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<document id="6667FB4035A90960EDB517B5B1FE8220" ID-CLB-Dataset="35127" ID-DOI="http://dx.doi.org/10.3897/zookeys.450.7452" ID-GBIF-Dataset="a3fc6ee3-7633-4bf5-9be4-f979ef9810c1" ID-PMC="PMC4233402" ID-Pensoft-Pub="1313-2970-450-1" ID-PubMed="25408617" ID-ZooBank="6EAFB9611C8C4A88BB84CBCE13CDE663" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-450-1" ModsDocOrigin="ZooKeys 450" ModsDocTitle="A taxonomic monograph of Nearctic Scolytus Geoffroy (Coleoptera, Curculionidae, Scolytinae)" checkinTime="1451245189022" checkinUser="pensoft" docAuthor="Smith, Sarah M. &amp; Cognato, Anthony I." docDate="2014" docId="EEDC40CA2E805259B0A8CE03348FEE3A" docLanguage="en" docName="ZooKeys 450: 1-182" docOrigin="ZooKeys 450" docSource="http://dx.doi.org/10.3897/zookeys.450.7452" docTitle="Scolytus oregoni Blackman 1934" docType="treatment" docVersion="5" lastPageNumber="41" masterDocId="FFF8FFEDC228FF81FFA4FFBD8B484705" masterDocTitle="A taxonomic monograph of Nearctic Scolytus Geoffroy (Coleoptera, Curculionidae, Scolytinae)" masterLastPageNumber="182" masterPageNumber="1" pageNumber="39" updateTime="1701374697345" updateUser="ExternalLinkService">
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<mods:title id="683D61C35E7244581D9469FE43EBD3CF">A taxonomic monograph of Nearctic Scolytus Geoffroy (Coleoptera, Curculionidae, Scolytinae)</mods:title>
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<mods:namePart id="1C5B67726B10EA33552CF07EAD47ACF1">Smith, Sarah M.</mods:namePart>
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<mods:namePart id="E73DC7274EEE8385985B82B2C3846D40">Cognato, Anthony I.</mods:namePart>
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<treatment id="EEDC40CA2E805259B0A8CE03348FEE3A" ID-GBIF-Taxon="152056107" LSID="urn:lsid:plazi:treatment:EEDC40CA2E805259B0A8CE03348FEE3A" httpUri="http://treatment.plazi.org/id/EEDC40CA2E805259B0A8CE03348FEE3A" lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39">
<subSubSection id="AC0EB39EA5FF3368D27A5FE339A97CFB" pageId="38" pageNumber="39" type="multiple">
<paragraph id="2EC26AC933A31A844D39B8F750B6898D" pageId="38" pageNumber="39">
<pageBreakToken id="8518D79E56755E02809CB880EFDCDC5D" pageId="38" pageNumber="39" start="start">Taxon</pageBreakToken>
classification Animalia Coleoptera Curculionidae
</paragraph>
</subSubSection>
<subSubSection id="004FC6C3D19FEDF31D5806649C9824B6" pageId="38" pageNumber="39" type="nomenclature">
<paragraph id="3DCD916597F4F94C85614F1D281A4B53" pageId="38" pageNumber="39">
<taxonomicName id="63506DD39AE265DD0DEB2C7C6C738BE7" ID-CoL="4VTMB" authority="Blackman, 1934" authorityName="Blackman" authorityYear="1934" class="Insecta" family="Curculionidae" genus="Scolytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolytus oregoni" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="oregoni">Scolytus oregoni Blackman, 1934</taxonomicName>
Figs 29, 42
</paragraph>
</subSubSection>
<subSubSection id="7159C11E46F6DA35152F2A4A9185E71D" pageId="39" pageNumber="40" type="reference_group">
<paragraph id="42197C55D3A33668288C2FAF9AA61BE0" pageId="39" pageNumber="40">
<taxonomicName id="5517C60963F361D9A1EE2BD8C5748A4D" class="Insecta" family="Curculionidae" genus="Scolytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolytus oregoni" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="oregoni">
<pageBreakToken id="812870E72DDA62C3A2D543CD703F847A" pageId="39" pageNumber="40" start="start">Scolytus</pageBreakToken>
oregoni
</taxonomicName>
Blackman, 1934: 18.
</paragraph>
</subSubSection>
<subSubSection id="46C632C6FB636C1508CAF7FB0BF96F9C" pageId="39" pageNumber="40" type="diagnosis">
<paragraph id="B4C3F864875C8149720445362CA02A9D" pageId="39" pageNumber="40">Diagnosis.</paragraph>
<paragraph id="4E4E1CC304AC031BC94F5E87EEC40178" pageId="39" pageNumber="40">
<taxonomicName id="76D58F4C1B0106E67CD93CB80284CF76" class="Insecta" family="Curculionidae" genus="Scolytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolytus oregoni" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="oregoni">Scolytus oregoni</taxonomicName>
is a rather distinctive species and both sexes are readily distinguished by having the apical margin of ventrite 1 thickened and on the surface of ventrite 2 and by the unarmed ventrite 2.
</paragraph>
</subSubSection>
<subSubSection id="05FC48143F763E09506E8E7B0F2B9AD3" pageId="39" pageNumber="40" type="description (male)">
<paragraph id="559455F97AC75B54FCF537D692EB45A8" pageId="39" pageNumber="40">Description (male).</paragraph>
<paragraph id="ACA91CAD01F12C96747F51ABBC2E296F" pageId="39" pageNumber="40">2.6-3.6 mm long (mean = 3.3 mm; n = 20); 2.0-2.4 times as long as wide. Color dark red-brown, antennae brown. Pronotum typically darker than elytra.</paragraph>
<paragraph id="FDBA389B33A00E3DCBB1011FD5C6CA47" pageId="39" pageNumber="40">Head. Epistoma weakly emarginate; epistomal process weakly developed; median area above mandibles bearing dense patch of long, yellow, hair-like setae. Frons appearing flattened when viewed laterally, slightly transversely impressed just above epistoma; moderately, coarsely, longitudinally aciculate-punctate; aciculations converging at epistoma; punctures sparse, small, coarse; moderately, uniformly covered by long, fine, yellow, erect, hair-like setae, these longer than width of midpoint of eye. Antennal scape short, elongate; club flattened, irregularly ovoid, setose with partial septum, two strongly arcuate sutures visible.</paragraph>
<paragraph id="D8DE8AA6CDF10014C32227FC6160F96F" pageId="39" pageNumber="40">Pronotum wider than long; apical margin broadly rounded, median area between eyes lined with scales; sides distinctly arcuate, strongly constricted near apex, forming a weak transverse impression near apical margin; surface smooth, shining, punctures on disc fine, shallow, moderately abundant, larger and more abundant laterally and on apical constriction; apical and anterolateral margins bearing sparse, erect, yellow, hair-like setae; base weakly bisinuate.</paragraph>
<paragraph id="89497E93BDE37B28531B24DBBED9B789" pageId="39" pageNumber="40">Elytra with sides sub-parallel on apical half, narrowing to subquadrate, smooth apex; apex weakly emarginated at suture. Margin of apical edge bearing large, coarse punctures. Disc glabrous, smooth, shining; interstriae not impressed, more than twice width of striae, punctures uniseriate, smaller than those of striae; striae weakly impressed. Declivity bearing sparse, short, erect, yellow setae. Metepimeron less than half-length of metanepisternum.</paragraph>
<paragraph id="DC81F440FFCCC3BAE3883CAE8F038186" pageId="39" pageNumber="40">Venter. Apical margin of ventrite 1 not elevated above base of ventrite 2. Basal margin of ventrite 2 strongly thickened, lip-like; ventrite 2 nearly perpendicular to ventrite 1; surface shagreened, dull, finely punctate; punctures small, fine, shallow; surface weakly concave, weakly to strongly medially impressed just above base; apical margin unarmed; setae small, less than 1 diameter of a puncture; lateral margins of ventrites 2-3 and ventrite 4 unarmed. Ventrite 5 carinate ridge closer to apical margin of segment; length of ventrite 5 equal to combined lengths of ventrites 3 and 4; setal patch and median depression absent.</paragraph>
<caption id="684E28B1B5B8E44DA51FAA8E96E4D50E" pageId="39" pageNumber="40">
<paragraph id="64563BEED8BDBEC2C0BA92EE2860F991" pageId="39" pageNumber="40">
Figure 42.
<taxonomicName id="6C7630BD2A43E08951E6571455615ADD" class="Insecta" family="Curculionidae" genus="Scolytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolytus oregoni" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="oregoni">Scolytus oregoni</taxonomicName>
A dorsal male habitus B lateral male habitus C male frons D male frons oblique E male venter F male venter oblique G dorsal female habitus H lateral female habitus I female frons J female frons oblique K female venter L female venter oblique.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="5BBC5E49BEA8D7D68E255E07BF7CB227" pageId="39" pageNumber="40" type="female">
<paragraph id="DBE5CBD01D5277AB17DE9F6BF93D93DC" pageId="39" pageNumber="40">Female.</paragraph>
<paragraph id="DB4DFD6D5E3C2FB4C10CFAC537DF86CA" pageId="39" pageNumber="40">2.8-4.0 mm long (mean = 3.23 mm; n = 20); 1.6-2.7 times as long as wide. Similar to male except epistoma entire, epistomal process absent, frons convex when viewed laterally, weakly longitudinally aciculate, setae sparser, shorter, less than width of eye; weakly transversely impressed between inner apices of eyes. Second ventrite unarmed, basal margin weakly thickened.</paragraph>
</subSubSection>
<subSubSection id="FD59AC7FF32BCD35EC77B0227EACFE29" pageId="39" pageNumber="40" type="specimens examined">
<paragraph id="71A1CA4151EA308EE72F920E3F98BC03" pageId="39" pageNumber="40">Specimens examined.</paragraph>
<paragraph id="EFE4622019358E0FB0B9EFAE96068C71" pageId="39" pageNumber="40">96.</paragraph>
</subSubSection>
<subSubSection id="D9B0FAE58290C6D96B718437F8FE876A" pageId="39" pageNumber="40" type="type material">
<paragraph id="56C1D3AF461F06DDEA2068B329DDA627" pageId="39" pageNumber="40">Type material.</paragraph>
<paragraph id="810D99C4556814E80FC7451C646CB91C" pageId="39" pageNumber="40">
Holotype, male, labeled &quot;Ashland Ore., May 20/19, Glendinning, WE Colr,
<taxonomicName id="F58C743B120B01C3F6A03DCC157B0B81" class="Pinopsida" family="Pinaceae" genus="Pseudotsuga" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pseudotsuga taxifolia" order="Pinales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="taxifolia">Pseudotsuga taxifolia</taxonomicName>
, Hopk. US. 13399a, Type No. 43834 USNM&quot; (USNM). Paratypes: UNITED STATES:Oregon: [Jackson Co.]: Ashland, Hopk. U.S. 14246-C, 10.III.[19]16, P.D. Sergent, ex.
<taxonomicName id="F6BED36E39EF56DE25396DDAB1C31BCD" class="Pinopsida" family="Pinaceae" genus="Pseudotsuga" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pseudotsuga taxifolia" order="Pinales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="taxifolia">Pseudotsuga taxifolia</taxonomicName>
(CNCI-2); II.[19]19, W.E Glendinning, Hopk. U.S. 1399-A (USNM-14).
</paragraph>
</subSubSection>
<subSubSection id="8D549148082DE019BBDB9EB4780046AF" pageId="39" pageNumber="40" type="non-type material">
<paragraph id="7A6B55C11B7AADA7AF70F6C4D8BA2E19" pageId="39" pageNumber="40">Non-type material.</paragraph>
<paragraph id="A99758A2947F279601D5D37B599594E0" pageId="39" pageNumber="40">
UNITED STATES:CALIFORNIA: [Del Norte Co.]: [Six Rivers National Forest], Gasquet R.S. [= Ranger Station], Hopk. U.S. 31722-H, 10.VII. R.L. Furniss, ex.
<taxonomicName id="C85E79D213B04C6C78654A270C2B5A29" class="Pinopsida" family="Pinaceae" genus="Pseudotsuga" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pseudotsuga taxifolia" order="Pinales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="taxifolia">Pseudotsuga taxifolia</taxonomicName>
(OSAC-23). Lake Co.: Middletown, 11.XI.1959, G.M. Thomas, ex.
<taxonomicName id="3D4E6867778691D4EEDE98CFC7A64EFF" class="Pinopsida" family="Pinaceae" genus="Pseudotsuga" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pseudotsuga menziesii" order="Pinales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="menziesii">Pseudotsuga menziesii</taxonomicName>
(OSAC-1). Los Angeles Co.: Angeles National Forest, Sawmill Mountain, 34.6926°N, 118.5499°W, V.28-VI.14.2007, Caterino, Chatzimanolis, ex. Lindgren trap (SBMN-4). Marin Co.: Alpine Lake, V.[19]57 (CASC-4). Mount Tamalpais 14.IX.[19]57, E.L. Smith, ex.
<taxonomicName id="3FD3A56764D3E81F5AE5DE0595BD5DB6" class="Pinopsida" family="Pinaceae" genus="Pseudotsuga" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pseudotsuga menziesii" order="Pinales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="menziesii">Pseudotsuga menziesii</taxonomicName>
(OSAC-1). Woodacre Creek, 9.VII.1951, P.S. Bartholomew (CASC-1). Napa Co.: Angwin, 2 mi N.N.E., N. side of Howell Mountain, 1300 ft, 16.VII.1974, H.B. Leech, ex. emerged from log of
<taxonomicName id="3D2E34BF93C395F98379D8D348B3A19A" class="Pinopsida" family="Pinaceae" genus="Pseudotsuga" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pseudotsuga menziesii" order="Pinales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="menziesii">Pseudotsuga menziesii</taxonomicName>
(CASC-1), 21.VII.1974 (CASC-4, USNM-5), 22.VII.1974 (CASC-1), 25.VII.1974 (CASC-4, USNM-1), 7.VIII.1983 (CASC-1), 12.IX.1983 (CASC-2). Callistoga, 5.X.1947, S.L. Wood, ex.
<taxonomicName id="7143DB4A663D3D87ADEE4597C9F8B7B0" class="Pinopsida" family="Pinaceae" genus="Pseudotsuga" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pseudotsuga taxifolia" order="Pinales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="taxifolia">Pseudotsuga taxifolia</taxonomicName>
(USNM-2). Mount Saint Helena, Hopk. U.S. 15401-A, F.B. Herbert, ex.
<taxonomicName id="5CC3BCEB958BE1C39F7CB2E165D835DE" class="Pinopsida" family="Pinaceae" genus="Pseudotsuga" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pseudotsuga taxifolia" order="Pinales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="taxifolia">Pseudotsuga taxifolia</taxonomicName>
(OSAC-1). Riverside Co.: Santa Barbara National Forest, Black Mountain Rd, 33.8395°N, 116.7306°W, 1.VII.2005, M. Caterino (SBMN-1). [Valyermo Co.]: Fenner Canyon, Hopk. U.S. 33853-A, 15.VIII.[19]51, A.D. Moore, ex.
<taxonomicName id="5E9AF4A21EFDC37E7F8031E2BC41F838" class="Pinopsida" family="Pinaceae" genus="Pseudotsuga" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pseudotsuga macrocarpa" order="Pinales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="macrocarpa">Pseudotsuga macrocarpa</taxonomicName>
(EMEC-3, OSAC-8, USNM-4). OREGON:Benton Co.: Marys Peak, Corvallis Watershed, 9.VIII.1963, ex. rotary traps (EMEC-1). Jackson Co.: Ashland, Hopk. U.S. 13363-B, 26.VI.1918, W.E.G, ex.
<taxonomicName id="684F1370C6305965B6F60A49DA10695C" class="Pinopsida" family="Pinaceae" genus="Pseudotsuga" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pseudotsuga taxifolia" order="Pinales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="taxifolia">Pseudotsuga taxifolia</taxonomicName>
, (OSAC-14). Mistletoe, Hopk. U.S. 15753-A, P.D. Sergent, ex.
<taxonomicName id="73B3867693CA64356DE2E958BCBE26A6" class="Pinopsida" family="Pinaceae" genus="Pseudotsuga" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pseudotsuga taxifolia" order="Pinales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="taxifolia">Pseudotsuga taxifolia</taxonomicName>
[=
<taxonomicName id="491971AC99E522B6075F372165424FCF" class="Pinopsida" family="Pinaceae" genus="Pseudotsuga" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pseudotsuga menziesii" order="Pinales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="menziesii">Pseudotsuga menziesii</taxonomicName>
] (OSAC-3). Rogue River National Forest, Rogue River Gorge viewpoint,
<geoCoordinate id="E2C0896D0DE057BBD22E78F60E6FAF99" direction="north" orientation="latitude" precision="1" value="42.909">N42°54.540'</geoCoordinate>
,
<geoCoordinate id="CAE275DBB3F4E3861907DE0E5D0980E4" direction="west" orientation="longitude" precision="1" value="-122.44555">W122°26.733'</geoCoordinate>
, 3489 ft, 21.VIII.2010, S.M. Smith, ex.
<taxonomicName id="0C73B45C962C42B6B098922505D23D4B" class="Pinopsida" family="Pinaceae" genus="Pseudotsuga" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pseudotsuga menziesii" order="Pinales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="menziesii">Pseudotsuga menziesii</taxonomicName>
(MSUC-3). Linn/Lane Co.: Blue River, 11 mi N.E., H.J. Andrews experimental forest, 5.VIII.1986, Log Decomp Study, Site 2, SE1/4 S15 T15S RSE, Trap 2WA (OSAC-1). [Tillamook Co.]: Woods, 18.XI.[19]38 (OSAC-1). Wasco Co.: The Dalles, ODA Port/Mill survey, Trap #65-01a, 14.VIII.1997, ex. Lindgren funnel with
<normalizedToken id="966671C6130063C2BC67ED45F861A780" originalValue="α-pinene">α-pinene</normalizedToken>
&amp; ethanol lure (MSUC-1).
</paragraph>
</subSubSection>
<subSubSection id="B066BBA364BFE976AD3BB052BA9D4FB3" pageId="39" pageNumber="40" type="distribution">
<paragraph id="91BFEAACC854B53FCEE3D8BF99CDE935" pageId="39" pageNumber="40">Distribution.</paragraph>
<paragraph id="47B6452422106320EB8D3EB1FBCC75A8" pageId="39" pageNumber="40">UNITED STATES: California, Oregon, Washington (Fig. 30).</paragraph>
</subSubSection>
<subSubSection id="B67278F1C1588DF9311E689C20235EE2" pageId="39" pageNumber="40" type="hosts">
<paragraph id="A902B0C49EB4FBD82035AE7E07A27573" pageId="39" pageNumber="40">Hosts.</paragraph>
<paragraph id="6448122A8F387DFBD24D7EB514A2FA1E" pageId="39" pageNumber="40">
<taxonomicName id="0D9B186952D826F5561AC62970AE2BDD" class="Pinopsida" family="Pinaceae" genus="Pseudotsuga" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pseudotsuga menziesii" order="Pinales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="menziesii">Pseudotsuga menziesii</taxonomicName>
(Mirb.) Franco (=
<taxonomicName id="00FFCE836693070572324FCF8EC46572" class="Pinopsida" family="Pinaceae" genus="Pseudotsuga" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pseudotsuga taxifolia" order="Pinales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="taxifolia">Pseudotsuga taxifolia</taxonomicName>
Britton) (Douglas fir) and
<taxonomicName id="FDDC60AB48E117F73CA41976DFA67A18" class="Pinopsida" family="Pinaceae" genus="Pseudotsuga" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pseudotsuga macrocarpa" order="Pinales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="macrocarpa">Pseudotsuga macrocarpa</taxonomicName>
(Vasey) Mayr (bigcone Douglas fir).
</paragraph>
</subSubSection>
<subSubSection id="9C33C49AF58C6404741A8EAC242D024D" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="biology">
<paragraph id="4473A078FE4207AA51EDF2C79C259D19" pageId="39" pageNumber="40">Biology.</paragraph>
<paragraph id="1C80366795531E0610ECAE11DB47B6A6" pageId="39" pageNumber="40">
<taxonomicName id="10EFD558B712DC050CE4D569C6F49263" class="Insecta" family="Curculionidae" genus="Scolytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolytus oregoni" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="oregoni">Scolytus oregoni</taxonomicName>
colonizes large limbs and tops of its host and also fresh slash (
<bibRefCitation id="70B15D048BB189922E48E80F56DD02C3" author="Edson, LJ" journalOrPublisher="Unpublished thesis, Humboldt State College, California" pageId="74" pageNumber="75" title="An annotated and illustrated key to the species of the genus Scolytus (Coleoptera, Scolytidae) attacking coniferous trees of the Nearctic region." year="1967">Edson 1967</bibRefCitation>
; Smith, pers. obs.).
</paragraph>
<paragraph id="B57363AD54FFC0DD8D0AA7455A92176C" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
The adult gallery is typically constructed parallel to the grain of the wood and has a central nuptial chamber (Fig. 24). The gallery structure is typically bayonet shaped but may also be longitudinal. Each egg gallery extends in opposite directions to the grain of the wood from the central nuptial chamber. The nuptial chamber is transverse to the egg galleries. The adult gallery deeply scores the sapwood and lightly scores the
<pageBreakToken id="2433B98BBD7DCCF2AF1329AE084350FA" pageId="40" pageNumber="41" start="start">cambium</pageBreakToken>
. The adult gallery averages 6.0-18.0 cm in length. Egg niches are closely spaced and deeply score the sapwood. Larvae extend their mines perpendicular to the egg gallery in a fan shaped pattern before terminating in pupation chambers, which are constructed in the sapwood (
<bibRefCitation id="6A20F23B67539C2085EDC59624E07DA3" author="Edson, LJ" journalOrPublisher="Unpublished thesis, Humboldt State College, California" pageId="74" pageNumber="75" title="An annotated and illustrated key to the species of the genus Scolytus (Coleoptera, Scolytidae) attacking coniferous trees of the Nearctic region." year="1967">Edson 1967</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="AC5D051FEAF64F1467AAED5B0EA8A443" pageId="40" pageNumber="41" type="remarks">
<paragraph id="B2AEB2C9DA8D9289A1476424A9EB97F3" pageId="40" pageNumber="41">Remarks.</paragraph>
<paragraph id="5EDF6846A948000AB85256F23C3AE131" pageId="40" pageNumber="41">
Specimens of
<taxonomicName id="05DBBD305BAF72C990B31702BB1089A8" class="Insecta" family="Curculionidae" genus="Scolytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolytus oregoni" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="oregoni">Scolytus oregoni</taxonomicName>
are very rarely collected and the species is perhaps the least common of the conifer-feeders in the United States. There are many gaps that occur in its distribution range, particularly between northern and southern California. Considerable variation is observed in the male ventrite 1 and 2 across the geographic range especially between northern California, Oregon and Washington and southern California. Southern California populations colonize
<taxonomicName id="1E507E405F6A8ED6D48E35F6B0A84335" class="Pinopsida" family="Pinaceae" genus="Pseudotsuga" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pseudotsuga macrocarpa" order="Pinales" pageId="40" pageNumber="41" phylum="Tracheophyta" rank="species" species="macrocarpa">Pseudotsuga macrocarpa</taxonomicName>
while individuals from the rest of the range colonize
<taxonomicName id="7E44EEC12A9CCA3D96211FBE633A6A8A" class="Pinopsida" family="Pinaceae" genus="Pseudotsuga" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pseudotsuga menziesii" order="Pinales" pageId="40" pageNumber="41" phylum="Tracheophyta" rank="species" species="menziesii">Pseudotsuga menziesii</taxonomicName>
. In addition,
<taxonomicName id="B2E35EB204C22A1159DC2B678DB91F46" class="Insecta" family="Curculionidae" genus="Scolytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scolytus oregoni" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="oregoni">Scolytus oregoni</taxonomicName>
males from California and particularly southern California have a greater impression of the second ventrite and thicker margin between ventrite 1 and 2 to specimens from Oregon and Washington.
</paragraph>
</subSubSection>
</treatment>
</document>