treatments-xml/data/8A/1E/B7/8A1EB74F897CFF92C7F36523FEBD908E.xml

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<mods:title id="302ED3D0E39F538DD977B51714694F55">A study of the genus Protosticta Selys, 1855, with descriptions of four new species from Vietnam (Odonata: Platystictidae)</mods:title>
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<mods:namePart id="7939B53177ABB3C36468311BFA6F410F">Phan, Quoc Toan</mods:namePart>
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<mods:namePart id="140FAD14A94F48DEBE7B54FBE8FB5B4C">Kompier, Tom</mods:namePart>
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<subSubSection id="4AAD55D2897CFF94C7F36523FDC99615" box="[151,592,1876,1902]" pageId="3" pageNumber="532" type="nomenclature">
<paragraph id="02080659897CFF94C7F36523FDC99615" blockId="3.[151,592,1876,1935]" box="[151,592,1876,1902]" pageId="3" pageNumber="532">
<heading id="5940B135897CFF94C7F36523FDC99615" bold="true" box="[151,592,1876,1902]" fontSize="11" level="1" pageId="3" pageNumber="532" reason="1">
<emphasis id="30C3DA4B897CFF94C7F36523FDC99615" bold="true" box="[151,592,1876,1902]" pageId="3" pageNumber="532">
3. 
<taxonomicName id="C5B77DDA897CFF94C7D06523FDC99615" ID-CoL="4N78D" authority="Asahina, 1985" authorityName="Asahina" authorityYear="1985" box="[180,592,1876,1902]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="532" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="30C3DA4B897CFF94C7D06523FE019615" bold="true" box="[180,408,1876,1902]" italics="true" pageId="3" pageNumber="532">Protosticta grandis</emphasis>
Asahina, 1985
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<paragraph id="02080659897CFF94C7F36501FEA796F4" blockId="3.[151,592,1876,1935]" box="[151,318,1910,1935]" pageId="3" pageNumber="532">
(
<figureCitation id="9A8C1ADC897CFF94C7FA6501FF6596F4" box="[158,252,1910,1935]" captionStart="FIGURE 3" captionStartId="4.[151,250,1912,1934]" captionTargetBox="[169,1425,470,1883]" captionTargetId="figure@4.[150,1436,465,1891]" captionTargetPageId="4" captionText="FIGURE 3. Protosticta grandis, Cao Bang. [A – E] male— (A) head &amp; thorax in lateral view; anal appendages in (B) dorsal &amp; (C) lateral view; (D) paraprocts in ventral view; penile organ in (E) dorsal view &amp; (F) lateral view; [G – H] female— (G) head &amp; thorax in lateral view; (H) abdomen tip in lateral view." httpUri="https://zenodo.org/record/257726/files/figure.png" pageId="3" pageNumber="532">Figure 3</figureCitation>
A–H)
</paragraph>
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<subSubSection id="4AAD55D2897CFF92C7F365B7FB0691AF" lastPageId="5" lastPageNumber="534" pageId="3" pageNumber="532" type="reference_group">
<paragraph id="02080659897CFF94C7F365B7FE079688" blockId="3.[151,1436,1983,2035]" pageId="3" pageNumber="532">
<taxonomicName id="C5B77DDA897CFF94C7F365B7FD9196AE" ID-CoL="4N78X" authority="Asahina (1984)" authorityName="Asahina" authorityYear="1984" box="[151,520,1983,2005]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="532" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="30C3DA4B897CFF94C7F365B7FEFA96AE" box="[151,355,1983,2005]" italics="true" pageId="3" pageNumber="532">Protosticta robusta:</emphasis>
<bibRefCitation id="66267BA8897CFF94C60F65C8FD9196AE" author="Asahina" box="[363,520,1983,2005]" pageId="3" pageNumber="532" refString="Asahina, S. (1984) A list of the Odonata from Thailand. Part III. Platystictidae. Kontyu, 52, 585 - 595. Asahina, S. (1997) Records of the Northern Vietnamese Odonata taken by the expedition members from the National Science Museum, Tokyo. 6. Platystictidae, Megapodagrionidae, Lestidae and Synlestidae. Bulletin of the National Science Museum, Series A (Zoology), 23, 107 - 113." type="journal article" year="1984">Asahina (1984)</bibRefCitation>
</taxonomicName>
[nec Fraser], pp. 590–591, 595, Figs. 16–20 [preoccupied]. 
<typeStatus id="DD0CB8FB897CFF94C31165B7FB3E96AE" box="[1141,1191,1984,2005]" pageId="3" pageNumber="532">Type</typeStatus>
locality: Khao Soi Dao mountains, 
<collectingCountry id="7AA046C9897CFF94C65965AAFE029688" box="[317,411,2013,2035]" name="Thailand" pageId="3" pageNumber="532">Thailand</collectingCountry>
.
</paragraph>
<paragraph id="02080659897BFF93C7F362EEFA8E9190" blockId="4.[151,1436,152,235]" pageId="4" pageNumber="533">
<taxonomicName id="C5B77DDA897BFF93C7F362EEFD8891D5" ID-CoL="4N78D" authority="Asahina (1985)" authorityName="Asahina" authorityYear="1985" box="[151,529,152,174]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="533" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="30C3DA4B897BFF93C7F362EEFEC491D5" box="[151,349,152,174]" italics="true" pageId="4" pageNumber="533">Protosticta grandis</emphasis>
: Asahina (1985)
</taxonomicName>
, p. 334 (new name for 
<taxonomicName id="C5B77DDA897BFF93C46A62EEFBEB91D5" ID-CoL="4N78X" authority="Asahina, 1984" authorityName="Asahina" authorityYear="1984" box="[782,1138,152,174]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="533" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="30C3DA4B897BFF93C46A62EEFC4D91D5" box="[782,980,152,174]" italics="true" pageId="4" pageNumber="533">Protosticta robusta</emphasis>
<bibRefCitation id="66267BA8897BFF93C4B962EFFBEB91D5" author="Asahina" box="[989,1138,152,174]" pageId="4" pageNumber="533" refString="Asahina, S. (1984) A list of the Odonata from Thailand. Part III. Platystictidae. Kontyu, 52, 585 - 595. Asahina, S. (1997) Records of the Northern Vietnamese Odonata taken by the expedition members from the National Science Museum, Tokyo. 6. Platystictidae, Megapodagrionidae, Lestidae and Synlestidae. Bulletin of the National Science Museum, Series A (Zoology), 23, 107 - 113." type="journal article" year="1984">Asahina, 1984</bibRefCitation>
</taxonomicName>
).— 
<bibRefCitation id="66267BA8897BFF93C3F362EFFE9491B7" author="Hamalainen" pageId="4" pageNumber="533" refString="Hamalainen, M. &amp; Pinratana, A. (1999) Atlas of the dragonflies of Thailand distribution maps by provinces. Brothers of St. Gabriel in Thailand, Bangkok, 176 pp." type="book" year="1999">Hämäläinen &amp; Pinratana (1999)</bibRefCitation>
, p. 60 (Chiang Mai, Kanchanaburi).— 
<bibRefCitation id="66267BA8897BFF93C5F162C1FCB991B7" author="Tsuda" box="[661,800,182,204]" pageId="4" pageNumber="533" refString="Tsuda, S. (2000) A distributional list of world Odonata 2000. Privately Published, Osaka, 430 pp." type="book" year="2000">Tsuda (2000)</bibRefCitation>
, p. 326 (catalogued 
<collectingCountry id="7AA046C9897BFF93C48B62C1FBD591B7" box="[1007,1100,182,204]" name="Thailand" pageId="4" pageNumber="533">Thailand</collectingCountry>
).—van 
<bibRefCitation id="66267BA8897BFF93C3F962C1FA9391B7" author="Tol" box="[1181,1290,182,204]" pageId="4" pageNumber="533" refString="van Tol, J. (2008) Notes on some species of the genus Protosticta from Vietnam (Odonata, Platystictidae). Zoologische Mededelingen Leiden, 82, 217 - 234." type="journal article" year="2008">Tol (2008)</bibRefCitation>
, pp. 219–220, 
<figureCitation id="9A8C1ADC897BFF93C7A362A2FEB89190" box="[199,289,213,235]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,250,1915,1937]" captionStartId-1="3.[151,250,1687,1709]" captionStartId-2="4.[151,250,1912,1934]" captionStartId-3="6.[151,250,1471,1493]" captionTargetBox-0="[157,1432,706,1896]" captionTargetBox-1="[159,1432,476,1674]" captionTargetBox-2="[169,1425,470,1883]" captionTargetBox-3="[156,1434,450,1445]" captionTargetId-0="figure@1.[151,1435,701,1909]" captionTargetId-1="figure@3.[151,1437,469,1675]" captionTargetId-2="figure@4.[150,1436,465,1891]" captionTargetId-3="figure@6.[151,1436,447,1449]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="6" captionText-0="FIGURE 1. Protosticta beaumonti, Mau Son. [A – D] male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; (C) right cercus &amp; paraproct in oblique-dorsal view; (D) penile organ in dorsal view; [E – F] female— (E) female in nature; (F) tip of abdomen (rearranged from photos taken in nature at Mau Son Mountain, Lang Son Province on 07. VI. 2014 by Tom Kompier, specimens were not collected)." captionText-1="FIGURE 2. Protosticta caroli, Ba Na. [A – G] male— (A) head &amp; thorax in lateral view; (B) prothorax in dorsal view; anal appendages in (C) dorsal &amp; (D) lateral view; (E) right cercus and paraproct in oblique-dorsal view; (F) male paraprocts in ventral view; (G) penile organ in dorsal view; [H – J] female— (H) head &amp; thorax in lateral view; (I) prothorax in dorsal view; (J) tip abdomen in lateral view." captionText-2="FIGURE 3. Protosticta grandis, Cao Bang. [A – E] male— (A) head &amp; thorax in lateral view; anal appendages in (B) dorsal &amp; (C) lateral view; (D) paraprocts in ventral view; penile organ in (E) dorsal view &amp; (F) lateral view; [G – H] female— (G) head &amp; thorax in lateral view; (H) abdomen tip in lateral view." captionText-3="FIGURE 4. Protosticta spp. [A – H] Protosticta ngoai spec. nov., holotype male— (A) head &amp; thorax in lateral view; anal appendages in (B) dorsal view, (C) lateral view &amp; (D) oblique-ventral view; (E) paraprocts in ventral view; penile organ in (F) dorsal view &amp; (G) lateral view; (H) wings; [I – J] Protosticta taipokauensis male—anal appendages in (I), lateral view &amp; (J), oblique-ventral view (rearranged from Asahina &amp; Dudgeon, 1987, p. 3, Figs. 6,7)." httpUri-0="https://zenodo.org/record/257724/files/figure.png" httpUri-1="https://zenodo.org/record/257725/files/figure.png" httpUri-2="https://zenodo.org/record/257726/files/figure.png" httpUri-3="https://zenodo.org/record/257727/files/figure.png" pageId="4" pageNumber="533">Figs. 1–4</figureCitation>
.— 
<bibRefCitation id="66267BA8897BFF93C65B62A2FD849190" box="[319,541,213,235]" pageId="4" pageNumber="533" refString="Makbun, N. et al. (2011) Additional records of Odonata from Doi Suthep-Pui National Park, Chiang Mai Province, Thailand. Notulae Odonatologicae, 7 (7), 61 - 68." type="journal article">
Makbun 
<emphasis id="30C3DA4B897BFF93C6FB62A1FE4A9190" box="[415,467,213,235]" italics="true" pageId="4" pageNumber="533">et al.</emphasis>
(2011)
</bibRefCitation>
, p. 63 (Chiang Mai).— 
<bibRefCitation id="66267BA8897BFF93C46162A2FBDD9190" author="Yokoi" box="[773,1092,213,235]" pageId="4" pageNumber="533" refString="Yokoi, N. &amp; Souphanthong, V. (2014) A list of Lao Dragonflies. Kyoei Printing Co. Ltd. Koriyama, 110 pp, 28 pls." type="book" year="2014">Yokoi &amp; Souphanthong (2014)</bibRefCitation>
, p. 25, Pl. 2, Fig. 13.
</paragraph>
<paragraph id="02080659897BFF93C7F36360FEB490E6" blockId="4.[151,1437,279,413]" pageId="4" pageNumber="533">
<emphasis id="30C3DA4B897BFF93C7F36360FE09904B" bold="true" box="[151,400,279,304]" pageId="4" pageNumber="533">Specimens examined</emphasis>
. 
<specimenCount id="14B1CDD0897BFF93C6C5636FFE69904B" box="[417,496,279,304]" pageId="4" pageNumber="533" type="male">1 male</specimenCount>
, 
<specimenCount id="14B1CDD0897BFF93C6996360FDF6904B" box="[509,623,279,304]" pageId="4" pageNumber="533" type="female">5 females</specimenCount>
, Cao Bang city, Cao Bang Province, 
<date id="76092099897BFF93C37E6360FB08904B" box="[1050,1169,279,304]" pageId="4" pageNumber="533" value="2015-05-20">20.V.2015</date>
; 
<specimenCount id="14B1CDD0897BFF93C3FA636FFB74904B" box="[1182,1261,279,304]" pageId="4" pageNumber="533" type="male">1 male</specimenCount>
, 
<specimenCount id="14B1CDD0897BFF93C39F636FFAF5904B" box="[1275,1388,279,304]" pageId="4" pageNumber="533" type="female">2 females</specimenCount>
, Pia Oac Nature Reserve, Cao Bang Province, 
<date id="76092099897BFF93C519634BFD6A902E" box="[637,755,316,341]" pageId="4" pageNumber="533" value="2015-05-18">18.V.2015</date>
; 
<specimenCount id="14B1CDD0897BFF93C465634BFCCB902E" box="[769,850,316,341]" pageId="4" pageNumber="533" type="male">1 male</specimenCount>
, 
<specimenCount id="14B1CDD0897BFF93C404634BFC5F902E" box="[864,966,316,341]" pageId="4" pageNumber="533" type="female">1 female</specimenCount>
, Bhalee, Tay Giang district, Quang Nam Province, 
<date id="76092099897BFF93C66D6328FEE79003" box="[265,382,351,376]" pageId="4" pageNumber="533" value="2015-05-27">27.V.2015</date>
, Quoc Toan Phan leg.; 
<specimenCount id="14B1CDD0897BFF93C5E06317FD719003" box="[644,744,351,376]" pageId="4" pageNumber="533" type="female">1 female</specimenCount>
, Tam Dao National Park, Vinh Phuc Province, 
<date id="76092099897BFF93C39F6328FAE99003" box="[1275,1392,351,376]" pageId="4" pageNumber="533" value="2015-05-12">12.V.2015</date>
, Vu Van Lien leg.
</paragraph>
<caption id="56C856D1897BFF93C7F3650FFD5896B0" httpUri="https://zenodo.org/record/257726/files/figure.png" pageId="4" pageNumber="533" targetBox="[169,1425,470,1883]" targetPageId="4">
<paragraph id="02080659897BFF93C7F3650FFD5896B0" blockId="4.[151,1436,1912,1995]" pageId="4" pageNumber="533">
<emphasis id="30C3DA4B897BFF93C7F3650FFE9496F6" bold="true" box="[151,269,1912,1934]" pageId="4" pageNumber="533">FIGURE 3</emphasis>
. 
<taxonomicName id="C5B77DDA897BFF93C67E650EFE4796F5" ID-CoL="4N78D" box="[282,478,1912,1934]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="533" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="30C3DA4B897BFF93C67E650EFE4796F5" box="[282,478,1912,1934]" italics="true" pageId="4" pageNumber="533">Protosticta grandis</emphasis>
</taxonomicName>
, Cao Bang. [A–E] male—(
<emphasis id="30C3DA4B897BFF93C593650FFC9096F6" bold="true" box="[759,777,1912,1933]" pageId="4" pageNumber="533">A</emphasis>
) head &amp; thorax in lateral view; anal appendages in (
<emphasis id="30C3DA4B897BFF93C247650FFAAD96F6" bold="true" box="[1315,1332,1912,1933]" pageId="4" pageNumber="533">B</emphasis>
) dorsal &amp; (
<emphasis id="30C3DA4B897BFF93C7FB65E0FF2896D6" bold="true" box="[159,177,1943,1965]" pageId="4" pageNumber="533">C</emphasis>
) lateral view; (
<emphasis id="30C3DA4B897BFF93C62865E0FEC796D7" bold="true" box="[332,350,1943,1964]" pageId="4" pageNumber="533">D</emphasis>
) paraprocts in ventral view; penile organ in (
<emphasis id="30C3DA4B897BFF93C44065E0FCAC96D7" bold="true" box="[804,821,1943,1964]" pageId="4" pageNumber="533">E</emphasis>
) dorsal view &amp; (
<emphasis id="30C3DA4B897BFF93C48565E0FC6996D7" bold="true" box="[993,1008,1943,1964]" pageId="4" pageNumber="533">F</emphasis>
) lateral view; [G–H] female—(
<emphasis id="30C3DA4B897BFF93C25465E0FADD96D6" bold="true" box="[1328,1348,1943,1965]" pageId="4" pageNumber="533">G</emphasis>
) head &amp; thorax in lateral view; (
<emphasis id="30C3DA4B897BFF93C6E065C1FE0196B0" bold="true" box="[388,408,1974,1995]" pageId="4" pageNumber="533">H</emphasis>
) abdomen tip in lateral view.
</paragraph>
</caption>
<paragraph id="02080659897AFF92C7A362E0FB0691AF" blockId="5.[151,1437,151,501]" pageId="5" pageNumber="534">
<emphasis id="30C3DA4B897AFF92C7A362E0FE9391CB" bold="true" box="[199,266,151,176]" pageId="5" pageNumber="534">Notes</emphasis>
. This is probably the largest 
<taxonomicName id="C5B77DDA897AFF92C50262EEFD7891CB" ID-CoL="6XZ8" box="[614,737,153,176]" class="Insecta" family="Platystictidae" genus="Protosticta" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="534" phylum="Arthropoda" rank="genus">
<emphasis id="30C3DA4B897AFF92C50262EEFD7891CB" box="[614,737,153,176]" italics="true" pageId="5" pageNumber="534">Protosticta</emphasis>
</taxonomicName>
species known. Both sexes have typically pale yellow anal appendages. It is found in wet forest with dense undergrowth around small covered streams.
</paragraph>
</subSubSection>
<subSubSection id="4AAD55D2897AFF92C7A362A8FDBB9067" pageId="5" pageNumber="534" type="description">
<paragraph id="02080659897AFF92C7A362A8FDBB9067" blockId="5.[151,1437,151,501]" pageId="5" pageNumber="534">
<emphasis id="30C3DA4B897AFF92C7A362A8FEEF9183" bold="true" box="[199,374,223,248]" pageId="5" pageNumber="534">Measurements</emphasis>
. Male. Hindwing 32; abdomen including appendages 51.5–52. Female. Hindwing 29, abdomen including appendages 41.
</paragraph>
</subSubSection>
<subSubSection id="4AAD55D2897AFF92C7A36350FEBD908E" pageId="5" pageNumber="534" type="distribution">
<paragraph id="02080659897AFF92C7A36350FEBD908E" blockId="5.[151,1437,151,501]" pageId="5" pageNumber="534">
<emphasis id="30C3DA4B897AFF92C7A36350FEC0903B" bold="true" box="[199,345,295,320]" pageId="5" pageNumber="534">Distribution</emphasis>
. 
<collectingCountry id="7AA046C9897AFF92C60F6350FE53903B" box="[363,458,295,320]" name="Vietnam" pageId="5" pageNumber="534">Vietnam</collectingCountry>
(Pia Oac Nature Reserve, Cao Bang Province; Tam Dao National Park, Vinh Phuc Province; Xuan Son National Park, Phu Tho Province; Thuong Tien Nature Reserve, Hoa Binh Province; Ba Vi National Park, Ha Noi; Cuc Phuong National Park, Ninh Binh Province; Son Kim, Huong Son district, Ha Tinh Province; Tay Giang district, Quang Nam Province; Chu Yang Sin National Park, Dak Lak Province; Bao Loc, Lam Dong Province), 
<collectingCountry id="7AA046C9897AFF92C6FF63C0FE6690AB" box="[411,511,439,464]" name="Thailand" pageId="5" pageNumber="534">Thailand</collectingCountry>
(Doi Suthep National Park, Chiang Mai Province), Lao PDR (LouangPhabang, Oudomxay).
</paragraph>
</subSubSection>
</treatment>
</document>