treatments-xml/data/3C/10/9A/3C109AC557765752B847663FF9039F68.xml

<document ID-CLB-Dataset="20767" ID-DOI="http://dx.doi.org/10.3897/phytokeys.189.49367" ID-GBIF-Dataset="49134428-816c-44b0-b17f-0e0711a9a714" ID-PMC="PMC8837500" ID-Pensoft-Pub="1314-2003-189-99" ID-Pensoft-UUID="9A120E648CC656AFAC749BE0FB1ABB37" ID-PubMed="35169386" ModsDocID="1314-2003-189-99" checkinTime="1644016132331" checkinUser="pensoft" docAuthor="Julius, Avelinah" docDate="2022" docId="3C109AC557765752B847663FF9039F68" docLanguage="en" docName="PhytoKeys 189: 99-127" docOrigin="PhytoKeys 189" docPubDate="2022-02-04" docSource="http://dx.doi.org/10.3897/phytokeys.189.49367" docTitle="Capparis trinervia subsp. var. var. chungiana Julius 2022, var. nov." docType="treatment" docVersion="7" id="9A120E648CC656AFAC749BE0FB1ABB37" lastPageNumber="99" masterDocId="9A120E648CC656AFAC749BE0FB1ABB37" masterDocTitle="Capparis (Capparaceae) in Peninsular Malaysia, including a new species and two new varieties" masterLastPageNumber="127" masterPageNumber="99" pageNumber="99" updateTime="1732824147665" updateUser="ExternalLinkService">
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<mods:title id="CC58FDD1E2B7A5649A9F3D063500FB27">Capparis (Capparaceae) in Peninsular Malaysia, including a new species and two new varieties</mods:title>
</mods:titleInfo>
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<mods:namePart id="7A884DAB054B872C4E75140710B67685">Julius, Avelinah</mods:namePart>
<mods:nameIdentifier id="0A4EE41F98BD9E2E10509FA560731A9A" type="ORCID">https://orcid.org/0000-0003-1991-1915</mods:nameIdentifier>
<mods:affiliation id="3A5F2D90C2C35BFC53D40754E5AA099E">Forest Research Institute Malaysia, Kepong, Selangor, 52109, Malaysia</mods:affiliation>
<mods:nameIdentifier id="6CAEDE6F059B5469E12D32B266622143" type="email">plagiovel80@yahoo.com</mods:nameIdentifier>
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<mods:title id="3F214B5F1C213689A5D451289A1F4FD2">PhytoKeys</mods:title>
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<mods:part id="B5C6C7797F1460C535B867EC6ACEFBD1">
<mods:date id="989A5EE99383269E4AA8FCA00CDC8BE1">2022</mods:date>
<mods:detail id="CF06287C80442DE3F4C1929824746D11" type="pubDate">
<mods:number id="379C56569DB3180B561B1A58DD8277D1">2022-02-04</mods:number>
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<mods:detail id="400A9C810D4C3BC73E8E1EE226EF7BD8" type="volume">
<mods:number id="DDD7E480958AFC03F8481D4533158980">189</mods:number>
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<mods:start id="66302572D9BA9F47B01F9D7FFC6C09E9">99</mods:start>
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<mods:url id="57E546F199B83F0680A9B9AA3DB20A55">http://dx.doi.org/10.3897/phytokeys.189.49367</mods:url>
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<mods:classification id="F66727A07230EA30497AABAE8730B138">journal article</mods:classification>
<mods:identifier id="8D4D460FF7A402FB7088CA3FC3ABE7E6" type="DOI">http://dx.doi.org/10.3897/phytokeys.189.49367</mods:identifier>
<mods:identifier id="C175AF86FC6CED822219E8731C71AE22" type="Pensoft-Pub">1314-2003-189-99</mods:identifier>
<mods:identifier id="5ADFBD4CAAADE471578FEA7DC0D84AC1" type="Pensoft-UUID">9A120E648CC656AFAC749BE0FB1ABB37</mods:identifier>
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<treatment id="3C109AC557765752B847663FF9039F68" ID-GBIF-Taxon="193278626" LSID="urn:lsid:plazi:treatment:3C109AC557765752B847663FF9039F68" httpUri="http://treatment.plazi.org/id/3C109AC557765752B847663FF9039F68" lastPageNumber="99" pageId="0" pageNumber="99" scope_family="Capparaceae">
<subSubSection id="3DF197DCA698002DAD2641E5B2069DA2" pageId="0" pageNumber="99" type="nomenclature">
<paragraph id="B7CFE1656E76A73C19EBC6A9670A2687" pageId="0" pageNumber="99">
<taxonomicName id="E3882652640F72CD59A9DAED90D980D9" ID-CoL="98XYM" LSID="3C109AC5-5776-5752-B847-663FF9039F68" authority="Julius" authorityName="Julius" authorityYear="2022" class="Magnoliopsida" family="Capparaceae" genus="Capparis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Capparis trinervia subsp. var. var. chungiana" order="Brassicales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="variety" species="trinervia" status="var. nov." subSpecies="var." variety="chungiana">Capparis trinervia var. chungiana Julius</taxonomicName>
<taxonomicNameLabel id="3A7280062AAC971565F5AE1AFD45C534" pageId="0" pageNumber="99">var. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="1661C77932D740982060FC8317772CDE" pageId="0" pageNumber="99" type="description">
<paragraph id="1274E6EBB1977849F68EF153888FB8D4" pageId="0" pageNumber="99">
<figureCitation id="17BB8F3BAE0FDB15BD0D8C666767EDA9" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Distribution of Capparis trinervia var. chungiana." figureDoi="10.3897/phytokeys.189.49367.figure6" httpUri="https://binary.pensoft.net/fig/642013" pageId="0" pageNumber="99">Figures 6</figureCitation>
<figureCitation id="7FE85A4711741DF7340E9AB64EB9F5C2" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Capparis trinervia var. chungiana Julius, var. nov. A the leafy branch B the flowering (bud) leafy twig C flower with sepals and petals removed exhibiting the filaments and ovary on gynophore D upper pedicel and base of calyx E the fruit in cross section F stipules close-up. (Drawn by Mohamad Aidil Noordin from FRI 64006 [A, E, F] and Mahmud Sidek s. n. [B-D])." figureDoi="10.3897/phytokeys.189.49367.figure7" httpUri="https://binary.pensoft.net/fig/642014" pageId="0" pageNumber="99">, 7</figureCitation>
<figureCitation id="13C393B1E5F43E82F876AE2A23FCF4DF" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Capparis trinervia var. chungiana Julius, var. nov. A fruiting twig B stipular thorns C young fruit on stout gynophore D leaf E young fruit, cross section. (Photographs by K. Imin)." figureDoi="10.3897/phytokeys.189.49367.figure8" httpUri="https://binary.pensoft.net/fig/642015" pageId="0" pageNumber="99">, 8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="89805DD64AA1283E37C0618A32011B00" pageId="0" pageNumber="99" type="diagnosis">
<paragraph id="AFB629774C6A109EFB64EA68C159A8E0" pageId="0" pageNumber="99">Diagnosis.</paragraph>
<paragraph id="9D998AA2516433FC4993E8FBCC6DA7B7" pageId="0" pageNumber="99">
Recognised in the genus 
<taxonomicName id="6F18C6D53A305AF84FC8CED6E25AA6C5" class="Magnoliopsida" family="Capparaceae" genus="Capparis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Capparis" order="Brassicales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="genus">
<emphasis id="887BB8D89C2C734832823901E8C15B54" italics="true" pageId="0" pageNumber="99">Capparis</emphasis>
</taxonomicName>
by the combination of the following characters: the leaves broadly ovate-elliptic, relatively large (13-16 
<normalizedToken id="D7A2FDDEB0AD91F4BBA27DE56AD96A0D" originalValue="×">x</normalizedToken>
5.5-8.5 cm) and leaves drying reddish-brown, rarely pale green with pale yellow and rarely dark red venation, the tertiary venation obscure, the inflorescence strictly terminal with flowers arranged racemosely and becoming crowded at the distal part of the inflorescence, the stamens 30-40 and the fruit globose with ca. 11 cm in diameter.
</paragraph>
</subSubSection>
<subSubSection id="7F97F7D3C38269DEAE3F72C6EBD94C6B" pageId="0" pageNumber="99" type="materials_examined">
<paragraph id="903A495E47F97A145F3C74153E0960E7" pageId="0" pageNumber="99">
<typeStatus id="8B397E955D2CA0CBB489B6C344FB6BFF">Type</typeStatus>
.
</paragraph>
<paragraph id="0FBD8F7D022652BD8229F53FC9001E13" pageId="0" pageNumber="99">
<materialsCitation id="CFAA2A2CF12A276538C33492063EACE9" collectingDate="1971-04-17" country="Malaysia" county="Peninsular" location="Fraser's Hill Road" municipality="Kuala Kubu Bharu" specimenCount="1" stateProvince="Selangor" typeStatus="holotype">
<collectingCounty id="BDC84DBB229AA3592A62EFF4B9AE8FBA">Peninsular</collectingCounty>
<collectingCountry id="CAA3604C2DA884BC45FBE231B236334A" name="Malaysia">Malaysia</collectingCountry>
. 
<collectingRegion id="2300FFF6B1D57D1990EFCCE69727A372" country="Malaysia" name="Selangor">Selangor</collectingRegion>
: 
<collectingMunicipality id="DB530BB05E41F738ABFED44F019C20CA">Kuala Kubu Bharu</collectingMunicipality>
to 
<location id="B6C3041DF497CA64E0A97F0077DD57F6" LSID="urn:lsid:plazi:treatment:3C109AC557765752B847663FF9039F68:B6C3041DF497CA64E0A97F0077DD57F6" country="Malaysia" county="Peninsular" municipality="Kuala Kubu Bharu" name="Fraser's Hill Road" stateProvince="Selangor">
<normalizedToken id="F24C95F7E01A0A526F739594B60921B7" originalValue="Fraser’s">Fraser's</normalizedToken>
Hill Road
</location>
, along the road side, 
<collectingDate id="A9CFEA200068C7B29D9E4FE74CB33703" value="1971-04-17">17 Apr 1971</collectingDate>
(fl.), 
<emphasis id="7631276D3C0E4EF9F499DBA1CDDC97D5" italics="true" pageId="0" pageNumber="99">Mahmud Sidek s.n.</emphasis>
(
<typeStatus id="AC2B6956399868CB5D219F10CA2DC660">holotype</typeStatus>
KLU!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="028D2A20E21781627D0C1E6E711EC519" pageId="0" pageNumber="99" type="description">
<paragraph id="3FDC1A10A78D85DB09E152BDEA5035B9" pageId="0" pageNumber="99">Description.</paragraph>
<paragraph id="7CC1D225ED0F2E78B7FC12121DDC79B8" pageId="0" pageNumber="99">
Woody climber 9-12 m long. 
<emphasis id="4406E35F63EBB757A8D63BD20A208965" bold="true" italics="true" pageId="0" pageNumber="99">Twigs</emphasis>
straight, covered with velvety, ferruginous hairs and glabrescent with age. 
<emphasis id="E7E9DD282D6B866A515A575F82FFAFF5" bold="true" italics="true" pageId="0" pageNumber="99">Stipular thorns</emphasis>
recurved downwards, tips 1.5-3 mm long. 
<emphasis id="87F3BAB9AA23038502BF761F42D3DCC9" bold="true" italics="true" pageId="0" pageNumber="99">Leaves</emphasis>
spirally arranged; petioles 15-20 
<normalizedToken id="9D4AEAD4B065979E9ECB795C460EAC98" originalValue="×">x</normalizedToken>
2-3 mm, stout, incrassate or thickened, velvety hairy becoming glabrous; lamina broadly ovate sometimes ovate-elliptic, 13-16 
<normalizedToken id="69AEAF4D5667ED11DC1508040C723757" originalValue="×">x</normalizedToken>
5.5-8.5 cm, coriaceous to subcoriaceous, surface rugose and usually reddish-brown when dry, rarely pale green, base cuneate or occasionally rounded, margin flat and entire, apex acute or shortly cuspidate, with acumen ca. 4 mm long, glabrous on both surfaces; mid-rib flat above, raised beneath; lateral veins 5-7(-8) pairs, flat above, distinct beneath, pale yellow when dry, looping towards the margin; intercostal veins obscure. 
<emphasis id="540891DE7A254C056CCF31958048CF0B" bold="true" italics="true" pageId="0" pageNumber="99">Inflorescences</emphasis>
terminal, flowers arranged racemosely and becoming crowded at the distal part of the inflorescence rachis slender and straight; bracts caducous. 
<emphasis id="084A6014E1CB22EDC1A3301C03D401D7" bold="true" italics="true" pageId="0" pageNumber="99">Flowers</emphasis>
[in bud] 4-7, buds globose, 1-1.8 
<normalizedToken id="539867F48C5B9CCE146D5A2E69540723" originalValue="×">x</normalizedToken>
1.3-1.8 cm; pedicels 3-3.5 cm long, swollen towards apex, velvety with ferruginous hairs; sepals 4, thick, whitish-green, orbicular, 1.2-1.5 
<normalizedToken id="FD0D9E730AB09F7B14CDC0F23251AC23" originalValue="×">x</normalizedToken>
1.2 cm, outer pair of sepals imbricate, covered with dense, velvety ferruginous hairs outside, glabrous inside, inner pair of sepals glabrous on both surfaces, except silky hairs at base outside; petals 4, cream or white with pink- or dark purple at base; stamens 30-40, unequal in length, anthers yellow; ovary green. 
<emphasis id="AB6E80EDDF258002E137B95AEA0F63E7" bold="true" italics="true" pageId="0" pageNumber="99">Fruits</emphasis>
young shiny and green, mature fruit n.v., globose to sub-globose on a stout gynophore, sometimes beaked when young, ca. 9 
<normalizedToken id="BE9A32E74202324C050BCFACCB9C3A52" originalValue="×">x</normalizedToken>
11 cm. 
<emphasis id="BB84CAB0BB8F5234D7C8A0350080A3E0" bold="true" italics="true" pageId="0" pageNumber="99">Seeds</emphasis>
(3-)12-15, sarcotesta whitish, testa thin and whitish-cream.
</paragraph>
</subSubSection>
<subSubSection id="725446206A68F25EE7EEC7321DC35E2C" pageId="0" pageNumber="99" type="vernacular names">
<paragraph id="EBF31202E7DCFFB5749E06B2AC57AFD3" pageId="0" pageNumber="99">Vernacular names.</paragraph>
<paragraph id="0A35B68445EF03589983C4521616B969" pageId="0" pageNumber="99">
<emphasis id="DC35712DA184F9FF7450962C9280F2A8" italics="true" pageId="0" pageNumber="99">Kuku lang</emphasis>
(Malay), 
<emphasis id="2C6866DBFB42BC5D864469582D4A3E6F" italics="true" pageId="0" pageNumber="99">mentimun</emphasis>
(Temuan).
</paragraph>
</subSubSection>
<subSubSection id="11615937DC9FE95C22941C10217E6DF6" pageId="0" pageNumber="99" type="distribution">
<paragraph id="F1FFF145B685DAC8DD4235892718F8E5" pageId="0" pageNumber="99">Distribution.</paragraph>
<paragraph id="55047D087FD01A979F5D30211B646801" pageId="0" pageNumber="99">
Endemic in Peninsular Malaysia, Negeri Sembilan, Selangor and Pahang (Fig. 
<figureCitation id="203DF5A3E025B820D60DF5E156DB2D6F" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Distribution of Capparis trinervia var. chungiana." figureDoi="10.3897/phytokeys.189.49367.figure6" httpUri="https://binary.pensoft.net/fig/642013" pageId="0" pageNumber="99">6</figureCitation>
).
</paragraph>
<caption id="8984F46CFC47417D8CA718BD4FAD326F" doi="10.3897/phytokeys.189.49367.figure6" httpUri="https://binary.pensoft.net/fig/642013" pageId="0" pageNumber="99" start="Figure 6" startId="F6">
<paragraph id="7FB1C25D81E6ADD9C9F0826BF5564AC8" pageId="0" pageNumber="99">
<emphasis id="D78DA68F24BEE9A312961E89E83EC6EE" bold="true" pageId="0" pageNumber="99">Figure 6.</emphasis>
Distribution of 
<taxonomicName id="BBBFC2AE7D5BCD3EFF3979EF15E53853" authorityName="Julius" authorityYear="2022" class="Magnoliopsida" family="Capparaceae" genus="Capparis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Capparis trinervia subsp. var. var. chungiana" order="Brassicales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="variety" species="trinervia" subSpecies="var." variety="chungiana">Capparis trinervia var. chungiana</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="B94848C30C32CC6A2E577E34C380ECE8" pageId="0" pageNumber="99" type="ecology">
<paragraph id="5DEA4C064E78B6EFB238F2D721B0B1C2" pageId="0" pageNumber="99">Ecology.</paragraph>
<paragraph id="ACDD09CF55C17A4DED113FFD2C560003" pageId="0" pageNumber="99">Lowland to lower montane forest, on forest edge or roadsides; 427-1067 m altitude.</paragraph>
</subSubSection>
<subSubSection id="27DB0AF206C5C3E77458563A0878B8B2" pageId="0" pageNumber="99" type="etymology">
<paragraph id="DA325207D2395044B4DB5B87F75A3AAD" pageId="0" pageNumber="99">Etymology.</paragraph>
<paragraph id="368C8EB1F9E8B741A333FAA3D919C595" pageId="0" pageNumber="99">The new variety is named after Dr Richard Chung Cheng Kong, Curator of the Kepong Herbarium (KEP) and Project Leader of Flora of Malaysia under 10thand 11th Malaysian Plans.</paragraph>
</subSubSection>
<subSubSection id="01333EA80BD0448144CBC976423E34EF" pageId="0" pageNumber="99" type="conservation status">
<paragraph id="EF0745CE83AE0F66CAC45A5A5ADEE27C" pageId="0" pageNumber="99">Conservation status.</paragraph>
<paragraph id="B5A3A5D551CCD732105C5F743033A404" pageId="0" pageNumber="99">
<taxonomicName id="A54EC54B5C0E61352E97428F9BCB6601" authorityName="Julius" authorityYear="2022" class="Magnoliopsida" family="Capparaceae" genus="Capparis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Capparis trinervia subsp. var. var. chungiana" order="Brassicales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="variety" species="trinervia" subSpecies="var." variety="chungiana">Capparis trinervia var. chungiana</taxonomicName>
inhabits forest margins, gaps or pathways near to roadsides, but is not a common species. This species has not been relocated in Selangor and Pahang even though the localities and adjacent areas have been revisited. This species, however, has been found and recorded from a new locality in Negeri Sembilan during a botanical survey in 2010, but it could not be relocated from two recent visits in 2014 and 2019. Recent field observations to Negeri Sembilan also show forest clearing pathways near the species habitat. Therefore, this species is assessed as Near Threatened following the IUCN Red List Categories and Criteria (
<bibRefCitation id="29BB5B4828EA78C756D50FF1A4CE16D3" author="IUCN" journalOrPublisher="Botany" pageId="0" pageNumber="99" refId="B18" refString="IUCN, 2001. IUCN Red List Categories and Criteria.Version 3.1. 1st edn. IUCN Species Survival Commission, IUCN, Gland, Switzerland and Cambridge, UK." title="IUCN Red List Categories and Criteria. Version 3.1. 1 st edn. IUCN Species Survival Commission, IUCN, Gland, Switzerland and Cambridge, UK." year="2001">IUCN 2001</bibRefCitation>
, 
<bibRefCitation id="E3D44BFD823479C3E1FE672D014B4F79" author="IUCN" journalOrPublisher="Botany" pageId="0" pageNumber="99" refId="B19" refString="IUCN, 2012. IUCN Red List Categories and Criteria. Version 3.1. 2nd edn. IUCN Species Survival Commission, IUCN, Gland, Switzerland and Cambridge, UK." title="IUCN Red List Categories and Criteria. Version 3.1. 2 nd edn. IUCN Species Survival Commission, IUCN, Gland, Switzerland and Cambridge, UK." year="2012">2012</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A4D5C58BB56A3945903B8D1D18959089" pageId="0" pageNumber="99" type="materials_examined">
<paragraph id="B20B2F3352C9D506C59DE785E87ACECF" pageId="0" pageNumber="99">Additional specimens examined.</paragraph>
<paragraph id="D298F1ED6E9B0306C2EBBB31E6188529" pageId="0" pageNumber="99">
<emphasis id="F91ED08D632378A4913E1862B04A1C7A" bold="true" pageId="0" pageNumber="99">
Peninsular 
<materialsCitation id="2A0B9DF9FD3E43E4C3F496000DF0B5C5" accessionNumber="FRI16263" collectingDate="1934-01-10" collectingDateMax="1970-12-11" collectingDateMin="1934-01-10" collectorName="Ulu Gombak FR, Mohd. Kasim, Mahmud Sidek, M. E. D. Poore, Kepong Plantation Field, Abdul Hamid, Ulu Langat, Gadoh KL, Genting Sempah, Kochummen" country="Malaysia" elevation="259" location="Gombak Forest" specimenCount="1" stateProvince="Selangor">
<collectingCountry id="4997E02660F976043FEABF702B208D11" name="Malaysia">Malaysia</collectingCountry>
. 
<collectingRegion id="97F3E9A23E33BDD46EF571A931DC4FE9" country="Malaysia" name="Selangor">Selangor</collectingRegion>
: 
<collectorName id="A18524C34F9C4379792E2AAA17E71F36">Ulu Gombak FR</collectorName>
, 
<quantity id="1AB4497CABB1FB4B8F5EE210456884B7" metricMagnitude="2" metricUnit="m" metricValue="2.59" unit="m" value="259.0">
<elevation id="36EF0B195CCA840E4AC9462E851733D9" metricMagnitude="2" metricUnit="m" metricValue="2.59" unit="m" value="259.0">259 m</elevation>
</quantity>
(
<quantity id="BB2E70F7F95C5172674020CC4F9EFAD2" metricMagnitude="2" metricUnit="m" metricValue="2.5908" unit="ft" value="850.0">
<elevation id="0D09FA68C837469351F0899EDA672779" metricMagnitude="2" metricUnit="m" metricValue="2.5908" unit="ft" value="850.0">850 ft</elevation>
</quantity>
) alt., 
<collectingDate id="9454496DDE545C782D52480DBA935577" value="1964-08-19">19 Aug 1964</collectingDate>
(fr.), 
<emphasis id="BF5EAB72A059EEA3E77AECD34E991B4D" italics="true" pageId="0" pageNumber="99">
<collectorName id="BF80C282B3E033A1121D0F205F85DDCC">Mohd. Kasim</collectorName>
&amp; 
<collectorName id="AAF26C1708278F55095CAA5B7D06D602">Mahmud Sidek</collectorName>
624
</emphasis>
(KLU!); 
<location id="F798E25818E4AF5178EA0BD57949C384" LSID="urn:lsid:plazi:treatment:3C109AC557765752B847663FF9039F68:F798E25818E4AF5178EA0BD57949C384" country="Malaysia" name="Gombak Forest" stateProvince="Selangor">Gombak Forest</location>
, 16th mile, followed path to stream and beyond, 
<quantity id="A93454F671E5B989B9407043444A707B" metricMagnitude="2" metricUnit="m" metricValue="4.27" unit="m" value="427.0">
<elevation id="5821B8C0C901CA3FBCA1DC018CB2521C" metricMagnitude="2" metricUnit="m" metricValue="4.27" unit="m" value="427.0">427 m</elevation>
</quantity>
(
<quantity id="4148FD3D3708F224211711E65937854A" metricMagnitude="2" metricUnit="m" metricValue="4.2672" unit="ft" value="1400.0">
<elevation id="5B76CE05F3A51AA7E9E30445095BCB14" metricMagnitude="2" metricUnit="m" metricValue="4.2672" unit="ft" value="1400.0">1400 ft</elevation>
</quantity>
) alt., 
<collectingDate id="BB48BA7F884BC83DCDBB6A3CAAAC88DA" value="1963-06-12">12 Jun 1963</collectingDate>
(fl.), 
<emphasis id="0F91A68C1987DEE12E5C94A04AE9ED90" italics="true" pageId="0" pageNumber="99">
<collectorName id="8B17CF91265366BDEE71C5FA43B82A95">M.E.D. Poore</collectorName>
1189
</emphasis>
(KLU!); 
<collectorName id="CA968CC07FCC96614547988266102AAB">Kepong Plantation Field</collectorName>
9J, hillslope, 
<collectingDate id="3EFB7B1F37413B5A45923F5CB92899D8" value="1934-01-10">10 Jan 1934</collectingDate>
(fl.), 
<emphasis id="5B5AF7C01EF319EAD04446DDD62EEA30" italics="true" pageId="0" pageNumber="99">
<collectorName id="120761619BCFBB307EEF7A21E38AC0E7">Abdul Hamid</collectorName>
33570
</emphasis>
(KEP!); 
<collectorName id="AA6DD20714BB47F72417FC779938220E">Ulu Langat</collectorName>
, 
<collectingDate id="D852B9DC546E2E0965C7080687BDDD3C" value="1960-03-30">30 Mar 1960</collectingDate>
(fl., fr.), 
<emphasis id="0C959AFA38293BBAD1FCE5A612D97982" italics="true" pageId="0" pageNumber="99">
<collectorName id="252405365C2495F233B8A99BB6BC5494">Gadoh KL</collectorName>
2072
</emphasis>
(KEP! 2 sheets); 
<collectorName id="A2ECED8D03AEA344B8DC49C773EF576B">Genting Sempah</collectorName>
, 
<collectingDate id="67E60D3EC54220FC0F198D9DADF560CB" value="1970-12-11">11 Dec 1970</collectingDate>
(fl., fr.), 
<emphasis id="7061C874E1A5D83E4A1E0CEB91D7E4C2" italics="true" pageId="0" pageNumber="99">
<collectorName id="D807EEFD46F1F376627D5251058243E0">Kochummen</collectorName>
<accessionNumber id="8BCF4532A5ADB4F79381C96894366704" httpUri="http://www.ncbi.nlm.nih.gov/protein/FRI16263">FRI16263</accessionNumber>
</emphasis>
(KEP!)
</materialsCitation>
</emphasis>
. 
<materialsCitation id="1151C1106E73226818A665B835C3930A" accessionNumber="FRI18471, FRI32242" collectingDate="1979-02-28" collectingDateMax="1981-08-26" collectingDateMin="1979-02-28" collectorName="K. M. Wong" country="Malaysia" elevation="1067" location="Kochummen" municipality="Fraser's Hill" specimenCount="1" stateProvince="Pahang">
<emphasis id="393050F8E6D4F18BACDDB64F094B6417" bold="true" pageId="0" pageNumber="99">
<collectingRegion id="9231C0CECC2FBD9D55267E714C93F5A1" country="Malaysia" name="Pahang">Pahang</collectingRegion>
</emphasis>
: 
<collectingMunicipality id="6C989B37CBC3F4109B8AA8FA28B10451">
<normalizedToken id="1DE51C846ED746E3F9FE4CFC31EEDFE4" originalValue="Fraser’s">Fraser's</normalizedToken>
Hill
</collectingMunicipality>
, near gate at gap, roadside, 
<quantity id="35CDC5B4429CABC4F892CBDD76B382E2" metricMagnitude="3" metricUnit="m" metricValue="1.067" unit="m" value="1067.0">
<elevation id="1F4C2E9CAB9F639B631CA2C70DDEC019" metricMagnitude="3" metricUnit="m" metricValue="1.067" unit="m" value="1067.0">1067 m</elevation>
</quantity>
(
<quantity id="7F0D63BCA2B9D9F0CEC1B1886673BD9A" metricMagnitude="3" metricUnit="m" metricValue="1.0668" unit="ft" value="3500.0">
<elevation id="6FD26A7058F64720E177AF982FD2D283" metricMagnitude="3" metricUnit="m" metricValue="1.0668" unit="ft" value="3500.0">3500 ft</elevation>
</quantity>
) alt., 
<collectingDate id="C11ADA30C395BC62203D6A686707A6B5" value="1979-02-28">28 Feb 1979</collectingDate>
(fl., fr.), 
<emphasis id="D8B4F3316FB9EABF0A4C6BD84E6375D7" italics="true" pageId="0" pageNumber="99">
<location id="A41A1014CF3EB04C16FF00F28CD0C24F" LSID="urn:lsid:plazi:treatment:3C109AC557765752B847663FF9039F68:A41A1014CF3EB04C16FF00F28CD0C24F" country="Malaysia" municipality="Fraser's Hill" name="Kochummen" stateProvince="Pahang">Kochummen</location>
<accessionNumber id="3ABAC207AD5480F7783FD97F07C84046" httpUri="http://www.ncbi.nlm.nih.gov/protein/FRI18471">FRI18471</accessionNumber>
</emphasis>
(KEP!); road to 
<normalizedToken id="4EA80A11EDD81193D856BA0E46235548" originalValue="Fraser’s">Fraser's</normalizedToken>
Hill, climber on Mahang tree beside the road, 
<quantity id="902F80EE71CEB7B3071F75B38A4E378F" metricMagnitude="2" metricUnit="m" metricValue="9.14" unit="m" value="914.0">
<elevation id="DC50036B63DE8089196789E7A0D8311E" metricMagnitude="2" metricUnit="m" metricValue="9.14" unit="m" value="914.0">914 m</elevation>
</quantity>
(
<quantity id="174CBA9A491B68E2D81A8061FF8D6B15" metricMagnitude="2" metricUnit="m" metricValue="9.144" unit="ft" value="3000.0">
<elevation id="11D1EBB09108A489EE1A94974D1B4B35" metricMagnitude="2" metricUnit="m" metricValue="9.144" unit="ft" value="3000.0">3000 ft</elevation>
</quantity>
) alt., 
<collectingDate id="AB3E53EF597201AF34BBCE903FDC56FD" value="1981-08-26">26 Aug 1981</collectingDate>
(fr.), 
<emphasis id="5EEE5447BFBAB4054D71CE43A5E16708" italics="true" pageId="0" pageNumber="99">
<collectorName id="04C0B9D00598B2B6E41D10D0D5029846">K.M. Wong</collectorName>
<accessionNumber id="D248A4F44DA0A7B04C49E725CE0C3BAA" httpUri="http://www.ncbi.nlm.nih.gov/protein/FRI32242">FRI32242</accessionNumber>
</emphasis>
(KEP!)
</materialsCitation>
. 
<materialsCitation id="8D54BE4046507045C760A6896EA2AF3E" accessionNumber="FRI64006" collectingDate="2010-05-06" collectorName="Kenaboi FR, G. Besar Hantu, Julius" country="Malaysia" elevation="500" latitude="3.1833334" location="Kg. London area" longLatPrecision="13" longitude="102.012665" municipality="Jelebu" specimenCount="1" stateProvince="Negeri Sembilan">
<emphasis id="3FCE856011475D4942B82739914A3A5D" bold="true" pageId="0" pageNumber="99">
<collectingRegion id="E11CBB98A93041FCED7EB22F027BA960" country="Malaysia" name="Negeri Sembilan">Negeri Sembilan</collectingRegion>
</emphasis>
: 
<collectingMunicipality id="FD3F3F91FE8299353D48DB44384DF9F6">Jelebu</collectingMunicipality>
, 
<collectorName id="E2E5D38BAE7B9739AC2A69E5BAB02437">Kenaboi FR</collectorName>
, 
<collectorName id="BECD1E14979E28F4BEF45DAC99964B8C">G. Besar Hantu</collectorName>
, near 
<location id="0E56BA4A22A6B16CFC32BC558471C185" LSID="urn:lsid:plazi:treatment:3C109AC557765752B847663FF9039F68:0E56BA4A22A6B16CFC32BC558471C185" country="Malaysia" latitude="3.1833334" longLatPrecision="13" longitude="102.012665" municipality="Jelebu" name="Kg. London area" stateProvince="Negeri Sembilan">Kg. London area</location>
, 
<geoCoordinate id="EA1742F00B8C07816A0E8F4E954B2253" degrees="3" direction="north" minutes="11.00" orientation="latitude" precision="9" value="3.1833334">3°11.00'N</geoCoordinate>
, 
<geoCoordinate id="AEB6113353AF0C874976714D1F157E96" degrees="102" direction="east" minutes="00.76" orientation="longitude" precision="9" value="102.012665">102°00.76'E</geoCoordinate>
, 
<quantity id="6320B48B7A5832AFB8739557458F06D3" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">
<elevation id="1DB9D1F302B65BA68EE4BD1884BA824D" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">500 m</elevation>
</quantity>
alt., 
<collectingDate id="A82E9504CA3243C9091B702EA924FA03" value="2010-05-06">6 May 2010</collectingDate>
(fr.), 
<emphasis id="F63DC8A0F49BE551F56FCEC2B46106B6" italics="true" pageId="0" pageNumber="99">
<collectorName id="D650714DF9A1E8E4277292F2AE5257F4">Julius</collectorName>
et al. 
<accessionNumber id="1675B734FBEED16716BD0ECD0257DCE9" httpUri="http://www.ncbi.nlm.nih.gov/protein/FRI64006">FRI64006</accessionNumber>
</emphasis>
(KEP!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="AD3A60FFBFF682691E144A032070E8F4" pageId="0" pageNumber="99" type="notes">
<paragraph id="A1748CF5B93FDF86606260A69D752418" pageId="0" pageNumber="99">Notes.</paragraph>
<paragraph id="C0C15B510B09D39D9DE0BDB5D3217D47" pageId="0" pageNumber="99">
The comparatively large flowers, with sepals up to 1.5 cm long, place 
<taxonomicName id="D0A78D9F11AC494F1A687DE6D2CBF309" authorityName="Julius" authorityYear="2022" class="Magnoliopsida" family="Capparaceae" genus="Capparis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Capparis trinervia subsp. var. var. chungiana" order="Brassicales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="variety" species="trinervia" subSpecies="var." variety="chungiana">Capparis trinervia var. chungiana</taxonomicName>
(Figs 
<figureCitation id="D758ECB6F6E32112E54478A325798524" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Capparis trinervia var. chungiana Julius, var. nov. A the leafy branch B the flowering (bud) leafy twig C flower with sepals and petals removed exhibiting the filaments and ovary on gynophore D upper pedicel and base of calyx E the fruit in cross section F stipules close-up. (Drawn by Mohamad Aidil Noordin from FRI 64006 [A, E, F] and Mahmud Sidek s. n. [B-D])." figureDoi="10.3897/phytokeys.189.49367.figure7" httpUri="https://binary.pensoft.net/fig/642014" pageId="0" pageNumber="99">7</figureCitation>
, 
<figureCitation id="8726788B8760CB8AC5D1A3621D74A250" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Capparis trinervia var. chungiana Julius, var. nov. A fruiting twig B stipular thorns C young fruit on stout gynophore D leaf E young fruit, cross section. (Photographs by K. Imin)." figureDoi="10.3897/phytokeys.189.49367.figure8" httpUri="https://binary.pensoft.net/fig/642015" pageId="0" pageNumber="99">8</figureCitation>
) in the Trinervia-Group (
<bibRefCitation id="4DF599B4E93109082868CBF12A6E7892" author="Jacobs, M" journalOrPublisher="Blumea" pageId="0" pageNumber="99" pagination="385 - 541" refId="B21" refString="Jacobs, M, 1965. The genus Capparis (Capparaceae) from the Indus to the Pacific. Blumea 12 (3): 385 - 541" title="The genus Capparis (Capparaceae) from the Indus to the Pacific." volume="12" year="1965">Jacobs 1965</bibRefCitation>
; 
<bibRefCitation id="0882A16AC4BE1EAB853F28D3BF24E5FF" author="Srisanga, P" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="99" refId="B28" refString="Srisanga, P, Chayamarit, K, 2004. Capparis trisonthiae (Capparaceae), a new species from Thailand. Adansonia Series 3 26(1): 63-66." title="Capparis trisonthiae (Capparaceae), a new species from Thailand. Adansonia Series 3 26 (1): 63 - 66." year="2004">Srisanga and Chayamarit 2004</bibRefCitation>
). Specimens of this new species were determined as 
<taxonomicName id="736CE4DC17BA7D53F9CDAEA3652B9BA9" class="Magnoliopsida" family="Capparaceae" kingdom="Plantae" lsidName="C. erycibe" order="Brassicales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="erycibe">
<emphasis id="620F6646940BDEF23C02611F1FCC116E" italics="true" pageId="0" pageNumber="99">C. erycibe</emphasis>
</taxonomicName>
, no doubt due to the comparatively large leaves, but it differs in having the flowers arranged racemosely and becoming crowded at the distal part of the inflorescence rather than the paniculate arrangement in 
<taxonomicName id="9CC0810B6FD4B504C7E8E6BD08A542CD" class="Magnoliopsida" family="Capparaceae" kingdom="Plantae" lsidName="C. erycibe" order="Brassicales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="erycibe">
<emphasis id="4868CE762553B4576472BB4B741EAB9D" italics="true" pageId="0" pageNumber="99">C. erycibe</emphasis>
</taxonomicName>
. In addition, 
<taxonomicName id="86B7A863E0EA42F7632B4634810BE479" class="Magnoliopsida" family="Capparaceae" kingdom="Plantae" lsidName="C. erycibe" order="Brassicales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="erycibe">
<emphasis id="81E3AD8A569EA093D8BC9EF29385822B" italics="true" pageId="0" pageNumber="99">C. erycibe</emphasis>
</taxonomicName>
has smaller flowers with sepals only 4-6 mm long and was placed within the Subumbellates-group (
<bibRefCitation id="88073986E70A057AB8AEF660E18D875D" author="Jacobs, M" journalOrPublisher="Blumea" pageId="0" pageNumber="99" pagination="385 - 541" refId="B21" refString="Jacobs, M, 1965. The genus Capparis (Capparaceae) from the Indus to the Pacific. Blumea 12 (3): 385 - 541" title="The genus Capparis (Capparaceae) from the Indus to the Pacific." volume="12" year="1965">Jacobs 1965</bibRefCitation>
: 411).
</paragraph>
<caption id="DFAC4DAF1EA2CD4925ABD29B8A985BA2" doi="10.3897/phytokeys.189.49367.figure7" httpUri="https://binary.pensoft.net/fig/642014" pageId="0" pageNumber="99" start="Figure 7" startId="F7">
<paragraph id="2FF275E01F4FDC35D913D021DEE177CE" pageId="0" pageNumber="99">
<emphasis id="F5177673E93A4E5391A803B14E709893" bold="true" pageId="0" pageNumber="99">Figure 7.</emphasis>
<taxonomicName id="CAD656BFDFB143F5D34EBD885515FD7D" authorityName="Julius" authorityYear="2022" class="Magnoliopsida" family="Capparaceae" genus="Capparis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Capparis trinervia subsp. var. var. chungiana" order="Brassicales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="variety" species="trinervia" subSpecies="var." variety="chungiana">Capparis trinervia var. chungiana</taxonomicName>
Julius, var. nov. 
<emphasis id="471B894FBBAB74289AEE76025D4D81B2" bold="true" pageId="0" pageNumber="99">A</emphasis>
the leafy branch 
<emphasis id="0AE49B311664F9AB4B4353FF487702EA" bold="true" pageId="0" pageNumber="99">B</emphasis>
the flowering (bud) leafy twig 
<emphasis id="2D9B445E05C99B05048006DB254D1ED1" bold="true" pageId="0" pageNumber="99">C</emphasis>
flower with sepals and petals removed exhibiting the filaments and ovary on gynophore 
<emphasis id="B8741BED278E4B647139E9145D23222D" bold="true" pageId="0" pageNumber="99">D</emphasis>
upper pedicel and base of calyx 
<emphasis id="34C3AAB233B121F6937BC1B324BFF8B3" bold="true" pageId="0" pageNumber="99">E</emphasis>
the fruit in cross section 
<emphasis id="6407A57F6F8188332FBD15E1A7167F44" bold="true" pageId="0" pageNumber="99">F</emphasis>
stipules close-up. (Drawn by Mohamad Aidil Noordin from 
<emphasis id="88F07FA0AF1122DD5CE923FF238344E8" italics="true" pageId="0" pageNumber="99">FRI64006</emphasis>
[
<emphasis id="F99FAEE265B4F90F02CFEC22C60AD976" bold="true" pageId="0" pageNumber="99">A, E, F</emphasis>
] and Mahmud Sidek s.n. [
<emphasis id="C78719B01428ADF228867D227298EE33" bold="true" pageId="0" pageNumber="99">B-D</emphasis>
]).
</paragraph>
</caption>
<caption id="DA1D9784CA41B9CDE91CE3D494680693" doi="10.3897/phytokeys.189.49367.figure8" httpUri="https://binary.pensoft.net/fig/642015" pageId="0" pageNumber="99" start="Figure 8" startId="F8">
<paragraph id="A5A9EF0FB989DAFED49BB8A7365219D0" pageId="0" pageNumber="99">
<emphasis id="F40D8528967B55EFD5CBA58E6DFA2E96" bold="true" pageId="0" pageNumber="99">Figure 8.</emphasis>
<taxonomicName id="33AB687DF7D9704933178671D6FF0F60" authorityName="Julius" authorityYear="2022" class="Magnoliopsida" family="Capparaceae" genus="Capparis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Capparis trinervia subsp. var. var. chungiana" order="Brassicales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="variety" species="trinervia" subSpecies="var." variety="chungiana">Capparis trinervia var. chungiana</taxonomicName>
Julius, var. nov. 
<emphasis id="26F83D3E9040F483124F1ACFCE382D22" bold="true" pageId="0" pageNumber="99">A</emphasis>
fruiting twig 
<emphasis id="B5D873DE32FB009EDC4005DEDD5948B9" bold="true" pageId="0" pageNumber="99">B</emphasis>
stipular thorns 
<emphasis id="52F5A9D5CCFDE2A6A45563F01FC8B7E9" bold="true" pageId="0" pageNumber="99">C</emphasis>
young fruit on stout gynophore 
<emphasis id="26EB1B292FAD5A7FC2A145D7EAD3482D" bold="true" pageId="0" pageNumber="99">D</emphasis>
leaf 
<emphasis id="03D949A4CE37B55D47F1DB67B3B92611" bold="true" pageId="0" pageNumber="99">E</emphasis>
young fruit, cross section. (Photographs by K. Imin).
</paragraph>
</caption>
<paragraph id="C31A64A72599DBE59C991CB3C4FFD09A" pageId="0" pageNumber="99">
The flower colour of this species has been described as 
<normalizedToken id="BEC01A010D847BEBA2051AA725B73059" originalValue="‘white’">'white'</normalizedToken>
or 
<normalizedToken id="524C513678A7629DF03BCB92DD7A1974" originalValue="‘cream’">'cream'</normalizedToken>
with pink or dark purple at base in the label notes. Unfortunately, the open flowers are missing from the specimen sheets. Additionally, there are no new flowering specimens from the recent collections and full details of the flowers, such as measurement of the mature flowers and bracts, remain incomplete. Moreover, the dissected parts of the rehydrated flower buds are in poor condition and mostly do not contain either androecium or gynoecium, the cause of which is unknown.
</paragraph>
<paragraph id="715CD1340DD83E04C4C2395F0A099411" pageId="0" pageNumber="99">
However, this new variety still can be distinguished from other closely related members of the Trinervia-Group (see Table 
<tableCitation id="996B7EA76706C50AEFD079178DB86446" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphological comparison of Capparis trinervia var. chungiana, C. trinervia var. trinervia and C. trisonthiae." httpUri="http://table.plazi.org/id/2C04E2908BF7FAAF9A0C623711577D4D" pageId="0" pageNumber="99" tableUuid="2C04E2908BF7FAAF9A0C623711577D4D">2</tableCitation>
for the comparison details) or species known from Peninsular Malaysia vegetatively. For example, the relatively large fruit (ca. 11 cm in diam.) of 
<taxonomicName id="DA83FBEF6B29ADED58B3B3A44349FBF4" authorityName="Julius" authorityYear="2022" class="Magnoliopsida" family="Capparaceae" genus="Capparis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Capparis trinervia subsp. var. var. chungiana" order="Brassicales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="variety" species="trinervia" subSpecies="var." variety="chungiana">Capparis trinervia var. chungiana</taxonomicName>
is similar to 
<taxonomicName id="14744A3A37CBCCFB391AE47511E2F1AA" class="Magnoliopsida" family="Capparaceae" kingdom="Plantae" lsidName="C. scortechinii" order="Brassicales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="scortechinii">
<emphasis id="6A196143D61C510F35DD7BA84FFA8054" italics="true" pageId="0" pageNumber="99">C. scortechinii</emphasis>
</taxonomicName>
(ca. 11 cm in diam.) and 
<taxonomicName id="619997249D8E325B1A4D8AE4FDE853BD" class="Magnoliopsida" family="Capparaceae" kingdom="Plantae" lsidName="C. trisonthiae" order="Brassicales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="trisonthiae">
<emphasis id="F916CFD6B2AFA8204FEE0DEA7CD20359" italics="true" pageId="0" pageNumber="99">C. trisonthiae</emphasis>
</taxonomicName>
from Thailand (6-8 cm in diam.), but the latter two differ in their densely racemose and paniculate inflorescences, respectively; and the flowers arranged racemosely and becoming crowded at the distal part of the inflorescence, though has similarity to typical var. 
<taxonomicName id="9822B272FDA57921E26E1981B25F7757" class="Magnoliopsida" family="Capparaceae" genus="Capparis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Capparis trisonthiae var. trinervia" order="Brassicales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="variety" species="trisonthiae" variety="trinervia">Capparis trisonthiae trinervia</taxonomicName>
, but var. 
<taxonomicName id="0AFF9AEC532C9072877E59F5DD099EF8" class="Magnoliopsida" family="Capparaceae" genus="Capparis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Capparis trisonthiae var. chungiana" order="Brassicales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="variety" species="trisonthiae" variety="chungiana">Capparis trisonthiae chungiana</taxonomicName>
differs by its subcoriaceous to coriaceous leaves with broadly ovate to ovate-elliptic lamina (compared to chartaceous with oblong-elliptic or broadly lanceolate lamina in var. 
<taxonomicName id="763DE0CE8D825DFD52A692D63037551B" class="Magnoliopsida" family="Capparaceae" genus="Capparis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Capparis trisonthiae var. trinervia" order="Brassicales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="variety" species="trisonthiae" variety="trinervia">Capparis trisonthiae trinervia</taxonomicName>
), the drying leaves are reddish-brown, rarely pale green with venation pale yellow, rarely dark red on both surfaces (compared to dull green with brownish main nerves on both surfaces in var. 
<taxonomicName id="4D6CF8A26B72A6923025D1A372F2B9C5" class="Magnoliopsida" family="Capparaceae" genus="Capparis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Capparis trisonthiae var. trinervia" order="Brassicales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="variety" species="trisonthiae" variety="trinervia">Capparis trisonthiae trinervia</taxonomicName>
), the intercostal veins are obscure (compared to irregular reticulate and distinct in var. 
<taxonomicName id="424E38399FDAEA3F5544ECB945856424" class="Magnoliopsida" family="Capparaceae" genus="Capparis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Capparis trisonthiae var. trinervia" order="Brassicales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="variety" species="trisonthiae" variety="trinervia">Capparis trisonthiae trinervia</taxonomicName>
) and the stamens are 30-40 (whereas in var. 
<taxonomicName id="B50787F8CA2BFFAE41A7BF3BE0241982" class="Magnoliopsida" family="Capparaceae" genus="Capparis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Capparis trisonthiae var. trinervia" order="Brassicales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="variety" species="trisonthiae" variety="trinervia">Capparis trisonthiae trinervia</taxonomicName>
, (30-)60-70).
</paragraph>
<caption id="2C04E2908BF7FAAF9A0C623711577D4D" ID-Table-UUID="2C04E2908BF7FAAF9A0C623711577D4D" httpUri="http://table.plazi.org/id/2C04E2908BF7FAAF9A0C623711577D4D" pageId="0" pageNumber="99" start="Table 2" startId="T2">
<paragraph id="2552811DD7EA647CD1A2A1FACA888944" pageId="0" pageNumber="99">
<emphasis id="EF8209121DA6C1B8CE12E424483A4C10" bold="true" pageId="0" pageNumber="99">Table 2.</emphasis>
Morphological comparison of 
<taxonomicName id="13BBD0A9F3CC0941C245416B422EB29A" authorityName="Julius" authorityYear="2022" class="Magnoliopsida" family="Capparaceae" genus="Capparis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Capparis trinervia subsp. var. var. chungiana" order="Brassicales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="variety" species="trinervia" subSpecies="var." variety="chungiana">Capparis trinervia var. chungiana</taxonomicName>
, 
<taxonomicName id="49A7F9A902442A8101FDF06057F872DC" authority="var. trinervia" class="Magnoliopsida" family="Capparaceae" kingdom="Plantae" lsidName="C. trinervia subsp. var. var. trinervia" order="Brassicales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="variety" species="trinervia" subSpecies="var." variety="trinervia">C. trinervia var. trinervia</taxonomicName>
and 
<taxonomicName id="9AABDED0765186B2DB9323F7B436BFF2" class="Magnoliopsida" family="Capparaceae" kingdom="Plantae" lsidName="C. trisonthiae" order="Brassicales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="trisonthiae">
<emphasis id="BA8B28975131C80C4F09A63E15580768" italics="true" pageId="0" pageNumber="99">C. trisonthiae</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="46FC66E298E610B2DD0B05B804B198D6" pageId="0" pageNumber="99">
<table id="ABA5B5A1B5384D8A3A04F56306087F91" pageId="0" pageNumber="99">
<tr id="D48AEE65634DDC4EA4B91330FC3B4068" pageId="0" pageNumber="99">
<th id="943EBCA3D6CBF5668F78D51641C85F71" colspan="1" pageId="0" pageNumber="99" rowspan="2">Characters</th>
<th id="ED2CE2E89C75B203720306AB3F928C37" colspan="3" pageId="0" pageNumber="99" rowspan="1">Species</th>
</tr>
<tr id="A5A22B7A80E001928E00989E40398947" pageId="0" pageNumber="99">
<th id="4BFE56E2B04FA8BA2380AE2FCAE0182E" colspan="1" pageId="0" pageNumber="99" rowspan="1">
<taxonomicName id="3E0617E09EC2444480D14BAF0FE26888" class="Magnoliopsida" family="Capparaceae" kingdom="Plantae" lsidName="C. trinervia subsp. var. var. chungiana" order="Brassicales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="variety" species="trinervia" subSpecies="var." variety="chungiana">C. trinervia var. chungiana</taxonomicName>
</th>
<th id="5B3B3F15F9BEE9822411653314966CF2" colspan="1" pageId="0" pageNumber="99" rowspan="1">
<taxonomicName id="65E14DE6F5CE89E0294E7E4F8B2DC054" authority="var. trinervia" class="Magnoliopsida" family="Capparaceae" kingdom="Plantae" lsidName="C. trinervia subsp. var. var. trinervia" order="Brassicales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="variety" species="trinervia" subSpecies="var." variety="trinervia">C. trinervia var. trinervia</taxonomicName>
</th>
<th id="5E513056C873558BDE63AACFE20B18E7" colspan="1" pageId="0" pageNumber="99" rowspan="1">
<taxonomicName id="A6F81995417A3C47360E0B0AAB1F0FA5" class="Magnoliopsida" family="Capparaceae" kingdom="Plantae" lsidName="C. trisonthiae" order="Brassicales" pageId="0" pageNumber="99" phylum="Tracheophyta" rank="species" species="trisonthiae">
<emphasis id="72AC41791C1558868590BA32D925F9EF" italics="true" pageId="0" pageNumber="99">C. trisonthiae</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="1A7E30ABD10149CA8A6E937FC8BF3788" pageId="0" pageNumber="99">
<td id="2AA70473BD98F716BCB4803E747E4B71" colspan="1" pageId="0" pageNumber="99" rowspan="1">Stipular thorns</td>
<td id="B8FF5B0616E3765E4A5D4F6B9D242704" colspan="1" pageId="0" pageNumber="99" rowspan="1">1.5-3 mm long, recurved downwards</td>
<td id="A98D9C47171ACA8F67ABC75029C126B6" colspan="1" pageId="0" pageNumber="99" rowspan="1">1-3(-6) mm long, patent or recurved, mostly downwards, rarely upwards</td>
<td id="6AA9F9CBE02AB5C532CDCEF04A3DA385" colspan="1" pageId="0" pageNumber="99" rowspan="1">2-4 mm long, recurved downwards</td>
</tr>
<tr id="678F505E03823C119DD9F793F1B29DD1" pageId="0" pageNumber="99">
<td id="055B02C78BE1285FEF199B2E22AA5FBF" colspan="1" pageId="0" pageNumber="99" rowspan="1">Petiole</td>
<td id="704701E4AD7B4FAB4ABABA61F39B3AEB" colspan="1" pageId="0" pageNumber="99" rowspan="1">1.5-2 cm long, 2-3 mm in diam., stout</td>
<td id="8B81ED61C6D9F7C84930AFE4DC47825B" colspan="1" pageId="0" pageNumber="99" rowspan="1">0.7-1.8 cm long, ca. 1.5 mm in diam., slender</td>
<td id="A9E5978D6B515873F5E4E2A08EE846F4" colspan="1" pageId="0" pageNumber="99" rowspan="1">ca. 2 cm long, ca. 4 mm in diam., stout</td>
</tr>
<tr id="526D3E24C608A665B1F7DAC75A3829E4" pageId="0" pageNumber="99">
<td id="372E7910C2C379A35F3181B2D3CDCDC0" colspan="1" pageId="0" pageNumber="99" rowspan="1">Leaf shape</td>
<td id="B231376E100CA94BE770DF4F545A0C4D" colspan="1" pageId="0" pageNumber="99" rowspan="1">Broadly ovate to ovate-elliptic</td>
<td id="8F79225B3C71E889AD225B60A1C21C33" colspan="1" pageId="0" pageNumber="99" rowspan="1">Oblong-elliptic or broadly lanceolate</td>
<td id="A805645FE6CC8980B1A263F2FD173F07" colspan="1" pageId="0" pageNumber="99" rowspan="1">Broadly ovate</td>
</tr>
<tr id="D5B1237337F5CC4F468C5DABACEE6580" pageId="0" pageNumber="99">
<td id="1429F2F6A2400982EF4BEDE006A374AD" colspan="1" pageId="0" pageNumber="99" rowspan="1">Leaf size</td>
<td id="37622211319ED88FB08D01A1EB737C86" colspan="1" pageId="0" pageNumber="99" rowspan="1">
13-16 
<normalizedToken id="E1038EB7BB138F1BCC53805F4E1E3E65" originalValue="×">x</normalizedToken>
5.5-8.5 cm
</td>
<td id="7C5112628D5D0C2F6B517020A8943D44" colspan="1" pageId="0" pageNumber="99" rowspan="1">
(5-)10-14(-19) 
<normalizedToken id="BA3779DF50604E88523FF498AC81246F" originalValue="×">x</normalizedToken>
(2-)3.5-8.5 cm
</td>
<td id="5CC1B1E92AC9FFC209BEBA192B9ACE33" colspan="1" pageId="0" pageNumber="99" rowspan="1">
(10-)12-18 
<normalizedToken id="71414FDB22AB60CE874477E1A1DF27DE" originalValue="×">x</normalizedToken>
(6.5-)8-11 cm
</td>
</tr>
<tr id="7B91CA8B162939FA33F9333286E7160F" pageId="0" pageNumber="99">
<td id="32B918AF25E9DA83FBA9D56BD3E90A05" colspan="1" pageId="0" pageNumber="99" rowspan="1">Leaf apex</td>
<td id="48D001C861E8A1746CDF1A1C9B6355F7" colspan="1" pageId="0" pageNumber="99" rowspan="1">Acute or shortly cuspidate</td>
<td id="980452C9E72BA0A13B8A77190E16259A" colspan="1" pageId="0" pageNumber="99" rowspan="1">Obtuse to shortly acuminate</td>
<td id="FCBBA5D4967CD030D4BA89F082644387" colspan="1" pageId="0" pageNumber="99" rowspan="1">Retuse to emarginate</td>
</tr>
<tr id="B641EDB76ABCD109F34E8E6BDDE7A85A" pageId="0" pageNumber="99">
<td id="80442CEA69D920F177DA78BDCC04E17A" colspan="1" pageId="0" pageNumber="99" rowspan="1">Leaf hairs</td>
<td id="100B8D31C1BEE580ABD5EF02E4EE6C2A" colspan="1" pageId="0" pageNumber="99" rowspan="1">Glabrous on both surfaces</td>
<td id="4C97E41CC8AD3C31486E690232B13EE3" colspan="1" pageId="0" pageNumber="99" rowspan="1">Glabrescent</td>
<td id="8D202FF546D5AF3A917FF05CD42D4040" colspan="1" pageId="0" pageNumber="99" rowspan="1">Glabrous on both surfaces</td>
</tr>
<tr id="65A5BFE3A51D07FF95935687D81F57DF" pageId="0" pageNumber="99">
<td id="0D7E894F5B6039392BF4394ECC9C1B02" colspan="1" pageId="0" pageNumber="99" rowspan="1">Leaf colour when dried</td>
<td id="3822A3D73EB1842C9A20D9B5E9A1D3E5" colspan="1" pageId="0" pageNumber="99" rowspan="1">Reddish-brown, rarely pale green with pale rarely dark red venation on both surfaces</td>
<td id="4C8F044D1E504D88E2399C722F96C7FC" colspan="1" pageId="0" pageNumber="99" rowspan="1">Dull green with brownish venation on both surfaces</td>
<td id="441256A274624FCD9C37065B4CEA2EA4" colspan="1" pageId="0" pageNumber="99" rowspan="1">Dull green above, brown beneath</td>
</tr>
<tr id="1D5245AECDAAFA49E4002A86A0273EEB" pageId="0" pageNumber="99">
<td id="BCD8972CF1C3D513569ECAE93627187A" colspan="1" pageId="0" pageNumber="99" rowspan="1">Intercostal veins</td>
<td id="488B4180C3A4B3935145DFC4363E10D8" colspan="1" pageId="0" pageNumber="99" rowspan="1">Obscure on both surfaces</td>
<td id="87C407EF485D5875A4D9D520C88B1B77" colspan="1" pageId="0" pageNumber="99" rowspan="1">Irregular reticulate, distinct on both surfaces</td>
<td id="95F939794070EF28B928912924C6984E" colspan="1" pageId="0" pageNumber="99" rowspan="1">Irregular reticulate, prominent on both surfaces</td>
</tr>
<tr id="422BE58646226831AB0841352233D1FA" pageId="0" pageNumber="99">
<td id="C98938F971A55700490DFEB5539FD99A" colspan="1" pageId="0" pageNumber="99" rowspan="1">Inflorescences</td>
<td id="37FEBD731A0B6919F28A986A74783BE5" colspan="1" pageId="0" pageNumber="99" rowspan="1">Racemes with flowers becoming crowded at the distal part of the inflorescence</td>
<td id="9D95E3CC684AEFA345032F99AE7F402D" colspan="1" pageId="0" pageNumber="99" rowspan="1">Racemes with flowers becoming crowded at the distal part of the inflorescence or subumbels</td>
<td id="73EC9A1317D63DC43462E416099AFF5A" colspan="1" pageId="0" pageNumber="99" rowspan="1">A leafy paniculate</td>
</tr>
<tr id="0324AC97EE8ACF5BA26B7D6029B6A601" pageId="0" pageNumber="99">
<td id="520058C92BED44FDCA65DA2858DBA21B" colspan="1" pageId="0" pageNumber="99" rowspan="1">Sepals shape</td>
<td id="312BC7FD6E0C4338DDE4F304BDD05221" colspan="1" pageId="0" pageNumber="99" rowspan="1">Orbicular</td>
<td id="3DC81F5A506F2C9A0026681789F4086E" colspan="1" pageId="0" pageNumber="99" rowspan="1">Orbicular</td>
<td id="4BD5717757EB7EEB657DE4FFDB27DB3D" colspan="1" pageId="0" pageNumber="99" rowspan="1">Lanceolate</td>
</tr>
<tr id="84FC25829455559CBCA9214C08F3C6A1" pageId="0" pageNumber="99">
<td id="ACD156E2BF52346E3028E74C164A1B51" colspan="1" pageId="0" pageNumber="99" rowspan="1">Sepals size</td>
<td id="67AF47B985CBC6A83DD855FB0B7E121E" colspan="1" pageId="0" pageNumber="99" rowspan="1">1.2-1.5 cm long</td>
<td id="159F3FBC17C4B08B5B7486590E59A169" colspan="1" pageId="0" pageNumber="99" rowspan="1">0.9-1.5 cm long</td>
<td id="CF011830109EE9CCDEC2D77428765F0F" colspan="1" pageId="0" pageNumber="99" rowspan="1">1.8-2.5 cm long</td>
</tr>
<tr id="54E6451D47FC592EE88967F4CE364E6D" pageId="0" pageNumber="99">
<td id="535062FA21D692A52196AEE76300F79F" colspan="1" pageId="0" pageNumber="99" rowspan="1">Sepals hairs</td>
<td id="27C72F5B8279AA8CF16AB92629A00A09" colspan="1" pageId="0" pageNumber="99" rowspan="1">Densely velvety ferruginous hairs abaxially, glabrous adaxially</td>
<td id="7AF55A73A17209CE76644B9749F319C8" colspan="1" pageId="0" pageNumber="99" rowspan="1">Densely orange-yellow puberulous abaxially, glabrous adaxially</td>
<td id="603ECE6994AF1021EF740E08D736BCA8" colspan="1" pageId="0" pageNumber="99" rowspan="1">Ferruginous puberulous abaxially, puberulous adaxially in the marginal parts</td>
</tr>
<tr id="5A82E0D5934A1AA577A9ED8C0CA968B4" pageId="0" pageNumber="99">
<td id="997F61E69DD43684EC3018824856FB7F" colspan="1" pageId="0" pageNumber="99" rowspan="1">Stamens</td>
<td id="FF81A401700E606C959551ACA633BEB8" colspan="1" pageId="0" pageNumber="99" rowspan="1">30-40</td>
<td id="035965E05E5CB306B556DB097FE913B1" colspan="1" pageId="0" pageNumber="99" rowspan="1">(30-)60-70</td>
<td id="4EBD007E07744FB1691BE4527FF6ACDF" colspan="1" pageId="0" pageNumber="99" rowspan="1">140-170</td>
</tr>
<tr id="0D9E5951910C1E316D253C4C69066185" pageId="0" pageNumber="99">
<td id="3F08E0AD7C80B91627C4FF552CA03509" colspan="1" pageId="0" pageNumber="99" rowspan="1">Fruit</td>
<td id="FC915347FEBFFE0E0FF120D56E8A20AE" colspan="1" pageId="0" pageNumber="99" rowspan="1">Globose, ca. 11 cm in diam.</td>
<td id="99890D2E2CF5C263F145F8811C928BA1" colspan="1" pageId="0" pageNumber="99" rowspan="1">Globose, 3.5-5 cm in diam.</td>
<td id="AF517BC33DC21D4EC8016F9A548622AB" colspan="1" pageId="0" pageNumber="99" rowspan="1">Ellipsoid, 6-8 cm in diam.</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>