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<document id="4C5D515E82159167E3E76CC0CB070F04" ID-DOI="10.5281/zenodo.202575" ID-GBIF-Dataset="e6fc0416-99c9-495a-af89-35115e9a1675" ID-ISSN="1175-5326" ID-Zenodo-Dep="202575" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460302833884" checkinUser="plazi" docAuthor="Pietrykowska-Tudruj, Ewa &amp; Staniec, Bernard" docDate="2011" docId="7E75879EFF97FFE06ACC054D0FEAF90C" docLanguage="en" docName="zt02865p067.pdf" docOrigin="Zootaxa 2865" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Bisnius fimetarius Gravenhorst" docType="treatment" docVersion="9" lastPageNumber="56" masterDocId="824CFFE6FF94FFE36A5B01220E7FFFC9" masterDocTitle="The morphology of the pupae of six species of Philonthina (Coleoptera, Staphylinidae, Staphylinini) with taxonomic remarks" masterLastPageNumber="67" masterPageNumber="53" pageNumber="56" updateTime="1698252825721" updateUser="plazi">
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<mods:title id="06B6FC00B9A35AE5BB48D6A2CBAA2022">The morphology of the pupae of six species of Philonthina (Coleoptera, Staphylinidae, Staphylinini) with taxonomic remarks</mods:title>
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<mods:namePart id="2F3DE9864D4F25EE8C81D26766594733">Pietrykowska-Tudruj, Ewa</mods:namePart>
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<mods:namePart id="6B6EFB7E86893F038A22EE25383DD129">Staniec, Bernard</mods:namePart>
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<mods:date id="B737E439AF9EE83307CE4BCFB9D66E60">2011</mods:date>
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<treatment id="7E75879EFF97FFE06ACC054D0FEAF90C" ID-DOI="http://doi.org/10.5281/zenodo.6183483" ID-GBIF-Taxon="119400051" ID-Zenodo-Dep="6183483" LSID="urn:lsid:plazi:treatment:7E75879EFF97FFE06ACC054D0FEAF90C" httpUri="http://treatment.plazi.org/id/7E75879EFF97FFE06ACC054D0FEAF90C" lastPageNumber="56" pageId="3" pageNumber="56">
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<paragraph id="F6633688FF97FFE06ACC054D0C50FB40" blockId="3.[151,559,1135,1194]" box="[151,559,1135,1161]" pageId="3" pageNumber="56">
<heading id="AD2B81E4FF97FFE06ACC054D0C50FB40" bold="true" box="[151,559,1135,1161]" fontSize="11" level="1" pageId="3" pageNumber="56" reason="1">
<taxonomicName id="31DC4D0BFF97FFE06ACC054D0C50FB40" ID-CoL="LWVR" authority="Gravenhorst" authorityName="Gravenhorst" box="[151,559,1135,1161]" class="Insecta" family="Staphylinidae" genus="Bisnius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="fimetarius">
<emphasis id="C4A8EA9AFF97FFE06ACC054D0C50FB40" bold="true" box="[151,559,1135,1161]" pageId="3" pageNumber="56">
<emphasis id="C4A8EA9AFF97FFE06ACC054D0F0BFB40" bold="true" box="[151,372,1135,1161]" italics="true" pageId="3" pageNumber="56">Bisnius fimetarius</emphasis>
(Gravenhorst)
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</heading>
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<paragraph id="F6633688FF97FFE06ACC05B00FA2FB63" blockId="3.[151,559,1135,1194]" box="[151,477,1170,1194]" pageId="3" pageNumber="56">
(
<figureCitation id="6EE72A0DFF97FFE06AC405B00F7CFB63" box="[159,259,1170,1194]" captionStart="FIGURES 1 6" captionStartId="4.[151,264,1962,1985]" captionTargetBox="[201,1371,187,1936]" captionTargetId="figure@4.[199,1376,187,1937]" captionTargetPageId="4" captionText="FIGURES 1 6. Pupa of Bisnius fimentarius (1 3) and Gabrius osseticus (4 6). (1, 4) Ventral aspect, (2, 5) lateral aspect, (3, 6) dorsal aspect." httpUri="https://zenodo.org/record/202576/files/figure.png" pageId="3" pageNumber="56">Figs 13</figureCitation>
, 4142, 52, 61, 62)
</paragraph>
<paragraph id="F6633688FF97FFE06ACC05F60FEAF90C" blockId="3.[151,1436,1236,1733]" pageId="3" pageNumber="56">
Body length:
<quantity id="31249B6DFF97FFE06B7405F60FADFB25" box="[303,466,1236,1260]" metricMagnitude="-3" metricUnit="m" metricValue="3.985" metricValueMax="4.4" metricValueMin="3.57" pageId="3" pageNumber="56" unit="mm" value="3.985" valueMax="4.4" valueMin="3.57">3.574.40 mm</quantity>
(mean
<quantity id="31249B6DFF97FFE0687D05F60CF1FB25" box="[550,654,1236,1260]" metricMagnitude="-3" metricUnit="m" metricValue="4.16" pageId="3" pageNumber="56" unit="mm" value="4.16">4.16 mm</quantity>
); body width:
<quantity id="31249B6DFF97FFE0697405F60DADFB25" box="[815,978,1236,1260]" metricMagnitude="-3" metricUnit="m" metricValue="1.635" metricValueMax="1.84" metricValueMin="1.43" pageId="3" pageNumber="56" unit="mm" value="1.635" valueMax="1.84" valueMin="1.43">1.431.84 mm</quantity>
(mean
<quantity id="31249B6DFF97FFE06E7D05F60AF1FB25" box="[1062,1166,1236,1260]" metricMagnitude="-3" metricUnit="m" metricValue="1.67" pageId="3" pageNumber="56" unit="mm" value="1.67">1.67 mm</quantity>
); head width:
<quantity id="31249B6DFF97FFE06F7705F60EBCFAC6" metricMagnitude="-3" metricUnit="m" metricValue="1.02" metricValueMax="1.24" metricValueMin="0.8" pageId="3" pageNumber="56" unit="mm" value="1.02" valueMax="1.24" valueMin="0.8">0.801.24 mm</quantity>
(mean
<quantity id="31249B6DFF97FFE06B4C05D50F01FAC6" box="[279,382,1271,1295]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="3" pageNumber="56" unit="mm" value="1.03">1.03 mm</quantity>
); pronotum width:
<quantity id="31249B6DFF97FFE0680F05D50C89FAC6" box="[596,758,1271,1295]" metricMagnitude="-3" metricUnit="m" metricValue="1.14" metricValueMax="1.17" metricValueMin="1.11" pageId="3" pageNumber="56" unit="mm" value="1.14" valueMax="1.17" valueMin="1.11">1.111.17 mm</quantity>
(mean
<quantity id="31249B6DFF97FFE0691105D50DCDFAC6" box="[842,946,1271,1295]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="3" pageNumber="56" unit="mm" value="1.15">1.15 mm</quantity>
), about 1.1 x as broad as long; body moderately stocky and weakly sclerotized. Antennae reaching to about half of elytra length (
<figureCitation id="6EE72A0DFF97FFE06E31043E0ACEFAFD" box="[1130,1201,1308,1332]" captionStart="FIGURES 1 6" captionStartId="4.[151,264,1962,1985]" captionTargetBox="[201,1371,187,1936]" captionTargetId="figure@4.[199,1376,187,1937]" captionTargetPageId="4" captionText="FIGURES 1 6. Pupa of Bisnius fimentarius (1 3) and Gabrius osseticus (4 6). (1, 4) Ventral aspect, (2, 5) lateral aspect, (3, 6) dorsal aspect." httpUri="https://zenodo.org/record/202576/files/figure.png" pageId="3" pageNumber="56">Fig. 2</figureCitation>
). Anterior margin of pronotum with 1014 projections (6/6, 7/7, 7/7, 5/5, 7/5, 6/5, 6/5 projections on sides in individuals). Wings almost reaching the posterior margin of 1st (morphologically 3rd) clearly visible abdominal segment (
<figureCitation id="6EE72A0DFF97FFE06EF604450B6CFAB6" box="[1197,1299,1383,1407]" captionStart="FIGURES 1 6" captionStartId="4.[151,264,1962,1985]" captionTargetBox="[201,1371,187,1936]" captionTargetId="figure@4.[199,1376,187,1937]" captionTargetPageId="4" captionText="FIGURES 1 6. Pupa of Bisnius fimentarius (1 3) and Gabrius osseticus (4 6). (1, 4) Ventral aspect, (2, 5) lateral aspect, (3, 6) dorsal aspect." httpUri="https://zenodo.org/record/202576/files/figure.png" pageId="3" pageNumber="56">Figs 1, 2</figureCitation>
). Each fore, middle, and hind tibiae with 5, 810, and 6 protuberances respectively. Tarsi of hind legs slightly protruding from anterior margin of 4th (morphologically 6th) clearly visible abdominal segment (
<figureCitation id="6EE72A0DFF97FFE06E7E04930AF0FA00" box="[1061,1167,1457,1481]" captionStart="FIGURES 1 6" captionStartId="4.[151,264,1962,1985]" captionTargetBox="[201,1371,187,1936]" captionTargetId="figure@4.[199,1376,187,1937]" captionTargetPageId="4" captionText="FIGURES 1 6. Pupa of Bisnius fimentarius (1 3) and Gabrius osseticus (4 6). (1, 4) Ventral aspect, (2, 5) lateral aspect, (3, 6) dorsal aspect." httpUri="https://zenodo.org/record/202576/files/figure.png" pageId="3" pageNumber="56">Figs 1, 2</figureCitation>
). Abdomen moderately and gradually widened from segment II to IV and narrowed to the terminal part of the body (
<figureCitation id="6EE72A0DFF97FFE06E9A04F70B79FA24" box="[1217,1286,1493,1517]" captionStart="FIGURES 1 6" captionStartId="4.[151,264,1962,1985]" captionTargetBox="[201,1371,187,1936]" captionTargetId="figure@4.[199,1376,187,1937]" captionTargetPageId="4" captionText="FIGURES 1 6. Pupa of Bisnius fimentarius (1 3) and Gabrius osseticus (4 6). (1, 4) Ventral aspect, (2, 5) lateral aspect, (3, 6) dorsal aspect." httpUri="https://zenodo.org/record/202576/files/figure.png" pageId="3" pageNumber="56">Fig. 3</figureCitation>
). Abdominal tergites with microstructure as in
<taxonomicName id="31DC4D0BFF97FFE0685404D80CECF9D8" box="[527,659,1530,1553]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="osseticus">
<emphasis id="C4A8EA9AFF97FFE0685404D80CECF9D8" box="[527,659,1530,1553]" italics="true" pageId="3" pageNumber="56">G. osseticus</emphasis>
</taxonomicName>
(
<figureCitation id="6EE72A0DFF97FFE068F804DB0C8BF9D8" box="[675,756,1529,1553]" captionStart="FIGURES 19 40" captionStartId="9.[151,264,1885,1908]" captionTargetBox="[232,1328,193,1860]" captionTargetId="figure@9.[216,1345,193,1863]" captionTargetPageId="9" captionText="FIGURES 19 40. Pupa of Gabrius osseticus (19, 20), Philonthus atratus (21 25), P. carbonarius (26 32), P. debilis (33 35) and P. decorus (36 40). (19, 21, 26, 27, 36) Lateral part of segment VII with microstructure (19 A, 26 A) and spine (27); (20, 22, 33, 39, 40) lateral part of segment VIII with microstructure (39 A) and spine (40); (23, 24, 28, 29) lateral part of segment IV with spine (24, 29); (25, 31, 32, 34, 35, 37, 38) lateral part of segment V with microstructure (34 A) and spine (32, 35, 38)." httpUri="https://zenodo.org/record/202579/files/figure.png" pageId="3" pageNumber="56">Fig. 19</figureCitation>
A). Abdominal segments VII and VIII each bearing a pair of setiform projections on sides, projections similar in length and structure to these one in
<taxonomicName id="31DC4D0BFF97FFE06E2C073C0A84F9FF" box="[1143,1275,1566,1590]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="osseticus">
<emphasis id="C4A8EA9AFF97FFE06E2C073C0A84F9FF" box="[1143,1275,1566,1590]" italics="true" pageId="3" pageNumber="56">G. osseticus</emphasis>
</taxonomicName>
(
<figureCitation id="6EE72A0DFF97FFE06F50073F0BEFF9FC" box="[1291,1424,1565,1589]" captionStart="FIGURES 19 40" captionStartId="9.[151,264,1885,1908]" captionTargetBox="[232,1328,193,1860]" captionTargetId="figure@9.[216,1345,193,1863]" captionTargetPageId="9" captionText="FIGURES 19 40. Pupa of Gabrius osseticus (19, 20), Philonthus atratus (21 25), P. carbonarius (26 32), P. debilis (33 35) and P. decorus (36 40). (19, 21, 26, 27, 36) Lateral part of segment VII with microstructure (19 A, 26 A) and spine (27); (20, 22, 33, 39, 40) lateral part of segment VIII with microstructure (39 A) and spine (40); (23, 24, 28, 29) lateral part of segment IV with spine (24, 29); (25, 31, 32, 34, 35, 37, 38) lateral part of segment V with microstructure (34 A) and spine (32, 35, 38)." httpUri="https://zenodo.org/record/202579/files/figure.png" pageId="3" pageNumber="56">Figs 19, 20</figureCitation>
). Terminal sternite of female pupa with two well-developed ventral prolongations (Fig. 41); in male pupa sternite IX as in Fig. 42; ventral and terminal prolongation with numerous, sharp, cuticular processes (Figs 41A, B). The first pair of functional spiracles elongated, gradually narrowed apically, almost as long as wide (
<figureCitation id="6EE72A0DFF97FFE06EC907AB0A9AF968" box="[1170,1253,1673,1697]" captionStart="FIGURES 52 70" captionStartId="11.[151,264,1916,1939]" captionTargetBox="[257,1311,181,1889]" captionTargetId="figure@11.[256,1312,181,1890]" captionTargetPageId="11" captionText="FIGURES 52 70. Pupa of Bisnius fimentarius (52, 61, 62), Gabrius osseticus (53, 58, 63, 64), Philonthus atratus (54, 59, 65), P. carbonarius (55, 66), P. debilis (56, 60, 68 70) and P. d e c o r u s (57, 67). (52 60) The first pair of functional spiracles; (61 70) atrophied spiracles." httpUri="https://zenodo.org/record/202580/files/figure.png" pageId="3" pageNumber="56">Fig. 52</figureCitation>
); atrophied spiracles as in
<figureCitation id="6EE72A0DFF97FFE06B4A078F0FEEF90C" box="[273,401,1709,1733]" captionStart="FIGURES 52 70" captionStartId="11.[151,264,1916,1939]" captionTargetBox="[257,1311,181,1889]" captionTargetId="figure@11.[256,1312,181,1890]" captionTargetPageId="11" captionText="FIGURES 52 70. Pupa of Bisnius fimentarius (52, 61, 62), Gabrius osseticus (53, 58, 63, 64), Philonthus atratus (54, 59, 65), P. carbonarius (55, 66), P. debilis (56, 60, 68 70) and P. d e c o r u s (57, 67). (52 60) The first pair of functional spiracles; (61 70) atrophied spiracles." httpUri="https://zenodo.org/record/202580/files/figure.png" pageId="3" pageNumber="56">Figs 61, 62</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>