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<document id="9D62ECF7AF6620D9E1DDA8E39EE96393" ID-DOI="10.11646/zootaxa.3619.2.5" ID-GBIF-Dataset="549f99ee-2153-4d61-a953-8def30c8159d" ID-ISSN="1175-5326" ID-Zenodo-Dep="216580" ID-ZooBank="DBB8DB78-09DF-4F68-AFB1-AAC8EBC95DD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460577749427" checkinUser="plazi" docAuthor="Ren, Li, Alonso-Zarazaga, Miguel A. &amp; Zhang, Runzhi" docDate="2013" docId="C62C2621100FC04AFF10FDD2FE316CB1" docLanguage="en" docName="zt03619p182.pdf" docOrigin="Zootaxa 3619 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Geotragus tuberculatus Chen 1990" docType="treatment" docVersion="8" lastPageNumber="167" masterDocId="3A155E59100CC04CFF87FFA8FF95686B" masterDocTitle="Revision of the Chinese Geotragus Schoenherr with description of three new species (Coleoptera: Curculionidae: Entiminae)" masterLastPageNumber="182" masterPageNumber="161" pageNumber="164" updateTime="1698323549919" updateUser="plazi">
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<mods:title id="70F48A8853714A6A7F924A9C85A1BF00">Revision of the Chinese Geotragus Schoenherr with description of three new species (Coleoptera: Curculionidae: Entiminae)</mods:title>
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<mods:namePart id="C5148D5E1AF4E7FD00C23ADAF8C07F12">Ren, Li</mods:namePart>
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<treatment id="C62C2621100FC04AFF10FDD2FE316CB1" ID-DOI="http://doi.org/10.5281/zenodo.6158915" ID-GBIF-Taxon="119560656" ID-Zenodo-Dep="6158915" LSID="urn:lsid:plazi:treatment:C62C2621100FC04AFF10FDD2FE316CB1" httpUri="http://treatment.plazi.org/id/C62C2621100FC04AFF10FDD2FE316CB1" lastPageId="6" lastPageNumber="167" pageId="3" pageNumber="164">
<subSubSection id="069FC4BC100FC04FFF10FDD2FE926ADD" pageId="3" pageNumber="164" type="nomenclature">
<paragraph id="4E3A9737100FC04FFF10FDD2FDA66AF8" blockId="3.[151,563,634,694]" box="[151,563,634,659]" pageId="3" pageNumber="164">
<heading id="1572205B100FC04FFF10FDD2FDA66AF8" bold="true" box="[151,563,634,659]" fontSize="10" level="2" pageId="3" pageNumber="164" reason="2">
<taxonomicName id="8985ECB4100FC04FFF10FDD2FDA66AF8" ID-CoL="3FT5Z" authority="Chen, 1990" authorityName="Chen" authorityYear="1990" box="[151,563,634,659]" class="Insecta" family="Curculionidae" genus="Geotragus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="164" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis id="7CF14B25100FC04FFF10FDD2FDA66AF8" bold="true" box="[151,563,634,659]" pageId="3" pageNumber="164">
<emphasis id="7CF14B25100FC04FFF10FDD2FE326AF8" bold="true" box="[151,423,634,659]" italics="true" pageId="3" pageNumber="164">Geotragus tuberculatus</emphasis>
<bibRefCitation id="2A14EAC6100FC04FFE2AFDD2FDA66AF8" author="Chen" box="[429,563,634,659]" pageId="3" pageNumber="164" refString="Chen, Y. Q. (1990) A new species of Geotragus from the Hengduan Mountains, Yunnan, China (Coleoptera: Curculionidae). Sinozoologia, 7, 149 - 150." type="journal article" year="1990">Chen, 1990</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="4E3A9737100FC04FFF10FD36FE926ADD" blockId="3.[151,563,634,694]" box="[151,263,670,694]" pageId="3" pageNumber="164">
(
<figureCitation id="D6BE8BB2100FC04FFF18FD36FF776ADD" box="[159,226,670,694]" captionStart="FIGURE 1" captionStartId="4.[151,250,841,864]" captionTargetBox="[151,1436,179,825]" captionTargetId="figure@4.[151,1436,175,825]" captionTargetPageId="4" captionText="FIGURE 1. Habitus of Geotragus tuberculatus Chen, male, paratype: A dorsal view B lateral view C head, dorsal view. Scales: 1 mm." httpUri="https://zenodo.org/record/216581/files/figure.png" pageId="3" pageNumber="164">Fig. 1</figureCitation>
,
<figureCitation id="D6BE8BB2100FC04FFF68FD36FF6B6ADD" box="[239,254,670,694]" captionStart="FIGURE 2" captionStartId="5.[151,250,1913,1936]" captionTargetBox="[263,1327,206,1864]" captionTargetId="figure@5.[237,1343,193,1884]" captionTargetPageId="5" captionText="FIGURE 2. Geotragus tuberculatus Chen: A male ventrites, ventral view B male metendosternite, dorsal view C male sternites VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view (g = groove) H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, lateral view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216582/files/figure.png" pageId="3" pageNumber="164">2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="069FC4BC100FC04FFF10FD4FFCC16A97" box="[151,852,742,765]" pageId="3" pageNumber="164" type="reference_group">
<paragraph id="4E3A9737100FC04FFF10FD4FFCC16A97" blockId="3.[151,852,742,765]" box="[151,852,742,765]" pageId="3" pageNumber="164">
<taxonomicName id="8985ECB4100FC04FFF10FD4FFD916A97" ID-CoL="3FT5Z" authority="Chen, 1990" authorityName="Chen" authorityYear="1990" box="[151,516,742,765]" class="Insecta" family="Curculionidae" genus="Geotragus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="164" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis id="7CF14B25100FC04FFF10FD4FFE136A96" box="[151,390,743,765]" italics="true" pageId="3" pageNumber="164">Geotragus tuberculatus</emphasis>
<bibRefCitation id="2A14EAC6100FC04FFE0BFD4EFD916A97" author="Chen" box="[396,516,742,764]" pageId="3" pageNumber="164" refString="Chen, Y. Q. (1990) A new species of Geotragus from the Hengduan Mountains, Yunnan, China (Coleoptera: Curculionidae). Sinozoologia, 7, 149 - 150." type="journal article" year="1990">Chen, 1990</bibRefCitation>
</taxonomicName>
. Sinozoologia, 7: 149150, fig.1.
</paragraph>
</subSubSection>
<subSubSection id="069FC4BC100FC04FFF10FC81FC866B0E" pageId="3" pageNumber="164" type="diagnosis">
<paragraph id="4E3A9737100FC04FFF10FC81FC866B0E" blockId="3.[151,1436,809,2021]" pageId="3" pageNumber="164">
<emphasis id="7CF14B25100FC04FFF10FC81FE846B29" bold="true" box="[151,273,809,834]" pageId="3" pageNumber="164">Diagnosis.</emphasis>
Elytra with interstria 3 with a row of three prominent tubercles before declivity; interstria 6 raised and forming a row of 34 rounded tubercles before declivity.
</paragraph>
</subSubSection>
<subSubSection id="069FC4BC100FC04AFF4EFCD9FE116ACC" lastPageId="6" lastPageNumber="167" pageId="3" pageNumber="164" type="description">
<paragraph id="4E3A9737100FC04FFF4EFCD9FB0B6BC6" blockId="3.[151,1436,809,2021]" pageId="3" pageNumber="164">
<emphasis id="7CF14B25100FC04FFF4EFCD9FDA66BE1" bold="true" box="[201,563,881,906]" pageId="3" pageNumber="164">Redescription. Holotype-male.</emphasis>
<emphasis id="7CF14B25100FC04FFDBDFCDAFD496BE2" box="[570,732,882,905]" italics="true" pageId="3" pageNumber="164">Measurements</emphasis>
(in mm): Standard length: 5.80; pronotal length: 1.70; pronotal width: 1.75; elytral length: 4.15; elytral width: 2.50; rostral length: 1.05; rostral width: 0.70.
</paragraph>
<paragraph id="4E3A9737100FC04FFF4EFC11FC4E6D52" blockId="3.[151,1436,809,2021]" pageId="3" pageNumber="164">
<emphasis id="7CF14B25100FC04FFF4EFC11FE096BBA" box="[201,412,953,977]" italics="true" pageId="3" pageNumber="164">Habitus and color</emphasis>
(
<figureCitation id="D6BE8BB2100FC04FFE29FC11FE646BBA" box="[430,497,953,977]" captionStart="FIGURE 1" captionStartId="4.[151,250,841,864]" captionTargetBox="[151,1436,179,825]" captionTargetId="figure@4.[151,1436,175,825]" captionTargetPageId="4" captionText="FIGURE 1. Habitus of Geotragus tuberculatus Chen, male, paratype: A dorsal view B lateral view C head, dorsal view. Scales: 1 mm." httpUri="https://zenodo.org/record/216581/files/figure.png" pageId="3" pageNumber="164">Fig. 1</figureCitation>
A, 1B): Body slender; color dark brown, antennae and legs reddish brown; with coppery metallic pearl white, pale brown to dark brown scales, scales moderately dense, tessellate, contiguous but not overlapping; scales on dorsal and lateral surfaces of rostrum moderately dense, elongate oval; scales behind epistome round and small, clearly different from others; posterior half of antennal scrobes glabrous; pronotum with polygonal scales, moderately dense, contiguous, some scales imbricate, with a longitudinal stripe of lighter scales on each side of dorsum; scales on elytra polygonal, dense, but not contiguous; scales on ventrites moderately dense, round to elongate-oval; scales on legs dense; with short, curved, recumbent to subrecumbent setae, sparsely distributed; rostrum covered with recumbent thick setae; setae of antennal scape and desmomeres 17 long, fine and sparse; disc and lateral surface of pronotum with sparse, recumbent setae; setae of interstriae lanceolate, curved, sparse and subrecumbent; setae on ventral surface moderately thick, long, recumbent and dense; setae of tibiae long and moderately dense, femora with recumbent lanceolate setae.
</paragraph>
<paragraph id="4E3A9737100FC04FFF4EFAEDFC626DCE" blockId="3.[151,1436,809,2021]" pageId="3" pageNumber="164">
<emphasis id="7CF14B25100FC04FFF4EFAEDFE936D36" box="[201,262,1349,1373]" italics="true" pageId="3" pageNumber="164">Head</emphasis>
(
<figureCitation id="D6BE8BB2100FC04FFE92FAEDFEC16D36" box="[277,340,1349,1373]" captionStart="FIGURE 1" captionStartId="4.[151,250,841,864]" captionTargetBox="[151,1436,179,825]" captionTargetId="figure@4.[151,1436,175,825]" captionTargetPageId="4" captionText="FIGURE 1. Habitus of Geotragus tuberculatus Chen, male, paratype: A dorsal view B lateral view C head, dorsal view. Scales: 1 mm." httpUri="https://zenodo.org/record/216581/files/figure.png" pageId="3" pageNumber="164">Fig. 1</figureCitation>
C): Moderately convex, dorsal surface corrugated, with many longitudinal fine and shallow sulci, and small and dense punctures; eyes strongly convex, with a deep and fine circumocular sulcus; frons slightly depressed, not elevated regarding to dorsal surface of rostrum in lateral view.
</paragraph>
<paragraph id="4E3A9737100FC04FFF4EFA1AFB266E16" blockId="3.[151,1436,809,2021]" pageId="3" pageNumber="164">
<emphasis id="7CF14B25100FC04FFF4EFA1AFEBD6DA2" box="[201,296,1458,1481]" italics="true" pageId="3" pageNumber="164">Rostrum</emphasis>
(
<figureCitation id="D6BE8BB2100FC04FFEB0FA19FEE26DA2" box="[311,375,1457,1481]" captionStart="FIGURE 1" captionStartId="4.[151,250,841,864]" captionTargetBox="[151,1436,179,825]" captionTargetId="figure@4.[151,1436,175,825]" captionTargetPageId="4" captionText="FIGURE 1. Habitus of Geotragus tuberculatus Chen, male, paratype: A dorsal view B lateral view C head, dorsal view. Scales: 1 mm." httpUri="https://zenodo.org/record/216581/files/figure.png" pageId="3" pageNumber="164">Fig. 1</figureCitation>
C): In dorsal view, 1.50× length of wide, apex narrower than base; base slightly narrower than frons; dorsal surface flat in median area of rostrum, punctured, with a narrow and deep median sulcus, extending longitudinally from antennal insertion to the vertex; posterior angle of epistome between 45° and 90°, with carina distinct and blunt; mandible scars round to rectangular; in dorsal view ventral margin of scrobes visible from apex to middle of rostrum; in lateral view, with a shallow triangular depression positioned laterally between eyes and antennal scrobes, close to ventral margin of rostrum, accompanied by subparallel deep sulcus.
</paragraph>
<paragraph id="4E3A9737100FC04FFF4EF921FBE66F5A" blockId="3.[151,1436,809,2021]" pageId="3" pageNumber="164">
<emphasis id="7CF14B25100FC04FFF4EF921FEA66ECA" box="[201,307,1673,1697]" italics="true" pageId="3" pageNumber="164">Antennae</emphasis>
: Scape slender, reaching middle of eyes at rest, 0.93× length of funicle; desmomere 1 1.32× length of 2, both segments elongate clavate; desmomere 3 short, clavate, 0.74× length of 2; desmomeres 46 moniliform, 4 0.93× length of 3, 5 0.93× length of 4; desmomeres 5 and 6 equal in length; desmomere 7 1.33× length of and wider than 6, elongate moniliform; pubescence of funicle similar throughout; club segment 1 1.13× length of 2, segment 2 0.73× length of segment 3; segment 3 with a marked
<emphasis id="7CF14B25100FC04FFCE2F8B1FC2B6F5A" box="[869,958,1817,1841]" italics="true" pageId="3" pageNumber="164">annulus</emphasis>
beyond middle.
</paragraph>
<paragraph id="4E3A9737100FC04FFF4EF896FBEA6FAA" blockId="3.[151,1436,809,2021]" pageId="3" pageNumber="164">
<emphasis id="7CF14B25100FC04FFF4EF896FEAD6F3E" box="[201,312,1854,1877]" italics="true" pageId="3" pageNumber="164">Pronotum</emphasis>
: 0.97× as long as wide, subquadrate; anterior margin truncate, posterior margin mesally slightly produced; disc with a fine, shallow, incomplete median longitudinal groove, slightly flattened on both sides of median groove; posterior margin not carinate; dorsal surface of pronotum with dense, large, and round granules, each granule with one recumbent lanceolate seta on top; postocular vibrissae blond, long.
</paragraph>
<paragraph id="4E3A9737100FC04FFF4EF865FCD86F8E" blockId="3.[151,1436,809,2021]" box="[201,845,1997,2021]" pageId="3" pageNumber="164">
<taxonomicName id="8985ECB4100FC04FFF4EF865FE1F6F8E" authority="Small" authorityName="Small" box="[201,394,1997,2021]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="3" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="7CF14B25100FC04FFF4EF865FEA26F8E" box="[201,311,1997,2021]" italics="true" pageId="3" pageNumber="164">Scutellum</emphasis>
: Small
</taxonomicName>
, reddish brown, apex broadly V-shaped.
</paragraph>
<caption id="1AFAC7BF1008C048FF10FCE1FEB36B14" httpUri="https://zenodo.org/record/216581/files/figure.png" pageId="4" pageNumber="165" targetBox="[151,1436,179,825]" targetPageId="4">
<paragraph id="4E3A97371008C048FF10FCE1FEB36B14" blockId="4.[151,1437,841,895]" pageId="4" pageNumber="165">
<emphasis id="7CF14B251008C048FF10FCE1FE836B34" bold="true" box="[151,278,841,864]" pageId="4" pageNumber="165">FIGURE 1.</emphasis>
Habitus of
<taxonomicName id="8985ECB41008C048FE1FFCE3FD596B0B" authority="Chen" authorityName="Chen" box="[408,716,842,864]" class="Insecta" family="Curculionidae" genus="Geotragus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="165" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis id="7CF14B251008C048FE1FFCE3FD1E6B0B" box="[408,651,842,864]" italics="true" pageId="4" pageNumber="165">Geotragus tuberculatus</emphasis>
Chen
</taxonomicName>
, male, paratype:
<emphasis id="7CF14B251008C048FC03FCE2FC036B34" bold="true" box="[900,918,842,863]" pageId="4" pageNumber="165">A</emphasis>
dorsal view
<emphasis id="7CF14B251008C048FBA4FCE2FBA16B34" bold="true" box="[1059,1076,842,863]" pageId="4" pageNumber="165">B</emphasis>
lateral view
<emphasis id="7CF14B251008C048FB45FCE1FB416B0B" bold="true" box="[1218,1236,841,864]" pageId="4" pageNumber="165">C</emphasis>
head, dorsal view. Scales: 1 mm.
</paragraph>
</caption>
<paragraph id="4E3A97371008C048FF4EFC0EFD3A6D69" blockId="4.[151,1436,934,2029]" pageId="4" pageNumber="165">
<emphasis id="7CF14B251008C048FF4EFC0EFE9A6BD5" box="[201,271,934,958]" italics="true" pageId="4" pageNumber="165">Elytra</emphasis>
: Base slightly emarginated; in lateral view moderately convex; in dorsal view, lateral margins clearly diverging from base to basal 1/4, then subparallel and slightly curved, reaching its greatest width at mid length, thence clearly converging towards the apex, apex widely rounded; interstriae tuberculate; interstria 1 moderately raised along basal 1/8, forming a longitudinal crest, raised at declivity forming a prominent longitudinal crest, apically rounded in lateral view; interstriae 3 distinctly raised and forming three consecutive conical tubercles before declivity; interstria 4 raised and forming a row of 23 small round tubercles before declivity; interstria 5 raised behind basal 1/8 and forming a row of 23 elongate tubercles before declivity, and a strongly protruding conical tubercle at declivity; interstria 6 raised and forming a row of 34 rounded tubercles before declivity; interstria 7 raised and forming 23 small elongate tubercles before declivity; interstriae 3 and 9 apically coalescent, raised at declivity forming a triangular tubercle.
</paragraph>
<paragraph id="4E3A97371008C048FF4EFAA6FB436D05" blockId="4.[151,1436,934,2029]" pageId="4" pageNumber="165">
<emphasis id="7CF14B251008C048FF4EFAA6FEA66D4D" box="[201,307,1294,1318]" italics="true" pageId="4" pageNumber="165">Abdomen</emphasis>
(
<figureCitation id="D6BE8BB21008C048FEC1FAA6FE1B6D4D" box="[326,398,1294,1318]" captionStart="FIGURE 2" captionStartId="5.[151,250,1913,1936]" captionTargetBox="[263,1327,206,1864]" captionTargetId="figure@5.[237,1343,193,1884]" captionTargetPageId="5" captionText="FIGURE 2. Geotragus tuberculatus Chen: A male ventrites, ventral view B male metendosternite, dorsal view C male sternites VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view (g = groove) H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, lateral view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216582/files/figure.png" pageId="4" pageNumber="165">Fig. 2</figureCitation>
A): Surface of ventrite 1 depressed at middle, slightly convex at sides; suture I strongly bisinuate; surface of ventrite 2 moderately convex, without apical transverse impression; mesal length of ventrite 1 1.28× length of 2, 2 1.09× length of 3 and 4 combined; ventrite 5 parabolic, with convex surface.
</paragraph>
<paragraph id="4E3A97371008C048FF4EFAD2FA116D97" blockId="4.[151,1436,934,2029]" pageId="4" pageNumber="165">
<emphasis id="7CF14B251008C048FF4EFAD2FEEA6DF9" box="[201,383,1402,1426]" italics="true" pageId="4" pageNumber="165">Metendosternite</emphasis>
(
<figureCitation id="D6BE8BB21008C048FE14FAD2FE496DF9" box="[403,476,1402,1426]" captionStart="FIGURE 2" captionStartId="5.[151,250,1913,1936]" captionTargetBox="[263,1327,206,1864]" captionTargetId="figure@5.[237,1343,193,1884]" captionTargetPageId="5" captionText="FIGURE 2. Geotragus tuberculatus Chen: A male ventrites, ventral view B male metendosternite, dorsal view C male sternites VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view (g = groove) H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, lateral view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216582/files/figure.png" pageId="4" pageNumber="165">Fig. 2</figureCitation>
B): Stalk 0.36× length of furcal arms, trapezoidal and transverse, anterior part of longitudinal flange slightly shorter than posterior; hemiductus wide, truncate, positioned at apical half of furcal arms but near the midpoint; anterior tendons weak, positioned almost in the middle of furcal arms, opposite to hemiductus; furcal arms thin, apically clearly bifurcate, diverging at nearly 60° in relation to longitudinal flange.
</paragraph>
<paragraph id="4E3A97371008C048FF4EF9A1FD966EE1" blockId="4.[151,1436,934,2029]" pageId="4" pageNumber="165">
<emphasis id="7CF14B251008C048FF4EF9A1FF6B6E4B" box="[201,254,1545,1568]" italics="true" pageId="4" pageNumber="165">Legs</emphasis>
: Protibiae sinuate, bent inwards at apical 1/4; inner margin of protibiae with eight sharp and large teeth, each tooth with a spiniform seta just behind it; teeth of meso- and metatibiae sparse and much smaller than those of the protibiae; protibial mucro moderately large, distinct, not covered by setae; tarsi long and broad; tarsomere 1 almost 2× length of tarsomere 2.
</paragraph>
<paragraph id="4E3A97371008C048FF4EF93EFDCD6F86" blockId="4.[151,1436,934,2029]" pageId="4" pageNumber="165">
<emphasis id="7CF14B251008C048FF4EF93EFDB56EC5" box="[201,544,1686,1710]" italics="true" pageId="4" pageNumber="165">Male genitalia and terminalia</emphasis>
: Sternite VIII (
<figureCitation id="D6BE8BB21008C048FD53F93EFC8D6EC5" box="[724,792,1686,1710]" captionStart="FIGURE 2" captionStartId="5.[151,250,1913,1936]" captionTargetBox="[263,1327,206,1864]" captionTargetId="figure@5.[237,1343,193,1884]" captionTargetPageId="5" captionText="FIGURE 2. Geotragus tuberculatus Chen: A male ventrites, ventral view B male metendosternite, dorsal view C male sternites VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view (g = groove) H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, lateral view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216582/files/figure.png" pageId="4" pageNumber="165">Fig. 2</figureCitation>
C) divided into two hemisternites, transversely oriented, subtriangular; each hemisternite laterally acuminate, anterior margin widely roundly emarginate, posterior margin strongly angulate; membrane between sternite VIII and IX with two very small, dot-like sclerotizations. Sternite IX (
<figureCitation id="D6BE8BB21008C048FF18F8A8FF766F73" box="[159,227,1792,1816]" captionStart="FIGURE 2" captionStartId="5.[151,250,1913,1936]" captionTargetBox="[263,1327,206,1864]" captionTargetId="figure@5.[237,1343,193,1884]" captionTargetPageId="5" captionText="FIGURE 2. Geotragus tuberculatus Chen: A male ventrites, ventral view B male metendosternite, dorsal view C male sternites VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view (g = groove) H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, lateral view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216582/files/figure.png" pageId="4" pageNumber="165">Fig. 2</figureCitation>
C) with basal plate bilobed, each lobe semi-trapezoidal; spiculum gastrale 0.77× length of penis, apodeme slender, almost straight, slightly curved anteriorly. Penis (
<figureCitation id="D6BE8BB21008C048FC9BF88BFCF56F50" box="[796,864,1827,1851]" captionStart="FIGURE 2" captionStartId="5.[151,250,1913,1936]" captionTargetBox="[263,1327,206,1864]" captionTargetId="figure@5.[237,1343,193,1884]" captionTargetPageId="5" captionText="FIGURE 2. Geotragus tuberculatus Chen: A male ventrites, ventral view B male metendosternite, dorsal view C male sternites VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view (g = groove) H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, lateral view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216582/files/figure.png" pageId="4" pageNumber="165">Fig. 2</figureCitation>
D, 2E) in dorsal view with tube
<quantity id="897D3AD21008C048FB40F88BFABE6F50" box="[1223,1323,1827,1851]" metricMagnitude="-3" metricUnit="m" metricValue="2.63" pageId="4" pageNumber="165" unit="mm" value="2.63">2.63 mm</quantity>
in length, ca. 6.9× length of wide, temones
<quantity id="897D3AD21008C048FD90F8EFFDE86F34" box="[535,637,1863,1887]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="4" pageNumber="165" unit="mm" value="0.88">0.88 mm</quantity>
long; lateral margins subparallel at middle, diverging at ostium level, thereafter roundly converging, apex ogival; in lateral view curved, more strongly so near base and at apex, greatest width at midpoint. Tegmen (
<figureCitation id="D6BE8BB21008C048FE58F826FDB16FCD" box="[479,548,1934,1958]" captionStart="FIGURE 2" captionStartId="5.[151,250,1913,1936]" captionTargetBox="[263,1327,206,1864]" captionTargetId="figure@5.[237,1343,193,1884]" captionTargetPageId="5" captionText="FIGURE 2. Geotragus tuberculatus Chen: A male ventrites, ventral view B male metendosternite, dorsal view C male sternites VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view (g = groove) H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, lateral view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216582/files/figure.png" pageId="4" pageNumber="165">Fig. 2</figureCitation>
F) 0.38× length of penis, with ring narrow, parameroid lobes developed with basal half more sclerotized; tegminal apodeme slender, forming a Y-shape with basal piece; apex of apodeme slightly expanded to form a round plate.
</paragraph>
<caption id="1AFAC7BF1009C049FF10F8D1FDD36F80" httpUri="https://zenodo.org/record/216582/files/figure.png" pageId="5" pageNumber="166" targetBox="[263,1327,206,1864]" targetPageId="5">
<paragraph id="4E3A97371009C049FF10F8D1FDD36F80" blockId="5.[151,1436,1913,2027]" pageId="5" pageNumber="166">
<emphasis id="7CF14B251009C049FF10F8D1FE876FE4" bold="true" box="[151,274,1913,1936]" pageId="5" pageNumber="166">FIGURE 2.</emphasis>
<taxonomicName id="8985ECB41009C049FE9FF8D2FDD06FE4" authority="Chen" authorityName="Chen" box="[280,581,1913,1935]" class="Insecta" family="Curculionidae" genus="Geotragus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="166" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis id="7CF14B251009C049FE9FF8D2FD906FE4" box="[280,517,1913,1935]" italics="true" pageId="5" pageNumber="166">Geotragus tuberculatus</emphasis>
Chen
</taxonomicName>
:
<emphasis id="7CF14B251009C049FDC9F8D1FDF56FE5" bold="true" box="[590,608,1913,1934]" pageId="5" pageNumber="166">A</emphasis>
male ventrites, ventral view
<emphasis id="7CF14B251009C049FC04F8D2FC016FE4" bold="true" box="[899,916,1914,1935]" pageId="5" pageNumber="166">B</emphasis>
male metendosternite, dorsal view
<emphasis id="7CF14B251009C049FB7EF8D1FA9E6FFB" bold="true" box="[1273,1291,1913,1936]" pageId="5" pageNumber="166">C</emphasis>
male sternites VIII and IX, dorsal view
<emphasis id="7CF14B251009C049FE27F830FE276FC6" bold="true" box="[416,434,1944,1965]" pageId="5" pageNumber="166">D</emphasis>
penis, dorsal view
<emphasis id="7CF14B251009C049FDF9F830FD1A6FC6" bold="true" box="[638,655,1944,1965]" pageId="5" pageNumber="166">E</emphasis>
penis, lateral view
<emphasis id="7CF14B251009C049FCDAF830FCF96FC6" bold="true" box="[861,876,1944,1965]" pageId="5" pageNumber="166">F</emphasis>
tegmen, dorsal view
<emphasis id="7CF14B251009C049FBCAF83FFBF46FC5" bold="true" box="[1101,1121,1943,1966]" pageId="5" pageNumber="166">G</emphasis>
female ventrites, ventral view (g=groove)
<emphasis id="7CF14B251009C049FE97F81FFEB16FA7" bold="true" box="[272,292,1975,1996]" pageId="5" pageNumber="166">H</emphasis>
sternite VIII and female genitalia, lateral view
<emphasis id="7CF14B251009C049FC8BF81FFC836FA7" bold="true" box="[780,790,1975,1996]" pageId="5" pageNumber="166">I</emphasis>
coxites and styli, dorsal view
<emphasis id="7CF14B251009C049FBC8F81FFBC96FA7" bold="true" box="[1103,1116,1975,1996]" pageId="5" pageNumber="166">J</emphasis>
bursal sclerites, lateral view
<emphasis id="7CF14B251009C049FA0FF81FFA096FA7" bold="true" box="[1416,1436,1975,1996]" pageId="5" pageNumber="166">K</emphasis>
spermatheca, lateral view. Scales: 0.5 mm.
</paragraph>
</caption>
<paragraph id="4E3A9737100AC04AFF4EFF3FFEC8689C" blockId="6.[151,1436,151,1242]" pageId="6" pageNumber="167">
<emphasis id="7CF14B25100AC04AFF4EFF3FFD9F68DB" bold="true" box="[201,522,151,176]" pageId="6" pageNumber="167">
Variation. Male
<typeStatus id="913E2995100AC04AFE0AFF30FD9068DB" box="[397,517,152,176]" pageId="6" pageNumber="167" type="paratype">paratypes</typeStatus>
.
</emphasis>
Measurements (in mm): Standard length: 5.606.10; pronotal length: 1.551.75; pronotal width: 1.651.95; elytral length: 4.054.20; elytral width: 1.602.60; rostral length: 1.051.13; rostral width: 0.700.80.
</paragraph>
<paragraph id="4E3A9737100AC04AFF57FEACFE116ACC" blockId="6.[151,1436,151,1242]" pageId="6" pageNumber="167">
<emphasis id="7CF14B25100AC04AFF57FEACFE356977" bold="true" box="[208,416,260,284]" pageId="6" pageNumber="167">
Female
<typeStatus id="913E2995100AC04AFEAAFEACFE0F6977" box="[301,410,260,284]" pageId="6" pageNumber="167" type="paratype">paratype</typeStatus>
.
</emphasis>
Measurements (in mm): Standard length: 6.80; pronotal length: 1.75; pronotal width: 2.05; elytral length: 5.30; elytral width: 3.50; rostral length: 1.05; rostral width: 0.88. Elytra 1.352.19× length of male, pronotum 0.85× length of male, ventrite 5 (
<figureCitation id="D6BE8BB2100AC04AFD05FEE4FD50690F" box="[642,709,332,356]" captionStart="FIGURE 2" captionStartId="5.[151,250,1913,1936]" captionTargetBox="[263,1327,206,1864]" captionTargetId="figure@5.[237,1343,193,1884]" captionTargetPageId="5" captionText="FIGURE 2. Geotragus tuberculatus Chen: A male ventrites, ventral view B male metendosternite, dorsal view C male sternites VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view (g = groove) H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, lateral view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216582/files/figure.png" pageId="6" pageNumber="167">Fig. 2</figureCitation>
G) parabolic, longer than ventrite 2. Sternite VIII (
<figureCitation id="D6BE8BB2100AC04AFA86FEE4FAD0690F" box="[1281,1349,332,356]" captionStart="FIGURE 2" captionStartId="5.[151,250,1913,1936]" captionTargetBox="[263,1327,206,1864]" captionTargetId="figure@5.[237,1343,193,1884]" captionTargetPageId="5" captionText="FIGURE 2. Geotragus tuberculatus Chen: A male ventrites, ventral view B male metendosternite, dorsal view C male sternites VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view (g = groove) H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, lateral view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216582/files/figure.png" pageId="6" pageNumber="167">Fig. 2</figureCitation>
H) with spiculum ventrale thin, rod-like and sinuate; lamina tightly folded, in lateral view triangular, ventral margin angulated close to speculum ventral, strongly sclerotized. Ovipositor (
<figureCitation id="D6BE8BB2100AC04AFC2EFE3CFC7969C7" box="[937,1004,404,428]" captionStart="FIGURE 2" captionStartId="5.[151,250,1913,1936]" captionTargetBox="[263,1327,206,1864]" captionTargetId="figure@5.[237,1343,193,1884]" captionTargetPageId="5" captionText="FIGURE 2. Geotragus tuberculatus Chen: A male ventrites, ventral view B male metendosternite, dorsal view C male sternites VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view (g = groove) H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, lateral view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216582/files/figure.png" pageId="6" pageNumber="167">Fig. 2</figureCitation>
H,
<date id="3A3BB1F7100AC04AFB88FE3CFBB269C7" box="[1039,1063,404,428]" pageId="6" pageNumber="167">2I</date>
) with coxites similar in length to sternite VIII, folded along middle, lateral margins strongly sclerotized, posteriorly slightly narrowed; styli relatively small, cylindrical, apically inserted two long setae. Bursal sclerites (
<figureCitation id="D6BE8BB2100AC04AFB93FE74FBCD699F" box="[1044,1112,476,500]" captionStart="FIGURE 2" captionStartId="5.[151,250,1913,1936]" captionTargetBox="[263,1327,206,1864]" captionTargetId="figure@5.[237,1343,193,1884]" captionTargetPageId="5" captionText="FIGURE 2. Geotragus tuberculatus Chen: A male ventrites, ventral view B male metendosternite, dorsal view C male sternites VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view (g = groove) H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, lateral view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216582/files/figure.png" pageId="6" pageNumber="167">Fig. 2</figureCitation>
J) near the point of union of spermathecal duct and of oviduct, facing each other at middle, each sclerite in lateral view chevron-shaped, with both arms curved, dorsal surface of mesal area with several spines. Spermatheca (
<figureCitation id="D6BE8BB2100AC04AFB11FD8CFB776A57" box="[1174,1250,548,572]" captionStart="FIGURE 2" captionStartId="5.[151,250,1913,1936]" captionTargetBox="[263,1327,206,1864]" captionTargetId="figure@5.[237,1343,193,1884]" captionTargetPageId="5" captionText="FIGURE 2. Geotragus tuberculatus Chen: A male ventrites, ventral view B male metendosternite, dorsal view C male sternites VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view (g = groove) H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, lateral view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216582/files/figure.png" pageId="6" pageNumber="167">Fig. 2</figureCitation>
K) with corpus subquadrate; cornu elongate, widely curved, U-shaped, apically gradually narrowed, apex narrowly rounded; nodulus prominent, subcontiguous with and angled almost 90° in relation to ramus; ramus trapezoid, prominent and moderately long.
</paragraph>
</subSubSection>
<subSubSection id="069FC4BC100AC04AFF4EFD1FFD7E6CF9" pageId="6" pageNumber="167" type="materials_examined">
<paragraph id="4E3A9737100AC04AFF4EFD1FFA8A6BB1" blockId="6.[151,1436,151,1242]" pageId="6" pageNumber="167">
<emphasis id="7CF14B25100AC04AFF4EFD1FFD936ABB" bold="true" box="[201,518,695,720]" pageId="6" pageNumber="167">
Material examined.
<typeStatus id="913E2995100AC04AFE46FD10FD936ABB" box="[449,518,696,720]" pageId="6" pageNumber="167">Types</typeStatus>
</emphasis>
: 13: (white): [Yúnnán, printed], [Lúshuǐ, handwritten] / [Yáojiāpíng, handwritten],
<quantity id="897D3AD2100AC04AFE5AFD77FDBE6A9C" box="[477,555,735,759]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="6" pageNumber="167" unit="m" value="2800.0">2800m</quantity>
[handwritten] / [
<collectingCountry id="3692D7A7100AC04AFC36FD77FBB26A9C" box="[945,1063,735,759]" name="China" pageId="6" pageNumber="167">Zhōngguó</collectingCountry>
Kēxuéyuàn, printed]; (white): 1981.
<date id="3A3BB1F7100AC04AFF52FCAFFE9D6B74" box="[213,264,775,799]" pageId="6" pageNumber="167">VI.6</date>
[handwritten] / [Cǎijízhĕ, Wáng Shūyǒng, printed]; (red, printed):
<typeStatus id="913E2995100AC04AFB2BFCAFFAD66B74" box="[1196,1347,775,799]" pageId="6" pageNumber="167" type="holotype">HOLOTYPE</typeStatus>
; (white, printed): IOZ(E) 905001. 1Ƥ: same data as
<typeStatus id="913E2995100AC04AFD0BFC83FD7B6B28" box="[652,750,811,835]" pageId="6" pageNumber="167" type="holotype">holotype</typeStatus>
except ALLOTYPE printed on sea-green paper and IOZ(E) 905002. 13: same data as
<typeStatus id="913E2995100AC04AFE48FCE7FDA46B0C" box="[463,561,847,871]" pageId="6" pageNumber="167" type="holotype">holotype</typeStatus>
except
<typeStatus id="913E2995100AC04AFD09FCF8FC8C6B03" box="[654,793,848,872]" pageId="6" pageNumber="167" type="paratype">PARATYPE</typeStatus>
printed on yellow paper and IOZ(E) 905005. 13: with same locality as
<typeStatus id="913E2995100AC04AFED8FCDBFE546BE0" box="[351,449,883,907]" pageId="6" pageNumber="167" type="holotype">holotype</typeStatus>
except
<quantity id="897D3AD2100AC04AFDA7FCDBFDE46BE0" box="[544,625,883,907]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="167" unit="m" value="2500.0">2500m</quantity>
, collecting date 1981.
<date id="3A3BB1F7100AC04AFCF5FCDBFC3C6BE0" box="[882,937,883,907]" pageId="6" pageNumber="167" value="1924-05">V.24</date>
,
<typeStatus id="913E2995100AC04AFC3DFCDBFBD06BE0" box="[954,1093,883,907]" pageId="6" pageNumber="167" type="paratype">PARATYPE</typeStatus>
printed on yellow paper and IOZ(E) 905004. 13: (white): [Yúnnán, printed], [Lúshuǐ, handwritten] / [Piànmǎ, handwritten],
<quantity id="897D3AD2100AC04AFF10FC6AFF706BB1" box="[151,229,962,986]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="167" unit="m" value="2300.0">2300m</quantity>
[handwritten] / [
<collectingCountry id="3692D7A7100AC04AFDB6FC6AFD326BB1" box="[561,679,962,986]" name="China" pageId="6" pageNumber="167">Zhōngguó</collectingCountry>
Kēxuéyuàn, printed]; (white): 1981.
<date id="3A3BB1F7100AC04AFBBAFC6AFBE66BB1" box="[1085,1139,962,986]" pageId="6" pageNumber="167" value="1931-05">V.31</date>
[handwritten] /
</paragraph>
<paragraph id="4E3A9737100AC04AFF5EFC42FE956C05" blockId="6.[151,1436,151,1242]" pageId="6" pageNumber="167">
[Cǎijízhĕ, Wáng Shūyǒng, printed]; (yellow, printed):
<typeStatus id="913E2995100AC04AFCB9FC43FC586C68" box="[830,973,1003,1027]" pageId="6" pageNumber="167" type="paratype">PARATYPE</typeStatus>
; (white, printed): IOZ(E) 905003.
<emphasis id="7CF14B25100AC04AFAD3FC42FE856C4C" bold="true" pageId="6" pageNumber="167">Other materials:</emphasis>
13:
<collectingCountry id="3692D7A7100AC04AFECAFBA6FE3C6C4D" box="[333,425,1038,1062]" name="China" pageId="6" pageNumber="167">CHINA</collectingCountry>
, Yunnan Prov. / Fugong, Maji Town / Majimi, plant beating /
<geoCoordinate id="2BB1F1F0100AC04AFBF1FBA6FA976C4D" box="[1142,1282,1038,1062]" direction="north" orientation="latitude" pageId="6" pageNumber="167" precision="1" value="27.40301">27.40301°N</geoCoordinate>
,
<geoCoordinate id="2BB1F1F0100AC04AFA88FBA6FA0D6C4D" box="[1295,1432,1038,1062]" direction="east" orientation="longitude" pageId="6" pageNumber="167" precision="1" value="98.82446">98.82446°E</geoCoordinate>
;
<quantity id="897D3AD2100AC04AFF10FB9AFF7D6C21" box="[151,232,1074,1098]" metricMagnitude="3" metricUnit="m" metricValue="1.505" pageId="6" pageNumber="167" unit="m" value="1505.0">1505m</quantity>
, 2005.8.26 day / Liang H.B, Zhang J.F. / Califormia Academy &amp; / IOZ., Chinese. Acad. Sci. / IOZ(E) 1798751.
</paragraph>
<paragraph id="4E3A9737100AC04AFF4EFBD2FD7E6CF9" blockId="6.[151,1436,151,1242]" box="[201,747,1146,1171]" pageId="6" pageNumber="167">
<typeStatus id="913E2995100AC04AFF4EFBD2FEA46CF9" box="[201,305,1146,1170]" pageId="6" pageNumber="167" type="holotype">Holotype</typeStatus>
and all
<typeStatus id="913E2995100AC04AFE0DFBD3FE606CF8" box="[394,501,1147,1171]" pageId="6" pageNumber="167" type="paratype">paratypes</typeStatus>
conserved in
<collectionCode id="28940FF2100AC04AFD15FBD2FD726CF9" box="[658,743,1146,1170]" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="6" pageNumber="167">IZCAS</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="069FC4BC100AC04AFF4EFB36FE316CB1" pageId="6" pageNumber="167" type="distribution">
<paragraph id="4E3A9737100AC04AFF4EFB36FE316CB1" blockId="6.[151,1436,151,1242]" pageId="6" pageNumber="167">
<emphasis id="7CF14B25100AC04AFF4EFB36FEF76CDC" bold="true" box="[201,354,1182,1207]" pageId="6" pageNumber="167">Distribution.</emphasis>
<collectingCountry id="3692D7A7100AC04AFEEEFB36FE256CDD" box="[361,432,1182,1206]" name="China" pageId="6" pageNumber="167">China</collectingCountry>
: Yunnan (Fugong, Lushui). Altitudinal range of this species is from
<quantity id="897D3AD2100AC04AFB21FB36FAA86CDD" box="[1190,1341,1182,1206]" metricMagnitude="3" metricUnit="m" metricValue="2.1525" metricValueMax="2.8" metricValueMin="1.505" pageId="6" pageNumber="167" unit="m" value="2152.5" valueMax="2800.0" valueMin="1505.0">15052800m</quantity>
. Fugong is a new locality record.
</paragraph>
</subSubSection>
</treatment>
</document>