexico" pageId="32" pageNumber="33">Mexico</collectingCountry>
:
<specimenCode box="[807,958,397,422]" collectionCode="GK-" pageId="32" pageNumber="33">GK-425253</specimenCode>
,
<specimenCode box="[974,1030,397,422]" collectionCode="GK-" pageId="32" pageNumber="33">4256</specimenCode>
(
<collectionCode box="[1050,1100,397,422]" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="32" pageNumber="33">IBH</collectionCode>
uncatalogued),
</materialsCitation>
<materialsCitation accessionNumber="SMF 96403, SMF 96407" collectingDate="2013-01-12" collectingDateMax="2013-01-13" collectingDateMin="2013-01-12" collectionCode="IBH, SMF" collectorName="All &amp; Raul Gomez Trejo Perez" country="Mexico" elevation="1120" latitude="17.05023" location="All" longLatPrecision="1" longitude="-98.68941" pageId="32" pageNumber="33" specimenCode="GK-4252-53, 4256, 96405-07, SMF 96397-99" specimenCount="3" stateProvince="Guerrero" typeStatus="paratype">
<accessionNumber box="[1291,1432,397,422]" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF96403" pageId="32" pageNumber="33">
<collectionCode box="[1291,1347,397,422]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="32" pageNumber="33">SMF</collectionCode>
96403
</accessionNumber>
,
<specimenCode box="[151,266,433,458]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="32" pageNumber="33">9640507</specimenCode>
, same collecting data as holotype.
</materialsCitation>
<materialsCitation accessionNumber="SMF 96403, SMF 96407" collectingDate="2013-01-12" collectingDateMax="2013-01-13" collectingDateMin="2013-01-12" collectionCode="IBH, SMF" collectorName="All &amp; Raul Gomez Trejo Perez" country="Mexico" elevation="1120" latitude="17.05023" location="All" longLatPrecision="1" longitude="-98.68941" pageId="32" pageNumber="33" specimenCode="GK-4252-53, 4256, 96405-07, SMF 96397-99" specimenCount="3" stateProvince="Guerrero" typeStatus="paratype">
<specimenCode box="[663,839,433,458]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="32" pageNumber="33">SMF 9639799</specimenCode>
from Iliatenco (
<geoCoordinate box="[1024,1164,433,458]" degrees="17.05023" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="1" value="17.05023">17.05023°N</geoCoordinate>
,
<geoCoordinate box="[1177,1324,433,458]" degrees="98.68941" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="1" value="-98.68941">98.68941°W</geoCoordinate>
),
<quantity box="[1344,1432,433,458]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="32" pageNumber="33" unit="m" value="1120.0">
<elevation box="[1344,1432,433,458]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="32" pageNumber="33" unit="m" value="1120.0">1120 m</elevation>
</quantity>
.
<collectorName box="[151,187,469,494]" pageId="32" pageNumber="33">All</collectorName>
paratypes are males, except for
<accessionNumber box="[556,693,469,494]" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF96407" pageId="32" pageNumber="33">
<collectionCode box="[556,612,469,494]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="32" pageNumber="33">SMF</collectionCode>
96407
</accessionNumber>
, which is a female.
<collectorName box="[921,957,469,494]" pageId="32" pageNumber="33">All</collectorName>
paratypes collected by
<collectorName pageId="32" pageNumber="33">Raúl Gómez Trejo Pérez</collectorName>
on
<date box="[255,527,505,530]" pageId="32" pageNumber="33" value="2013-01-12" valueMax="2013-01-13" valueMin="2013-01-12">
<collectingDate box="[255,527,505,530]" pageId="32" pageNumber="33" value="2013-01-12" valueMax="2013-01-13" valueMin="2013-01-12">12 and 13 January 2013</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph blockId="32.[151,1437,325,602]" pageId="32" pageNumber="33">
<emphasis bold="true" box="[199,438,541,566]" pageId="32" pageNumber="33">Referred specimens</emphasis>
<materialsCitation accessionNumber="SMF 96389" box="[439,1348,540,566]" collectionCode="SMF" collectorName="Cascada de Santa Cruz el Rincon" country="Mexico" elevation="620" location="Cascada de Santa Cruz" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Guerrero">
<emphasis bold="true" box="[467,563,541,566]" pageId="32" pageNumber="33">
<collectingCountry box="[467,558,541,566]" name="Mexico" pageId="32" pageNumber="33">Mexico</collectingCountry>
:
</emphasis>
<collectingRegion box="[575,681,541,566]" country="Mexico" name="Guerrero" pageId="32" pageNumber="33">Guerrero</collectingRegion>
:
<collectorName box="[695,1095,541,566]" pageId="32" pageNumber="33">
<location LSID="urn:lsid:plazi:treatment:8A288798FFE0E71F7EC3FF61FCC8C141:075E6055FFE0E7197CE3FDEBFC14C407" box="[695,967,541,566]" country="Mexico" name="Cascada de Santa Cruz" pageId="32" pageNumber="33" stateProvince="Guerrero">Cascada de Santa Cruz</location>
el Rincón
</collectorName>
,
<quantity box="[1111,1191,541,566]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="32" pageNumber="33" unit="m" value="620.0">
<elevation box="[1111,1191,541,566]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="32" pageNumber="33" unit="m" value="620.0">620 m</elevation>
</quantity>
:
<accessionNumber box="[1206,1348,541,566]" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF96389" pageId="32" pageNumber="33">
<collectionCode box="[1206,1262,541,566]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="32" pageNumber="33">SMF</collectionCode>
96389
</accessionNumber>
</materialsCitation>
;
<materialsCitation collectionCode="SMF" collectorName="Sta. Cruz El Rincon" country="Mexico" elevation="580" location="Rio la Tejeria" pageId="32" pageNumber="33" specimenCode="SMF 96390-94" specimenCount="1" stateProvince="Guerrero">
<location LSID="urn:lsid:plazi:treatment:8A288798FFE0E71F7EC3FF61FCC8C141:075E6055FFE0E7197B00FDEBFF3BC46B" country="Mexico" name="Rio la Tejeria" pageId="32" pageNumber="33" stateProvince="Guerrero">Río la Tejería</location>
, near
<collectorName box="[297,524,577,602]" pageId="32" pageNumber="33">Sta. Cruz El Rincón</collectorName>
,
<quantity box="[535,610,577,602]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="32" pageNumber="33" unit="m" value="580.0">
<elevation box="[535,610,577,602]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="32" pageNumber="33" unit="m" value="580.0">580 m</elevation>
</quantity>
:
<specimenCode box="[620,795,577,602]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="32" pageNumber="33">SMF 96390-94</specimenCode>
</materialsCitation>
;
<materialsCitation box="[806,1318,577,602]" collectionCode="SMF" collectorName="Santa Cruz el Rincon" country="Mexico" elevation="705" pageId="32" pageNumber="33" specimenCode="SMF 96395-96" specimenCount="1" stateProvince="Guerrero">
<collectorName box="[806,1047,577,602]" pageId="32" pageNumber="33">Santa Cruz el Rincón</collectorName>
,
<quantity box="[1058,1133,577,602]" metricMagnitude="2" metricUnit="m" metricValue="7.05" pageId="32" pageNumber="33" unit="m" value="705.0">
<elevation box="[1058,1133,577,602]" metricMagnitude="2" metricUnit="m" metricValue="7.05" pageId="32" pageNumber="33" unit="m" value="705.0">705 m</elevation>
</quantity>
:
<specimenCode box="[1144,1318,577,602]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="32" pageNumber="33">SMF 96395-96</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931003" ID-Zenodo-Dep="4931003" httpUri="https://zenodo.org/record/4931003/files/figure.png" pageId="32" pageNumber="33" startId="32.[151,250,1968,1990]" targetBox="[157,1436,658,1943]" targetPageId="32">
<paragraph blockId="32.[151,1436,1968,2021]" pageId="32" pageNumber="33">
<emphasis bold="true" box="[151,290,1968,1990]" pageId="32" pageNumber="33">FIGURE 20.</emphasis>
Dewlaps of
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[425,557,1968,1990]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="nietoi">
<emphasis box="[425,557,1968,1990]" italics="true" pageId="32" pageNumber="33">Anolis nietoi</emphasis>
</taxonomicName>
in life. (a) SMF 96406; (b) SMF 96403; (c) SMF 96402; (d) SMF 96407. (ac) are males, (d) is a female.
</paragraph>
</caption>
<subSubSection pageId="33" pageNumber="34" type="diagnosis">
<paragraph blockId="33.[151,1437,151,743]" pageId="33" pageNumber="34">
<emphasis bold="true" box="[199,314,151,176]" pageId="33" pageNumber="34">Diagnosis</emphasis>
. A small species (our Species B of the
<taxonomicName authorityName=": Ahl" authorityYear="1934" box="[758,910,152,176]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="nebuloides">
<emphasis box="[758,910,152,176]" italics="true" pageId="33" pageNumber="34">A. nebuloides</emphasis>
</taxonomicName>
complex, see above; SVL in largest male 50.0 mm, largest female
<quantity box="[384,488,185,210]" metricMagnitude="-2" metricUnit="m" metricValue="3.85" pageId="33" pageNumber="34" unit="mm" value="38.5">38.5 mm</quantity>
) of the genus
<taxonomicName authority="(sensu Poe 2004)" baseAuthorityName="sensu Poe" baseAuthorityYear="2004" box="[661,944,185,210]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="genus">
<emphasis box="[661,732,185,209]" italics="true" pageId="33" pageNumber="34">Anolis</emphasis>
(sensu
<bibRefCitation author="Poe, S." box="[825,936,185,210]" pageId="33" pageNumber="34" pagination="37 - 89" refId="ref95161" refString="Poe, S. (2004) Phylogeny of anoles. Herpetological Monographs, 18, 37 - 89. http: // dx. doi. org / 10.1655 / 0733 - 1347 (2004) 018 [0037: poa] 2.0. co; 2" type="journal article" year="2004">Poe 2004</bibRefCitation>
)
</taxonomicName>
that differs from all Mexican and Central American anoles except
<emphasis box="[428,813,219,243]" italics="true" pageId="33" pageNumber="34">
<taxonomicName authorityName=": Ahl" authorityYear="1934" box="[428,584,219,243]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="nebuloides">A. nebuloides</taxonomicName>
,
<taxonomicName authorityName="Smith" authorityYear="1933" box="[595,809,219,243]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="megapholidotus">A. megapholidotus</taxonomicName>
,
</emphasis>
and two species described below (our Species D and E of the
<taxonomicName authorityName=": Ahl" authorityYear="1934" box="[223,375,252,276]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="nebuloides">
<emphasis box="[223,375,252,276]" italics="true" pageId="33" pageNumber="34">A. nebuloides</emphasis>
</taxonomicName>
complex) by having a combination of (1) strongly keeled ventral scales; (2) usually a patch of three greatly enlarged supraocular scales; (3) 1012 rows of greatly enlarged dorsal scales that are larger than ventral scales; (4) short hind legs, longest toe of adpressed hind leg usually reaching to posterior margin of eye, occasionally to a point between levels of ear opening and eye or to mideye (to ear opening in a single female), ratio of shank length/SVL 0.240.28; (5) circumnasal usually in contact with first supralabial; and (6) a large pink dewlap in males and a very small pink dewlap in females (
<figureCitation box="[818,903,418,443]" captionStart="FIGURE 20" captionStartId="32.[151,250,1968,1990]" captionTargetBox="[157,1436,658,1943]" captionTargetId="figure-185@32.[157,1442,658,1943]" captionTargetPageId="32" captionText="FIGURE 20. Dewlaps of Anolis nietoi in life. (a) SMF 96406; (b) SMF 96403; (c) SMF 96402; (d) SMF 96407. (ac) are males, (d) is a female." figureDoi="http://doi.org/10.5281/zenodo.4931003" httpUri="https://zenodo.org/record/4931003/files/figure.png" pageId="33" pageNumber="34">Fig. 20</figureCitation>
).
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[923,1067,419,443]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="nietoi">
<emphasis box="[923,1067,419,443]" italics="true" pageId="33" pageNumber="34">Anolis nietoi</emphasis>
</taxonomicName>
differs from Species D and E of the
<taxonomicName authorityName=": Ahl" authorityYear="1934" box="[194,348,452,476]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="nebuloides">
<emphasis box="[194,348,452,476]" italics="true" pageId="33" pageNumber="34">A. nebuloides</emphasis>
</taxonomicName>
complex by having (1) a rather abrupt transition between the enlarged dorsal scales and lateral body scales (vs. a more gradual transition over 3 to 4 scales); (2) the edges of the field of enlarged dorsal scale rows well defined and in a more or less straight line (vs. the edges of this field forming an undulating line due to single enlarged scales or groups thereof outside the main field) (3) and by having a more reddish male dewlap (vs. pink).
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[151,299,585,609]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="nietoi">
<emphasis box="[151,299,585,609]" italics="true" pageId="33" pageNumber="34">Anolis nietoi</emphasis>
</taxonomicName>
differs from
<taxonomicName authorityName=": Ahl" authorityYear="1934" box="[464,622,585,609]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="nebuloides">
<emphasis box="[464,622,585,609]" italics="true" pageId="33" pageNumber="34">A. nebuloides</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Smith" authorityYear="1933" box="[688,903,585,609]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="megapholidotus">
<emphasis box="[688,903,585,609]" italics="true" pageId="33" pageNumber="34">A. megapholidotus</emphasis>
</taxonomicName>
in the ranges and average values of several morphometric and pholidotic characters (see
<tableCitation box="[658,742,618,643]" captionStart="Table 8" captionStartId="29.[147,206,563,584]" captionTargetBox="[149,1864,690,1549]" captionText="Table 8. Selected measurements, proportions and scale characters of the species related to Anolis nebuloides. Range is followed by mean value and standard deviation in parentheses. For abbreviations see text." httpUri="http://table.plazi.org/id/56FE6606FFDDE7247EC7FDC5FE47C4B1" pageId="33" pageNumber="34" tableUuid="56FE6606FFDDE7247EC7FDC5FE47C4B1">Table 8</tableCitation>
), most obvious in (1) number of middorsal scales in one head length; and (2) number of subdigital lamellae on Phalanges IIIV of Toe IV. Also, the incompletely everted hemipenis of the
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[350,447,686,709]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="nietoi">
<emphasis box="[350,447,686,709]" italics="true" pageId="33" pageNumber="34">A. nietoi</emphasis>
</taxonomicName>
<typeStatus box="[456,554,685,710]" pageId="33" pageNumber="34">holotype</typeStatus>
appears to be unilobed (vs. bilobed in
<taxonomicName authorityName=": Ahl" authorityYear="1934" box="[1004,1157,685,709]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="nebuloides">
<emphasis box="[1004,1157,685,709]" italics="true" pageId="33" pageNumber="34">A. nebuloides</emphasis>
</taxonomicName>
and Species E of the
<taxonomicName authorityName=": Ahl" authorityYear="1934" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="nebuloides">
<emphasis italics="true" pageId="33" pageNumber="34">A. nebuloides</emphasis>
</taxonomicName>
complex; the hemipenis morphology in
<taxonomicName authorityName="Smith" authorityYear="1933" box="[726,936,718,742]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="megapholidotus">
<emphasis box="[726,936,718,742]" italics="true" pageId="33" pageNumber="34">A. megapholidotus</emphasis>
</taxonomicName>
is unknown).
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931005" ID-Zenodo-Dep="4931005" httpUri="https://zenodo.org/record/4931005/files/figure.png" pageId="33" pageNumber="34" startId="33.[151,250,1709,1731]" targetBox="[151,1436,823,1687]" targetPageId="33">
<paragraph blockId="33.[151,1436,1709,1762]" pageId="33" pageNumber="34">
<emphasis bold="true" box="[151,288,1709,1731]" pageId="33" pageNumber="34">FIGURE 21.</emphasis>
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[294,423,1709,1731]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="34" phylum="Chordata" rank="species" species="nietoi">
<emphasis box="[294,423,1709,1731]" italics="true" pageId="33" pageNumber="34">Anolis nietoi</emphasis>
</taxonomicName>
in life. (a) SMF 96406; (b) IBH 27051; (c) SMF 96398; (d) SMF 96403. (ab) are males, (cd) are females.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931007" ID-Zenodo-Dep="4931007" httpUri="https://zenodo.org/record/4931007/files/figure.png" pageId="34" pageNumber="35" startId="34.[151,250,1537,1559]" targetBox="[153,1436,199,1506]" targetPageId="34">
<paragraph blockId="34.[151,1435,1537,1590]" pageId="34" pageNumber="35">
<emphasis bold="true" box="[151,290,1537,1559]" pageId="34" pageNumber="35">FIGURE 22.</emphasis>
Holotype of
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[429,560,1537,1559]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="nietoi">
<emphasis box="[429,560,1537,1559]" italics="true" pageId="34" pageNumber="35">Anolis nietoi</emphasis>
</taxonomicName>
(SMF 96404): (a) dorsal view; (b) lateral view; (c) ventral view; (d) lateral view of head; (e) dorsal view of head; (f) ventral view of head. Scale bars equal 5.0 mm in (ac) and 1.0 mm in (df), respectively.
</paragraph>
</caption>
<subSubSection lastPageId="38" lastPageNumber="39" pageId="34" pageNumber="35" type="description">
<paragraph blockId="34.[151,1436,1665,2014]" lastBlockId="37.[151,1436,1581,2037]" lastPageId="37" lastPageNumber="38" pageId="34" pageNumber="35">
<emphasis bold="true" box="[199,528,1665,1690]" pageId="34" pageNumber="35">
Description of the
<typeStatus box="[426,528,1665,1690]" pageId="34" pageNumber="35">holotype</typeStatus>
</emphasis>
. Adult male, as indicated by everted hemipenes and presence of large dewlap; SVL 50.0 mm; tail incomplete; tail moderately compressed in cross section, tail height
<quantity box="[1131,1216,1701,1726]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="34" pageNumber="35" unit="mm" value="2.7">2.7 mm</quantity>
and width
<quantity box="[1343,1432,1701,1726]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="34" pageNumber="35" unit="mm" value="1.8">1.8 mm</quantity>
; axilla to groin distance
<quantity box="[422,528,1737,1762]" metricMagnitude="-2" metricUnit="m" metricValue="2.09" pageId="34" pageNumber="35" unit="mm" value="20.9">20.9 mm</quantity>
; head length
<quantity box="[683,787,1737,1762]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="34" pageNumber="35" unit="mm" value="13.4">13.4 mm</quantity>
, head length/SVL ratio 0.27; snout length
<quantity box="[1278,1370,1737,1762]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="34" pageNumber="35" unit="mm" value="5.8">5.8 mm</quantity>
; head width
<quantity box="[223,313,1773,1798]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="34" pageNumber="35" unit="mm" value="7.6">7.6 mm</quantity>
; longest toe of adpressed hind limb reaching to a point between ear opening and eye; shank length
<quantity box="[151,253,1809,1834]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="34" pageNumber="35" unit="mm" value="12.5">12.5 mm</quantity>
, shank length/head length ratio 0.93; longest finger of extended forelimb reaching to a point between eye and nostril; longest finger of adpressed forelimb reaching to a point
<quantity box="[931,1019,1845,1870]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="34" pageNumber="35" unit="mm" value="1.8">1.8 mm</quantity>
in front of anterior insertion of hind limbs. Scales on snout keeled, unicarinate; posterior head scales rugose or keeled; 6 postrostrals; 7 scales between nasals; 1 elongate prenasal scale on each side, in contact with both rostral and first supralabial; circumnasal in contact with first supralabial; scales in shallow prefrontal depression mostly keeled, keels mostly oriented longitudinally; supraorbital semicircles well developed, separated medially by one scale row at narrowest point; supraorbital disc composed of an oval patch of 3 greatly enlarged, weakly keeled scales; circumorbital row complete, therefore, enlarged supraorbital scales completely separated from supraorbital semicircles; 3 elongated, strongly overlapping superciliaries, anterior one larger than posterior ones, followed posteriorly by 3 to 4 roundish to squarish scales of moderate size; 2 to 3 rows of small smooth scales extending between enlarged supraorbitals and superciliaries; parietal depression shallow; interparietal scale well developed, 1.6 x
<quantity box="[1150,1238,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="35" pageNumber="36" unit="mm" value="1.6">1.6 mm</quantity>
(length x width), surrounded by scales of moderate size; 2 scales present between interparietal and supraorbital semicircles; canthal ridge distinct, composed of 3 large (second largest) and 3 small anterior canthal scales; 6 scales present between second canthals; 7 scales present between posterior canthals; 25 (right) 29 (left) mostly keeled loreal scales in a maximum of 5 horizontal rows; 4 keeled subocular scales arranged in a single row; 6 supralabials to level below center of eye; 3 suboculars broadly in contact with 3 supralabials; ear opening 0.5 x
<quantity box="[1138,1227,476,501]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="35" pageNumber="36" unit="mm" value="1.6">1.6 mm</quantity>
(length x height); mental distinctly wider than long, completely divided medially, bordered posteriorly by 5 postmentals, outer ones slightly larger than median ones; 6 infralabials to level below center of eye; sublabials not differentiated; keeled granular scales present on chin and throat; dewlap large, extending from level beyond anterior margin of eye to level of chest; 89 horizontal gorgetal-sternal rows with 914 scales per row; modal number of marginal pairs 24; a nuchal crest and a dorsal ridge present; dorsum of body with strongly keeled, imbricate, non-mucronate scales; about 12 medial rows greatly enlarged; largest dorsal scales about 0.80 x
<quantity box="[1015,1119,1652,1677]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="37" pageNumber="38" unit="mm" value="0.8">0.80 mm</quantity>
(length x width); about 18 medial dorsal scales in one head length; about 33 medial dorsal scales between levels of axilla and groin; lateral scales smooth, granular and more or less homogeneous in size, average size
<quantity box="[1053,1157,1724,1749]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="37" pageNumber="38" unit="mm" value="0.3">0.30 mm</quantity>
in diameter; ventrals at midbody strongly keeled, imbricate, mucronate, about 0.70 x
<quantity box="[855,956,1760,1785]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="37" pageNumber="38" unit="mm" value="0.65">0.65 mm</quantity>
(length x width); about 30 medial ventral scales in one head length; about 37 medial ventral scales between levels of axilla and groin; 96 scales around midbody; all caudal scales keeled; middorsal caudal scales moderately enlarged, not forming a crest; lateral caudal scales without whorls of enlarged scales, although an indistinct division in segments is discernible; a pair of moderately enlarged postcloacal scales present, about
<quantity box="[759,845,1904,1929]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="37" pageNumber="38" unit="mm" value="0.7">0.7 mm</quantity>
wide; no tube-like axillary pocket present; scales on dorsal surface of forelimb weakly keeled; digital pads dilated, dilated pad about 3 times width of non-dilated distal phalanx; distal phalanx narrower than and raised from dilated pad; 21 (right)22 (left) lamellae under phalanges IIIV of fourth toe; 7 scales under distal phalanx of fourth toe.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931009" ID-Zenodo-Dep="4931009" box="[151,1273,2000,2022]" httpUri="https://zenodo.org/record/4931009/files/figure.png" pageId="35" pageNumber="36" startId="35.[151,250,2000,2022]" targetBox="[151,1436,651,1957]" targetPageId="35">
<paragraph blockId="35.[151,1273,2000,2022]" box="[151,1273,2000,2022]" pageId="35" pageNumber="36">
<emphasis bold="true" box="[151,288,2000,2022]" pageId="35" pageNumber="36">FIGURE 23.</emphasis>
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[293,422,2000,2022]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="nietoi">
<emphasis box="[293,422,2000,2022]" italics="true" pageId="35" pageNumber="36">Anolis nietoi</emphasis>
</taxonomicName>
: head scalation in (a,c,e) SMF 96406; (b,d,f) SMF 96398. Scale bars equal 1.0 mm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931011" ID-Zenodo-Dep="4931011" httpUri="https://zenodo.org/record/4931011/files/figure.png" pageId="36" pageNumber="37" startId="36.[151,250,1945,1967]" targetBox="[151,1436,179,1926]" targetPageId="36">
<paragraph blockId="36.[151,1436,1945,2028]" pageId="36" pageNumber="37">
<emphasis bold="true" box="[151,289,1945,1967]" pageId="36" pageNumber="37">FIGURE 24.</emphasis>
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[296,427,1945,1967]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="nietoi">
<emphasis box="[296,427,1945,1967]" italics="true" pageId="36" pageNumber="37">Anolis nietoi</emphasis>
</taxonomicName>
: nasal region in (a) SMF 96406; (b) SMF 96398; superciliary region in (c) SMF 96403; (d) SMF 96406; chin region in (e) SMF 96403; (f) SMF 96406; cloacal region in (g) SMF 96395; (h) SMF 96390. Scale bars equal 1.0 mm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931013" ID-Zenodo-Dep="4931013" httpUri="https://zenodo.org/record/4931013/files/figure.png" pageId="37" pageNumber="38" startId="37.[151,250,1498,1520]" targetBox="[151,1426,155,1461]" targetPageId="37">
<paragraph blockId="37.[151,1436,1498,1551]" pageId="37" pageNumber="38">
<emphasis bold="true" box="[151,289,1498,1520]" pageId="37" pageNumber="38">FIGURE 25.</emphasis>
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[296,426,1498,1520]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="nietoi">
<emphasis box="[296,426,1498,1520]" italics="true" pageId="37" pageNumber="38">Anolis nietoi</emphasis>
</taxonomicName>
: dorsal scales in (a) IBH 26573; (b) SMF 96213; flank scales in (c) SMF 96406; (d) SMF 96398; ventral scales in (e) SMF 96406; (f) SMF 96398. Scale bars equal 1.0 mm.
</paragraph>
</caption>
<paragraph blockId="38.[151,1437,151,1904]" box="[199,1412,151,176]" pageId="38" pageNumber="39">The incompletely everted hemipenis appears to be a small unilobate organ, possibly with rudimentary lobes.</paragraph>
<paragraph blockId="38.[151,1437,151,1904]" pageId="38" pageNumber="39">Coloration after six months preservation in 70% ethanol was recorded as follows: Dorsal surfaces of head, body, limbs, and tail Drab (19) with a suffusion of Dark Neutral Gray (299) at middorsum and with an indistinct Dark Neutral Gray (299) lateral longitudinal stripe between levels of axilla and groin; ventral surfaces of head, body, limbs, and tail Cream White (52); finger and toe pads Vandyke Brown (282); dewlap Flesh Color (249).</paragraph>
<paragraph blockId="38.[151,1437,151,1904]" pageId="38" pageNumber="39">
<emphasis bold="true" box="[199,311,332,357]" pageId="38" pageNumber="39">Variation</emphasis>
. The
<typeStatus box="[382,489,332,356]" pageId="38" pageNumber="39">paratypes</typeStatus>
agree well with the
<typeStatus box="[732,830,332,357]" pageId="38" pageNumber="39">holotype</typeStatus>
in general appearance, morphometrics and scalation (see
<tableCitation box="[205,291,367,392]" captionStart="Table 8" captionStartId="29.[147,206,563,584]" captionTargetBox="[149,1864,690,1549]" captionText="Table 8. Selected measurements, proportions and scale characters of the species related to Anolis nebuloides. Range is followed by mean value and standard deviation in parentheses. For abbreviations see text." httpUri="http://table.plazi.org/id/56FE6606FFDDE7247EC7FDC5FE47C4B1" pageId="38" pageNumber="39" tableUuid="56FE6606FFDDE7247EC7FDC5FE47C4B1">Table 8</tableCitation>
). Variation is evident in some scalation characters as follows: The number of slightly to moderately enlarged dorsal scale rows varies from 10 to 13. All specimens have a single elongate prenasal. Suboculars in contact with supralabials (23 suboculars in contact with 25 supralabials) in all specimens examined. The longest toe of adpressed hind leg reaching to a point between posterior and anterior border of eye in most individuals except in
<specimenCount box="[267,437,511,536]" pageId="38" pageNumber="39" type="generic">two specimens</specimenCount>
(
<specimenCode box="[455,638,511,536]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="38" pageNumber="39">SMF 9638990</specimenCode>
) that have the longest toe of adpressed hind leg reaching to a point between ear opening and eye. Usually males have a pair of greatly enlarged postcloacal scales, no enlarged postcloacals in
<specimenCount box="[324,425,583,608]" pageId="38" pageNumber="39" type="male">one male</specimenCount>
specimen (
<accessionNumber box="[553,692,583,608]" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF96396" pageId="38" pageNumber="39">
<collectionCode box="[553,611,583,608]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="38" pageNumber="39">SMF</collectionCode>
96396
</accessionNumber>
).
</paragraph>
<paragraph blockId="38.[151,1437,151,1904]" pageId="38" pageNumber="39">
The coloration in life of an adult male (
<accessionNumber box="[643,780,620,645]" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF96397" pageId="38" pageNumber="39">SMF 96397</accessionNumber>
) was recorded as follows: Dorsal surface of body Antique Brown (24) with suffusions of Marsh Brown (25); lateral ground color Tawny Olive (17); dorsal surface of head Raw Umber (23); ventral surface of body Cream Color (12); ventral surface of legs Verona Brown (37); chin with Medium Neutral Gray (298) and Pink (242) speckles; dewlap Vinaceous Color (244) and Cream White (52) gorgetals; iris Raw Umber (22). The coloration in life of another adult male (
<accessionNumber box="[1014,1143,764,789]" httpUri="http://www.ncbi.nlm.nih.gov/protein/IBH27050" pageId="38" pageNumber="39">IBH 27050</accessionNumber>
) was recorded as follows: Dorsal ground color Amber (51) grading into Cinnamon Drab (50) towards tail with Russet (44) chevrons, Cream Color (12) lateral spots and Light Buff (2) lateral line; coloration between lateral line and venter Drab (19); ventral surface of body Cream Color (12); ventral surface of limbs Drab (19); ventral surface of tail Clay Color (18); dewlap Vinaceous (244); iris Cinnamon (255). The coloration in life of another adult male (
<accessionNumber box="[1240,1374,908,933]" httpUri="http://www.ncbi.nlm.nih.gov/protein/IBH27051" pageId="38" pageNumber="39">IBH 27051</accessionNumber>
) was recorded as follows: Dorsal ground color Cinnamon (255) with Medium Neutral Gray (298) vertebral line, barely visible Antique Brown (24) chevrons and a Light Buff (2) lateral line that is edged by Vinaceous (244); ventral surface of body Chamois (84) with Salmon Color (83) suffusions; ventral surface of limbs Prouts Brown (47); dewlap Spinel Pink (253); iris Olive Yellow (117). The coloration in life of another adult male (
<accessionNumber box="[1239,1377,1052,1077]" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF46406" pageId="38" pageNumber="39">SMF 46406</accessionNumber>
) was recorded as follows: Dorsal ground color Grayish Horn Color (268) with suffusions of Light Neutral Gray (297) and a Cream White (52) lateral stripe edged by Flesh Ocher (57); interorbital bar Verona Brown (37); dorsal surface of limbs and tail Tawny (60); ventral surface of body Pale Buff (1); ventral surface of limbs Clay Color (18); dewlap Pinkish Rose (220); iris Ground Cinnamon (270).
</paragraph>
<paragraph blockId="38.[151,1437,151,1904]" pageId="38" pageNumber="39">
The coloration in life of an adult female of
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[683,825,1232,1256]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="nietoi">
<emphasis box="[683,825,1232,1256]" italics="true" pageId="38" pageNumber="39">Anolis nietoi</emphasis>
</taxonomicName>
(
<accessionNumber box="[840,976,1231,1256]" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF96397" pageId="38" pageNumber="39">SMF 96397</accessionNumber>
) was recorded as follows: Dorsal ground color of body and tail Verona Brown (37) with Dusky Brown (258) Chevrons, lateral stripe and reticulations in head; tail with Cream White (12) chevrons; Ventral surface of body Pale Buff (1) with suffusions of Pink (242); ventral surface of tail and legs Light Russet Vinaceous (246); dewlap Vinaceous (244); iris Sayal Brown (41). The coloration in life of another adult female (
<accessionNumber box="[630,768,1375,1400]" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF96407" pageId="38" pageNumber="39">SMF 96407</accessionNumber>
) was recorded as follows: Dorsal ground color Drab (119) with Dark Neutral Gray (299) and Olive Brown (278) diamonds that are edged by Micado Brown (42) and Burnt Umber (48); dorsal surface of limbs and tail Russet (44); ventral surface of body Pale Pinkish Buff (3); ventral surface of limbs and tail Antique Brown (24); dewlap Pale Pinkish Purple (223); iris Clay Color (18).
</paragraph>
</subSubSection>
<subSubSection pageId="38" pageNumber="39" type="etymology">
<paragraph blockId="38.[151,1437,151,1904]" pageId="38" pageNumber="39">
<emphasis bold="true" box="[199,327,1519,1544]" pageId="38" pageNumber="39">Etymology</emphasis>
. The name
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[475,539,1520,1543]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="nietoi">
<emphasis box="[475,539,1520,1543]" italics="true" pageId="38" pageNumber="39">nietoi</emphasis>
</taxonomicName>
is a patronym honoring Adrián Nieto Montes de Oca who has contributed substantially to our knowledge on Mexican anoles.
</paragraph>
<paragraph blockId="38.[151,1437,151,1904]" pageId="38" pageNumber="39">
<emphasis bold="true" box="[199,463,1591,1616]" pageId="38" pageNumber="39">Natural History Notes</emphasis>
. All
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[519,661,1592,1616]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="nietoi">
<emphasis box="[519,661,1592,1616]" italics="true" pageId="38" pageNumber="39">Anolis nietoi</emphasis>
</taxonomicName>
from Iliatenco where encountered during the day along a river shore. They were active and near the base of coffee trees, banana palms, and small shrubs.
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[1099,1242,1628,1652]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="nietoi">
<emphasis box="[1099,1242,1628,1652]" italics="true" pageId="38" pageNumber="39">Anolis nietoi</emphasis>
</taxonomicName>
from Río Tejería where found at night while they were sleeping on big leaves hanging over the river at
<quantity box="[1133,1273,1663,1688]" metricMagnitude="0" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="38" pageNumber="39" unit="cm" value="125.0" valueMax="150.0" valueMin="100.0">100150 cm</quantity>
above ground level.
</paragraph>
<paragraph blockId="38.[151,1437,151,1904]" pageId="38" pageNumber="39">
<emphasis bold="true" box="[199,720,1735,1760]" pageId="38" pageNumber="39">Geographic Distribution and Conservation</emphasis>
. As currently known,
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[983,1129,1736,1760]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="nietoi">
<emphasis box="[983,1129,1736,1760]" italics="true" pageId="38" pageNumber="39">Anolis nietoi</emphasis>
</taxonomicName>
is restricted to the Pacific versant of the Sierra Madre del Sur in the southeastern portion of the Mexican State of
<collectingRegion box="[1180,1281,1772,1797]" country="Mexico" name="Guerrero" pageId="38" pageNumber="39">Guerrero</collectingRegion>
at elevations between 580 and
<quantity box="[352,468,1807,1832]" metricMagnitude="3" metricUnit="m" metricValue="1.185" pageId="38" pageNumber="39" unit="m" value="1185.0">1185 masl</quantity>
(
<figureCitation box="[485,570,1807,1832]" captionStart="FIGURE 40" captionStartId="54.[151,250,1951,1973]" captionTargetBox="[280,1301,1023,1911]" captionTargetId="figure-399@54.[269,1311,1013,1921]" captionTargetPageId="54" captionText="FIGURE 40. Map indicating collecting localities of species related to Anolis nebuloides. Each symbol can represent one or more adjacent localities. Black squares: Anolis megapholidotus; white squares: A. nietoi; black circles: A. nebuloides; black triangles: A. stevepoei; white circles: A. zapotecorum." figureDoi="http://doi.org/10.5281/zenodo.4931043" httpUri="https://zenodo.org/record/4931043/files/figure.png" pageId="38" pageNumber="39">Fig. 40</figureCitation>
). Given its presumably small geographic range, degraded habitat, and threat from deforestation it seems justified to classify
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[696,792,1844,1868]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="nietoi">
<emphasis box="[696,792,1844,1868]" italics="true" pageId="38" pageNumber="39">A. nietoi</emphasis>
</taxonomicName>
as Critically Endangered based on criterion B1ab(iii) of the IUCN Red List Categories and Criteria (
<bibRefCitation author="IUCN" box="[644,783,1879,1904]" pageId="38" pageNumber="39" refId="ref91603" refString="IUCN (2012) IUCN Red List of Threatened Species. Version 2012.2. Available from: http: // www. iucnredlist. org (accessed 11 January 2013)" type="url" year="2012">IUCN 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>