512 lines
58 KiB
XML
512 lines
58 KiB
XML
<document ID-DOI="http://dx.doi.org/10.11646/zootaxa.3862.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="4930959" ID-ZooBank="3FA375FE-E4E0-4509-BE02-EE5E786B07C6" approvalRequired="20" approvalRequired_for_treatments="20" checkinTime="1623452360688" checkinUser="felipe" docAuthor="Köhler, Gunther, Pérez, Raúl Gómez Trejo, Petersen, Claus Bo P., Méndez, Fausto R. & Cruz, De La" docDate="2014" docId="8A288798FF85E7757EC3FDAAFEE7C7EA" docLanguage="en" docName="zt03862p210.pdf.imf" docOrigin="Zootaxa 3862 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.10:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="10" docTitle="Anolis carlliebi Köhler & Pérez & Petersen & Méndez & Cruz 2014, sp. nov." docType="treatment" docVersion="6" lastPageNumber="77" masterDocId="7611FFE0FFC0E7397E54FFF6FFD3C631" masterDocTitle="A revision of the Mexican Anolis (Reptilia, Squamata, Dactyloidae) from the Pacific versant west of the Isthmus de Tehuantepec in the states of Oaxaca, Guerrero, and Puebla, with the description of six new species" masterLastPageNumber="210" masterPageNumber="1" pageNumber="70" updateTime="1673621398369" updateUser="juliana">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of the Mexican Anolis (Reptilia, Squamata, Dactyloidae) from the Pacific versant west of the Isthmus de Tehuantepec in the states of Oaxaca, Guerrero, and Puebla, with the description of six new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Köhler, Gunther</mods:namePart>
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<mods:nameIdentifier type="email">gkoehler@senckenberg.de</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pérez, Raúl Gómez Trejo</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Petersen, Claus Bo P.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Méndez, Fausto R.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cruz, De La</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2014-09-19</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>3862</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>210</mods:end>
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</mods:extent>
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</mods:part>
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||
</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.11646/zootaxa.3862.1.1</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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||
<mods:identifier type="Zenodo-Dep">4930959</mods:identifier>
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<mods:identifier type="ZooBank">3FA375FE-E4E0-4509-BE02-EE5E786B07C6</mods:identifier>
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</mods:mods>
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||
<treatment LSID="urn:lsid:plazi:treatment:8A288798FF85E7757EC3FDAAFEE7C7EA" httpUri="http://treatment.plazi.org/id/8A288798FF85E7757EC3FDAAFEE7C7EA" lastPageId="76" lastPageNumber="77" pageId="69" pageNumber="70">
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<subSubSection box="[151,436,604,630]" pageId="69" pageNumber="70" type="nomenclature">
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<paragraph blockId="69.[151,436,604,664]" box="[151,436,604,630]" pageId="69" pageNumber="70">
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<heading bold="true" box="[151,436,604,630]" fontSize="11" level="2" pageId="69" pageNumber="70" reason="4">
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<emphasis bold="true" box="[151,436,604,630]" pageId="69" pageNumber="70">
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<taxonomicName authority="Köhler & Pérez & Petersen & Méndez & Cruz, 2014" authorityName="Köhler & Pérez & Petersen & Méndez & Cruz" authorityYear="2014" box="[151,335,604,630]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="69" pageNumber="70" phylum="Chordata" rank="species" species="carlliebi" status="sp. nov.">
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<emphasis bold="true" box="[151,335,604,630]" italics="true" pageId="69" pageNumber="70">Anolis carlliebi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[343,436,604,630]" pageId="69" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,283,639,664]" pageId="69" pageNumber="70" type="description">
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<paragraph blockId="69.[151,436,604,664]" box="[151,283,639,664]" pageId="69" pageNumber="70">
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<figureCitation box="[151,283,639,664]" captionStart-0="FIGURE 52" captionStart-1="FIGURE 53" captionStart-2="FIGURE 54" captionStart-3="FIGURE 55" captionStart-4="FIGURE 56" captionStart-5="FIGURE 57" captionStartId-0="70.[151,250,1885,1907]" captionStartId-1="71.[151,250,1528,1550]" captionStartId-2="72.[151,250,2014,2036]" captionStartId-3="73.[151,250,1962,1984]" captionStartId-4="74.[151,250,1515,1537]" captionStartId-5="75.[151,250,1390,1412]" captionTargetBox-0="[218,1352,315,1864]" captionTargetBox-1="[155,1436,199,1505]" captionTargetBox-2="[157,1436,679,1986]" captionTargetBox-3="[151,1434,193,1942]" captionTargetBox-4="[159,1436,187,1493]" captionTargetBox-5="[257,1330,613,1369]" captionTargetId-0="figure-107@70.[218,1352,315,1864]" captionTargetId-1="figure-255@71.[155,1440,199,1505]" captionTargetId-2="figure-278@72.[157,1442,679,1986]" captionTargetId-3="figure-65@73.[149,1434,193,1942]" captionTargetId-4="figure-237@74.[159,1444,187,1493]" captionTargetId-5="figure-455@75.[257,1330,613,1369]" captionTargetPageId-0="70" captionTargetPageId-1="71" captionTargetPageId-2="72" captionTargetPageId-3="73" captionTargetPageId-4="74" captionTargetPageId-5="75" captionText-0="FIGURE 52. Anolis carlliebi in life: (a) two males from Ixtlán de Juárez, Oaxaca; (b) a male from San Juan Tepeuxila, Oaxaca. Photos by Christopher Ehecatl Vega Flores (a) and Luis Canseco Marquéz (b)." captionText-1="FIGURE 53. Holotype of Anolis carlliebi (SMF 96714): (a) dorsal view; (b) lateral view; (c) ventral view; (d) lateral view of head; (e) dorsal view of head; (f) ventral view of head. Scale bars equal 5.0 mm in (a–c) and 1.0 mm in (d–f), respectively." captionText-2="FIGURE 54. Anolis carlliebi: head scalation in (a,c,e) MVZ 144181; (b,d,f) UTA R-52816. Scale bars equal 1.0 mm." captionText-3="FIGURE 55. Anolis carlliebi: nasal region in (a) UTA R-52818; (b) SMF 96714; superciliary region in (c) UTA R-52818; (d) MVZ 144181; chin region in (e) MVZ 164275; (f) SMF 96714; cloacal region in (g) UTA R-52816. Scale bars equal 1.0 mm." captionText-4="FIGURE 56. Anolis carlliebi: dorsal scales in (a) UTA R-52818; (b) SMF 96714; flank scales in (c) UTA R-52818; (d) SMF 96714; ventral scales in (e) UTA R-52816; (f) SMF 96714. Scale bars equal 1.0 mm." captionText-5="FIGURE 57. Hemipenis of Anolis carlliebi (UTA R-52816): (left) sulcate view; (right) asulcate view. Scale bar equals 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4931067" figureDoi-1="http://doi.org/10.5281/zenodo.4931069" figureDoi-2="http://doi.org/10.5281/zenodo.4931071" figureDoi-3="http://doi.org/10.5281/zenodo.4931073" figureDoi-4="http://doi.org/10.5281/zenodo.4931075" figureDoi-5="http://doi.org/10.5281/zenodo.4931077" httpUri-0="https://zenodo.org/record/4931067/files/figure.png" httpUri-1="https://zenodo.org/record/4931069/files/figure.png" httpUri-2="https://zenodo.org/record/4931071/files/figure.png" httpUri-3="https://zenodo.org/record/4931073/files/figure.png" httpUri-4="https://zenodo.org/record/4931075/files/figure.png" httpUri-5="https://zenodo.org/record/4931077/files/figure.png" pageId="69" pageNumber="70">Figs. 52–57</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="69" pageNumber="70" type="reference_group">
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<paragraph blockId="69.[151,1436,711,794]" box="[151,462,711,733]" pageId="69" pageNumber="70">
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<taxonomicName authority=": Smith 1939" authorityName="Smith" authorityYear="1939" box="[151,462,711,733]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="69" pageNumber="70" phylum="Chordata" rank="species" species="nebuloides">
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<emphasis box="[151,332,711,733]" italics="true" pageId="69" pageNumber="70">Anolis nebuloides</emphasis>
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:
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<bibRefCitation author="Smith, H. M." box="[345,462,711,733]" pageId="69" pageNumber="70" pagination="15 - 35" refId="ref96249" refString="Smith, H. M. (1939) Notes on Mexican reptiles and amphibians. Zoological Series of Field Museum of Natural History, 24 (4), 15 - 35." type="journal article" year="1939">Smith 1939</bibRefCitation>
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</taxonomicName>
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</paragraph>
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<paragraph blockId="69.[151,1436,711,794]" pageId="69" pageNumber="70">
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<taxonomicName authority=": Ramirez-Bautista et al. 2002 a" authorityName="Ramirez-Bautista" authorityYear="2002" box="[151,673,742,764]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="69" pageNumber="70" phylum="Chordata" rank="species" species="quercorum">
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<emphasis box="[151,336,742,764]" italics="true" pageId="69" pageNumber="70">Anolis quercorum</emphasis>
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:
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<bibRefCitation author="Ramirez-Bautista, A. & Oliver-Lopez, L. & Mata-Silva, V." box="[354,673,742,764]" pageId="69" pageNumber="70" pagination="203 - 204" refId="ref95502" refString="Ramirez-Bautista, A., Oliver-Lopez, L. & Mata-Silva, V. (2002 a) Anolis quercorum (Gray Anole). General Ecology. Herpetological Review, 33, 203 - 204" type="journal article" year="2002">
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Ramirez-Bautista
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<emphasis box="[542,597,742,764]" italics="true" pageId="69" pageNumber="70">et al.</emphasis>
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2002a
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</bibRefCitation>
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</taxonomicName>
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,b,
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<bibRefCitation author="Ramirez-Bautista, A." box="[705,945,742,764]" pageId="69" pageNumber="70" pagination="328 - 331" refId="ref95388" refString="Ramirez-Bautista, A. (2003) Some reproductive characteristics of a tropical arid lizard assemblage from Zapotitlan Salinas, Puebla, Mexico. Herpetological Review, 34 (4), 328 - 331." type="journal article" year="2003">Ramirez-Bautista 2003</bibRefCitation>
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,
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<bibRefCitation author="Canseco-Marquez, L. & Gutierrrez-Mayen, M. G." box="[960,1431,742,764]" pageId="69" pageNumber="70" refId="ref88500" refString="Canseco-Marquez, L. & Gutierrrez-Mayen, M. G. (2006) Guia del campo de los anfibios y reptiles del Valle de Zapotitlan, Puebla. Sociedad Herpetologica Mexicana, A. C. Escuela de Biologia, BUAP, 78 pp." type="book" year="2006">Canseco-Márquez & Gutiérrrez-Mayén 2006</bibRefCitation>
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, 2010
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</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="69" pageNumber="70" type="materials_examined">
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||
<paragraph blockId="69.[151,1437,837,2021]" pageId="69" pageNumber="70">
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||
<materialsCitation accessionNumber="SMF 96714" collectingDate="2013-06-13" collectionCode="SMF" collectorName="de Juarez" country="Mexico" elevation="1945" latitude="17.31635" location="Ixtlan de Juarez" longLatPrecision="1" longitude="-96.48347" pageId="69" pageNumber="70" specimenCode="WGS84, GK-4501" specimenCount="1" stateProvince="Oaxaca" typeStatus="holotype">
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||
<emphasis bold="true" box="[151,260,837,862]" pageId="69" pageNumber="70">
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||
<typeStatus box="[151,260,837,862]" pageId="69" pageNumber="70">Holotype</typeStatus>
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</emphasis>
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.
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<accessionNumber box="[276,415,837,862]" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF96714" pageId="69" pageNumber="70">
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<collectionCode box="[276,332,837,862]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="69" pageNumber="70">SMF</collectionCode>
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96714
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</accessionNumber>
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, an adult female from
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<location LSID="urn:lsid:plazi:treatment:8A288798FF85E7757EC3FDAAFEE7C7EA:075E6055FF85E77C7CFEFCB3FCB0C56F" box="[682,867,837,862]" country="Mexico" latitude="17.31635" longLatPrecision="1" longitude="-96.48347" name="Ixtlan de Juarez" pageId="69" pageNumber="70" stateProvince="Oaxaca">
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Ixtlán
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<collectorName box="[757,867,837,862]" pageId="69" pageNumber="70">de Juárez</collectorName>
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</location>
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, university campus (
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<geoCoordinate box="[1106,1261,837,862]" degrees="17.316350" direction="north" orientation="latitude" pageId="69" pageNumber="70" precision="1" value="17.31635">17.316350°N</geoCoordinate>
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,
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<geoCoordinate box="[1275,1432,837,862]" degrees="96.483470" direction="west" orientation="longitude" pageId="69" pageNumber="70" precision="1" value="-96.48347">96.483470°W</geoCoordinate>
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,
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<specimenCode box="[151,245,875,900]" collectionCode="WGS" pageId="69" pageNumber="70">WGS84</specimenCode>
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),
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<quantity box="[268,359,875,900]" metricMagnitude="3" metricUnit="m" metricValue="1.9449999999999998" pageId="69" pageNumber="70" unit="m" value="1945.0">
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||
<elevation box="[268,359,875,900]" metricMagnitude="3" metricUnit="m" metricValue="1.9449999999999998" pageId="69" pageNumber="70" unit="m" value="1945.0">1945 m</elevation>
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</quantity>
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,
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<collectingRegion box="[375,587,875,900]" country="Mexico" name="Oaxaca" pageId="69" pageNumber="70">Estado de Oaxaca</collectingRegion>
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,
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<collectingCountry box="[602,691,875,900]" name="Mexico" pageId="69" pageNumber="70">Mexico</collectingCountry>
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; collected
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<date box="[819,976,875,900]" pageId="69" pageNumber="70" value="2013-06-13">
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<collectingDate box="[819,976,875,900]" pageId="69" pageNumber="70" value="2013-06-13">13 June 2013</collectingDate>
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</date>
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by Raúl Gómez Trejo Pérez. Field tag number
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<specimenCode box="[243,353,912,937]" collectionCode="GK-" pageId="69" pageNumber="70">GK-4501</specimenCode>
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.
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</materialsCitation>
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</paragraph>
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<paragraph blockId="69.[151,1437,837,2021]" pageId="69" pageNumber="70">
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<materialsCitation accessionNumber="SMF 96714" collectingDate="1947-05-11" collectionCode="IBH" collectorName="de Juarez & Jonathan A. Campbell" country="Mexico" county="Ixtlan de Juarez" elevation="1619" latitude="17.732" location="Oaxaca" longLatPrecision="76" longitude="-96.866" municipality="Hellmuth Wagner" pageId="69" pageNumber="70" specimenCode="GK-4502" specimenCount="1" stateProvince="Oaxaca" typeStatus="paratype">
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||
<emphasis bold="true" box="[199,319,949,974]" pageId="69" pageNumber="70">
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<typeStatus box="[199,319,949,974]" pageId="69" pageNumber="70">Paratypes</typeStatus>
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</emphasis>
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. All from the Mexican state of
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<collectingRegion box="[700,789,949,974]" country="Mexico" name="Oaxaca" pageId="69" pageNumber="70">Oaxaca</collectingRegion>
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:
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<specimenCode box="[803,909,949,974]" collectionCode="GK-" pageId="69" pageNumber="70">GK-4502</specimenCode>
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(
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<collectionCode box="[928,978,949,974]" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="69" pageNumber="70">IBH</collectionCode>
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uncatalogued), same collecting data as holotype;
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</materialsCitation>
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<materialsCitation accessionNumber="SMF 96714" box="[267,1436,987,1012]" collectingDate="1947-05-11" collectionCode="IBH" collectorName="de Juarez & Jonathan A. Campbell" country="Mexico" county="Ixtlan de Juarez" elevation="1619" latitude="17.732" location="Oaxaca" longLatPrecision="76" longitude="-96.866" municipality="Hellmuth Wagner" pageId="69" pageNumber="70" specimenCode="GK-4502" specimenCount="1" stateProvince="Oaxaca" typeStatus="paratype">
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<specimenCode box="[267,434,987,1012]" collectionCode="UMMZ" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="69" pageNumber="70" type="University or college">UMMZ 99820</specimenCode>
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(
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<specimenCount box="[451,626,987,1012]" pageId="69" pageNumber="70" type="generic">two specimens</specimenCount>
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), from
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<collectingCounty box="[712,899,987,1012]" pageId="69" pageNumber="70">
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Ixtlán
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<collectorName box="[788,899,987,1012]" pageId="69" pageNumber="70">de Juárez</collectorName>
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</collectingCounty>
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, collected
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<date box="[1024,1178,987,1012]" pageId="69" pageNumber="70" value="1947-05-11">
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<collectingDate box="[1024,1178,987,1012]" pageId="69" pageNumber="70" value="1947-05-11">11 May 1947</collectingDate>
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</date>
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by
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<collectingMunicipality box="[1226,1432,987,1012]" pageId="69" pageNumber="70">Hellmuth Wagner</collectingMunicipality>
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||
;
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</materialsCitation>
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<materialsCitation accessionNumber="SMF 96714" collectingDate="1947-05-11" collectionCode="IBH" collectorName="de Juarez & Jonathan A. Campbell" country="Mexico" county="Ixtlan de Juarez" elevation="1619" latitude="17.732" location="Oaxaca" longLatPrecision="76" longitude="-96.866" municipality="Hellmuth Wagner" pageId="69" pageNumber="70" specimenCode="GK-4502" specimenCount="1" stateProvince="Oaxaca" typeStatus="paratype">
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<collectingRegion box="[151,206,1024,1049]" country="United States of America" name="Utah" pageId="69" pageNumber="70">UTA</collectingRegion>
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R-8421, from 0.9 mi N (by road)
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<location LSID="urn:lsid:plazi:treatment:8A288798FF85E7757EC3FDAAFEE7C7EA:075E6055FF85E77C7C3BFBF6FC6CC228" box="[623,959,1024,1049]" county="Ixtlan de Juarez" latitude="17.732" longLatPrecision="76" longitude="-96.866" municipality="Hellmuth Wagner" name="El Cerezal on Mexican Hwy" pageId="69" pageNumber="70" stateProvince="Utah">El Cerezal on Mexican Hwy</location>
|
||
175,
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||
<location LSID="urn:lsid:plazi:treatment:8A288798FF85E7757EC3FDAAFEE7C7EA:075E6055FF85E77C7A52FBF6FB08C228" box="[1030,1243,1024,1049]" county="Ixtlan de Juarez" latitude="17.732" longLatPrecision="76" longitude="-96.866" municipality="Hellmuth Wagner" name="Sierra Aloapaneca" pageId="69" pageNumber="70" stateProvince="Utah">Sierra Aloapaneca</location>
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, by
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<collectorName pageId="69" pageNumber="70">Jonathan A. Campbell</collectorName>
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||
;
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</materialsCitation>
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<materialsCitation accessionNumber="SMF 96714" collectingDate="1947-05-11" collectionCode="IBH" collectorName="de Juarez & Jonathan A. Campbell" country="Mexico" county="Ixtlan de Juarez" elevation="1619" latitude="17.732" location="Oaxaca" longLatPrecision="76" longitude="-96.866" municipality="Hellmuth Wagner" pageId="69" pageNumber="70" specimenCode="GK-4502" specimenCount="1" stateProvince="Oaxaca" typeStatus="paratype">
|
||
<collectingRegion box="[276,331,1061,1086]" country="United States of America" name="Utah" pageId="69" pageNumber="70">UTA</collectingRegion>
|
||
R-52816–17, from
|
||
<location LSID="urn:lsid:plazi:treatment:8A288798FF85E7757EC3FDAAFEE7C7EA:075E6055FF85E77C7C7BFBD0FD6CC20F" box="[559,703,1062,1086]" county="Ixtlan de Juarez" latitude="17.732" longLatPrecision="76" longitude="-96.866" municipality="Hellmuth Wagner" name="Tutepetongo" pageId="69" pageNumber="70" stateProvince="Utah">Tutepetongo</location>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:8A288798FF85E7757EC3FDAAFEE7C7EA:075E6055FF85E77C7C9FFBD3FC65C20F" box="[715,950,1061,1086]" county="Ixtlan de Juarez" latitude="17.732" longLatPrecision="76" longitude="-96.866" municipality="Hellmuth Wagner" name="Municipio Cuicatlan" pageId="69" pageNumber="70" stateProvince="Utah">Municipio Cuicatlán</location>
|
||
,
|
||
<geoCoordinate box="[963,1074,1061,1086]" degrees="17.732" direction="north" orientation="latitude" pageId="69" pageNumber="70" precision="55" value="17.732">17.732°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[1087,1202,1061,1086]" degrees="96.866" direction="west" orientation="longitude" pageId="69" pageNumber="70" precision="55" value="-96.866">96.866°W</geoCoordinate>
|
||
, 1619 m, by
|
||
<collectorName pageId="69" pageNumber="70">Eric N. Smith</collectorName>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation accessionNumber="SMF 96714" box="[232,1235,1099,1124]" collectingDate="1947-05-11" collectionCode="IBH" collectorName="de Juarez & Jonathan A. Campbell" country="Mexico" county="Ixtlan de Juarez" elevation="1619" latitude="17.732" location="Oaxaca" longLatPrecision="76" longitude="-96.866" municipality="Hellmuth Wagner" pageId="69" pageNumber="70" specimenCode="GK-4502" specimenCount="1" stateProvince="Oaxaca" typeStatus="paratype">
|
||
<specimenCode box="[232,431,1099,1124]" collectionCode="MVZ" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34777" lsid="urn:lsid:biocol.org:col:34777" name="Museum of Vertebrate Zoology, University of California Berkeley" pageId="69" pageNumber="70" type="Museum">MVZ 164275–76</specimenCode>
|
||
, from
|
||
<collectingMunicipality box="[503,683,1099,1124]" pageId="69" pageNumber="70">Ixtlán de Juárez</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:8A288798FF85E7757EC3FDAAFEE7C7EA:075E6055FF85E77C7CE3FBBDFC4AC255" box="[695,921,1099,1124]" country="Mexico" latitude="17.330557" longLatPrecision="21" longitude="-96.48722" municipality="Ixtlan de Juarez" name="Vivero Rancho Teja" pageId="69" pageNumber="70" stateProvince="Utah">Vivero Rancho Teja</location>
|
||
,
|
||
<geoCoordinate box="[932,1075,1099,1124]" degrees="17" direction="north" minutes="19" orientation="latitude" pageId="69" pageNumber="70" precision="15" seconds="50" value="17.330557">17°19’50’’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[1087,1235,1099,1124]" degrees="96" direction="west" minutes="29" orientation="longitude" pageId="69" pageNumber="70" precision="15" seconds="14" value="-96.48722">96°29’14’’W</geoCoordinate>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="69.[151,1437,837,2021]" pageId="69" pageNumber="70">
|
||
<collectionCode box="[151,200,1136,1161]" pageId="69" pageNumber="70">GK-</collectionCode>
|
||
4502 (
|
||
<collectionCode box="[273,323,1136,1161]" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="69" pageNumber="70">IBH</collectionCode>
|
||
uncatalogued),
|
||
<collectionCode box="[505,567,1137,1160]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34777" lsid="urn:lsid:biocol.org:col:34777" name="Museum of Vertebrate Zoology, University of California Berkeley" pageId="69" pageNumber="70" type="Museum">MVZ</collectionCode>
|
||
144181, 164275–76, and
|
||
<collectionCode box="[864,951,1137,1160]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="69" pageNumber="70" type="University or college">UMMZ</collectionCode>
|
||
99820 are adult females,
|
||
<collectingRegion box="[1248,1303,1136,1161]" country="United States of America" name="Utah" pageId="69" pageNumber="70">UTA</collectingRegion>
|
||
R-52816 is an adult male, and
|
||
<collectingRegion box="[363,418,1173,1198]" country="United States of America" name="Utah" pageId="69" pageNumber="70">UTA</collectingRegion>
|
||
R-8421 is a subadult male.
|
||
</paragraph>
|
||
<paragraph blockId="69.[151,1437,837,2021]" box="[199,1273,1210,1236]" pageId="69" pageNumber="70">
|
||
<emphasis bold="true" box="[199,434,1211,1236]" pageId="69" pageNumber="70">Referred specimens</emphasis>
|
||
<materialsCitation box="[434,948,1210,1236]" collectionCode="USNM" country="Mexico" location="Cuicatlan" pageId="69" pageNumber="70" specimenCode="USNM 47614" specimenCount="1" stateProvince="Oaxaca">
|
||
⎯
|
||
<emphasis bold="true" box="[462,558,1211,1236]" pageId="69" pageNumber="70">
|
||
<collectingCountry box="[462,553,1211,1236]" name="Mexico" pageId="69" pageNumber="70">Mexico</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<collectingRegion box="[565,654,1211,1236]" country="Mexico" name="Oaxaca" pageId="69" pageNumber="70">Oaxaca</collectingRegion>
|
||
:
|
||
<location LSID="urn:lsid:plazi:treatment:8A288798FF85E7757EC3FDAAFEE7C7EA:075E6055FF85E77C7CCCFB4DFCDBC2E5" box="[664,776,1211,1236]" country="Mexico" name="Cuicatlan" pageId="69" pageNumber="70" stateProvince="Oaxaca">Cuicatlán</location>
|
||
:
|
||
<specimenCode box="[786,948,1211,1236]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="69" pageNumber="70" type="Museum">USNM 47614</specimenCode>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation accessionNumber="KU 320768" box="[959,1269,1211,1236]" collectionCode="KU" country="Mexico" location="Ixtepeji" pageId="69" pageNumber="70" specimenCount="1" stateProvince="Oaxaca">
|
||
<quantity box="[959,1002,1211,1236]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="69" pageNumber="70" unit="mi" value="2.0">2mi</quantity>
|
||
E
|
||
<location LSID="urn:lsid:plazi:treatment:8A288798FF85E7757EC3FDAAFEE7C7EA:075E6055FF85E77C7A5DFB4DFBB0C2E5" box="[1033,1123,1211,1236]" country="Mexico" name="Ixtepeji" pageId="69" pageNumber="70" stateProvince="Oaxaca">Ixtepeji</location>
|
||
:
|
||
<accessionNumber box="[1134,1269,1211,1236]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/KU320768" pageId="69" pageNumber="70">
|
||
<collectionCode box="[1134,1174,1212,1235]" country="USA" name="Biodiversity Institute, University of Kansas" pageId="69" pageNumber="70" type="University or college">KU</collectionCode>
|
||
320768
|
||
</accessionNumber>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="69" pageNumber="70" type="diagnosis">
|
||
<paragraph blockId="69.[151,1437,837,2021]" pageId="69" pageNumber="70">
|
||
<emphasis bold="true" box="[199,314,1248,1273]" pageId="69" pageNumber="70">Diagnosis</emphasis>
|
||
. A small species (our Species B of the
|
||
<taxonomicName box="[762,964,1249,1273]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="69" pageNumber="70" phylum="Chordata" rank="species" species="quercorum">
|
||
<emphasis box="[762,964,1249,1273]" italics="true" pageId="69" pageNumber="70">Anolis quercorum</emphasis>
|
||
</taxonomicName>
|
||
complex, see above; SVL in largest male 46.0 mm, largest female 41.0 mm) of the genus
|
||
<taxonomicName authority="(sensu Poe 2004)" baseAuthorityName="sensu Poe" baseAuthorityYear="2004" box="[693,966,1285,1310]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="69" pageNumber="70" phylum="Chordata" rank="genus">
|
||
<emphasis box="[693,765,1286,1310]" italics="true" pageId="69" pageNumber="70">Anolis</emphasis>
|
||
(sensu
|
||
<bibRefCitation author="Poe, S." box="[851,958,1285,1310]" pageId="69" pageNumber="70" pagination="37 - 89" refId="ref95161" refString="Poe, S. (2004) Phylogeny of anoles. Herpetological Monographs, 18, 37 - 89. http: // dx. doi. org / 10.1655 / 0733 - 1347 (2004) 018 [0037: poa] 2.0. co; 2" type="journal article" year="2004">Poe 2004</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
that differs from all Mexican and Central American anoles except
|
||
<taxonomicName box="[426,579,1324,1347]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="69" pageNumber="70" phylum="Chordata" rank="species" species="quercorum">
|
||
<emphasis box="[426,579,1324,1347]" italics="true" pageId="69" pageNumber="70">A. quercorum</emphasis>
|
||
</taxonomicName>
|
||
and a species described below (our Species C of the
|
||
<taxonomicName box="[1171,1323,1324,1347]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="69" pageNumber="70" phylum="Chordata" rank="species" species="quercorum">
|
||
<emphasis box="[1171,1323,1324,1347]" italics="true" pageId="69" pageNumber="70">A. quercorum</emphasis>
|
||
</taxonomicName>
|
||
complex) by having a combination of (1) strongly keeled ventral scales; (2) usually a patch of three greatly enlarged supraocular scales; (3) 10–12 rows of slightly to moderately enlarged dorsal scales that are smaller than ventral scales; (4) short hind legs, longest toe of adpressed hind leg usually reaching to ear opening or to a point between ear opening and eye, ratio shank length/SVL 0.23–0.25; (5) circumnasal usually in contact with first supralabial; and (6) a large pink dewlap in males and a very small red or pink dewlap in females.
|
||
<taxonomicName authorityName="Köhler & Pérez & Petersen & Méndez & Cruz" authorityYear="2014" box="[1093,1266,1510,1534]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="69" pageNumber="70" phylum="Chordata" rank="species" species="carlliebi">
|
||
<emphasis box="[1093,1266,1510,1534]" italics="true" pageId="69" pageNumber="70">Anolis carlliebi</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="69" pageNumber="70" phylum="Chordata" rank="species" species="quercorum">
|
||
<emphasis italics="true" pageId="69" pageNumber="70">A. quercorum</emphasis>
|
||
</taxonomicName>
|
||
and the species described below (our Species C of the
|
||
<taxonomicName box="[883,1035,1548,1571]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="69" pageNumber="70" phylum="Chordata" rank="species" species="quercorum">
|
||
<emphasis box="[883,1035,1548,1571]" italics="true" pageId="69" pageNumber="70">A. quercorum</emphasis>
|
||
</taxonomicName>
|
||
complex) in the ranges and average values of several morphometric and pholidotic characters (see
|
||
<tableCitation box="[861,947,1584,1609]" captionStart="TABLE 9" captionStartId="68.[151,241,152,175]" captionTargetPageId="68" captionText="TABLE 9. Selected measurements, proportions and scale characters of the species related to Anolis quercorum. Range is followed by mean value and standard deviation in parentheses. For abbreviations see text." httpUri="http://table.plazi.org/id/56FE6606FF84E77D7EC3FF6EFB86C6E3" pageId="69" pageNumber="70" tableUuid="56FE6606FF84E77D7EC3FF6EFB86C6E3">Table 9</tableCitation>
|
||
), most obvious in (1) number of middorsal scales in one head length; and (2) ratio shank length/HL. Also,
|
||
<taxonomicName authorityName="Köhler & Pérez & Petersen & Méndez & Cruz" authorityYear="2014" box="[861,1036,1622,1646]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="69" pageNumber="70" phylum="Chordata" rank="species" species="carlliebi">
|
||
<emphasis box="[861,1036,1622,1646]" italics="true" pageId="69" pageNumber="70">Anolis carlliebi</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName box="[1189,1344,1622,1646]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="69" pageNumber="70" phylum="Chordata" rank="species" species="quercorum">
|
||
<emphasis box="[1189,1344,1622,1646]" italics="true" pageId="69" pageNumber="70">A. quercorum</emphasis>
|
||
</taxonomicName>
|
||
and the species described below (our Species C of the
|
||
<taxonomicName box="[713,871,1660,1683]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="69" pageNumber="70" phylum="Chordata" rank="species" species="quercorum">
|
||
<emphasis box="[713,871,1660,1683]" italics="true" pageId="69" pageNumber="70">A. quercorum</emphasis>
|
||
</taxonomicName>
|
||
complex) in hemipenis morphology (hemipenis unilobed in
|
||
<taxonomicName authorityName="Köhler & Pérez & Petersen & Méndez & Cruz" authorityYear="2014" box="[294,426,1698,1722]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="69" pageNumber="70" phylum="Chordata" rank="species" species="carlliebi">
|
||
<emphasis box="[294,426,1698,1722]" italics="true" pageId="69" pageNumber="70">A. carlliebi</emphasis>
|
||
</taxonomicName>
|
||
vs. slightly to distinctly bilobed in the other two species) and by substantial genetic distances (see above).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="75" lastPageNumber="76" pageId="69" pageNumber="70" type="description">
|
||
<paragraph blockId="69.[151,1437,837,2021]" lastBlockId="72.[151,1437,151,627]" lastPageId="72" lastPageNumber="73" pageId="69" pageNumber="70">
|
||
<emphasis bold="true" box="[199,525,1772,1797]" pageId="69" pageNumber="70">
|
||
Description of the
|
||
<typeStatus box="[422,525,1772,1797]" pageId="69" pageNumber="70">holotype</typeStatus>
|
||
</emphasis>
|
||
. Adult female, as indicated by the shape of the base of tail and the absence of a large dewlap; SVL 45.0 mm; tail incomplete; tail moderately compressed in cross section, tail height
|
||
<quantity box="[1301,1388,1810,1835]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="69" pageNumber="70" unit="mm" value="2.2">2.2 mm</quantity>
|
||
and width
|
||
<quantity box="[221,309,1847,1872]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="69" pageNumber="70" unit="mm" value="1.8">1.8 mm</quantity>
|
||
; axilla to groin distance
|
||
<quantity box="[581,683,1847,1872]" metricMagnitude="-2" metricUnit="m" metricValue="2.02" pageId="69" pageNumber="70" unit="mm" value="20.2">20.2 mm</quantity>
|
||
; head length
|
||
<quantity box="[829,930,1847,1872]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="69" pageNumber="70" unit="mm" value="11.1">11.1 mm</quantity>
|
||
, head length/SVL ratio 0.25; snout length
|
||
<quantity metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="69" pageNumber="70" unit="mm" value="4.9">4.9 mm</quantity>
|
||
; head width
|
||
<quantity box="[339,428,1884,1909]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="69" pageNumber="70" unit="mm" value="6.6">6.6 mm</quantity>
|
||
; longest toe of adpressed hind limb reaching to ear opening; shank length
|
||
<quantity box="[1257,1359,1884,1909]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="69" pageNumber="70" unit="mm" value="10.8">10.8 mm</quantity>
|
||
, shank length/head length ratio 0.97; longest finger of extended forelimb reaching to tip of snout; longest finger of adpressed forelimb reaching to a point
|
||
<quantity box="[599,687,1959,1984]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="69" pageNumber="70" unit="mm" value="5.3">5.3 mm</quantity>
|
||
in front of anterior insertion of hind limbs. Dorsal head scales in internasal region keeled, unicarinate; other dorsal head scales rugose or keeled; 6 postrostrals; 6 scales between nasals; 1 elongate prenasal scale on each side, in contact with both rostral and first supralabial; circumnasal separated from supralabial by one scale row; prefrontal depression absent; supraorbital semicircles well developed, narrowly in contact medially; supraorbital disc composed of an oval patch of 3 greatly enlarged, weakly keeled scales; circumorbital row complete on right side, incomplete on left side with on enlarged supraorbital scale in contact with supraorbital semicircles; 3 elongated, strongly overlapping superciliaries, the two anterior ones much larger than posterior one, followed posteriorly by 5 to 6 roundish to squarish scales of moderate size; 1 to 2 rows of small keeled scales extending between enlarged supraorbitals and superciliaries; parietal depression absent; interparietal scale well developed, 1.8 x
|
||
<quantity box="[612,700,1800,1825]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="71" pageNumber="72" unit="mm" value="1.3">1.3 mm</quantity>
|
||
(length x width), surrounded by scales of moderate size; 1 scale present between interparietal and supraorbital semicircles; canthal ridge distinct, composed of 3 large (second largest) and 3 small anterior canthal scales; 5 scales present between second canthals; 8 scales present between posterior canthals; 19 (right)– 20 (left) mostly keeled loreal scales in a maximum of 5 horizontal rows; 4 keeled subocular scales arranged in a single row; 6 (right)– 5 (left) supralabials to level below center of eye; 3 suboculars broadly in contact with 4 supralabials; ear opening 0.5 x 1.0 mm (length x height); mental distinctly wider than long, completely divided medially, bordered posteriorly by 4 postmentals, outer ones much larger than median ones, a single small scale interspersed on each side between postmental, infralabial, and mental; 6 infralabials to level below center of eye; one greatly enlarged sublabial on each side; keeled granular scales present on chin and throat; dewlap very small; dorsum of body with weakly keeled, subimbricate, non-mucronate scales; about 18 medial rows slightly to moderately enlarged; largest dorsal scales about 0.39 x
|
||
<quantity box="[1037,1137,255,280]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="72" pageNumber="73" unit="mm" value="0.34">0.34 mm</quantity>
|
||
(length x width); about 32 medial dorsal scales in one head length; about 62 medial dorsal scales between levels of axilla and groin; lateral scales keeled, granular and more or less homogeneous in size, average size
|
||
<quantity box="[1050,1154,325,350]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="72" pageNumber="73" unit="mm" value="0.2">0.20 mm</quantity>
|
||
in diameter; ventrals at midbody strongly keeled, imbricate, mucronate, about 0.47 x
|
||
<quantity box="[855,956,359,384]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" pageId="72" pageNumber="73" unit="mm" value="0.49">0.49 mm</quantity>
|
||
(length x width); about 22 medial ventral scales in one head length; about 51 medial ventral scales between levels of axilla and groin; 124 scales around midbody; all caudal scales keeled; middorsal caudal scales moderately enlarged, not forming a crest; lateral caudal scales without whorls of enlarged scales, although an indistinct division in segments is discernible; no tube-like axillary pocket present; scales on dorsal surface of forelimb weakly keeled; digital pads dilated, dilated pad about 3 times width of non-dilated distal phalanx; distal phalanx narrower than and raised from dilated pad; 24 (right)–22 (left) lamellae under phalanges II–IV of Toe IV of hind limbs; 7 (right) scales under distal phalanx of Toe IV of hind limbs.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931067" ID-Zenodo-Dep="4931067" httpUri="https://zenodo.org/record/4931067/files/figure.png" pageId="70" pageNumber="71" startId="70.[151,250,1885,1907]" targetBox="[218,1352,315,1864]" targetPageId="70">
|
||
<paragraph blockId="70.[151,1436,1885,1938]" pageId="70" pageNumber="71">
|
||
<emphasis bold="true" box="[151,287,1885,1907]" pageId="70" pageNumber="71">FIGURE 52.</emphasis>
|
||
<taxonomicName authorityName="Köhler & Pérez & Petersen & Méndez & Cruz" authorityYear="2014" box="[292,449,1885,1907]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="70" pageNumber="71" phylum="Chordata" rank="species" species="carlliebi">
|
||
<emphasis box="[292,449,1885,1907]" italics="true" pageId="70" pageNumber="71">Anolis carlliebi</emphasis>
|
||
</taxonomicName>
|
||
in life: (a) two males from Ixtlán de Juárez, Oaxaca; (b) a male from San Juan Tepeuxila, Oaxaca. Photos by Christopher Ehecatl Vega Flores (a) and Luis Canseco Marquéz (b).
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931069" ID-Zenodo-Dep="4931069" httpUri="https://zenodo.org/record/4931069/files/figure.png" pageId="71" pageNumber="72" startId="71.[151,250,1528,1550]" targetBox="[155,1436,199,1505]" targetPageId="71">
|
||
<paragraph blockId="71.[151,1435,1528,1580]" pageId="71" pageNumber="72">
|
||
<emphasis bold="true" box="[151,288,1528,1550]" pageId="71" pageNumber="72">FIGURE 53.</emphasis>
|
||
Holotype of
|
||
<taxonomicName authorityName="Köhler & Pérez & Petersen & Méndez & Cruz" authorityYear="2014" box="[424,581,1528,1550]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="71" pageNumber="72" phylum="Chordata" rank="species" species="carlliebi">
|
||
<emphasis box="[424,581,1528,1550]" italics="true" pageId="71" pageNumber="72">Anolis carlliebi</emphasis>
|
||
</taxonomicName>
|
||
(SMF 96714): (a) dorsal view; (b) lateral view; (c) ventral view; (d) lateral view of head; (e) dorsal view of head; (f) ventral view of head. Scale bars equal 5.0 mm in (a–c) and 1.0 mm in (d–f), respectively.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931071" ID-Zenodo-Dep="4931071" box="[151,1344,2014,2036]" httpUri="https://zenodo.org/record/4931071/files/figure.png" pageId="72" pageNumber="73" startId="72.[151,250,2014,2036]" targetBox="[157,1436,679,1986]" targetPageId="72">
|
||
<paragraph blockId="72.[151,1344,2014,2036]" box="[151,1344,2014,2036]" pageId="72" pageNumber="73">
|
||
<emphasis bold="true" box="[151,288,2014,2036]" pageId="72" pageNumber="73">FIGURE 54.</emphasis>
|
||
<taxonomicName authorityName="Köhler & Pérez & Petersen & Méndez & Cruz" authorityYear="2014" box="[293,450,2014,2036]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="72" pageNumber="73" phylum="Chordata" rank="species" species="carlliebi">
|
||
<emphasis box="[293,450,2014,2036]" italics="true" pageId="72" pageNumber="73">Anolis carlliebi</emphasis>
|
||
</taxonomicName>
|
||
: head scalation in (a,c,e) MVZ 144181; (b,d,f) UTA R-52816. Scale bars equal 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931073" ID-Zenodo-Dep="4931073" httpUri="https://zenodo.org/record/4931073/files/figure.png" pageId="73" pageNumber="74" startId="73.[151,250,1962,1984]" targetBox="[151,1434,193,1942]" targetPageId="73">
|
||
<paragraph blockId="73.[151,1436,1962,2014]" pageId="73" pageNumber="74">
|
||
<emphasis bold="true" box="[151,288,1962,1984]" pageId="73" pageNumber="74">FIGURE 55.</emphasis>
|
||
<taxonomicName authorityName="Köhler & Pérez & Petersen & Méndez & Cruz" authorityYear="2014" box="[294,452,1962,1984]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="73" pageNumber="74" phylum="Chordata" rank="species" species="carlliebi">
|
||
<emphasis box="[294,452,1962,1984]" italics="true" pageId="73" pageNumber="74">Anolis carlliebi</emphasis>
|
||
</taxonomicName>
|
||
: nasal region in (a) UTA R-52818; (b) SMF 96714; superciliary region in (c) UTA R-52818; (d) MVZ 144181; chin region in (e) MVZ 164275; (f) SMF 96714; cloacal region in (g) UTA R-52816. Scale bars equal 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931075" ID-Zenodo-Dep="4931075" httpUri="https://zenodo.org/record/4931075/files/figure.png" pageId="74" pageNumber="75" startId="74.[151,250,1515,1537]" targetBox="[159,1436,187,1493]" targetPageId="74">
|
||
<paragraph blockId="74.[151,1436,1515,1568]" pageId="74" pageNumber="75">
|
||
<emphasis bold="true" box="[151,289,1515,1537]" pageId="74" pageNumber="75">FIGURE 56.</emphasis>
|
||
<taxonomicName authorityName="Köhler & Pérez & Petersen & Méndez & Cruz" authorityYear="2014" box="[295,453,1515,1537]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="74" pageNumber="75" phylum="Chordata" rank="species" species="carlliebi">
|
||
<emphasis box="[295,453,1515,1537]" italics="true" pageId="74" pageNumber="75">Anolis carlliebi</emphasis>
|
||
</taxonomicName>
|
||
: dorsal scales in (a) UTA R-52818; (b) SMF 96714; flank scales in (c) UTA R-52818; (d) SMF 96714; ventral scales in (e) UTA R-52816; (f) SMF 96714. Scale bars equal 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="74.[151,1436,1637,2009]" pageId="74" pageNumber="75">Coloration in life was recorded as follows: Dorsal ground color Drab (19); dorsal surface of head Clay Color (18); with Raw Umber (23) interorbital bar and Cream Color (12) lines from eyes to neck; vertebral region Light Buff (2) with Mars Brown (25) chevrons; dorsal surface of limbs and tail Antique Brown (24); ventral surface of body, limbs and tail Pale Horn Color (11) with Pale Pinkish Buff (3) speckles; dewlap Spectrum Orange (9) with Pale Buff (1) gorgetals; iris Cinnamon-Rufous (31).</paragraph>
|
||
<paragraph blockId="74.[151,1436,1637,2009]" pageId="74" pageNumber="75">Coloration after four months preservation in 70% ethanol was recorded as follows: Dorsal surfaces of head and body Medium Neutral Gray (298) with large confluent Dusky Brown (285) blotches on middorsum and with a Grayish Horn Color (268) postocular stripe, edged with Burnt Umber (48); limb Medium Neutral Gray (298) suffused with Dusky Brown (285); tail Drab (19) with faint Hair Brown (277) vertical bands; ventral surfaces of head, body, limbs, and tail Pale Buff (1) with a suffusion of Medium Neutral Gray (298) on venter; finger and toe pads Vandyke Brown (282).</paragraph>
|
||
<paragraph blockId="75.[151,1437,151,563]" pageId="75" pageNumber="76">
|
||
<emphasis bold="true" box="[199,311,151,176]" pageId="75" pageNumber="76">Variation</emphasis>
|
||
. The
|
||
<typeStatus box="[381,488,152,176]" pageId="75" pageNumber="76">paratypes</typeStatus>
|
||
agree well with the
|
||
<typeStatus box="[728,826,151,176]" pageId="75" pageNumber="76">holotype</typeStatus>
|
||
in general appearance, morphometrics, and scalation (
|
||
<tableCitation box="[158,249,189,214]" captionStart="TABLE 9" captionStartId="68.[151,241,152,175]" captionTargetPageId="68" captionText="TABLE 9. Selected measurements, proportions and scale characters of the species related to Anolis quercorum. Range is followed by mean value and standard deviation in parentheses. For abbreviations see text." httpUri="http://table.plazi.org/id/56FE6606FF84E77D7EC3FF6EFB86C6E3" pageId="75" pageNumber="76" tableUuid="56FE6606FF84E77D7EC3FF6EFB86C6E3">Table 9</tableCitation>
|
||
). They differ in some scalation characters as follows: The number of slightly to moderately enlarged dorsal scale rows varies from 14 to 20. All specimens have a single elongate prenasal. Suboculars in contact with supralabials (2–4 suboculars in contact with 1–4 supralabials) in all specimens examined. All males have a pair of greatly enlarged postcloacal scales.
|
||
</paragraph>
|
||
<paragraph blockId="75.[151,1437,151,563]" pageId="75" pageNumber="76">
|
||
The coloration in life of an adult female
|
||
<typeStatus box="[682,778,339,363]" pageId="75" pageNumber="76">paratype</typeStatus>
|
||
from Ixtlán de Juárez (GK-4502, IBH) was recorded as follows: Dorsal ground color Drab (19) with Drab-Gray (256) vertebral line, Olive Brown (278) dots and a Light Buff (2) lateral line; dorsal surface of head Verona Brown (37) with Brussels Brown (33) interorbital bar; dorsal surface of limbs Cinnamon-Rufous (31) with Yellow Ocher (14) bars; dorsal surface of tail Drab (19) with suffusion of Robin Rufous (29); ventral surface of body, limbs and tail Light Buff (2) with Pale Pinkish Buff (3) suffusions; dewlap Orange Yellow (8) grading into Burnt Orange (19) towards front and Pale Buff (1) gorgetals; iris Natal Brown (49).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.4931077" ID-Zenodo-Dep="4931077" box="[151,1436,1390,1412]" httpUri="https://zenodo.org/record/4931077/files/figure.png" pageId="75" pageNumber="76" startId="75.[151,250,1390,1412]" targetBox="[257,1330,613,1369]" targetPageId="75">
|
||
<paragraph blockId="75.[151,1436,1390,1412]" box="[151,1436,1390,1412]" pageId="75" pageNumber="76">
|
||
<emphasis bold="true" box="[151,287,1390,1412]" pageId="75" pageNumber="76">FIGURE 57.</emphasis>
|
||
Hemipenis of
|
||
<taxonomicName authorityName="Köhler & Pérez & Petersen & Méndez & Cruz" authorityYear="2014" box="[435,592,1390,1412]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="75" pageNumber="76" phylum="Chordata" rank="species" species="carlliebi">
|
||
<emphasis box="[435,592,1390,1412]" italics="true" pageId="75" pageNumber="76">Anolis carlliebi</emphasis>
|
||
</taxonomicName>
|
||
(UTA R-52816): (left) sulcate view; (right) asulcate view. Scale bar equals 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="75.[151,1437,1486,2034]" pageId="75" pageNumber="76">
|
||
The completely everted hemipenis of UTA-R 52816 (
|
||
<figureCitation box="[811,894,1486,1511]" captionStart="FIGURE 57" captionStartId="75.[151,250,1390,1412]" captionTargetBox="[257,1330,613,1369]" captionTargetId="figure-455@75.[257,1330,613,1369]" captionTargetPageId="75" captionText="FIGURE 57. Hemipenis of Anolis carlliebi (UTA R-52816): (left) sulcate view; (right) asulcate view. Scale bar equals 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4931077" httpUri="https://zenodo.org/record/4931077/files/figure.png" pageId="75" pageNumber="76">Fig. 57</figureCitation>
|
||
) is a small unilobate organ; sulcus spermaticus bordered by weakly developed sulcal lips and opening into a single apical field; no asulcate processus present; no surface ornamentation discernable.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="76" lastPageNumber="77" pageId="75" pageNumber="76" type="etymology">
|
||
<paragraph blockId="75.[151,1437,1486,2034]" pageId="75" pageNumber="76">
|
||
<emphasis bold="true" box="[199,327,1598,1623]" pageId="75" pageNumber="76">Etymology</emphasis>
|
||
. The name
|
||
<taxonomicName authorityName="Köhler & Pérez & Petersen & Méndez & Cruz" authorityYear="2014" box="[461,556,1598,1622]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="75" pageNumber="76" phylum="Chordata" rank="species" species="carlliebi">
|
||
<emphasis box="[461,556,1598,1622]" italics="true" pageId="75" pageNumber="76">carlliebi</emphasis>
|
||
</taxonomicName>
|
||
is a patronym honoring Carl S. Lieb who has contributed substantially to our knowledge of Mexican anoles.
|
||
</paragraph>
|
||
<paragraph blockId="75.[151,1437,1486,2034]" lastBlockId="76.[151,1436,151,475]" lastPageId="76" lastPageNumber="77" pageId="75" pageNumber="76">
|
||
<emphasis bold="true" box="[199,467,1673,1698]" pageId="75" pageNumber="76">Natural History Notes</emphasis>
|
||
. The two SMF
|
||
<typeStatus box="[652,700,1674,1698]" pageId="75" pageNumber="76">type</typeStatus>
|
||
specimens of
|
||
<taxonomicName authorityName="Köhler & Pérez & Petersen & Méndez & Cruz" authorityYear="2014" box="[868,1043,1673,1697]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="75" pageNumber="76" phylum="Chordata" rank="species" species="carlliebi">
|
||
<emphasis box="[868,1043,1673,1697]" italics="true" pageId="75" pageNumber="76">Anolis carlliebi</emphasis>
|
||
</taxonomicName>
|
||
were collected during a 4.5 hours night search in a pine oak forest at the campus of the Ixtlán University. The specimens were found sleeping on twigs between 0.3 and 1.0 m above the ground. Most of the following is summarized from the information provided in
|
||
<bibRefCitation author="Ramirez-Bautista, A. & Oliver-Lopez, L. & Mata-Silva, V." box="[287,638,1785,1810]" pageId="75" pageNumber="76" pagination="203 - 204" refId="ref95502" refString="Ramirez-Bautista, A., Oliver-Lopez, L. & Mata-Silva, V. (2002 a) Anolis quercorum (Gray Anole). General Ecology. Herpetological Review, 33, 203 - 204" type="journal article" year="2002">
|
||
Ramirez-Bautista
|
||
<emphasis box="[490,547,1785,1809]" italics="true" pageId="75" pageNumber="76">et al.</emphasis>
|
||
(2002a
|
||
</bibRefCitation>
|
||
,b). The habitat of
|
||
<taxonomicName authorityName="Köhler & Pérez & Petersen & Méndez & Cruz" authorityYear="2014" box="[842,1016,1785,1809]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="75" pageNumber="76" phylum="Chordata" rank="species" species="carlliebi">
|
||
<emphasis box="[842,1016,1785,1809]" italics="true" pageId="75" pageNumber="76">Anolis carlliebi</emphasis>
|
||
</taxonomicName>
|
||
includes xerophilic brush land, thorn forest, and deciduous tropical forest, heavily altered in most of the species’ range by human activities. During daytime, this lizard is mostly found on the leaf litter covered floor, but also on rocks and low vegetation. It is also found in disturbed areas, dump-yards, and along house fences, and is easy to spot at night sleeping on shrub twigs. It feeds on small beetles (
|
||
<taxonomicName authorityName="Linnaeus" authorityYear="1758" box="[458,586,1934,1959]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
|
||
, family
|
||
<taxonomicName authorityName="Latreille" authorityYear="1802" box="[685,848,1934,1959]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="family">Curculionidae</taxonomicName>
|
||
), true bugs (Homoptera, family
|
||
<taxonomicName authorityName="Latreille" authorityYear="1825" box="[1228,1373,1934,1959]" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="family">Cicadellidae</taxonomicName>
|
||
), and
|
||
<taxonomicName box="[151,303,1972,1996]" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
|
||
(families
|
||
<taxonomicName authorityName="Latreille" authorityYear="1802" box="[432,605,1972,1997]" class="Insecta" family="Ichneumonidae" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="family">Ichneumonidae</taxonomicName>
|
||
and
|
||
<taxonomicName box="[674,803,1972,1997]" class="Insecta" family="Braconidae" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="family">Braconidae</taxonomicName>
|
||
) (
|
||
<bibRefCitation author="Canseco-Marquez, L. & Gutierrrez-Mayen, M. G." box="[832,1367,1972,1997]" pageId="75" pageNumber="76" refId="ref88546" refString="Canseco-Marquez, L., & Gutierrrez-Mayen, M. G. (2010) Anfibios y reptiles del Valle de Tehuacan-Cuicatlan. Comision Nacional para el conocimiento y uso de la biodiversidad, Mexico D. F., Mexico, 302 pp." type="book" year="2010">Canseco-Márquez & Gutiérrrez-Mayén 2010</bibRefCitation>
|
||
). The limited available data suggest seasonal reproduction in
|
||
<taxonomicName authority="(Ramirez-Bautista et al. 2002 b)" baseAuthorityName="Ramirez-Bautista" baseAuthorityYear="2002" box="[803,1316,2009,2034]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="75" pageNumber="76" phylum="Chordata" rank="species" species="carlliebi">
|
||
<emphasis box="[803,933,2009,2033]" italics="true" pageId="75" pageNumber="76">A. carlliebi</emphasis>
|
||
(
|
||
<bibRefCitation author="Ramirez-Bautista, A. & Mata-Silva, V. & Oliver-Lopez, L." box="[952,1308,2009,2034]" pageId="75" pageNumber="76" pagination="269 - 271" refId="ref95594" refString="Ramirez-Bautista, A., Mata-Silva, V. & Oliver-Lopez, L. (2002 b) Reproductive data for Anolis quercorum, a little-known anole from Zapotitlan de las Salinas, Puebla, Mexico. Herpetological Review, 33, 269 - 271." type="journal article" year="2002">
|
||
Ramirez-Bautista
|
||
<emphasis box="[1162,1222,2009,2033]" italics="true" pageId="75" pageNumber="76">et al.</emphasis>
|
||
2002b
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. CansecoMárquez & Gutiérrrez-Mayén (2010) noted that
|
||
<specimenCount box="[698,822,151,176]" pageId="76" pageNumber="77" type="female">one female</specimenCount>
|
||
collected in May in Santiago Dominguillo laid an egg and that a male collected in October in San Lucas Teteletitlán had enlarged testicles.
|
||
<bibRefCitation author="Ramirez-Bautista, A. & Oliver-Lopez, L. & Mata-Silva, V." pageId="76" pageNumber="77" pagination="203 - 204" refId="ref95502" refString="Ramirez-Bautista, A., Oliver-Lopez, L. & Mata-Silva, V. (2002 a) Anolis quercorum (Gray Anole). General Ecology. Herpetological Review, 33, 203 - 204" type="journal article" year="2002">
|
||
Ramirez-Bautista
|
||
<emphasis box="[1375,1436,189,213]" italics="true" pageId="76" pageNumber="77">et al.</emphasis>
|
||
(2002a)
|
||
</bibRefCitation>
|
||
reported a male courting a female on a
|
||
<taxonomicName box="[681,780,227,250]" class="Magnoliopsida" family="Fabaceae" genus="Prosopis" kingdom="Animalia" order="Fabales" pageId="76" pageNumber="77" phylum="Tracheophyta" rank="genus">
|
||
<emphasis box="[681,780,227,250]" italics="true" pageId="76" pageNumber="77">Prosopis</emphasis>
|
||
</taxonomicName>
|
||
tree in September.
|
||
<bibRefCitation author="Canseco-Marquez, L. & Gutierrrez-Mayen, M. G." pageId="76" pageNumber="77" refId="ref88546" refString="Canseco-Marquez, L., & Gutierrrez-Mayen, M. G. (2010) Anfibios y reptiles del Valle de Tehuacan-Cuicatlan. Comision Nacional para el conocimiento y uso de la biodiversidad, Mexico D. F., Mexico, 302 pp." type="book" year="2010">Canseco-Márquez & Gutiérrrez-Mayén (2010)</bibRefCitation>
|
||
reported observations of juveniles in November in Santa María Texcatitlán.
|
||
</paragraph>
|
||
<paragraph blockId="76.[151,1436,151,475]" pageId="76" pageNumber="77">
|
||
<emphasis bold="true" box="[199,735,301,326]" pageId="76" pageNumber="77">Geographic Distribution and Conservation</emphasis>
|
||
. As currently known,
|
||
<taxonomicName box="[1016,1198,301,325]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="carlliebi">
|
||
<emphasis box="[1016,1198,301,325]" italics="true" pageId="76" pageNumber="77">Anolis carlliebi</emphasis>
|
||
</taxonomicName>
|
||
is restricted to the Tehuacán-Cuicatlán Valley and the adjacent Ixtlán Valley in the northern central portion of the Mexican State of
|
||
<collectingRegion box="[151,235,375,400]" country="Mexico" name="Oaxaca" pageId="76" pageNumber="77">Oaxaca</collectingRegion>
|
||
at elevations of
|
||
<quantity box="[430,638,375,400]" metricMagnitude="3" metricUnit="m" metricValue="1.5825" metricValueMax="2.5149999999999997" metricValueMin="0.65" pageId="76" pageNumber="77" unit="m" value="1582.5" valueMax="2515.0" valueMin="650.0">650 to 2,515 masl</quantity>
|
||
(
|
||
<figureCitation box="[656,742,375,400]" captionStart="FIGURE 71" captionStartId="90.[151,250,1975,1997]" captionTargetBox="[286,1307,1067,1956]" captionTargetId="figure-229@90.[275,1317,1056,1964]" captionTargetPageId="90" captionText="FIGURE 71. Map indicating collecting localities of species related to Anolis quercorum. Each symbol can represent one or more adjacent localities. Black circles: Anolis carlliebi; white circles: A. quercorum; white squares: A. sacamecatensis." figureDoi="http://doi.org/10.5281/zenodo.4931105" httpUri="https://zenodo.org/record/4931105/files/figure.png" pageId="76" pageNumber="77">Fig. 71</figureCitation>
|
||
). Given its usual abundance wherever this species occurs, it seems justified to classify
|
||
<taxonomicName box="[463,593,413,437]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="76" pageNumber="77" phylum="Chordata" rank="species" species="carlliebi">
|
||
<emphasis box="[463,593,413,437]" italics="true" pageId="76" pageNumber="77">A. carlliebi</emphasis>
|
||
</taxonomicName>
|
||
as Least Concern based on the IUCN Red List Categories and Criteria (
|
||
<bibRefCitation author="IUCN" box="[158,297,450,475]" pageId="76" pageNumber="77" refId="ref91603" refString="IUCN (2012) IUCN Red List of Threatened Species. Version 2012.2. Available from: http: // www. iucnredlist. org (accessed 11 January 2013)" type="url" year="2012">IUCN 2012</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |