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<document id="D33A66C5C2E156BB11272CAC96307CE5" ID-CLB-Dataset="4934" ID-DOI="10.11646/zootaxa.4985.1.2" ID-GBIF-Dataset="bd94b499-c7a6-418e-bb8f-3c5d59675260" ID-ISSN="1175-5326" ID-Zenodo-Dep="4929647" ID-ZooBank="5B23C14A-C7AB-4F22-B0A8-79E700F29FFF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623439671578" checkinUser="plazi" docAuthor="Ramos-Pastrana, Yardany &amp; Rafael, José Albertino" docDate="2021" docId="507C87C0CD601C41EEEA3BD9FBCAF9FB" docLanguage="en" docName="zootaxa.4985.1.2.pdf" docOrigin="Zootaxa 4985 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tomosvaryella mexicanensis Ale-Rocha &amp; Rafael 1995" docType="treatment" docVersion="11" lastPageNumber="49" masterDocId="AC45FFB8CD6C1C4DEE7D390FFFCEFFBC" masterDocTitle="Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species" masterLastPageNumber="68" masterPageNumber="37" pageNumber="49" updateTime="1699012508356" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3B2140ED885ADA1B9677EE2ED225C618">Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species</mods:title>
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<mods:namePart id="879532DA9C357C255112F590D879B0A7">Ramos-Pastrana, Yardany</mods:namePart>
<mods:nameIdentifier id="BB25F8A1C3902A9B32A20E7EA954330C" type="ORCID">0000-0002-3193-6659</mods:nameIdentifier>
<mods:affiliation id="A11328020D77662274BDA27BD0078291">Universidad de la Amazonia, Grupo de Investigación en Entomología Universidad de la Amazonia - GIEU-, Laboratório de Entomologia, Av, 11 5 - 69 Juan XXIII, Florencia, Caquetá, Colombia. ya. ramos @ udla. edu. co; https: // orcid. org / 0000 - 0002 - 3193 - 6659 Instituto Nacional de Pesquisas da Amazônia, INPA, Manaus, Amazonas, Brazil.</mods:affiliation>
<mods:nameIdentifier id="05FDD9F7458AC12A7D8DEF440B4B9D08" type="email">ya.ramos@udla.edu.co</mods:nameIdentifier>
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<mods:namePart id="04C7229461565C0FC8FA150B3F95A30B">Rafael, José Albertino</mods:namePart>
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<mods:title id="6E60C4F65F4503CF9DE61A50FA8787E0">Zootaxa</mods:title>
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<mods:date id="79283C4772E048CEAD0614492C240254">2021</mods:date>
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<mods:number id="38CE08782EE8CFEF3C20E26E2E2D9C14">2021-06-11</mods:number>
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<mods:number id="DB7302915393456B1E242DF3CDC31713">4985</mods:number>
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<treatment id="507C87C0CD601C41EEEA3BD9FBCAF9FB" ID-DOI="http://doi.org/10.5281/zenodo.4929074" ID-GBIF-Taxon="184587569" ID-Zenodo-Dep="4929074" LSID="urn:lsid:plazi:treatment:507C87C0CD601C41EEEA3BD9FBCAF9FB" httpUri="http://treatment.plazi.org/id/507C87C0CD601C41EEEA3BD9FBCAF9FB" lastPageNumber="49" pageId="12" pageNumber="49">
<subSubSection id="90CF655DCD601C41EEEA3BD9FCF2FD4C" box="[151,828,726,753]" pageId="12" pageNumber="49" type="nomenclature">
<paragraph id="D86A36D6CD601C41EEEA3BD9FCF2FD4C" blockId="12.[151,828,726,789]" box="[151,828,726,753]" pageId="12" pageNumber="49">
<heading id="832281BACD601C41EEEA3BD9FCF2FD4C" bold="true" box="[151,828,726,753]" fontSize="11" level="1" pageId="12" pageNumber="49" reason="1">
<taxonomicName id="1FD54D55CD601C41EEEA3BD9FCF2FD4C" ID-CoL="57DZX" authority="Ale-Rocha &amp; Rafael, 1995" authorityName="Ale-Rocha &amp; Rafael" authorityYear="1995" box="[151,828,726,753]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="mexicanensis">
<emphasis id="EAA1EAC4CD601C41EEEA3BD9FCF2FD4C" bold="true" box="[151,828,726,753]" pageId="12" pageNumber="49">
<emphasis id="EAA1EAC4CD601C41EEEA3BD9FE3EFD4D" bold="true" box="[151,496,726,753]" italics="true" pageId="12" pageNumber="49">Tomosvaryella mexicanensis</emphasis>
<bibRefCitation id="BC444B27CD601C41EF8B3BD9FCF2FD4C" author="Ale-Rocha, R. &amp; Rafael, J. A." box="[502,828,726,752]" pageId="12" pageNumber="49" pagination="407 - 427" refId="ref21517" refString="Ale-Rocha, R. &amp; Rafael, J. A. (1995) Tomosvaryella Aczel do Mexico, America Central e Antilhas (Diptera, Pipunculidae). Revista Brasileira de Zoologia, 12, 407 - 427. https: // doi. org / 10.1590 / S 0101 - 81751995000200017" type="journal article" year="1995">Ale-Rocha &amp; Rafael, 1995</bibRefCitation>
</emphasis>
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<subSubSection id="90CF655DCD601C41EEEA3BF4FE82FCA9" box="[151,332,762,789]" pageId="12" pageNumber="49" type="description">
<paragraph id="D86A36D6CD601C41EEEA3BF4FE82FCA9" blockId="12.[151,828,726,789]" box="[151,332,762,789]" pageId="12" pageNumber="49">
<figureCitation id="40EE2A53CD601C41EEEA3BF4FED9FCA9" box="[151,279,762,789]" captionStart="FIGURES 53–62" captionStartId="14.[151,264,1887,1911]" captionTargetBox="[221,1366,181,1863]" captionTargetId="figure-29@14.[221,1366,181,1863]" captionTargetPageId="14" captionText="FIGURES 53–62. Tomosvaryella mexicanensis Ale-Rocha &amp; Rafael, 1995 (IAvH–2019). Male. 53, Habitus, left lateral view; 54, Left hind trochanter, lateral view; 55, Tergites and sternites 6 and 7, ventral view; 56, Tip of abdomen with syntergosternite 8, ventral view; 57, Surstyli, dorsal view; 58, Left surstylus, lateral view; 59, Right surstylus, lateral view; 60, Phallic guide and phallus, left lateral view; 61, Ejaculatory apodeme, lateral view; 62, Hypandrium and gonopods, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4929671" httpUri="https://zenodo.org/record/4929671/files/figure.png" pageId="12" pageNumber="49">Figs 53–62</figureCitation>
,
<figureCitation id="40EE2A53CD601C41EF5F3BF4FE82FCA9" box="[290,332,763,789]" captionStart="FIGURE 132" captionStartId="27.[151,250,1121,1145]" captionTargetBox="[264,1323,417,1097]" captionTargetId="figure-114@27.[264,1323,417,1097]" captionTargetPageId="27" captionText="FIGURE 132. Geographical records of Tomosvaryella species in Colombia. T. tuberculata, T. martae sp. nov., T. galapagensis, T. prostata and T. mexicanensis." figureDoi="http://doi.org/10.5281/zenodo.4930413" httpUri="https://zenodo.org/record/4930413/files/figure.png" pageId="12" pageNumber="49">132</figureCitation>
</paragraph>
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<subSubSection id="90CF655DCD601C41EEEA3A30FDCEFCCA" pageId="12" pageNumber="49" type="reference_group">
<paragraph id="D86A36D6CD601C41EEEA3A30FDCEFCCA" blockId="12.[151,1437,831,886]" pageId="12" pageNumber="49">
<treatmentCitationGroup id="F8C511F8CD601C41EEEA3A30FDCEFCCA" pageId="12" pageNumber="49">
<treatmentCitation id="597410C7CD601C41EEEA3A30FCCDFCEB" author="Ale-Rocha, R. &amp; Rafael, J. A." box="[151,771,831,855]" page="416" pageId="12" pageNumber="49" year="1995">
<taxonomicName id="1FD54D55CD601C41EEEA3A30FCCDFCEB" ID-CoL="57DZX" authority="Ale-Rocha &amp; Rafael, 1995: 416" authorityName="Ale-Rocha &amp; Rafael" authorityPageNumber="416" authorityYear="1995" box="[151,771,831,855]" class="Insecta" family="Pipunculidae" genus="Tomosvaryella" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="49" phylum="Arthropoda" rank="species" species="mexicanensis">
<emphasis id="EAA1EAC4CD601C41EEEA3A30FE77FCEB" box="[151,441,831,855]" italics="true" pageId="12" pageNumber="49">Tomosvaryella mexicanensis</emphasis>
<bibRefCitation id="BC444B27CD601C41EFC33A30FCCDFCEB" author="Ale-Rocha, R. &amp; Rafael, J. A." box="[446,771,831,855]" pageId="12" pageNumber="49" pagination="407 - 427" refId="ref21517" refString="Ale-Rocha, R. &amp; Rafael, J. A. (1995) Tomosvaryella Aczel do Mexico, America Central e Antilhas (Diptera, Pipunculidae). Revista Brasileira de Zoologia, 12, 407 - 427. https: // doi. org / 10.1590 / S 0101 - 81751995000200017" type="journal article" year="1995">Ale-Rocha &amp; Rafael, 1995: 416</bibRefCitation>
</taxonomicName>
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, figs 29–35;
<treatmentCitation id="597410C7CD601C41EDFA3A4FFB9BFCEB" author="De Meyer, M." box="[903,1109,831,856]" page="90" pageId="12" pageNumber="49" year="1996">
<bibRefCitation id="BC444B27CD601C41EDFA3A4FFB9BFCEB" author="De Meyer, M." box="[903,1109,831,856]" pageId="12" pageNumber="49" pagination="1 - 127" refId="ref21998" refString="De Meyer, M. (1996) World catalogue of Pipunculidae (Diptera). Documents de Travail de l' Institut Royal des Sciences Naturelles de Belgique, 86, 1 - 127." type="journal article" year="1996">De Meyer, 1996: 90</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="597410C7CD601C41EA1D3A30FA56FCEB" ID-DOI="http://doi.org/10.5281/zenodo.6269530" ID-Zenodo-Dep="6269530" author="Rafael, J. A. &amp; Ale-Rocha, R." box="[1120,1432,831,855]" httpUri="http://treatment.plazi.org/id/C779878C9A4A2224FECECD53C94EFD02" page="16" pageId="12" pageNumber="49" year="2004">
<bibRefCitation id="BC444B27CD601C41EA1D3A30FA56FCEB" author="Rafael, J. A. &amp; Ale-Rocha, R." box="[1120,1432,831,855]" pageId="12" pageNumber="49" pagination="1 - 18" refId="ref23196" refString="Rafael, J. A. &amp; Ale-Rocha, R. (2004) Nicaraguan Pipunculidae (Diptera): new records and description of new species. Zootaxa, 529 (1), 1 - 18. https: // doi. org / 10.11646 / zootaxa. 529.1.1" type="journal article" year="2004">Rafael &amp; Ale-Rocha, 2004: 16</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="597410C7CD601C41EEBA3A51FE32FCCA" author="Rodriguez, H. C. &amp; Rafael, J. A." box="[199,508,862,886]" page="33" pageId="12" pageNumber="49" year="2012">
<bibRefCitation id="BC444B27CD601C41EEBA3A51FE32FCCA" author="Rodriguez, H. C. &amp; Rafael, J. A." box="[199,508,862,886]" pageId="12" pageNumber="49" refId="ref23353" refString="Rodriguez, H. C. &amp; Rafael, J. A. (2012) Pipunculidae (Diptera) of the Latin America and the Caribbean: A catalog of Species with Notes on Biology and Pipunculid-Host Associations. LAP Lambert Academic Publishing GmbH &amp; Co. KG, Saarbrucken, 48 pp." type="book" year="2012">Rodriguez &amp; Rafael, 2012: 33</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="90CF655DCD601C41EEEA3AABFAFDFB2B" pageId="12" pageNumber="49" type="diagnosis">
<paragraph id="D86A36D6CD601C41EEEA3AABFAFDFB2B" blockId="12.[151,1437,932,1607]" pageId="12" pageNumber="49">
<emphasis id="EAA1EAC4CD601C41EEEA3AABFEDCFC02" bold="true" box="[151,274,932,958]" pageId="12" pageNumber="49">Diagnosis.</emphasis>
Male. Postcranium black, dorsally brownish-pruinose, ventrally grayish-pruinose (
<figureCitation id="40EE2A53CD601C41EAC43AABFAC3FC03" box="[1209,1293,932,959]" captionStart="FIGURES 53–62" captionStartId="14.[151,264,1887,1911]" captionTargetBox="[221,1366,181,1863]" captionTargetId="figure-29@14.[221,1366,181,1863]" captionTargetPageId="14" captionText="FIGURES 53–62. Tomosvaryella mexicanensis Ale-Rocha &amp; Rafael, 1995 (IAvH–2019). Male. 53, Habitus, left lateral view; 54, Left hind trochanter, lateral view; 55, Tergites and sternites 6 and 7, ventral view; 56, Tip of abdomen with syntergosternite 8, ventral view; 57, Surstyli, dorsal view; 58, Left surstylus, lateral view; 59, Right surstylus, lateral view; 60, Phallic guide and phallus, left lateral view; 61, Ejaculatory apodeme, lateral view; 62, Hypandrium and gonopods, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4929671" httpUri="https://zenodo.org/record/4929671/files/figure.png" pageId="12" pageNumber="49">Fig. 53</figureCitation>
). Frontal triangle pale-yellowish pruinose. Hind trochanter with somewhat acute protuberance (
<figureCitation id="40EE2A53CD601C41EA393AC7FB06FC5F" box="[1092,1224,968,995]" captionStart="FIGURES 53–62" captionStartId="14.[151,264,1887,1911]" captionTargetBox="[221,1366,181,1863]" captionTargetId="figure-29@14.[221,1366,181,1863]" captionTargetPageId="14" captionText="FIGURES 53–62. Tomosvaryella mexicanensis Ale-Rocha &amp; Rafael, 1995 (IAvH–2019). Male. 53, Habitus, left lateral view; 54, Left hind trochanter, lateral view; 55, Tergites and sternites 6 and 7, ventral view; 56, Tip of abdomen with syntergosternite 8, ventral view; 57, Surstyli, dorsal view; 58, Left surstylus, lateral view; 59, Right surstylus, lateral view; 60, Phallic guide and phallus, left lateral view; 61, Ejaculatory apodeme, lateral view; 62, Hypandrium and gonopods, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4929671" httpUri="https://zenodo.org/record/4929671/files/figure.png" pageId="12" pageNumber="49">Figs 56–57</figureCitation>
). Tergites and sternites 6 and 7 as in
<figureCitation id="40EE2A53CD601C41EF113AE3FE71FBBB" box="[364,447,1004,1031]" captionStart="FIGURES 53–62" captionStartId="14.[151,264,1887,1911]" captionTargetBox="[221,1366,181,1863]" captionTargetId="figure-29@14.[221,1366,181,1863]" captionTargetPageId="14" captionText="FIGURES 53–62. Tomosvaryella mexicanensis Ale-Rocha &amp; Rafael, 1995 (IAvH–2019). Male. 53, Habitus, left lateral view; 54, Left hind trochanter, lateral view; 55, Tergites and sternites 6 and 7, ventral view; 56, Tip of abdomen with syntergosternite 8, ventral view; 57, Surstyli, dorsal view; 58, Left surstylus, lateral view; 59, Right surstylus, lateral view; 60, Phallic guide and phallus, left lateral view; 61, Ejaculatory apodeme, lateral view; 62, Hypandrium and gonopods, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4929671" httpUri="https://zenodo.org/record/4929671/files/figure.png" pageId="12" pageNumber="49">Fig. 55</figureCitation>
. Surstyli similar in shape, with hooked-tip, right surstylus slightly shorter (
<figureCitation id="40EE2A53CD601C41EB703AE3FA5EFBBB" box="[1293,1424,1004,1031]" captionStart="FIGURES 53–62" captionStartId="14.[151,264,1887,1911]" captionTargetBox="[221,1366,181,1863]" captionTargetId="figure-29@14.[221,1366,181,1863]" captionTargetPageId="14" captionText="FIGURES 53–62. Tomosvaryella mexicanensis Ale-Rocha &amp; Rafael, 1995 (IAvH–2019). Male. 53, Habitus, left lateral view; 54, Left hind trochanter, lateral view; 55, Tergites and sternites 6 and 7, ventral view; 56, Tip of abdomen with syntergosternite 8, ventral view; 57, Surstyli, dorsal view; 58, Left surstylus, lateral view; 59, Right surstylus, lateral view; 60, Phallic guide and phallus, left lateral view; 61, Ejaculatory apodeme, lateral view; 62, Hypandrium and gonopods, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4929671" httpUri="https://zenodo.org/record/4929671/files/figure.png" pageId="12" pageNumber="49">Figs 58–59</figureCitation>
), both surstyli with tips downward-directed when examined in lateral view (
<figureCitation id="40EE2A53CD601C41EDAA3D1FFB94FB97" box="[983,1114,1040,1067]" captionStart="FIGURES 53–62" captionStartId="14.[151,264,1887,1911]" captionTargetBox="[221,1366,181,1863]" captionTargetId="figure-29@14.[221,1366,181,1863]" captionTargetPageId="14" captionText="FIGURES 53–62. Tomosvaryella mexicanensis Ale-Rocha &amp; Rafael, 1995 (IAvH–2019). Male. 53, Habitus, left lateral view; 54, Left hind trochanter, lateral view; 55, Tergites and sternites 6 and 7, ventral view; 56, Tip of abdomen with syntergosternite 8, ventral view; 57, Surstyli, dorsal view; 58, Left surstylus, lateral view; 59, Right surstylus, lateral view; 60, Phallic guide and phallus, left lateral view; 61, Ejaculatory apodeme, lateral view; 62, Hypandrium and gonopods, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4929671" httpUri="https://zenodo.org/record/4929671/files/figure.png" pageId="12" pageNumber="49">Figs 58–59</figureCitation>
). Apex of phallic guide short with stout dorsal spine, angled at about 25 degrees (
<figureCitation id="40EE2A53CD601C41EC993D3BFCF7FBF3" box="[740,825,1076,1103]" captionStart="FIGURES 53–62" captionStartId="14.[151,264,1887,1911]" captionTargetBox="[221,1366,181,1863]" captionTargetId="figure-29@14.[221,1366,181,1863]" captionTargetPageId="14" captionText="FIGURES 53–62. Tomosvaryella mexicanensis Ale-Rocha &amp; Rafael, 1995 (IAvH–2019). Male. 53, Habitus, left lateral view; 54, Left hind trochanter, lateral view; 55, Tergites and sternites 6 and 7, ventral view; 56, Tip of abdomen with syntergosternite 8, ventral view; 57, Surstyli, dorsal view; 58, Left surstylus, lateral view; 59, Right surstylus, lateral view; 60, Phallic guide and phallus, left lateral view; 61, Ejaculatory apodeme, lateral view; 62, Hypandrium and gonopods, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4929671" httpUri="https://zenodo.org/record/4929671/files/figure.png" pageId="12" pageNumber="49">Fig. 60</figureCitation>
). Phallus branches with central branch bearing row of small spines, one of the lateral branches with one small spine (
<figureCitation id="40EE2A53CD601C41ED1F3D57FC76FBCF" box="[866,952,1112,1139]" captionStart="FIGURES 53–62" captionStartId="14.[151,264,1887,1911]" captionTargetBox="[221,1366,181,1863]" captionTargetId="figure-29@14.[221,1366,181,1863]" captionTargetPageId="14" captionText="FIGURES 53–62. Tomosvaryella mexicanensis Ale-Rocha &amp; Rafael, 1995 (IAvH–2019). Male. 53, Habitus, left lateral view; 54, Left hind trochanter, lateral view; 55, Tergites and sternites 6 and 7, ventral view; 56, Tip of abdomen with syntergosternite 8, ventral view; 57, Surstyli, dorsal view; 58, Left surstylus, lateral view; 59, Right surstylus, lateral view; 60, Phallic guide and phallus, left lateral view; 61, Ejaculatory apodeme, lateral view; 62, Hypandrium and gonopods, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4929671" httpUri="https://zenodo.org/record/4929671/files/figure.png" pageId="12" pageNumber="49">Fig. 60</figureCitation>
). Ejaculatory apodeme sandal-shaped with apex rounded (
<figureCitation id="40EE2A53CD601C41EF433D73FE5FFB2B" box="[318,401,1148,1175]" captionStart="FIGURES 53–62" captionStartId="14.[151,264,1887,1911]" captionTargetBox="[221,1366,181,1863]" captionTargetId="figure-29@14.[221,1366,181,1863]" captionTargetPageId="14" captionText="FIGURES 53–62. Tomosvaryella mexicanensis Ale-Rocha &amp; Rafael, 1995 (IAvH–2019). Male. 53, Habitus, left lateral view; 54, Left hind trochanter, lateral view; 55, Tergites and sternites 6 and 7, ventral view; 56, Tip of abdomen with syntergosternite 8, ventral view; 57, Surstyli, dorsal view; 58, Left surstylus, lateral view; 59, Right surstylus, lateral view; 60, Phallic guide and phallus, left lateral view; 61, Ejaculatory apodeme, lateral view; 62, Hypandrium and gonopods, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4929671" httpUri="https://zenodo.org/record/4929671/files/figure.png" pageId="12" pageNumber="49">Fig. 61</figureCitation>
). Gonopods subsymmetrical, apexes acute, right gonopod slightly longer (
<figureCitation id="40EE2A53CD601C41EAAF3D73FAE9FB2B" box="[1234,1319,1148,1175]" captionStart="FIGURES 53–62" captionStartId="14.[151,264,1887,1911]" captionTargetBox="[221,1366,181,1863]" captionTargetId="figure-29@14.[221,1366,181,1863]" captionTargetPageId="14" captionText="FIGURES 53–62. Tomosvaryella mexicanensis Ale-Rocha &amp; Rafael, 1995 (IAvH–2019). Male. 53, Habitus, left lateral view; 54, Left hind trochanter, lateral view; 55, Tergites and sternites 6 and 7, ventral view; 56, Tip of abdomen with syntergosternite 8, ventral view; 57, Surstyli, dorsal view; 58, Left surstylus, lateral view; 59, Right surstylus, lateral view; 60, Phallic guide and phallus, left lateral view; 61, Ejaculatory apodeme, lateral view; 62, Hypandrium and gonopods, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4929671" httpUri="https://zenodo.org/record/4929671/files/figure.png" pageId="12" pageNumber="49">Fig. 62</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="90CF655DCD601C41EEBA3DAFFF3CFAF6" pageId="12" pageNumber="49" type="materials_examined">
<paragraph id="D86A36D6CD601C41EEBA3DAFFF3CFAF6" blockId="12.[151,1437,932,1607]" pageId="12" pageNumber="49">
<emphasis id="EAA1EAC4CD601C41EEBA3DAFFE7FFB06" bold="true" box="[199,433,1184,1210]" pageId="12" pageNumber="49">Material examined.</emphasis>
<materialsCitation id="68BD3C8BCD601C41EFC43DAFFF20FAF6" ID-GBIF-Occurrence="3335985357" collectingDate="2001-07-01" collectingDateMax="2001-12-16" collectingDateMin="2001-07-01" collectionCode="LEUA, INPA" collectorName="Malaise &amp; R. Henriquez" country="Colombia" elevation="10" latitude="11.333333" location="Neguanje" longLatPrecision="1295" longitude="-74.03333" municipality="Tayrona" pageId="12" pageNumber="49" specimenCount="9" specimenCount-male="9" stateProvince="Magdalena">
<collectingCountry id="A0C27646CD601C41EFC43DAFFD9FFB06" box="[441,593,1184,1210]" name="Colombia" pageId="12" pageNumber="49">COLOMBIA</collectingCountry>
,
<emphasis id="EAA1EAC4CD601C41EC203DAFFD2DFB06" bold="true" box="[605,739,1184,1210]" pageId="12" pageNumber="49">
<collectingRegion id="1A11F834CD601C41EC203DAFFD2DFB06" box="[605,739,1184,1210]" country="Colombia" name="Magdalena" pageId="12" pageNumber="49">Magdalena</collectingRegion>
</emphasis>
, PNN[Parque Nacional Natural]
<collectingMunicipality id="380EACACCD601C41EA263DAEFB77FB07" box="[1115,1209,1185,1211]" pageId="12" pageNumber="49">Tayrona</collectingMunicipality>
,
<location id="DD0A600DCD601C41EAB83DAFFAFDFB06" LSID="urn:lsid:plazi:treatment:507C87C0CD601C41EEEA3BD9FBCAF9FB:DD0A600DCD601C41EAB83DAFFAFDFB06" box="[1221,1331,1184,1210]" country="Colombia" latitude="11.333333" longLatPrecision="1295" longitude="-74.03333" municipality="Tayrona" name="Neguanje" pageId="12" pageNumber="49" stateProvince="Magdalena">Neguanje</location>
,
<geoCoordinate id="BDE15011CD601C41EB423DAFFA53FB06" box="[1343,1437,1184,1210]" degrees="11" direction="north" minutes="20" orientation="latitude" pageId="12" pageNumber="49" precision="925" value="11.333333">11º20’N</geoCoordinate>
/
<geoCoordinate id="BDE15011CD601C41EED53DCBFEC3FB62" box="[168,269,1220,1246]" degrees="74" direction="west" minutes="02" orientation="longitude" pageId="12" pageNumber="49" precision="925" value="-74.03333">74º02’W</geoCoordinate>
,
<quantity id="1F2D9B33CD601C41EF663DCBFE96FB62" box="[283,344,1219,1246]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="49" unit="m" value="10.0">
<elevation id="53F8D1E5CD601C41EF663DCBFE96FB62" box="[283,344,1219,1246]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="49" unit="m" value="10.0">10 m</elevation>
</quantity>
[eters],
<collectorName id="75205300CD601C41EFD33DCBFDC7FB62" box="[430,521,1220,1246]" pageId="12" pageNumber="49">Malaise</collectorName>
,
<date id="AC6B1016CD601C41EC6A3DCBFD75FB62" box="[535,699,1220,1246]" pageId="12" pageNumber="49" value="2001-09-17" valueMax="2001-09-27" valueMin="2001-09-17">
<collectingDate id="BC2FE9FECD601C41EC6A3DCBFD75FB62" box="[535,699,1220,1246]" pageId="12" pageNumber="49" value="2001-09-17" valueMax="2001-09-27" valueMin="2001-09-17">17–27.ix.2001</collectingDate>
</date>
,
<collectorName id="75205300CD601C41ECB43DCAFCAFFB62" box="[713,865,1220,1247]" pageId="12" pageNumber="49">R. Henriquez</collectorName>
leg, “M2136” (
<specimenCount id="CED3FD5FCD601C41EA643DCBFBF0FB62" box="[1049,1086,1220,1246]" pageId="12" pageNumber="49" type="male">1♂</specimenCount>
, IAvH); “
<emphasis id="EAA1EAC4CD601C41EACC3DCBFB20FB62" box="[1201,1262,1220,1246]" italics="true" pageId="12" pageNumber="49">idem,</emphasis>
(
<specimenCount id="CED3FD5FCD601C41EB7C3DCBFAE8FB62" box="[1281,1318,1220,1246]" pageId="12" pageNumber="49" type="male">1♂</specimenCount>
,
<collectionCode id="BEC4AE13CD601C41EB493DCBFA48FB62" box="[1332,1414,1220,1246]" pageId="12" pageNumber="49">LEUA</collectionCode>
)”; “
<emphasis id="EAA1EAC4CD601C41EED93DE7FF2FFABE" box="[164,225,1256,1282]" italics="true" pageId="12" pageNumber="49">idem,</emphasis>
(
<specimenCount id="CED3FD5FCD601C41EE8B3DE7FED4FABE" box="[246,282,1256,1282]" pageId="12" pageNumber="49" type="male">1♂</specimenCount>
,
<collectionCode id="BEC4AE13CD601C41EF573DE7FEBEFABE" box="[298,368,1256,1282]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="12" pageNumber="49">INPA</collectionCode>
)”; “
<emphasis id="EAA1EAC4CD601C41EFE33DE7FE15FABE" box="[414,475,1256,1282]" italics="true" pageId="12" pageNumber="49">idem,</emphasis>
<date id="AC6B1016CD601C41EF9A3DE7FD71FABE" box="[487,703,1256,1282]" pageId="12" pageNumber="49" value="2001-07-28" valueMax="2001-08-18" valueMin="2001-07-28">
<collectingDate id="BC2FE9FECD601C41EF9A3DE7FD71FABE" box="[487,703,1256,1282]" pageId="12" pageNumber="49" value="2001-07-28" valueMax="2001-08-18" valueMin="2001-07-28">28.vii–18.viii.2001</collectingDate>
</date>
, “M2019” (
<specimenCount id="CED3FD5FCD601C41ED313DE7FCBCFABE" box="[844,882,1256,1282]" pageId="12" pageNumber="49" type="male">2♂</specimenCount>
, IAvH)” (photographed specimen); “
<emphasis id="EAA1EAC4CD601C41EB5D3DE7FA90FABE" box="[1312,1374,1256,1282]" italics="true" pageId="12" pageNumber="49">idem,</emphasis>
(
<specimenCount id="CED3FD5FCD601C41EB0E3DE7FA59FABE" box="[1395,1431,1256,1282]" pageId="12" pageNumber="49" type="male">1♂</specimenCount>
,
<collectionCode id="BEC4AE13CD601C41EEEA3C03FF27FA9A" box="[151,233,1292,1318]" pageId="12" pageNumber="49">LEUA</collectionCode>
)”; “
<emphasis id="EAA1EAC4CD601C41EF6F3C03FE81FA9A" box="[274,335,1292,1318]" italics="true" pageId="12" pageNumber="49">idem,</emphasis>
(
<specimenCount id="CED3FD5FCD601C41EF223C03FE4DFA9A" box="[351,387,1292,1318]" pageId="12" pageNumber="49" type="male">1♂</specimenCount>
,
<collectionCode id="BEC4AE13CD601C41EFF33C03FE1AFA9A" box="[398,468,1292,1318]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="12" pageNumber="49">INPA</collectionCode>
)”; “
<emphasis id="EAA1EAC4CD601C41EF803C03FDF4FA9A" box="[509,570,1292,1318]" italics="true" pageId="12" pageNumber="49">idem,</emphasis>
<date id="AC6B1016CD601C41EC3D3C03FD22FA9A" box="[576,748,1292,1318]" pageId="12" pageNumber="49" value="2001-12-01" valueMax="2001-12-16" valueMin="2001-12-01">
<collectingDate id="BC2FE9FECD601C41EC3D3C03FD22FA9A" box="[576,748,1292,1318]" pageId="12" pageNumber="49" value="2001-12-01" valueMax="2001-12-16" valueMin="2001-12-01">01–16.xii.2001</collectingDate>
</date>
, “M2754” (
<specimenCount id="CED3FD5FCD601C41ED0C3C03FC5BFA9A" box="[881,917,1292,1318]" pageId="12" pageNumber="49" type="male">1♂</specimenCount>
, IAvH)”; “
<emphasis id="EAA1EAC4CD601C41EA703C03FB84FA9A" box="[1037,1098,1292,1318]" italics="true" pageId="12" pageNumber="49">idem,</emphasis>
<date id="AC6B1016CD601C41EA2C3C03FB20FA9A" box="[1105,1262,1292,1318]" pageId="12" pageNumber="49" value="2001-07-01" valueMax="2001-07-14" valueMin="2001-07-01">
<collectingDate id="BC2FE9FECD601C41EA2C3C03FB20FA9A" box="[1105,1262,1292,1318]" pageId="12" pageNumber="49" value="2001-07-01" valueMax="2001-07-14" valueMin="2001-07-01">1–14.vii.2001</collectingDate>
</date>
, “M1928” (
<specimenCount id="CED3FD5FCD601C41EB0E3C03FA59FA9A" box="[1395,1431,1292,1318]" pageId="12" pageNumber="49" type="male">1♂</specimenCount>
, IAvH)”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="90CF655DCD601C41EEBA3C5BFB82FA2E" pageId="12" pageNumber="49" type="distribution">
<paragraph id="D86A36D6CD601C41EEBA3C5BFB82FA2E" blockId="12.[151,1437,932,1607]" pageId="12" pageNumber="49">
<emphasis id="EAA1EAC4CD601C41EEBA3C5BFDC6FAD2" bold="true" box="[199,520,1364,1390]" pageId="12" pageNumber="49">Geographical distribution.</emphasis>
<collectingCountry id="A0C27646CD601C41EC6F3C5BFDB2FAD2" box="[530,636,1364,1390]" name="Bahamas" pageId="12" pageNumber="49">Bahamas</collectingCountry>
;
<collectingCountry id="A0C27646CD601C41ECF63C5BFD1DFAD2" box="[651,723,1364,1390]" name="Brazil" pageId="12" pageNumber="49">Brazil</collectingCountry>
;
<collectingCountry id="A0C27646CD601C41EC9C3C5BFCACFAD2" box="[737,866,1364,1390]" name="Costa Rica" pageId="12" pageNumber="49">Costa Rica</collectingCountry>
;
<collectingCountry id="A0C27646CD601C41ED0C3C5BFC7EFAD2" box="[881,944,1364,1390]" name="Cuba" pageId="12" pageNumber="49">Cuba</collectingCountry>
;
<collectingCountry id="A0C27646CD601C41EDC33C5BFC34FAD2" box="[958,1018,1364,1390]" name="Haiti" pageId="12" pageNumber="49">Haiti</collectingCountry>
;
<collectingCountry id="A0C27646CD601C41EA753C5BFBA8FAD2" box="[1032,1126,1364,1390]" name="Jamaica" pageId="12" pageNumber="49">Jamaica</collectingCountry>
;
<collectingCountry id="A0C27646CD601C41EA093C5BFB03FAD2" box="[1140,1229,1364,1390]" name="Mexico" pageId="12" pageNumber="49">Mexico</collectingCountry>
;
<collectingCountry id="A0C27646CD601C41EAA13C5BFA9DFAD2" box="[1244,1363,1364,1390]" name="Nicaragua" pageId="12" pageNumber="49">Nicaragua</collectingCountry>
;
<collectingCountry id="A0C27646CD601C41EB1C3C5AFA56FAD3" box="[1377,1432,1365,1391]" name="Peru" pageId="12" pageNumber="49">Peru</collectingCountry>
;
<collectingCountry id="A0C27646CD601C41EEEA3C77FE4EFA2E" box="[151,384,1400,1426]" name="Dominican Republic" pageId="12" pageNumber="49">Dominican Republic</collectingCountry>
(
<bibRefCitation id="BC444B27CD601C41EFF33C77FD78FA2E" author="Rodriguez, H. C. &amp; Rafael, J. A." box="[398,694,1400,1426]" pageId="12" pageNumber="49" refId="ref23353" refString="Rodriguez, H. C. &amp; Rafael, J. A. (2012) Pipunculidae (Diptera) of the Latin America and the Caribbean: A catalog of Species with Notes on Biology and Pipunculid-Host Associations. LAP Lambert Academic Publishing GmbH &amp; Co. KG, Saarbrucken, 48 pp." type="book" year="2012">Rodriguez &amp; Rafael 2012</bibRefCitation>
);
<collectingCountry id="A0C27646CD601C41ECB43C77FCF9FA2E" box="[713,823,1400,1426]" name="Colombia" pageId="12" pageNumber="49">Colombia</collectingCountry>
(new record) (
<figureCitation id="40EE2A53CD601C41EDA03C77FB8EFA2E" box="[989,1088,1400,1426]" captionStart="FIGURE 132" captionStartId="27.[151,250,1121,1145]" captionTargetBox="[264,1323,417,1097]" captionTargetId="figure-114@27.[264,1323,417,1097]" captionTargetPageId="27" captionText="FIGURE 132. Geographical records of Tomosvaryella species in Colombia. T. tuberculata, T. martae sp. nov., T. galapagensis, T. prostata and T. mexicanensis." figureDoi="http://doi.org/10.5281/zenodo.4930413" httpUri="https://zenodo.org/record/4930413/files/figure.png" pageId="12" pageNumber="49">Fig. 132</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="90CF655DCD601C41EEBA3C93FF23FA42" pageId="12" pageNumber="49" type="biology_ecology">
<paragraph id="D86A36D6CD601C41EEBA3C93FF23FA42" blockId="12.[151,1437,932,1607]" pageId="12" pageNumber="49">
<emphasis id="EAA1EAC4CD601C41EEBA3C93FEE4FA0A" bold="true" box="[199,298,1436,1462]" pageId="12" pageNumber="49">Habitat.</emphasis>
The specimens were collected with Malaise traps at ground level at the reserve Parque Nacional Natrual Tayrona. This area has mainly thorny bushes and dry tropical forest typical of the Caribbean region of
<collectingCountry id="A0C27646CD601C41EB0F3CCFFF26FA42" name="Colombia" pageId="12" pageNumber="49">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="90CF655DCD601C41EEBA3F07FBCAF9FB" pageId="12" pageNumber="49" type="discussion">
<paragraph id="D86A36D6CD601C41EEBA3F07FBCAF9FB" blockId="12.[151,1437,932,1607]" pageId="12" pageNumber="49">
<emphasis id="EAA1EAC4CD601C41EEBA3F07FE81F99E" bold="true" box="[199,335,1544,1570]" pageId="12" pageNumber="49">Comments.</emphasis>
The analyzed specimens are larger and have the protuberance of the hind trochanter acuter apically compared to the
<typeStatus id="076E8874CD601C41EF2E3F23FE7BF9FA" box="[339,437,1580,1606]" pageId="12" pageNumber="49" type="holotype">holotype</typeStatus>
specimen illustrated by
<bibRefCitation id="BC444B27CD601C41ECB83F23FBCEF9FB" author="Ale-Rocha, R. &amp; Rafael, J. A." box="[709,1024,1580,1607]" pageId="12" pageNumber="49" pagination="407 - 427" refId="ref21517" refString="Ale-Rocha, R. &amp; Rafael, J. A. (1995) Tomosvaryella Aczel do Mexico, America Central e Antilhas (Diptera, Pipunculidae). Revista Brasileira de Zoologia, 12, 407 - 427. https: // doi. org / 10.1590 / S 0101 - 81751995000200017" type="journal article" year="1995">Ale-Rocha &amp; Rafael (1995)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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