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<document id="BDDCAF562F9D8BB8F514D33E38126455" ID-DOI="10.11646/zootaxa.3619.2.5" ID-GBIF-Dataset="549f99ee-2153-4d61-a953-8def30c8159d" ID-ISSN="1175-5326" ID-Zenodo-Dep="216580" ID-ZooBank="DBB8DB78-09DF-4F68-AFB1-AAC8EBC95DD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460577749427" checkinUser="plazi" docAuthor="Ren, Li, Alonso-Zarazaga, Miguel A. &amp; Zhang, Runzhi" docDate="2013" docId="C62C26211000C043FF10FF3FFA936ADC" docLanguage="en" docName="zt03619p182.pdf" docOrigin="Zootaxa 3619 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Geotragus rugosus Ren, Alonso-Zarazaga &amp; Zhang, 2013, sp. nov." docType="treatment" docVersion="8" lastPageNumber="176" masterDocId="3A155E59100CC04CFF87FFA8FF95686B" masterDocTitle="Revision of the Chinese Geotragus Schoenherr with description of three new species (Coleoptera: Curculionidae: Entiminae)" masterLastPageNumber="182" masterPageNumber="161" pageNumber="173" updateTime="1698323549919" updateUser="plazi">
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<mods:title id="B98BC44AD60CBEA3EE586FE8615D64A7">Revision of the Chinese Geotragus Schoenherr with description of three new species (Coleoptera: Curculionidae: Entiminae)</mods:title>
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<mods:namePart id="C32F20350B0C07B62FAFDF0C0D9D12BD">Ren, Li</mods:namePart>
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<mods:namePart id="927BCEB077B9260ACD77054F007955F0">Alonso-Zarazaga, Miguel A.</mods:namePart>
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<mods:namePart id="83EC1CE8B66D1634DBD44E3E25360D24">Zhang, Runzhi</mods:namePart>
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<mods:title id="36DF46B22F71C2A79468D8824C072345">Zootaxa</mods:title>
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<mods:date id="9C9758882C5DD555C2CDD9BCF3B91451">2013</mods:date>
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<treatment id="C62C26211000C043FF10FF3FFA936ADC" ID-DOI="http://doi.org/10.5281/zenodo.6158925" ID-GBIF-Taxon="119560661" ID-Zenodo-Dep="6158925" LSID="urn:lsid:plazi:treatment:C62C26211000C043FF10FF3FFA936ADC" httpUri="http://treatment.plazi.org/id/C62C26211000C043FF10FF3FFA936ADC" lastPageId="15" lastPageNumber="176" pageId="12" pageNumber="173">
<subSubSection id="069FC4BC1000C040FF10FF3FFE9368BF" pageId="12" pageNumber="173" type="nomenclature">
<paragraph id="4E3A97371000C040FF10FF3FFE4768DB" blockId="12.[151,466,151,212]" box="[151,466,151,176]" pageId="12" pageNumber="173">
<heading id="1572205B1000C040FF10FF3FFE4768DB" bold="true" box="[151,466,151,176]" fontSize="10" level="2" pageId="12" pageNumber="173" reason="2">
<emphasis id="7CF14B251000C040FF10FF3FFE4768DB" bold="true" box="[151,466,151,176]" pageId="12" pageNumber="173">
<taxonomicName id="8985ECB41000C040FF10FF3FFEE768C4" ID-CoL="3FT5X" box="[151,370,151,176]" class="Insecta" family="Curculionidae" genus="Geotragus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="173" phylum="Arthropoda" rank="species" species="rugosus" status="sp. nov.">
<emphasis id="7CF14B251000C040FF10FF3FFEE768C4" bold="true" box="[151,370,151,176]" italics="true" pageId="12" pageNumber="173">Geotragus rugosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67C2F65E1000C040FEFFFF30FE4768DB" box="[376,466,152,176]" pageId="12" pageNumber="173" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="4E3A97371000C040FF10FF14FE9368BF" blockId="12.[151,466,151,212]" box="[151,262,188,212]" pageId="12" pageNumber="173">
(
<figureCitation id="D6BE8BB21000C040FF18FF14FF7768BF" box="[159,226,188,212]" captionStart="FIGURE 6" captionStartId="12.[151,250,1439,1462]" captionTargetBox="[151,1436,805,1418]" captionTargetId="figure@12.[151,1436,805,1418]" captionTargetPageId="12" captionText="FIGURE 6. Habitus of Geotragus rugosus sp. nov., male, holotype: A dorsal view B lateral view C head, dorsal view. Scales: 1 mm." httpUri="https://zenodo.org/record/216586/files/figure.png" pageId="12" pageNumber="173">Fig. 6</figureCitation>
,
<figureCitation id="D6BE8BB21000C040FF68FF15FF6868BF" box="[239,253,189,212]" captionStart="FIGURE 7" captionStartId="13.[151,250,1904,1927]" captionTargetBox="[302,1315,219,1853]" captionTargetId="figure@13.[274,1322,193,1878]" captionTargetPageId="13" captionText="FIGURE 7. Geotragus rugosus sp. nov.: A male ventrites, ventral view B male metendosternite, dorsal view C male sternite VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, dorsal view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216587/files/figure.png" pageId="12" pageNumber="173">7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="069FC4BC1000C040FF10FEABFC01690F" pageId="12" pageNumber="173" type="diagnosis">
<paragraph id="4E3A97371000C040FF10FEABFC01690F" blockId="12.[151,1436,259,752]" pageId="12" pageNumber="173">
<emphasis id="7CF14B251000C040FF10FEABFE846977" bold="true" box="[151,273,259,284]" pageId="12" pageNumber="173">Diagnosis.</emphasis>
This species can be easily distinguished from other species by the following characters: dorsal part of rostrum strongly corrugated, with a deep wide central furrow and many subparallel furrows; preocular area with a wide, deep and oblique impression laterally on each side of rostrum.
</paragraph>
</subSubSection>
<subSubSection id="069FC4BC1000C042FF4EFEC7FB406F99" lastPageId="14" lastPageNumber="175" pageId="12" pageNumber="173" type="description">
<paragraph id="4E3A97371000C040FF4EFEC7FB0B69C7" blockId="12.[151,1436,259,752]" pageId="12" pageNumber="173">
<emphasis id="7CF14B251000C040FF4EFEC7FD8F69E3" bold="true" box="[201,538,367,392]" pageId="12" pageNumber="173">Description. Holotype-male.</emphasis>
<emphasis id="7CF14B251000C040FDA4FED9FD5069E3" box="[547,709,369,392]" italics="true" pageId="12" pageNumber="173">Measurements</emphasis>
(in mm): Standard length: 6.50; pronotal length: 2.20; pronotal width: 2.05; elytral length: 4.80; elytral width: 2.60; rostral length: 1.04; rostral width: 0.80.
</paragraph>
<paragraph id="4E3A97371000C040FF4EFE10FB206A9B" blockId="12.[151,1436,259,752]" pageId="12" pageNumber="173">
<emphasis id="7CF14B251000C040FF4EFE10FE0069BB" box="[201,405,440,464]" italics="true" pageId="12" pageNumber="173">Habitus and color</emphasis>
(
<figureCitation id="D6BE8BB21000C040FE22FE1FFE7C69A4" box="[421,489,439,463]" captionStart="FIGURE 6" captionStartId="12.[151,250,1439,1462]" captionTargetBox="[151,1436,805,1418]" captionTargetId="figure@12.[151,1436,805,1418]" captionTargetPageId="12" captionText="FIGURE 6. Habitus of Geotragus rugosus sp. nov., male, holotype: A dorsal view B lateral view C head, dorsal view. Scales: 1 mm." httpUri="https://zenodo.org/record/216586/files/figure.png" pageId="12" pageNumber="173">Fig. 6</figureCitation>
A, 6B): Body slender; integument reddish brown, only head and sides of elytra dark red, antennae and legs pale reddish brown; with coppery metallic green to yellowish brown scales, scales moderately dense, tessellate, contiguous but not overlapping; scales on dorsal and lateral surfaces of rostrum sparse, oval; posterior half of antennal scrobes glabrous; pronotum with round scales, moderately dense; scales on elytra round, moderately dense; scales on ventrites dense, round to elongate-oval from ventrites 15, ventrite 5 with lanceolate scales; legs densely covered by scales; body with recumbent to subrecumbent sparse and slightly thick setae; rostrum covered with recumbent to subrecumbent fine setae; setae of antennal scape and desmomeres 17 long, fine and sparse; setae of interstriae lanceolate, sparse and subrecumbent; setae on ventral surface thick and sparse; setae of tibiae long and moderately dense, femora with subrecumbent lanceolate setae.
</paragraph>
<caption id="1AFAC7BF1000C040FF10FA37FF4C6DBF" httpUri="https://zenodo.org/record/216586/files/figure.png" pageId="12" pageNumber="173" targetBox="[151,1436,805,1418]" targetPageId="12">
<paragraph id="4E3A97371000C040FF10FA37FF4C6DBF" blockId="12.[151,1437,1439,1492]" pageId="12" pageNumber="173">
<emphasis id="7CF14B251000C040FF10FA37FE866DDE" bold="true" box="[151,275,1439,1462]" pageId="12" pageNumber="173">FIGURE 6.</emphasis>
Habitus of
<taxonomicName id="8985ECB41000C040FE0DFA08FDDE6DDD" box="[394,587,1440,1462]" class="Insecta" family="Curculionidae" genus="Geotragus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="173" phylum="Arthropoda" rank="species" species="rugosus" status="sp. nov.">
<emphasis id="7CF14B251000C040FE0DFA08FDDE6DDD" box="[394,587,1440,1462]" italics="true" pageId="12" pageNumber="173">Geotragus rugosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67C2F65E1000C040FDD5FA08FD376DDE" box="[594,674,1440,1461]" pageId="12" pageNumber="173" rank="species">
<emphasis id="7CF14B251000C040FDD5FA08FD376DDE" bold="true" box="[594,674,1440,1461]" pageId="12" pageNumber="173">sp. nov.</emphasis>
</taxonomicNameLabel>
, male, holotype:
<emphasis id="7CF14B251000C040FCD3FA08FCF36DDE" bold="true" box="[852,870,1440,1461]" pageId="12" pageNumber="173">A</emphasis>
dorsal view
<emphasis id="7CF14B251000C040FC6EFA08FC6F6DDE" bold="true" box="[1001,1018,1440,1461]" pageId="12" pageNumber="173">B</emphasis>
lateral view
<emphasis id="7CF14B251000C040FBF9FA37FB056DDD" bold="true" box="[1150,1168,1439,1462]" pageId="12" pageNumber="173">C</emphasis>
head, dorsal view. Scales: 1 mm.
</paragraph>
</caption>
<paragraph id="4E3A97371000C040FF4EF9A9FE0E6E0A" blockId="12.[151,1437,1537,2030]" pageId="12" pageNumber="173">
<emphasis id="7CF14B251000C040FF4EF9A9FE936E72" box="[201,262,1537,1561]" italics="true" pageId="12" pageNumber="173">Head</emphasis>
(
<figureCitation id="D6BE8BB21000C040FE9FF9A9FECB6E72" box="[280,350,1537,1561]" captionStart="FIGURE 6" captionStartId="12.[151,250,1439,1462]" captionTargetBox="[151,1436,805,1418]" captionTargetId="figure@12.[151,1436,805,1418]" captionTargetPageId="12" captionText="FIGURE 6. Habitus of Geotragus rugosus sp. nov., male, holotype: A dorsal view B lateral view C head, dorsal view. Scales: 1 mm." httpUri="https://zenodo.org/record/216586/files/figure.png" pageId="12" pageNumber="173">Fig. 6</figureCitation>
C): Moderately flat, dorsal part with many longitudinal sulci, corrugated, punctures small and dense; eyes strongly convex, with a deep circumocular sulcus; frons moderately flat, not higher than dorsal part of rostrum in lateral view.
</paragraph>
<paragraph id="4E3A97371000C040FF4EF9C6FBC46F36" blockId="12.[151,1437,1537,2030]" pageId="12" pageNumber="173">
<emphasis id="7CF14B251000C040FF4EF9C6FEBD6EEE" box="[201,296,1646,1669]" italics="true" pageId="12" pageNumber="173">Rostrum</emphasis>
(
<figureCitation id="D6BE8BB21000C040FEB0F9C5FEEF6EEE" box="[311,378,1645,1669]" captionStart="FIGURE 6" captionStartId="12.[151,250,1439,1462]" captionTargetBox="[151,1436,805,1418]" captionTargetId="figure@12.[151,1436,805,1418]" captionTargetPageId="12" captionText="FIGURE 6. Habitus of Geotragus rugosus sp. nov., male, holotype: A dorsal view B lateral view C head, dorsal view. Scales: 1 mm." httpUri="https://zenodo.org/record/216586/files/figure.png" pageId="12" pageNumber="173">Fig. 6</figureCitation>
C): In dorsal view, 1.3× length of wide; apex narrower than base; base wider than frons; dorsal surface depressed in median area of rostrum, corrugated, with a wide and deep median sulcus, extending from antennal insertion to vertex, along the median sulcus, with many subparallel longitudinal sulci, short and deep; epistome broad, posterior angle of epistome between 45° and 90°, with carina distinct; mandible scars quadrate to rectangular; in dorsal view ventral margin of scrobes visible from apex to middle of rostrum; in lateral view, with a deep and wide groove positioned laterally between eyes and antennal scrobes, perpendicular to antennal scrobes, close to ventral margin of rostrum accompanied by subparallel deep and wide sulcus.
</paragraph>
<paragraph id="4E3A97371000C042FF4EF8C1FD5368B8" blockId="12.[151,1437,1537,2030]" lastBlockId="14.[151,1437,151,2034]" lastPageId="14" lastPageNumber="175" pageId="12" pageNumber="173">
<emphasis id="7CF14B251000C040FF4EF8C1FEA66FEA" box="[201,307,1897,1921]" italics="true" pageId="12" pageNumber="173">Antennae</emphasis>
: Scape slender, reaching middle of eyes, 0.82× length of funicle; desmomere 1 1.24× length of 2, both segments elongate clavate, desmomere 1 apical rather stout, strongly narrowed toward base; desmomere 3 as long as 4, both segments elongate moniliform, 0.67× length of desmomere 2; desmomere 5 as long as 6, both segments moniliform and 0.93× length of desmomere 4; desmomere 7 much wider than and 1.15× length of 6, moniliform; pubescence of funicle similar throughout; club segment 1 1.6× length of 2, segment 2 0.6× length of 3; segment 3 with a marked
<emphasis id="7CF14B251002C042FE30FF14FD8568BF" box="[439,528,188,212]" italics="true" pageId="14" pageNumber="175">annulus</emphasis>
beyond middle.
</paragraph>
<caption id="1AFAC7BF1001C041FF10F8D8FE2E6F88" httpUri="https://zenodo.org/record/216587/files/figure.png" pageId="13" pageNumber="174" targetBox="[302,1315,219,1853]" targetPageId="13">
<paragraph id="4E3A97371001C041FF10F8D8FE2E6F88" blockId="13.[151,1436,1904,2019]" pageId="13" pageNumber="174">
<emphasis id="7CF14B251001C041FF10F8D8FE866FED" bold="true" box="[151,275,1904,1927]" pageId="13" pageNumber="174">FIGURE 7.</emphasis>
<taxonomicName id="8985ECB41001C041FE9CF8DAFE486FEC" box="[283,477,1906,1927]" class="Insecta" family="Curculionidae" genus="Geotragus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="174" phylum="Arthropoda" rank="species" species="rugosus" status="sp. nov.">
<emphasis id="7CF14B251001C041FE9CF8DAFE486FEC" box="[283,477,1906,1927]" italics="true" pageId="13" pageNumber="174">Geotragus rugosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67C2F65E1001C041FE63F8D9FDA36FED" box="[484,566,1905,1926]" pageId="13" pageNumber="174" rank="species">
<emphasis id="7CF14B251001C041FE63F8D9FDA36FED" bold="true" box="[484,566,1905,1926]" pageId="13" pageNumber="174">sp. nov.</emphasis>
</taxonomicNameLabel>
:
<emphasis id="7CF14B251001C041FDC3F8D9FDC36FED" bold="true" box="[580,598,1905,1926]" pageId="13" pageNumber="174">A</emphasis>
male ventrites, ventral view
<emphasis id="7CF14B251001C041FC05F8D9FC066FED" bold="true" box="[898,915,1905,1926]" pageId="13" pageNumber="174">B</emphasis>
male metendosternite, dorsal view
<emphasis id="7CF14B251001C041FB78F8D8FA846FEC" bold="true" box="[1279,1297,1904,1927]" pageId="13" pageNumber="174">C</emphasis>
male sternite VIII and IX, dorsal view
<emphasis id="7CF14B251001C041FE1DF838FE396FCE" bold="true" box="[410,428,1936,1957]" pageId="13" pageNumber="174">D</emphasis>
penis, dorsal view
<emphasis id="7CF14B251001C041FDF4F838FD116FCE" bold="true" box="[627,644,1936,1957]" pageId="13" pageNumber="174">E</emphasis>
penis, lateral view
<emphasis id="7CF14B251001C041FCCBF838FCCE6FCE" bold="true" box="[844,859,1936,1957]" pageId="13" pageNumber="174">F</emphasis>
tegmen, dorsal view
<emphasis id="7CF14B251001C041FBBFF827FBD96FCD" bold="true" box="[1080,1100,1935,1958]" pageId="13" pageNumber="174">G</emphasis>
female ventrites, ventral view
<emphasis id="7CF14B251001C041FA0FF838FA096FCE" bold="true" box="[1416,1436,1936,1957]" pageId="13" pageNumber="174">H</emphasis>
sternite VIII and female genitalia, lateral view
<emphasis id="7CF14B251001C041FDFCF807FD106FAF" bold="true" box="[635,645,1967,1988]" pageId="13" pageNumber="174">I</emphasis>
coxites and styli, dorsal view
<emphasis id="7CF14B251001C041FC46F807FC5B6FAF" bold="true" box="[961,974,1967,1988]" pageId="13" pageNumber="174">J</emphasis>
bursal sclerites, dorsal view
<emphasis id="7CF14B251001C041FB7BF807FA856FAF" bold="true" box="[1276,1296,1967,1988]" pageId="13" pageNumber="174">K</emphasis>
spermatheca, lateral view. Scales: 0.5 mm.
</paragraph>
</caption>
<paragraph id="4E3A97371002C042FF4EFF48FCA16909" blockId="14.[151,1437,151,2034]" pageId="14" pageNumber="175">
<emphasis id="7CF14B251002C042FF4EFF48FEAD689C" box="[201,312,224,247]" italics="true" pageId="14" pageNumber="175">Pronotum</emphasis>
: 1.07× as long as wide, elongate; anterior and posterior margin truncate; disc with a fine, shallow, incomplete median longitudinal groove; posterior margin distinct carinate, linear; dorsal surface of pronotum with slightly dense, large and round granules, space between granules moderately convex, each granule with one recumbent lanceolate seta on top; postocular vibrissae long.
</paragraph>
<paragraph id="4E3A97371002C042FF4EFEC6FCD769ED" blockId="14.[151,1437,151,2034]" box="[201,834,366,391]" pageId="14" pageNumber="175">
<taxonomicName id="8985ECB41002C042FF4EFEC6FE1F69EC" authority="Large" authorityName="Large" box="[201,394,366,391]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="14" pageNumber="175" phylum="Arthropoda" rank="genus">
<emphasis id="7CF14B251002C042FF4EFEC6FEA269ED" box="[201,311,366,390]" italics="true" pageId="14" pageNumber="175">Scutellum</emphasis>
: Large
</taxonomicName>
, reddish black, apex broadly V-shaped.
</paragraph>
<paragraph id="4E3A97371002C042FF4EFE3AFB266ACF" blockId="14.[151,1437,151,2034]" pageId="14" pageNumber="175">
<emphasis id="7CF14B251002C042FF4EFE3AFE9A69C1" box="[201,271,402,426]" italics="true" pageId="14" pageNumber="175">Elytra</emphasis>
: Base truncate; in lateral view moderately convex; in dorsal view, lateral margins clearly diverging from base to basal 1/5, then subparallel, reaching its greatest width nearly at declivity, thence clearly converging towards the apex, apex blunt, W-shaped; odd interstriae slightly more raised than even ones; interstria 1 moderately raised along basal 1/8, forming a longitudinal crest, not raised at declivity, top of declivity normally rounded in lateral view; interstria 3 much wider than remaining interstriae, distinctly raised and expanded at basal 1/4, forming a elongate crest, slightly raised and forming 13 round tubercles just before declivity; interstria 5 raised after basal 1/ 8 and forming a row of several small tubercles, more strongly raised at declivity, forming a cone-like tubercle; interstriae 3 and 9 coalescing, raised at declivity forming a V-shaped tubercle on each elytron.
</paragraph>
<paragraph id="4E3A97371002C042FF4EFD18FBA66B64" blockId="14.[151,1437,151,2034]" pageId="14" pageNumber="175">
<emphasis id="7CF14B251002C042FF4EFD18FEA66AA3" box="[201,307,688,712]" italics="true" pageId="14" pageNumber="175">Abdomen</emphasis>
(
<figureCitation id="D6BE8BB21002C042FEC4FD18FE126AA3" box="[323,391,688,712]" captionStart="FIGURE 7" captionStartId="13.[151,250,1904,1927]" captionTargetBox="[302,1315,219,1853]" captionTargetId="figure@13.[274,1322,193,1878]" captionTargetPageId="13" captionText="FIGURE 7. Geotragus rugosus sp. nov.: A male ventrites, ventral view B male metendosternite, dorsal view C male sternite VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, dorsal view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216587/files/figure.png" pageId="14" pageNumber="175">Fig. 7</figureCitation>
A): Surface of ventrite 1 depressed at middle and convex on lateral areas; suture I bisinuate; ventrite 2 moderately convex medially, with apical transverse impression; mesal length of ventrite 1 1.4× length of 2, ventrite 2 1.1× length of 3 and 4 combined; ventrite 5 convex, widely parabolic.
</paragraph>
<paragraph id="4E3A97371002C042FF4EFCB3FE086BA9" blockId="14.[151,1437,151,2034]" pageId="14" pageNumber="175">
<emphasis id="7CF14B251002C042FF4EFCB3FEEA6B58" box="[201,383,795,819]" italics="true" pageId="14" pageNumber="175">Metendosternite</emphasis>
(
<figureCitation id="D6BE8BB21002C042FE09FCB3FE446B58" box="[398,465,795,819]" captionStart="FIGURE 7" captionStartId="13.[151,250,1904,1927]" captionTargetBox="[302,1315,219,1853]" captionTargetId="figure@13.[274,1322,193,1878]" captionTargetPageId="13" captionText="FIGURE 7. Geotragus rugosus sp. nov.: A male ventrites, ventral view B male metendosternite, dorsal view C male sternite VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, dorsal view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216587/files/figure.png" pageId="14" pageNumber="175">Fig. 7</figureCitation>
B): Stalk 0.4× length of furcal arms, trapezoidal and slightly transverse, anterior part of longitudinal flange very short, only 1/5 as long as posterior; hemiductus weak, not very clear, wide and truncate; anterior tendons positioned almost in the middle of furcal arms, opposite to hemiductus; furcal arms strong, apically slightly bifurcate, diverging at nearly 60° in relation to longitudinal flange, with an oblique keel near basal part of each furcal arm.
</paragraph>
<paragraph id="4E3A97371002C042FF4EFC67FB3D6C3A" blockId="14.[151,1437,151,2034]" pageId="14" pageNumber="175">
<emphasis id="7CF14B251002C042FF4EFC67FF6B6B8D" box="[201,254,975,998]" italics="true" pageId="14" pageNumber="175">Legs</emphasis>
: Protibiae bent inwards at apical 1/4, inner margin not bisinuate, almost straight, outer margin slightly arcuate inwards; inner margin of protibiae distinctly with 11 sharp and large teeth, each tooth with a spiniform seta just behind it; teeth of meso- and metatibiae sparse and much smaller than protibiae; protibial mucro moderately large, distinct, not covered by setae; tarsi slender; tarsomere 1 much longer and wider than 2.
</paragraph>
<paragraph id="4E3A97371002C042FF4EFBF5FAF26DDD" blockId="14.[151,1437,151,2034]" pageId="14" pageNumber="175">
<emphasis id="7CF14B251002C042FF4EFBF5FDB56C1E" box="[201,544,1117,1141]" italics="true" pageId="14" pageNumber="175">Male genitalia and terminalia</emphasis>
: Sternite VIII (
<figureCitation id="D6BE8BB21002C042FD53FBF4FC8D6C1F" box="[724,792,1116,1140]" captionStart="FIGURE 7" captionStartId="13.[151,250,1904,1927]" captionTargetBox="[302,1315,219,1853]" captionTargetId="figure@13.[274,1322,193,1878]" captionTargetPageId="13" captionText="FIGURE 7. Geotragus rugosus sp. nov.: A male ventrites, ventral view B male metendosternite, dorsal view C male sternite VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, dorsal view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216587/files/figure.png" pageId="14" pageNumber="175">Fig. 7</figureCitation>
C) divided into two hemisternites, transversely oriented, crescent shape; each hemisternite laterally acuminate, anterior margin widely roundly emarginate, posterior margin arc shaped; membrane between sternite VIII and IX with two very small, dot-like sclerotizations. Sternite IX (
<figureCitation id="D6BE8BB21002C042FAE9FB0CFF306C8B" captionStart="FIGURE 7" captionStartId="13.[151,250,1904,1927]" captionTargetBox="[302,1315,219,1853]" captionTargetId="figure@13.[274,1322,193,1878]" captionTargetPageId="13" captionText="FIGURE 7. Geotragus rugosus sp. nov.: A male ventrites, ventral view B male metendosternite, dorsal view C male sternite VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, dorsal view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216587/files/figure.png" pageId="14" pageNumber="175">Fig. 7</figureCitation>
C) with basal plate bilobed, each lobe semi-trapezoidal; spiculum gastrale 0.68× length of penis, more sclerotized at middle, basal 1/2 almost straight, slightly curved anteriorly. Penis (
<figureCitation id="D6BE8BB21002C042FC43FB43FB9F6D68" box="[964,1034,1259,1283]" captionStart="FIGURE 7" captionStartId="13.[151,250,1904,1927]" captionTargetBox="[302,1315,219,1853]" captionTargetId="figure@13.[274,1322,193,1878]" captionTargetPageId="13" captionText="FIGURE 7. Geotragus rugosus sp. nov.: A male ventrites, ventral view B male metendosternite, dorsal view C male sternite VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, dorsal view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216587/files/figure.png" pageId="14" pageNumber="175">Fig. 7</figureCitation>
D, 7E) in dorsal view
<quantity id="897D3AD21002C042FA93FB43FAE96D68" box="[1300,1404,1259,1283]" metricMagnitude="-3" metricUnit="m" metricValue="2.74" pageId="14" pageNumber="175" unit="mm" value="2.74">2.74 mm</quantity>
in length, ca. 4.5× length of wide, temones
<quantity id="897D3AD21002C042FDE4FAA7FD5D6D4C" box="[611,712,1295,1319]" metricMagnitude="-3" metricUnit="m" metricValue="1.37" pageId="14" pageNumber="175" unit="mm" value="1.37">1.37 mm</quantity>
long; lateral margins subparallel at middle, slightly expanded at ostium level, thereafter converging straight but asymmetry, apex obliquely truncated; in lateral view strongly curved, after ostium strongly constricted toward apex, almost flat at apex. Tegmen (
<figureCitation id="D6BE8BB21002C042FBC4FAFFFB136D04" box="[1091,1158,1367,1391]" captionStart="FIGURE 7" captionStartId="13.[151,250,1904,1927]" captionTargetBox="[302,1315,219,1853]" captionTargetId="figure@13.[274,1322,193,1878]" captionTargetPageId="13" captionText="FIGURE 7. Geotragus rugosus sp. nov.: A male ventrites, ventral view B male metendosternite, dorsal view C male sternite VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, dorsal view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216587/files/figure.png" pageId="14" pageNumber="175">Fig. 7</figureCitation>
F) 0.53× length of penis, with ring narrow, parameroid lobes developed, narrowly spaced at base, basal half more sclerotized area triangular; tegminal apodeme slender, forming a Y-shape with basal piece, distal end of apodeme not expanded, rounded.
</paragraph>
<paragraph id="4E3A97371002C042FF4EFA69FB7E6E2E" blockId="14.[151,1437,151,2034]" pageId="14" pageNumber="175">
<emphasis id="7CF14B251002C042FF4EFA69FD9F6DB1" bold="true" box="[201,522,1473,1498]" pageId="14" pageNumber="175">
Variation. Male
<typeStatus id="913E29951002C042FE0AFA6AFD906DB1" box="[397,517,1474,1498]" pageId="14" pageNumber="175" type="paratype">paratypes</typeStatus>
.
</emphasis>
Measurements (in mm): Standard length: 6.207.20; pronotal length: 1.902.45; pronotal width: 1.802.40; elytral length: 4.205.50; elytral width: 2.503.00; rostral length: 1.001.22; rostral width: 0.801.10. Inner margin of protibiae distinctly with 1114 sharp and large teeth. Among examined specimens, there is one male with a darker penis and with the apical margin of the ostium rounded.
</paragraph>
<paragraph id="4E3A97371002C042FF4EF9F9FB406F99" blockId="14.[151,1437,151,2034]" pageId="14" pageNumber="175">
<emphasis id="7CF14B251002C042FF4EF9F9FE306E02" bold="true" box="[201,421,1617,1641]" pageId="14" pageNumber="175">
Female
<typeStatus id="913E29951002C042FEA0F9F9FE356E02" box="[295,416,1617,1641]" pageId="14" pageNumber="175" type="paratype">paratypes</typeStatus>
.
</emphasis>
Measurements (in mm): Standard length: 7.407.80; pronotal length: 2.002.10; pronotal width: 2.302.40; elytral length: 4.905.80; elytral width: 3.503.60; rostral length: 1.201.23; rostral width: 1.111.13. Elytra much wider than male, pronotum transverse, ventrite 5 (
<figureCitation id="D6BE8BB21002C042FC5CF930FB8B6EDB" box="[987,1054,1688,1712]" captionStart="FIGURE 7" captionStartId="13.[151,250,1904,1927]" captionTargetBox="[302,1315,219,1853]" captionTargetId="figure@13.[274,1322,193,1878]" captionTargetPageId="13" captionText="FIGURE 7. Geotragus rugosus sp. nov.: A male ventrites, ventral view B male metendosternite, dorsal view C male sternite VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, dorsal view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216587/files/figure.png" pageId="14" pageNumber="175">Fig. 7</figureCitation>
G) parabolic, slightly shorter than ventrite 2. Sternite VIII (
<figureCitation id="D6BE8BB21002C042FE36F914FE606EBF" box="[433,501,1724,1748]" captionStart="FIGURE 7" captionStartId="13.[151,250,1904,1927]" captionTargetBox="[302,1315,219,1853]" captionTargetId="figure@13.[274,1322,193,1878]" captionTargetPageId="13" captionText="FIGURE 7. Geotragus rugosus sp. nov.: A male ventrites, ventral view B male metendosternite, dorsal view C male sternite VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, dorsal view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216587/files/figure.png" pageId="14" pageNumber="175">Fig. 7</figureCitation>
H) with spiculum ventrale “baseball bat” shape and slightly curved; lamina tightly folded, in lateral view triangular, ventral margin angulated at midpoint, strongly sclerotized. Ovipositor (
<figureCitation id="D6BE8BB21002C042FABAF948FA146E93" box="[1341,1409,1760,1784]" captionStart="FIGURE 7" captionStartId="13.[151,250,1904,1927]" captionTargetBox="[302,1315,219,1853]" captionTargetId="figure@13.[274,1322,193,1878]" captionTargetPageId="13" captionText="FIGURE 7. Geotragus rugosus sp. nov.: A male ventrites, ventral view B male metendosternite, dorsal view C male sternite VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, dorsal view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216587/files/figure.png" pageId="14" pageNumber="175">Fig. 7</figureCitation>
H,
<date id="3A3BB1F71002C042FF10F8ABFF3A6F70" box="[151,175,1795,1819]" pageId="14" pageNumber="175">7I</date>
) with coxites shorter than sternite VIII, cone-like, lateral margins strongly sclerotized, posteriorly slightly narrowed with rounded apex; styli relatively small, cylindrical, with 23 long setae, more sclerotized than coxites. Bursal sclerites (
<figureCitation id="D6BE8BB21002C042FED3F8E3FE026F08" box="[340,407,1867,1891]" captionStart="FIGURE 7" captionStartId="13.[151,250,1904,1927]" captionTargetBox="[302,1315,219,1853]" captionTargetId="figure@13.[274,1322,193,1878]" captionTargetPageId="13" captionText="FIGURE 7. Geotragus rugosus sp. nov.: A male ventrites, ventral view B male metendosternite, dorsal view C male sternite VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, dorsal view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216587/files/figure.png" pageId="14" pageNumber="175">Fig. 7</figureCitation>
J) near the point of union of spermathecal duct and of oviduct, facing each other at middle, each sclerite in lateral view T-shaped with margins curved, dorsal surface of mesal area with many sharp spines. Spermatheca (
<figureCitation id="D6BE8BB21002C042FEBEF83AFEEB6FC1" box="[313,382,1938,1962]" captionStart="FIGURE 7" captionStartId="13.[151,250,1904,1927]" captionTargetBox="[302,1315,219,1853]" captionTargetId="figure@13.[274,1322,193,1878]" captionTargetPageId="13" captionText="FIGURE 7. Geotragus rugosus sp. nov.: A male ventrites, ventral view B male metendosternite, dorsal view C male sternite VIII and IX, dorsal view D penis, dorsal view E penis, lateral view F tegmen, dorsal view G female ventrites, ventral view H sternite VIII and female genitalia, lateral view I coxites and styli, dorsal view J bursal sclerites, dorsal view K spermatheca, lateral view. Scales: 0.5 mm." httpUri="https://zenodo.org/record/216587/files/figure.png" pageId="14" pageNumber="175">Fig. 7</figureCitation>
K) with corpus nearly rounded; cornu slightly elongate, strongly curved as broad V-shaped, apically gradually narrowed with apex rounded; nodulus prominent, curved tube-like, subcontiguous with and strongly angled at about 20° in relation to ramus; ramus trapezoid, prominent with truncate apex.
</paragraph>
</subSubSection>
<subSubSection id="069FC4BC1003C043FF4EFF3FFCC76A04" pageId="15" pageNumber="176" type="materials_examined">
<paragraph id="4E3A97371003C043FF4EFF3FFAC86A48" blockId="15.[151,1437,151,695]" pageId="15" pageNumber="176">
<emphasis id="7CF14B251003C043FF4EFF3FFDAE68DB" bold="true" box="[201,571,151,176]" pageId="15" pageNumber="176">
Material examined.
<typeStatus id="913E29951003C043FE42FF30FDA368DB" box="[453,566,152,176]" pageId="15" pageNumber="176" type="holotype">Holotype</typeStatus>
:
</emphasis>
3: (white, printed):
<collectingCountry id="3692D7A71003C043FCBBFF3FFC0268C4" box="[828,919,151,175]" name="China" pageId="15" pageNumber="176">CHINA</collectingCountry>
, Yunnan Provin. / Gongshan, Cikai town / Dabadi, riverside. /
<geoCoordinate id="2BB1F1F01003C043FEF1FF14FD9768BF" box="[374,514,188,212]" direction="north" orientation="latitude" pageId="15" pageNumber="176" precision="15" value="27.796667">N27°4748”</geoCoordinate>
,
<geoCoordinate id="2BB1F1F01003C043FD89FF14FD0368BF" box="[526,662,188,212]" direction="east" orientation="longitude" pageId="15" pageNumber="176" precision="15" value="98.50584">E98°3021”</geoCoordinate>
;
<quantity id="897D3AD21003C043FD24FF14FD6668BF" box="[675,755,188,212]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="176" unit="m" value="3000.0">3000m</quantity>
, 2002.9.30, day /
<collectionCode id="28940FF21003C043FC3BFF14FB8668BF" box="[956,1043,188,212]" pageId="15" pageNumber="176">LIANG</collectionCode>
Hongbin. / Sino-American Exped. / Institute Zoology,
<collectionCode id="28940FF21003C043FEF2FF77FE3A689C" LSID="urn:lsid:biocol.org:col:34700" box="[373,431,223,247]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="15" pageNumber="176">CAS</collectionCode>
; (red, printed):
<typeStatus id="913E29951003C043FDD9FF77FD63689C" box="[606,758,223,247]" pageId="15" pageNumber="176" type="holotype">HOLOTYPE</typeStatus>
; (white, printed): IOZ(E) 905051.
<typeStatus id="913E29951003C043FBFBFF48FB616893" box="[1148,1268,224,248]" pageId="15" pageNumber="176" type="paratype">
<emphasis id="7CF14B251003C043FBFBFF48FB616893" bold="true" box="[1148,1268,224,248]" pageId="15" pageNumber="176">Paratypes</emphasis>
</typeStatus>
(43, 5Ƥ): 1Ƥ: same data as
<typeStatus id="913E29951003C043FEADFEACFE196977" box="[298,396,260,284]" pageId="15" pageNumber="176" type="holotype">holotype</typeStatus>
except
<typeStatus id="913E29951003C043FE64FEACFDFB6977" box="[483,622,260,284]" pageId="15" pageNumber="176" type="paratype">PARATYPE</typeStatus>
printed on yellow paper and IOZ(E) 905053. 13: with same locality and elevation as
<typeStatus id="913E29951003C043FEAAFE8FFE1A6954" box="[301,399,295,319]" pageId="15" pageNumber="176" type="holotype">holotype</typeStatus>
except collecting date 2002.9.27 night, light,
<typeStatus id="913E29951003C043FC31FE80FBD4692B" box="[950,1089,296,320]" pageId="15" pageNumber="176" type="paratype">PARATYPE</typeStatus>
printed on yellow paper and IOZ(E) 905052. 33: (white, printed):
<collectingCountry id="3692D7A71003C043FDEEFEE4FD51690F" box="[617,708,332,356]" name="China" pageId="15" pageNumber="176">CHINA</collectingCountry>
, Yunnan Provin., / Gongshan, Dulongjiang, / Maku. Wood, vegetation /
<geoCoordinate id="2BB1F1F01003C043FEA0FEC7FE2669EC" box="[295,435,367,391]" direction="north" orientation="latitude" pageId="15" pageNumber="176" precision="1" value="27.68875">27.68875°N</geoCoordinate>
,
<geoCoordinate id="2BB1F1F01003C043FE46FEC7FDDF69EC" box="[449,586,367,391]" direction="east" orientation="longitude" pageId="15" pageNumber="176" precision="1" value="98.31303">98.31303°E</geoCoordinate>
;
<quantity id="897D3AD21003C043FDDFFEC7FD3C69EC" box="[600,681,367,391]" metricMagnitude="3" metricUnit="m" metricValue="1.677" pageId="15" pageNumber="176" unit="m" value="1677.0">1677m</quantity>
, 2006.9.3 day / Liu Ye collector / California Academy &amp; / IOZ., Chinese. Acad. Sci.; (yellow, printed):
<typeStatus id="913E29951003C043FDD3FE3CFD7669C7" box="[596,739,404,428]" pageId="15" pageNumber="176" type="paratype">PARATYPE</typeStatus>
; (white, printed): IOZ(E) 1798744, IOZ(E) 1798745, IOZ(E) 1798747. 2Ƥ: ditto, IOZ(E) 1798746, IOZ(E) 1798748. 2Ƥ: (white, printed): [Yúnnán Gòngshān Dúlóngjiāng Xiāng] /! &quot;# [Mǐlǐwáng Lùbiān Wǎnshàng] / 2004.
<date id="3A3BB1F71003C043FB29FE4BFB766990" box="[1198,1251,483,507]" pageId="15" pageNumber="176" value="1931-10">X.31</date>
$%&amp; [Liáng Hóngbīn] / [
<collectingCountry id="3692D7A71003C043FE37FDA2FDB36A49" box="[432,550,522,546]" name="China" pageId="15" pageNumber="176">Zhōngguó</collectingCountry>
Kēxuéyuàn];
<quantity id="897D3AD21003C043FD44FDA3FC846A48" box="[707,785,523,547]" metricMagnitude="3" metricUnit="m" metricValue="1.956" pageId="15" pageNumber="176" unit="m" value="1956.0">1956m</quantity>
/ '([Wĕidù] 27.72383 /)([Jīngdù] 98.36117 /
</paragraph>
<paragraph id="4E3A97371003C043FF75FD9AFA086A21" blockId="15.[151,1437,151,695]" box="[242,1437,562,587]" pageId="15" pageNumber="176">
[
<collectingCountry id="3692D7A71003C043FF7FFD9AFEFB6A21" box="[248,366,562,586]" name="China" pageId="15" pageNumber="176">Zhōngguó</collectingCountry>
Kēxuéyuàn]; (yellow, printed):
<typeStatus id="913E29951003C043FD51FD9BFCF06A20" box="[726,869,563,587]" pageId="15" pageNumber="176" type="paratype">PARATYPE</typeStatus>
; (white, printed): IOZ(E) 905050, IOZ(E) 905054.
</paragraph>
<paragraph id="4E3A97371003C043FF4EFDFFFCC76A04" blockId="15.[151,1437,151,695]" box="[201,850,599,624]" pageId="15" pageNumber="176">
<typeStatus id="913E29951003C043FF4EFDFFFEA46A04" box="[201,305,599,623]" pageId="15" pageNumber="176" type="holotype">Holotype</typeStatus>
and all
<typeStatus id="913E29951003C043FE0DFDF0FE606A1B" box="[394,501,600,624]" pageId="15" pageNumber="176" type="paratype">paratypes</typeStatus>
are to be conserved in
<collectionCode id="28940FF21003C043FD7EFDFFFCDB6A04" box="[761,846,599,623]" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="15" pageNumber="176">IZCAS</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="069FC4BC1003C043FF4EFDD2FA936ADC" pageId="15" pageNumber="176" type="distribution">
<paragraph id="4E3A97371003C043FF4EFDD2FA936ADC" blockId="15.[151,1437,151,695]" pageId="15" pageNumber="176">
<emphasis id="7CF14B251003C043FF4EFDD2FEF76AF8" bold="true" box="[201,354,634,659]" pageId="15" pageNumber="176">Distribution.</emphasis>
<collectingCountry id="3692D7A71003C043FEEEFDD2FE256AF9" box="[361,432,634,658]" name="China" pageId="15" pageNumber="176">China</collectingCountry>
: Yunnan (Gongshan). Altitudinal range of this species is from
<quantity id="897D3AD21003C043FBECFDD2FB2C6AF9" box="[1131,1209,634,658]" metricMagnitude="3" metricUnit="m" metricValue="1.677" pageId="15" pageNumber="176" unit="m" value="1677.0">1677m</quantity>
to
<quantity id="897D3AD21003C043FB5AFDD2FABF6AF9" box="[1245,1322,634,658]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="176" unit="m" value="3000.0">3000m</quantity>
high.
<emphasis id="7CF14B251003C043FF40FD37FEDB6ADC" bold="true" box="[199,334,671,695]" pageId="15" pageNumber="176">Etymology.</emphasis>
The specific epithet is referring to the strongly corrugated dorsal part of the rostrum.
</paragraph>
</subSubSection>
</treatment>
</document>