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<document id="DED3129618EF845EECE02B845AE5DB03" ID-DOI="10.11646/zootaxa.3981.4.1" ID-GBIF-Dataset="15c3bf52-4481-4a75-8643-7c919bdde6c4" ID-ISSN="1175-5326" ID-Zenodo-Dep="235846" ID-ZooBank="EF71D83B-17B4-49CA-826E-D3A8E7979750" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461248924452" checkinUser="plazi" docAuthor="Zwick, Peter" docDate="2015" docId="5C5BE52CFF88BC622CB5FBD791A60FBA" docLanguage="en" docName="zt03981p490.pdf" docOrigin="Zootaxa 3981 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptocyphon Zwick, 2015, n. gen." docType="treatment" docVersion="8" lastPageNumber="476" masterDocId="A0629D54FF91BC7B2C22FFE99334080F" masterDocTitle="Australian Marsh Beetles (Coleoptera: Scirtidae). 8. The new genera Cygnocyphon, Eximiocyphon, Paracyphon, Leptocyphon, Tectocyphon, and additions to Contacyphon de Gozis, Nanocyphon Zwick and Eurycyphon Watts" masterLastPageNumber="490" masterPageNumber="451" pageNumber="476" updateTime="1698607092210" updateUser="plazi">
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<mods:title id="EAAA6FDAF4E2D392731EAECD15DCE906">Australian Marsh Beetles (Coleoptera: Scirtidae). 8. The new genera Cygnocyphon, Eximiocyphon, Paracyphon, Leptocyphon, Tectocyphon, and additions to Contacyphon de Gozis, Nanocyphon Zwick and Eurycyphon Watts</mods:title>
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<mods:date id="5182E5C8D9436A8D936CE42D4F7653F6">2015</mods:date>
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<treatment id="5C5BE52CFF88BC622CB5FBD791A60FBA" ID-DOI="http://doi.org/10.5281/zenodo.6110574" ID-GBIF-Taxon="119635593" ID-Zenodo-Dep="6110574" LSID="urn:lsid:plazi:treatment:5C5BE52CFF88BC622CB5FBD791A60FBA" httpUri="http://treatment.plazi.org/id/5C5BE52CFF88BC622CB5FBD791A60FBA" lastPageNumber="476" pageId="25" pageNumber="476">
<subSubSection id="9CE807B1FF88BC622CB5FBD792A70C57" box="[151,403,1086,1112]" pageId="25" pageNumber="476" type="nomenclature">
<paragraph id="D44D543AFF88BC622CB5FBD792A70C57" blockId="25.[151,403,1086,1112]" box="[151,403,1086,1112]" pageId="25" pageNumber="476">
<heading id="8F05E356FF88BC622CB5FBD792A70C57" bold="true" box="[151,403,1086,1112]" fontSize="11" level="1" pageId="25" pageNumber="476" reason="1">
<emphasis id="E6868828FF88BC622CB5FBD792A70C57" bold="true" box="[151,403,1086,1112]" pageId="25" pageNumber="476">
<emphasis id="E6868828FF88BC622CB5FBD7920E0C57" bold="true" box="[151,314,1086,1112]" italics="true" pageId="25" pageNumber="476">
<taxonomicName id="13F22FB9FF88BC622CB5FBD792000C57" box="[151,308,1086,1112]" class="Insecta" family="Scirtidae" genus="Leptocyphon" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="genus" status="gen. nov.">Leptocyphon</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="FDB53553FF88BC622D60FBD792A70C57" box="[322,403,1086,1112]" pageId="25" pageNumber="476" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9CE807B1FF88BC622CB5FB6C91AC0C91" box="[151,664,1157,1182]" pageId="25" pageNumber="476" type="materials_examined">
<paragraph id="D44D543AFF88BC622CB5FB6C91AC0C91" blockId="25.[151,1437,1157,1974]" box="[151,664,1157,1182]" pageId="25" pageNumber="476">
<emphasis id="E6868828FF88BC622CB5FB6C921F0C91" bold="true" box="[151,299,1157,1182]" pageId="25" pageNumber="476">
<typeStatus id="0B49EA98FF88BC622CB5FB6C93E60C91" box="[151,210,1157,1182]" pageId="25" pageNumber="476">Type</typeStatus>
species
</emphasis>
:
<emphasis id="E6868828FF88BC622D18FB6C91660C91" box="[314,594,1157,1182]" italics="true" pageId="25" pageNumber="476">
<taxonomicName id="13F22FB9FF88BC622D18FB6C917A0C91" box="[314,590,1157,1182]" class="Insecta" family="Scirtidae" genus="Leptocyphon" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="furcalonga" status="sp. nov.">Leptocyphon furcalonga</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="FDB53553FF88BC622E7BFB6C91AC0C91" box="[601,664,1157,1182]" pageId="25" pageNumber="476" rank="species">
<emphasis id="E6868828FF88BC622E7BFB6C91AC0C91" bold="true" box="[601,664,1157,1182]" pageId="25" pageNumber="476">n. sp.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="9CE807B1FF88BC622CE5FB4097220F61" pageId="25" pageNumber="476" type="diagnosis">
<paragraph id="D44D543AFF88BC622CE5FB4090860E25" blockId="25.[151,1437,1157,1974]" pageId="25" pageNumber="476">
<emphasis id="E6868828FF88BC622CE5FB40920E0CCD" bold="true" box="[199,314,1193,1218]" pageId="25" pageNumber="476">Diagnosis</emphasis>
. Small, BL
<emphasis id="E6868828FF88BC622DE1FB4392EA0CCE" box="[451,478,1194,1217]" italics="true" pageId="25" pageNumber="476">ca</emphasis>
<quantity id="130AF9DFFF88BC622DC6FB4391150CCD" box="[484,545,1194,1218]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="25" pageNumber="476" unit="mm" value="3.0">3mm</quantity>
. Slender, flat (
<figureCitation id="4CC948BFFF88BC622EE1FB4090790CCD" box="[707,845,1193,1218]" captionStart="FIGURES 76 79" captionStartId="26.[151,264,1961,1984]" captionTargetBox="[150,1436,1070,1933]" captionTargetId="figure@26.[150,1436,1070,1940]" captionTargetPageId="26" captionText="FIGURES 76 79. Leptocyphon furcalonga, n. gen., n. sp.: 76, 77, habitus dorsal and lateral; 78, forebody, oblique frontodorsal view; 79, cleared head, ventral. Not to scale." httpUri="https://zenodo.org/record/235858/files/figure.png" pageId="25" pageNumber="476">Figs. 7678</figureCitation>
). Pronotum transverse, short, little narrower than the elytra. Base sinuous, projecting towards scutellum, laterally straight, rear angles almost rectangular, blunt. Sides converging towards the rounded front corners between which the front margin is straight. Sides of elytra little curved, epipleura narrow. Head a little wider than long, no antennal groove. Eyes moderately projecting. Subgenal ridge joining a branch from the gular suture, together they meet the edge of the oral cavity (
<figureCitation id="4CC948BFFF88BC622898FAD096250D5D" box="[1210,1297,1337,1362]" captionStart="FIGURES 76 79" captionStartId="26.[151,264,1961,1984]" captionTargetBox="[150,1436,1070,1933]" captionTargetId="figure@26.[150,1436,1070,1940]" captionTargetPageId="26" captionText="FIGURES 76 79. Leptocyphon furcalonga, n. gen., n. sp.: 76, 77, habitus dorsal and lateral; 78, forebody, oblique frontodorsal view; 79, cleared head, ventral. Not to scale." httpUri="https://zenodo.org/record/235858/files/figure.png" pageId="25" pageNumber="476">Fig. 79</figureCitation>
). Only right mandible with small tooth, or both toothless, no spinules in molar area. Palpi straight, antennae and legs unmodified. Prosternal epipleura reaching to level of front coxa. Prosternum very small (
<figureCitation id="4CC948BFFF88BC622898FA6896260D95" box="[1210,1298,1409,1434]" captionStart="FIGURES 80 88" captionStartId="27.[151,264,1910,1933]" captionTargetBox="[194,1393,193,1857]" captionTargetId="figure@27.[182,1393,193,1859]" captionTargetPageId="27" captionText="FIGURES 80 88. Leptocyphon furcalonga, n. gen., n. sp.: 80, 81, ventral views of prothorax and mesoventrite; male: 82, T 8; 83, S 8; 84, T 9 and S 9, superimposed; 85, penis; 86, tegmen and parameres, with enlarged detail of paramere structure. Female: 87, cleared terminalia, with contours of sternites 6 and 7; apodemes of S 8 shown in black; 88, prehensor. 80 84 and 85, 86 and 88 to the same scales, respectively. aT 8, apodemes of T 8; ba, bacula of ovipositor; lc, laterocervicalia." httpUri="https://zenodo.org/record/235859/files/figure.png" pageId="25" pageNumber="476">Fig. 80</figureCitation>
), prosternal process reduced, not reaching back beyond the coxae, extremely narrow. Receiving mesoventral groove indistinct, mesoventral process very narrow but complete (
<figureCitation id="4CC948BFFF88BC622EE5FA20902A0DED" box="[711,798,1481,1506]" captionStart="FIGURES 80 88" captionStartId="27.[151,264,1910,1933]" captionTargetBox="[194,1393,193,1857]" captionTargetId="figure@27.[182,1393,193,1859]" captionTargetPageId="27" captionText="FIGURES 80 88. Leptocyphon furcalonga, n. gen., n. sp.: 80, 81, ventral views of prothorax and mesoventrite; male: 82, T 8; 83, S 8; 84, T 9 and S 9, superimposed; 85, penis; 86, tegmen and parameres, with enlarged detail of paramere structure. Female: 87, cleared terminalia, with contours of sternites 6 and 7; apodemes of S 8 shown in black; 88, prehensor. 80 84 and 85, 86 and 88 to the same scales, respectively. aT 8, apodemes of T 8; ba, bacula of ovipositor; lc, laterocervicalia." httpUri="https://zenodo.org/record/235859/files/figure.png" pageId="25" pageNumber="476">Fig. 81</figureCitation>
). Male S8 large, Y-shaped, with long unpaired sclerite (
<figureCitation id="4CC948BFFF88BC622CBDFA0492340E09" box="[159,256,1517,1542]" captionStart="FIGURES 80 88" captionStartId="27.[151,264,1910,1933]" captionTargetBox="[194,1393,193,1857]" captionTargetId="figure@27.[182,1393,193,1859]" captionTargetPageId="27" captionText="FIGURES 80 88. Leptocyphon furcalonga, n. gen., n. sp.: 80, 81, ventral views of prothorax and mesoventrite; male: 82, T 8; 83, S 8; 84, T 9 and S 9, superimposed; 85, penis; 86, tegmen and parameres, with enlarged detail of paramere structure. Female: 87, cleared terminalia, with contours of sternites 6 and 7; apodemes of S 8 shown in black; 88, prehensor. 80 84 and 85, 86 and 88 to the same scales, respectively. aT 8, apodemes of T 8; ba, bacula of ovipositor; lc, laterocervicalia." httpUri="https://zenodo.org/record/235859/files/figure.png" pageId="25" pageNumber="476">Figs. 83</figureCitation>
,
<figureCitation id="4CC948BFFF88BC622D32FA0792040E09" box="[272,304,1518,1542]" captionStart="FIGURES 89 93" captionStartId="29.[151,264,1962,1985]" captionTargetBox="[174,1428,650,1918]" captionTargetId="figure@29.[153,1434,623,1940]" captionTargetPageId="29" captionText="FIGURES 89 93. Leptocyphon quadricornutus, n. sp., male: 89, T 8, with enlarged detail; 90, S 8; 91, S 9; 92, tegmen and parameres; 93, penis. 89, 90 and 91 93 to the same scales, respectively." httpUri="https://zenodo.org/record/235860/files/figure.png" pageId="25" pageNumber="476">90</figureCitation>
). No stylus. Female (known only of
<taxonomicName id="13F22FB9FF88BC622EC7FA0790B50E09" box="[741,897,1517,1542]" class="Insecta" family="Scirtidae" genus="Leptocyphon" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="furcalonga">
<emphasis id="E6868828FF88BC622EC7FA0790B50E09" box="[741,897,1517,1542]" italics="true" pageId="25" pageNumber="476">L. furcalonga</emphasis>
</taxonomicName>
): bacula firmly connected to gonocoxites, not articulated, gonostyle wart-like (
<figureCitation id="4CC948BFFF88BC622E21F9F891630E25" box="[515,599,1553,1578]" captionStart="FIGURES 80 88" captionStartId="27.[151,264,1910,1933]" captionTargetBox="[194,1393,193,1857]" captionTargetId="figure@27.[182,1393,193,1859]" captionTargetPageId="27" captionText="FIGURES 80 88. Leptocyphon furcalonga, n. gen., n. sp.: 80, 81, ventral views of prothorax and mesoventrite; male: 82, T 8; 83, S 8; 84, T 9 and S 9, superimposed; 85, penis; 86, tegmen and parameres, with enlarged detail of paramere structure. Female: 87, cleared terminalia, with contours of sternites 6 and 7; apodemes of S 8 shown in black; 88, prehensor. 80 84 and 85, 86 and 88 to the same scales, respectively. aT 8, apodemes of T 8; ba, bacula of ovipositor; lc, laterocervicalia." httpUri="https://zenodo.org/record/235859/files/figure.png" pageId="25" pageNumber="476">Fig. 87</figureCitation>
); prehensor complex (
<figureCitation id="4CC948BFFF88BC622F73F9F890900E25" box="[849,932,1553,1578]" captionStart="FIGURES 80 88" captionStartId="27.[151,264,1910,1933]" captionTargetBox="[194,1393,193,1857]" captionTargetId="figure@27.[182,1393,193,1859]" captionTargetPageId="27" captionText="FIGURES 80 88. Leptocyphon furcalonga, n. gen., n. sp.: 80, 81, ventral views of prothorax and mesoventrite; male: 82, T 8; 83, S 8; 84, T 9 and S 9, superimposed; 85, penis; 86, tegmen and parameres, with enlarged detail of paramere structure. Female: 87, cleared terminalia, with contours of sternites 6 and 7; apodemes of S 8 shown in black; 88, prehensor. 80 84 and 85, 86 and 88 to the same scales, respectively. aT 8, apodemes of T 8; ba, bacula of ovipositor; lc, laterocervicalia." httpUri="https://zenodo.org/record/235859/files/figure.png" pageId="25" pageNumber="476">Fig. 88</figureCitation>
).
</paragraph>
<paragraph id="D44D543AFF88BC622CE5F9DC92780EB5" blockId="25.[151,1437,1157,1974]" pageId="25" pageNumber="476">
<emphasis id="E6868828FF88BC622CE5F9DC923E0E41" bold="true" box="[199,266,1589,1614]" pageId="25" pageNumber="476">Notes</emphasis>
. The two included species are very similar in habitus and body structure but differ strikingly in the shape of the male genitalia. I include them with hesitation in the same genus. When the female of the second species becomes known a decision will be easier. Both species occur in the extreme SW of Western
<collectingCountry id="ACE514AAFF88BC6228E6F994961F0E99" box="[1220,1323,1661,1686]" name="Australia" pageId="25" pageNumber="476">Australia</collectingCountry>
and were rarely collected.
</paragraph>
<paragraph id="D44D543AFF88BC622CE5F92C97220F61" blockId="25.[151,1437,1157,1974]" pageId="25" pageNumber="476">
Describing and naming a small slender Australian marsh beetle with blunt pronotal front angles is difficult. The available generic key (
<bibRefCitation id="B06329CBFF88BC622DE5F903917C0F0D" author="Watts" box="[455,584,1770,1794]" pageId="25" pageNumber="476" refString="Watts, C. H. S. (2011) Revision of Australian Scirtidae of the genera Chameloscyphon gen. nov., Daploeuros gen. nov., Dasyscyphon gen. nov., Eurycyphon gen. nov., Macrodascillus Carter, Petrocyphon gen. nov. and Spaniosdascillus gen. nov. (Coleoptera). Transactions of the Royal Society of South Australia, 135 (1), 66 - 110." type="journal article" year="2011">Watts 2011</bibRefCitation>
) leads to
<taxonomicName id="13F22FB9FF88BC622E97F900904F0F0D" box="[693,891,1769,1794]" class="Insecta" family="Scirtidae" genus="Pseudomicrocara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="genus">
<emphasis id="E6868828FF88BC622E97F900904F0F0D" box="[693,891,1769,1794]" italics="true" pageId="25" pageNumber="476">Pseudomicrocara</emphasis>
</taxonomicName>
or
<taxonomicName id="13F22FB9FF88BC622F83F90090CC0F0D" box="[929,1016,1769,1794]" class="Insecta" family="Scirtidae" genus="Cyphon" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="genus">
<emphasis id="E6868828FF88BC622F83F90090CC0F0D" box="[929,1016,1769,1794]" italics="true" pageId="25" pageNumber="476">Cyphon</emphasis>
</taxonomicName>
(in the former wide sense) which are separated by size (overlapping) and small spines in the molar area, which varies in
<taxonomicName id="13F22FB9FF88BC622873F8E496230F29" box="[1105,1303,1805,1830]" class="Insecta" family="Scirtidae" genus="Pseudomicrocara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="genus">
<emphasis id="E6868828FF88BC622873F8E496230F29" box="[1105,1303,1805,1830]" italics="true" pageId="25" pageNumber="476">Pseudomicrocara</emphasis>
</taxonomicName>
. Therefore, new taxa resembling
<taxonomicName id="13F22FB9FF88BC622DACF8D891600F45" box="[398,596,1841,1866]" class="Insecta" family="Scirtidae" genus="Pseudomicrocara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="genus">
<emphasis id="E6868828FF88BC622DACF8D891600F45" box="[398,596,1841,1866]" italics="true" pageId="25" pageNumber="476">Pseudomicrocara</emphasis>
</taxonomicName>
(s. l.) need to be compared to the
<typeStatus id="0B49EA98FF88BC622FCEF8DB97280F45" box="[1004,1052,1842,1866]" pageId="25" pageNumber="476">type</typeStatus>
species,
<taxonomicName id="13F22FB9FF88BC6228AAF8DB93E60F61" authority="Armstrong, 1953" authorityName="Armstrong" authorityYear="1953" class="Insecta" family="Scirtidae" genus="Pseudomicrocara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="E6868828FF88BC6228AAF8DB96250F45" box="[1160,1297,1841,1866]" italics="true" pageId="25" pageNumber="476">P. orientalis</emphasis>
<bibRefCitation id="B06329CBFF88BC62293EF8DB93E60F61" author="Armstrong" pageId="25" pageNumber="476" refString="Armstrong, J. W. T. (1953) On Australian Helodidae (Coleoptera). 1. Descriptions of new genera and species. 1. Proceedings of the Linnean Society of New South Wales, 78, 19 - 32." type="journal article" year="1953">Armstrong, 1953</bibRefCitation>
</taxonomicName>
, see below. What the closest relatives of
<taxonomicName id="13F22FB9FF88BC622EBEF8BC90190F61" box="[668,813,1877,1902]" class="Insecta" family="Scirtidae" genus="Leptocyphon" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E6868828FF88BC622EBEF8BC90190F61" box="[668,813,1877,1902]" italics="true" pageId="25" pageNumber="476">Leptocyphon</emphasis>
</taxonomicName>
may be is uncertain.
</paragraph>
</subSubSection>
<subSubSection id="9CE807B1FF88BC622CE5F89091A60FBA" pageId="25" pageNumber="476" type="etymology">
<paragraph id="D44D543AFF88BC622CE5F89091A60FBA" blockId="25.[151,1437,1157,1974]" pageId="25" pageNumber="476">
<emphasis id="E6868828FF88BC622CE5F89092720F9D" bold="true" box="[199,326,1913,1938]" pageId="25" pageNumber="476">Etymology</emphasis>
. The generic name refers to the thin prosternal and mesoventral processes and connects Gree
<emphasis id="E6868828FF88BC6229ADF89093F50FB9" italics="true" pageId="25" pageNumber="476">k lept</emphasis>
os, thin, with the classical name,
<emphasis id="E6868828FF88BC622E16F874915B0FB9" box="[564,623,1949,1974]" italics="true" pageId="25" pageNumber="476">Cyph</emphasis>
on.
</paragraph>
</subSubSection>
</treatment>
</document>