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<document id="E5AB58EFB5F7DBD532A03A4F2DD4852E" ID-CLB-Dataset="223919" ID-DOI="10.11646/zootaxa.3862.1.1" ID-GBIF-Dataset="bebe1ef9-e8d6-459a-a102-8b9989b465f1" ID-ISSN="1175-5326" ID-Zenodo-Dep="4930959" ID-ZooBank="3FA375FE-E4E0-4509-BE02-EE5E786B07C6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623452360688" checkinUser="felipe" docAuthor="Köhler, Gunther, Pérez, Raúl Gómez Trejo, Petersen, Claus Bo P., Méndez, Fausto R. &amp; Cruz, De La" docDate="2014" docId="8A288798FF94E7657EC3FE2AFEBCC156" docLanguage="en" docName="zt03862p210.pdf.imf" docOrigin="Zootaxa 3862 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Anolis quercorum Fitch 1978" docType="treatment" docVersion="12" lastPageNumber="93" masterDocId="7611FFE0FFC0E7397E54FFF6FFD3C631" masterDocTitle="A revision of the Mexican Anolis (Reptilia, Squamata, Dactyloidae) from the Pacific versant west of the Isthmus de Tehuantepec in the states of Oaxaca, Guerrero, and Puebla, with the description of six new species" masterLastPageNumber="210" masterPageNumber="1" pageNumber="85" updateTime="1732602322999" updateUser="ExternalLinkService">
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<mods:title id="16692A108F6425AAE7B54A39FDE938F5">A revision of the Mexican Anolis (Reptilia, Squamata, Dactyloidae) from the Pacific versant west of the Isthmus de Tehuantepec in the states of Oaxaca, Guerrero, and Puebla, with the description of six new species</mods:title>
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<mods:namePart id="1D4C3C3457FA037E428D8B3E18CF1890">Pérez, Raúl Gómez Trejo</mods:namePart>
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<mods:namePart id="A12C4FA0F4483C48354630C7D8BA3F45">Petersen, Claus Bo P.</mods:namePart>
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<mods:namePart id="104F7F460D8537432DE73EBE16AA93E4">Méndez, Fausto R.</mods:namePart>
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<taxonomicName id="C5814D0DFF94E76D7EC3FE2AFE2FC7C7" authority="Fitch 1978" authorityName="Fitch" authorityYear="1978" box="[151,508,476,502]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="quercorum">
<emphasis id="30F5EA9CFF94E76D7EC3FE2AFE2FC7C7" bold="true" box="[151,508,476,502]" pageId="84" pageNumber="85">
<emphasis id="30F5EA9CFF94E76D7EC3FE2AFEA1C7C7" bold="true" box="[151,370,476,502]" italics="true" pageId="84" pageNumber="85">Anolis quercorum</emphasis>
<bibRefCitation id="66104B7FFF94E76D7F2DFE2AFE2FC7C7" author="Fitch, H. S." box="[377,508,476,502]" pageId="84" pageNumber="85" pagination="1 - 15" refId="ref89954" refString="Fitch, H. S. (1978) Two new anoles (Reptilia: Iguanidae) from Oaxaca with comments on other Mexican species. Contributions in Biology and Geology Milwaukee Public Museum, 20, 1 - 15." type="journal article" year="1978">Fitch 1978</bibRefCitation>
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<taxonomicName id="C5814D0DFF94E76D7EC3FDD4FE37C409" authority="Fitch 1978: 6" authorityName="Fitch" authorityPageNumber="6" authorityYear="1978" box="[151,484,546,568]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="quercorum">
<emphasis id="30F5EA9CFF94E76D7EC3FDD4FE9DC409" box="[151,334,546,568]" italics="true" pageId="84" pageNumber="85">Anolis quercorum</emphasis>
<bibRefCitation id="66104B7FFF94E76D7F03FDD4FE37C409" author="Fitch, H. S." box="[343,484,546,568]" pageId="84" pageNumber="85" pagination="1 - 15" refId="ref89954" refString="Fitch, H. S. (1978) Two new anoles (Reptilia: Iguanidae) from Oaxaca with comments on other Mexican species. Contributions in Biology and Geology Milwaukee Public Museum, 20, 1 - 15." type="journal article" year="1978">Fitch 1978: 6</bibRefCitation>
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; type locality: “
<quantity id="C5799B6BFF94E76D7CD8FDD4FD1CC409" box="[652,719,546,568]" metricMagnitude="4" metricUnit="m" metricValue="2.6" pageId="84" pageNumber="85" unit="km" value="26.0">26 km</quantity>
SE Nochixtlán (
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NW Cuesta Blanca, Highway 190),
<collectingRegion id="C045F86CFF94E76D7B1DFDD4FA4BC409" box="[1353,1432,546,568]" country="Mexico" name="Oaxaca" pageId="84" pageNumber="85">Oaxaca</collectingRegion>
,
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”.
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: KU 176050.
<bibRefCitation id="66104B7FFF94E76D7C4AFDB7FD5AC466" author="Lieb, C. S." box="[542,649,577,599]" pageId="84" pageNumber="85" refId="ref93186" refString="Lieb, C. S. (1981) Biochemical and Karyological Systematics of the Mexican Lizards of the Anolis gadovi and A. nebulosus Species Groups (Reptilia: Iguanidae). Unpublished Ph. D. Dissertation, University of California, Los Angeles, 308 pp." type="book" year="1981">Lieb 1981</bibRefCitation>
,
<bibRefCitation id="66104B7FFF94E76D7CC0FDB7FC8CC466" author="Fitch, H. S. &amp; Hillis, D. M." box="[660,863,577,599]" pageId="84" pageNumber="85" pagination="315 - 323" refId="ref90154" refString="Fitch, H. S. &amp; Hillis, D. M. (1984) The Anolis dewlap: Interspecific variability and morphological associations with habitat. Copeia, 1984 (2), 315 - 323. http: // dx. doi. org / 10.2307 / 1445187" type="journal article" year="1984">Fitch &amp; Hillis 1984</bibRefCitation>
,
<bibRefCitation id="66104B7FFF94E76D7D3EFDB7FBE0C466" author="Gorman, G. C. &amp; Lieb, C. S. &amp; Harwood, R. H." box="[874,1075,577,599]" pageId="84" pageNumber="85" pagination="145 - 152" refId="ref91071" refString="Gorman, G. C., Lieb, C. S. &amp; Harwood, R. H. (1984) The relationships of Anolis gadovi: Albumin immunological evidence. Caribbean Journal of Science, 20 (3 - 4), 145 - 152." type="journal article" year="1984">
Gorman
<emphasis id="30F5EA9CFF94E76D7D90FDB4FC2BC466" box="[964,1016,577,599]" italics="true" pageId="84" pageNumber="85">et al.</emphasis>
1984
</bibRefCitation>
, Flores-Villela 1993,
<bibRefCitation id="66104B7FFF94E76D7B45FDB7FE8DC447" author="Flores-Villela, O. &amp; Gerez, P." pageId="84" pageNumber="85" refId="ref90370" refString="Flores-Villela, O. &amp; Gerez, P. (1994) Biodiversidad y conservacion en Mexico: vertebrados, vegetacion y uso del suelo. Secunda Edicion. Conision Nacional para el Conocimiento y uso de la Biodiversidad y Universidad Nacional Autonoma de Mexico, Mexico, D. F., 439 pp." type="book" year="1994">Flores-Villela &amp; Gerez 1994</bibRefCitation>
,
<bibRefCitation id="66104B7FFF94E76D7F3FFD96FE0BC447" author="Lieb, C. S." box="[363,472,608,630]" pageId="84" pageNumber="85" pagination="146 - 157" refId="ref93236" refString="Lieb, C. S. (1995) Preliminary key to the anole lizards of Mexico. In: Flores-Villela, O. A., Mendoza Quijano, F. &amp; Gonzalez Porter, G. (Eds.), Recopilacion de Caves para la Determinacion de Anfibios y Reptiles de Mexico. Publicaciones Especiales del Museo de Zoologia, Universidad Nacional Autonoma de Mexico, pp. 146 - 157." type="book chapter" year="1995">Lieb 1995</bibRefCitation>
,
<bibRefCitation id="66104B7FFF94E76D7FB1FD96FD89C447" author="Liner, E. A." box="[485,602,608,630]" pageId="84" pageNumber="85" pagination="1 - 60" refId="ref93458" refString="Liner, E. A. (2007) A checklist of the amphibians and reptiles of Mexico. Occasional Papers of the Museum of Natural Science, Louisiana State University, 80, 1 - 60." type="journal article" year="2007">Liner 2007</bibRefCitation>
,
<bibRefCitation id="66104B7FFF94E76D7C3CFD96FC6BC447" author="Flaschendrager, A. &amp; Wijffels, L. C. M." box="[616,952,608,630]" pageId="84" pageNumber="85" refId="ref90663" refString="Flaschendrager, A. &amp; Wijffels, L. C. M. (2009) Anolis. Im Biotop und Terrarium. Natur und Tier - Verlag, Munster, 319 pp." type="book" year="2009">Fläschendräger &amp; Wijffels 2009</bibRefCitation>
,
<bibRefCitation id="66104B7FFF94E76D7D91FD96FB1EC447" author="Flores-Villela, O. &amp; Canseco-Marquez, L. &amp; Ochoa-Ochoa, L. M." box="[965,1229,608,630]" pageId="84" pageNumber="85" pagination="303 - 321" refId="ref90290" refString="Flores-Villela, O., Canseco-Marquez, L. &amp; Ochoa-Ochoa, L. M. (2010) Geographic distribution and conservation of the herpetofauna of northern Mexico. In: Wilson, L. D., Townsend, J. H. &amp; Johnson, J. D., (Eds.), Conservation of Mesoamerican Amphibians and Reptiles. Eagle Mountain Press, Eagle Mountain, Utah, pp. 303 - 321." type="book chapter" year="2010">
Flores-Villela
<emphasis id="30F5EA9CFF94E76D7A0EFD97FB43C447" box="[1114,1168,608,630]" italics="true" pageId="84" pageNumber="85">et al.</emphasis>
2010
</bibRefCitation>
,
<bibRefCitation id="66104B7FFF94E76D7A8FFD96FE5EC4A5" author="Urbina-Cardona, J. N. &amp; Flores-Villela, O." pageId="84" pageNumber="85" pagination="1031 - 1041" refId="ref97767" refString="Urbina-Cardona, J. N. &amp; Flores-Villela, O. (2010) Ecological-niche modeling and prioritization of conservation-area networks for Mexican herpetofauna. Conservation Biology, 24 (84), 1031 - 1041. http: // dx. doi. org / 10.1111 / j. 1523 - 1739.2009.01432. x" type="journal article" year="2010">Urbina-Cardona &amp; Flores-Villela 2010</bibRefCitation>
,
<bibRefCitation id="66104B7FFF94E76D7FC3FD88FD70C4A5" author="Wilson, L. D. &amp; Townsend, J. H." box="[407,675,638,660]" pageId="84" pageNumber="85" pagination="761 - 812" refId="ref98308" refString="Wilson, L. D., Townsend, J. H. (2010) The herpetofauna of Mesoamerica: biodiversity significance, conservation status, and future challenges. In: Wilson, L. D., Townsend, J. H. &amp; Johnson, J. D., (Eds.), Conservation of Mesoamerican Amphibians and Reptiles. Eagle Mountain Press, Eagle Mountain, Utah, pp. 761 - 812." type="book chapter" year="2010">Wilson &amp; Townsend 2010</bibRefCitation>
,
<bibRefCitation id="66104B7FFF94E76D7CF9FD88FCBAC4A5" author="Wilson, L. D. &amp; Johnson, J. D. &amp; Mata-Silva, V." box="[685,873,638,660]" pageId="84" pageNumber="85" pagination="1 - 47" refId="ref98259" refString="Wilson, L. D. &amp; Johnson, J. D. &amp; Mata-Silva, V. (2013) A conservation reassessment of the reptiles of Mexico based on the EVS measure. Amphibian &amp; Reptile Conservation, 7 (1), 1 - 47." type="journal article" year="2013">
Wilson
<emphasis id="30F5EA9CFF94E76D7CA8FD89FCFCC4A5" box="[764,815,638,660]" italics="true" pageId="84" pageNumber="85">et al.</emphasis>
2013
</bibRefCitation>
,
<bibRefCitation id="66104B7FFF94E76D7D20FD88FBFDC4A5" author="Kohler, G. &amp; Gomez-Trejo-Perez, R. &amp; Petersen, C. B. P. &amp; Mendez de la Cruz, F." box="[884,1070,638,660]" pageId="84" pageNumber="85" pagination="453 - 468" refId="ref92525" refString="Kohler, G., Gomez-Trejo-Perez, R., Petersen, C. B. P. &amp; Mendez de la Cruz, F. (2014) A new species of pine anole from the Sierra Madre del Sur in Oaxaca, Mexico (Reptilia, Dactyloidae: Anolis). Zootaxa, 3753 (5), 453 - 468." type="journal article" year="2014">
Köhler
<emphasis id="30F5EA9CFF94E76D7D94FD89FC27C4A5" box="[960,1012,638,660]" italics="true" pageId="84" pageNumber="85">et al.</emphasis>
2014
</bibRefCitation>
, Köhler 2014
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</paragraph>
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<paragraph id="023E368EFF94E76D7EC3FD6BFD8EC482" blockId="84.[151,1436,546,722]" box="[151,605,669,691]" pageId="84" pageNumber="85">
<taxonomicName id="C5814D0DFF94E76D7EC3FD6BFD8EC482" authority=": Gehlbach &amp; Colette 1957" authorityName=": Gehlbach &amp; Colette" authorityYear="1957" box="[151,605,669,691]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="nebuloides">
<emphasis id="30F5EA9CFF94E76D7EC3FD6BFE9FC482" box="[151,332,669,691]" italics="true" pageId="84" pageNumber="85">Anolis nebuloides</emphasis>
:
<bibRefCitation id="66104B7FFF94E76D7F0DFD6BFD8EC482" author="Gehlbach, F. &amp; Colette, B. B." box="[345,605,669,691]" pageId="84" pageNumber="85" pagination="227 - 231" refId="ref90960" refString="Gehlbach, F. R &amp; Colette, B. B. (1957) A Contribution to the herpetofauna of the highlands of Oaxaca and Puebla, Mexico. Herpetologica, 13 (3), 227 - 231." type="journal article" year="1957">Gehlbach &amp; Colette 1957</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="023E368EFF94E76D7EC3FD4AFB61C4E3" blockId="84.[151,1436,546,722]" box="[151,1202,700,722]" pageId="84" pageNumber="85">
<taxonomicName id="C5814D0DFF94E76D7EC3FD4AFD94C4E3" authority=": Savage &amp; Guyer 1989" authorityName="Savage &amp; Guyer" authorityYear="1989" box="[151,583,700,722]" family="Polychrotidae" genus="Norops" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" rank="species" species="quercorum">
<emphasis id="30F5EA9CFF94E76D7EC3FD4AFE8DC4E0" box="[151,350,700,721]" italics="true" pageId="84" pageNumber="85">Norops quercorum:</emphasis>
<bibRefCitation id="66104B7FFF94E76D7F30FD4AFD94C4E3" author="Savage, J. M. &amp; Guyer, C." box="[356,583,700,722]" pageId="84" pageNumber="85" pagination="105 - 116" refId="ref95903" refString="Savage, J. M. &amp; Guyer, C. (1989) Infrageneric classification and species composition of the anole genera, Anolis, Ctenonotus, Dactyloa, Norops, and Semiurus (Sauria: Iguanidae). Amphibia-Reptilia, 10 (2), 105 - 116. http: // dx. doi. org / 10.1163 / 156853889 x 00142" type="journal article" year="1989">Savage &amp; Guyer 1989</bibRefCitation>
</taxonomicName>
,
<bibRefCitation id="66104B7FFF94E76D7C06FD4AFD26C4E3" author="Nicholson, K. E." box="[594,757,700,722]" pageId="84" pageNumber="85" pagination="93 - 120" refId="ref94419" refString="Nicholson, K. E. (2002) Phylogenetic analysis and a test of the current infrageneric classification of Norops (beta Anolis). Herpetological Monographs, 16, 93 - 120. http: // dx. doi. org / 10.1655 / 0733 - 1347 (2002) 016 [0093: paaato] 2.0. co; 2" type="journal article" year="2002">Nicholson 2002</bibRefCitation>
, Castiglia
<emphasis id="30F5EA9CFF94E76D7D34FD4AFC40C4E3" box="[864,915,700,722]" italics="true" pageId="84" pageNumber="85">et al.</emphasis>
2010,
<bibRefCitation id="66104B7FFF94E76D7D8CFD4AFB61C4E3" author="Nicholson, K. E. &amp; Crother, B. I. &amp; Guyer, C. &amp; Savage, J. M." box="[984,1202,700,722]" pageId="84" pageNumber="85" pagination="1 - 108" refId="ref94482" refString="Nicholson, K. E., Crother, B. I., Guyer, C. &amp; Savage, J. M. (2012) It is time for a new classification of anoles (Squamata: Dactyloidae). Zootaxa, 3477, 1 - 108." type="journal article" year="2012">
Nicholson
<emphasis id="30F5EA9CFF94E76D7A12FD4AFBA9C4E3" box="[1094,1146,700,722]" italics="true" pageId="84" pageNumber="85">et al.</emphasis>
2012
</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="4A9B6505FF94E76D7EC3FD08FB28C2F1" pageId="84" pageNumber="85" type="diagnosis">
<paragraph id="023E368EFF94E76D7EC3FD08FB28C2F1" blockId="84.[151,1437,766,2029]" pageId="84" pageNumber="85">
<emphasis id="30F5EA9CFF94E76D7EC3FD08FED9C526" bold="true" box="[151,266,766,791]" pageId="84" pageNumber="85">Diagnosis</emphasis>
. A small species (SVL in largest male 46.0 mm, largest female
<quantity id="C5799B6BFF94E76D7D87FD08FBEBC526" box="[979,1080,766,791]" metricMagnitude="-2" metricUnit="m" metricValue="4.35" pageId="84" pageNumber="85" unit="mm" value="43.5">43.5 mm</quantity>
) of the genus
<taxonomicName id="C5814D0DFF94E76D7A81FD08FF0BC50F" authority="(sensu Poe 2004)" baseAuthorityName="sensu Poe" baseAuthorityYear="2004" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="genus">
<emphasis id="30F5EA9CFF94E76D7A81FD08FACEC527" box="[1237,1309,766,790]" italics="true" pageId="84" pageNumber="85">Anolis</emphasis>
(sensu
<bibRefCitation id="66104B7FFF94E76D7B26FD09FF02C50F" author="Poe, S." pageId="84" pageNumber="85" pagination="37 - 89" refId="ref95161" refString="Poe, S. (2004) Phylogeny of anoles. Herpetological Monographs, 18, 37 - 89. http: // dx. doi. org / 10.1655 / 0733 - 1347 (2004) 018 [0037: poa] 2.0. co; 2" type="journal article" year="2004">Poe 2004</bibRefCitation>
)
</taxonomicName>
that differs from all Mexican and Central American anoles except
<taxonomicName id="C5814D0DFF94E76D7DBCFCD0FBBAC50C" box="[1000,1129,805,829]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="carlliebi">
<emphasis id="30F5EA9CFF94E76D7DBCFCD0FBBAC50C" box="[1000,1129,805,829]" italics="true" pageId="84" pageNumber="85">A. carlliebi</emphasis>
</taxonomicName>
and
<taxonomicName id="C5814D0DFF94E76D7AF1FCD0FAA5C50C" authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[1189,1398,806,829]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="sacamecatensis">
<emphasis id="30F5EA9CFF94E76D7AF1FCD0FAA5C50C" box="[1189,1398,806,829]" italics="true" pageId="84" pageNumber="85">A. sacamecatensis</emphasis>
</taxonomicName>
by having a combination of (1) strongly keeled ventral scales; (2) usually a patch of three greatly enlarged supraocular scales; (3) 12–20 rows of slightly to moderately enlarged dorsal scales that are smaller than ventral scales; (4) short hind legs, longest toe of adpressed hind leg usually reaching to ear opening or to a point between ear opening and eye, ratio shank length/SVL 0.22–0.26; (5) circumnasal usually in contact with first supralabial; and (6) a large pink dewlap in males and a very small pinkish orange-red dewlap in females (
<figureCitation id="9ABA2A0BFF94E76D7A42FC10FBB8C5CE" box="[1046,1131,998,1023]" captionStart="FIGURE 65" captionStartId="85.[151,250,1966,1988]" captionTargetBox="[151,1436,1085,1939]" captionTargetId="figure-444@85.[151,1436,1085,1939]" captionTargetPageId="85" captionText="FIGURE 65. Dewlaps of Anolis quercorum in life. (a) SMF 96451; (b) IBH 26974; (c) IBH 26977; (d) SMF 96458; (e) IBH 26983; (f) IBH 26975. (a–d) are males, (e–f) are females." figureDoi="http://doi.org/10.5281/zenodo.4931093" httpUri="https://zenodo.org/record/4931093/files/figure.png" pageId="84" pageNumber="85">Fig. 65</figureCitation>
).
<taxonomicName id="C5814D0DFF94E76D7AD5FC10FA98C5CF" authorityName="Fitch" authorityYear="1978" box="[1153,1355,998,1022]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="quercorum">
<emphasis id="30F5EA9CFF94E76D7AD5FC10FA98C5CF" box="[1153,1355,998,1022]" italics="true" pageId="84" pageNumber="85">Anolis quercorum</emphasis>
</taxonomicName>
differs from
<taxonomicName id="C5814D0DFF94E76D7E81FBF8FE87C214" box="[213,340,1037,1061]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="carlliebi">
<emphasis id="30F5EA9CFF94E76D7E81FBF8FE87C214" box="[213,340,1037,1061]" italics="true" pageId="84" pageNumber="85">A. carlliebi</emphasis>
</taxonomicName>
and
<taxonomicName id="C5814D0DFF94E76D7FD9FBF8FD88C214" authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[397,603,1038,1061]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="sacamecatensis">
<emphasis id="30F5EA9CFF94E76D7FD9FBF8FD88C214" box="[397,603,1038,1061]" italics="true" pageId="84" pageNumber="85">A. sacamecatensis</emphasis>
</taxonomicName>
in the ranges and average values of several morphometric and pholidotic characters (see
<tableCitation id="4F030335FF94E76D7F17FBC2FE4BC27C" box="[323,408,1076,1101]" captionStart="TABLE 9" captionStartId="68.[151,241,152,175]" captionTargetPageId="68" captionText="TABLE 9. Selected measurements, proportions and scale characters of the species related to Anolis quercorum. Range is followed by mean value and standard deviation in parentheses. For abbreviations see text." httpUri="http://table.plazi.org/id/56FE6606FF84E77D7EC3FF6EFB86C6E3" pageId="84" pageNumber="85" tableUuid="56FE6606FF84E77D7EC3FF6EFB86C6E3">Table 9</tableCitation>
), most obvious in (1) number of middorsal scales in one head length; (2) ratio shank length/ HL; and (3) subdigital lamellae on Phalanges II–IV of Toe IV. Also,
<taxonomicName id="C5814D0DFF94E76D7DC0FBADFB8FC243" authorityName="Fitch" authorityYear="1978" box="[916,1116,1115,1139]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="quercorum">
<emphasis id="30F5EA9CFF94E76D7DC0FBADFB8FC243" box="[916,1116,1115,1139]" italics="true" pageId="84" pageNumber="85">Anolis quercorum</emphasis>
</taxonomicName>
differs from
<taxonomicName id="C5814D0DFF94E76D7ABBFBADFABEC242" box="[1263,1389,1115,1139]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="carlliebi">
<emphasis id="30F5EA9CFF94E76D7ABBFBADFABEC242" box="[1263,1389,1115,1139]" italics="true" pageId="84" pageNumber="85">A. carlliebi</emphasis>
</taxonomicName>
and
<taxonomicName id="C5814D0DFF94E76D7EC3FB74FEBBC2A8" authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[151,360,1154,1177]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="sacamecatensis">
<emphasis id="30F5EA9CFF94E76D7EC3FB74FEBBC2A8" box="[151,360,1154,1177]" italics="true" pageId="84" pageNumber="85">A. sacamecatensis</emphasis>
</taxonomicName>
in hemipenis morphology (hemipenis distinctly bilobed in
<taxonomicName id="C5814D0DFF94E76D7A75FB74FB6CC2A8" authorityName="Fitch" authorityYear="1978" box="[1057,1215,1154,1177]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="quercorum">
<emphasis id="30F5EA9CFF94E76D7A75FB74FB6CC2A8" box="[1057,1215,1154,1177]" italics="true" pageId="84" pageNumber="85">A. quercorum</emphasis>
</taxonomicName>
vs. unilobed in
<taxonomicName id="C5814D0DFF94E76D7BD0FB74FF25C2F1" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="carlliebi">
<emphasis id="30F5EA9CFF94E76D7BD0FB74FF25C2F1" italics="true" pageId="84" pageNumber="85">A. carlliebi</emphasis>
</taxonomicName>
and slightly bilobed in
<taxonomicName id="C5814D0DFF94E76D7FAAFB5EFD1FC28E" authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[510,716,1192,1215]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="sacamecatensis">
<emphasis id="30F5EA9CFF94E76D7FAAFB5EFD1FC28E" box="[510,716,1192,1215]" italics="true" pageId="84" pageNumber="85">A. sacamecatensis</emphasis>
</taxonomicName>
) and by substantial genetic distances (see above).
</paragraph>
</subSubSection>
<subSubSection id="4A9B6505FF94E76E7E93FB38FE9CC766" lastPageId="87" lastPageNumber="88" pageId="84" pageNumber="85" type="description">
<paragraph id="023E368EFF94E76C7E93FB38FF23C21B" blockId="84.[151,1437,766,2029]" lastBlockId="85.[151,1437,151,1066]" lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85">
<emphasis id="30F5EA9CFF94E76D7E93FB38FE8AC2D6" bold="true" box="[199,345,1230,1255]" pageId="84" pageNumber="85">Description.</emphasis>
<taxonomicName id="C5814D0DFF94E76D7F37FB38FDFCC2D7" authorityName="Fitch" authorityYear="1978" box="[355,559,1230,1254]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="84" pageNumber="85" phylum="Chordata" rank="species" species="quercorum">
<emphasis id="30F5EA9CFF94E76D7F37FB38FDFCC2D7" box="[355,559,1230,1254]" italics="true" pageId="84" pageNumber="85">Anolis quercorum</emphasis>
</taxonomicName>
is a small anole (maximum recorded SVL 46.0 mm in males,
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in females); dorsal head scales in internasal region smooth to weakly keeled, occasionally strongly keeled, unicarinate, other dorsal head scales smooth to weakly keeled in prefrontal region, in parietal region smooth to rugose; shallow to deep prefrontal depression present, shallow parietal depression; 4–7 postrostrals; anterior nasal usually single, exceptionally divided, the lower scale in contact with rostral and first supralabial; 4–8 internasals; canthal ridge sharply defined; scales comprising supraorbital semicircles well defined, smooth to strongly keeled, largest scale in semicircles usually subequal or larger than largest supraocular scale; supraorbital semicircles usually broadly in contact, occasionally separated by a complete row of scales; 1–4 scales separating supraorbital semicircles and interparietal at narrowest point; interparietal well defined, greatly enlarged relative to adjacent scales, surrounded by scales of moderate size, longer than wide, usually subequal to ear opening; enlarged supraoculars usually a patch of 3 greatly enlarged scales in a single row (occasionally a patch of 4–7 moderately to greatly enlarged, somewhat irregular arranged scales in two rows) rugose to weakly keeled, separated from supraorbital semicircles by a complete row of small scales, or these scales narrowly in contact (broadly in contact in SMF 96456); 2–3 scales between enlarged supraoculars and superciliaries; usually 2 occasionally 3 elongate superciliaries, anterior one longest, followed posteriorly by a series of 3–4 rounded or squarish keeled scales of moderate size; usually 3 enlarged canthals, the second canthal largest; 4–9 scales between second canthals; 6–10 scales between posterior canthals; loreal region slightly concave, 11–29 mostly keeled (some smooth to rugose) loreal scales in a maximum of 3–5 horizontal rows; 5–7 supralabials to level below center of eye; suboculars keeled, broadly in contact with supralabials (2–3 suboculars in contact with 2–4 supralabials); ear opening vertically oval; scales anterior to ear opening keeled, juxtaposed, about four times larger than granulars posterior to ear opening; 5–8 infralabials to level below center of eye; 3–5 postmentals (usually 4), outer pair slightly to distinctly larger than adjacent median postmental scales; 0–3(commonly 0 or 1) enlarged sublabials in contact with infralabials on each side; keeled granular scales present on chin and throat; male dewlap of large size extending onto chest; 7–8 horizontal gorgetal-sternal rows with 14–18 scales per row; modal number of marginal pairs 2–3;female dewlap very small or small; a nuchal crest and a dorsal ridge present in males; scales on middorsum weakly to strongly keeled, subimbricate with rounded posterior margins; 12–20 middorsal scale rows slightly to moderately enlarged, with a gradual transition to lateral scales; lateral scales keeled, more or less homogeneous; 48–67 dorsal scales along vertebral midline between levels of axilla and groin in males,
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females; 22–36 dorsal scales along vertebral midline contained in one head length in males,
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females; ventral scales on midsection larger than largest dorsal scales; scales on midventer usually strongly keeled, imbricate, usually mucronate, occasionally weakly keeled, subimbricate, non-mucronate; 33–52 ventral scales along midventral line between levels of axilla and groin in males,
<quantity id="C5799B6BFF95E76C7CD0FDECFD35C402" box="[644,742,538,563]" metricMagnitude="0" metricUnit="m" metricValue="1.0413999999999999" metricValueMax="1.2446000000000002" metricValueMin="0.8382000000000002" pageId="85" pageNumber="86" unit="in" value="41.0" valueMax="49.0" valueMin="33.0">33–49 in</quantity>
females; 22–40 ventral scales contained in one head length in males,
<quantity id="C5799B6BFF95E76C7EB3FDB7FE98C46B" box="[231,331,577,602]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" metricValueMax="7.112" metricValueMin="4.572" pageId="85" pageNumber="86" unit="in" value="23.0" valueMax="28.0" valueMin="18.0">18–28 in</quantity>
females; 106–140 scales around midbody in males,
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females; tube-like axillary pocket absent; precloacal scales smooth to weakly keeled; males with a pair of greatly enlarged postcloacal scales; tail moderately compressed in cross section, tail height/tail width
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males,
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females; all caudal scales strongly keeled, homogeneous, although an indistinct division in segments is discernible; dorsal medial caudal scale row hardly enlarged, strongly keeled, not forming a crest; scales on anterodorsal surface of brachium subimbricate to imbricate, strongly keeled, unicarinate, non-mucronate; scales on dorsal surface of antebrachium strongly keeled, unicarinate, subimbricate to imbricate, some slightly mucronate; 19–26 subdigital lamellae on Phalanges II–IV of Toe IV of hind limbs; 5–7 subdigital lamellae on distal phalanx of Toe IV of hind limbs; digital pads dilated, slightly more than twice the size of distal phalanx. In all specimens examined, the longest toe of the adpressed hind leg reaches to level of tympanum or to a point between shoulder and tympanum. Of 14 individuals, the longest toe of adpressed hind leg reaching almost to ear opening in 3.3%, to ear opening in 60.0%, and to a point slightly beyond ear opening in 10.0%. For variation in selected scalation and morphometric characters see
<tableCitation id="4F030335FF95E76C7EC3FBE7FF3EC21B" box="[151,237,1041,1066]" captionStart="TABLE 9" captionStartId="68.[151,241,152,175]" captionTargetPageId="68" captionText="TABLE 9. Selected measurements, proportions and scale characters of the species related to Anolis quercorum. Range is followed by mean value and standard deviation in parentheses. For abbreviations see text." httpUri="http://table.plazi.org/id/56FE6606FF84E77D7EC3FF6EFB86C6E3" pageId="85" pageNumber="86" tableUuid="56FE6606FF84E77D7EC3FF6EFB86C6E3">Table 9</tableCitation>
.
</paragraph>
<caption id="56FE6606FF95E76C7EC3F858FD0FC1D2" ID-DOI="http://doi.org/10.5281/zenodo.4931093" ID-Zenodo-Dep="4931093" httpUri="https://zenodo.org/record/4931093/files/figure.png" pageId="85" pageNumber="86" startId="85.[151,250,1966,1988]" targetBox="[151,1436,1085,1939]" targetPageId="85">
<paragraph id="023E368EFF95E76C7EC3F858FD0FC1D2" blockId="85.[151,1436,1966,2019]" pageId="85" pageNumber="86">
<emphasis id="30F5EA9CFF95E76C7EC3F858FEF3C1F2" bold="true" box="[151,288,1966,1988]" pageId="85" pageNumber="86">FIGURE 65.</emphasis>
Dewlaps of
<taxonomicName id="C5814D0DFF95E76C7FF0F858FD89C1F5" authorityName="Fitch" authorityYear="1978" box="[420,602,1966,1988]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="85" pageNumber="86" phylum="Chordata" rank="species" species="quercorum">
<emphasis id="30F5EA9CFF95E76C7FF0F858FD89C1F5" box="[420,602,1966,1988]" italics="true" pageId="85" pageNumber="86">Anolis quercorum</emphasis>
</taxonomicName>
in life. (a) SMF 96451; (b) IBH 26974; (c) IBH 26977; (d) SMF 96458; (e) IBH 26983; (f) IBH 26975. (a–d) are males, (e–f) are females.
</paragraph>
</caption>
<caption id="56FE6606FF96E76F7EC3FA07FDA4C017" ID-DOI="http://doi.org/10.5281/zenodo.4931095" ID-Zenodo-Dep="4931095" httpUri="https://zenodo.org/record/4931095/files/figure.png" pageId="86" pageNumber="87" startId="86.[151,250,1521,1543]" targetBox="[155,1436,187,1493]" targetPageId="86">
<paragraph id="023E368EFF96E76F7EC3FA07FDA4C017" blockId="86.[151,1435,1521,1574]" pageId="86" pageNumber="87">
<emphasis id="30F5EA9CFF96E76F7EC3FA07FEF2C037" bold="true" box="[151,289,1521,1543]" pageId="86" pageNumber="87">FIGURE 66.</emphasis>
<taxonomicName id="C5814D0DFF96E76F7F7CFA07FE0DC036" authorityName="Fitch" authorityYear="1978" box="[296,478,1521,1543]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="86" pageNumber="87" phylum="Chordata" rank="species" species="quercorum">
<emphasis id="30F5EA9CFF96E76F7F7CFA07FE0DC036" box="[296,478,1521,1543]" italics="true" pageId="86" pageNumber="87">Anolis quercorum</emphasis>
</taxonomicName>
in life. (a) SMF 96448; (b) SMF 96438; (c) SMF 96447; (d) IBH 26983; (e) IBH 26976; (f) SMF 96456. (a–c) are males, (d–f) are females.
</paragraph>
</caption>
<paragraph id="023E368EFF96E76F7E93F99AFD74C0DE" blockId="86.[151,1437,1644,2011]" pageId="86" pageNumber="87">
The coloration in life of an adult male from
<quantity id="C5799B6BFF96E76F7C92F99AFCD2C0B4" box="[710,769,1644,1669]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="86" pageNumber="87" unit="km" value="2.0">2 km</quantity>
SE Cuesta Blanca (SMF 96457) was recorded as follows: Dorsal ground color of body, head, limbs and tail Drab (19); limbs with Tawny Olive (17) bands; tail with Olive Brown (278) bands; ventral surfaces of body, limbs and tail Light Buff (2); dewlap Pale Purple (223) with Pale Pinkish Buff (3) gorgetals; iris Dark Drab (45).
</paragraph>
<paragraph id="023E368EFF96E76E7E93F90CFB4AC6E3" blockId="86.[151,1437,1644,2011]" lastBlockId="87.[151,1437,151,343]" lastPageId="87" lastPageNumber="88" pageId="86" pageNumber="87">
The coloration in life of an adult female from
<quantity id="C5799B6BFF96E76F7C81F90CFCDCC122" box="[725,783,1786,1811]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="86" pageNumber="87" unit="km" value="2.0">2 km</quantity>
SE Cuesta Blanca (IBH 26982) was recorded as follows: Dorsal ground color Mikado Brown (42) with a suffusion of Cinnamon Brown (43) on head; vertebral line Dark Salmon Color (59) grading into Orange-Rufous (56) on tail and edged by Cinnamon Brown (43) on dorsum; ventral surface of head Light Buff (2) ventral surface of limbs Drab (19); ventral surface of tail Salmon Color (58); dewlap Spinel Pink (235) with Pale Pinkish Buff (3) gorgetals; iris Sprout´s Brown (47). The coloration in life of another adult female from
<quantity id="C5799B6BFF96E76F7F90F856FE2EC188" box="[452,509,1952,1977]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="86" pageNumber="87" unit="km" value="2.0">2 km</quantity>
SE Cuesta Blanca (IBH 26983;
<figureCitation id="9ABA2A0BFF96E76F7D39F856FC1FC188" box="[877,972,1952,1977]" captionStart="FIGURE 66" captionStartId="86.[151,250,1521,1543]" captionTargetBox="[155,1436,187,1493]" captionTargetId="figure-253@86.[155,1440,187,1493]" captionTargetPageId="86" captionText="FIGURE 66. Anolis quercorum in life. (a) SMF 96448; (b) SMF 96438; (c) SMF 96447; (d) IBH 26983; (e) IBH 26976; (f) SMF 96456. (a–c) are males, (d–f) are females." figureDoi="http://doi.org/10.5281/zenodo.4931095" httpUri="https://zenodo.org/record/4931095/files/figure.png" pageId="86" pageNumber="87">Fig. 66d</figureCitation>
) was recorded as follows: Dorsal ground color Drab (19) with a weakly defined Medium Fawn Color (257) zigzag band; dorsal surface of limbs True Cinnamon (260) with Cinnamon Drab (259) bands; ventral surface of head and body Pale Buff (1); ventral surface of limbs Pale Pinkish Buff (3); ventral surface of tail Beige (254); iris Sprout´s Brown (47).
</paragraph>
<paragraph id="023E368EFF97E76E7E93FF2CFE9CC766" blockId="87.[151,1437,151,343]" pageId="87" pageNumber="88">
The almost completely everted hemipenis of SMF 96439 (
<figureCitation id="9ABA2A0BFF97E76E7D26FF2CFC1BC6C2" box="[882,968,218,243]" captionStart="FIGURE 70" captionStartId="90.[151,250,954,976]" captionTargetBox="[151,1436,483,928]" captionTargetId="figure-220@90.[151,1436,483,929]" captionTargetPageId="90" captionText="FIGURE 70. Hemipenis of Anolis quercorum (SMF 96439): (left) sulcate view; (center) asulcate view; (right) lateral view. Scale bar equals 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4931103" httpUri="https://zenodo.org/record/4931103/files/figure.png" pageId="87" pageNumber="88">Fig. 70</figureCitation>
) is a bilobate organ; sulcus spermaticus bordered by well developed sulcal lips and opening into two apical fields void of ornamentation, one on each lobe; a large finger-like asulcate processus and a weak asulcate ridge present; apex strongly calyculate, truncus with transverse folds.
</paragraph>
</subSubSection>
<caption id="56FE6606FF97E76E7EC3F93FFAFFC0EE" ID-DOI="http://doi.org/10.5281/zenodo.4931097" ID-Zenodo-Dep="4931097" box="[151,1324,1737,1759]" httpUri="https://zenodo.org/record/4931097/files/figure.png" pageId="87" pageNumber="88" startId="87.[151,250,1737,1759]" targetBox="[155,1436,403,1709]" targetPageId="87">
<paragraph id="023E368EFF97E76E7EC3F93FFAFFC0EE" blockId="87.[151,1324,1737,1759]" box="[151,1324,1737,1759]" pageId="87" pageNumber="88">
<emphasis id="30F5EA9CFF97E76E7EC3F93FFEF3C0EF" bold="true" box="[151,288,1737,1759]" pageId="87" pageNumber="88">FIGURE 67.</emphasis>
<taxonomicName id="C5814D0DFF97E76E7F71F93FFE08C0EE" authorityName="Fitch" authorityYear="1978" box="[293,475,1737,1759]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="87" pageNumber="88" phylum="Chordata" rank="species" species="quercorum">
<emphasis id="30F5EA9CFF97E76E7F71F93FFE08C0EE" box="[293,475,1737,1759]" italics="true" pageId="87" pageNumber="88">Anolis quercorum</emphasis>
</taxonomicName>
: head scalation in (a,c,e) KU 176051; (b,d,f) SMF 96444. Scale bars equal 1.0 mm.
</paragraph>
</caption>
<caption id="56FE6606FF98E7617EC3F869FF16C1C2" ID-DOI="http://doi.org/10.5281/zenodo.4931099" ID-Zenodo-Dep="4931099" httpUri="https://zenodo.org/record/4931099/files/figure.png" pageId="88" pageNumber="89" startId="88.[151,250,1951,1973]" targetBox="[155,1436,181,1930]" targetPageId="88">
<paragraph id="023E368EFF98E7617EC3F869FF16C1C2" blockId="88.[151,1436,1951,2035]" pageId="88" pageNumber="89">
<emphasis id="30F5EA9CFF98E7617EC3F869FECCC185" bold="true" box="[151,287,1951,1973]" pageId="88" pageNumber="89">FIGURE 68.</emphasis>
<taxonomicName id="C5814D0DFF98E7617F71F869FE09C184" authorityName="Fitch" authorityYear="1978" box="[293,474,1951,1973]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="88" pageNumber="89" phylum="Chordata" rank="species" species="quercorum">
<emphasis id="30F5EA9CFF98E7617F71F869FE09C184" box="[293,474,1951,1973]" italics="true" pageId="88" pageNumber="89">Anolis quercorum</emphasis>
</taxonomicName>
: nasal region in (a) SMF 96441; (b) KU 176051; superciliary region in (c) SMF 96444; (d) KU 176051; chin region in (e) SMF 96444; (f) SMF 96441; cloacal region in (g) IBH 26973; (h) KU 176051. Scale bars equal 1.0 mm.
</paragraph>
</caption>
<caption id="56FE6606FF99E7607EC3FA04FC04C017" ID-DOI="http://doi.org/10.5281/zenodo.4931101" ID-Zenodo-Dep="4931101" httpUri="https://zenodo.org/record/4931101/files/figure.png" pageId="89" pageNumber="90" startId="89.[151,250,1522,1544]" targetBox="[151,1436,192,1498]" targetPageId="89">
<paragraph id="023E368EFF99E7607EC3FA04FC04C017" blockId="89.[151,1436,1522,1575]" pageId="89" pageNumber="90">
<emphasis id="30F5EA9CFF99E7607EC3FA04FEF1C036" bold="true" box="[151,290,1522,1544]" pageId="89" pageNumber="90">FIGURE 69.</emphasis>
<taxonomicName id="C5814D0DFF99E7607F7EFA04FE31C039" authorityName="Fitch" authorityYear="1978" box="[298,482,1522,1544]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="89" pageNumber="90" phylum="Chordata" rank="species" species="quercorum">
<emphasis id="30F5EA9CFF99E7607F7EFA04FE31C039" box="[298,482,1522,1544]" italics="true" pageId="89" pageNumber="90">Anolis quercorum</emphasis>
</taxonomicName>
: dorsal scales in (a) KU 176051; (b) SMF 96444; flank scales in (c) KU 176051; (d) SMF 96444; ventral scales in (e) KU 176051; (f) SMF 96444. Scale bars equal 1.0 mm.
</paragraph>
</caption>
<subSubSection id="4A9B6505FF99E7607E93F9A5FBBAC0CB" pageId="89" pageNumber="90" type="biology_ecology">
<paragraph id="023E368EFF99E7607E93F9A5FBBAC0CB" blockId="89.[151,1436,1619,2036]" pageId="89" pageNumber="90">
<emphasis id="30F5EA9CFF99E7607E93F9A5FE08C05D" bold="true" box="[199,475,1619,1644]" pageId="89" pageNumber="90">Natural History Notes:</emphasis>
The habitat of
<taxonomicName id="C5814D0DFF99E7607CD8F9A2FC85C05A" authorityName="Fitch" authorityYear="1978" box="[652,854,1620,1644]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="89" pageNumber="90" phylum="Chordata" rank="species" species="quercorum">
<emphasis id="30F5EA9CFF99E7607CD8F9A2FC85C05A" box="[652,854,1620,1644]" italics="true" pageId="89" pageNumber="90">Anolis quercorum</emphasis>
</taxonomicName>
is leaf litter in oak forest. Here this typical ground anole can be found in amazing population densities. In the vicinity of the
<typeStatus id="DD3A882CFF99E7607DBFF98EFBC8C0A1" box="[1003,1051,1656,1680]" pageId="89" pageNumber="90">type</typeStatus>
locality of this species (
<quantity id="C5799B6BFF99E7607B63F981FAA0C0A1" box="[1335,1395,1655,1680]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="89" pageNumber="90" unit="km" value="2.0">2 km</quantity>
SE Cuesta Blanca,
<geoCoordinate id="67B55049FF99E7607F00F96CFE0CC082" box="[340,479,1690,1715]" degrees="17.33065" direction="north" orientation="latitude" pageId="89" pageNumber="90" precision="1" value="17.33065">17.33065°N</geoCoordinate>
,
<geoCoordinate id="67B55049FF99E7607FA5F96CFD5DC082" box="[497,654,1690,1715]" degrees="97.158420" direction="west" orientation="longitude" pageId="89" pageNumber="90" precision="1" value="-97.15842">97.158420°W</geoCoordinate>
,
<quantity id="C5799B6BFF99E7607CF4F96CFCCCC082" box="[672,799,1690,1715]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="89" pageNumber="90" unit="m" value="2250.0">2250 masl</quantity>
), we saw at least 50 individuals within one hour of searching in a relatively small area (&lt;1 ha). When disturbed, these lizards darted a meter or so across the leaf litter and then usually froze and were well camouflaged against the vivid background colors.
</paragraph>
</subSubSection>
<subSubSection id="4A9B6505FF99E7607E93F8F3FAB6C1B9" pageId="89" pageNumber="90" type="distribution">
<paragraph id="023E368EFF99E7607E93F8F3FAB6C1B9" blockId="89.[151,1436,1619,2036]" pageId="89" pageNumber="90">
<emphasis id="30F5EA9CFF99E7607E93F8F3FD00C12F" bold="true" box="[199,723,1797,1822]" pageId="89" pageNumber="90">Geographic Distribution and Conservation</emphasis>
. As currently known,
<taxonomicName id="C5814D0DFF99E7607D88F8F3FB7BC12C" authorityName="Fitch" authorityYear="1978" box="[988,1192,1797,1821]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="89" pageNumber="90" phylum="Chordata" rank="species" species="quercorum">
<emphasis id="30F5EA9CFF99E7607D88F8F3FB7BC12C" box="[988,1192,1797,1821]" italics="true" pageId="89" pageNumber="90">Anolis quercorum</emphasis>
</taxonomicName>
is distributed on the Pacific versant of the western central and northwestern portions of the Mexican State of
<collectingRegion id="C045F86CFF99E7607AF8F8DEFAD3C170" box="[1196,1280,1832,1857]" country="Mexico" name="Oaxaca" pageId="89" pageNumber="90">Oaxaca</collectingRegion>
at elevations between 1616 and
<quantity id="C5799B6BFF99E7607F38F8BAFE32C154" box="[364,481,1868,1893]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="89" pageNumber="90" unit="m" value="2250.0">2250 masl</quantity>
(
<figureCitation id="9ABA2A0BFF99E7607FA5F8BAFD96C154" box="[497,581,1868,1893]" captionStart="FIGURE 71" captionStartId="90.[151,250,1975,1997]" captionTargetBox="[286,1307,1067,1956]" captionTargetId="figure-229@90.[275,1317,1056,1964]" captionTargetPageId="90" captionText="FIGURE 71. Map indicating collecting localities of species related to Anolis quercorum. Each symbol can represent one or more adjacent localities. Black circles: Anolis carlliebi; white circles: A. quercorum; white squares: A. sacamecatensis." figureDoi="http://doi.org/10.5281/zenodo.4931105" httpUri="https://zenodo.org/record/4931105/files/figure.png" pageId="89" pageNumber="90">Fig. 71</figureCitation>
). Given its usual abundance wherever this species occurs, it seems justified to classify
<taxonomicName id="C5814D0DFF99E7607F44F886FE7AC1B6" authorityName="Fitch" authorityYear="1978" box="[272,425,1904,1927]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="89" pageNumber="90" phylum="Chordata" rank="species" species="quercorum">
<emphasis id="30F5EA9CFF99E7607F44F886FE7AC1B6" box="[272,425,1904,1927]" italics="true" pageId="89" pageNumber="90">A. quercorum</emphasis>
</taxonomicName>
as Least Concern based on the IUCN Red List Categories and Criteria (
<bibRefCitation id="66104B7FFF99E7607A9BF899FA8AC1B9" author="IUCN" box="[1231,1369,1903,1928]" pageId="89" pageNumber="90" refId="ref91603" refString="IUCN (2012) IUCN Red List of Threatened Species. Version 2012.2. Available from: http: // www. iucnredlist. org (accessed 11 January 2013)" type="url" year="2012">IUCN 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4A9B6505FF99E7637E93F865FBBFC79C" lastPageId="90" lastPageNumber="91" pageId="89" pageNumber="90" type="materials_examined">
<paragraph id="023E368EFF99E7637E93F865FBBFC79C" blockId="89.[151,1436,1619,2036]" lastBlockId="90.[151,1437,151,429]" lastPageId="90" lastPageNumber="91" pageId="89" pageNumber="90">
<emphasis id="30F5EA9CFF99E7607E93F865FE31C19D" bold="true" box="[199,482,1939,1964]" pageId="89" pageNumber="90">Specimens examined—</emphasis>
<materialsCitation id="B2E93CD3FF99E7607FB6F865FDB5C1FE" ID-GBIF-Occurrence="4013201514" collectionCode="UTA" country="Mexico" elevation="2250" location="El" pageId="89" pageNumber="90" specimenCode="IBH 26973-76, 26982-83, SMF 96441-55, 96457-58, 96747-51, 98021, 98735" specimenCount="1" stateProvince="Oaxaca">
<emphasis id="30F5EA9CFF99E7607FB6F865FDECC19D" bold="true" box="[482,575,1939,1964]" pageId="89" pageNumber="90">
<collectingCountry id="7A96761EFF99E7607FB6F865FDE8C19D" box="[482,571,1939,1964]" name="Mexico" pageId="89" pageNumber="90">Mexico</collectingCountry>
:
</emphasis>
<collectingRegion id="C045F86CFF99E7607C1EF865FD71C19D" box="[586,674,1939,1964]" country="Mexico" name="Oaxaca" pageId="89" pageNumber="90">Oaxaca</collectingRegion>
: 0.9 mi N (by road)
<collectorName id="AF745358FF99E7607DC9F865FBCAC19D" box="[925,1049,1939,1964]" pageId="89" pageNumber="90">
<location id="075E6055FF99E7607DC9F865FC65C19D" LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF99E7607DC9F865FC65C19D" box="[925,950,1939,1964]" country="Mexico" name="El" pageId="89" pageNumber="90" stateProvince="Oaxaca">El</location>
Cerezal
</collectorName>
,
<collectorName id="AF745358FF99E7607A7CF865FB29C19D" box="[1064,1274,1939,1964]" pageId="89" pageNumber="90">Sierra Aloapaneca</collectorName>
(N of city of Oaxaca)
<collectorName id="AF745358FF99E7607F56F841FEAAC1FE" box="[258,377,1975,1999]" pageId="89" pageNumber="90">Mex Hwy</collectorName>
175:
<collectionCode id="6490AE4BFF99E7607F92F840FE2EC1FE" box="[454,509,1974,1999]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34873" lsid="urn:lsid:biocol.org:col:34873" name="University of Texas at Arlington" pageId="89" pageNumber="90">UTA</collectionCode>
R-8421;
</materialsCitation>
<materialsCitation id="B2E93CD3FF99E7607C21F840FD61C1C5" ID-GBIF-Occurrence="4013201421" collectionCode="UTA" country="Mexico" elevation="2250" location="El" pageId="89" pageNumber="90" specimenCode="IBH 26973-76, 26982-83, SMF 96441-55, 96457-58, 96747-51, 98021, 98735" specimenCount="1" stateProvince="Oaxaca">
<locationDeviation id="138C2FACFF99E7607C21F840FC48C1FE" box="[629,923,1974,1999]" location="Cuesta Blanca" pageId="89" pageNumber="90">2 km SE Cuesta Blanca</locationDeviation>
, 2250 m:
<specimenCode id="52279EF5FF99E7607A49F841FB1EC1FE" box="[1053,1229,1974,1999]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="89" pageNumber="90">IBH 26973–76</specimenCode>
,
<specimenCode id="52279EF5FF99E7607AB4F840FA81C1FE" box="[1248,1362,1974,1999]" collectionCode="IBH" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="89" pageNumber="90">26982–83</specimenCode>
,
<specimenCode id="52279EF5FF99E7607B30F840FED9C1C5" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="89" pageNumber="90">SMF 96441–55</specimenCode>
,
<specimenCode id="52279EF5FF99E7607F41F82DFE54C1C5" box="[277,391,2011,2036]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="89" pageNumber="90">96457–58</specimenCode>
,
<specimenCode id="52279EF5FF99E7607FC7F82DFDD6C1C5" box="[403,517,2011,2036]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="89" pageNumber="90">96747–51</specimenCode>
,
<specimenCode id="52279EF5FF99E7607C44F82DFD8AC1C5" box="[528,601,2011,2036]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="89" pageNumber="90">98021</specimenCode>
,
<specimenCode id="52279EF5FF99E7607C31F82DFD7CC1C5" box="[613,687,2011,2036]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="89" pageNumber="90">98735</specimenCode>
;
</materialsCitation>
<materialsCitation id="B2E93CD3FF99E7607CEDF82DFB4CC1C5" ID-GBIF-Occurrence="4013201332" box="[697,1183,2011,2036]" collectionCode="UTA" country="Mexico" elevation="2250" location="El" pageId="89" pageNumber="90" specimenCode="IBH 26973-76, 26982-83, SMF 96441-55, 96457-58, 96747-51, 98021, 98735" specimenCount="1" stateProvince="Oaxaca">
1.7 mi SE
<location id="075E6055FF99E7607D7BF82DFC63C1C5" LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF99E7607D7BF82DFC63C1C5" box="[815,944,2011,2036]" country="Mexico" name="Nochixtlan" pageId="89" pageNumber="90">Nochixtlán</location>
:
<collectionCode id="6490AE4BFF99E7607DEEF82DFC25C1C5" box="[954,1014,2011,2036]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="89" pageNumber="90" type="Museum">MCZ</collectionCode>
R-133097–98;
</materialsCitation>
<materialsCitation id="B2E93CD3FF99E7637AF3F82DFD41C681" ID-GBIF-Occurrence="4013201510" accessionNumber="KU 176051" collectionCode="UTA" country="Mexico" elevation="2250" lastPageId="90" lastPageNumber="91" location="El" pageId="89" pageNumber="90" specimenCode="IBH 26973-76, 26982-83, SMF 96441-55, 96457-58, 96747-51, 98021, 98735" specimenCount="1" stateProvince="Oaxaca">
26 km SE
<location id="075E6055FF99E7607B48F82DFA4BC1C5" LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF99E7607B48F82DFA4BC1C5" box="[1308,1432,2011,2036]" country="Mexico" name="Nochistlan" pageId="89" pageNumber="90">Nochistlán</location>
, 2.5 km NW
<location id="075E6055FF9AE7637F73FF61FE1CC681" LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF9AE7637F73FF61FE1CC681" box="[295,463,151,176]" name="Cuesta Blanca" pageId="90" pageNumber="91">Cuesta Blanca</location>
:
<specimenCode id="52279EF5FF9AE7637F8FFF6EFD5DC681" box="[475,654,151,176]" collectionCode="KU" country="USA" name="Biodiversity Institute, University of Kansas" pageId="90" pageNumber="91" type="University or college">KU 176051–57</specimenCode>
;
</materialsCitation>
<materialsCitation id="B2E93CD3FF9AE7637CCFFF61FB9EC681" ID-GBIF-Occurrence="4013201405" accessionNumber="KU 176051" box="[667,1101,151,176]" collectionCode="KU" elevation="2164" location="Cuesta Blanca" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
3.5 WSW
<location id="075E6055FF9AE7637D41FF61FCA8C681" LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF9AE7637D41FF61FCA8C681" box="[789,891,151,176]" name="Tlaxiaco" pageId="90" pageNumber="91">Tlaxiaco</location>
:
<collectionCode id="6490AE4BFF9AE7637DD3FF61FC6DC681" box="[903,958,151,176]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34873" lsid="urn:lsid:biocol.org:col:34873" name="University of Texas at Arlington" pageId="90" pageNumber="91">UTA</collectionCode>
R-3568–71;
</materialsCitation>
<materialsCitation id="B2E93CD3FF9AE7637A02FF61FE9CC6E4" ID-GBIF-Occurrence="4013201418" accessionNumber="KU 176051" collectionCode="KU" elevation="2164" location="Cuesta Blanca" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
32 mi NW Oaxaca, 2164 m:
<specimenCode id="52279EF5FF9AE7637EC3FF4AFE9FC6E4" box="[151,332,188,213]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="90" pageNumber="91" type="University or college">UMMZ 115057</specimenCode>
;
</materialsCitation>
<materialsCitation id="B2E93CD3FF9AE7637F02FF4AFBCBC6E4" ID-GBIF-Occurrence="4013201340" accessionNumber="KU 176051" box="[342,1048,188,213]" collectionCode="KU" elevation="2164" location="Cuesta Blanca" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
32.5 mi NW Oaxaca, 2134 m:
<specimenCode id="52279EF5FF9AE7637CFAFF4AFC8CC6E4" box="[686,863,188,213]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="90" pageNumber="91" type="University or college">UMMZ 112636</specimenCode>
(
<specimenCount id="1487FD07FF9AE7637D3BFF4AFBDFC6E4" box="[879,1036,188,213]" pageId="90" pageNumber="91" type="generic">12 specimens</specimenCount>
);
</materialsCitation>
<materialsCitation id="B2E93CD3FF9AE7637A4BFF4AFEDEC6C9" ID-GBIF-Occurrence="4013201386" accessionNumber="KU 176051" collectionCode="KU" country="Mexico" elevation="2164" location="Cuesta Blanca" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
3.1 km NW
<location id="075E6055FF9AE7637AFEFF4AFAEBC6E4" LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF9AE7637AFEFF4AFAEBC6E4" box="[1194,1336,188,213]" name="Llano Verde" pageId="90" pageNumber="91" stateProvince="Oaxaca">Llano Verde</location>
:
<collectionCode id="6490AE4BFF9AE7637B16FF4AFAAAC6E4" box="[1346,1401,188,213]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34873" lsid="urn:lsid:biocol.org:col:34873" name="University of Texas at Arlington" pageId="90" pageNumber="91">UTA</collectionCode>
R-11502–03;
</materialsCitation>
<materialsCitation id="B2E93CD3FF9AE7637F41FF29FCE2C6C9" ID-GBIF-Occurrence="4013201475" accessionNumber="KU 176051" box="[277,817,223,248]" collectionCode="KU" country="Mexico" elevation="2164" location="Cuesta Blanca" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
5.6 km E
<location id="075E6055FF9AE7637FD5FF29FDC3C6C9" LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF9AE7637FD5FF29FDC3C6C9" box="[385,528,223,248]" name="Llano Verde" pageId="90" pageNumber="91" stateProvince="Oaxaca">Llano Verde</location>
:
<collectionCode id="6490AE4BFF9AE7637C4EFF16FD91C6C6" box="[538,578,224,247]" country="USA" name="Biodiversity Institute, University of Kansas" pageId="90" pageNumber="91" type="University or college">KU</collectionCode>
193181,
<specimenCode id="52279EF5FF9AE7637CFFFF29FCFEC6C9" box="[683,813,223,248]" collectionCode="KU" country="USA" name="Biodiversity Institute, University of Kansas" pageId="90" pageNumber="91" type="University or college">193188–89</specimenCode>
;
</materialsCitation>
<materialsCitation id="B2E93CD3FF9AE7637D6CFF29FAEAC6C9" ID-GBIF-Occurrence="4013201499" accessionNumber="KU 176051" box="[824,1337,223,248]" collectionCode="KU" country="Mexico" elevation="2164" location="Cuesta Blanca" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
12.8 km W
<location id="075E6055FF9AE7637DE8FF29FB98C6C9" LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF9AE7637DE8FF29FB98C6C9" box="[956,1099,223,248]" name="Llano Verde" pageId="90" pageNumber="91" stateProvince="Oaxaca">Llano Verde</location>
:
<collectionCode id="6490AE4BFF9AE7637A01FF29FB42C6C9" box="[1109,1169,223,248]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="90" pageNumber="91" type="Museum">MCZ</collectionCode>
R-171142–44;
</materialsCitation>
<materialsCitation id="B2E93CD3FF9AE7637B14FF29FBC8C72C" ID-GBIF-Occurrence="4013201316" accessionNumber="KU 176051" collectionCode="KU" country="Mexico" elevation="2164" location="Cuesta Blanca" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
52 mi N Oaxaca on
<collectingCountry id="7A96761EFF9AE7637F4EFEF2FE8BC72A" box="[282,344,260,283]" name="Mexico" pageId="90" pageNumber="91">MEX</collectingCountry>
190
<locationDeviation id="138C2FACFF9AE7637FC1FEF2FD5BC72C" box="[405,648,260,285]" location="El Cumbre" pageId="90" pageNumber="91">2.4 mi N El Cumbre</locationDeviation>
:
<specimenCode id="52279EF5FF9AE7637CC0FEF2FC9AC72C" box="[660,841,260,285]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="90" pageNumber="91" type="University or college">UMMZ 131485</specimenCode>
(
<specimenCount id="1487FD07FF9AE7637D0EFEF2FBDCC72C" box="[858,1039,260,285]" pageId="90" pageNumber="91" type="generic">nine specimens</specimenCount>
);
</materialsCitation>
<materialsCitation id="B2E93CD3FF9AE7637A70FEF2FF37C771" ID-GBIF-Occurrence="4013201412" accessionNumber="KU 176051" collectionCode="KU" country="Mexico" elevation="2164" location="Cuesta Blanca" municipality="Motor Court" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
<location id="075E6055FF9AE7637A70FEF2FB1AC72C" LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF9AE7637A70FEF2FB1AC72C" box="[1060,1225,260,285]" country="Mexico" name="Hotel Victoria" pageId="90" pageNumber="91" stateProvince="Oaxaca">Hotel Victoria</location>
, 1616 m:
<collectionCode id="6490AE4BFF9AE7637B6BFEF2FAA5C72C" box="[1343,1398,260,285]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34873" lsid="urn:lsid:biocol.org:col:34873" name="University of Texas at Arlington" pageId="90" pageNumber="91">UTA</collectionCode>
R-25789;
</materialsCitation>
<materialsCitation id="B2E93CD3FF9AE7637EBBFED1FDE2C771" ID-GBIF-Occurrence="4013201477" accessionNumber="KU 176051" box="[239,561,295,320]" collectionCode="KU" country="Mexico" elevation="2164" location="Cuesta Blanca" municipality="Motor Court" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
Oaxaca:
<specimenCode id="52279EF5FF9AE7637F01FEDEFDFEC771" box="[341,557,295,320]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="90" pageNumber="91" type="Museum">FMNH 108789-92</specimenCode>
;
</materialsCitation>
<materialsCitation id="B2E93CD3FF9AE7637C68FED1FB64C771" ID-GBIF-Occurrence="4013201464" accessionNumber="KU 176051" box="[572,1207,295,320]" collectionCode="KU" country="Mexico" elevation="2164" location="Cuesta Blanca" municipality="Motor Court" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
Oaxaca
<collectingMunicipality id="E25AACF4FF9AE7637CCEFEDEFCFDC771" box="[666,814,295,320]" pageId="90" pageNumber="91">Motor Court</collectingMunicipality>
: Oaxaca
<location id="075E6055FF9AE7637DCEFED1FC1DC771" LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF9AE7637DCEFED1FC1DC771" box="[922,974,295,320]" country="Mexico" municipality="Motor Court" name="City" pageId="90" pageNumber="91" stateProvince="Oaxaca">City</location>
:
<collectionCode id="6490AE4BFF9AE7637D89FED1FBCAC771" box="[989,1049,295,320]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="90" pageNumber="91" type="Museum">MCZ</collectionCode>
R-86374–75;
</materialsCitation>
<materialsCitation id="B2E93CD3FF9AE7637A96FED1FE29C754" ID-GBIF-Occurrence="4013201357" accessionNumber="KU 176051" collectionCode="KU" country="Mexico" elevation="2164" location="Cuesta Blanca" municipality="Motor Court" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
grounds of Oaxaca
<location id="075E6055FF9AE7637EC3FEBAFEC6C754" LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF9AE7637EC3FEBAFEC6C754" box="[151,277,332,357]" country="Mexico" name="Port Motel" pageId="90" pageNumber="91" stateProvince="Oaxaca">Port Motel</location>
:
<collectionCode id="6490AE4BFF9AE7637F75FEBAFE8EC754" box="[289,349,332,357]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="90" pageNumber="91" type="Museum">MCZ</collectionCode>
R-79562–64;
</materialsCitation>
<materialsCitation id="B2E93CD3FF9AE7637C57FEBAFBB2C754" ID-GBIF-Occurrence="4013201380" accessionNumber="KU 176051" box="[515,1121,332,357]" collectionCode="KU" country="Mexico" elevation="2164" location="Cuesta Blanca" municipality="Motor Court" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
3 mi N
<location id="075E6055FF9AE7637C08FEBAFD3CC754" LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF9AE7637C08FEBAFD3CC754" box="[604,751,332,357]" country="Mexico" name="La Heradura" pageId="90" pageNumber="91" stateProvince="Oaxaca">La Heradura</location>
:
<collectionCode id="6490AE4BFF9AE7637CAEFEBAFCE4C754" box="[762,823,332,357]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="90" pageNumber="91" type="Museum">MCZ</collectionCode>
R-177946–49, 54954–58;
</materialsCitation>
<materialsCitation id="B2E93CD3FF9AE7637A3DFEBAFD41C7B9" ID-GBIF-Occurrence="4013201424" accessionNumber="KU 176051" collectionCode="KU" country="Mexico" elevation="2164" location="Cuesta Blanca" municipality="Motor Court" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
<location id="075E6055FF9AE7637A3DFEBAFA4BC755" LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF9AE7637A3DFEBAFA4BC755" box="[1129,1432,332,357]" country="Mexico" name="Tamazulapan del Progreso" pageId="90" pageNumber="91" stateProvince="Oaxaca">Tamazulapán del Progreso</location>
, 1990 m: IBH 26977, SMF 96438–40,
<specimenCode id="52279EF5FF9AE7637C10FE99FD5DC7B9" box="[580,654,367,392]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="90" pageNumber="91">96456</specimenCode>
;
</materialsCitation>
<materialsCitation id="B2E93CD3FF9AE7637CCDFE99FBEEC7B9" ID-GBIF-Occurrence="4013201360" accessionNumber="KU 176051" box="[665,1085,367,392]" collectionCode="KU" country="Mexico" elevation="2164" location="Cuesta Blanca" municipality="Motor Court" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
<location id="075E6055FF9AE7637CCDFE99FC88C7B9" LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF9AE7637CCDFE99FC88C7B9" box="[665,859,367,392]" country="Mexico" name="Cerro San Felipe" pageId="90" pageNumber="91" stateProvince="Oaxaca">Cerro San Felipe</location>
:
<collectionCode id="6490AE4BFF9AE7637D32FE99FC70C7B9" box="[870,931,367,392]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="90" pageNumber="91" type="Museum">MCZ</collectionCode>
R-93671–72;
</materialsCitation>
<materialsCitation id="B2E93CD3FF9AE7637A11FE99FD9EC79C" ID-GBIF-Occurrence="4013201448" accessionNumber="KU 176051" collectionCode="KU" country="Mexico" elevation="2164" location="Cuesta Blanca" municipality="Motor Court" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
15 km NW (on road)
<collectingMunicipality id="E25AACF4FF9AE7637B6CFE99FEDCC79C" pageId="90" pageNumber="91">San Juan del Estado</collectingMunicipality>
, distrito
<location id="075E6055FF9AE7637F25FE62FE71C79C" LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF9AE7637F25FE62FE71C79C" box="[369,418,404,429]" country="Mexico" municipality="San Juan del Estado" name="Etla" pageId="90" pageNumber="91" stateProvince="Oaxaca">Etla</location>
:
<specimenCode id="52279EF5FF9AE7637FF9FE62FD9AC79C" box="[429,585,404,429]" collectionCode="MVZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34777" lsid="urn:lsid:biocol.org:col:34777" name="Museum of Vertebrate Zoology, University of California Berkeley" pageId="90" pageNumber="91" type="Museum">MVZ 144181</specimenCode>
;
</materialsCitation>
<materialsCitation id="B2E93CD3FF9AE7637C01FE62FBBFC79C" ID-GBIF-Occurrence="4013201435" accessionNumber="KU 176051" box="[597,1132,404,429]" collectionCode="KU" country="Mexico" elevation="2164" location="Cuesta Blanca" municipality="Motor Court" pageId="90" pageNumber="91" specimenCode="KU 176051-57" specimenCount="1" stateProvince="Oaxaca">
<location id="075E6055FF9AE7637C01FE62FDBDC79C" LSID="urn:lsid:plazi:treatment:8A288798FF94E7657EC3FE2AFEBCC156:075E6055FF9AE7637C01FE62FDBDC79C" box="[597,622,404,429]" country="Mexico" name="El" pageId="90" pageNumber="91" stateProvince="Oaxaca">El</location>
Tejocote, 2078 m:
<collectionCode id="6490AE4BFF9AE7637D12FE62FCAEC79C" box="[838,893,404,429]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34873" lsid="urn:lsid:biocol.org:col:34873" name="University of Texas at Arlington" pageId="90" pageNumber="91">UTA</collectionCode>
R-25788, 52818–23.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="56FE6606FF9AE7637EC3FC4CFE45C5DF" ID-DOI="http://doi.org/10.5281/zenodo.4931103" ID-Zenodo-Dep="4931103" httpUri="https://zenodo.org/record/4931103/files/figure.png" pageId="90" pageNumber="91" startId="90.[151,250,954,976]" targetBox="[151,1436,483,928]" targetPageId="90">
<paragraph id="023E368EFF9AE7637EC3FC4CFE45C5DF" blockId="90.[151,1437,954,1006]" pageId="90" pageNumber="91">
<emphasis id="30F5EA9CFF9AE7637EC3FC4CFEF1C5FE" bold="true" box="[151,290,954,976]" pageId="90" pageNumber="91">FIGURE 70.</emphasis>
Hemipenis of
<taxonomicName id="C5814D0DFF9AE7637FEAFC4CFDA6C5E1" authorityName="Fitch" authorityYear="1978" box="[446,629,954,976]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="90" pageNumber="91" phylum="Chordata" rank="species" species="quercorum">
<emphasis id="30F5EA9CFF9AE7637FEAFC4CFDA6C5E1" box="[446,629,954,976]" italics="true" pageId="90" pageNumber="91">Anolis quercorum</emphasis>
</taxonomicName>
(SMF 96439): (left) sulcate view; (center) asulcate view; (right) lateral view. Scale bar equals 1.0 mm.
</paragraph>
</caption>
<caption id="56FE6606FF9AE7637EC3F841FA9FC1DD" ID-DOI="http://doi.org/10.5281/zenodo.4931105" ID-Zenodo-Dep="4931105" httpUri="https://zenodo.org/record/4931105/files/figure.png" pageId="90" pageNumber="91" startId="90.[151,250,1975,1997]" targetBox="[286,1307,1067,1956]" targetPageId="90">
<paragraph id="023E368EFF9AE7637EC3F841FA9FC1DD" blockId="90.[151,1436,1975,2028]" pageId="90" pageNumber="91">
<emphasis id="30F5EA9CFF9AE7637EC3F841FEF2C1FD" bold="true" box="[151,289,1975,1997]" pageId="90" pageNumber="91">FIGURE 71.</emphasis>
Map indicating collecting localities of species related to
<taxonomicName id="C5814D0DFF9AE7637D22F841FBFEC1FC" authorityName="Fitch" authorityYear="1978" box="[886,1069,1975,1997]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="90" pageNumber="91" phylum="Chordata" rank="species" species="quercorum">
<emphasis id="30F5EA9CFF9AE7637D22F841FBFEC1FC" box="[886,1069,1975,1997]" italics="true" pageId="90" pageNumber="91">Anolis quercorum</emphasis>
</taxonomicName>
. Each symbol can represent one or more adjacent localities. Black circles:
<taxonomicName id="C5814D0DFF9AE7637C70F820FD11C1DD" box="[548,706,2006,2028]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="90" pageNumber="91" phylum="Chordata" rank="species" species="carlliebi">
<emphasis id="30F5EA9CFF9AE7637C70F820FD11C1DD" box="[548,706,2006,2028]" italics="true" pageId="90" pageNumber="91">Anolis carlliebi</emphasis>
</taxonomicName>
; white circles:
<taxonomicName id="C5814D0DFF9AE7637D09F820FC34C1DD" authorityName="Fitch" authorityYear="1978" box="[861,999,2006,2028]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="90" pageNumber="91" phylum="Chordata" rank="species" species="quercorum">
<emphasis id="30F5EA9CFF9AE7637D09F820FC34C1DD" box="[861,999,2006,2028]" italics="true" pageId="90" pageNumber="91">A. quercorum</emphasis>
</taxonomicName>
; white squares:
<taxonomicName id="C5814D0DFF9AE7637AD8F820FA96C1DD" authorityName="Köhler &amp; Pérez &amp; Petersen &amp; Méndez &amp; Cruz" authorityYear="2014" box="[1164,1349,2006,2028]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="90" pageNumber="91" phylum="Chordata" rank="species" species="sacamecatensis">
<emphasis id="30F5EA9CFF9AE7637AD8F820FA96C1DD" box="[1164,1349,2006,2028]" italics="true" pageId="90" pageNumber="91">A. sacamecatensis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="56FE6606FF9BE7627EC3FA00FF1DC07B" ID-DOI="http://doi.org/10.5281/zenodo.4931107" ID-Zenodo-Dep="4931107" httpUri="https://zenodo.org/record/4931107/files/figure.png" pageId="91" pageNumber="92" startId="91.[151,250,1526,1548]" targetBox="[151,1436,195,1502]" targetPageId="91">
<paragraph id="023E368EFF9BE7627EC3FA00FF1DC07B" blockId="91.[151,1435,1526,1610]" pageId="91" pageNumber="92">
<emphasis id="30F5EA9CFF9BE7627EC3FA00FEF1C03D" bold="true" box="[151,290,1526,1548]" pageId="91" pageNumber="92">FIGURE 72.</emphasis>
Syntypes of
<taxonomicName id="C5814D0DFF9BE7627FFAFA00FD47C03D" authorityName="Thominot" authorityYear="1887" box="[430,660,1526,1548]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="91" pageNumber="92" phylum="Chordata" rank="species" species="boulengerianus">
<emphasis id="30F5EA9CFF9BE7627FFAFA00FD47C03D" box="[430,660,1526,1548]" italics="true" pageId="91" pageNumber="92">Anolis boulengerianus</emphasis>
</taxonomicName>
: (a) MNHN 6554 dorsal view; (b) MNHN 6554 ventral view; (c) MNHN 1994.1671 ventral view; (d) MNHN 1994.1671 lateral view; (e) MNHN 1994.1670 dorsal view; (f) MNHN 1994.1670 ventral view.
</paragraph>
</caption>
<subSubSection id="4A9B6505FF9BE7657EC3F959FEBCC156" lastPageId="92" lastPageNumber="93" pageId="91" pageNumber="92" type="discussion">
<paragraph id="023E368EFF9BE7627EC3F959FB5EC0F8" blockId="91.[151,1165,1710,1737]" box="[151,1165,1710,1737]" pageId="91" pageNumber="92">
<emphasis id="30F5EA9CFF9BE7627EC3F959FB5EC0F8" bold="true" box="[151,1165,1710,1737]" pageId="91" pageNumber="92">
Taxonomy of the Mexican anoles related to
<taxonomicName id="C5814D0DFF9BE7627CE1F958FB5EC0F8" authority="Thominot 1887" authorityName="Thominot" authorityYear="1887" box="[693,1165,1710,1737]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="91" pageNumber="92" phylum="Chordata" rank="species" species="boulengerianus">
<emphasis id="30F5EA9CFF9BE7627CE1F958FC1BC0F9" bold="true" box="[693,968,1710,1736]" italics="true" pageId="91" pageNumber="92">Anolis boulengerianus</emphasis>
<bibRefCitation id="66104B7FFF9BE7627D9BF959FB5EC0F8" author="Thominot, A." box="[975,1165,1711,1737]" pageId="91" pageNumber="92" pagination="182 - 190" refId="ref97703" refString="Thominot, A. (1887) Description de trois especes nouvelles d' Anolis et d'un amphisbaenien. Bulletin de la Societe Philomatique de Paris, Series 7, 11, 182 - 190." type="journal article" year="1887">Thominot 1887</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="023E368EFF9BE7657EC3F903FE96C72A" blockId="91.[151,1436,1781,2022]" lastBlockId="92.[151,1436,151,284]" lastPageId="92" lastPageNumber="93" pageId="91" pageNumber="92">
<bibRefCitation id="66104B7FFF9BE7627EC3F903FE8AC13F" author="Thominot, A." box="[151,345,1781,1806]" pageId="91" pageNumber="92" pagination="182 - 190" refId="ref97703" refString="Thominot, A. (1887) Description de trois especes nouvelles d' Anolis et d'un amphisbaenien. Bulletin de la Societe Philomatique de Paris, Series 7, 11, 182 - 190." type="journal article" year="1887">Thominot (1887)</bibRefCitation>
described
<taxonomicName id="C5814D0DFF9BE7627F82F903FD06C13C" authorityName="Thominot" authorityYear="1887" box="[470,725,1781,1805]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="91" pageNumber="92" phylum="Chordata" rank="species" species="boulengerianus">
<emphasis id="30F5EA9CFF9BE7627F82F903FD06C13C" box="[470,725,1781,1805]" italics="true" pageId="91" pageNumber="92">Anolis boulengerianus</emphasis>
</taxonomicName>
based on
<specimenCount id="1487FD07FF9BE7627D18F903FBD1C13F" box="[844,1026,1781,1806]" pageId="91" pageNumber="92" type="generic">three specimens</specimenCount>
(now MNHN 6554, 1994.1670–71;
<figureCitation id="9ABA2A0BFF9BE7627EC3F8EFFF3FC103" box="[151,236,1817,1842]" captionStart="FIGURE 72" captionStartId="91.[151,250,1526,1548]" captionTargetBox="[151,1436,195,1502]" captionTargetId="figure-187@91.[151,1436,195,1502]" captionTargetPageId="91" captionText="FIGURE 72. Syntypes of Anolis boulengerianus: (a) MNHN 6554 dorsal view; (b) MNHN 6554 ventral view; (c) MNHN 1994.1671 ventral view; (d) MNHN 1994.1671 lateral view; (e) MNHN 1994.1670 dorsal view; (f) MNHN 1994.1670 ventral view." figureDoi="http://doi.org/10.5281/zenodo.4931107" httpUri="https://zenodo.org/record/4931107/files/figure.png" pageId="91" pageNumber="92">Fig. 72</figureCitation>
and
<figureCitation id="9ABA2A0BFF9BE7627F7BF8EFFE9DC103" box="[303,334,1817,1842]" captionStart="FIGURE 73" captionStartId="92.[151,250,1244,1266]" captionTargetBox="[151,1436,353,1217]" captionTargetId="figure-386@92.[151,1436,353,1217]" captionTargetPageId="92" captionText="FIGURE 73. Syntypes of Anolis boulengerianus: (a) MNHN 6554 dorsal view of head; (b) MNHN 6554 lateral view of head; (c) MNHN 6554 ventral view of head; (d) MNHN 6554 details of dorsum." figureDoi="http://doi.org/10.5281/zenodo.4931109" httpUri="https://zenodo.org/record/4931109/files/figure.png" pageId="91" pageNumber="92">73</figureCitation>
), examined by GK, from “Téhuantépec (
<collectingCountry id="7A96761EFF9BE7627D69F8EFFC7AC103" box="[829,937,1817,1842]" name="Mexico" pageId="91" pageNumber="92">Mexique</collectingCountry>
).” For most of the time since its original description, this nominal species had remained in the synonymy of
<taxonomicName id="C5814D0DFF9BE7627DCBF8C8FBE2C164" baseAuthorityName="Wiegmann" baseAuthorityYear="1834" box="[927,1073,1853,1877]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="91" pageNumber="92" phylum="Chordata" rank="species" species="nebulosus">
<emphasis id="30F5EA9CFF9BE7627DCBF8C8FBE2C164" box="[927,1073,1853,1877]" italics="true" pageId="91" pageNumber="92">A. nebulosus</emphasis>
</taxonomicName>
or
<taxonomicName id="C5814D0DFF9BE7627A0FF8C8FB26C164" authorityName=": Gehlbach &amp; Colette" authorityYear="1957" box="[1115,1269,1853,1877]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="91" pageNumber="92" phylum="Chordata" rank="species" species="nebuloides">
<emphasis id="30F5EA9CFF9BE7627A0FF8C8FB26C164" box="[1115,1269,1853,1877]" italics="true" pageId="91" pageNumber="92">A. nebuloides</emphasis>
</taxonomicName>
(e.g.,
<bibRefCitation id="66104B7FFF9BE7627B14F8CBFF01C14B" author="Barbour, T." pageId="91" pageNumber="92" pagination="121 - 155" refId="ref87861" refString="Barbour, T. (1934) The anoles. II. The mainland species from Mexico southward. Bulletin of the Museum of Comparative Zoology at Harvard College, 77, 121 - 155." type="journal article" year="1934">Barbour 1934</bibRefCitation>
,
<bibRefCitation id="66104B7FFF9BE7627E89F897FE3CC14B" author="Smith, H. M. &amp; Taylor, E. H." box="[221,495,1889,1914]" pageId="91" pageNumber="92" pagination="1 - 253" refId="ref97125" refString="Smith, H. M. &amp; Taylor, E. H. (1950 a) An annotated checklist and key to the reptiles of Mexico exclusive of the snakes. Bulletin of the United States National Museum, 199, 1 - 253." type="journal article" year="1950">
Smith and Taylor 1950
<bibRefCitation id="66104B7FFF9BE7627FB6F897FE3CC14B" author="Peters, J. A." box="[482,495,1889,1914]" pageId="91" pageNumber="92" pagination="1 - 37" refId="ref94820" refString="Peters, J. A. (1954) The amphibians and reptiles of the coast and coastal sierra of Michoacan, Mexico. Occasional Papers of the Museum of Zoology University of Michigan, 554, 1 - 37." type="journal article" year="1954">a</bibRefCitation>
</bibRefCitation>
, Guibé 1954,
<bibRefCitation id="66104B7FFF9BE7627CD9F894FCF6C14B" author="Brygoo, E. R." box="[653,805,1889,1914]" pageId="91" pageNumber="92" pagination="1 - 112" refId="ref88294" refString="Brygoo, E. R. (1989) Les types d'Iguanides (Reptiles, Sauriens) du Museum national d'Histoire naturelle Catalogue critique. Bulletin Museum National d'Histoire Naturelle, 4, Series 11, Section A, No. 3 (Supplement), 1 - 112." type="book chapter" year="1989">Brygoo 1989</bibRefCitation>
).
<bibRefCitation id="66104B7FFF9BE7627D6CF897FC1FC14B" author="Davis, W. B." box="[824,972,1889,1914]" pageId="91" pageNumber="92" pagination="1 - 6" refId="ref89150" refString="Davis, W. B. (1954) Three new anoles from Mexico. Herpetologica, 10 (1), 1 - 6." type="journal article" year="1954">Davis (1954)</bibRefCitation>
described
<taxonomicName id="C5814D0DFF9BE7627A12F894FB3AC148" authorityName="Davis" authorityYear="1954" box="[1094,1257,1889,1913]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="91" pageNumber="92" phylum="Chordata" rank="species" species="subocularis">
<emphasis id="30F5EA9CFF9BE7627A12F894FB3AC148" box="[1094,1257,1889,1913]" italics="true" pageId="91" pageNumber="92">A. subocularis</emphasis>
</taxonomicName>
from “one mile southwest of Tierra Colorado,
<quantity id="C5799B6BFF9BE7627FBAF873FDE9C1AF" box="[494,570,1925,1950]" metricMagnitude="2" metricUnit="m" metricValue="2.7432" pageId="91" pageNumber="92" unit="ft" value="900.0">900 ft.</quantity>
,
<collectingRegion id="C045F86CFF9BE7627C12F873FD7EC1AF" box="[582,685,1925,1950]" country="Mexico" name="Guerrero" pageId="91" pageNumber="92">Guerrero</collectingRegion>
” (
<typeStatus id="DD3A882CFF9BE7627C93F873FCF8C1AF" box="[711,811,1925,1950]" pageId="91" pageNumber="92" type="holotype">holotype</typeStatus>
now TCWC 8675,
<figureCitation id="9ABA2A0BFF9BE7627A5CF873FB88C1AF" box="[1032,1115,1925,1950]" captionStart="FIGURE 74" captionStartId="93.[151,250,1552,1574]" captionTargetBox="[155,1436,202,1508]" captionTargetId="figure-67@93.[155,1440,202,1508]" captionTargetPageId="93" captionText="FIGURE 74. Holotype of Anolis subocularis (TCWC 8675): (a) dorsal view; (b) lateral view; (c) ventral view; (d) lateral view of head; (e) dorsal view of head; (f) ventral view of head. Scale bars equal 5.0 mm in (a–c) and 1.0 mm in (d–f), respectively." figureDoi="http://doi.org/10.5281/zenodo.4931111" httpUri="https://zenodo.org/record/4931111/files/figure.png" pageId="91" pageNumber="92">Fig. 74</figureCitation>
), examined by GK. In 1978, Fitch described
<taxonomicName id="C5814D0DFF9BE7627F1DF85CFE01C1F0" authorityName="Fitch" authorityYear="1978" box="[329,466,1961,1985]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="91" pageNumber="92" phylum="Chordata" rank="species" species="isthmicus">
<emphasis id="30F5EA9CFF9BE7627F1DF85CFE01C1F0" box="[329,466,1961,1985]" italics="true" pageId="91" pageNumber="92">A. isthmicus</emphasis>
</taxonomicName>
based on specimens from “
<quantity id="C5799B6BFF9BE7627D52F85FFCB7C1F3" box="[774,868,1961,1986]" metricMagnitude="4" metricUnit="m" metricValue="1.95" pageId="91" pageNumber="92" unit="km" value="19.5">19.5 km</quantity>
WNW Tehuantepec,
<collectingRegion id="C045F86CFF9BE7627A02F85FFB7EC1F3" box="[1110,1197,1961,1986]" country="Mexico" name="Oaxaca" pageId="91" pageNumber="92">Oaxaca</collectingRegion>
,
<collectingCountry id="7A96761EFF9BE7627AE3F85FFADCC1F3" box="[1207,1295,1961,1986]" name="Mexico" pageId="91" pageNumber="92">Mexico</collectingCountry>
” and nearby localities. In his comparisons, he differentiated it from a selection of anole species from southern
<collectingCountry id="7A96761EFF9BE7627ABCF83BFA93C1D7" box="[1256,1344,1997,2022]" name="Mexico" pageId="91" pageNumber="92">Mexico</collectingCountry>
, but did not include
<taxonomicName id="C5814D0DFF9CE7657F4EFF6EFE34C681" authorityName="Thominot" authorityYear="1887" box="[282,487,152,176]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="boulengerianus">
<emphasis id="30F5EA9CFF9CE7657F4EFF6EFE34C681" box="[282,487,152,176]" italics="true" pageId="92" pageNumber="93">A. boulengerianus</emphasis>
</taxonomicName>
in his considerations. Recently,
<bibRefCitation id="66104B7FFF9CE7657D04FF61FB1DC681" author="Nieto Montes de Oca, A. &amp; Kohler, G. &amp; Feria-Ortiz, M." box="[848,1230,151,176]" pageId="92" pageNumber="93" pagination="125 - 133" refId="ref94533" refString="Nieto Montes de Oca, A., Kohler, G. &amp; Feria-Ortiz, M. (2014) Anolis boulengerianus Thominot 1887, a senior synonym of Anolis isthmicus Fitch 1978 (Squamata: Dactyloidae). Zootaxa, 3794 (1), 125 - 133. http: // dx. doi. org / 10.11646 / zootaxa. 3794.1.5" type="journal article" year="2014">
Nieto Montes de Oca
<emphasis id="30F5EA9CFF9CE7657A11FF6FFBAEC681" box="[1093,1149,152,176]" italics="true" pageId="92" pageNumber="93">et al.</emphasis>
(2014)
</bibRefCitation>
demonstrated that the type material of
<taxonomicName id="C5814D0DFF9CE7657F23FF4AFD97C6E5" authorityName="Thominot" authorityYear="1887" box="[375,580,188,212]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="boulengerianus">
<emphasis id="30F5EA9CFF9CE7657F23FF4AFD97C6E5" box="[375,580,188,212]" italics="true" pageId="92" pageNumber="93">A. boulengerianus</emphasis>
</taxonomicName>
is conspecific with that of
<taxonomicName id="C5814D0DFF9CE7657D27FF4AFC2FC6E5" authorityName="Fitch" authorityYear="1978" box="[883,1020,188,212]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="isthmicus">
<emphasis id="30F5EA9CFF9CE7657D27FF4AFC2FC6E5" box="[883,1020,188,212]" italics="true" pageId="92" pageNumber="93">A. isthmicus</emphasis>
</taxonomicName>
and therefore synonymized the latter with
<taxonomicName id="C5814D0DFF9CE7657E84FF16FE4EC6C9" authorityName="Thominot" authorityYear="1887" box="[208,413,224,248]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="boulengerianus">
<emphasis id="30F5EA9CFF9CE7657E84FF16FE4EC6C9" box="[208,413,224,248]" italics="true" pageId="92" pageNumber="93">A. boulengerianus</emphasis>
</taxonomicName>
. Because of priority,
<taxonomicName id="C5814D0DFF9CE7657CDAFF16FC8FC6C9" authorityName="Thominot" authorityYear="1887" box="[654,860,224,248]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="boulengerianus">
<emphasis id="30F5EA9CFF9CE7657CDAFF16FC8FC6C9" box="[654,860,224,248]" italics="true" pageId="92" pageNumber="93">A. boulengerianus</emphasis>
</taxonomicName>
is the valid name for the species previously known as
<taxonomicName id="C5814D0DFF9CE7657EE1FEF2FEEDC72D" authorityName="Fitch" authorityYear="1978" box="[181,318,260,284]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="isthmicus">
<emphasis id="30F5EA9CFF9CE7657EE1FEF2FEEDC72D" box="[181,318,260,284]" italics="true" pageId="92" pageNumber="93">A. isthmicus</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="56FE6606FF9CE7657EC3FB2AFC59C321" ID-DOI="http://doi.org/10.5281/zenodo.4931109" ID-Zenodo-Dep="4931109" httpUri="https://zenodo.org/record/4931109/files/figure.png" pageId="92" pageNumber="93" startId="92.[151,250,1244,1266]" targetBox="[151,1436,353,1217]" targetPageId="92">
<paragraph id="023E368EFF9CE7657EC3FB2AFC59C321" blockId="92.[151,1436,1244,1296]" pageId="92" pageNumber="93">
<emphasis id="30F5EA9CFF9CE7657EC3FB2AFEF3C2C0" bold="true" box="[151,288,1244,1266]" pageId="92" pageNumber="93">FIGURE 73.</emphasis>
Syntypes of
<taxonomicName id="C5814D0DFF9CE7657FF0FB2AFD5AC2C3" authorityName="Thominot" authorityYear="1887" box="[420,649,1244,1266]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="boulengerianus">
<emphasis id="30F5EA9CFF9CE7657FF0FB2AFD5AC2C3" box="[420,649,1244,1266]" italics="true" pageId="92" pageNumber="93">Anolis boulengerianus</emphasis>
</taxonomicName>
: (a) MNHN 6554 dorsal view of head; (b) MNHN 6554 lateral view of head; (c) MNHN 6554 ventral view of head; (d) MNHN 6554 details of dorsum.
</paragraph>
</caption>
<paragraph id="023E368EFF9CE7657E93FAA1FEBCC156" blockId="92.[151,1437,1366,1895]" pageId="92" pageNumber="93">
For our analysis, we defined three OTUs in this complex: OTU1 is formed by specimens from south-central and southeastern
<collectingRegion id="C045F86CFF9CE7657F0EFA8CFE10C3A2" box="[346,451,1402,1427]" country="Mexico" name="Guerrero" pageId="92" pageNumber="93">Guerrero</collectingRegion>
; OTU2 by specimens from the coastal plain between Huatulco and Puerto Escondido in south-central
<collectingRegion id="C045F86CFF9CE7657F65FA68FE5AC386" box="[305,393,1438,1463]" country="Mexico" name="Oaxaca" pageId="92" pageNumber="93">Oaxaca</collectingRegion>
; and OTU 3 by specimens from the hills west and northwest of Tehuantepec in southeastern
<collectingRegion id="C045F86CFF9CE7657EC3FA34FF3CC3EA" box="[151,239,1474,1499]" country="Mexico" name="Oaxaca" pageId="92" pageNumber="93">Oaxaca</collectingRegion>
. The analysis of the CO1 and 16s gene fragments revealed the existence of three genetic clusters (
<figureCitation id="9ABA2A0BFF9CE7657B68FA34FA43C3EA" box="[1340,1424,1474,1499]" captionStart="FIGURE 75" captionStartId="94.[151,250,1931,1953]" captionTargetBox="[251,1331,200,1904]" captionTargetId="figure-82@94.[344,1242,195,1227]" captionTargetPageId="94" captionText="FIGURE 75. Phylogenetic tree of specimens of the species related to Anolis boulengerianus from a maximum-likelihood analysis of DNA sequences of two mitochondrial genes: (a) 16S rRNA and (b) CO1. A scale bar is indicated. The numbers at nodes are bootstrap values. The tree is rooted with the species Anolis macrinii. For designation of OTUs see text." figureDoi="http://doi.org/10.5281/zenodo.4931113" httpUri="https://zenodo.org/record/4931113/files/figure.png" pageId="92" pageNumber="93">Fig. 75</figureCitation>
). Genetic distances between our OTUs 2 and 3 are low in both genes (5.7–7.7% in CO1;
<quantity id="C5799B6BFF9CE7657A3DFA10FB09C3CE" box="[1129,1242,1510,1535]" metricMagnitude="-2" metricUnit="m" metricValue="5.460999999999999" metricValueMax="7.111999999999999" metricValueMin="3.81" pageId="92" pageNumber="93" unit="in" value="2.15" valueMax="2.8" valueMin="1.5">1.5–2.8 in</quantity>
16s). In contrast, our OTU1 has a genetic distance to OTU 2 and 3, respectively, of 13.9–17.6 (CO1) and 6.9–7.8 (16s), respectively. We observed differences in male dewlap coloration between OTUs 2 and 3: in OTU 2, the male dewlap is pinkish red without paler areas around gorgetals whereas in OTU 3 the male dewlap is orange yellow with paler areas around gorgetals. Also, the eastern populations (our OTU 3) occur in a much more arid habitat than the western populations (our OTU 2). These findings are evidence for the existence of three species in this complex. Our OTU 1 contains the
<typeStatus id="DD3A882CFF9CE7657F6BF949FEBCC0E6" box="[319,367,1727,1751]" pageId="92" pageNumber="93">type</typeStatus>
locality of
<taxonomicName id="C5814D0DFF9CE7657FA2F949FD49C0E7" authorityName="Davis" authorityYear="1954" box="[502,666,1726,1750]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="subocularis">
<emphasis id="30F5EA9CFF9CE7657FA2F949FD49C0E7" box="[502,666,1726,1750]" italics="true" pageId="92" pageNumber="93">A. subocularis</emphasis>
</taxonomicName>
and therefore this name has to be applied to this species. The
<typeStatus id="DD3A882CFF9CE7657B38F949FA4FC0E6" box="[1388,1436,1727,1751]" pageId="92" pageNumber="93">type</typeStatus>
localities of
<taxonomicName id="C5814D0DFF9CE7657F75F915FE3EC0CA" authorityName="Thominot" authorityYear="1887" box="[289,493,1763,1787]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="boulengerianus">
<emphasis id="30F5EA9CFF9CE7657F75F915FE3EC0CA" box="[289,493,1763,1787]" italics="true" pageId="92" pageNumber="93">A. boulengerianus</emphasis>
</taxonomicName>
and
<taxonomicName id="C5814D0DFF9CE7657C70F915FD7FC0CA" authorityName="Fitch" authorityYear="1978" box="[548,684,1763,1787]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="isthmicus">
<emphasis id="30F5EA9CFF9CE7657C70F915FD7FC0CA" box="[548,684,1763,1787]" italics="true" pageId="92" pageNumber="93">A. isthmicus</emphasis>
</taxonomicName>
, respectively, are both contained in our OTU3; thus the older name (i.e.,
<taxonomicName id="C5814D0DFF9CE7657E80F8F1FE77C12F" authorityName="Thominot" authorityYear="1887" box="[212,420,1798,1822]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="boulengerianus">
<emphasis id="30F5EA9CFF9CE7657E80F8F1FE77C12F" box="[212,420,1798,1822]" italics="true" pageId="92" pageNumber="93">A. boulengerianus</emphasis>
</taxonomicName>
) is the valid name for this species and the nominal species
<taxonomicName id="C5814D0DFF9CE7657A35F8F1FB3EC12F" authorityName="Fitch" authorityYear="1978" box="[1121,1261,1798,1822]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="isthmicus">
<emphasis id="30F5EA9CFF9CE7657A35F8F1FB3EC12F" box="[1121,1261,1798,1822]" italics="true" pageId="92" pageNumber="93">A. isthmicus</emphasis>
</taxonomicName>
remains in the synonymy of
<taxonomicName id="C5814D0DFF9CE7657F67F8DDFDD1C172" authorityName="Thominot" authorityYear="1887" box="[307,514,1835,1859]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="92" pageNumber="93" phylum="Chordata" rank="species" species="boulengerianus">
<emphasis id="30F5EA9CFF9CE7657F67F8DDFDD1C172" box="[307,514,1835,1859]" italics="true" pageId="92" pageNumber="93">A. boulengerianus</emphasis>
</taxonomicName>
. No scientific name is available for our OTU 2, and we therefore describe it as a new species below.
</paragraph>
</subSubSection>
</treatment>
</document>
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