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<document ID-DOI="http://dx.doi.org/10.11646/zootaxa.3862.1.1" ID-GBIF-Dataset="bebe1ef9-e8d6-459a-a102-8b9989b465f1" ID-ISSN="1175-5326" ID-Zenodo-Dep="4930959" ID-ZooBank="3FA375FE-E4E0-4509-BE02-EE5E786B07C6" approvalRequired="20" approvalRequired_for_treatments="20" checkinTime="1623452360688" checkinUser="felipe" docAuthor="Köhler, Gunther, Pérez, Raúl Gómez Trejo, Petersen, Claus Bo P., Méndez, Fausto R. &amp; Cruz, De La" docDate="2014" docId="8A288798FF9FE75F7EC3FB81FE13C07E" docLanguage="en" docName="zt03862p210.pdf.imf" docOrigin="Zootaxa 3862 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.10:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="10" docTitle="Anolis immaculogularis Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz 2014, sp. nov." docType="treatment" docVersion="6" lastPageNumber="103" masterDocId="7611FFE0FFC0E7397E54FFF6FFD3C631" masterDocTitle="A revision of the Mexican Anolis (Reptilia, Squamata, Dactyloidae) from the Pacific versant west of the Isthmus de Tehuantepec in the states of Oaxaca, Guerrero, and Puebla, with the description of six new species" masterLastPageNumber="210" masterPageNumber="1" pageNumber="96" updateTime="1673622526097" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the Mexican Anolis (Reptilia, Squamata, Dactyloidae) from the Pacific versant west of the Isthmus de Tehuantepec in the states of Oaxaca, Guerrero, and Puebla, with the description of six new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Köhler, Gunther</mods:namePart>
<mods:nameIdentifier type="email">gkoehler@senckenberg.de</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pérez, Raúl Gómez Trejo</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Petersen, Claus Bo P.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Méndez, Fausto R.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cruz, De La</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="pubDate">
<mods:number>2014-09-19</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>3862</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>210</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.11646/zootaxa.3862.1.1</mods:identifier>
<mods:identifier type="GBIF-Dataset">bebe1ef9-e8d6-459a-a102-8b9989b465f1</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">4930959</mods:identifier>
<mods:identifier type="ZooBank">3FA375FE-E4E0-4509-BE02-EE5E786B07C6</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4930969" ID-Zenodo-Dep="4930969" LSID="urn:lsid:plazi:treatment:8A288798FF9FE75F7EC3FB81FE13C07E" httpUri="http://treatment.plazi.org/id/8A288798FF9FE75F7EC3FB81FE13C07E" lastPageId="102" lastPageNumber="103" pageId="95" pageNumber="96">
<subSubSection box="[151,542,1143,1170]" pageId="95" pageNumber="96" type="nomenclature">
<paragraph blockId="95.[151,542,1143,1203]" box="[151,542,1143,1170]" pageId="95" pageNumber="96">
<heading bold="true" box="[151,542,1143,1170]" fontSize="11" level="2" pageId="95" pageNumber="96" reason="4">
<emphasis bold="true" box="[151,542,1143,1170]" pageId="95" pageNumber="96">
<taxonomicName authority="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz, 2014" authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[151,441,1143,1169]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="95" pageNumber="96" phylum="Chordata" rank="species" species="immaculogularis" status="sp. nov.">
<emphasis bold="true" box="[151,441,1143,1169]" italics="true" pageId="95" pageNumber="96">Anolis immaculogularis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[449,542,1144,1170]" pageId="95" pageNumber="96" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,284,1178,1203]" pageId="95" pageNumber="96" type="description">
<paragraph blockId="95.[151,542,1143,1203]" box="[151,284,1178,1203]" pageId="95" pageNumber="96">
<figureCitation box="[151,284,1178,1203]" captionStart-0="FIGURE 77" captionStart-1="FIGURE 78" captionStart-2="FIGURE 79" captionStart-3="FIGURE 80" captionStart-4="FIGURE 81" captionStart-5="FIGURE 82" captionStart-6="FIGURE 83" captionStartId-0="96.[151,250,1949,1971]" captionStartId-1="97.[151,250,1526,1548]" captionStartId-2="98.[151,250,1785,1807]" captionStartId-3="99.[151,250,1530,1552]" captionStartId-4="100.[151,250,1949,1971]" captionStartId-5="101.[151,250,1525,1547]" captionStartId-6="102.[151,250,793,815]" captionTargetBox-0="[252,1338,1193,1915]" captionTargetBox-1="[155,1436,198,1504]" captionTargetBox-2="[157,1436,459,1765]" captionTargetBox-3="[153,1436,195,1502]" captionTargetBox-4="[151,1434,181,1930]" captionTargetBox-5="[156,1436,193,1499]" captionTargetBox-6="[151,1436,193,772]" captionTargetId-0="figure-560@96.[252,1338,1193,1915]" captionTargetId-1="figure-249@97.[155,1440,198,1504]" captionTargetId-2="figure-203@98.[157,1442,459,1765]" captionTargetId-3="figure-236@99.[153,1438,195,1502]" captionTargetId-4="figure-71@100.[149,1434,181,1930]" captionTargetId-5="figure-192@101.[156,1441,193,1499]" captionTargetId-6="figure-465@102.[151,1436,193,772]" captionTargetPageId-0="96" captionTargetPageId-1="97" captionTargetPageId-2="98" captionTargetPageId-3="99" captionTargetPageId-4="100" captionTargetPageId-5="101" captionTargetPageId-6="102" captionText-0="FIGURE 77. Dewlaps of Anolis immaculogularis in life. (a) SMF 96264; (b) IBH 26562; (c) SMF 96263; (d) SMF 96467; (e) IBH 26564; (f) IBH 26560. (ad) are males, (ef) are females." captionText-1="FIGURE 78. Anolis immaculogularis in life. (a) SMF 96263; (b) SMF 96467; (c) SMF 96467; (d) IBH 26564; (e) IBH 26560; (f) IBH 26693. (ac) are males, (df) are females." captionText-2="FIGURE 79. Holotype of Anolis immaculogularis (SMF 96266): (a) dorsal view; (b) lateral view; (c) ventral view; (d) lateral view of head; (e) dorsal view of head; (f) ventral view of head. Scale bars equal 5.0 mm in (ac) and 1.0 mm in (df), respectively." captionText-3="FIGURE 80. Anolis immaculogularis: head scalation in (a,c,e) IBH 26564; (b,d,f) IBH 26562. Scale bars equal 1.0 mm." captionText-4="FIGURE 81. Anolis immaculogularis: nasal region in (a) IBH 26562; (b) IBH 26564; superciliary region in (c) IBH 26560; (d) IBH 26564; chin region in (e) IBH 26560; (f) IBH 26564; cloacal region in (g) SMF 93843; (h) SMF 93842. Scale bars equal 1.0 mm." captionText-5="FIGURE 82. Anolis immaculogularis: dorsal scales in (a) IBH 26562; (b) IBH 26560; flank scales in (c) IBH 26564; (d) IBH 26560; ventral scales in (e) IBH 26564; (f) IBH 26560. Scale bars equal 1.0 mm." captionText-6="FIGURE 83. Hemipenis of Anolis immaculogularis (SMF 93842): (left) sulcate view; (center) asulcate view; (right) lateral view. Scale bar equals 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4931117" figureDoi-1="http://doi.org/10.5281/zenodo.4931119" figureDoi-2="http://doi.org/10.5281/zenodo.4931121" figureDoi-3="http://doi.org/10.5281/zenodo.4931123" figureDoi-4="http://doi.org/10.5281/zenodo.4931125" figureDoi-5="http://doi.org/10.5281/zenodo.4931127" figureDoi-6="http://doi.org/10.5281/zenodo.4931129" httpUri-0="https://zenodo.org/record/4931117/files/figure.png" httpUri-1="https://zenodo.org/record/4931119/files/figure.png" httpUri-2="https://zenodo.org/record/4931121/files/figure.png" httpUri-3="https://zenodo.org/record/4931123/files/figure.png" httpUri-4="https://zenodo.org/record/4931125/files/figure.png" httpUri-5="https://zenodo.org/record/4931127/files/figure.png" httpUri-6="https://zenodo.org/record/4931129/files/figure.png" pageId="95" pageNumber="96">Figs. 7783</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="95" pageNumber="96" type="materials_examined">
<paragraph blockId="95.[151,1437,1250,2038]" pageId="95" pageNumber="96">
<materialsCitation accessionNumber="SMF 96266" collectingDate="2012-11-14" collectionCode="SMF" country="Mexico" county="Gunther Kohler" elevation="40" latitude="15.87038" location="Puerto Escondido" longLatPrecision="1" longitude="-97.10152" municipality="Punta Colorada" pageId="95" pageNumber="96" specimenCode="WGS84, GK-4116" specimenCount="1" stateProvince="Oaxaca" typeStatus="holotype">
<emphasis bold="true" box="[151,260,1250,1275]" pageId="95" pageNumber="96">
<typeStatus box="[151,260,1250,1275]" pageId="95" pageNumber="96">Holotype</typeStatus>
</emphasis>
.
<accessionNumber box="[282,426,1250,1275]" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF96266" pageId="95" pageNumber="96">
<collectionCode box="[282,338,1250,1275]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="95" pageNumber="96">SMF</collectionCode>
96266
</accessionNumber>
, an adult male from
<location LSID="urn:lsid:plazi:treatment:8A288798FF9FE75F7EC3FB81FE13C07E:075E6055FF9FE7667CECFB15FC5AC2CA" box="[696,905,1250,1275]" country="Mexico" county="Gunther Kohler" latitude="15.87038" longLatPrecision="1" longitude="-97.10152" municipality="Punta Colorada" name="Puerto Escondido" pageId="95" pageNumber="96" stateProvince="Oaxaca">Puerto Escondido</location>
,
<collectingMunicipality box="[925,1106,1250,1275]" pageId="95" pageNumber="96">Punta Colorada</collectingMunicipality>
(
<geoCoordinate box="[1129,1269,1250,1275]" degrees="15.87038" direction="north" orientation="latitude" pageId="95" pageNumber="96" precision="1" value="15.87038">15.87038°N</geoCoordinate>
,
<geoCoordinate box="[1289,1432,1250,1275]" degrees="97.10152" direction="west" orientation="longitude" pageId="95" pageNumber="96" precision="1" value="-97.10152">97.10152°W</geoCoordinate>
,
<specimenCode box="[151,245,1285,1310]" collectionCode="WGS" pageId="95" pageNumber="96">WGS84</specimenCode>
),
<quantity box="[267,329,1285,1310]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="95" pageNumber="96" unit="m" value="40.0">
<elevation box="[267,329,1285,1310]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="95" pageNumber="96" unit="m" value="40.0">40 m</elevation>
</quantity>
,
<collectingRegion box="[344,554,1285,1310]" country="Mexico" name="Oaxaca" pageId="95" pageNumber="96">Estado de Oaxaca</collectingRegion>
,
<collectingCountry box="[568,657,1285,1310]" name="Mexico" pageId="95" pageNumber="96">Mexico</collectingCountry>
; collected
<date box="[782,1004,1285,1310]" pageId="95" pageNumber="96" value="2012-11-14">
<collectingDate box="[782,1004,1285,1310]" pageId="95" pageNumber="96" value="2012-11-14">14 November 2012</collectingDate>
</date>
by
<collectingCounty box="[1052,1232,1285,1310]" pageId="95" pageNumber="96">Gunther Köhler</collectingCounty>
and Jesús García Grajales. Field tag number
<specimenCode box="[456,565,1320,1345]" collectionCode="GK-" pageId="95" pageNumber="96">GK-4116</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph blockId="95.[151,1437,1250,2038]" pageId="95" pageNumber="96">
<materialsCitation accessionNumber="SMF 96267, IBH 26567" collectingDate="2012-11-14" collectionCode="SMF, IBH" country="Mexico" county="Universidad del Mar" elevation="80" latitude="15.88934" location="All" longLatPrecision="1" longitude="-97.07703" municipality="Puerto Escondido" pageId="95" pageNumber="96" specimenCode="WGS84" specimenCount="1" stateProvince="Oaxaca" typeStatus="paratype">
<emphasis bold="true" box="[199,319,1354,1379]" pageId="95" pageNumber="96">
<typeStatus box="[199,319,1354,1379]" pageId="95" pageNumber="96">Paratypes</typeStatus>
</emphasis>
.
<location LSID="urn:lsid:plazi:treatment:8A288798FF9FE75F7EC3FB81FE13C07E:075E6055FF9FE7667F1AFABCFEA1C352" box="[334,370,1354,1379]" country="Mexico" county="Universidad del Mar" latitude="15.88934" longLatPrecision="1" longitude="-97.07703" municipality="Puerto Escondido" name="All" pageId="95" pageNumber="96" stateProvince="Oaxaca">All</location>
from
<collectingRegion box="[442,650,1354,1379]" country="Mexico" name="Oaxaca" pageId="95" pageNumber="96">Estado de Oaxaca</collectingRegion>
,
<collectingCountry box="[662,752,1354,1379]" name="Mexico" pageId="95" pageNumber="96">Mexico</collectingCountry>
:
<accessionNumber box="[764,902,1354,1379]" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF96267" pageId="95" pageNumber="96">
<collectionCode box="[764,820,1354,1379]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="95" pageNumber="96">SMF</collectionCode>
96267
</accessionNumber>
, same collecting data as holotype.
<accessionNumber box="[1302,1432,1354,1379]" httpUri="http://www.ncbi.nlm.nih.gov/protein/IBH26567" pageId="95" pageNumber="96">
<collectionCode box="[1302,1350,1355,1378]" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="95" pageNumber="96">IBH</collectionCode>
26567
</accessionNumber>
, from
<collectingMunicipality box="[213,415,1389,1414]" pageId="95" pageNumber="96">Puerto Escondido</collectingMunicipality>
,
<collectingCounty box="[428,662,1389,1414]" pageId="95" pageNumber="96">Universidad del Mar</collectingCounty>
(
<geoCoordinate box="[677,818,1389,1414]" degrees="15.88934" direction="north" orientation="latitude" pageId="95" pageNumber="96" precision="1" value="15.88934">15.88934°N</geoCoordinate>
,
<geoCoordinate box="[830,973,1389,1414]" degrees="97.07703" direction="west" orientation="longitude" pageId="95" pageNumber="96" precision="1" value="-97.07703">97.07703°W</geoCoordinate>
,
<specimenCode box="[985,1080,1389,1414]" collectionCode="WGS" pageId="95" pageNumber="96">WGS84</specimenCode>
),
<quantity box="[1100,1160,1389,1414]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="95" pageNumber="96" unit="m" value="80.0">
<elevation box="[1100,1160,1389,1414]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="95" pageNumber="96" unit="m" value="80.0">80 m</elevation>
</quantity>
</materialsCitation>
,
<materialsCitation accessionNumber="IBH 26560, SMF 96263" collectingDate="2012-11-14" collectionCode="IBH, SMF" collectorName="Huatulco" country="Mexico" elevation="90" latitude="15.78191" location="Gunther Kohler" longLatPrecision="1" longitude="-96.16257" municipality="Jesus Garcia Grajales." pageId="95" pageNumber="96" specimenCode="IBH 26560-64, SMF 96263-64" specimenCount="1" stateProvince="Oaxaca" typeStatus="paratype">
collected
<date pageId="95" pageNumber="96" value="2012-11-14">
<collectingDate pageId="95" pageNumber="96" value="2012-11-14">14 November 2012</collectingDate>
</date>
by
<location LSID="urn:lsid:plazi:treatment:8A288798FF9FE75F7EC3FB81FE13C07E:075E6055FF9FE7667F5CFA66FE13C398" box="[264,448,1424,1449]" country="Mexico" latitude="15.78191" longLatPrecision="1" longitude="-96.16257" municipality="Jesus Garcia Grajales." name="Gunther Kohler" pageId="95" pageNumber="96" stateProvince="Oaxaca">Gunther Köhler</location>
and
<collectingMunicipality box="[517,781,1424,1449]" pageId="95" pageNumber="96">Jesús García Grajales.</collectingMunicipality>
<specimenCode box="[795,973,1424,1449]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="95" pageNumber="96">
<accessionNumber box="[795,929,1424,1449]" httpUri="http://www.ncbi.nlm.nih.gov/protein/IBH26560" pageId="95" pageNumber="96">IBH 26560</accessionNumber>
64
</specimenCode>
,
<specimenCode box="[991,1176,1424,1449]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="95" pageNumber="96">
<accessionNumber box="[991,1133,1424,1449]" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF96263" pageId="95" pageNumber="96">SMF 96263</accessionNumber>
64
</specimenCode>
, from near
<collectorName box="[1327,1432,1424,1449]" pageId="95" pageNumber="96">Huatulco</collectorName>
,
<geoCoordinate box="[151,289,1458,1483]" degrees="15.78191" direction="north" orientation="latitude" pageId="95" pageNumber="96" precision="1" value="15.78191">15.78191°N</geoCoordinate>
,
<geoCoordinate box="[303,446,1458,1483]" degrees="96.16257" direction="west" orientation="longitude" pageId="95" pageNumber="96" precision="1" value="-96.16257">96.16257°W</geoCoordinate>
,
<quantity box="[459,521,1458,1483]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="95" pageNumber="96" unit="m" value="90.0">
<elevation box="[459,521,1458,1483]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="95" pageNumber="96" unit="m" value="90.0">90 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation accessionNumber="IBH 26993, SMF 93842" collectingDate="2012-11-12" collectionCode="IBH, SMF" collectorName="Parque National Huatulco" country="Mexico" elevation="10" latitude="15.73268" location="Gunther Kohler" longLatPrecision="1" longitude="-96.16357" municipality="Raul Gomez Trejo Perez." pageId="95" pageNumber="96" specimenCode="IBH 26993-94, SMF 93842-43, 96467-68" specimenCount="1" stateProvince="Oaxaca" typeStatus="paratype">
collected
<date box="[644,865,1458,1483]" pageId="95" pageNumber="96" value="2012-11-12">
<collectingDate box="[644,865,1458,1483]" pageId="95" pageNumber="96" value="2012-11-12">12 November 2012</collectingDate>
</date>
by
<location LSID="urn:lsid:plazi:treatment:8A288798FF9FE75F7EC3FB81FE13C07E:075E6055FF9FE7667DDAFA44FB92C3FA" box="[910,1089,1458,1483]" country="Mexico" latitude="15.73268" longLatPrecision="1" longitude="-96.16357" municipality="Raul Gomez Trejo Perez." name="Gunther Kohler" pageId="95" pageNumber="96" stateProvince="Oaxaca">Gunther Köhler</location>
and
<collectingMunicipality box="[1147,1437,1458,1483]" pageId="95" pageNumber="96">Raúl Gómez Trejo Pérez.</collectingMunicipality>
<specimenCode box="[151,326,1493,1518]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="95" pageNumber="96">
<accessionNumber box="[151,283,1493,1518]" httpUri="http://www.ncbi.nlm.nih.gov/protein/IBH26993" pageId="95" pageNumber="96">IBH 26993</accessionNumber>
94
</specimenCode>
,
<specimenCode box="[342,525,1493,1518]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="95" pageNumber="96">
<accessionNumber box="[342,483,1493,1518]" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF93842" pageId="95" pageNumber="96">SMF 93842</accessionNumber>
43
</specimenCode>
,
<specimenCode box="[541,654,1493,1518]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="95" pageNumber="96">9646768</specimenCode>
from
<collectorName box="[732,1036,1493,1518]" pageId="95" pageNumber="96">Parque National Huatulco</collectorName>
,
<geoCoordinate box="[1052,1191,1493,1518]" degrees="15.73268" direction="north" orientation="latitude" pageId="95" pageNumber="96" precision="1" value="15.73268">15.73268°N</geoCoordinate>
,
<geoCoordinate box="[1207,1350,1493,1518]" degrees="96.16357" direction="west" orientation="longitude" pageId="95" pageNumber="96" precision="1" value="-96.16357">96.16357°W</geoCoordinate>
,
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<elevation box="[1367,1432,1493,1518]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="95" pageNumber="96" unit="m" value="10.0">10 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation accessionNumber="IBH 26562, SMF 93842, IBH 26560, SMF 96267, IBH 26561" collectingDate="2013-02-28" collectionCode="IBH, SMF" country="Mexico" location="Gunther Kohler" municipality="Raul Gomez Trejo Perez." pageId="95" pageNumber="96" specimenCode="IBH 26562-63, SMF 93842-43, 96263-64, 96467, 26564, 26993, 96468, 26567, 26569" specimenCount="1" stateProvince="Oaxaca" typeStatus="paratype">
collected
<date box="[262,468,1528,1553]" pageId="95" pageNumber="96" value="2013-02-28">
<collectingDate box="[262,468,1528,1553]" pageId="95" pageNumber="96" value="2013-02-28">28 February 2013</collectingDate>
</date>
by
<location LSID="urn:lsid:plazi:treatment:8A288798FF9FE75F7EC3FB81FE13C07E:075E6055FF9FE7667C51FA0EFD6AC020" box="[517,697,1528,1553]" country="Mexico" municipality="Raul Gomez Trejo Perez." name="Gunther Kohler" pageId="95" pageNumber="96" stateProvince="Oaxaca">Gunther Köhler</location>
and
<collectingMunicipality box="[758,1053,1528,1553]" pageId="95" pageNumber="96">Raúl Gómez Trejo Pérez.</collectingMunicipality>
<specimenCode box="[1063,1236,1528,1553]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="95" pageNumber="96">
<accessionNumber box="[1063,1194,1528,1553]" httpUri="http://www.ncbi.nlm.nih.gov/protein/IBH26562" pageId="95" pageNumber="96">IBH 26562</accessionNumber>
63
</specimenCode>
,
<specimenCode box="[1251,1432,1528,1553]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="95" pageNumber="96">
<accessionNumber box="[1251,1390,1528,1553]" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF93842" pageId="95" pageNumber="96">SMF 93842</accessionNumber>
43
</specimenCode>
,
<specimenCode box="[151,266,1562,1587]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="95" pageNumber="96">9626364</specimenCode>
,
<specimenCode box="[277,347,1562,1587]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="95" pageNumber="96">96467</specimenCode>
are males,
<accessionNumber box="[473,601,1562,1587]" httpUri="http://www.ncbi.nlm.nih.gov/protein/IBH26560" pageId="95" pageNumber="96">
<collectionCode box="[473,521,1563,1586]" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="95" pageNumber="96">IBH</collectionCode>
26560
</accessionNumber>
,
<specimenCode box="[612,685,1562,1587]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="95" pageNumber="96">26564</specimenCode>
,
<specimenCode box="[695,768,1562,1587]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="95" pageNumber="96">26993</specimenCode>
,
<accessionNumber box="[779,915,1562,1587]" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF96267" pageId="95" pageNumber="96">
<collectionCode box="[779,835,1562,1587]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="95" pageNumber="96">SMF</collectionCode>
96267
</accessionNumber>
,
<specimenCode box="[926,996,1562,1587]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="95" pageNumber="96">96468</specimenCode>
are females,
<accessionNumber box="[1144,1272,1562,1587]" httpUri="http://www.ncbi.nlm.nih.gov/protein/IBH26561" pageId="95" pageNumber="96">
<collectionCode box="[1144,1192,1563,1586]" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="95" pageNumber="96">IBH</collectionCode>
26561
</accessionNumber>
,
<specimenCode box="[1283,1356,1562,1587]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="95" pageNumber="96">26567</specimenCode>
,
<specimenCode box="[1366,1436,1562,1587]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="95" pageNumber="96">26569</specimenCode>
are juveniles
</materialsCitation>
.
</paragraph>
<paragraph blockId="95.[151,1437,1250,2038]" pageId="95" pageNumber="96">
<emphasis bold="true" box="[199,438,1632,1657]" pageId="95" pageNumber="96">Referred specimens</emphasis>
<materialsCitation box="[438,1209,1631,1657]" collectionCode="UMMZ" collectorName="Puerto Angel" country="Mexico" location="Angeles Bay" pageId="95" pageNumber="96" specimenCode="UMMZ 84239" specimenCount="1" stateProvince="Oaxaca">
<emphasis bold="true" box="[466,562,1632,1657]" pageId="95" pageNumber="96">
<collectingCountry box="[466,557,1632,1657]" name="Mexico" pageId="95" pageNumber="96">Mexico</collectingCountry>
:
</emphasis>
<collectingRegion box="[573,662,1632,1657]" country="Mexico" name="Oaxaca" pageId="95" pageNumber="96">Oaxaca</collectingRegion>
:
<location LSID="urn:lsid:plazi:treatment:8A288798FF9FE75F7EC3FB81FE13C07E:075E6055FF9FE7667CF0F996FCEBC048" box="[676,824,1632,1657]" country="Mexico" name="Angeles Bay" pageId="95" pageNumber="96" stateProvince="Oaxaca">Angeles Bay</location>
[
<collectorName box="[841,1004,1632,1657]" pageId="95" pageNumber="96">Puerto Angel</collectorName>
?]:
<specimenCode box="[1037,1209,1632,1657]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="95" pageNumber="96" type="University or college">UMMZ 84239</specimenCode>
</materialsCitation>
;
<materialsCitation collectionCode="KU" country="Mexico" location="Puerto Angel" pageId="95" pageNumber="96" specimenCode="KU 38262" specimenCount="1" stateProvince="Oaxaca">
<location LSID="urn:lsid:plazi:treatment:8A288798FF9FE75F7EC3FB81FE13C07E:075E6055FF9FE7667A9CF997FAB6C048" box="[1224,1381,1632,1657]" country="Mexico" name="Puerto Angel" pageId="95" pageNumber="96" stateProvince="Oaxaca">Puerto Angel</location>
:
<specimenCode collectionCode="KU" country="USA" name="Biodiversity Institute, University of Kansas" pageId="95" pageNumber="96" type="University or college">KU 38262</specimenCode>
</materialsCitation>
;
<materialsCitation box="[236,788,1666,1691]" collectionCode="UMMZ" country="Mexico" location="Escondido Bay" pageId="95" pageNumber="96" specimenCode="UMMZ 84238" specimenCount="1" stateProvince="Oaxaca">
<location LSID="urn:lsid:plazi:treatment:8A288798FF9FE75F7EC3FB81FE13C07E:075E6055FF9FE7667EB8F974FE48C0AA" box="[236,411,1666,1691]" country="Mexico" name="Escondido Bay" pageId="95" pageNumber="96" stateProvince="Oaxaca">Escondido Bay</location>
:
<specimenCode box="[422,587,1666,1691]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="95" pageNumber="96" type="University or college">UMMZ 84238</specimenCode>
(
<specimenCount box="[602,780,1666,1691]" pageId="95" pageNumber="96" type="generic">four specimens</specimenCount>
)
</materialsCitation>
;
<materialsCitation box="[799,1127,1666,1691]" collectionCode="USNM" country="Mexico" location="Cacalotepec" pageId="95" pageNumber="96" specimenCode="USNM 123703" specimenCount="1" stateProvince="Oaxaca">
<location LSID="urn:lsid:plazi:treatment:8A288798FF9FE75F7EC3FB81FE13C07E:075E6055FF9FE7667D4BF974FC7FC0AA" box="[799,940,1666,1691]" country="Mexico" name="Cacalotepec" pageId="95" pageNumber="96" stateProvince="Oaxaca">Cacalotepec</location>
:
<specimenCode box="[951,1127,1666,1691]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="95" pageNumber="96" type="Museum">USNM 123703</specimenCode>
</materialsCitation>
;
<materialsCitation collectionCode="UIMNH" collectorName="Plan de la Campana" country="Mexico" location="Potrero Itierba Santa" pageId="95" pageNumber="96" specimenCode="UIMNH 40932" specimenCount="1" stateProvince="Oaxaca">
<location LSID="urn:lsid:plazi:treatment:8A288798FF9FE75F7EC3FB81FE13C07E:075E6055FF9FE7667A27F975FA8CC0AA" box="[1139,1375,1666,1691]" country="Mexico" name="Potrero Itierba Santa" pageId="95" pageNumber="96" stateProvince="Oaxaca">Potrero Itierba Santa</location>
,
<collectorName pageId="95" pageNumber="96">Plan de la Campana</collectorName>
:
<specimenCode box="[332,507,1701,1726]" collectionCode="UIMNH" country="USA" httpUri="http://grbio.org/cool/zrdp-fspx" name="University of Illinois, Museum of Natural History" pageId="95" pageNumber="96">UIMNH 40932</specimenCode>
</materialsCitation>
;
<materialsCitation box="[518,1328,1701,1726]" collectionCode="UIMNH" country="Mexico" elevation="174" location="Cuajinicuilapa" pageId="95" pageNumber="96" specimenCode="UIMNH 52898" specimenCount="1" stateProvince="Oaxaca">
<locationDeviation box="[518,751,1701,1726]" location="Rio Canoa" pageId="95" pageNumber="96">
<quantity box="[518,568,1701,1726]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="95" pageNumber="96" unit="mi" value="5.0">5 mi</quantity>
NW Río Canoa
</locationDeviation>
,
<quantity box="[762,826,1701,1726]" metricMagnitude="4" metricUnit="m" metricValue="5.3108352" pageId="95" pageNumber="96" unit="mi" value="33.0">33 mi</quantity>
ESE
<location LSID="urn:lsid:plazi:treatment:8A288798FF9FE75F7EC3FB81FE13C07E:075E6055FF9FE7667D2EF953FBF1C08F" box="[890,1058,1701,1726]" country="Mexico" name="Cuajinicuilapa" pageId="95" pageNumber="96" stateProvince="Oaxaca">Cuajinicuilapa</location>
:
<quantity box="[1068,1142,1701,1726]" metricMagnitude="2" metricUnit="m" metricValue="1.7399999999999998" pageId="95" pageNumber="96" unit="m" value="174.0">
<elevation box="[1068,1142,1701,1726]" metricMagnitude="2" metricUnit="m" metricValue="1.7399999999999998" pageId="95" pageNumber="96" unit="m" value="174.0">174 m</elevation>
</quantity>
:
<specimenCode box="[1153,1328,1701,1726]" collectionCode="UIMNH" country="USA" httpUri="http://grbio.org/cool/zrdp-fspx" name="University of Illinois, Museum of Natural History" pageId="95" pageNumber="96">UIMNH 52898</specimenCode>
</materialsCitation>
;
<materialsCitation collectionCode="KU" collectorName="N Pochutla" country="Mexico" elevation="230" location="San Jose Chacalapa" pageId="95" pageNumber="96" specimenCode="KU 38261" specimenCount="1" stateProvince="Oaxaca">
<location LSID="urn:lsid:plazi:treatment:8A288798FF9FE75F7EC3FB81FE13C07E:075E6055FF9FE7667B6FF953FEDCC0D0" country="Mexico" name="San Jose Chacalapa" pageId="95" pageNumber="96" stateProvince="Oaxaca">San José Chacalapa</location>
,
<quantity box="[281,352,1736,1761]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="95" pageNumber="96" unit="km" value="13.0">13 km</quantity>
on road
<collectorName box="[449,577,1736,1761]" pageId="95" pageNumber="96">N Pochutla</collectorName>
,
<quantity box="[588,662,1736,1761]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="95" pageNumber="96" unit="m" value="230.0">
<elevation box="[588,662,1736,1761]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="95" pageNumber="96" unit="m" value="230.0">230 m</elevation>
</quantity>
:
<specimenCode box="[673,793,1736,1761]" collectionCode="KU" country="USA" name="Biodiversity Institute, University of Kansas" pageId="95" pageNumber="96" type="University or college">KU 38261</specimenCode>
,
</materialsCitation>
<materialsCitation accessionNumber="SMF 96265" box="[803,940,1736,1761]" collectionCode="SMF" collectorName="N Pochutla" country="Mexico" elevation="230" isTailEnumeration="true" location="San Jose Chacalapa" pageId="95" pageNumber="96" specimenCount="1" stateProvince="Oaxaca">
<accessionNumber box="[803,940,1736,1761]" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF96265" pageId="95" pageNumber="96">
<collectionCode box="[803,859,1736,1761]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="95" pageNumber="96">SMF</collectionCode>
96265
</accessionNumber>
</materialsCitation>
;
<materialsCitation box="[950,1304,1736,1761]" collectionCode="UIMNH" country="Mexico" location="Pochutla" pageId="95" pageNumber="96" specimenCode="UIMNH 8903" specimenCount="1" stateProvince="Oaxaca">
<quantity box="[950,1001,1736,1761]" metricMagnitude="4" metricUnit="m" metricValue="1.2874752" pageId="95" pageNumber="96" unit="mi" value="8.0">8 mi</quantity>
E
<location LSID="urn:lsid:plazi:treatment:8A288798FF9FE75F7EC3FB81FE13C07E:075E6055FF9FE7667A53F93EFBBEC0D0" box="[1031,1133,1736,1761]" country="Mexico" name="Pochutla" pageId="95" pageNumber="96" stateProvince="Oaxaca">Pochutla</location>
:
<specimenCode box="[1143,1304,1736,1761]" collectionCode="UIMNH" country="USA" httpUri="http://grbio.org/cool/zrdp-fspx" name="University of Illinois, Museum of Natural History" pageId="95" pageNumber="96">UIMNH 8903</specimenCode>
</materialsCitation>
;
<materialsCitation collectionCode="UMMZ" collectorName="Puerto Escondido" country="Mexico" elevation="250" pageId="95" pageNumber="96" specimenCode="UMMZ 130953" specimenCount="1" stateProvince="Oaxaca">
<quantity box="[1315,1365,1736,1761]" metricMagnitude="3" metricUnit="m" metricValue="9.656064" pageId="95" pageNumber="96" unit="mi" value="6.0">6 mi</quantity>
N (on road)
<collectorName box="[218,421,1770,1795]" pageId="95" pageNumber="96">Puerto Escondido</collectorName>
,
<quantity box="[434,510,1770,1795]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="95" pageNumber="96" unit="m" value="250.0">
<elevation box="[434,510,1770,1795]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="95" pageNumber="96" unit="m" value="250.0">250 m</elevation>
</quantity>
:
<specimenCode box="[522,702,1770,1795]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="95" pageNumber="96" type="University or college">UMMZ 130953</specimenCode>
(
<specimenCount box="[718,892,1770,1795]" pageId="95" pageNumber="96" type="generic">two specimens</specimenCount>
)
</materialsCitation>
;
<materialsCitation box="[912,1431,1770,1795]" collectionCode="TCWC" country="Mexico" location="San Jose Chacalapa" pageId="95" pageNumber="96" specimenCode="TCWC 17348" specimenCount="1" stateProvince="Oaxaca">
<quantity box="[912,992,1770,1795]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="95" pageNumber="96" unit="km" value="1.5">1.5 km</quantity>
N
<location LSID="urn:lsid:plazi:treatment:8A288798FF9FE75F7EC3FB81FE13C07E:075E6055FF9FE7667A51F91CFB3AC132" box="[1029,1257,1770,1795]" country="Mexico" name="San Jose Chacalapa" pageId="95" pageNumber="96" stateProvince="Oaxaca">San José Chacalapa</location>
:
<specimenCode box="[1269,1431,1770,1795]" collectionCode="TCWC" country="USA" httpUri="http://grbio.org/cool/dj6p-xxc3" name="Texas Cooperative Wildlife Collection" pageId="95" pageNumber="96">TCWC 17348</specimenCode>
</materialsCitation>
;
<materialsCitation box="[151,648,1805,1830]" collectionCode="UIMNH" country="Mexico" location="Mira Leon" pageId="95" pageNumber="96" specimenCode="UIMNH 8904" specimenCount="1" stateProvince="Oaxaca">
<location LSID="urn:lsid:plazi:treatment:8A288798FF9FE75F7EC3FB81FE13C07E:075E6055FF9FE7667EC3F8FBFEC3C114" box="[151,272,1805,1830]" country="Mexico" name="Mira Leon" pageId="95" pageNumber="96" stateProvince="Oaxaca">Mira Leon</location>
,
<locationDeviation box="[284,476,1805,1830]" location="Huatulco" pageId="95" pageNumber="96">
<quantity box="[284,335,1805,1830]" metricMagnitude="3" metricUnit="m" metricValue="1.609344" pageId="95" pageNumber="96" unit="mi" value="1.0">1 mi</quantity>
N Huatulco
</locationDeviation>
:
<specimenCode box="[486,648,1805,1830]" collectionCode="UIMNH" country="USA" httpUri="http://grbio.org/cool/zrdp-fspx" name="University of Illinois, Museum of Natural History" pageId="95" pageNumber="96">UIMNH 8904</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="96" lastPageNumber="97" pageId="95" pageNumber="96" type="diagnosis">
<paragraph blockId="95.[151,1437,1250,2038]" lastBlockId="96.[151,1437,151,1147]" lastPageId="96" lastPageNumber="97" pageId="95" pageNumber="96">
<emphasis bold="true" box="[199,314,1840,1865]" pageId="95" pageNumber="96">Diagnosis</emphasis>
. A small to moderate-sized species (SVL in largest male 43.0 mm, largest female
<quantity box="[1252,1354,1840,1865]" metricMagnitude="-2" metricUnit="m" metricValue="4.95" pageId="95" pageNumber="96" unit="mm" value="49.5">49.5 mm</quantity>
) of the genus
<taxonomicName authority="(sensu Poe 2004)" baseAuthorityName="sensu Poe" baseAuthorityYear="2004" box="[234,540,1874,1899]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="95" pageNumber="96" phylum="Chordata" rank="genus">
<emphasis box="[234,306,1875,1899]" italics="true" pageId="95" pageNumber="96">Anolis</emphasis>
(sensu
<bibRefCitation author="Poe, S." box="[414,532,1874,1899]" pageId="95" pageNumber="96" pagination="37 - 89" refId="ref95161" refString="Poe, S. (2004) Phylogeny of anoles. Herpetological Monographs, 18, 37 - 89. http: // dx. doi. org / 10.1655 / 0733 - 1347 (2004) 018 [0037: poa] 2.0. co; 2" type="journal article" year="2004">Poe 2004</bibRefCitation>
)
</taxonomicName>
that differs from all Mexican and Central American anoles except
<emphasis italics="true" pageId="95" pageNumber="96">
<taxonomicName authorityName="Thominot" authorityYear="1887" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="95" pageNumber="96" phylum="Chordata" rank="species" species="boulengerianus">A. boulengerianus</taxonomicName>
,
<taxonomicName box="[340,469,1909,1933]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="95" pageNumber="96" phylum="Chordata" rank="species" species="carlliebi">A. carlliebi</taxonomicName>
,
<taxonomicName authorityName="Fitch" authorityYear="1978" box="[480,636,1910,1933]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="95" pageNumber="96" phylum="Chordata" rank="species" species="quercorum">A. quercorum</taxonomicName>
,
<taxonomicName authorityName="Davis" authorityYear="1954" box="[648,812,1909,1933]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="95" pageNumber="96" phylum="Chordata" rank="species" species="subocularis">A. subocularis</taxonomicName>
,
</emphasis>
and
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[873,1078,1910,1933]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="95" pageNumber="96" phylum="Chordata" rank="species" species="sacamecatensis">
<emphasis box="[873,1078,1910,1933]" italics="true" pageId="95" pageNumber="96">A. sacamecatensis</emphasis>
</taxonomicName>
by having a combination of (1) strongly keeled ventral scales; (2) usually a patch of three greatly enlarged supraocular scales; (3) 1320 rows of slightly to moderately enlarged dorsal scales that are smaller than ventral scales; (4) moderately long hind legs, longest toe of adpressed hind leg usually reaching to level of posterior border of eye or beyond, ratio shank length/ SVL 0.240.31; and (5) a large pinkish red dewlap in males and a small red or pink dewlap in females.
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="96" pageNumber="97" phylum="Chordata" rank="species" species="immaculogularis">
<emphasis italics="true" pageId="96" pageNumber="97">Anolis immaculogularis</emphasis>
</taxonomicName>
differs from
<emphasis box="[497,805,187,211]" italics="true" pageId="96" pageNumber="97">
<taxonomicName box="[497,628,187,211]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="96" pageNumber="97" phylum="Chordata" rank="species" species="carlliebi">A. carlliebi</taxonomicName>
,
<taxonomicName authorityName="Fitch" authorityYear="1978" box="[642,801,187,210]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="96" pageNumber="97" phylum="Chordata" rank="species" species="quercorum">A. quercorum</taxonomicName>
,
</emphasis>
and
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[865,1073,187,210]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="96" pageNumber="97" phylum="Chordata" rank="species" species="sacamecatensis">
<emphasis box="[865,1073,187,210]" italics="true" pageId="96" pageNumber="97">A. sacamecatensis</emphasis>
</taxonomicName>
by having longer legs with the longest toe of adpressed hind leg usually reaching to level of posterior border of eye or beyond, ratio shank length/ SVL 0.240.31 (vs. to ear opening or to a point between ear opening and eye, ratio shank length/SVL 0.200.26).
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[151,422,291,315]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="96" pageNumber="97" phylum="Chordata" rank="species" species="immaculogularis">
<emphasis box="[151,422,291,315]" italics="true" pageId="96" pageNumber="97">Anolis immaculogularis</emphasis>
</taxonomicName>
differs from
<taxonomicName authorityName="Davis" authorityYear="1954" box="[577,742,291,315]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="96" pageNumber="97" phylum="Chordata" rank="species" species="subocularis">
<emphasis box="[577,742,291,315]" italics="true" pageId="96" pageNumber="97">A. subocularis</emphasis>
</taxonomicName>
in male dewlap coloration in life (pinkish red without paler areas around gorgetals in
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[441,663,325,349]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="96" pageNumber="97" phylum="Chordata" rank="species" species="immaculogularis">
<emphasis box="[441,663,325,349]" italics="true" pageId="96" pageNumber="97">A. immaculogularis</emphasis>
</taxonomicName>
vs. pinkish red with paler areas around gorgetals in
<taxonomicName authorityName="Davis" authorityYear="1954" box="[1257,1420,325,349]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="96" pageNumber="97" phylum="Chordata" rank="species" species="subocularis">
<emphasis box="[1257,1420,325,349]" italics="true" pageId="96" pageNumber="97">A. subocularis</emphasis>
</taxonomicName>
). Also, in
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[245,466,360,384]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="96" pageNumber="97" phylum="Chordata" rank="species" species="immaculogularis">
<emphasis box="[245,466,360,384]" italics="true" pageId="96" pageNumber="97">A. immaculogularis</emphasis>
</taxonomicName>
, the subocular scales are usually broadly in contact with the supralabials (vs. subocular scales separated from supralabials by one scale row or these scales in contact in
<taxonomicName authorityName="Davis" authorityYear="1954" box="[1163,1333,395,419]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="96" pageNumber="97" phylum="Chordata" rank="species" species="subocularis">
<emphasis box="[1163,1333,395,419]" italics="true" pageId="96" pageNumber="97">A. subocularis</emphasis>
</taxonomicName>
).
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="96" pageNumber="97" phylum="Chordata" rank="species" species="immaculogularis">
<emphasis italics="true" pageId="96" pageNumber="97">Anolis immaculogularis</emphasis>
</taxonomicName>
differs from
<taxonomicName authorityName="Thominot" authorityYear="1887" box="[490,696,429,453]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="96" pageNumber="97" phylum="Chordata" rank="species" species="boulengerianus">
<emphasis box="[490,696,429,453]" italics="true" pageId="96" pageNumber="97">A. boulengerianus</emphasis>
</taxonomicName>
in male dewlap coloration in life (pinkish red without paler areas around gorgetals in
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[373,594,464,488]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="96" pageNumber="97" phylum="Chordata" rank="species" species="immaculogularis">
<emphasis box="[373,594,464,488]" italics="true" pageId="96" pageNumber="97">A. immaculogularis</emphasis>
</taxonomicName>
vs. orange yellow with paler areas around gorgetals in
<taxonomicName authorityName="Thominot" authorityYear="1887" box="[1213,1418,464,488]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="96" pageNumber="97" phylum="Chordata" rank="species" species="boulengerianus">
<emphasis box="[1213,1418,464,488]" italics="true" pageId="96" pageNumber="97">A. boulengerianus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="102" lastPageNumber="103" pageId="96" pageNumber="97" type="description">
<paragraph blockId="96.[151,1437,151,1147]" lastBlockId="98.[151,1436,151,410]" lastPageId="98" lastPageNumber="99" pageId="96" pageNumber="97">
<emphasis bold="true" box="[199,533,498,523]" pageId="96" pageNumber="97">
Description of the
<typeStatus box="[423,528,498,523]" pageId="96" pageNumber="97">holotype</typeStatus>
.
</emphasis>
Adult male as indicated by almost completely everted hemipenes and presence of large dewlap; SVL 44.0 mm; tail incomplete; tail moderately compressed in cross section, tail height
<quantity box="[1349,1437,533,558]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="96" pageNumber="97" unit="mm" value="1.7">1.7 mm</quantity>
and width
<quantity box="[276,368,567,592]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="96" pageNumber="97" unit="mm" value="1.3">1.3 mm</quantity>
; axilla to groin distance
<quantity box="[659,765,567,592]" metricMagnitude="-2" metricUnit="m" metricValue="1.77" pageId="96" pageNumber="97" unit="mm" value="17.7">17.7 mm</quantity>
; head length
<quantity box="[923,1028,567,592]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="96" pageNumber="97" unit="mm" value="11.8">11.8 mm</quantity>
, head length/SVL ratio 0.27; snout length
<quantity box="[228,318,602,627]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="96" pageNumber="97" unit="mm" value="5.2">5.2 mm</quantity>
; head width
<quantity box="[461,551,602,627]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="96" pageNumber="97" unit="mm" value="6.9">6.9 mm</quantity>
; longest toe of adpressed hind limb reaching to ear opening; shank length
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="96" pageNumber="97" unit="mm" value="11.8">11.8 mm</quantity>
, shank length/head length ratio 1.00; longest finger of extended forelimb reaching to mideye; longest finger of adpressed forelimb reaching to a point
<quantity box="[611,701,671,696]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="96" pageNumber="97" unit="mm" value="2.5">2.5 mm</quantity>
beyond anterior insertion of hind limbs. Dorsal head scales in internasal region keeled, uni- to tricarinate; other dorsal head scales rugose or keeled; 6 postrostrals; 7 scales between nasals; 1 elongate prenasal scale on each side, in contact with both rostral and first supralabial; circumnasal in contact with first supralabial on left side, separated from supralabial by one scale row on right side; scales in moderate prefrontal depression mostly rugose or weakly keeled; supraorbital semicircles well developed, 3 scales of each side in broad contact with each other; supraorbital disc composed of an oval patch of 3 greatly enlarged, rugose keeled scales, arranged in a single row; two complete rows of circumorbital scales separating enlarged supraorbital scale from supraorbital semicircles; 2 elongated, strongly overlapping superciliaries, anterior one much larger than posterior one, followed posteriorly by 3 to 4 roundish to squarish scales of moderate size; 2 to 3 rows of small keeled scales extending between enlarged supraorbitals and superciliaries; parietal depression shallow; interparietal scale well developed, 1.7 x
<quantity box="[709,796,1018,1043]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="96" pageNumber="97" unit="mm" value="1.2">1.2 mm</quantity>
(length x width), surrounded by scales of moderate size; 1 scale present between interparietal and supraorbital semicircles; canthal ridge distinct, composed of 3 large (second largest) and 4 small anterior canthal scales, posteriormost canthal scale divided on left side; 8 scales present between second canthals; 11 scales present between posterior canthals; 41 (right) 41 (left) mostly keeled loreal scales in a maximum of 6 horizontal rows; 4 to 6 keeled subocular scales arranged in a single row, one scale horizontally divided on right side; 7 (right) 6 (left) supralabials to level below center of eye; 1 subocular in contact with 1 supralabial; ear opening 0.5 x
<quantity box="[595,685,1715,1740]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="97" pageNumber="98" unit="mm" value="1.1">1.1 mm</quantity>
(length x height); mental distinctly wider than long, completely divided medially, bordered posteriorly by 4 postmentals, outer ones moderately larger than median ones; 6 infralabials to level below center of eye; no greatly enlarged sublabials on each side; keeled granular scales present on chin and throat; dewlap large, extending from level below anterior margin of eye to level of chest; 89 horizontal gorgetal-sternal rows with 816 scales per row; modal number of marginal pairs 24; dorsum of body with strongly keeled, subimbricate, non-mucronate scales; about 14 medial rows slightly to moderately enlarged; largest dorsal scales about 0.48 x
<quantity box="[524,623,1915,1940]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" pageId="97" pageNumber="98" unit="mm" value="0.39">0.39 mm</quantity>
(length x width); about 24 medial dorsal scales in one head length; about 37 medial dorsal scales between levels of axilla and groin; lateral scales keeled, granular and more or less homogeneous in size, average size
<quantity box="[586,694,1982,2007]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="97" pageNumber="98" unit="mm" value="0.25">0.25 mm</quantity>
in diameter; ventrals at midbody strongly keeled, imbricate, mucronate, about 0.55 x
<quantity box="[438,540,151,176]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="98" pageNumber="99" unit="mm" value="0.4">0.40 mm</quantity>
(length x width); about 34 medial ventral scales in one head length; about 40 medial ventral scales between levels of axilla and groin; 106 scales around midbody; all caudal scales keeled; caudal middorsal scales slightly enlarged, not forming a crest; lateral caudal scales without whorls of enlarged scales, although an indistinct division in segments is discernible; a pair of greatly enlarged postcloacal scales present, about
<quantity box="[323,412,285,310]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="98" pageNumber="99" unit="mm" value="0.9">0.9 mm</quantity>
wide; no tube-like axillary pocket present; scales on dorsal surface of forelimb weakly keeled; digital pads dilated, dilated pad about 3 times width of non-dilated distal phalanx; distal phalanx narrower than and raised from dilated pad; 22 lamellae under phalanges IIIV of Toe IV of hind limbs; 7 (right)6 (left) scales under distal phalanx of Toe IV of hind limbs.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931117" ID-Zenodo-Dep="4931117" httpUri="https://zenodo.org/record/4931117/files/figure.png" pageId="96" pageNumber="97" startId="96.[151,250,1949,1971]" targetBox="[252,1338,1193,1915]" targetPageId="96">
<paragraph blockId="96.[151,1436,1948,2001]" pageId="96" pageNumber="97">
<emphasis bold="true" box="[151,288,1949,1971]" pageId="96" pageNumber="97">FIGURE 77.</emphasis>
Dewlaps of
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[416,659,1948,1970]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="96" pageNumber="97" phylum="Chordata" rank="species" species="immaculogularis">
<emphasis box="[416,659,1948,1970]" italics="true" pageId="96" pageNumber="97">Anolis immaculogularis</emphasis>
</taxonomicName>
in life. (a) SMF 96264; (b) IBH 26562; (c) SMF 96263; (d) SMF 96467; (e) IBH 26564; (f) IBH 26560. (ad) are males, (ef) are females.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931119" ID-Zenodo-Dep="4931119" httpUri="https://zenodo.org/record/4931119/files/figure.png" pageId="97" pageNumber="98" startId="97.[151,250,1526,1548]" targetBox="[155,1436,198,1504]" targetPageId="97">
<paragraph blockId="97.[151,1436,1526,1579]" pageId="97" pageNumber="98">
<emphasis bold="true" box="[151,287,1526,1548]" pageId="97" pageNumber="98">FIGURE 78.</emphasis>
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[293,536,1526,1548]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="97" pageNumber="98" phylum="Chordata" rank="species" species="immaculogularis">
<emphasis box="[293,536,1526,1548]" italics="true" pageId="97" pageNumber="98">Anolis immaculogularis</emphasis>
</taxonomicName>
in life. (a) SMF 96263; (b) SMF 96467; (c) SMF 96467; (d) IBH 26564; (e) IBH 26560; (f) IBH 26693. (ac) are males, (df) are females.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931121" ID-Zenodo-Dep="4931121" httpUri="https://zenodo.org/record/4931121/files/figure.png" pageId="98" pageNumber="99" startId="98.[151,250,1785,1807]" targetBox="[157,1436,459,1765]" targetPageId="98">
<paragraph blockId="98.[151,1435,1785,1868]" pageId="98" pageNumber="99">
<emphasis bold="true" box="[151,288,1785,1807]" pageId="98" pageNumber="99">FIGURE 79.</emphasis>
Holotype of
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[423,667,1785,1807]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="98" pageNumber="99" phylum="Chordata" rank="species" species="immaculogularis">
<emphasis box="[423,667,1785,1807]" italics="true" pageId="98" pageNumber="99">Anolis immaculogularis</emphasis>
</taxonomicName>
(SMF 96266): (a) dorsal view; (b) lateral view; (c) ventral view; (d) lateral view of head; (e) dorsal view of head; (f) ventral view of head. Scale bars equal 5.0 mm in (ac) and 1.0 mm in (df), respectively.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931123" ID-Zenodo-Dep="4931123" box="[151,1372,1530,1552]" httpUri="https://zenodo.org/record/4931123/files/figure.png" pageId="99" pageNumber="100" startId="99.[151,250,1530,1552]" targetBox="[153,1436,195,1502]" targetPageId="99">
<paragraph blockId="99.[151,1372,1530,1552]" box="[151,1372,1530,1552]" pageId="99" pageNumber="100">
<emphasis bold="true" box="[151,288,1530,1552]" pageId="99" pageNumber="100">FIGURE 80.</emphasis>
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[293,536,1530,1552]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="99" pageNumber="100" phylum="Chordata" rank="species" species="immaculogularis">
<emphasis box="[293,536,1530,1552]" italics="true" pageId="99" pageNumber="100">Anolis immaculogularis</emphasis>
</taxonomicName>
: head scalation in (a,c,e) IBH 26564; (b,d,f) IBH 26562. Scale bars equal 1.0 mm.
</paragraph>
</caption>
<paragraph blockId="99.[151,1437,1595,1968]" pageId="99" pageNumber="100">Coloration after eleven months preservation in 70% ethanol was recorded as follows: Dorsal surfaces of head, body, limbs, and tail Drab (19); eye region Grayish Horn Color (268); ventral surfaces of head and body Pale Buff (1); ventral surfaces of limbs and tail Pale Pinkish Buff (3); finger and toe pads Drab (19); anterior portion of dewlap Salmon Color (251) grading into Pale Buff (1) on posterior portion.</paragraph>
<paragraph blockId="99.[151,1437,1595,1968]" pageId="99" pageNumber="100">
<emphasis bold="true" box="[199,317,1735,1760]" pageId="99" pageNumber="100">Variation.</emphasis>
The
<typeStatus box="[374,480,1736,1760]" pageId="99" pageNumber="100">paratypes</typeStatus>
agree well with the
<typeStatus box="[705,802,1735,1760]" pageId="99" pageNumber="100">holotype</typeStatus>
in general appearance, morphometrics and scalation (see Table 16). Variation was evident in some scalation characters as follows: The number of slightly to moderately enlarged dorsal scale rows varies from 12 to 17. All but
<specimenCount box="[770,922,1804,1829]" pageId="99" pageNumber="100" type="generic">one specimen</specimenCount>
have a single elongate prenasal. Suboculars in contact with supralabials (14 suboculars in contact with 15 supralabials) in all specimens examined. Of 14 individuals, the longest toe of adpressed hind leg reaching to a point between ear opening and eye in 14.3%, to posterior border of eye in 50.0%, to mideye in 28.6%, and to anterior border of eye in 7.1%. All males have a pair of greatly enlarged postcloacal scales. For variation in selected scalation and morphometric characters see
<tableCitation box="[1299,1398,1943,1968]" captionStart="TABLE 10" captionStartId="109.[151,239,152,175]" captionTargetPageId="109" captionText="TABLE 10. Selected measurements, proportions and scale characters of the species related to Anolis boulengerianus. Range is followed by mean value and standard deviation in parentheses. For abbreviations see text." httpUri="http://table.plazi.org/id/56FE6606FFADE7547EC3FF6EFB6BC6E3" pageId="99" pageNumber="100" tableUuid="56FE6606FFADE7547EC3FF6EFB6BC6E3">Table 10</tableCitation>
.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931125" ID-Zenodo-Dep="4931125" httpUri="https://zenodo.org/record/4931125/files/figure.png" pageId="100" pageNumber="101" startId="100.[151,250,1949,1971]" targetBox="[151,1434,181,1930]" targetPageId="100">
<paragraph blockId="100.[151,1436,1949,2033]" pageId="100" pageNumber="101">
<emphasis bold="true" box="[151,287,1949,1971]" pageId="100" pageNumber="101">FIGURE 81.</emphasis>
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[293,536,1949,1971]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="100" pageNumber="101" phylum="Chordata" rank="species" species="immaculogularis">
<emphasis box="[293,536,1949,1971]" italics="true" pageId="100" pageNumber="101">Anolis immaculogularis</emphasis>
</taxonomicName>
: nasal region in (a) IBH 26562; (b) IBH 26564; superciliary region in (c) IBH 26560; (d) IBH 26564; chin region in (e) IBH 26560; (f) IBH 26564; cloacal region in (g) SMF 93843; (h) SMF 93842. Scale bars equal 1.0 mm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931127" ID-Zenodo-Dep="4931127" httpUri="https://zenodo.org/record/4931127/files/figure.png" pageId="101" pageNumber="102" startId="101.[151,250,1525,1547]" targetBox="[156,1436,193,1499]" targetPageId="101">
<paragraph blockId="101.[151,1436,1525,1578]" pageId="101" pageNumber="102">
<emphasis bold="true" box="[151,288,1525,1547]" pageId="101" pageNumber="102">FIGURE 82.</emphasis>
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[294,538,1525,1547]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="101" pageNumber="102" phylum="Chordata" rank="species" species="immaculogularis">
<emphasis box="[294,538,1525,1547]" italics="true" pageId="101" pageNumber="102">Anolis immaculogularis</emphasis>
</taxonomicName>
: dorsal scales in (a) IBH 26562; (b) IBH 26560; flank scales in (c) IBH 26564; (d) IBH 26560; ventral scales in (e) IBH 26564; (f) IBH 26560. Scale bars equal 1.0 mm.
</paragraph>
</caption>
<paragraph blockId="101.[151,1436,1648,1906]" pageId="101" pageNumber="102">
The coloration in life of an adult male from near Huatulco (
<accessionNumber box="[904,1045,1648,1673]" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF96263" pageId="101" pageNumber="102">SMF 96263</accessionNumber>
) was recorded as follows: Dorsal ground color Cinnamon Drab (50) with Raw Umber (280) chevrons and a Brownish Olive (276) interorbital bar; lateral stripes Tawny Olive (17) edged above and below by series of Hazel (26) streaks; dorsal surface of limbs and tail Drab (19) with Buff (15) bands; ventral surfaces of body, limbs and tail Pale Buff (1) with Burnt Sienna mottling on ventral surface of head; dewlap Pinkish Flesh Color (253) with Pale Buff (1) gorgetals and Pale Pinkish Buff (3) marginal; iris Raw Umber (22). The coloration in life of the dewlap of an adult male from Puerto Escondido (
<accessionNumber box="[289,431,1848,1873]" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF96266" pageId="101" pageNumber="102">SMF 96266</accessionNumber>
) was recorded as follows: Dewlap ground color Peach Red (70) with Chamois (84) gorgetals and marginals.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931129" ID-Zenodo-Dep="4931129" httpUri="https://zenodo.org/record/4931129/files/figure.png" pageId="102" pageNumber="103" startId="102.[151,250,793,815]" targetBox="[151,1436,193,772]" targetPageId="102">
<paragraph blockId="102.[151,1436,793,845]" pageId="102" pageNumber="103">
<emphasis bold="true" box="[151,290,793,815]" pageId="102" pageNumber="103">FIGURE 83.</emphasis>
Hemipenis of
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[446,691,793,815]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="102" pageNumber="103" phylum="Chordata" rank="species" species="immaculogularis">
<emphasis box="[446,691,793,815]" italics="true" pageId="102" pageNumber="103">Anolis immaculogularis</emphasis>
</taxonomicName>
(SMF 93842): (left) sulcate view; (center) asulcate view; (right) lateral view. Scale bar equals 1.0 mm.
</paragraph>
</caption>
<paragraph blockId="102.[151,1436,926,1615]" pageId="102" pageNumber="103">
The coloration in life of and adult female from near Huatulco (
<accessionNumber box="[922,1053,926,951]" httpUri="http://www.ncbi.nlm.nih.gov/protein/IBH26560" pageId="102" pageNumber="103">IBH 26560</accessionNumber>
) was recorded as follows: Dorsal ground color Drab (19) with Yellow Ocher (14) middorsal stripe and a Clay Color (18) interorbital bar; dorsal surface of limbs Amber (51) with faint Salmon Color (83) grading into Cinnamon-Drab (50) distally; ventral surface of limbs and tail Tawny Olive; dewlap Scarlet (69) with Pale Buff (1) gorgetals and Chamois (84) marginal; iris Burnt Sienna (38).
</paragraph>
<paragraph blockId="102.[151,1436,926,1615]" pageId="102" pageNumber="103">
The completely everted hemipenis of
<accessionNumber box="[660,801,1101,1126]" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF93842" pageId="102" pageNumber="103">SMF 93842</accessionNumber>
(
<figureCitation box="[824,912,1101,1126]" captionStart="FIGURE 83" captionStartId="102.[151,250,793,815]" captionTargetBox="[151,1436,193,772]" captionTargetId="figure-465@102.[151,1436,193,772]" captionTargetPageId="102" captionText="FIGURE 83. Hemipenis of Anolis immaculogularis (SMF 93842): (left) sulcate view; (center) asulcate view; (right) lateral view. Scale bar equals 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4931129" httpUri="https://zenodo.org/record/4931129/files/figure.png" pageId="102" pageNumber="103">Fig. 83</figureCitation>
) is a small, slightly bilobate organ; sulcus spermaticus bordered by well developed sulcal lips and opening into two apical fields void of ornamentation, one on each lobe; a finger-like asulcate processus and an indistinct asulcate ridge present; no surface ornamentation discernible.
</paragraph>
</subSubSection>
<subSubSection pageId="102" pageNumber="103" type="etymology">
<paragraph blockId="102.[151,1436,926,1615]" pageId="102" pageNumber="103">
<emphasis bold="true" box="[199,327,1240,1265]" pageId="102" pageNumber="103">Etymology</emphasis>
. The name
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[469,659,1241,1265]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="102" pageNumber="103" phylum="Chordata" rank="species" species="immaculogularis">
<emphasis box="[469,659,1241,1265]" italics="true" pageId="102" pageNumber="103">immaculogularis</emphasis>
</taxonomicName>
is a compound adjective derived from
<emphasis box="[1121,1267,1241,1265]" italics="true" pageId="102" pageNumber="103">immaculosus</emphasis>
(Latin for not spotted) and
<emphasis box="[295,345,1276,1300]" italics="true" pageId="102" pageNumber="103">gula</emphasis>
(Latin for throat) referring to male dewlap that lacks pale spots around the gorgetals, a diagnostic character to distinguish
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[419,687,1311,1335]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="102" pageNumber="103" phylum="Chordata" rank="species" species="immaculogularis">
<emphasis box="[419,687,1311,1335]" italics="true" pageId="102" pageNumber="103">Anolis immaculogularis</emphasis>
</taxonomicName>
from its supposed closest relative,
<taxonomicName authorityName="Thominot" authorityYear="1887" box="[1082,1287,1311,1335]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="102" pageNumber="103" phylum="Chordata" rank="species" species="boulengerianus">
<emphasis box="[1082,1287,1311,1335]" italics="true" pageId="102" pageNumber="103">A. boulengerianus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph blockId="102.[151,1436,926,1615]" pageId="102" pageNumber="103">
<emphasis bold="true" box="[199,475,1345,1370]" pageId="102" pageNumber="103">Natural History Notes:</emphasis>
Around Huatulco and Puerto Escondido we encountered
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[1129,1351,1346,1370]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="102" pageNumber="103" phylum="Chordata" rank="species" species="immaculogularis">
<emphasis box="[1129,1351,1346,1370]" italics="true" pageId="102" pageNumber="103">A. immaculogularis</emphasis>
</taxonomicName>
mostly in forest edge situations and in open, bushy forest habitats; here most specimens were found while these were sleeping on branches and leaves at 0.4 to 2.0 m above the ground.
</paragraph>
<paragraph blockId="102.[151,1436,926,1615]" pageId="102" pageNumber="103">
<emphasis bold="true" box="[199,711,1450,1475]" pageId="102" pageNumber="103">Geographic Distribution and Conservation</emphasis>
. As currently known,
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[961,1229,1450,1474]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="102" pageNumber="103" phylum="Chordata" rank="species" species="immaculogularis">
<emphasis box="[961,1229,1450,1474]" italics="true" pageId="102" pageNumber="103">Anolis immaculogularis</emphasis>
</taxonomicName>
is restricted to the Pacific versant of the Mexican State of
<collectingRegion box="[607,691,1485,1510]" country="Mexico" name="Oaxaca" pageId="102" pageNumber="103">Oaxaca</collectingRegion>
from about the town of Huatulco to at least the region of Puerto Escondido at elevations between 10 and
<quantity box="[632,738,1520,1545]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="102" pageNumber="103" unit="m" value="230.0">230 masl</quantity>
(
<figureCitation box="[758,846,1520,1545]" captionStart="FIGURE 76" captionStartId="95.[151,250,993,1015]" captionTargetBox="[328,1272,154,976]" captionTargetId="figure-414@95.[317,1281,145,985]" captionTargetPageId="95" captionText="FIGURE 76. Map indicating collecting localities of species related to Anolis boulengerianus. Each symbol can represent one or more adjacent localities. Black circles: Anolis boulengerianus; white circles: A. immaculogularis; black squares: A. subocularis." figureDoi="http://doi.org/10.5281/zenodo.4931115" httpUri="https://zenodo.org/record/4931115/files/figure.png" pageId="102" pageNumber="103">Fig. 76</figureCitation>
). Given its usual abundance wherever this species occurs, it seems justified to classify
<taxonomicName authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[558,780,1555,1579]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="102" pageNumber="103" phylum="Chordata" rank="species" species="immaculogularis">
<emphasis box="[558,780,1555,1579]" italics="true" pageId="102" pageNumber="103">A. immaculogularis</emphasis>
</taxonomicName>
as Least Concern based on the IUCN Red List Categories and Criteria (
<bibRefCitation author="IUCN" box="[298,436,1590,1615]" pageId="102" pageNumber="103" refId="ref91603" refString="IUCN (2012) IUCN Red List of Threatened Species. Version 2012.2. Available from: http: // www. iucnredlist. org (accessed 11 January 2013)" type="url" year="2012">IUCN 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>