treatments-xml/data/C6/1B/4E/C61B4E432563FFCAFF31FBF02A5CF687.xml

<document id="B3AEB0D809F3C3CB3F51EC2134A46286" ID-CLB-Dataset="24557" ID-DOI="10.11646/zootaxa.4712.1.3" ID-GBIF-Dataset="5557b2c6-99ea-4be4-bcfa-eb014aaa8932" ID-ISSN="1175-5326" ID-Zenodo-Dep="3586826" ID-ZooBank="C831FB1F-CA1D-4AF8-9130-6649D151ACE3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1576824226905" checkinUser="plazi" docAuthor="Bui, Van Bac, Ziegler, Thomas &amp; Bonkowski, Michael" docDate="2019" docId="C61B4E432563FFCAFF31FBF02A5CF687" docLanguage="en" docName="zootaxa.4712.1.3.pdf" docOrigin="Zootaxa 4712 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Copris (Paracopris) javanensis Bui, Ziegler &amp; Bonkowski 2019, new species" docType="treatment" docVersion="8" lastPageNumber="57" masterDocId="3A22363B2560FFCCFFA6FFC62E6DF138" masterDocTitle="Checklist of beetles in the subgenus Copris (Paracopris) Balthasar from Asia with description of a new species, and redescription of Copris (Paracopris) punctulatus Wiedemann (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="64" masterPageNumber="51" pageNumber="54" updateTime="1698775412311" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="78C40FF04C80BE484C09AAD652925A7A">Checklist of beetles in the subgenus Copris (Paracopris) Balthasar from Asia with description of a new species, and redescription of Copris (Paracopris) punctulatus Wiedemann (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="EF1582D3EE0A1BD529B622DCA80EB3E2">Bui, Van Bac</mods:namePart>
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<mods:namePart id="E8C30BEB01E9B3851C39E1E6027D622A">Ziegler, Thomas</mods:namePart>
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<mods:namePart id="98F4D74FBDCB6D1CF4D70509FE81AF3A">Bonkowski, Michael</mods:namePart>
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<mods:title id="0283145E209CD91121CC4A9CE7E334A4">Zootaxa</mods:title>
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<mods:date id="0AC175EDAF555DE41C029C170C4EF78C">2019</mods:date>
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<mods:number id="065EEE1259783E2E084CFDAD6520E164">2019-12-18</mods:number>
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<paragraph id="4E0DFF552563FFCFFF31FBF02D95F568" blockId="3.[151,1016,1078,1140]" box="[151,1016,1078,1105]" pageId="3" pageNumber="54">
<heading id="154548392563FFCFFF31FBF02D95F568" bold="true" box="[151,1016,1078,1105]" fontSize="11" level="1" pageId="3" pageNumber="54" reason="1">
<emphasis id="7CC623472563FFCFFF31FBF02D95F568" bold="true" box="[151,1016,1078,1105]" pageId="3" pageNumber="54">
<taxonomicName id="89B284D62563FFCFFF31FBF02D36F568" ID-CoL="Y5QH" authority="Bui, Ziegler &amp; Bonkowski" authorityName="Bui, Ziegler &amp; Bonkowski" authorityYear="2019" box="[151,859,1078,1104]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="javanensis" status="sp. nov." subGenus="Paracopris">
<emphasis id="7CC623472563FFCFFF31FBF02E8AF568" bold="true" box="[151,231,1078,1104]" italics="true" pageId="3" pageNumber="54">Copris</emphasis>
(
<emphasis id="7CC623472563FFCFFF5EFBF02F11F568" bold="true" box="[248,380,1078,1104]" italics="true" pageId="3" pageNumber="54">Paracopris</emphasis>
) 
<emphasis id="7CC623472563FFCFFE2BFBF02C61F568" bold="true" box="[397,524,1078,1104]" italics="true" pageId="3" pageNumber="54">javanensis</emphasis>
Bui, Ziegler &amp; Bonkowski
</taxonomicName>
, 
<taxonomicNameLabel id="67F59E3C2563FFCFFCCEFBF12D95F568" box="[872,1016,1078,1105]" pageId="3" pageNumber="54" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<paragraph id="4E0DFF552563FFCFFF31FB9C2FA1F54C" blockId="3.[151,1016,1078,1140]" box="[151,460,1114,1140]" pageId="3" pageNumber="54">
(
<figureCitation id="D689E3D02563FFCFFF39FB9C2E85F54C" box="[159,232,1114,1140]" captionStart="FIGURES 1" captionStartId="5.[151,264,1965,1990]" captionTargetBox="[215,1374,179,1936]" captionTargetId="figure@5.[206,1381,166,1945]" captionTargetPageId="5" captionText="FIGURES 1A–D. Holotype (A, B) and female paratype (C, D) of Copris (Paracopris) javanensis Bui, Ziegler &amp; Bonkowski, new species. A, C, habitus, dorsal view; B, D, habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3586828" httpUri="https://zenodo.org/record/3586828/files/figure.png" pageId="3" pageNumber="54">Figs. 1</figureCitation>
A–D, 2A, C–E, 3A)
</paragraph>
</subSubSection>
<subSubSection id="06A8ACDE2563FFCFFF31FB642DD6F43C" pageId="3" pageNumber="54" type="materials_examined">
<paragraph id="4E0DFF552563FFCFFF31FB642C13F584" blockId="3.[151,1437,1185,2041]" box="[151,638,1185,1213]" pageId="3" pageNumber="54">
<materialsCitation id="FEDAF5082563FFCFFF31FB642C16F584" ID-GBIF-Occurrence="2521263207" box="[151,635,1185,1213]" country="Indonesia" location="Mt. Moeria" pageId="3" pageNumber="54" specimenCount="1" stateProvince="Jawa Timur" typeStatus="holotype">
<emphasis id="7CC623472563FFCFFF31FB642F5BF584" bold="true" box="[151,310,1185,1212]" pageId="3" pageNumber="54">
<typeStatus id="910941F72563FFCFFF31FB642EBFF584" box="[151,210,1186,1212]" pageId="3" pageNumber="54">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="36A5BFC52563FFCFFE9AFB642FC1F584" box="[316,428,1186,1212]" name="Indonesia" pageId="3" pageNumber="54">Indonesia</collectingCountry>
, 
<collectingRegion id="8C7631B72563FFCFFE11FB652F86F585" box="[439,491,1187,1213]" country="Indonesia" name="Jawa Timur" pageId="3" pageNumber="54">Java</collectingRegion>
, 
<location id="4B6DA98E2563FFCFFE51FB652C16F584" LSID="urn:lsid:plazi:treatment:C61B4E432563FFCAFF31FBF02A5CF687:4B6DA98E2563FFCFFE51FB652C16F584" box="[503,635,1186,1213]" country="Indonesia" name="Mt. Moeria" pageId="3" pageNumber="54" stateProvince="Jawa Timur">Mt. Moeria</location>
</materialsCitation>
.
</paragraph>
<paragraph id="4E0DFF552563FFCFFF61FB002DD6F43C" blockId="3.[151,1437,1185,2041]" pageId="3" pageNumber="54">
<emphasis id="7CC623472563FFCFFF61FB002FECF5D8" bold="true" box="[199,385,1221,1248]" pageId="3" pageNumber="54">Type specimen.</emphasis>
<materialsCitation id="FEDAF5082563FFCFFE2CFB002BF4F5D8" ID-GBIF-Occurrence="2521263204" box="[394,1433,1222,1249]" collectionCode="MNHN, R" country="Indonesia" location="Oberthur" municipality="Mt. Moeria" pageId="3" pageNumber="54" specimenCount="1" stateProvince="Jawa Timur" typeStatus="holotype">
<emphasis id="7CC623472563FFCFFE2CFB002C6DF5D8" bold="true" box="[394,512,1222,1248]" pageId="3" pageNumber="54">
<typeStatus id="910941F72563FFCFFE2CFB002F96F5D8" box="[394,507,1222,1248]" pageId="3" pageNumber="54" type="holotype">Holotype</typeStatus>
:
</emphasis>
male (
<collectionCode id="28A367902563FFCFFDF7FB002CC1F5D8" box="[593,684,1222,1248]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="54">MNHN</collectionCode>
), “ 
<collectingCountry id="36A5BFC52563FFCFFD6AFB002D53F5D8" box="[716,830,1222,1248]" name="Indonesia" pageId="3" pageNumber="54">Indonesia</collectingCountry>
, 
<collectingRegion id="8C7631B72563FFCFFCEAFB012D13F5D9" box="[844,894,1223,1249]" country="Indonesia" name="Jawa Timur" pageId="3" pageNumber="54">Java</collectingRegion>
| 
<collectingMunicipality id="AE69652F2563FFCFFC30FB012A76F5D8" box="[918,1051,1222,1249]" pageId="3" pageNumber="54">Mt. Moeria</collectingMunicipality>
, 3–4000’ | ex Coll. 
<collectionCode id="28A367902563FFCFFAA7FB012B7AF5D9" box="[1281,1303,1223,1249]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="54">R</collectionCode>
. 
<location id="4B6DA98E2563FFCFFA82FB002BFDF5D8" LSID="urn:lsid:plazi:treatment:C61B4E432563FFCAFF31FBF02A5CF687:4B6DA98E2563FFCFFA82FB002BFDF5D8" box="[1316,1424,1222,1248]" country="Indonesia" municipality="Mt. Moeria" name="Oberthur" pageId="3" pageNumber="54" stateProvince="Jawa Timur">Oberthür</location>
”
</materialsCitation>
. 
<materialsCitation id="FEDAF5082563FFCFFF31FB2C2DDAF43C" ID-GBIF-Occurrence="2521263206" box="[151,951,1258,1285]" collectionCode="MNHN" country="Indonesia" location="Oberthur" municipality="Mt. Moeria" pageId="3" pageNumber="54" specimenCount="11" specimenCount-female="7" specimenCount-male="4" stateProvince="Jawa Timur" typeStatus="paratype">
<emphasis id="7CC623472563FFCFFF31FB2C2F75F43C" bold="true" box="[151,280,1258,1284]" pageId="3" pageNumber="54">
<typeStatus id="910941F72563FFCFFF31FB2C2F7EF43C" box="[151,275,1258,1284]" pageId="3" pageNumber="54" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="58B434DC2563FFCFFEB9FB2C2F18F43C" box="[287,373,1258,1284]" pageId="3" pageNumber="54" type="male">4 males</specimenCount>
and 
<specimenCount id="58B434DC2563FFCFFE0AFB2D2C75F43C" box="[428,536,1258,1285]" pageId="3" pageNumber="54" type="female">7 females</specimenCount>
(
<collectionCode id="28A367902563FFCFFD81FB2C2CEFF43C" box="[551,642,1258,1284]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="54">MNHN</collectionCode>
), same data as for holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="06A8ACDE2563FFC8FF61FACB2ADCF267" lastPageId="4" lastPageNumber="55" pageId="3" pageNumber="54" type="description">
<paragraph id="4E0DFF552563FFCFFF61FACB2B39F474" blockId="3.[151,1437,1185,2041]" pageId="3" pageNumber="54">
<emphasis id="7CC623472563FFCFFF61FACB2F87F410" bold="true" box="[199,490,1293,1320]" pageId="3" pageNumber="54">
Description of 
<typeStatus id="910941F72563FFCFFEDAFAC82F88F410" box="[380,485,1294,1320]" pageId="3" pageNumber="54" type="holotype">holotype</typeStatus>
.
</emphasis>
Male. Total body length 
<quantity id="894A52B02563FFCFFCB7FACB2D17F411" box="[785,890,1293,1321]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="3" pageNumber="54" unit="mm" value="15.5">15.5 mm</quantity>
, maximum body width 
<quantity id="894A52B02563FFCFFB29FACB2A87F411" box="[1167,1258,1293,1321]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="54" unit="mm" value="7.5">7.5 mm</quantity>
. Whole surface chocolate brown. Mouthparts and antennomeres 1–6 reddish brown. Antennal club and setae on legs yellow.
</paragraph>
<paragraph id="4E0DFF552563FFCFFF61FA902B22F738" blockId="3.[151,1437,1185,2041]" pageId="3" pageNumber="54">
Head 
<quantity id="894A52B02563FFCFFEACFA902F0CF449" box="[266,353,1366,1393]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="3" pageNumber="54" unit="mm" value="4.2">4.2 mm</quantity>
long and 5.0 mm wide. Clypeal surface smooth and impunctate; clypeal apex strongly upwards protruded, and widely and shallowly emarginate. Genae rectangular, distinctly separated from clypeus by well-defined suture; most surface of genae smooth, except for small areas in middle sparsely and weakly punctate. Surface of vertex distinctly punctate; punctures denser and deeper at areas near eyes. Cephalic horn small but distinct. Antenna with 9 antennomeres; antennomere 1 longer than antennomeres 2–6 combined and also antennal club.
</paragraph>
<paragraph id="4E0DFF552563FFCFFF61F9CD2B77F7A8" blockId="3.[151,1437,1185,2041]" pageId="3" pageNumber="54">
Pronotum 
<quantity id="894A52B02563FFCFFE9BF9CF2FFEF71D" box="[317,403,1545,1573]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="3" pageNumber="54" unit="mm" value="5.2">5.2 mm</quantity>
long and 
<quantity id="894A52B02563FFCFFDA4F9CC2C34F71D" box="[514,601,1546,1573]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="3" pageNumber="54" unit="mm" value="6.8">6.8 mm</quantity>
wide, and without anterior declivity. Pronotal anterior margin slightly forwards curved in middle; anterolateral angles obtuse. Pronotal disc with weak median sulcus; sulcus not extending to pronotal anterior margin. Whole surface of pronotum deeply and densely punctate; most punctures equal in size, except for punctures near sulcus slightly larger. Each pronotal side with one deep and large excavation.
</paragraph>
<paragraph id="4E0DFF552563FFCFFF61F95C2AE4F78C" blockId="3.[151,1437,1185,2041]" box="[199,1161,1690,1717]" pageId="3" pageNumber="54">Anterior angles of prothorax flat, and shallowly and sparsely punctate on ventral side.</paragraph>
<paragraph id="4E0DFF552563FFCFFF61F9782C94F618" blockId="3.[151,1437,1185,2041]" pageId="3" pageNumber="54">
Elytra (ElyL 
<quantity id="894A52B02563FFCFFEFFF97B2FDDF7E1" box="[345,432,1725,1753]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="3" pageNumber="54" unit="mm" value="8.5">8.5 mm</quantity>
) deeply striate; striae weakly, sparsely and equally punctate. Elytral intervals convex, and deeply and densely punctate; interval punctures evenly distributed and relatively equal in size. Each elytron with one rather abrupt vertical flattening near sutural angle.
</paragraph>
<paragraph id="4E0DFF552563FFCFFF61F8EC2EBAF6B5" blockId="3.[151,1437,1185,2041]" pageId="3" pageNumber="54">Mesepimeron and metepisternum flat and weakly punctate. Mesosternum strongly rugose. Metasternum relatively smooth, and with median distinct groove; sides of metasternum sparsely and shallowly punctate, and without setae.</paragraph>
<paragraph id="4E0DFF552563FFC8FF61F8512FBDF0EB" blockId="3.[151,1437,1185,2041]" lastBlockId="4.[151,1437,153,1007]" lastPageId="4" lastPageNumber="55" pageId="3" pageNumber="54">Legs. Ventral surface of profemora strongly punctate; punctures unevenly distributed, and unequal in size, becoming denser and smaller towards anterior margin of profemora; a few punctures at posterior half of profemora with yellow and long setae. Anterior margin of profemora with yellow, dense and long setae. Protibia with four lateral teeth, and protibial surface strongly and unevenly punctate. Protibial spurs pointed, slightly curved outwards near apex, and equal in length to protarsomeres 2–4 combined. Protarsomere 1 and 5 equal in length, and both slightly longer than individual protarsomere 2–4. Mesofemora smooth and impunctate on ventral side. Mesotibia smooth and broadly dilated at posterior end; mesotibia with two spurs, one of which blunt, one longer and pointed. Mesotarsus with five tarsomeres, and all mesotarsomeres with yellow scanty setae; basal mesotarsomere elongated, almost equal in length to mesotibial spurs. Most ventral surface of metafemora smooth and impunctate, except for apical third sparsely punctate. Metatibia smooth and broadly dilated at posterior end, and with one spur pointed. Metatarsus with five tarsomeres covered with yellow scantly setae; basal metatarsomere apically enlarged, and shorter than metatibial spur.</paragraph>
<paragraph id="4E0DFF552564FFC8FF61FE1B2FEAF307" blockId="4.[151,1437,153,1007]" pageId="4" pageNumber="55">
Abdomen and pygidium. Abdominal ventrites smooth, impunctate, and narrower at midline. Pygidium (PyL 
<quantity id="894A52B02564FFC8FADFFE1A2EABF323" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="55" unit="mm" value="1.5">1.5 mm</quantity>
; PyW 
<quantity id="894A52B02564FFC8FEB4FDC72F04F323" box="[274,361,513,539]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="4" pageNumber="55" unit="mm" value="3.1">3.1 mm</quantity>
) with margin complete; pygidial surface slightly convex, and deeply punctate; punctures dense but clearly separated.
</paragraph>
<paragraph id="4E0DFF552564FFC8FF61FD8F2A7FF3BF" blockId="4.[151,1437,153,1007]" pageId="4" pageNumber="55">Aedeagus. Phallobase slightly longer and wider than parameres. A broad angle almost reaching 180º formed by phallobase and parameres. Parameres slightly curved downwards in apical part.</paragraph>
<paragraph id="4E0DFF552564FFC8FF61FD562A16F3CB" blockId="4.[151,1437,153,1007]" pageId="4" pageNumber="55">
<emphasis id="7CC623472564FFC8FF61FD562FDAF393" bold="true" box="[199,439,656,683]" pageId="4" pageNumber="55">Sexual dimorphism.</emphasis>
Female 
<typeStatus id="910941F72564FFC8FDBEFD572CEEF393" box="[536,643,657,683]" pageId="4" pageNumber="55" type="paratype">paratypes</typeStatus>
differ from the male 
<typeStatus id="910941F72564FFC8FCD3FD572DBAF393" box="[885,983,657,683]" pageId="4" pageNumber="55" type="holotype">holotype</typeStatus>
in the following characters: Clypeal anterior margin not protruded in middle, and without emargination. Head with one slightly curved transverse carina between clypeus and frons. Pronotal anterior margin strongly forwards curved in middle.
</paragraph>
<paragraph id="4E0DFF552564FFC8FF61FD3A2ADCF267" blockId="4.[151,1437,153,1007]" pageId="4" pageNumber="55">
<emphasis id="7CC623472564FFC8FF61FD3A2F2AF22F" bold="true" box="[199,327,764,791]" pageId="4" pageNumber="55">Variability</emphasis>
. Body length 
<quantity id="894A52B02564FFC8FE4EFD3A2CE1F22F" box="[488,652,764,791]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.65" metricValueMin="1.15" pageId="4" pageNumber="55" unit="mm" value="14.0" valueMax="16.5" valueMin="11.5">11.5–16.5 mm</quantity>
, body width 6.0– 
<quantity id="894A52B02564FFC8FCF7FD3A2DCAF22F" box="[849,935,764,791]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="55" unit="mm" value="7.5">7.5 mm</quantity>
(n=12). The two males are larger, compared to the 
<typeStatus id="910941F72564FFC8FF79FCE72F2CF203" box="[223,321,801,827]" pageId="4" pageNumber="55" type="holotype">holotype</typeStatus>
possess a more strongly protruded clypeus upwards but similar in the remaining characters. There are no significant differences in external morphology among the female specimens examined.
</paragraph>
</subSubSection>
<subSubSection id="06A8ACDE2564FFC8FF61FCAE2C82F640" pageId="4" pageNumber="55" type="diagnosis">
<paragraph id="4E0DFF552564FFC8FF61FCAE2C37F2D7" blockId="4.[151,1437,153,1007]" pageId="4" pageNumber="55">
<emphasis id="7CC623472564FFC8FF61FCAE2FA3F2BB" bold="true" box="[199,462,872,899]" pageId="4" pageNumber="55">Differential diagnosis.</emphasis>
The new species strongly resembles 
<taxonomicName id="89B284D62564FFC8FCC8FCAF2AB1F2BB" authorityName="Wiedemann" authorityYear="1823" box="[878,1244,873,899]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" sensu="lato" species="punctulatus" subGenus="Paracopris">
<emphasis id="7CC623472564FFC8FCC8FCAF2DD7F2BB" box="[878,954,873,899]" italics="true" pageId="4" pageNumber="55">Copris</emphasis>
(
<emphasis id="7CC623472564FFC8FC6CFCAC2A25F2BB" box="[970,1096,874,899]" italics="true" pageId="4" pageNumber="55">Paracopris</emphasis>
) 
<emphasis id="7CC623472564FFC8FBFFFCAF2AB1F2BB" box="[1113,1244,873,899]" italics="true" pageId="4" pageNumber="55">punctulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67F59E3C2564FFC8FB44FCAF2B3FF2BB" box="[1250,1362,873,899]" pageId="4" pageNumber="55" sensu="lato">sensu lato</taxonomicNameLabel>
that is widely distributed in Asia: 
<collectingCountry id="36A5BFC52564FFC8FE6DFC4B2C40F29F" box="[459,557,909,935]" name="Vietnam" pageId="4" pageNumber="55">Vietnam</collectingCountry>
, 
<collectingCountry id="36A5BFC52564FFC8FD9FFC4B2C1FF29F" box="[569,626,909,935]" name="Laos" pageId="4" pageNumber="55">Laos</collectingCountry>
, 
<collectingCountry id="36A5BFC52564FFC8FDD8FC4B2C99F29F" box="[638,756,909,935]" name="Cambodia" pageId="4" pageNumber="55">Cambodia</collectingCountry>
, 
<collectingCountry id="36A5BFC52564FFC8FCA6FC4B2D0BF29F" box="[768,870,909,935]" name="Thailand" pageId="4" pageNumber="55">Thailand</collectingCountry>
, 
<collectingCountry id="36A5BFC52564FFC8FCD5FC4B2D8EF29F" box="[883,995,909,935]" name="Indonesia" pageId="4" pageNumber="55">Indonesia</collectingCountry>
, 
<collectingCountry id="36A5BFC52564FFC8FC49FC4B2A35F29F" box="[1007,1112,909,935]" name="Malaysia" pageId="4" pageNumber="55">Malaysia</collectingCountry>
, 
<collectingCountry id="36A5BFC52564FFC8FBC3FC4B2AB9F29F" box="[1125,1236,909,935]" name="Myanmar" pageId="4" pageNumber="55">Myanmar</collectingCountry>
, 
<collectingCountry id="36A5BFC52564FFC8FB46FC4B2B4EF29F" box="[1248,1315,909,935]" name="China" pageId="4" pageNumber="55">China</collectingCountry>
and 
<collectingCountry id="36A5BFC52564FFC8FAFAFC4B2BF5F29F" box="[1372,1432,909,935]" name="India" pageId="4" pageNumber="55">India</collectingCountry>
, but has a consistently different pygidial surface and anterior pronotal margin (see detailed diagnostic characters separating these two species in 
<tableCitation id="0330CAEE2564FFC8FE51FC132C24F2D7" box="[503,585,981,1007]" captionStart="TABLE 1" captionStartId="4.[151,239,1054,1078]" captionTargetBox="[166,1420,1126,1518]" captionText="TABLE 1. Diagnostic characters separating the new species from Copris (Paracopris) punctulatus sensu lato (see also Figs. 2A–F)." httpUri="http://table.plazi.org/id/1ACDAFDD2564FFC8FF31FBD82F72F563" pageId="4" pageNumber="55" tableUuid="1ACDAFDD2564FFC8FF31FBD82F72F563">Table 1</tableCitation>
).
</paragraph>
<caption id="1ACDAFDD2564FFC8FF31FBD82F72F563" ID-Table-UUID="1ACDAFDD2564FFC8FF31FBD82F72F563" httpUri="http://table.plazi.org/id/1ACDAFDD2564FFC8FF31FBD82F72F563" pageId="4" pageNumber="55" targetBox="[166,1420,1126,1518]" targetIsTable="true" targetPageId="4">
<paragraph id="4E0DFF552564FFC8FF31FBD82F72F563" blockId="4.[151,1436,1054,1115]" pageId="4" pageNumber="55">
<emphasis id="7CC623472564FFC8FF31FBD82F66F50E" bold="true" box="[151,267,1054,1078]" pageId="4" pageNumber="55">TABLE 1.</emphasis>
Diagnostic characters separating the new species from 
<taxonomicName id="89B284D62564FFC8FCC3FBD92AAEF50F" authorityName="Wiedemann" authorityYear="1823" box="[869,1219,1055,1079]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" sensu="lato" species="punctulatus" subGenus="Paracopris">
<emphasis id="7CC623472564FFC8FCC3FBD92DC3F50E" box="[869,942,1055,1078]" italics="true" pageId="4" pageNumber="55">Copris</emphasis>
(
<emphasis id="7CC623472564FFC8FC19FBD92A5AF50E" box="[959,1079,1055,1078]" italics="true" pageId="4" pageNumber="55">Paracopris</emphasis>
) 
<emphasis id="7CC623472564FFC8FBE1FBD92AAEF50F" box="[1095,1219,1055,1079]" italics="true" pageId="4" pageNumber="55">punctulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67F59E3C2564FFC8FB6AFBD92B5AF50F" box="[1228,1335,1055,1079]" pageId="4" pageNumber="55" sensu="lato">sensu lato</taxonomicNameLabel>
(see also Figs. 2A–F).
</paragraph>
</caption>
<paragraph id="4E0DFF552564FFC8FF00FBA02A53F4D6" pageId="4" pageNumber="55">
<table id="3CB20DF525640033FF00FBA02BE1F4D6" box="[166,1420,1126,1518]" gridcols="3" gridrows="12" pageId="4" pageNumber="55">
<tr id="F082FD1725640033FF00FBA02BE1F547" box="[166,1420,1126,1151]" gridrow="0" pageId="4" pageNumber="55">
<th id="B353946B25640033FF00FBA02C43F547" box="[166,558,1126,1151]" gridcol="0" gridrow="0" pageId="4" pageNumber="55">Characters</th>
<th id="B353946B25640033FDE9FBA02DAFF547" box="[591,962,1126,1151]" gridcol="1" gridrow="0" pageId="4" pageNumber="55">
<taxonomicName id="89B284D62564FFC8FDE9FBA12D09F547" authority="Bui &amp; Ziegler &amp; Bonkowski, 2019" authorityName="new" authorityYear="2019" box="[591,868,1126,1151]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="javanensis" status="new" subGenus="Paracopris">
<emphasis id="7CC623472564FFC8FDE9FBA12CF9F547" bold="true" box="[591,660,1127,1151]" italics="true" pageId="4" pageNumber="55">Copris</emphasis>
(
<emphasis id="7CC623472564FFC8FD04FBA12CD8F546" bold="true" box="[674,693,1127,1150]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
) 
<emphasis id="7CC623472564FFC8FD62FBA12D5FF547" bold="true" box="[708,818,1127,1151]" italics="true" pageId="4" pageNumber="55">javanensis</emphasis>
<taxonomicNameLabel id="67F59E3C2564FFC8FC9EFBA12D09F547" box="[824,868,1127,1151]" pageId="4" pageNumber="55" rank="species">new</taxonomicNameLabel>
</taxonomicName>
species
</th>
<th id="B353946B25640033FC42FBA02BE1F547" box="[996,1420,1126,1151]" gridcol="2" gridrow="0" pageId="4" pageNumber="55">
<emphasis id="7CC623472564FFC8FC42FBA12B75F547" bold="true" box="[996,1304,1126,1151]" pageId="4" pageNumber="55">
<taxonomicName id="89B284D62564FFC8FC42FBA12AC5F547" authorityName="Wiedemann" authorityYear="1823" box="[996,1192,1126,1151]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" sensu="lato" species="punctulatus" subGenus="Paracopris">
<emphasis id="7CC623472564FFC8FC42FBA12D96F547" bold="true" box="[996,1019,1127,1151]" italics="true" pageId="4" pageNumber="55">C.</emphasis>
(
<emphasis id="7CC623472564FFC8FBACFBA12A70F546" bold="true" box="[1034,1053,1127,1150]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
) 
<emphasis id="7CC623472564FFC8FB8AFBA02AC5F547" bold="true" box="[1068,1192,1126,1151]" italics="true" pageId="4" pageNumber="55">punctulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67F59E3C2564FFC8FB08FBA12B75F547" box="[1198,1304,1127,1151]" pageId="4" pageNumber="55" sensu="lato">sensu lato</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="F082FD1725640033FF00FB402BE1F5A7" box="[166,1420,1158,1183]" gridrow="1" pageId="4" pageNumber="55" rowspan-0="1">
<td id="B353946B25640033FDE9FB402DAFF5A7" box="[591,962,1158,1183]" gridcol="1" gridrow="1" pageId="4" pageNumber="55">
(
<specimenCount id="58B434DC2564FFC8FDF0FB402C88F5A7" box="[598,741,1158,1183]" pageId="4" pageNumber="55" type="generic">12 specimens</specimenCount>
examined)
</td>
<td id="B353946B25640033FC42FB402BE1F5A7" box="[996,1420,1158,1183]" gridcol="2" gridrow="1" pageId="4" pageNumber="55">
<emphasis id="7CC623472564FFC8FC42FB402A88F5A7" bold="true" box="[996,1253,1158,1183]" pageId="4" pageNumber="55">
(
<specimenCount id="58B434DC2564FFC8FC4AFB402A03F5A7" box="[1004,1134,1158,1183]" pageId="4" pageNumber="55" type="generic">9 specimens</specimenCount>
examined)
</emphasis>
</td>
</tr>
<tr id="F082FD1725640033FF00FB612BE1F5F8" box="[166,1420,1191,1216]" gridrow="2" pageId="4" pageNumber="55">
<th id="B353946B25640033FF00FB612C43F5F8" box="[166,558,1191,1216]" gridcol="0" gridrow="2" pageId="4" pageNumber="55">Coloration</th>
<td id="B353946B25640033FDE9FB612DAFF5F8" box="[591,962,1191,1216]" gridcol="1" gridrow="2" pageId="4" pageNumber="55">Chocolate brown</td>
<td id="B353946B25640033FC42FB612BE1F5F8" box="[996,1420,1191,1216]" gridcol="2" gridrow="2" pageId="4" pageNumber="55">Reddish brown</td>
</tr>
<tr id="F082FD1725640033FF00FB0A2BE1F5DD" box="[166,1420,1228,1253]" gridrow="3" pageId="4" pageNumber="55">
<th id="B353946B25640033FF00FB0A2C43F5DD" box="[166,558,1228,1253]" gridcol="0" gridrow="3" pageId="4" pageNumber="55">Anterior margin of pronotum in males</th>
<td id="B353946B25640033FDE9FB0A2DAFF5DD" box="[591,962,1228,1253]" gridcol="1" gridrow="3" pageId="4" pageNumber="55">Slightly forwards curved in middle</td>
<td id="B353946B25640033FC42FB0A2BE1F5DD" box="[996,1420,1228,1253]" gridcol="2" gridrow="3" pageId="4" pageNumber="55">Slightly backwards curved in middle</td>
</tr>
<tr id="F082FD1725640033FF00FB2A2BE1F43D" box="[166,1420,1260,1285]" gridrow="4" pageId="4" pageNumber="55" rowspan-0="1">
<td id="B353946B25640033FDE9FB2A2DAFF43D" box="[591,962,1260,1285]" gridcol="1" gridrow="4" pageId="4" pageNumber="55">
(
<figureCitation id="D689E3D02564FFC8FDF0FB2A2CCBF43D" box="[598,678,1260,1285]" captionStart="FIGURES 2" captionStartId="8.[151,264,1889,1914]" captionTargetBox="[164,1427,351,1859]" captionTargetId="figure@8.[151,1435,337,1869]" captionTargetPageId="8" captionText="FIGURES 2A–F. Diagnostic characters separating Copris (Paracopris) javanensis Bui, Ziegler &amp; Bonkowski, new species (A, C, E) from C. (P.) punctulatus punctulatus (B, D, F). A, B, anterior pronotal margin; C, D, pygidium; E, F, aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3586830" httpUri="https://zenodo.org/record/3586830/files/figure.png" pageId="4" pageNumber="55">Fig. 2A</figureCitation>
)
</td>
<td id="B353946B25640033FC42FB2A2BE1F43D" box="[996,1420,1260,1285]" gridcol="2" gridrow="4" pageId="4" pageNumber="55">
(
<figureCitation id="D689E3D02564FFC8FC4AFB2A2A57F43D" box="[1004,1082,1260,1285]" captionStart="FIGURES 2" captionStartId="8.[151,264,1889,1914]" captionTargetBox="[164,1427,351,1859]" captionTargetId="figure@8.[151,1435,337,1869]" captionTargetPageId="8" captionText="FIGURES 2A–F. Diagnostic characters separating Copris (Paracopris) javanensis Bui, Ziegler &amp; Bonkowski, new species (A, C, E) from C. (P.) punctulatus punctulatus (B, D, F). A, B, anterior pronotal margin; C, D, pygidium; E, F, aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3586830" httpUri="https://zenodo.org/record/3586830/files/figure.png" pageId="4" pageNumber="55">Fig. 2B</figureCitation>
)
</td>
</tr>
<tr id="F082FD1725640033FF00FACB2BE1F41E" box="[166,1420,1293,1318]" gridrow="5" pageId="4" pageNumber="55">
<th id="B353946B25640033FF00FACB2C43F41E" box="[166,558,1293,1318]" gridcol="0" gridrow="5" pageId="4" pageNumber="55">Anterior margin of pronotum in</th>
<td id="B353946B25640033FDE9FACB2DAFF41E" box="[591,962,1293,1318]" gridcol="1" gridrow="5" pageId="4" pageNumber="55">Strongly forwards curved in middle</td>
<td id="B353946B25640033FC42FACB2BE1F41E" box="[996,1420,1293,1318]" gridcol="2" gridrow="5" pageId="4" pageNumber="55">Slightly forwards curved in middle (Fig.</td>
</tr>
<tr id="F082FD1725640033FF00FAEB2BE1F47E" box="[166,1420,1325,1350]" gridrow="6" pageId="4" pageNumber="55">
<th id="B353946B25640033FF00FAEB2C43F47E" box="[166,558,1325,1350]" gridcol="0" gridrow="6" pageId="4" pageNumber="55">females</th>
<td id="B353946B25640033FDE9FAEB2DAFF47E" box="[591,962,1325,1350]" gridcol="1" gridrow="6" pageId="4" pageNumber="55">
(
<figureCitation id="D689E3D02564FFC8FDF0FAEB2CCEF47E" box="[598,675,1325,1350]" captionStart="FIGURES 1" captionStartId="5.[151,264,1965,1990]" captionTargetBox="[215,1374,179,1936]" captionTargetId="figure@5.[206,1381,166,1945]" captionTargetPageId="5" captionText="FIGURES 1A–D. Holotype (A, B) and female paratype (C, D) of Copris (Paracopris) javanensis Bui, Ziegler &amp; Bonkowski, new species. A, C, habitus, dorsal view; B, D, habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3586828" httpUri="https://zenodo.org/record/3586828/files/figure.png" pageId="4" pageNumber="55">Fig. 1C</figureCitation>
)
</td>
<td id="B353946B25640033FC42FAEB2BE1F47E" box="[996,1420,1325,1350]" gridcol="2" gridrow="6" pageId="4" pageNumber="55">4C)</td>
</tr>
<tr id="F082FD1725640033FF00FA892BE1F450" box="[166,1420,1359,1384]" gridrow="7" pageId="4" pageNumber="55">
<th id="B353946B25640033FF00FA892C43F450" box="[166,558,1359,1384]" gridcol="0" gridrow="7" pageId="4" pageNumber="55">Punctures on pygidium surface</th>
<td id="B353946B25640033FDE9FA892DAFF450" box="[591,962,1359,1384]" gridcol="1" gridrow="7" pageId="4" pageNumber="55">
Clearly separated (
<figureCitation id="D689E3D02564FFC8FCABFA892D36F45F" box="[781,859,1359,1384]" captionStart="FIGURES 2" captionStartId="8.[151,264,1889,1914]" captionTargetBox="[164,1427,351,1859]" captionTargetId="figure@8.[151,1435,337,1869]" captionTargetPageId="8" captionText="FIGURES 2A–F. Diagnostic characters separating Copris (Paracopris) javanensis Bui, Ziegler &amp; Bonkowski, new species (A, C, E) from C. (P.) punctulatus punctulatus (B, D, F). A, B, anterior pronotal margin; C, D, pygidium; E, F, aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3586830" httpUri="https://zenodo.org/record/3586830/files/figure.png" pageId="4" pageNumber="55">Fig. 2C</figureCitation>
)
</td>
<td id="B353946B25640033FC42FA892BE1F450" box="[996,1420,1359,1384]" gridcol="2" gridrow="7" pageId="4" pageNumber="55">
Contiguous (
<figureCitation id="D689E3D02564FFC8FBC1FA892ADBF45F" box="[1127,1206,1359,1384]" captionStart="FIGURES 2" captionStartId="8.[151,264,1889,1914]" captionTargetBox="[164,1427,351,1859]" captionTargetId="figure@8.[151,1435,337,1869]" captionTargetPageId="8" captionText="FIGURES 2A–F. Diagnostic characters separating Copris (Paracopris) javanensis Bui, Ziegler &amp; Bonkowski, new species (A, C, E) from C. (P.) punctulatus punctulatus (B, D, F). A, B, anterior pronotal margin; C, D, pygidium; E, F, aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3586830" httpUri="https://zenodo.org/record/3586830/files/figure.png" pageId="4" pageNumber="55">Fig. 2D</figureCitation>
)
</td>
</tr>
<tr id="F082FD1725640033FF00FAB22BE1F4B5" box="[166,1420,1396,1421]" gridrow="8" pageId="4" pageNumber="55">
<th id="B353946B25640033FF00FAB22C43F4B5" box="[166,558,1396,1421]" gridcol="0" gridrow="8" pageId="4" pageNumber="55">Phallobase width relative to parameres</th>
<td id="B353946B25640033FDE9FAB22DAFF4B5" box="[591,962,1396,1421]" gridcol="1" gridrow="8" pageId="4" pageNumber="55">PhaW slightly wider than PaW</td>
<td id="B353946B25640033FC42FAB22BE1F4B5" box="[996,1420,1396,1421]" gridcol="2" gridrow="8" pageId="4" pageNumber="55">
PhaW greatly wider than PaW (
<figureCitation id="D689E3D02564FFC8FA83FAB22B1CF4B5" box="[1317,1393,1396,1421]" captionStart="FIGURES 2" captionStartId="8.[151,264,1889,1914]" captionTargetBox="[164,1427,351,1859]" captionTargetId="figure@8.[151,1435,337,1869]" captionTargetPageId="8" captionText="FIGURES 2A–F. Diagnostic characters separating Copris (Paracopris) javanensis Bui, Ziegler &amp; Bonkowski, new species (A, C, E) from C. (P.) punctulatus punctulatus (B, D, F). A, B, anterior pronotal margin; C, D, pygidium; E, F, aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3586830" httpUri="https://zenodo.org/record/3586830/files/figure.png" pageId="4" pageNumber="55">Fig. 2F</figureCitation>
)
</td>
</tr>
<tr id="F082FD1725640033FF00FA522BE1F495" box="[166,1420,1428,1453]" gridrow="9" pageId="4" pageNumber="55" rowspan-2="1">
<th id="B353946B25640033FF00FA522C43F495" box="[166,558,1428,1453]" gridcol="0" gridrow="9" pageId="4" pageNumber="55">width</th>
<td id="B353946B25640033FDE9FA522DAFF495" box="[591,962,1428,1453]" gridcol="1" gridrow="9" pageId="4" pageNumber="55">
(
<figureCitation id="D689E3D02564FFC8FDF0FA522CCEF495" box="[598,675,1428,1453]" captionStart="FIGURES 2" captionStartId="8.[151,264,1889,1914]" captionTargetBox="[164,1427,351,1859]" captionTargetId="figure@8.[151,1435,337,1869]" captionTargetPageId="8" captionText="FIGURES 2A–F. Diagnostic characters separating Copris (Paracopris) javanensis Bui, Ziegler &amp; Bonkowski, new species (A, C, E) from C. (P.) punctulatus punctulatus (B, D, F). A, B, anterior pronotal margin; C, D, pygidium; E, F, aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3586830" httpUri="https://zenodo.org/record/3586830/files/figure.png" pageId="4" pageNumber="55">Fig. 2E</figureCitation>
)
</td>
</tr>
<tr id="F082FD1725640033FF00FA732BE1F4F6" box="[166,1420,1461,1486]" gridrow="10" pageId="4" pageNumber="55">
<th id="B353946B25640033FF00FA732C43F4F6" box="[166,558,1461,1486]" gridcol="0" gridrow="10" pageId="4" pageNumber="55">Shape of parameres</th>
<td id="B353946B25640033FDE9FA732DAFF4F6" box="[591,962,1461,1486]" gridcol="1" gridrow="10" pageId="4" pageNumber="55">Slightly curved downwards in apical</td>
<td id="B353946B25640033FC42FA732BE1F4F6" box="[996,1420,1461,1486]" gridcol="2" gridrow="10" pageId="4" pageNumber="55">Strongly curved downwards in apical part</td>
</tr>
<tr id="F082FD1725640033FF00FA132BE1F4D6" box="[166,1420,1493,1518]" gridrow="11" pageId="4" pageNumber="55" rowspan-0="1">
<td id="B353946B25640033FDE9FA132DAFF4D6" box="[591,962,1493,1518]" gridcol="1" gridrow="11" pageId="4" pageNumber="55">
part (
<figureCitation id="D689E3D02564FFC8FD22FA132CBCF4D6" box="[644,721,1493,1518]" captionStart="FIGURES 2" captionStartId="8.[151,264,1889,1914]" captionTargetBox="[164,1427,351,1859]" captionTargetId="figure@8.[151,1435,337,1869]" captionTargetPageId="8" captionText="FIGURES 2A–F. Diagnostic characters separating Copris (Paracopris) javanensis Bui, Ziegler &amp; Bonkowski, new species (A, C, E) from C. (P.) punctulatus punctulatus (B, D, F). A, B, anterior pronotal margin; C, D, pygidium; E, F, aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3586830" httpUri="https://zenodo.org/record/3586830/files/figure.png" pageId="4" pageNumber="55">Fig. 2E</figureCitation>
)
</td>
<td id="B353946B25640033FC42FA132BE1F4D6" box="[996,1420,1493,1518]" gridcol="2" gridrow="11" pageId="4" pageNumber="55">
(
<figureCitation id="D689E3D02564FFC8FC4AFA132A5AF4D6" box="[1004,1079,1493,1518]" captionStart="FIGURES 2" captionStartId="8.[151,264,1889,1914]" captionTargetBox="[164,1427,351,1859]" captionTargetId="figure@8.[151,1435,337,1869]" captionTargetPageId="8" captionText="FIGURES 2A–F. Diagnostic characters separating Copris (Paracopris) javanensis Bui, Ziegler &amp; Bonkowski, new species (A, C, E) from C. (P.) punctulatus punctulatus (B, D, F). A, B, anterior pronotal margin; C, D, pygidium; E, F, aedeagus in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3586830" httpUri="https://zenodo.org/record/3586830/files/figure.png" pageId="4" pageNumber="55">Fig. 2F</figureCitation>
)
</td>
</tr>
</table>
</paragraph>
<paragraph id="4E0DFF552564FFC8FF61F9DC2C82F640" blockId="4.[151,1437,1562,1949]" pageId="4" pageNumber="55">
The males of both 
<taxonomicName id="89B284D62564FFC8FE3FF9DC2CABF70C" authority="Bui &amp; Ziegler &amp; Bonkowski, 2019" authorityName="new" authorityYear="2019" box="[409,710,1562,1588]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="javanensis" status="new" subGenus="Paracopris">
<emphasis id="7CC623472564FFC8FE3FF9DC2F88F70C" box="[409,485,1562,1588]" italics="true" pageId="4" pageNumber="55">Copris</emphasis>
(
<emphasis id="7CC623472564FFC8FE53F9DD2C67F70C" box="[501,522,1563,1588]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
) 
<emphasis id="7CC623472564FFC8FDBCF9DD2CFDF70C" box="[538,656,1563,1588]" italics="true" pageId="4" pageNumber="55">javanensis</emphasis>
<taxonomicNameLabel id="67F59E3C2564FFC8FD31F9DC2CABF70C" box="[663,710,1562,1588]" pageId="4" pageNumber="55" rank="species">new</taxonomicNameLabel>
</taxonomicName>
species and 
<taxonomicName id="89B284D62564FFC8FCF5F9DC2A48F70C" authorityName="Wiedemann" authorityYear="1823" box="[851,1061,1562,1588]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" sensu="lato" species="punctulatus" subGenus="Paracopris">
<emphasis id="7CC623472564FFC8FCF5F9DC2D00F70C" box="[851,877,1562,1588]" italics="true" pageId="4" pageNumber="55">C.</emphasis>
(
<emphasis id="7CC623472564FFC8FCDBF9DD2DFEF70C" box="[893,915,1563,1588]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
) 
<emphasis id="7CC623472564FFC8FC04F9DC2A48F70C" box="[930,1061,1562,1588]" italics="true" pageId="4" pageNumber="55">punctulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67F59E3C2564FFC8FB8DF9DC2AF6F70C" box="[1067,1179,1562,1588]" pageId="4" pageNumber="55" sensu="lato">sensu lato</taxonomicNameLabel>
possess a clypeal apex strongly upwards protruded and shallowly and widely emarginated, making these two species distinct from most Asian members of the subgenus 
<taxonomicName id="89B284D62564FFC8FDA1F9A52CE8F744" authorityName="Balthasar" authorityYear="1939" box="[519,645,1635,1660]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="subGenus" subGenus="Paracopris">
<emphasis id="7CC623472564FFC8FDA1F9A52CE8F744" box="[519,645,1635,1660]" italics="true" pageId="4" pageNumber="55">Paracopris</emphasis>
</taxonomicName>
(for details see 
<tableCitation id="0330CAEE2564FFC8FC9DF9A42DFDF744" box="[827,912,1634,1660]" captionStart="TABLE 2" captionStartId="6.[151,239,154,178]" captionTargetBox="[166,1421,261,1584]" captionText="TABLE 2. Diagnostic characters of clypeus separating the new species from the other Copris (Paracopris) species (based on the specimens examined by us in MNHN and NMPC, and data published by Wiedemann 1823; Arrow 1931; Balthasar 1939, 1963; Ochi &amp; Kon 1996; Hanboonsong et al. 2003; Ochi et al. 2019a,b)" httpUri="http://table.plazi.org/id/1ACDAFDD2566FFCAFF31FF5C2DB7F1C2" pageId="4" pageNumber="55" tableUuid="1ACDAFDD2566FFCAFF31FF5C2DB7F1C2">Table 2</tableCitation>
), except for 
<taxonomicName id="89B284D62564FFC8FBB9F9A42A9BF744" authorityName="Lansberge" authorityYear="1886" box="[1055,1270,1634,1660]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="pedarioides" subGenus="Paracopris">
<emphasis id="7CC623472564FFC8FBB9F9A42A54F744" box="[1055,1081,1634,1660]" italics="true" pageId="4" pageNumber="55">C.</emphasis>
(
<emphasis id="7CC623472564FFC8FBECF9A52A32F744" box="[1098,1119,1635,1660]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
) 
<emphasis id="7CC623472564FFC8FBC9F9A42A9BF744" box="[1135,1270,1634,1660]" italics="true" pageId="4" pageNumber="55">pedarioides</emphasis>
</taxonomicName>
<emphasis id="7CC623472564FFC8FB50F9A42A97F744" box="[1270,1274,1634,1660]" italics="true" pageId="4" pageNumber="55">,</emphasis>
<taxonomicName id="89B284D62564FFC8FAA4F9A42BF0F744" authorityName="Arrow" authorityYear="1931" box="[1282,1437,1634,1660]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="surdus" subGenus="Paracopris">
<emphasis id="7CC623472564FFC8FAA4F9A42B71F744" box="[1282,1308,1634,1660]" italics="true" pageId="4" pageNumber="55">C.</emphasis>
(
<emphasis id="7CC623472564FFC8FA8BF9A52B2FF744" box="[1325,1346,1635,1660]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
) 
<emphasis id="7CC623472564FFC8FAF5F9A42BF0F744" box="[1363,1437,1634,1660]" italics="true" pageId="4" pageNumber="55">surdus</emphasis>
</taxonomicName>
and 
<taxonomicName id="89B284D62564FFC8FF60F9402FE7F798" authorityName="Felsche" authorityYear="1910" box="[198,394,1670,1696]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="cariniceps" subGenus="Paracopris">
<emphasis id="7CC623472564FFC8FF60F9402E8DF798" box="[198,224,1670,1696]" italics="true" pageId="4" pageNumber="55">C.</emphasis>
(
<emphasis id="7CC623472564FFC8FF56F9412F68F798" box="[240,261,1671,1696]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
) 
<emphasis id="7CC623472564FFC8FEB3F9412FE7F798" box="[277,394,1671,1696]" italics="true" pageId="4" pageNumber="55">cariniceps</emphasis>
</taxonomicName>
. The three latter can be clearly distinguished from the two former due to having very densely to contiguously punctate elytra and very densely and deeply punctate genae (
<figureCitation id="D689E3D02564FFC8FBA2F96C2A3DF7FC" box="[1028,1104,1706,1732]" captionStart="FIGURES 3" captionStartId="9.[151,264,1806,1831]" captionTargetBox="[163,1427,949,1787]" captionTargetId="figure@9.[151,1435,949,1787]" captionTargetPageId="9" captionText="FIGURES 3A–C. Surface of elytral intervals of Copris (Paracopris) species. A, Copris (P.) javanensis Bui, Ziegler &amp; Bonkowski, new species; B, Copris (P.) punctulatus punctulatus; C, Copris (P.) cariniceps." figureDoi="http://doi.org/10.5281/zenodo.3586832" httpUri="https://zenodo.org/record/3586832/files/figure.png" pageId="4" pageNumber="55">Figs. 3</figureCitation>
A–C). In addition, characters on the male cephalic horn clearly differentiate 
<taxonomicName id="89B284D62564FFC8FD0AF9082DECF7D0" authorityName="Lansberge" authorityYear="1886" box="[684,897,1742,1768]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="pedarioides" subGenus="Paracopris">
<emphasis id="7CC623472564FFC8FD0AF9082CABF7D0" box="[684,710,1742,1768]" italics="true" pageId="4" pageNumber="55">C.</emphasis>
(
<emphasis id="7CC623472564FFC8FD71F9092C81F7D0" box="[727,748,1743,1768]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
) 
<emphasis id="7CC623472564FFC8FD5BF9082DECF7D0" box="[765,897,1742,1768]" italics="true" pageId="4" pageNumber="55">pedarioides</emphasis>
</taxonomicName>
and 
<taxonomicName id="89B284D62564FFC8FC1DF9082A3BF7D0" authorityName="Arrow" authorityYear="1931" box="[955,1110,1742,1768]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="surdus" subGenus="Paracopris">
<emphasis id="7CC623472564FFC8FC1DF9082DB8F7D0" box="[955,981,1742,1768]" italics="true" pageId="4" pageNumber="55">C.</emphasis>
(
<emphasis id="7CC623472564FFC8FC40F9092D91F7D0" box="[998,1020,1743,1768]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
) 
<emphasis id="7CC623472564FFC8FBAAF9082A3BF7D0" box="[1036,1110,1742,1768]" italics="true" pageId="4" pageNumber="55">surdus</emphasis>
</taxonomicName>
from 
<taxonomicName id="89B284D62564FFC8FB3BF9082B08F7D0" authority="Bui &amp; Ziegler &amp; Bonkowski, 2019" authorityName="Bui &amp; Ziegler &amp; Bonkowski" authorityYear="2019" box="[1181,1381,1742,1768]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="javanensis" status="sp. nov." subGenus="Paracopris">
<emphasis id="7CC623472564FFC8FB3BF9082ADAF7D0" box="[1181,1207,1742,1768]" italics="true" pageId="4" pageNumber="55">C.</emphasis>
(
<emphasis id="7CC623472564FFC8FB6FF9092AB3F7D0" box="[1225,1246,1743,1768]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
) 
<emphasis id="7CC623472564FFC8FB49F9092B08F7D0" box="[1263,1381,1743,1768]" italics="true" pageId="4" pageNumber="55">javanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67F59E3C2564FFC8FAC8F9082E85F634" pageId="4" pageNumber="55" rank="species">new species</taxonomicNameLabel>
and 
<taxonomicName id="89B284D62564FFC8FE86F9342F9EF634" authorityName="Wiedemann" authorityYear="1823" box="[288,499,1778,1804]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" sensu="lato" species="punctulatus" subGenus="Paracopris">
<emphasis id="7CC623472564FFC8FE86F9342F57F634" box="[288,314,1778,1804]" italics="true" pageId="4" pageNumber="55">C.</emphasis>
(
<emphasis id="7CC623472564FFC8FEEDF9352F0DF634" box="[331,352,1779,1804]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
) 
<emphasis id="7CC623472564FFC8FED6F9342F9EF634" box="[368,499,1778,1804]" italics="true" pageId="4" pageNumber="55">punctulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67F59E3C2564FFC8FE5DF9342C03F634" box="[507,622,1778,1804]" pageId="4" pageNumber="55" sensu="lato">sensu lato</taxonomicNameLabel>
. In the two former, the male cephalic horn is relatively long, whereas in the two latter, the male cephalic horn is small. The males of 
<taxonomicName id="89B284D62564FFC8FC96F8D02D9EF608" authorityName="Felsche" authorityYear="1910" box="[816,1011,1814,1840]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="cariniceps" subGenus="Paracopris">
<emphasis id="7CC623472564FFC8FC96F8D02D27F608" box="[816,842,1814,1840]" italics="true" pageId="4" pageNumber="55">C.</emphasis>
(
<emphasis id="7CC623472564FFC8FCFCF8D12D02F608" box="[858,879,1815,1840]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
) 
<emphasis id="7CC623472564FFC8FCD8F8D12D9EF608" box="[894,1011,1815,1840]" italics="true" pageId="4" pageNumber="55">cariniceps</emphasis>
</taxonomicName>
bear a clypeus strongly longitudinally carinate in middle, distinctly separating this species from 
<taxonomicName id="89B284D62564FFC8FC96F8FC2D94F66C" authority="Bui &amp; Ziegler &amp; Bonkowski, 2019" authorityName="Bui &amp; Ziegler &amp; Bonkowski" authorityYear="2019" box="[816,1017,1850,1876]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="javanensis" status="sp. nov." subGenus="Paracopris">
<emphasis id="7CC623472564FFC8FC96F8FC2D24F66C" box="[816,841,1850,1876]" italics="true" pageId="4" pageNumber="55">C.</emphasis>
(
<emphasis id="7CC623472564FFC8FCFAF8FD2D1CF66C" box="[860,881,1851,1876]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
) 
<emphasis id="7CC623472564FFC8FC25F8FD2D94F66C" box="[899,1017,1851,1876]" italics="true" pageId="4" pageNumber="55">javanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67F59E3C2564FFC8FBA4F8FC2AE6F66C" box="[1026,1163,1850,1876]" pageId="4" pageNumber="55" rank="species">new species</taxonomicNameLabel>
and 
<taxonomicName id="89B284D62564FFC8FB60F8FC2BF0F66C" authorityName="Wiedemann" authorityYear="1823" box="[1222,1437,1850,1876]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" sensu="lato" species="punctulatus" subGenus="Paracopris">
<emphasis id="7CC623472564FFC8FB60F8FC2A8DF66C" box="[1222,1248,1850,1876]" italics="true" pageId="4" pageNumber="55">C.</emphasis>
(
<emphasis id="7CC623472564FFC8FB54F8FD2B65F66C" box="[1266,1288,1851,1876]" italics="true" pageId="4" pageNumber="55">P.</emphasis>
) 
<emphasis id="7CC623472564FFC8FABCF8FC2BF0F66C" box="[1306,1437,1850,1876]" italics="true" pageId="4" pageNumber="55">punctulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67F59E3C2564FFC8FF31F8982F6BF640" box="[151,262,1886,1912]" pageId="4" pageNumber="55" sensu="lato">sensu lato</taxonomicNameLabel>
whose clypeus has no longitudinal carinae.
</paragraph>
</subSubSection>
<subSubSection id="06A8ACDE2564FFCAFF61F8442A5CF687" lastPageId="6" lastPageNumber="57" pageId="4" pageNumber="55" type="etymology">
<paragraph id="4E0DFF552564FFC8FF61F8442B51F6A5" blockId="4.[151,1437,1562,1949]" box="[199,1340,1922,1949]" pageId="4" pageNumber="55">
<emphasis id="7CC623472564FFC8FF61F8442F23F6A4" bold="true" box="[199,334,1922,1948]" pageId="4" pageNumber="55">Etymology.</emphasis>
The specific epithet “javanensis” refers to the name of the 
<typeStatus id="910941F72564FFC8FC42F8442A79F6A4" box="[996,1044,1922,1948]" pageId="4" pageNumber="55">type</typeStatus>
locality, 
<collectingRegion id="8C7631B72564FFC8FBDAF8442AC3F6A4" box="[1148,1198,1922,1948]" country="Indonesia" name="Jawa Timur" pageId="4" pageNumber="55">Java</collectingRegion>
(
<collectingCountry id="36A5BFC52564FFC8FB1AF8442B5DF6A5" box="[1212,1328,1922,1949]" name="Indonesia" pageId="4" pageNumber="55">Indonesia</collectingCountry>
).
</paragraph>
<caption id="1ACDAFDD2565FFC9FF31F86B2D2AF6D2" ID-DOI="http://doi.org/10.5281/zenodo.3586828" ID-Zenodo-Dep="3586828" httpUri="https://zenodo.org/record/3586828/files/figure.png" pageId="5" pageNumber="56" startId="5.[151,264,1965,1990]" targetBox="[215,1374,179,1936]" targetPageId="5">
<paragraph id="4E0DFF552565FFC9FF31F86B2D2AF6D2" blockId="5.[151,1437,1965,2026]" pageId="5" pageNumber="56">
<emphasis id="7CC623472565FFC9FF31F86B2F3EF6FE" bold="true" box="[151,339,1965,1990]" pageId="5" pageNumber="56">FIGURES 1A–D.</emphasis>
Holotype (A, B) and female paratype (C, D) of 
<taxonomicName id="89B284D62565FFC9FC99F8692BF5F6FE" authority="Bui, Ziegler &amp; Bonkowski" authorityName="Bui, Ziegler &amp; Bonkowski" authorityYear="2019" box="[831,1432,1965,1990]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="javanensis" status="sp. nov." subGenus="Paracopris">
<emphasis id="7CC623472565FFC9FC99F8692DE9F6FE" box="[831,900,1967,1990]" italics="true" pageId="5" pageNumber="56">Copris</emphasis>
(
<emphasis id="7CC623472565FFC9FC35F8692A68F6FE" box="[915,1029,1967,1990]" italics="true" pageId="5" pageNumber="56">Paracopris</emphasis>
) 
<emphasis id="7CC623472565FFC9FBB3F8692AEDF6FE" box="[1045,1152,1967,1990]" italics="true" pageId="5" pageNumber="56">javanensis</emphasis>
Bui, Ziegler &amp; Bonkowski
</taxonomicName>
, 
<taxonomicNameLabel id="67F59E3C2565FFC9FF31F8152F7CF6D2" box="[151,273,2001,2026]" pageId="5" pageNumber="56" rank="species">new species</taxonomicNameLabel>
. A, C, habitus, dorsal view; B, D, habitus, ventral view.
</paragraph>
</caption>
<caption id="1ACDAFDD2566FFCAFF31FF5C2DB7F1C2" ID-Table-UUID="1ACDAFDD2566FFCAFF31FF5C2DB7F1C2" httpUri="http://table.plazi.org/id/1ACDAFDD2566FFCAFF31FF5C2DB7F1C2" pageId="6" pageNumber="57" targetBox="[166,1421,261,1584]" targetIsTable="true" targetPageId="6">
<paragraph id="4E0DFF552566FFCAFF31FF5C2DB7F1C2" blockId="6.[151,1436,154,250]" pageId="6" pageNumber="57">
<emphasis id="7CC623472566FFCAFF31FF5C2F64F18A" bold="true" box="[151,265,154,178]" pageId="6" pageNumber="57">
<tableCitation id="0330CAEE2566FFCAFF31FF5C2F69F18A" box="[151,260,154,178]" captionStart="TABLE 2" captionStartId="6.[151,239,154,178]" captionTargetBox="[166,1421,261,1584]" captionText="TABLE 2. Diagnostic characters of clypeus separating the new species from the other Copris (Paracopris) species (based on the specimens examined by us in MNHN and NMPC, and data published by Wiedemann 1823; Arrow 1931; Balthasar 1939, 1963; Ochi &amp; Kon 1996; Hanboonsong et al. 2003; Ochi et al. 2019a,b)" httpUri="http://table.plazi.org/id/1ACDAFDD2566FFCAFF31FF5C2DB7F1C2" pageId="6" pageNumber="57" tableUuid="1ACDAFDD2566FFCAFF31FF5C2DB7F1C2">TABLE 2</tableCitation>
.
</emphasis>
Diagnostic characters of clypeus separating the new species from the other 
<taxonomicName id="89B284D62566FFCAFB80FF5C2A92F18A" authority="" authorityName="Balthasar" authorityYear="1939" box="[1062,1279,154,178]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="subGenus" subGenus="Paracopris">
<emphasis id="7CC623472566FFCAFB80FF5C2A02F189" box="[1062,1135,154,177]" italics="true" pageId="6" pageNumber="57">Copris</emphasis>
(
<emphasis id="7CC623472566FFCAFBD8FF5C2A9BF189" box="[1150,1270,154,177]" italics="true" pageId="6" pageNumber="57">Paracopris</emphasis>
)
</taxonomicName>
species (based on the specimens examined by us in MNHN and NMPC, and data published by 
<bibRefCitation id="2A2382A42566FFCAFC44FF782ACDF1EE" author="Wiedemann, C. R. W." box="[994,1184,190,214]" pageId="6" pageNumber="57" pagination="1 - 164" refId="ref9330" refString="Wiedemann, C. R. W. (1823) Zweihundert neue Kafer von Java, Bengalen und dem Borgebirge der guten Hoffnung. Zoolo- gisches Magazin, 2, 1 - 164." type="journal article" year="1823">Wiedemann 1823</bibRefCitation>
; 
<bibRefCitation id="2A2382A42566FFCAFB0FFF782B40F1EE" author="Arrow, G. J." box="[1193,1325,190,214]" pageId="6" pageNumber="57" refId="ref7267" refString="Arrow, G. J. (1931) The fauna of British India, including Ceylon and Burma. Coleoptera Lamellicornia. Part III (Coprinae). Taylor and Francis, London, 428 pp." type="book" year="1931">Arrow 1931</bibRefCitation>
; 
<bibRefCitation id="2A2382A42566FFCAFA91FF782BF1F1EE" author="Balthasar, V." box="[1335,1436,190,214]" pageId="6" pageNumber="57" pagination="1 - 36" refId="ref7382" refString="Balthasar, V. (1939) Neue Arten der coprophagen Scarabaeiden aus dem Museo Zoologico delle R. Universita di Firenze. Redia, 25, 1 - 36." type="journal article" year="1939">Balthasar</bibRefCitation>
1939, 
<bibRefCitation id="2A2382A42566FFCAFF7CFF242F7FF1C2" author="Balthasar, V." box="[218,274,226,250]" pageId="6" pageNumber="57" refId="ref7444" refString="Balthasar, V. (1963) Monographie der Scarabaeidae und Aphodiidae der Palaearktischen und Orientalischen Region. Coleo- ptera: Lamellicornia. Band 1. Allgemeiner Teil, Systematischer Teil: 1. Scarabaeinae, 2. Coprinae (Pinotini, Coprini). Verlag der Tschechoslowakischen Akademie der Wissenschaften, Prag, 391 pp." type="book" year="1963">1963</bibRefCitation>
; 
<bibRefCitation id="2A2382A42566FFCAFEBBFF242F8CF1C2" author="Ochi, T. &amp; Kon, M." box="[285,481,226,250]" pageId="6" pageNumber="57" pagination="29 - 35" refId="ref8462" refString="Ochi, T. &amp; Kon, M. (1996) Studies on the coprophagous scarab beetles from East Asia (IV) (Coleoptera: Scarabaeidae). Gior- nale Italiano di Entomologia, 8, 29 - 35." type="journal article" year="1996">Ochi &amp; Kon 1996</bibRefCitation>
; 
<bibRefCitation id="2A2382A42566FFCAFE4DFF242C96F1C2" author="Hanboonsong, Y. &amp; Masumoto, K. &amp; Ochi, T." box="[491,763,226,250]" pageId="6" pageNumber="57" pagination="103 - 124" refId="ref7902" refString="Hanboonsong, Y., Masumoto, K. &amp; Ochi, T. (2003) Dung beetles (Coleoptera, Scarabaeidae) of Thailand. Part 5. Genera Copris and Microcopris (Coprini). Elytra, 31, 103 - 124." type="journal article" year="2003">
Hanboonsong 
<emphasis id="7CC623472566FFCAFD20FF252CD8F1C2" box="[646,693,226,250]" italics="true" pageId="6" pageNumber="57">et al</emphasis>
. 2003
</bibRefCitation>
; 
<bibRefCitation id="2A2382A42566FFCAFCA3FF242DACF1C2" author="Ochi, T. &amp; Kon, M. &amp; Kawahara, M." box="[773,961,226,250]" pageId="6" pageNumber="57" pagination="469 - 478" refId="ref8655" refString="Ochi, T., Kon, M. &amp; Kawahara, M. (2019 a). Notes on the Coprophagous Scarab beetles from Southeast Asia. XXXII Four new taxa of the genus Copris from Myanmar (Coleoptera, Scarabaeidae). Giornale Italiano di Entomologia, 15 (64), 469 - 478." type="journal article" year="2019">
Ochi 
<emphasis id="7CC623472566FFCAFCE6FF252D02F1C2" box="[832,879,226,250]" italics="true" pageId="6" pageNumber="57">et al</emphasis>
. 2019a
</bibRefCitation>
,b)
</paragraph>
</caption>
<paragraph id="4E0DFF552566FFCAFF00FEC32A1CF717" pageId="6" pageNumber="57">
<table id="3CB20DF525660033FF00FEC32BE0F708" box="[166,1421,261,1584]" gridcols="2" gridrows="26" pageId="6" pageNumber="57">
<tr id="F082FD1725660033FF00FEC32BE0F026" box="[166,1421,261,286]" gridrow="0" pageId="6" pageNumber="57">
<th id="B353946B25660033FF00FEC32FD1F026" box="[166,444,261,286]" gridcol="0" gridrow="0" pageId="6" pageNumber="57">Species</th>
<th id="B353946B25660033FDAEFEC32BE0F026" box="[520,1421,261,286]" gridcol="1" gridrow="0" pageId="6" pageNumber="57">
<emphasis id="7CC623472566FFCAFDAEFEC32D53F026" bold="true" box="[520,830,261,286]" pageId="6" pageNumber="57">Character of clypeus (males)</emphasis>
</th>
</tr>
<tr id="F082FD1725660033FF00FEED2BE0F07C" box="[166,1421,299,324]" gridrow="1" pageId="6" pageNumber="57">
<td id="B353946B25660033FF00FEED2FD1F07C" box="[166,444,299,324]" gridcol="0" gridrow="1" pageId="6" pageNumber="57">
<taxonomicName id="89B284D62566FFCAFF00FEEA2FD1F07B" authority="Bui &amp; Ziegler &amp; Bonkowski, 2019" authorityName="new" authorityYear="2019" box="[166,444,300,324]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="javanensis" status="new" subGenus="Paracopris">
<emphasis id="7CC623472566FFCAFF00FEEA2E86F07B" box="[166,235,300,323]" italics="true" pageId="6" pageNumber="57">Copris</emphasis>
(
<emphasis id="7CC623472566FFCAFEA6FEEB2F7EF07C" box="[256,275,301,324]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFE87FEEB2FE1F07C" box="[289,396,301,324]" italics="true" pageId="6" pageNumber="57">javanensis</emphasis>
new
</taxonomicName>
</td>
<td id="B353946B25660033FDAEFEED2BE0F07C" box="[520,1421,299,324]" gridcol="1" gridrow="1" pageId="6" pageNumber="57">Clypeal apex distinctly upwards protruded, and widely and shallowly emarginated.</td>
</tr>
<tr id="F082FD1725660033FF00FE892BE0F050" box="[166,1421,335,360]" gridrow="2" pageId="6" pageNumber="57" rowspan-1="1">
<td id="B353946B25660033FF00FE892FD1F050" box="[166,444,335,360]" gridcol="0" gridrow="2" pageId="6" pageNumber="57">species</td>
</tr>
<tr id="F082FD1725660033FF00FEB62BE0F0B1" box="[166,1421,368,393]" gridrow="3" pageId="6" pageNumber="57">
<td id="B353946B25660033FF00FEB62FD1F0B1" box="[166,444,368,393]" gridcol="0" gridrow="3" pageId="6" pageNumber="57">
<emphasis id="7CC623472566FFCAFF00FEB72ED0F0B0" box="[166,189,369,392]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
(
<emphasis id="7CC623472566FFCAFF6AFEB72EB2F0B0" box="[204,223,369,392]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFF4BFEB72FFDF0B1" box="[237,400,369,393]" italics="true" pageId="6" pageNumber="57">anomiopseoides</emphasis>
</td>
<td id="B353946B25660033FDAEFEB62BE0F0B1" box="[520,1421,368,393]" gridcol="1" gridrow="3" pageId="6" pageNumber="57">
<taxonomicName id="89B284D62566FFCAFDAEFEB62C3AF0B1" authority="Clypeal" authorityName="Clypeal" box="[520,599,368,393]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="anomiopseoides" subGenus="Paracopris">Clypeal</taxonomicName>
apex deeply and almost semicircularly emarginated; middle part of 
<tr id="F082FD1725660033FF00FE522BE0F0F6" box="[166,1421,404,462]" gridrow="4" pageId="6" pageNumber="57">
<td id="B353946B25660033FF00FE522FD1F0F6" box="[166,444,404,462]" gridcol="0" gridrow="4" pageId="6" pageNumber="57">
<emphasis id="7CC623472566FFCAFAAFFEB62ED0F0F5" italics="true" pageId="6" pageNumber="57">emarginaC.</emphasis>
(
<emphasis id="7CC623472566FFCAFF6AFE702EB2F0F5" box="[204,223,438,461]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFF4BFE732F39F0F5" box="[237,340,437,461]" italics="true" pageId="6" pageNumber="57">andrewesi</emphasis>
</td>
<td id="B353946B25660033FDAEFE522BE0F0F6" box="[520,1421,404,462]" gridcol="1" gridrow="4" pageId="6" pageNumber="57">
tion strongly protruded upwards, and bifurcated. 
<taxonomicName id="89B284D62566FFCAFDAEFE732C3AF0F6" authority="Clypeal" authorityName="Clypeal" box="[520,599,437,462]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="andrewesi" subGenus="Paracopris">Clypeal</taxonomicName>
apex strongly reflexed and emarginated; middle part of emargination protruded,
</td>
</tr>
</td>
</tr>
<tr id="F082FD1725660033FF00FE1F2BE0F32A" box="[166,1421,473,530]" gridrow="5" pageId="6" pageNumber="57">
<td id="B353946B25660033FF00FE1F2FD1F32A" box="[166,444,473,530]" gridcol="0" gridrow="5" pageId="6" pageNumber="57">
<taxonomicName id="89B284D62566FFCAFF00FE3C2F34F32A" authorityName="Ochi, Kon &amp; Pham" authorityYear="2019" box="[166,345,506,530]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="bartolozzii" subGenus="Paracopris">
<emphasis id="7CC623472566FFCAFF00FE3C2ED0F329" box="[166,189,506,529]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
(
<emphasis id="7CC623472566FFCAFF6AFE3D2EB2F32A" box="[204,223,507,530]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFF4BFE3C2F34F32A" box="[237,345,506,530]" italics="true" pageId="6" pageNumber="57">bartolozzii</emphasis>
</taxonomicName>
(**)
</td>
<td id="B353946B25660033FDAEFE1F2BE0F32A" box="[520,1421,473,530]" gridcol="1" gridrow="5" pageId="6" pageNumber="57">
forming a rounded lobe. “Clypeal margin rather broadly but shallowly emarginate at apex, each side of 
<tr id="F082FD1725660033FF00FDDB2BE0F30E" box="[166,1421,541,566]" gridrow="6" pageId="6" pageNumber="57">
<td id="B353946B25660033FF00FDDB2BE0F30E" box="[166,1421,541,566]" colspan="2" colspanRight="1" gridcol="0" gridrow="6" pageId="6" pageNumber="57">
emargination weakly produced forward, forming small, rounded and reflexed tooth” (
<bibRefCitation id="2A2382A42566FFCAFAA8FDD82B14F30E" author="Ochi, T. &amp; Kon, M. &amp; Pham, H. T." box="[1294,1401,542,566]" pageId="6" pageNumber="57" pagination="435 - 446" refId="ref8783" refString="Ochi, T., Kon, M. &amp; Pham, H. T. (2019 b) Five new taxa of Copris (Coleoptera: Scarabaeidae) from Vietnam and Laos. Giornale Italiano di Entomologia, 15 (64), 435 - 446." type="journal article" year="2019">
<emphasis id="7CC623472566FFCAFAA8FDD82B14F30E" box="[1294,1401,542,566]" italics="true" pageId="6" pageNumber="57">Ochi et al.</emphasis>
</bibRefCitation>
</td>
</tr>
</td>
</tr>
<tr id="F082FD1725660033FF00FD872BE0F343" box="[166,1421,577,635]" gridrow="7" pageId="6" pageNumber="57">
<td id="B353946B25660033FF00FD872FD1F343" box="[166,444,577,635]" gridcol="0" gridrow="7" pageId="6" pageNumber="57">
<taxonomicName id="89B284D62566FFCAFF00FDA52F16F343" authorityName="Ochi, Kon &amp; Kawahara" authorityYear="2019" box="[166,379,611,635]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="chinmontanus" subGenus="Paracopris">
<emphasis id="7CC623472566FFCAFF00FDA52ED0F342" box="[166,189,611,634]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
(
<emphasis id="7CC623472566FFCAFF6AFDA52EB2F342" box="[204,223,611,634]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFF4BFDA52F16F343" box="[237,379,611,635]" italics="true" pageId="6" pageNumber="57">chinmontanus</emphasis>
</taxonomicName>
(**)
</td>
<td id="B353946B25660033FDAEFD872BE0F343" box="[520,1421,577,635]" gridcol="1" gridrow="7" pageId="6" pageNumber="57">2019b) “Clypeus with anterior margin suddenly produced upward as a clypeal horn at middle,</td>
</tr>
<tr id="F082FD1725660033FF00FD402BE0F3A7" box="[166,1421,646,671]" gridrow="8" pageId="6" pageNumber="57">
<td id="B353946B25660033FF00FD402BE0F3A7" box="[166,1421,646,671]" colspan="2" colspanRight="1" gridcol="0" gridrow="8" pageId="6" pageNumber="57">
which is curved backward near the pointed apex, ca. 
<quantity id="894A52B02566FFCAFB87FD412A03F3A6" box="[1057,1134,647,671]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="57" unit="mm" value="1.2">1.2 mm</quantity>
in length in lateral view…”
</td>
</tr>
<tr id="F082FD1725660033FF00FD6C2BE0F3DC" box="[166,1421,682,740]" gridrow="9" pageId="6" pageNumber="57">
<td id="B353946B25660033FF00FD6C2FD1F3DC" box="[166,444,682,740]" gridcol="0" gridrow="9" pageId="6" pageNumber="57">
<emphasis id="7CC623472566FFCAFF00FD0A2ED0F3DB" box="[166,189,716,739]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
(
<emphasis id="7CC623472566FFCAFF6AFD0A2EB2F3DB" box="[204,223,716,739]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFF4BFD0A2FF8F3DB" box="[237,405,716,739]" italics="true" pageId="6" pageNumber="57">compressipennis</emphasis>
</td>
<td id="B353946B25660033FDAEFD6C2BE0F3DC" box="[520,1421,682,740]" gridcol="1" gridrow="9" pageId="6" pageNumber="57">
(
<bibRefCitation id="2A2382A42566FFCAFDA9FD6C2CD2F3FB" author="Ochi, T. &amp; Kon, M. &amp; Kawahara, M." box="[527,703,682,707]" pageId="6" pageNumber="57" pagination="469 - 478" refId="ref8655" refString="Ochi, T., Kon, M. &amp; Kawahara, M. (2019 a). Notes on the Coprophagous Scarab beetles from Southeast Asia. XXXII Four new taxa of the genus Copris from Myanmar (Coleoptera, Scarabaeidae). Giornale Italiano di Entomologia, 15 (64), 469 - 478." type="journal article" year="2019">
Ochi 
<emphasis id="7CC623472566FFCAFDEEFD6D2C16F3FB" box="[584,635,683,707]" italics="true" pageId="6" pageNumber="57">et al.</emphasis>
2019a
</bibRefCitation>
) 
<taxonomicName id="89B284D62566FFCAFDAEFD0D2C3AF3DC" authority="Clypeal" authorityName="Clypeal" box="[520,599,715,740]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="compressipennis" subGenus="Paracopris">Clypeal</taxonomicName>
apex widely emarginated, and not protruded.
</td>
</tr>
<tr id="F082FD1725660033FF00FD362BE0F231" box="[166,1421,752,777]" gridrow="10" pageId="6" pageNumber="57">
<td id="B353946B25660033FF00FD362FD1F231" box="[166,444,752,777]" gridcol="0" gridrow="10" pageId="6" pageNumber="57">
<emphasis id="7CC623472566FFCAFF00FD342ED0F231" box="[166,189,754,777]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
(
<emphasis id="7CC623472566FFCAFF6AFD342EB2F231" box="[204,223,754,777]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFF4BFD372F27F231" box="[237,330,753,777]" italics="true" pageId="6" pageNumber="57">cribratus</emphasis>
</td>
<td id="B353946B25660033FDAEFD362BE0F231" box="[520,1421,752,777]" gridcol="1" gridrow="10" pageId="6" pageNumber="57">
<taxonomicName id="89B284D62566FFCAFDAEFD362C3AF231" authority="Clypeal" authorityName="Clypeal" box="[520,599,752,777]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="cribratus" subGenus="Paracopris">Clypeal</taxonomicName>
apex distinctly emarginated, and not protruded.
</td>
</tr>
<tr id="F082FD1725660033FF00FCD02BE0F217" box="[166,1421,790,815]" gridrow="11" pageId="6" pageNumber="57">
<td id="B353946B25660033FF00FCD02FD1F217" box="[166,444,790,815]" gridcol="0" gridrow="11" pageId="6" pageNumber="57">
<emphasis id="7CC623472566FFCAFF00FCD12ED0F216" box="[166,189,791,814]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
(
<emphasis id="7CC623472566FFCAFF6AFCDE2EB2F217" box="[204,223,792,815]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFF4BFCD12F2EF217" box="[237,323,791,815]" italics="true" pageId="6" pageNumber="57">davisoni</emphasis>
</td>
<td id="B353946B25660033FDAEFCD02BE0F217" box="[520,1421,790,815]" gridcol="1" gridrow="11" pageId="6" pageNumber="57">
<taxonomicName id="89B284D62566FFCAFDAEFCD02C3AF217" authority="Clypeal" authorityName="Clypeal" box="[520,599,790,815]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="davisoni" subGenus="Paracopris">Clypeal</taxonomicName>
apex upwards protruded, forming one backwardly curved and pointed horn.
</td>
</tr>
<tr id="F082FD1725660033FF00FCFA2BE0F26D" box="[166,1421,828,853]" gridrow="12" pageId="6" pageNumber="57">
<td id="B353946B25660033FF00FCFA2FD1F26D" box="[166,444,828,853]" gridcol="0" gridrow="12" pageId="6" pageNumber="57">
<emphasis id="7CC623472566FFCAFF00FCFB2ED0F26C" box="[166,189,829,852]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
(
<emphasis id="7CC623472566FFCAFF6AFCFB2EB2F26C" box="[204,223,829,852]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFF4BFCFB2F5BF26C" box="[237,310,829,852]" italics="true" pageId="6" pageNumber="57">excisus</emphasis>
</td>
<td id="B353946B25660033FDAEFCFA2BE0F26D" box="[520,1421,828,853]" gridcol="1" gridrow="12" pageId="6" pageNumber="57">
<taxonomicName id="89B284D62566FFCAFDAEFCFA2C3AF26D" authority="Clypeal" authorityName="Clypeal" box="[520,599,828,853]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="excisus" subGenus="Paracopris">Clypeal</taxonomicName>
apex widely and deeply emarginated, forming a pair of slender lateral processes
</td>
</tr>
<tr id="F082FD1725660033FF00FCA62BE0F2A2" box="[166,1421,864,922]" gridrow="13" pageId="6" pageNumber="57">
<td id="B353946B25660033FF00FCA62FD1F2A2" box="[166,444,864,922]" gridcol="0" gridrow="13" pageId="6" pageNumber="57">
<emphasis id="7CC623472566FFCAFF00FC442ED0F2A1" box="[166,189,898,921]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
(
<emphasis id="7CC623472566FFCAFF6AFC442EB2F2A1" box="[204,223,898,921]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFF4BFC442F24F2A2" box="[237,329,898,922]" italics="true" pageId="6" pageNumber="57">furciceps</emphasis>
</td>
<td id="B353946B25660033FDAEFCA62BE0F2A2" box="[520,1421,864,922]" gridcol="1" gridrow="13" pageId="6" pageNumber="57">
forwards directed. 
<taxonomicName id="89B284D62566FFCAFDAEFC472C3AF2A2" authority="Clypeal" authorityName="Clypeal" box="[520,599,897,922]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="furciceps" subGenus="Paracopris">Clypeal</taxonomicName>
apex deeply and widely emarginated, and with one backwards and pointed horn
</td>
</tr>
<tr id="F082FD1725660033FF00FC632BE0F2E6" box="[166,1421,933,990]" gridrow="14" pageId="6" pageNumber="57">
<td id="B353946B25660033FF00FC632FD1F2E6" box="[166,444,933,990]" gridcol="0" gridrow="14" pageId="6" pageNumber="57">
<emphasis id="7CC623472566FFCAFF00FC012ED0F2E6" box="[166,189,967,990]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
(
<emphasis id="7CC623472566FFCAFF6AFC012EB2F2E6" box="[204,223,967,990]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFF4BFC012F55F2E6" box="[237,312,967,990]" italics="true" pageId="6" pageNumber="57">imitans</emphasis>
</td>
<td id="B353946B25660033FDAEFC632BE0F2E6" box="[520,1421,933,990]" gridcol="1" gridrow="14" pageId="6" pageNumber="57">
in middle. 
<taxonomicName id="89B284D62566FFCAFDAEFC032C3AF2E6" authority="Clypeal" authorityName="Clypeal" box="[520,599,965,990]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="imitans" subGenus="Paracopris">Clypeal</taxonomicName>
apex reflexed, and weakly emarginated; middle part of emargination upwards
</td>
</tr>
<tr id="F082FD1725660033FF00FC2F2BE0F51B" box="[166,1421,1001,1059]" gridrow="15" pageId="6" pageNumber="57">
<td id="B353946B25660033FF00FC2F2FD1F51B" box="[166,444,1001,1059]" gridcol="0" gridrow="15" pageId="6" pageNumber="57">
<taxonomicName id="89B284D62566FFCAFF00FBCD2F5BF51B" authorityName="Ochi &amp; Kon" authorityYear="1996" box="[166,310,1035,1059]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="kasagii" subGenus="Paracopris">
<emphasis id="7CC623472566FFCAFF00FBCD2ED0F51A" box="[166,189,1035,1058]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
(
<emphasis id="7CC623472566FFCAFF6AFBCA2EB2F51B" box="[204,223,1036,1059]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFF4BFBCD2F5BF51B" box="[237,310,1035,1059]" italics="true" pageId="6" pageNumber="57">kasagii</emphasis>
</taxonomicName>
(*)
</td>
<td id="B353946B25660033FDAEFC2F2BE0F51B" box="[520,1421,1001,1059]" gridcol="1" gridrow="15" pageId="6" pageNumber="57">protruded, forming one short erect process. Clypeal anterior margin distinctly bidentate in middle, and not protruded.</td>
</tr>
<tr id="F082FD1725660033FF00FBF62BE0F571" box="[166,1421,1072,1097]" gridrow="16" pageId="6" pageNumber="57">
<td id="B353946B25660033FF00FBF62FD1F571" box="[166,444,1072,1097]" gridcol="0" gridrow="16" pageId="6" pageNumber="57">
<taxonomicName id="89B284D62566FFCAFF00FBF72F3BF571" authorityName="Ochi, Kon &amp; Pham" authorityYear="2019" box="[166,342,1073,1097]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="montreuili" subGenus="Paracopris">
<emphasis id="7CC623472566FFCAFF00FBF72ED0F570" box="[166,189,1073,1096]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
(
<emphasis id="7CC623472566FFCAFF6AFBF72EB2F570" box="[204,223,1073,1096]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFF4BFBF72F3BF571" box="[237,342,1073,1097]" italics="true" pageId="6" pageNumber="57">montreuili</emphasis>
</taxonomicName>
(**)
</td>
<td id="B353946B25660033FDAEFBF62BE0F571" box="[520,1421,1072,1097]" gridcol="1" gridrow="16" pageId="6" pageNumber="57">“Clypeus with antero-median portion noticeably broadly incised, posterior portion of the</td>
</tr>
<tr id="F082FD1725660033FF00FB922BE0F555" box="[166,1421,1108,1133]" gridrow="17" pageId="6" pageNumber="57">
<td id="B353946B25660033FF00FB922BE0F555" box="[166,1421,1108,1133]" colspan="2" colspanRight="1" gridcol="0" gridrow="17" pageId="6" pageNumber="57">incision developed as a short straight prominence which is strongly inclined backward”</td>
</tr>
<tr id="F082FD1725660033FF00FBBE2BE0F58A" box="[166,1421,1144,1202]" gridrow="18" pageId="6" pageNumber="57">
<td id="B353946B25660033FF00FBBE2FD1F58A" box="[166,444,1144,1202]" gridcol="0" gridrow="18" pageId="6" pageNumber="57">
<taxonomicName id="89B284D62566FFCAFF00FB5C2F5BF589" authorityName="Ochi, Kon &amp; Kawahara" authorityYear="2019" box="[166,310,1177,1201]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="olivieri" subGenus="Paracopris">
<emphasis id="7CC623472566FFCAFF00FB5C2ED0F589" box="[166,189,1178,1201]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
(
<emphasis id="7CC623472566FFCAFF6AFB5C2EB2F589" box="[204,223,1178,1201]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFF4BFB5F2F5BF589" box="[237,310,1177,1201]" italics="true" pageId="6" pageNumber="57">olivieri</emphasis>
</taxonomicName>
(**)
</td>
<td id="B353946B25660033FDAEFBBE2BE0F58A" box="[520,1421,1144,1202]" gridcol="1" gridrow="18" pageId="6" pageNumber="57">
(
<bibRefCitation id="2A2382A42566FFCAFDA9FBBE2CA8F5A9" author="Ochi, T. &amp; Kon, M. &amp; Pham, H. T." box="[527,709,1144,1169]" pageId="6" pageNumber="57" pagination="435 - 446" refId="ref8783" refString="Ochi, T., Kon, M. &amp; Pham, H. T. (2019 b) Five new taxa of Copris (Coleoptera: Scarabaeidae) from Vietnam and Laos. Giornale Italiano di Entomologia, 15 (64), 435 - 446." type="journal article" year="2019">
Ochi 
<emphasis id="7CC623472566FFCAFDEEFBBF2C16F5A9" box="[584,635,1145,1169]" italics="true" pageId="6" pageNumber="57">et al.</emphasis>
2019b
</bibRefCitation>
). “Clypeus with a slender horn situated at middle just behind clypeal margin, which is ca.
</td>
</tr>
<tr id="F082FD1725660033FF00FB7B2BE0F5EE" box="[166,1421,1213,1238]" gridrow="19" pageId="6" pageNumber="57">
<td id="B353946B25660033FF00FB7B2BE0F5EE" box="[166,1421,1213,1238]" colspan="2" colspanRight="1" gridcol="0" gridrow="19" pageId="6" pageNumber="57">
<quantity id="894A52B02566FFCAFDAEFB7B2C38F5ED" box="[520,597,1213,1237]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="57" unit="mm" value="1.7">1.7 mm</quantity>
long in lateral view, slightly curved backward, another short point a little behind
</td>
</tr>
<tr id="F082FD1725660033FF00FB272BE0F5C2" box="[166,1421,1249,1274]" gridrow="20" pageId="6" pageNumber="57">
<td id="B353946B25660033FF00FB272BE0F5C2" box="[166,1421,1249,1274]" colspan="2" colspanRight="1" gridcol="0" gridrow="20" pageId="6" pageNumber="57">
the cephalic horn is present, which is ca. 
<quantity id="894A52B02566FFCAFC0FFB272D9BF5C1" box="[937,1014,1249,1273]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="57" unit="mm" value="0.3">0.3 mm</quantity>
in height and slightly inclined forward”
</td>
</tr>
<tr id="F082FD1725660033FF00FAC32BE0F406" box="[166,1421,1285,1342]" gridrow="21" pageId="6" pageNumber="57">
<td id="B353946B25660033FF00FAC32FD1F406" box="[166,444,1285,1342]" gridcol="0" gridrow="21" pageId="6" pageNumber="57">
<taxonomicName id="89B284D62566FFCAFF00FAE02F3FF406" authorityName="Ochi, Kon &amp; Kawahara" authorityYear="2019" box="[166,338,1318,1342]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="popaensis" subGenus="Paracopris">
<emphasis id="7CC623472566FFCAFF00FAE02ED0F405" box="[166,189,1318,1341]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
(
<emphasis id="7CC623472566FFCAFF6AFAE12EB2F406" box="[204,223,1319,1342]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFF4BFAE12F3FF406" box="[237,338,1319,1342]" italics="true" pageId="6" pageNumber="57">popaensis</emphasis>
</taxonomicName>
(**)
</td>
<td id="B353946B25660033FDAEFAC32BE0F406" box="[520,1421,1285,1342]" gridcol="1" gridrow="21" pageId="6" pageNumber="57">
(
<bibRefCitation id="2A2382A42566FFCAFDA9FAC32CAEF425" author="Ochi, T. &amp; Kon, M. &amp; Kawahara, M." box="[527,707,1285,1310]" pageId="6" pageNumber="57" pagination="469 - 478" refId="ref8655" refString="Ochi, T., Kon, M. &amp; Kawahara, M. (2019 a). Notes on the Coprophagous Scarab beetles from Southeast Asia. XXXII Four new taxa of the genus Copris from Myanmar (Coleoptera, Scarabaeidae). Giornale Italiano di Entomologia, 15 (64), 469 - 478." type="journal article" year="2019">
Ochi 
<emphasis id="7CC623472566FFCAFDEEFAC02C16F425" box="[584,635,1285,1309]" italics="true" pageId="6" pageNumber="57">et al.</emphasis>
2019a
</bibRefCitation>
). “Clypeus also with another stout subtriangular horn which is situated at median portion
</td>
</tr>
<tr id="F082FD1725660033FF00FA8F2BE0F45A" box="[166,1421,1353,1378]" gridrow="22" pageId="6" pageNumber="57">
<td id="B353946B25660033FF00FA8F2BE0F45A" box="[166,1421,1353,1378]" colspan="2" colspanRight="1" gridcol="0" gridrow="22" pageId="6" pageNumber="57">
of clypeus, ca. 
<quantity id="894A52B02566FFCAFD07FA8C2C83F459" box="[673,750,1354,1378]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="57" unit="mm" value="2.1">2.1 mm</quantity>
in length in lateral view, flat on anterior face, strongly 
<tr id="F082FD1725660033FF00FAAB2BE0F49F" box="[166,1421,1389,1447]" gridrow="23" pageId="6" pageNumber="57">
<td id="B353946B25660033FF00FAAB2FD1F49F" box="[166,444,1389,1447]" gridcol="0" gridrow="23" pageId="6" pageNumber="57">
<emphasis id="7CC623472566FFCAFABBFA8C2ED0F49E" italics="true" pageId="6" pageNumber="57">comC.</emphasis>
(
<emphasis id="7CC623472566FFCAFF6AFA492EB2F49E" box="[204,223,1423,1446]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFF4BFA492F0BF49F" box="[237,358,1423,1447]" italics="true" pageId="6" pageNumber="57">punjabensis</emphasis>
</td>
<td id="B353946B25660033FDAEFAAB2BE0F49F" box="[520,1421,1389,1447]" gridcol="1" gridrow="23" pageId="6" pageNumber="57">
pressed on both sides and bidentate at apex” (
<bibRefCitation id="2A2382A42566FFCAFC73FAAB2AE4F4BE" author="Ochi, T. &amp; Kon, M. &amp; Kawahara, M." box="[981,1161,1389,1414]" pageId="6" pageNumber="57" pagination="469 - 478" refId="ref8655" refString="Ochi, T., Kon, M. &amp; Kawahara, M. (2019 a). Notes on the Coprophagous Scarab beetles from Southeast Asia. XXXII Four new taxa of the genus Copris from Myanmar (Coleoptera, Scarabaeidae). Giornale Italiano di Entomologia, 15 (64), 469 - 478." type="journal article" year="2019">
Ochi 
<emphasis id="7CC623472566FFCAFBA9FAA92A2FF4BE" box="[1039,1090,1390,1414]" italics="true" pageId="6" pageNumber="57">et al.</emphasis>
2019a
</bibRefCitation>
). 
<taxonomicName id="89B284D62566FFCAFDAEFA482C3AF49F" authority="Clypeal" authorityName="Clypeal" box="[520,599,1422,1447]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="punjabensis" subGenus="Paracopris">Clypeal</taxonomicName>
apex shallowly emarginated; middle part of emargination protruded, forming a
</td>
</tr>
</td>
</tr>
<tr id="F082FD1725660033FF00FA742BE0F4D4" box="[166,1421,1458,1516]" gridrow="24" pageId="6" pageNumber="57">
<td id="B353946B25660033FF00FA742FD1F4D4" box="[166,444,1458,1516]" gridcol="0" gridrow="24" pageId="6" pageNumber="57">
<emphasis id="7CC623472566FFCAFF00FA122ED0F4D3" box="[166,189,1492,1515]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
(
<emphasis id="7CC623472566FFCAFF6AFA122EB2F4D3" box="[204,223,1492,1515]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFF4BFA122F0DF4D3" box="[237,352,1492,1515]" italics="true" pageId="6" pageNumber="57">ramosiceps</emphasis>
</td>
<td id="B353946B25660033FDAEFA742BE0F4D4" box="[520,1421,1458,1516]" gridcol="1" gridrow="24" pageId="6" pageNumber="57">
rounded tongue-like lobe. 
<taxonomicName id="89B284D62566FFCAFDAEFA152C3AF4D4" authority="Clypeal" authorityName="Clypeal" box="[520,599,1491,1516]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="ramosiceps" subGenus="Paracopris">Clypeal</taxonomicName>
apex protruded, forming one upwards protruded horn which widely bifurcated;
</td>
</tr>
<tr id="F082FD1725660033FF00FA312BE0F708" box="[166,1421,1527,1584]" gridrow="25" pageId="6" pageNumber="57">
<td id="B353946B25660033FF00FA312FD1F708" box="[166,444,1527,1584]" gridcol="0" gridrow="25" pageId="6" pageNumber="57">
<emphasis id="7CC623472566FFCAFF00F9DE2ED0F717" box="[166,189,1560,1583]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
(
<emphasis id="7CC623472566FFCAFF6AF9DF2EB2F708" box="[204,223,1561,1584]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFF4BF9DF2F2CF708" box="[237,321,1561,1584]" italics="true" pageId="6" pageNumber="57">signatus</emphasis>
</td>
<td id="B353946B25660033FDAEFA312BE0F708" box="[520,1421,1527,1584]" gridcol="1" gridrow="25" pageId="6" pageNumber="57">
two branches upwards and backwards curved. 
<taxonomicName id="89B284D62566FFCAFDAEF9D12C3AF708" authority="Clypeal" authorityName="Clypeal" box="[520,599,1559,1584]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="signatus" subGenus="Paracopris">Clypeal</taxonomicName>
apex with two narrow and almost parallel processes.
</td>
</tr>
</table>
</paragraph>
<paragraph id="4E0DFF552566FFCAFF31F9F92A5CF687" blockId="6.[151,1437,1599,1983]" pageId="6" pageNumber="57">
(*) 
<bibRefCitation id="2A2382A42566FFCAFF66F9F92FABF76F" author="Ochi, T. &amp; Kon, M." box="[192,454,1599,1623]" pageId="6" pageNumber="57" pagination="29 - 35" refId="ref8462" refString="Ochi, T. &amp; Kon, M. (1996) Studies on the coprophagous scarab beetles from East Asia (IV) (Coleoptera: Scarabaeidae). Gior- nale Italiano di Entomologia, 8, 29 - 35." type="journal article" year="1996">Ochi &amp; Kon (1996: 25)</bibRefCitation>
originally showed the clypeus of 
<taxonomicName id="89B284D62566FFCAFCE7F9F92A1CF76F" authorityName="Ochi &amp; Kon" authorityYear="1996" box="[833,1137,1599,1623]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="kasagii" subGenus="Paracopris">
<emphasis id="7CC623472566FFCAFCE7F9F92DE7F76E" box="[833,906,1599,1622]" italics="true" pageId="6" pageNumber="57">Copris</emphasis>
(
<emphasis id="7CC623472566FFCAFC3FF9F92A7CF76E" box="[921,1041,1599,1622]" italics="true" pageId="6" pageNumber="57">Paracopris</emphasis>
) 
<emphasis id="7CC623472566FFCAFB82F9F92A1CF76F" box="[1060,1137,1599,1623]" italics="true" pageId="6" pageNumber="57">kasagii</emphasis>
</taxonomicName>
with distinct emargination: “Clypeus produced as a reflexed point in the middle, which is medially notched as obtuse angle and distinctly bidentate, from anterior aspect”, though the illustrated photos (
<bibRefCitation id="2A2382A42566FFCAFD6AF9412DD1F7A7" author="Ochi, T. &amp; Kon, M." box="[716,956,1671,1695]" pageId="6" pageNumber="57" pagination="29 - 35" refId="ref8462" refString="Ochi, T. &amp; Kon, M. (1996) Studies on the coprophagous scarab beetles from East Asia (IV) (Coleoptera: Scarabaeidae). Gior- nale Italiano di Entomologia, 8, 29 - 35." type="journal article" year="1996">Ochi &amp; Kon 1996: 26</bibRefCitation>
) showed an indistinctly emarginated clypeal anterior margin. However, both the description and photos of 
<taxonomicName id="89B284D62566FFCAFCE5F96D2DB3F7FB" authorityName="Ochi &amp; Kon" authorityYear="1996" box="[835,990,1707,1731]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="kasagii" subGenus="Paracopris">
<emphasis id="7CC623472566FFCAFCE5F96D2D31F7FA" box="[835,860,1707,1730]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
(
<emphasis id="7CC623472566FFCAFCCDF96A2D12F7FB" box="[875,895,1708,1731]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFC37F96D2DB3F7FB" box="[913,990,1707,1731]" italics="true" pageId="6" pageNumber="57">kasagii</emphasis>
</taxonomicName>
by 
<bibRefCitation id="2A2382A42566FFCAFBAAF96D2A8BF7FB" author="Ochi, T. &amp; Kon, M." box="[1036,1254,1707,1731]" pageId="6" pageNumber="57" pagination="29 - 35" refId="ref8462" refString="Ochi, T. &amp; Kon, M. (1996) Studies on the coprophagous scarab beetles from East Asia (IV) (Coleoptera: Scarabaeidae). Gior- nale Italiano di Entomologia, 8, 29 - 35." type="journal article" year="1996">Ochi &amp; Kon (1996)</bibRefCitation>
showed that the clypeal anterior margin was not protruded, clearly distinguishing 
<taxonomicName id="89B284D62566FFCAFCF7F9092D84F7DF" authorityName="Ochi &amp; Kon" authorityYear="1996" box="[849,1001,1743,1767]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="kasagii" subGenus="Paracopris">
<emphasis id="7CC623472566FFCAFCF7F9092D07F7DE" box="[849,874,1743,1766]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
(
<emphasis id="7CC623472566FFCAFCDFF9162DE3F7DF" box="[889,910,1744,1767]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFC3AF9092D84F7DF" box="[924,1001,1743,1767]" italics="true" pageId="6" pageNumber="57">kasagii</emphasis>
</taxonomicName>
from the new species. (**) We have not been able to examine the following five newly discovered species: 
<taxonomicName id="89B284D62566FFCAFB99F9352BF8F633" authorityName="Ochi, Kon &amp; Pham" authorityYear="2019" box="[1087,1429,1779,1803]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="bartolozzii" subGenus="Paracopris">
<emphasis id="7CC623472566FFCAFB99F9352AE5F632" box="[1087,1160,1779,1802]" italics="true" pageId="6" pageNumber="57">Copris</emphasis>
(
<emphasis id="7CC623472566FFCAFB3CF9352B7FF632" box="[1178,1298,1779,1802]" italics="true" pageId="6" pageNumber="57">Paracopris</emphasis>
) 
<emphasis id="7CC623472566FFCAFA85F9352BF8F633" box="[1315,1429,1779,1803]" italics="true" pageId="6" pageNumber="57">bartolozzii</emphasis>
</taxonomicName>
, 
<taxonomicName id="89B284D62566FFCAFF31F8D12F14F617" authorityName="Ochi, Kon &amp; Kawahara" authorityYear="2019" box="[151,377,1815,1839]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="chinmontanus" subGenus="Paracopris">
<emphasis id="7CC623472566FFCAFF31F8D12EDDF616" box="[151,176,1815,1838]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
(
<emphasis id="7CC623472566FFCAFF66F8DE2EB9F617" box="[192,212,1816,1839]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFF45F8D12F14F617" box="[227,377,1815,1839]" italics="true" pageId="6" pageNumber="57">chinmontanus</emphasis>
</taxonomicName>
, 
<taxonomicName id="89B284D62566FFCAFE20F8D12C2DF617" authorityName="Ochi, Kon &amp; Pham" authorityYear="2019" box="[390,576,1815,1839]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="montreuili" subGenus="Paracopris">
<emphasis id="7CC623472566FFCAFE20F8D12FF2F616" box="[390,415,1815,1838]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
(
<emphasis id="7CC623472566FFCAFE09F8DE2FAEF617" box="[431,451,1816,1839]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFE74F8D12C2DF617" box="[466,576,1815,1839]" italics="true" pageId="6" pageNumber="57">montreuili</emphasis>
</taxonomicName>
, 
<taxonomicName id="89B284D62566FFCAFDE8F8D12C8BF617" authorityName="Ochi, Kon &amp; Kawahara" authorityYear="2019" box="[590,742,1815,1839]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="olivieri" subGenus="Paracopris">
<emphasis id="7CC623472566FFCAFDE8F8D12C0BF616" box="[590,614,1815,1838]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
(
<emphasis id="7CC623472566FFCAFDD0F8DE2CE6F617" box="[630,651,1816,1839]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFD3FF8D12C8BF617" box="[665,742,1815,1839]" italics="true" pageId="6" pageNumber="57">olivieri</emphasis>
</taxonomicName>
and 
<taxonomicName id="89B284D62566FFCAFCBDF8D12DBFF616" authorityName="Ochi, Kon &amp; Kawahara" authorityYear="2019" box="[795,978,1815,1839]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="popaensis" subGenus="Paracopris">
<emphasis id="7CC623472566FFCAFCBDF8D12D59F616" box="[795,820,1815,1838]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
(
<emphasis id="7CC623472566FFCAFCE5F8DE2D35F617" box="[835,856,1816,1839]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFCC1F8D12DBFF616" box="[871,978,1815,1838]" italics="true" pageId="6" pageNumber="57">popaensis</emphasis>
</taxonomicName>
. The original descriptions and illustrations showed clear separation in clypeal morphological characters between these five species and 
<taxonomicName id="89B284D62566FFCAFBDEF8FC2BF1F66A" authority="Bui &amp; Ziegler &amp; Bonkowski, 2019" authorityName="new" authorityYear="2019" box="[1144,1436,1850,1876]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="javanensis" status="new" subGenus="Paracopris">
<emphasis id="7CC623472566FFCAFBDEF8FC2AA9F66C" box="[1144,1220,1850,1876]" italics="true" pageId="6" pageNumber="57">Copris</emphasis>
(
<emphasis id="7CC623472566FFCAFB72F8FA2A84F66B" box="[1236,1257,1852,1875]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFB5EF8FD2B04F66A" box="[1272,1385,1851,1874]" italics="true" pageId="6" pageNumber="57">javanensis</emphasis>
new
</taxonomicName>
species (
<tableCitation id="0330CAEE2566FFCAFF55F8992F2AF64F" box="[243,327,1887,1911]" captionStart="TABLE 2" captionStartId="6.[151,239,154,178]" captionTargetBox="[166,1421,261,1584]" captionText="TABLE 2. Diagnostic characters of clypeus separating the new species from the other Copris (Paracopris) species (based on the specimens examined by us in MNHN and NMPC, and data published by Wiedemann 1823; Arrow 1931; Balthasar 1939, 1963; Ochi &amp; Kon 1996; Hanboonsong et al. 2003; Ochi et al. 2019a,b)" httpUri="http://table.plazi.org/id/1ACDAFDD2566FFCAFF31FF5C2DB7F1C2" pageId="6" pageNumber="57" tableUuid="1ACDAFDD2566FFCAFF31FF5C2DB7F1C2">Table 2</tableCitation>
). In addition, the densely to contiguously punctate elytra of 
<taxonomicName id="89B284D62566FFCAFC69F8992AE0F64F" authorityName="Ochi, Kon &amp; Pham" authorityYear="2019" box="[975,1165,1887,1911]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="bartolozzii" subGenus="Paracopris">
<emphasis id="7CC623472566FFCAFC69F8992D8AF64E" box="[975,999,1887,1910]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
(
<emphasis id="7CC623472566FFCAFC51F8A62A61F64F" box="[1015,1036,1888,1911]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFBBDF8992AE0F64F" box="[1051,1165,1887,1911]" italics="true" pageId="6" pageNumber="57">bartolozzii</emphasis>
</taxonomicName>
, 
<emphasis id="7CC623472566FFCAFB3DF8992BF0F64E" box="[1179,1437,1887,1911]" italics="true" pageId="6" pageNumber="57">
<taxonomicName id="89B284D62566FFCAFB3DF8992B15F64F" authorityName="Ochi, Kon &amp; Kawahara" authorityYear="2019" box="[1179,1400,1887,1911]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="chinmontanus" subGenus="Paracopris">C.(P.) chinmontanus</taxonomicName>
, 
<taxonomicName id="89B284D62566FFCAFA22F8992BF0F64E" authorityName="Ochi, Kon &amp; Pham" authorityYear="2019" box="[1412,1437,1887,1910]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="montreuili" subGenus="Paracopris">C.</taxonomicName>
</emphasis>
(
<emphasis id="7CC623472566FFCAFF06F8422ED8F6A3" box="[160,181,1924,1947]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFF62F8452F5FF6A3" box="[196,306,1923,1947]" italics="true" pageId="6" pageNumber="57">montreuili</emphasis>
and 
<taxonomicName id="89B284D62566FFCAFEC0F8452C70F6A2" authorityName="Ochi, Kon &amp; Kawahara" authorityYear="2019" box="[358,541,1923,1947]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="popaensis" subGenus="Paracopris">
<emphasis id="7CC623472566FFCAFEC0F8452F12F6A2" box="[358,383,1923,1946]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
(
<emphasis id="7CC623472566FFCAFE29F8422FCEF6A3" box="[399,419,1924,1947]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFE14F8452C70F6A2" box="[434,541,1923,1946]" italics="true" pageId="6" pageNumber="57">popaensis</emphasis>
</taxonomicName>
differentiate these species from 
<taxonomicName id="89B284D62566FFCAFCDCF8442AF3F6A2" authority="Bui &amp; Ziegler &amp; Bonkowski, 2019" authorityName="new" authorityYear="2019" box="[890,1182,1922,1948]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="javanensis" status="new" subGenus="Paracopris">
<emphasis id="7CC623472566FFCAFCDCF8442DABF6A4" box="[890,966,1922,1948]" italics="true" pageId="6" pageNumber="57">Copris</emphasis>
(
<emphasis id="7CC623472566FFCAFC70F8422D86F6A3" box="[982,1003,1924,1947]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFC5CF8452A06F6A2" box="[1018,1131,1923,1946]" italics="true" pageId="6" pageNumber="57">javanensis</emphasis>
<taxonomicNameLabel id="67F59E3C2566FFCAFBD4F8452AF3F6A2" box="[1138,1182,1923,1946]" pageId="6" pageNumber="57" rank="species">new</taxonomicNameLabel>
</taxonomicName>
species, while the deep median sulcus of pronotal disc of 
<taxonomicName id="89B284D62566FFCAFDA7F8612CF4F687" authorityName="Ochi, Kon &amp; Kawahara" authorityYear="2019" box="[513,665,1959,1983]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="olivieri" subGenus="Paracopris">
<emphasis id="7CC623472566FFCAFDA7F8612C74F686" box="[513,537,1959,1982]" italics="true" pageId="6" pageNumber="57">C.</emphasis>
(
<emphasis id="7CC623472566FFCAFD8FF86E2C50F687" box="[553,573,1960,1983]" italics="true" pageId="6" pageNumber="57">P.</emphasis>
) 
<emphasis id="7CC623472566FFCAFDEAF8612CF4F687" box="[588,665,1959,1983]" italics="true" pageId="6" pageNumber="57">olivieri</emphasis>
</taxonomicName>
differentiates it from the new species.
</paragraph>
</subSubSection>
</treatment>
</document>