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<document id="295F2DC2F4292FF61112554A1680562C" ID-DOI="10.11646/zootaxa.3981.4.1" ID-GBIF-Dataset="15c3bf52-4481-4a75-8643-7c919bdde6c4" ID-ISSN="1175-5326" ID-Zenodo-Dep="235846" ID-ZooBank="EF71D83B-17B4-49CA-826E-D3A8E7979750" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461248924452" checkinUser="plazi" docAuthor="Zwick, Peter" docDate="2015" docId="5C5BE52CFFB2BC5F2CB5F8E793C50FE0" docLanguage="en" docName="zt03981p490.pdf" docOrigin="Zootaxa 3981 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Contacyphon forcipatus Zwick, 2015, n. sp." docType="treatment" docVersion="8" lastPageNumber="487" masterDocId="A0629D54FF91BC7B2C22FFE99334080F" masterDocTitle="Australian Marsh Beetles (Coleoptera: Scirtidae). 8. The new genera Cygnocyphon, Eximiocyphon, Paracyphon, Leptocyphon, Tectocyphon, and additions to Contacyphon de Gozis, Nanocyphon Zwick and Eurycyphon Watts" masterLastPageNumber="490" masterPageNumber="451" pageNumber="486" updateTime="1698607092210" updateUser="plazi">
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<mods:title id="B0557B4537D11530B0B1B9F02D9AE625">Australian Marsh Beetles (Coleoptera: Scirtidae). 8. The new genera Cygnocyphon, Eximiocyphon, Paracyphon, Leptocyphon, Tectocyphon, and additions to Contacyphon de Gozis, Nanocyphon Zwick and Eurycyphon Watts</mods:title>
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<mods:date id="55877163B51CD0AA0A4CFFA03FCB1B8B">2015</mods:date>
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<paragraph id="D44D543AFFB2BC582CB5F8E7913E0F27" blockId="35.[151,522,1806,1866]" box="[151,522,1806,1832]" pageId="35" pageNumber="486">
<heading id="8F05E356FFB2BC582CB5F8E7913E0F27" bold="true" box="[151,522,1806,1832]" fontSize="11" level="1" pageId="35" pageNumber="486" reason="1">
<emphasis id="E6868828FFB2BC582CB5F8E7913E0F27" bold="true" box="[151,522,1806,1832]" pageId="35" pageNumber="486">
<emphasis id="E6868828FFB2BC582CB5F8E792F50F27" bold="true" box="[151,449,1806,1832]" italics="true" pageId="35" pageNumber="486">
<taxonomicName id="13F22FB9FFB2BC582CB5F8E7928F0F27" box="[151,443,1806,1832]" class="Insecta" family="Scirtidae" genus="Contacyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="486" phylum="Arthropoda" rank="species" species="forcipatus" status="sp. nov.">Contacyphon forcipatus</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="FDB53553FFB2BC582DEAF8E7913E0F27" box="[456,522,1806,1832]" pageId="35" pageNumber="486" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="D44D543AFFB2BC582CB5F8D8927E0F45" blockId="35.[151,522,1806,1866]" box="[151,330,1841,1866]" pageId="35" pageNumber="486">
(
<figureCitation id="4CC948BFFFB2BC582CBDF8D892750F45" box="[159,321,1841,1866]" captionStart="FIGURES 103 106" captionStartId="36.[151,264,905,928]" captionTargetBox="[160,1428,209,872]" captionTargetId="figure@36.[153,1434,193,885]" captionTargetPageId="36" captionText="FIGURES 103 106. Contacyphon forcipatus, n. sp., male. 103, segments 8 and 9, superimposed; 104, tegmen; 105, penis (103 105, holotype); 106, penis and tegmen of the paratype, superimposed. All to the same scale." httpUri="https://zenodo.org/record/235863/files/figure.png" pageId="35" pageNumber="486">Figs. 103106</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="9CE807B1FFB2BC5F2CB5F89093E40D85" lastPageId="36" lastPageNumber="487" pageId="35" pageNumber="486" type="materials_examined">
<paragraph id="D44D543AFFB2BC582CB5F89096600FB9" blockId="35.[151,1436,1913,1974]" pageId="35" pageNumber="486">
<emphasis id="E6868828FFB2BC582CB5F89092D90F9D" bold="true" box="[151,493,1913,1938]" pageId="35" pageNumber="486">
<typeStatus id="0B49EA98FFB2BC582CB5F89093E60F9D" box="[151,210,1913,1938]" pageId="35" pageNumber="486">Type</typeStatus>
material:
<specimenCount id="C2F49FB3FFB2BC582D71F89092420F9D" box="[339,374,1913,1938]" pageId="35" pageNumber="486" type="generic">1♂</specimenCount>
<typeStatus id="0B49EA98FFB2BC582DA2F89092D90F9D" box="[384,493,1913,1938]" pageId="35" pageNumber="486" type="holotype">Holotype</typeStatus>
</emphasis>
: Jardine R. Xing
<quantity id="130AF9DFFFB2BC582E9CF893903C0F9D" box="[702,776,1913,1938]" metricMagnitude="4" metricUnit="m" metricValue="2.9" pageId="35" pageNumber="486" unit="km" value="29.0">29 km</quantity>
S Bamaga
<emphasis id="E6868828FFB2BC582FB2F89090FC0F9D" bold="true" box="[912,968,1913,1938]" pageId="35" pageNumber="486">N.Q.</emphasis>
<date id="A04C72FAFFB2BC582FF0F890978D0F9D" box="[978,1209,1913,1938]" pageId="35" pageNumber="486" value="1985-08-28" valueMax="1985-09-05" valueMin="1985-08-28">28.Aug.5.Sep.1985</date>
E.N.Marks coll. ex light trap (
<collectionCode id="B2E3CCFFFFB2BC582D2EF87792630FB9" LSID="urn:lsid:biocol.org:col:32981" box="[268,343,1950,1974]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="35" pageNumber="486">ANIC</collectionCode>
).
<emphasis id="E6868828FFB2BC582D4BF874913D0FB9" bold="true" box="[361,521,1949,1974]" pageId="35" pageNumber="486">
<specimenCount id="C2F49FB3FFB2BC582D4BF87492B80FB9" box="[361,396,1949,1974]" pageId="35" pageNumber="486" type="generic">1♂</specimenCount>
<typeStatus id="0B49EA98FFB2BC582DB1F87491300FB9" box="[403,516,1949,1974]" pageId="35" pageNumber="486" type="paratype">Paratype</typeStatus>
:
</emphasis>
Cow Bay N of Daintree, N.Qld.
<date id="A04C72FAFFB2BC582F59F874973D0FB9" box="[891,1033,1949,1974]" pageId="35" pageNumber="486" value="1984-02-07" valueMax="1984-02-20" valueMin="1984-02-07">720.ii.1984</date>
J.C.Cunningham (QDPC-C).
</paragraph>
<caption id="808D04B2FFB5BC5F2CB5FC6097410BB1" httpUri="https://zenodo.org/record/235863/files/figure.png" pageId="36" pageNumber="487" targetBox="[160,1428,209,872]" targetPageId="36">
<paragraph id="D44D543AFFB5BC5F2CB5FC6097410BB1" blockId="36.[151,1435,905,958]" pageId="36" pageNumber="487">
<emphasis id="E6868828FFB5BC5F2CB5FC6092440B90" bold="true" box="[151,368,905,928]" pageId="36" pageNumber="487">FIGURES 103106.</emphasis>
<emphasis id="E6868828FFB5BC5F2D5AFC6391410BAF" box="[376,629,906,928]" italics="true" pageId="36" pageNumber="487">
<taxonomicName id="13F22FB9FFB5BC5F2D5AFC6391450BAF" box="[376,625,906,928]" class="Insecta" family="Scirtidae" genus="Contacyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="forcipatus" status="sp. nov.">Contacyphon forcipatus</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="FDB53553FFB5BC5F2E5FFC63918D0B90" box="[637,697,906,927]" pageId="36" pageNumber="487" rank="species">
<emphasis id="E6868828FFB5BC5F2E5FFC63918D0B90" bold="true" box="[637,697,906,927]" pageId="36" pageNumber="487">n. sp.</emphasis>
</taxonomicNameLabel>
, male. 103, segments 8 and 9, superimposed; 104, tegmen; 105, penis (103105, holotype); 106, penis and tegmen of the paratype, superimposed. All to the same scale.
</paragraph>
</caption>
<paragraph id="D44D543AFFB5BC5F2CE5FC0F96410C49" blockId="36.[151,1437,998,2031]" pageId="36" pageNumber="487">
<emphasis id="E6868828FFB5BC5F2CE5FC0F92120BF0" bold="true" box="[199,294,998,1023]" pageId="36" pageNumber="487">Habitus</emphasis>
. Oval, BL
<quantity id="130AF9DFFFB5BC5F2D87FC0E92CA0BF1" box="[421,510,999,1023]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="36" pageNumber="487" unit="mm" value="1.7">1.7 mm</quantity>
, BL/BW ~1.5. HCW
<quantity id="130AF9DFFFB5BC5F2ED7FC0F907A0BF1" box="[757,846,998,1023]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="36" pageNumber="487" unit="mm" value="0.6">0.6 mm</quantity>
, PW
<quantity id="130AF9DFFFB5BC5F2FAEFC0E90E90BF0" box="[908,989,999,1023]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="36" pageNumber="487" unit="mm" value="0.8">0.8mm</quantity>
, PL
<quantity id="130AF9DFFFB5BC5F2830FC0E975D0BF1" box="[1042,1129,999,1023]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="36" pageNumber="487" unit="mm" value="0.4">0.4 mm</quantity>
in midline. Rather convex. Pronotum anterolaterally rough through coarse granular punctures. The granular punctures in the centre of disc are very fine. Elytral punctation very fine. Uniformly yellow-ochre. Blond pilosity somewhat shaggy, almost erect.
</paragraph>
<paragraph id="D44D543AFFB5BC5F2CE5FBBB963A0CD8" blockId="36.[151,1437,998,2031]" pageId="36" pageNumber="487">Head very large, oral area narrow, labrum a small approximately square lobe with rounded margin. Mandible with wide cutting edge, no tooth. Terminal segment of maxillary palpus shorter than penultimate, slightly concave, distal half narrow, pointed. Labial palpus straight, with short, pointed terminal segment. Antennae slender, scape large, without sharp edge, pedicel much thinner, similar in length to antennomere 3. Legs unmodified.</paragraph>
<paragraph id="D44D543AFFB5BC5F2CE5FB0A93E40D85" blockId="36.[151,1437,998,2031]" pageId="36" pageNumber="487">Lower side. Head with well delimited small buttonhole. Prosternal process short, tear-shaped, mesoventral groove very short, transverse with angular, slightly projecting anterior margin. The rear margin is part of the regularly curved front edge of the mesoventrite, not angular as in most congeners. Mesoventral process fairly wide, caudally incised, separating the coxae completely and meeting the metaventral process. Metaventral discrimen not seen.</paragraph>
</subSubSection>
<subSubSection id="9CE807B1FFB5BC5F2CE5FA7F91E60FA9" pageId="36" pageNumber="487" type="description">
<paragraph id="D44D543AFFB5BC5F2CE5FA7F96740EA4" blockId="36.[151,1437,998,2031]" pageId="36" pageNumber="487">
<emphasis id="E6868828FFB5BC5F2CE5FA7F92300DA0" bold="true" box="[199,260,1430,1455]" pageId="36" pageNumber="487">Male</emphasis>
. T8 short, a tranverse arch supported by strong straight apodemes 3 times longer than the plate. S8 not developed. T9 unsclerotized, smaller than T8, apodemes shorter and thinner, a little curved. T8 and T9 are completely bare, no hairs or microtrichia. S9 is a narrow hairpin-like sclerite which first widens, then suddenly narrows and is longitudinally divided. Each distal part supports a large sharp barb and then tapers to a delicate tip (
<figureCitation id="4CC948BFFFB5BC5F2CBDF9CF93C80E30" box="[159,252,1574,1599]" captionStart="FIGURES 103 106" captionStartId="36.[151,264,905,928]" captionTargetBox="[160,1428,209,872]" captionTargetId="figure@36.[153,1434,193,885]" captionTargetPageId="36" captionText="FIGURES 103 106. Contacyphon forcipatus, n. sp., male. 103, segments 8 and 9, superimposed; 104, tegmen; 105, penis (103 105, holotype); 106, penis and tegmen of the paratype, superimposed. All to the same scale." httpUri="https://zenodo.org/record/235863/files/figure.png" pageId="36" pageNumber="487">Fig. 103</figureCitation>
).Tegmen and parameres form an elongate oval sclerite which is caudally narrowly open (
<figureCitation id="4CC948BFFFB5BC5F28D0F9CF96600E30" box="[1266,1364,1574,1599]" captionStart="FIGURES 103 106" captionStartId="36.[151,264,905,928]" captionTargetBox="[160,1428,209,872]" captionTargetId="figure@36.[153,1434,193,885]" captionTargetPageId="36" captionText="FIGURES 103 106. Contacyphon forcipatus, n. sp., male. 103, segments 8 and 9, superimposed; 104, tegmen; 105, penis (103 105, holotype); 106, penis and tegmen of the paratype, superimposed. All to the same scale." httpUri="https://zenodo.org/record/235863/files/figure.png" pageId="36" pageNumber="487">Fig. 104</figureCitation>
), each paramere with a sharp straight subapical spine appressed to the main sclerite. Penis (
<figureCitation id="4CC948BFFFB5BC5F286CF9A3979A0E6C" box="[1102,1198,1610,1635]" captionStart="FIGURES 103 106" captionStartId="36.[151,264,905,928]" captionTargetBox="[160,1428,209,872]" captionTargetId="figure@36.[153,1434,193,885]" captionTargetPageId="36" captionText="FIGURES 103 106. Contacyphon forcipatus, n. sp., male. 103, segments 8 and 9, superimposed; 104, tegmen; 105, penis (103 105, holotype); 106, penis and tegmen of the paratype, superimposed. All to the same scale." httpUri="https://zenodo.org/record/235863/files/figure.png" pageId="36" pageNumber="487">Fig. 105</figureCitation>
) with large oval pala full of muscles attaching to the common sinew of a pair of long sinuous prosthemes that cross over each other. They are in a different position in the
<typeStatus id="0B49EA98FFB5BC5F2E1DF97A91940EA4" box="[575,672,1683,1707]" pageId="36" pageNumber="487" type="paratype">paratype</typeStatus>
(
<figureCitation id="4CC948BFFFB5BC5F2E8CF97B903A0EA4" box="[686,782,1682,1707]" captionStart="FIGURES 103 106" captionStartId="36.[151,264,905,928]" captionTargetBox="[160,1428,209,872]" captionTargetId="figure@36.[153,1434,193,885]" captionTargetPageId="36" captionText="FIGURES 103 106. Contacyphon forcipatus, n. sp., male. 103, segments 8 and 9, superimposed; 104, tegmen; 105, penis (103 105, holotype); 106, penis and tegmen of the paratype, superimposed. All to the same scale." httpUri="https://zenodo.org/record/235863/files/figure.png" pageId="36" pageNumber="487">Fig. 106</figureCitation>
) and can apparently be moved like scissor blades.
</paragraph>
<paragraph id="D44D543AFFB5BC5F2CE5F95F92850EC0" blockId="36.[151,1437,998,2031]" box="[199,433,1718,1743]" pageId="36" pageNumber="487">
<emphasis id="E6868828FFB5BC5F2CE5F95F922A0EC0" bold="true" box="[199,286,1718,1743]" pageId="36" pageNumber="487">Female</emphasis>
. Not known.
</paragraph>
<paragraph id="D44D543AFFB5BC5F2CE5F93391E60FA9" blockId="36.[151,1437,998,2031]" pageId="36" pageNumber="487">
<emphasis id="E6868828FFB5BC5F2CE5F93393CB0EFC" bold="true" box="[199,255,1754,1779]" pageId="36" pageNumber="487">Note</emphasis>
. The new species belongs to genus
<taxonomicName id="13F22FB9FFB5BC5F2EAFF93390160EFC" box="[653,802,1754,1779]" class="Insecta" family="Scirtidae" genus="Contacyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="genus">
<emphasis id="E6868828FFB5BC5F2EAFF93390160EFC" box="[653,802,1754,1779]" italics="true" pageId="36" pageNumber="487">Contacyphon</emphasis>
</taxonomicName>
but close relatives are unknown. The lack of mandibular teeth and the rounded rear margin of the mesoventral groove are minor deviations from the common pattern and are possibly related to small body size. The fully developed but bare plates of T8 and T9, the modified narrow S9, and the shape of the penis are reminiscent of species in the
<emphasis id="E6868828FFB5BC5F2F34F8AE90410F51" box="[790,885,1863,1886]" italics="true" pageId="36" pageNumber="487">apoanus</emphasis>
-subgroup, e.g.
<taxonomicName id="13F22FB9FFB5BC5F280AF8AE93EC0F8D" authority="Klausnitzer, 2012" authorityName="Klausnitzer" authorityYear="2012" class="Insecta" family="Scirtidae" genus="Contacyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="rotundosternalis">
<emphasis id="E6868828FFB5BC5F280AF8AE96370F50" box="[1064,1283,1862,1887]" italics="true" pageId="36" pageNumber="487">C. rotundosternalis</emphasis>
(
<bibRefCitation id="B06329CBFFB5BC5F2931F8AF93E50F8D" author="Klausnitzer" pageId="36" pageNumber="487" refString="Klausnitzer, B. (2012) Zur Kenntnis vo n Hydrocypho n L. Redtenbacher, 1858 un d Cypho n Paykull, 1799 aus Nepal und der Orientalischen Region (Insecta: Coleoptera: Scirtidae). (169. Beitrag zur Kenntnis der Scirtidae). In: Hartmann, M. &amp; Weipert, J. (Eds.), Biodiversitat und Naturausstattung im Himalaya IV. Verein der Freunde &amp; Forderer des Naturkundemuseums Erfurt e. V., Erfurt, pp. 329 - 347." type="book chapter" year="2012">Klausnitzer, 2012</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="ACE514AAFFB5BC5F2D3DF88392560F8C" box="[287,354,1898,1923]" name="Nepal" pageId="36" pageNumber="487">Nepal</collectingCountry>
and
<taxonomicName id="13F22FB9FFB5BC5F2DB8F882903E0F8D" authority="Yoshitomi, 2009" authorityName="Yoshitomi" authorityYear="2009" box="[410,778,1898,1923]" class="Insecta" family="Scirtidae" genus="Contacyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="sagadanus">
<emphasis id="E6868828FFB5BC5F2DB8F88291000F8C" box="[410,564,1898,1923]" italics="true" pageId="36" pageNumber="487">C. sagadanus</emphasis>
(
<bibRefCitation id="B06329CBFFB5BC5F2E66F88390360F8D" author="Yoshitomi" box="[580,770,1898,1923]" pageId="36" pageNumber="487" refString="Yoshitomi, H. (2009) Scirtid beetles (Coleoptera, Scirtidae) of the Oriental region Part 10. New species and new record of Cyphon variabilis species-group. Elytra, Tokyo, 37 (1), 87 - 97." type="journal article" year="2009">Yoshitomi, 2009</bibRefCitation>
)
</taxonomicName>
from Luzon. However, their tegmen is a pilose plate very different from the unusual tegmen of
<taxonomicName id="13F22FB9FFB5BC5F2E18F86691FF0FA9" box="[570,715,1934,1958]" class="Insecta" family="Scirtidae" genus="Contacyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="forcipatus">
<emphasis id="E6868828FFB5BC5F2E18F86691FF0FA9" box="[570,715,1934,1958]" italics="true" pageId="36" pageNumber="487">C. forcipatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="9CE807B1FFB5BC5F2CE5F85B93C50FE0" pageId="36" pageNumber="487" type="etymology">
<paragraph id="D44D543AFFB5BC5F2CE5F85B93C50FE0" blockId="36.[151,1437,998,2031]" pageId="36" pageNumber="487">
<emphasis id="E6868828FFB5BC5F2CE5F85B92720FC4" bold="true" box="[199,326,1970,1995]" pageId="36" pageNumber="487">Etymology</emphasis>
. The name is an adjective alluding to the shape of the genitalia: Latin
<taxonomicName id="13F22FB9FFB5BC5F2848F85B97EE0FC5" box="[1130,1242,1970,1994]" class="Insecta" family="Scirtidae" genus="Contacyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="487" phylum="Arthropoda" rank="species" species="forcipatus">
<emphasis id="E6868828FFB5BC5F2848F85B97EE0FC5" box="[1130,1242,1970,1994]" italics="true" pageId="36" pageNumber="487">forcipatus</emphasis>
</taxonomicName>
, provided with a forceps.
</paragraph>
</subSubSection>
</treatment>
</document>