257 lines
35 KiB
XML
257 lines
35 KiB
XML
<document ID-CLB-Dataset="33297" ID-DOI="http://dx.doi.org/10.3897/zookeys.665.11609" ID-GBIF-Dataset="fb32460f-a523-4df5-9866-f88848316b07" ID-PMC="PMC5523170" ID-Pensoft-Pub="1313-2970-665-121" ID-Pensoft-UUID="FFF18360FFDEFFE0143AFFD6C055A851" ID-PubMed="28769630" ID-Zenodo-Dep="579504" ID-ZooBank="222DBEEE143E4E0BBA4FC3484A230961" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-665-121" ModsDocOrigin="ZooKeys 665" ModsDocTitle="A new species group in the genus Dichaetophora, with descriptions of six new species from the Oriental region (Diptera, Drosophilidae)" checkinTime="1491378299097" checkinUser="pensoft" docAuthor="Yang, Jin-Hua, Toda, Masanori J., Suwito, Awit, Hashim, Rosli & Gao, Jian-Jun" docDate="2017" docId="3ED161995F59D6D07250C6B39BA436C5" docLanguage="en" docName="ZooKeys 665: 121-146" docOrigin="ZooKeys 665" docPubDate="2017-04-04" docSource="http://dx.doi.org/10.3897/zookeys.665.11609" docTitle="Dichaetophora trilobita Yang & Gao 2017, sp. n." docType="treatment" docUuid="BAF11C36-EB50-4E98-A8A1-537AE7AD715A" docUuidSource="ZooBank" docVersion="9" id="FFF18360FFDEFFE0143AFFD6C055A851" lastPageNumber="124" masterDocId="FFF18360FFDEFFE0143AFFD6C055A851" masterDocTitle="A new species group in the genus Dichaetophora, with descriptions of six new species from the Oriental region (Diptera, Drosophilidae)" masterLastPageNumber="146" masterPageNumber="121" pageNumber="124" updateTime="1705949547935" updateUser="pensoft">
|
|
<mods:mods id="41058DC75C931F4DF3A6660FC1CC48C8" xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo id="7E8FE06183AE6D7F041DA0E038FAE578">
|
|
<mods:title id="EEC493734450BA3F8C319A6E2868134A">A new species group in the genus Dichaetophora, with descriptions of six new species from the Oriental region (Diptera, Drosophilidae)</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name id="D3403D85C9BEC9553871F1A0250D1C26" type="personal">
|
|
<mods:role id="8E66101E61B102BEDF922E87DBAC3E0B">
|
|
<mods:roleTerm id="0C0011E33DBE5F3D1DE3B9CF5E388DDF">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="9A32F302C3E8941C33AAF5666148B2BA">Yang, Jin-Hua</mods:namePart>
|
|
<mods:affiliation id="DEE1EB91B5BBEEE54182F6B4851FFBE9">State Key Laboratory for Conservation and Utilization of Bioresources in Yunnan, Yunnan University, 2 Cuihubeilu, Kunming 650091, China</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name id="45CC9ECA3F8CEF06723C61C8F4098560" type="personal">
|
|
<mods:role id="BC00B158D3D2B09C5DF28C76560A375D">
|
|
<mods:roleTerm id="CD609C1090A6A8DDF6345F40582E6F66">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="417047CBCFD709DC97936178CBEA09F5">Toda, Masanori J.</mods:namePart>
|
|
<mods:affiliation id="24912A4B6C3460A270EBFD10B162556B">Hokkaido University Museum, Hokkaido University, N 10, W 8, Kita-ku, Sapporo 060 - 0810, Japan</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name id="C36BEF5038E3A5D84D294DEB5587302C" type="personal">
|
|
<mods:role id="7E2B344121FE0673F076BFB50B5B34F2">
|
|
<mods:roleTerm id="81C980286FBC1F2BA6085493EDD161BA">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="65986CD7AFDD42A44FF3AE27861A16E3">Suwito, Awit</mods:namePart>
|
|
<mods:affiliation id="63B4E962DB31A79646C8D9C32852A559">Zoology Division (Museum Zoologicum Bogoriense), Research Center for Biology-LIPI, Cibinong, Bogor 16911, Indonesia</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name id="631F9967675E09629ADD6FCB56D5E3E1" type="personal">
|
|
<mods:role id="52A797C017495ACED57BCD8D82C76CEA">
|
|
<mods:roleTerm id="63DD4E8486C7A7F02C6410230E7C3772">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="8DB1A99BA06529653663FE218CDC36E2">Hashim, Rosli</mods:namePart>
|
|
<mods:affiliation id="5E8CA42D668BD87893CB3D20D6EB8EF8">Institute of Biological Science, University of Malaya, 50603 Kuala Lumpur, Malaysia</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name id="8DF414B1BB3988C3AA340764B70DB214" type="personal">
|
|
<mods:role id="EE09A6791C17A96F86C3153997BA7189">
|
|
<mods:roleTerm id="557422E89AB2553236D394BCD9A9A0CC">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="DA9B87BBB307458B6AD1DA2F33B46D90">Gao, Jian-Jun</mods:namePart>
|
|
<mods:affiliation id="411EEB0E04BF9303039AEA0EC4F9F83C">State Key Laboratory for Conservation and Utilization of Bioresources in Yunnan, Yunnan University, 2 Cuihubeilu, Kunming 650091, China</mods:affiliation>
|
|
<mods:nameIdentifier id="CB04D90F5DC1E9513EB51D824FF10943" type="email">gao-leyun@263.net</mods:nameIdentifier>
|
|
</mods:name>
|
|
<mods:typeOfResource id="B71AAA7EF6F8FBC29227E8ED6B8DF941">text</mods:typeOfResource>
|
|
<mods:relatedItem id="96574E673E719333ECD63BA1559202F5" type="host">
|
|
<mods:titleInfo id="4A6324C80C308480FCA4070677B03F77">
|
|
<mods:title id="EF87D301A9683B434FB43AC79F297E35">ZooKeys</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part id="2C9B6E43A3E64A30A29AEAB65CF503A7">
|
|
<mods:date id="63C9F35C01E59F7E7154629102B6FAC0">2017</mods:date>
|
|
<mods:detail id="072BC6980C9E911CED49A2D836F20E83" type="pubDate">
|
|
<mods:number id="E4ACE195923CF1A7CEC12149295A324E">2017-04-04</mods:number>
|
|
</mods:detail>
|
|
<mods:detail id="94A1683C321AC466147FAECC062018BB" type="volume">
|
|
<mods:number id="570399440063F3B3A88013DB659F3A2D">665</mods:number>
|
|
</mods:detail>
|
|
<mods:extent id="935571E7907DA2A40CF84BA71BD49563" unit="page">
|
|
<mods:start id="DF83E7C33FE24FD3A916458EF96C85B1">121</mods:start>
|
|
<mods:end id="EA534EF46D470492093ABBA3C66923AD">146</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location id="E6355DF1222C39056A85FC61FC9DA679">
|
|
<mods:url id="198843BD9745EACB36037CBA8E8524F8">http://dx.doi.org/10.3897/zookeys.665.11609</mods:url>
|
|
</mods:location>
|
|
<mods:classification id="09A17063E653A146A3D8A728553AE78B">journal article</mods:classification>
|
|
<mods:identifier id="4E2F02E62E53BFA79B2AA37896853435" type="DOI">http://dx.doi.org/10.3897/zookeys.665.11609</mods:identifier>
|
|
<mods:identifier id="7AC04B0B6927924E07AEC2587255203B" type="Pensoft-Pub">1313-2970-665-121</mods:identifier>
|
|
<mods:identifier id="B48970A33413FA204A15CF04ED117B9B" type="ZooBank">222DBEEE143E4E0BBA4FC3484A230961</mods:identifier>
|
|
<mods:identifier id="7AB51D54D6BB75E51CC9C883BA915BF2" type="Pensoft-UUID">FFF18360FFDEFFE0143AFFD6C055A851</mods:identifier>
|
|
<mods:identifier id="DFD6F8E9E67C8ED342A36B57E3C825B3" type="Zenodo-Dep">579504</mods:identifier>
|
|
</mods:mods>
|
|
<treatment id="3ED161995F59D6D07250C6B39BA436C5" ID-GBIF-Taxon="128229125" LSID="urn:lsid:zoobank.org:act:BAF11C36-EB50-4E98-A8A1-537AE7AD715A" httpUri="http://treatment.plazi.org/id/3ED161995F59D6D07250C6B39BA436C5" lastPageNumber="124" pageId="3" pageNumber="124">
|
|
<subSubSection id="E51A83186336FE85F80F518197D3D63B" pageId="3" pageNumber="124" type="nomenclature">
|
|
<paragraph id="72F1412799484C092FC559381B888326" pageId="3" pageNumber="124">
|
|
<taxonomicName id="30347C762EB8A5F3DE5CFC998AD52528" ID-CoL="TRL" LSID="http://zoobank.org/BAF11C36-EB50-4E98-A8A1-537AE7AD715A" authority="Yang & Gao" authorityName="Yang & Gao" authorityYear="2017" class="Insecta" family="Drosophilidae" genus="Dichaetophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichaetophora trilobita" order="Diptera" pageId="3" pageNumber="124" phylum="Arthropoda" rank="species" species="trilobita" status="sp. n.">Dichaetophora trilobita Yang & Gao</taxonomicName>
|
|
<taxonomicNameLabel id="E3C299BBC9DD9A33BDDC4B703AF23789" pageId="3" pageNumber="124">sp. n.</taxonomicNameLabel>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="5352C8B2AA84E2C1A3ECC2ADC016E6CD" pageId="3" pageNumber="124" type="description">
|
|
<paragraph id="8906B2D3C5C3E31176AEF6B97A17D172" pageId="3" pageNumber="124">
|
|
<figureCitation id="35D75E4F3F724D6EFAE042CCA5C0F917" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Left lateral habitus, head and thorax (dorsal view), wing (left, ventral view), and abdomen (dorsal view). A-D Dichaetophora tirlobita sp. n. (# 03877) E-G D. heterochroma sp. n. (# 03879) H-J D. flatosternata sp. n. (# 04172) K-M D. borneoensis sp. n. (# 03895) N-P D. javaensis sp. n. (# 03892) Q-S D. sumatraensis sp. n. (# 03890). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.665.11609.figure3" httpUri="https://binary.pensoft.net/fig/130047" pageId="3" pageNumber="124">Figs 3A-D</figureCitation>
|
|
<figureCitation id="0C276D20C0D31897ECBB6562A820835F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Head (anterior view), postocciput, palpus, and prementum (ventral and lateral view, respectively). A-E Dichaetophora tirlobita sp. n. (# 03877) F-J D. heterochroma sp. n. (# 03879) K-O D. flatosternata sp. n. (# 04172) P-T D. borneoensis sp. n. (# 03895) U-Y D. javaensis sp. n. (# 03892) Z-D D. sumatraensis sp. n. (# 03890). Scale bars: 0.1 mm except for A, F, K, P, U, Z (0.5 mm)." figureDoi="10.3897/zookeys.665.11609.figure4" httpUri="https://binary.pensoft.net/fig/130048" pageId="3" pageNumber="124">, 4A-E</figureCitation>
|
|
<figureCitation id="404CE0F0F0136074D41E4356A9826DE2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dichaetophora trilobita sp. n. (A-H # 03877 I-K paratype # 03878). A Periphallic organs (posterior view), with red arrow indicating the median process on the caudoventral bridge of cerci B periphallic organs (posterolateral view), with red arrows indicating the prominent setae on the cercus and the anteroventral fusion of cercus (sclerotized, marginal plate) with the epandrium C surstyli (ventral view) D surstylus (inner side) E-G phallic organs (ventral, ventrolateral and lateral view, respectively) H paramedian setae (indicated with red arrows), and apical portion of paramere I, J oviscapt (lateral and ventral view, respectively) K spermathecae. Abbreviations: aed = aedeagus, aed a = aedeagal apodeme, cerc = cercus, epand = epandrium, hypd = hypandrium, pm = paramere, sur = surstylus, 10 S = tenth sternite. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.665.11609.figure5" httpUri="https://binary.pensoft.net/fig/130049" pageId="3" pageNumber="124">, 5</figureCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="96E57763CA2FB6D02CB6D94DE34933B8" pageId="3" pageNumber="124" type="type material">
|
|
<paragraph id="B6B9956C8749DE781F8393B2ABD99A4F" pageId="3" pageNumber="124">Type material.</paragraph>
|
|
<paragraph id="F16AFDB8AA04B9A95620FAE1CBA6077C" pageId="3" pageNumber="124">Holotype ♂ (#03877): MALAYSIA: Ulu Gombak, Selangor, 8.xii.2013, MJ Toda (UMKL).</paragraph>
|
|
<paragraph id="B9B41040FAF9FB9B9976DC34D3F32DFC" pageId="3" pageNumber="124">Paratypes: same data as holotype (1♀: #03878, UMKL); Kubah, Sarawak, Malaysia, 19.i.1999 (1♂, 1♀, SEHU); Park Headquarters, Mt. Kinabalu, Sabah, Malaysia, 11.iii.2008, MJ Toda (1♂: #03876, KIZ); Poring, Mt. Kinabalu, Sabah, Malaysia, 20.iii.2008, MJ Toda (1♀: #03882, KIZ).</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C7AC3FCCDB64DB37F5086A626C0992C5" pageId="3" pageNumber="124" type="diagnosis">
|
|
<paragraph id="A152A609FE08F5B2ACC235ABC97EC2D5" pageId="3" pageNumber="124">Diagnosis.</paragraph>
|
|
<paragraph id="A3E11009691E4BC7880EC4E80E05D41C" pageId="3" pageNumber="124">
|
|
Postocellar setae absent; wing without distinct cloud (Fig.
|
|
<figureCitation id="5CBE0C209A60F80B8FDE373006865639" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Left lateral habitus, head and thorax (dorsal view), wing (left, ventral view), and abdomen (dorsal view). A-D Dichaetophora tirlobita sp. n. (# 03877) E-G D. heterochroma sp. n. (# 03879) H-J D. flatosternata sp. n. (# 04172) K-M D. borneoensis sp. n. (# 03895) N-P D. javaensis sp. n. (# 03892) Q-S D. sumatraensis sp. n. (# 03890). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.665.11609.figure3" httpUri="https://binary.pensoft.net/fig/130047" pageId="3" pageNumber="124">3C</figureCitation>
|
|
); sclerotized, caudoventral bridge of cerci with small, narrow, apically pointed, median process and a pair of lateral, broad, apically rounded lobes (Fig.
|
|
<figureCitation id="42DB25873D7916FAFA99760092C257C2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dichaetophora trilobita sp. n. (A-H # 03877 I-K paratype # 03878). A Periphallic organs (posterior view), with red arrow indicating the median process on the caudoventral bridge of cerci B periphallic organs (posterolateral view), with red arrows indicating the prominent setae on the cercus and the anteroventral fusion of cercus (sclerotized, marginal plate) with the epandrium C surstyli (ventral view) D surstylus (inner side) E-G phallic organs (ventral, ventrolateral and lateral view, respectively) H paramedian setae (indicated with red arrows), and apical portion of paramere I, J oviscapt (lateral and ventral view, respectively) K spermathecae. Abbreviations: aed = aedeagus, aed a = aedeagal apodeme, cerc = cercus, epand = epandrium, hypd = hypandrium, pm = paramere, sur = surstylus, 10 S = tenth sternite. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.665.11609.figure5" httpUri="https://binary.pensoft.net/fig/130049" pageId="3" pageNumber="124">5A</figureCitation>
|
|
); aedeagus apically curved, hook-like, and finely wrinkled all over, subapically with numerous, coarse serrations, submedially not swollen dorsally (Fig.
|
|
<figureCitation id="3B7F97C16F85375617CA56E0514EF576" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dichaetophora trilobita sp. n. (A-H # 03877 I-K paratype # 03878). A Periphallic organs (posterior view), with red arrow indicating the median process on the caudoventral bridge of cerci B periphallic organs (posterolateral view), with red arrows indicating the prominent setae on the cercus and the anteroventral fusion of cercus (sclerotized, marginal plate) with the epandrium C surstyli (ventral view) D surstylus (inner side) E-G phallic organs (ventral, ventrolateral and lateral view, respectively) H paramedian setae (indicated with red arrows), and apical portion of paramere I, J oviscapt (lateral and ventral view, respectively) K spermathecae. Abbreviations: aed = aedeagus, aed a = aedeagal apodeme, cerc = cercus, epand = epandrium, hypd = hypandrium, pm = paramere, sur = surstylus, 10 S = tenth sternite. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.665.11609.figure5" httpUri="https://binary.pensoft.net/fig/130049" pageId="3" pageNumber="124">5G</figureCitation>
|
|
); spermatheca without distinct introvert (Fig.
|
|
<figureCitation id="1008A56328A5637E1BCA6952A6CEABEB" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dichaetophora trilobita sp. n. (A-H # 03877 I-K paratype # 03878). A Periphallic organs (posterior view), with red arrow indicating the median process on the caudoventral bridge of cerci B periphallic organs (posterolateral view), with red arrows indicating the prominent setae on the cercus and the anteroventral fusion of cercus (sclerotized, marginal plate) with the epandrium C surstyli (ventral view) D surstylus (inner side) E-G phallic organs (ventral, ventrolateral and lateral view, respectively) H paramedian setae (indicated with red arrows), and apical portion of paramere I, J oviscapt (lateral and ventral view, respectively) K spermathecae. Abbreviations: aed = aedeagus, aed a = aedeagal apodeme, cerc = cercus, epand = epandrium, hypd = hypandrium, pm = paramere, sur = surstylus, 10 S = tenth sternite. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.665.11609.figure5" httpUri="https://binary.pensoft.net/fig/130049" pageId="3" pageNumber="124">5K</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption id="46E564250A434A9F4B2ED07DF3BE4297" ID-Zenodo-Dep="989995" doi="10.3897/zookeys.665.11609.figure3" httpUri="https://binary.pensoft.net/fig/130047" pageId="3" pageNumber="124" start="Figure 3" startId="F3">
|
|
<paragraph id="9217823C656D115377D7602D710E7A81" pageId="3" pageNumber="124">
|
|
<emphasis id="F3A94B776D00AEEB8182BDE3599227D0" bold="true" pageId="3" pageNumber="124">Figure 3.</emphasis>
|
|
Left lateral habitus, head and thorax (dorsal view), wing (left, ventral view), and abdomen (dorsal view).
|
|
<emphasis id="7A5F2DBF4148F930C250F78FCD373885" bold="true" pageId="3" pageNumber="124">A-D</emphasis>
|
|
<taxonomicName id="FEFEFF6527DF6415AE134CDD3CF4A0B6" authorityName="Yang & Gao" authorityYear="2017" class="Insecta" family="Drosophilidae" genus="Dichaetophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichaetophora tirlobita" order="Diptera" pageId="3" pageNumber="124" phylum="Arthropoda" rank="species" species="tirlobita">
|
|
<emphasis id="5E849224A1F316798A1E1E74A5BBD9F0" italics="true" pageId="3" pageNumber="124">Dichaetophora tirlobita</emphasis>
|
|
</taxonomicName>
|
|
sp. n. (#03877)
|
|
<emphasis id="873EB6227CE9028BA8AECFDE1B971811" bold="true" pageId="3" pageNumber="124">E-G</emphasis>
|
|
<taxonomicName id="097169D88E060FB3CF59EFBD5530CA4D" lsidName="D. heterochroma" pageId="3" pageNumber="124" rank="species" species="heterochroma">
|
|
<emphasis id="5E8BD7F95DF4763902C82813AB4AE336" italics="true" pageId="3" pageNumber="124">D. heterochroma</emphasis>
|
|
</taxonomicName>
|
|
sp. n. (#03879)
|
|
<emphasis id="3A1EACE1CEA3AA15AE47EE7D141AF85E" bold="true" pageId="3" pageNumber="124">H-J</emphasis>
|
|
<taxonomicName id="930907312EAB13D75BA3A96B5A6FB6DD" lsidName="D. flatosternata" pageId="3" pageNumber="124" rank="species" species="flatosternata">D. flatosternata</taxonomicName>
|
|
sp. n. (#04172)
|
|
<emphasis id="80980A49DC31247086474DC42C3876D8" bold="true" pageId="3" pageNumber="124">K-M</emphasis>
|
|
<taxonomicName id="CDA74FAA4467EE59C0FE334000E51EF0" lsidName="D. borneoensis" pageId="3" pageNumber="124" rank="species" species="borneoensis">
|
|
<emphasis id="955D5AFBC514C7DDB4AA45B9C38435CB" italics="true" pageId="3" pageNumber="124">D. borneoensis</emphasis>
|
|
</taxonomicName>
|
|
sp. n. (#03895)
|
|
<emphasis id="3F7E5AD066C720F6FA65658477167E48" bold="true" pageId="3" pageNumber="124">N-P</emphasis>
|
|
<taxonomicName id="A51FE5E071E0825985BA8B916C585B30" lsidName="D. javaensis" pageId="3" pageNumber="124" rank="species" species="javaensis">
|
|
<emphasis id="3B32C61CC887A4A595C87E6132ECEE21" italics="true" pageId="3" pageNumber="124">D. javaensis</emphasis>
|
|
</taxonomicName>
|
|
sp. n. (#03892)
|
|
<emphasis id="72DB2DB9872EC2389DE2594A8343B1A9" bold="true" pageId="3" pageNumber="124">Q-S</emphasis>
|
|
<taxonomicName id="912D817CFAC30DE17D11D27590209785" lsidName="D. sumatraensis" pageId="3" pageNumber="124" rank="species" species="sumatraensis">
|
|
<emphasis id="92543FAAC3336DB2F74264E7D7382E7E" italics="true" pageId="3" pageNumber="124">D. sumatraensis</emphasis>
|
|
</taxonomicName>
|
|
sp. n. (#03890). Scale bars: 1.0 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="149345B1D81D038EA965C6D2F301E6B4" ID-Zenodo-Dep="989997" doi="10.3897/zookeys.665.11609.figure4" httpUri="https://binary.pensoft.net/fig/130048" pageId="3" pageNumber="124" start="Figure 4" startId="F4">
|
|
<paragraph id="CBD060CD8AF97073CBC077EAEAC83562" pageId="3" pageNumber="124">
|
|
<emphasis id="15495149178B3F8B32360C721789E211" bold="true" pageId="3" pageNumber="124">Figure 4.</emphasis>
|
|
Head (anterior view), postocciput, palpus, and prementum (ventral and lateral view, respectively).
|
|
<emphasis id="25365CE67ED97E095880CE9E1088D30D" bold="true" pageId="3" pageNumber="124">A-E</emphasis>
|
|
<taxonomicName id="1A59C1837958B25F7859A669E67A05AF" authorityName="Yang & Gao" authorityYear="2017" class="Insecta" family="Drosophilidae" genus="Dichaetophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichaetophora tirlobita" order="Diptera" pageId="3" pageNumber="124" phylum="Arthropoda" rank="species" species="tirlobita">
|
|
<emphasis id="36274C84C7EA446E955079271469B7B2" italics="true" pageId="3" pageNumber="124">Dichaetophora tirlobita</emphasis>
|
|
</taxonomicName>
|
|
sp. n. (#03877)
|
|
<emphasis id="2F77E0E1FBACABE1CCDB869F70DFF8BA" bold="true" pageId="3" pageNumber="124">F-J</emphasis>
|
|
<taxonomicName id="A6839AE9F0199F68C69542D0B1AB7F70" lsidName="D. heterochroma" pageId="3" pageNumber="124" rank="species" species="heterochroma">
|
|
<emphasis id="FF376CFE89C491BCCB6E94636DE47EAF" italics="true" pageId="3" pageNumber="124">D. heterochroma</emphasis>
|
|
</taxonomicName>
|
|
sp. n. (#03879)
|
|
<emphasis id="35C9E605525CAEFEA051477986432F05" bold="true" pageId="3" pageNumber="124">K-O</emphasis>
|
|
<taxonomicName id="60879252DD8EC13392FAD2C0CA6C5466" lsidName="D. flatosternata" pageId="3" pageNumber="124" rank="species" species="flatosternata">
|
|
<emphasis id="36F25AB6A42D754D0D97F95269FA9DFA" italics="true" pageId="3" pageNumber="124">D. flatosternata</emphasis>
|
|
</taxonomicName>
|
|
sp. n. (#04172)
|
|
<emphasis id="B8A8ADE4D3C440732D291C881D55EE1C" bold="true" pageId="3" pageNumber="124">P-T</emphasis>
|
|
<taxonomicName id="9DAF2EE14051AB74669B029DACD2A887" lsidName="D. borneoensis" pageId="3" pageNumber="124" rank="species" species="borneoensis">
|
|
<emphasis id="A0CB182C286B781C2B290B0FA2A95F68" italics="true" pageId="3" pageNumber="124">D. borneoensis</emphasis>
|
|
</taxonomicName>
|
|
sp. n. (#03895)
|
|
<emphasis id="E4086B8ACEEC3C8FCED6698740195E78" bold="true" pageId="3" pageNumber="124">U-Y</emphasis>
|
|
<taxonomicName id="885F20F8D5EEDADAFCA8C592DC910D08" lsidName="D. javaensis" pageId="3" pageNumber="124" rank="species" species="javaensis">
|
|
<emphasis id="E31AAD710981B64586BE7D3ACB5E75D3" italics="true" pageId="3" pageNumber="124">D. javaensis</emphasis>
|
|
</taxonomicName>
|
|
sp. n. (#03892)
|
|
<emphasis id="2F1F989FBAF6081087F22CDE57B2EC47" bold="true" pageId="3" pageNumber="124">Z-D</emphasis>
|
|
<taxonomicName id="0D07995E2DCA6E22FFFDBCC26CE774E9" lsidName="D. sumatraensis" pageId="3" pageNumber="124" rank="species" species="sumatraensis">
|
|
<emphasis id="58858D267085976148E818B2D77037DA" italics="true" pageId="3" pageNumber="124">D. sumatraensis</emphasis>
|
|
</taxonomicName>
|
|
sp. n. (#03890). Scale bars: 0.1 mm except for
|
|
<emphasis id="C02EA6C2235A2000C9D154237467E5B9" bold="true" pageId="3" pageNumber="124">A, F, K, P, U, Z</emphasis>
|
|
(0.5 mm).
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="482C001EAB6DF734CA5AB2CE806DBE04" ID-Zenodo-Dep="989999" doi="10.3897/zookeys.665.11609.figure5" httpUri="https://binary.pensoft.net/fig/130049" pageId="3" pageNumber="124" start="Figure 5" startId="F5">
|
|
<paragraph id="21C2D18095729E1A5514F5A69C5CAC7C" pageId="3" pageNumber="124">
|
|
<emphasis id="833EAAC397D5869A454F2A6C5FBD59E4" bold="true" pageId="3" pageNumber="124">Figure 5.</emphasis>
|
|
<taxonomicName id="4B41075762366F772B166CEEEEEC62F6" authorityName="Yang & Gao" authorityYear="2017" class="Insecta" family="Drosophilidae" genus="Dichaetophora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dichaetophora trilobita" order="Diptera" pageId="3" pageNumber="124" phylum="Arthropoda" rank="species" species="trilobita">
|
|
<emphasis id="00189D518EFE9AF4880AFC67FB70C067" italics="true" pageId="3" pageNumber="124">Dichaetophora trilobita</emphasis>
|
|
</taxonomicName>
|
|
sp. n. (
|
|
<emphasis id="0B903402865C6A62E2A79737573ABA98" bold="true" pageId="3" pageNumber="124">A-H</emphasis>
|
|
#03877
|
|
<emphasis id="E6930178FD8099B5B27C7F849F49824F" bold="true" pageId="3" pageNumber="124">I-K</emphasis>
|
|
paratype #03878).
|
|
<emphasis id="E8AD522EC224D8F0ACD9775F124DE15D" bold="true" pageId="3" pageNumber="124">A</emphasis>
|
|
Periphallic organs (posterior view), with red arrow indicating the median process on the caudoventral bridge of cerci
|
|
<emphasis id="2F658BCD3F5C51240C54C140686505B6" bold="true" pageId="3" pageNumber="124">B</emphasis>
|
|
periphallic organs (posterolateral view), with red arrows indicating the prominent setae on the cercus and the anteroventral fusion of cercus (sclerotized, marginal plate) with the epandrium
|
|
<emphasis id="3363EDD4904E2DF3C71A7FEE09B14D78" bold="true" pageId="3" pageNumber="124">C</emphasis>
|
|
surstyli (ventral view)
|
|
<emphasis id="14F7020979233EA7D33CBE4378A18E91" bold="true" pageId="3" pageNumber="124">D</emphasis>
|
|
surstylus (inner side)
|
|
<emphasis id="EFAD7CDD9820BE9ED00A9FFA39859AC3" bold="true" pageId="3" pageNumber="124">E-G</emphasis>
|
|
phallic organs (ventral, ventrolateral and lateral view, respectively)
|
|
<emphasis id="376B6B42EA2D92E47D04D47F75D8028B" bold="true" pageId="3" pageNumber="124">H</emphasis>
|
|
paramedian setae (indicated with red arrows), and apical portion of paramere
|
|
<emphasis id="2FDA1C6E16695E3B1957CA0635EA20B7" bold="true" pageId="3" pageNumber="124">I, J</emphasis>
|
|
oviscapt (lateral and ventral view, respectively)
|
|
<emphasis id="251DAF72149307E58105EC4FEFF227B5" bold="true" pageId="3" pageNumber="124">K</emphasis>
|
|
spermathecae. Abbreviations: aed = aedeagus, aed a = aedeagal apodeme, cerc = cercus, epand = epandrium, hypd = hypandrium, pm = paramere, sur = surstylus, 10S = tenth sternite. Scale bars: 0.1 mm.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="270A143535612B0100A95E9A38346A17" pageId="3" pageNumber="124" type="description">
|
|
<paragraph id="38C6CD0BBC4718D4E447A00D26485F87" pageId="3" pageNumber="124">Description.</paragraph>
|
|
<paragraph id="7B0F5875970B86194A03608E1B4DA0A2" pageId="3" pageNumber="124">
|
|
<emphasis id="CC64963FCBC4D615205FEA50A83A521D" italics="true" pageId="3" pageNumber="124">Head</emphasis>
|
|
(Figs
|
|
<figureCitation id="9DF53F5C329825958A8E6D536B3F5BB7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Left lateral habitus, head and thorax (dorsal view), wing (left, ventral view), and abdomen (dorsal view). A-D Dichaetophora tirlobita sp. n. (# 03877) E-G D. heterochroma sp. n. (# 03879) H-J D. flatosternata sp. n. (# 04172) K-M D. borneoensis sp. n. (# 03895) N-P D. javaensis sp. n. (# 03892) Q-S D. sumatraensis sp. n. (# 03890). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.665.11609.figure3" httpUri="https://binary.pensoft.net/fig/130047" pageId="3" pageNumber="124">3A, B</figureCitation>
|
|
,
|
|
<figureCitation id="C1C0F9B96B7300802AD2668FDF501A63" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Head (anterior view), postocciput, palpus, and prementum (ventral and lateral view, respectively). A-E Dichaetophora tirlobita sp. n. (# 03877) F-J D. heterochroma sp. n. (# 03879) K-O D. flatosternata sp. n. (# 04172) P-T D. borneoensis sp. n. (# 03895) U-Y D. javaensis sp. n. (# 03892) Z-D D. sumatraensis sp. n. (# 03890). Scale bars: 0.1 mm except for A, F, K, P, U, Z (0.5 mm)." figureDoi="10.3897/zookeys.665.11609.figure4" httpUri="https://binary.pensoft.net/fig/130048" pageId="3" pageNumber="124">4A-E</figureCitation>
|
|
): First flagellomere grayish yellow. Dorsolateral tentorial apodemes slightly divergent in basal 2/5 but strongly divergent in distal 3/5; supracervical setae 21-23 per side; postocular setae 22-23 per side. Palpus with two subprominent, lateromedial setae. Cibarium with ca. 70 medial and ca. 8 posterior sensilla per side.
|
|
</paragraph>
|
|
<paragraph id="FEBEFFDD88DCAABA4F815B6562E3CF93" pageId="3" pageNumber="124">
|
|
<emphasis id="771A5DD44DE89C82960A1DE86DABCF4D" italics="true" pageId="3" pageNumber="124">Wings</emphasis>
|
|
(Fig.
|
|
<figureCitation id="B716B7AE0C9FC2D1BF5EA2B83D81FEB5" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Left lateral habitus, head and thorax (dorsal view), wing (left, ventral view), and abdomen (dorsal view). A-D Dichaetophora tirlobita sp. n. (# 03877) E-G D. heterochroma sp. n. (# 03879) H-J D. flatosternata sp. n. (# 04172) K-M D. borneoensis sp. n. (# 03895) N-P D. javaensis sp. n. (# 03892) Q-S D. sumatraensis sp. n. (# 03890). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.665.11609.figure3" httpUri="https://binary.pensoft.net/fig/130047" pageId="3" pageNumber="124">3C</figureCitation>
|
|
): Veins pale brown to brown. Halter pale gray; stem darker.
|
|
</paragraph>
|
|
<paragraph id="C236DE88E34A9B195D940AED535756A6" pageId="3" pageNumber="124">
|
|
<emphasis id="B287B52EAE0E52BFC29D9D1CD2609825" italics="true" pageId="3" pageNumber="124">Legs</emphasis>
|
|
(Fig.
|
|
<figureCitation id="2BB4D78CC78396C5DC5D18694EDEC435" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Left lateral habitus, head and thorax (dorsal view), wing (left, ventral view), and abdomen (dorsal view). A-D Dichaetophora tirlobita sp. n. (# 03877) E-G D. heterochroma sp. n. (# 03879) H-J D. flatosternata sp. n. (# 04172) K-M D. borneoensis sp. n. (# 03895) N-P D. javaensis sp. n. (# 03892) Q-S D. sumatraensis sp. n. (# 03890). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.665.11609.figure3" httpUri="https://binary.pensoft.net/fig/130047" pageId="3" pageNumber="124">3A</figureCitation>
|
|
) pale grayish yellow; mid- and hindleg femora distally, foreleg tibia and tarsi proximally to medially darker. Apical setae present on all tibiae; hindleg apical seta short, stout. Mid-leg first tarsomere longer than total length of four succeeding tarsomeres; hindleg first tarsomere as long as total length of four succeeding tarsomeres.
|
|
</paragraph>
|
|
<paragraph id="9FBD58033204E1F46DDD7D1342C1EF6A" pageId="3" pageNumber="124">
|
|
<emphasis id="169BF55E0974442A282E713F1C531C5C" italics="true" pageId="3" pageNumber="124">Male terminalia</emphasis>
|
|
(Fig.
|
|
<figureCitation id="A093AC631A5F75B0AA35A0806FF36D75" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dichaetophora trilobita sp. n. (A-H # 03877 I-K paratype # 03878). A Periphallic organs (posterior view), with red arrow indicating the median process on the caudoventral bridge of cerci B periphallic organs (posterolateral view), with red arrows indicating the prominent setae on the cercus and the anteroventral fusion of cercus (sclerotized, marginal plate) with the epandrium C surstyli (ventral view) D surstylus (inner side) E-G phallic organs (ventral, ventrolateral and lateral view, respectively) H paramedian setae (indicated with red arrows), and apical portion of paramere I, J oviscapt (lateral and ventral view, respectively) K spermathecae. Abbreviations: aed = aedeagus, aed a = aedeagal apodeme, cerc = cercus, epand = epandrium, hypd = hypandrium, pm = paramere, sur = surstylus, 10 S = tenth sternite. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.665.11609.figure5" httpUri="https://binary.pensoft.net/fig/130049" pageId="3" pageNumber="124">5A-H</figureCitation>
|
|
): Epandrium pubescent except for anterior to ventral margin, with 1-3 dorsal and ca. 22 ventral, long setae per side. Surstylus with peg-like prensisetae in sinuate row on caudal margin, 2-5 apically pointed spines and medial patch of pubescence on outer surface and ca. 26 apically pointed spines (ventral ones shorter, thicker and straight, but medial to dorsal ones trichoid and recurved) on inner, caudal portion. Tenth sternite nearly flat. Cercus pubescent except for caudal margin, with 11-12 long setae near dorsal to posterior margin and 7-11 short setae in cluster on ventral portion of sclerotized marginal plate. Paramere with sensilla apically.
|
|
</paragraph>
|
|
<paragraph id="D0A53AEE838CA8F539B27CB1359A68EF" pageId="3" pageNumber="124">
|
|
<emphasis id="552676533F5577A79C3310CBA7EFD2B3" italics="true" pageId="3" pageNumber="124">Female terminalia</emphasis>
|
|
(Fig.
|
|
<figureCitation id="EDB34F7D0AEBA505FD8490CA72D8DEC3" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Dichaetophora trilobita sp. n. (A-H # 03877 I-K paratype # 03878). A Periphallic organs (posterior view), with red arrow indicating the median process on the caudoventral bridge of cerci B periphallic organs (posterolateral view), with red arrows indicating the prominent setae on the cercus and the anteroventral fusion of cercus (sclerotized, marginal plate) with the epandrium C surstyli (ventral view) D surstylus (inner side) E-G phallic organs (ventral, ventrolateral and lateral view, respectively) H paramedian setae (indicated with red arrows), and apical portion of paramere I, J oviscapt (lateral and ventral view, respectively) K spermathecae. Abbreviations: aed = aedeagus, aed a = aedeagal apodeme, cerc = cercus, epand = epandrium, hypd = hypandrium, pm = paramere, sur = surstylus, 10 S = tenth sternite. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.665.11609.figure5" httpUri="https://binary.pensoft.net/fig/130049" pageId="3" pageNumber="124">5I-K</figureCitation>
|
|
): Oviscapt valve dorsomedially narrowly extended, with 3-4 lateral and 9-13 marginal, peg-like ovisensilla.
|
|
</paragraph>
|
|
<paragraph id="BE2E712B97F960E52B4410D5AFE6BE28" pageId="3" pageNumber="124">
|
|
<emphasis id="21E8E63019380BA99F912D51890D7ACA" italics="true" pageId="3" pageNumber="124">Measurements</emphasis>
|
|
(in mm): BL (straight distance from anterior edge of pedicel to tip of abdomen) = 2.03 in holotype (1♂ paratype: 2.00; range in 2♀ paratypes: 2.00-2.21), ThL (distance from anterior notal margin to apex of scutellum) = 0.84 (0.82; 0.86-0.88), WL (distance from humeral cross vein to wing apex) = 1.62 (1.69; 1.72-1.73), WW (maximum wing width) = 0.71 (0.72; 0.74-0.79).
|
|
</paragraph>
|
|
<paragraph id="19325E797EBF1A802A0E92E751D2CD74" pageId="3" pageNumber="124">
|
|
<emphasis id="14F93CCAC7A7595CD5B844BE2FF5DE8B" italics="true" pageId="3" pageNumber="124">Indices</emphasis>
|
|
: FW/HW (frontal width/head width) = 0.53 (range in 1♂, 2♀, or less if noted, paratypes: 0.38-0.50), ch/o (maximum width of gena/maximum diameter of eye) = 0.18 (0.19-0.26), prorb (proclinate orbital seta/posterior reclinate orbital seta in length) = 0.72 (2♀: 0.71-0.80), rcorb (anterior reclinate orbital seta/posterior reclinate orbital seta in length) = 0.30 (2♀: 0.31-0.33), orbito (distance between proclinate and posterior reclinate orbital setae / distance between inner vertical and posterior reclinate orbital setae) = 0.60 (0.58-0.68), dcl (anterior dorsocentral seta/posterior dorsocentral seta in length) = 0.78 (2♀: 0.64-0.74), sctl (basal scutellar seta/apical scutellar seta in length) = 0.67 (1♀: 0.60), sterno (anterior katepisternal seta/posterior katepisternal seta in length) = 0.51 (0.57-0.60), dcp (distance between ipsilateral dorsocentral setae/distance between anterior dorsocentral setae) = 0.49 (0.53-0.60), sctlp (distance between ipsilateral scutellar setae/distance between apical scutellar setae) = 0.79 (0.68-0.77), C (2nd costal section between subcostal break and R2+3/3rd costal section between R2+3 and R4+5) = 1.57 (1.35-1.53), 4c (3rd costal section between R2+3 and R4+5/M1 between r-m and dm-cu) = 1.54 (1.46-1.66), 4v (M1 between dm-cu and wing margin/M1 between r-m and dm-cu) = 2.19 (2.11-2.33), 5x (CuA1 between dm-cu and wing margin/dm-cu between M1 and CuA1) = 2.19 (2.00-2.38), ac (3rd costal section between R2+3 and R4+5/distance between distal ends of R4+5 and M1) = 3.58 (3.66-3.76), M (CuA1 between dm-cu and wing margin/M1 between r-m and dm-cu) = 0.72 (0.66-0.70), C3F (length of heavy setation in 3rd costal section/length of 3rd costal section) = 0.70 (0.66-0.68).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="71934E656C6E00F19B83DCB028E5424E" pageId="3" pageNumber="124" type="etymology">
|
|
<paragraph id="C67092B1D186A6F131054F51DEA68218" pageId="3" pageNumber="124">Etymology.</paragraph>
|
|
<paragraph id="C817330AFB4117845C1BB40E2DD3AD0F" pageId="3" pageNumber="124">Referring to the trilobed, caudoventral bridge of cercal sclerotized plates.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C4C54F36C76ADB6581F1380A4715046A" pageId="3" pageNumber="124" type="distribution">
|
|
<paragraph id="791F94983431640D471503E10E38468C" pageId="3" pageNumber="124">Distribution.</paragraph>
|
|
<paragraph id="3D7A681FD50C8D618C7D302658ECD26B" pageId="3" pageNumber="124">Malaysia (Peninsular Malaysia, Sarawak, Sabah).</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |