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<document id="0DAF5167431E73F569408CE9801791F4" ID="10.11646/zootaxa.4132.4.12" ID-DOI="10.11646/zootaxa.4132.4.12" ID-GBIF-Dataset="4ac2937d-dcf8-46d7-be2d-33bba5e6bc9f" ID-ISSN="1175-5326" ID-Zenodo-Dep="271906" ID-ZooBank="CB0F3D90-DCE6-445C-9163-C450F9FEEF7F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1467372964119" checkinUser="plazi" docAuthor="Pivar, Robert J., Moulton, John K. &amp; Sinclair, Bradley J." docDate="2016" docId="2322594EFFD5C637E5CBF2982DE8F94C" docLanguage="en" docName="zootaxa.4132.4.12.pdf" docOrigin="Zootaxa 4132 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Austrothaumalea spiculata Pivar, sp. nov." docType="treatment" docVersion="6" lastPageNumber="595" masterDocId="DF1B2136FFD4C636E55CF20F2873FFF8" masterDocTitle="A new species of Austrothaumalea Tonnoir from Australia (Diptera: Thaumaleidae)" masterLastPageNumber="597" masterPageNumber="594" pageNumber="595" updateTime="1698363117957" updateUser="plazi">
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<mods:title id="52076A23889FDC06F594F1D05D5E2E27">A new species of Austrothaumalea Tonnoir from Australia (Diptera: Thaumaleidae)</mods:title>
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<mods:namePart id="EE2DE718EBB2F310BEB3CCFF40DFE264">Pivar, Robert J.</mods:namePart>
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<mods:namePart id="550B38BD33FC6374153A2B9B0D28AE00">Moulton, John K.</mods:namePart>
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<mods:name id="EA123BD9467FFE4A344CB275325D46E6" type="personal">
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<mods:namePart id="2C37913B790922EC473CFC9579129D98">Sinclair, Bradley J.</mods:namePart>
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<mods:typeOfResource id="1460E72FF073CF6896E2BC196F6D5211">text</mods:typeOfResource>
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<mods:title id="0841614342C6BEFB2358B45DB9A5D099">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="03715CE5A4151C8ACF8CD554DD25F89E">
<mods:date id="D0CEE7211575BE75BA18769C8438129B">2016</mods:date>
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<mods:number id="043473066E8FDE2D4DCAE4EF6DC38CA5">4132</mods:number>
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<mods:number id="4E245F27316E9EAD19DFAE518A373ACC">4</mods:number>
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<mods:classification id="50BAB122D9AFCDB0E53F8171E6EB8B04">journal article</mods:classification>
<mods:identifier id="C49194DB8177B6E5A0FBB518F387265F" type="DOI">10.11646/zootaxa.4132.4.12</mods:identifier>
<mods:identifier id="582B0C97061AC3DBDC6CB3908E5DD8AA" type="GBIF-Dataset">4ac2937d-dcf8-46d7-be2d-33bba5e6bc9f</mods:identifier>
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<treatment id="2322594EFFD5C637E5CBF2982DE8F94C" ID-DOI="http://doi.org/10.5281/zenodo.6062778" ID-GBIF-Taxon="121268820" ID-Zenodo-Dep="6062778" LSID="urn:lsid:plazi:treatment:2322594EFFD5C637E5CBF2982DE8F94C" httpUri="http://treatment.plazi.org/id/2322594EFFD5C637E5CBF2982DE8F94C" lastPageNumber="595" pageId="1" pageNumber="595">
<subSubSection id="E391BBD3FFD5C637E5CBF2982AFEFF4A" box="[151,653,151,178]" pageId="1" pageNumber="595" type="nomenclature">
<paragraph id="AB34E858FFD5C637E5CBF2982AFEFF4A" blockId="1.[151,1436,151,1433]" box="[151,653,151,178]" pageId="1" pageNumber="595">
<heading id="F07C5F34FFD5C637E5CBF2982AFEFF4A" bold="true" box="[151,653,151,178]" fontSize="11" level="1" pageId="1" pageNumber="595" reason="1">
<emphasis id="99FF344AFFD5C637E5CBF2982AFEFF4A" bold="true" box="[151,653,151,178]" pageId="1" pageNumber="595">
<taxonomicName id="6C8B93DBFFD5C637E5CBF2982A5AFF4A" authority="Pivar" authorityName="Pivar" box="[151,553,151,178]" class="Insecta" family="Thaumaleidae" genus="Austrothaumalea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="595" phylum="Arthropoda" rank="species" species="spiculata" status="sp. nov.">
<emphasis id="99FF344AFFD5C637E5CBF29829AEFF49" bold="true" box="[151,477,151,177]" italics="true" pageId="1" pageNumber="595">Austrothaumalea spiculata</emphasis>
Pivar
</taxonomicName>
<taxonomicNameLabel id="82CC8931FFD5C637E76CF2972AFEFF4A" box="[560,653,152,178]" pageId="1" pageNumber="595" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E391BBD3FFD5C637E5CBF2B52BF2FE59" pageId="1" pageNumber="595" type="materials_examined">
<paragraph id="AB34E858FFD5C637E5CBF2B52A67FEA4" blockId="1.[151,1436,151,1433]" pageId="1" pageNumber="595">
<emphasis id="99FF344AFFD5C637E5CBF2B52930FF29" bold="true" box="[151,323,186,209]" pageId="1" pageNumber="595">
<typeStatus id="743056FAFFD5C637E5CBF2B528BCFF29" box="[151,207,186,209]" pageId="1" pageNumber="595">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="743056FAFFD5C637E401F2B52987FF29" box="[349,500,186,209]" pageId="1" pageNumber="595" type="holotype">
<emphasis id="99FF344AFFD5C637E401F2B52987FF29" bold="true" box="[349,500,186,209]" pageId="1" pageNumber="595">HOLOTYPE</emphasis>
</typeStatus>
♂, labelled: “AUS: Victoria: Grampians N.P./ MacKenzie Falls,
<date id="DF35CE98FFD5C637E07EF2B52DE0FF29" box="[1314,1427,186,209]" pageId="1" pageNumber="595" value="2014-09-14">14.ix.2014</date>
/ S37°06ʹ37.05″ E142°24ʹ31.39″/ ex. side seep along foot bridge/ to MacKenzie Falls, elev.
<quantity id="6C7345BDFFD5C637E1F8F2D32C93FF0C" box="[1188,1248,220,244]" metricMagnitude="2" metricUnit="m" metricValue="3.78" pageId="1" pageNumber="595" unit="m" value="378.0">378m</quantity>
/ J.K. Moulton”; “
<typeStatus id="743056FAFFD5C637E5FCF30F295CFEEF" box="[160,303,256,279]" pageId="1" pageNumber="595" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="6C8B93DBFFD5C637E462F30F2987FEEF" box="[318,500,256,279]" class="Insecta" family="Thaumaleidae" genus="Austrothaumalea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="595" phylum="Arthropoda" rank="genus">Austrothaumalea</taxonomicName>
/
<taxonomicName id="6C8B93DBFFD5C637E75EF30F2A13FEEF" box="[514,608,256,279]" class="Insecta" family="Thaumaleidae" genus="Austrothaumalea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="595" phylum="Arthropoda" rank="species" species="spiculata">spiculata</taxonomicName>
/ Pivar [red label]” (AMS).
<typeStatus id="743056FAFFD5C637E6D7F30F2C58FEEF" box="[907,1067,256,279]" pageId="1" pageNumber="595" type="paratype">
<emphasis id="99FF344AFFD5C637E6D7F30F2C58FEEF" bold="true" box="[907,1067,256,279]" pageId="1" pageNumber="595">PARATYPES</emphasis>
</typeStatus>
:
<specimenCount id="BD8D23D1FFD5C637E165F30F2C10FEEF" box="[1081,1123,256,279]" pageId="1" pageNumber="595" type="generic">2 ♂</specimenCount>
, same collection information as
<typeStatus id="743056FAFFD5C637E5E4F32D2966FEC1" box="[184,277,290,313]" pageId="1" pageNumber="595" type="holotype">holotype</typeStatus>
(
<collectionCode id="CD9A709DFFD5C637E47BF32D2911FEC1" box="[295,354,290,313]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="1" pageNumber="595">CNC</collectionCode>
,
<collectionCode id="CD9A709DFFD5C637E42DF32D29B0FEC1" box="[369,451,290,313]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="595">USNM</collectionCode>
). One additional ♂
<typeStatus id="743056FAFFD5C637E7FFF32D2A8CFEC1" box="[675,767,290,313]" pageId="1" pageNumber="595" type="paratype">paratype</typeStatus>
was used for molecular study and the voucher specimen is housed in the UTK Insect Museum.
</paragraph>
<paragraph id="AB34E858FFD5C637E59BF3672BF2FE59" blockId="1.[151,1436,151,1433]" pageId="1" pageNumber="595">
<emphasis id="99FF344AFFD5C637E59BF367293CFE87" bold="true" box="[199,335,360,383]" pageId="1" pageNumber="595">Recognition</emphasis>
.
<taxonomicName id="6C8B93DBFFD5C637E43CF3682AF7FE87" box="[352,644,359,383]" class="Insecta" family="Thaumaleidae" genus="Austrothaumalea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="595" phylum="Arthropoda" rank="species" species="spiculata">
<emphasis id="99FF344AFFD5C637E43CF3682AF7FE87" box="[352,644,359,383]" italics="true" pageId="1" pageNumber="595">Austrothaumalea spiculata</emphasis>
</taxonomicName>
is readily distinguished from all other species by its spearhead-shaped paramere, arched gonostyli and absence of apical epandrial processes.
</paragraph>
</subSubSection>
<subSubSection id="E391BBD3FFD5C637E59BF3A22BF7FB5E" pageId="1" pageNumber="595" type="description">
<paragraph id="AB34E858FFD5C637E59BF3A22A04FE3C" blockId="1.[151,1436,151,1433]" box="[199,631,429,452]" pageId="1" pageNumber="595">
<emphasis id="99FF344AFFD5C637E59BF3A229D6FE3C" bold="true" box="[199,421,429,452]" pageId="1" pageNumber="595">Description of male</emphasis>
(female unknown).
</paragraph>
<paragraph id="AB34E858FFD5C637E59BF3C02966FDB7" blockId="1.[151,1436,151,1433]" pageId="1" pageNumber="595">
<emphasis id="99FF344AFFD5C637E59BF3C02921FE1F" box="[199,338,463,487]" italics="true" pageId="1" pageNumber="595">Colouration:</emphasis>
Head dull, dark brown; pronotum dark brown; postpronotum pale brown; mesonotum and pleura brown and somewhat shiny; katepisternum reddish-brown; legs pale yellow, becoming darker on apical tarsal segments; scutellum and mediotergite pale brown, shiny; halter stalk cream colored, knob dark; abdomen dark brown; terminalia pale brown.
</paragraph>
<paragraph id="AB34E858FFD5C637E59BF05529F9FD1B" blockId="1.[151,1436,151,1433]" pageId="1" pageNumber="595">
<emphasis id="99FF344AFFD5C637E59BF055288EFD89" box="[199,253,602,625]" italics="true" pageId="1" pageNumber="595">Wing</emphasis>
(Fig. 1A): Wing length:
<quantity id="6C7345BDFFD5C637E75BF0552AF9FD89" box="[519,650,602,625]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" pageId="1" pageNumber="595" unit="mm" value="2.8" valueMax="2.9" valueMin="2.7">2.72.9 mm</quantity>
. Slightly more infuscate dorsal to vein M1, with darkening where R4+5 and M1 meet wing margin, and slightly basal to R2+3; R1(+R2+3) with macrotrichia along entire length, remaining veins bare; R 2+3 situated towards apex of R 1; bend in R 4+5 well defined; R 4+5 and M 1 running parallel toward wing margin; CuA with short basal appendage.
</paragraph>
<paragraph id="AB34E858FFD5C637E59BF0E12AFAFC69" blockId="1.[151,1436,151,1433]" pageId="1" pageNumber="595">
<emphasis id="99FF344AFFD5C637E59BF0E1295FFCFE" box="[199,300,750,774]" italics="true" pageId="1" pageNumber="595">Abdomen</emphasis>
(Fig. 1B): Abdominal sternite 1 narrow, spectacle-shaped; sternite 2 reduced to slender, median sclerite, with pair of small, posterolateral sclerites, setae restricted to apical fourth and between posterolateral sclerites; sternites 36 rectangular, anterior margin well sclerotized, with pair of small, posterolateral sclerites, sternite 3 with setae on posterior two-thirds, sternites 46 with setae on median third; sternite 7 trapezoidal-shaped, with posterior marginal setae; sternite 8 triangle-shaped, without setae.
</paragraph>
<paragraph id="AB34E858FFD5C637E59BF1932BF7FB5E" blockId="1.[151,1436,151,1433]" pageId="1" pageNumber="595">
<emphasis id="99FF344AFFD5C637E59BF193290BFC4C" box="[199,376,924,948]" italics="true" pageId="1" pageNumber="595">Male terminalia</emphasis>
(
<figureCitation id="33B0F4DDFFD5C637E4D7F19329A7FC4B" box="[395,468,924,947]" captionStart="FIGURE 2" captionStartId="2.[151,244,1871,1893]" captionTargetBox="[183,1404,1171,1846]" captionTargetId="figure@2.[176,1410,1165,1854]" captionTargetPageId="2" captionText="FIGURE 2. Male terminalia of Austrothaumalea spiculata sp. nov. in (A) ventral view and (B) lateral view. Scale bar = 0.1 mm. Abbreviations: cerc, cercus; epand, epandrium; gcx, gonocoxite; gcx pl, gonocoxal plate; gst, gonostylus; hypd, hypandrium; pm, paramere." httpUri="https://zenodo.org/record/271907/files/figure.png" pageId="1" pageNumber="595">Figs 2</figureCitation>
A, B): Epandrium broadly triangular from ventral aspect; apical third slightly tapered; posterior margin broad and rounded; lacking pointed posterolateral processes; light scar visible in lateral view when cleared. Hypandrium broad, nearly subequal to width of base of gonocoxites; anterior margin slightly concave. Gonocoxite broad, inner margin of basal two-thirds slightly serrated before becoming widest two-thirds of way down; apical third strongly tapered. Gonostylus nearly subequal in length to gonocoxite, curved strongly before mid-length; thickest at mid-point; gradually tapered to blunt tip. Parameres distally fused, half length of epandrium, rod-like with subapical lateral expansions creating spearhead-like apex. Gonocoxal plate broad basally; apical margin concave with short, pointed posterolateral process; basally not fused to hypandrium.
</paragraph>
</subSubSection>
<subSubSection id="E391BBD3FFD5C637E59BF6BD2904FAAC" pageId="1" pageNumber="595" type="distribution">
<paragraph id="AB34E858FFD5C637E59BF6BD2904FAAC" blockId="1.[151,1436,151,1433]" pageId="1" pageNumber="595">
<emphasis id="99FF344AFFD5C637E59BF6BD2921FB31" bold="true" box="[199,338,1202,1225]" pageId="1" pageNumber="595">Distribution</emphasis>
. Known only from the
<typeStatus id="743056FAFFD5C637E704F6BD2AF5FB31" box="[600,646,1202,1225]" pageId="1" pageNumber="595">type</typeStatus>
locality (Fig. 1C). Grampians National Park is located in the southwest region of Victoria, roughly
<quantity id="6C7345BDFFD5C637E494F6DA2A6FFB14" box="[456,540,1237,1260]" metricMagnitude="5" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="595" unit="km" value="260.0">260 km</quantity>
west of Melbourne. It consists of sandstone mountain ranges and eight different vegetation communities. Situated in the northern half of the park, Mackenzie Falls is Victorias highest waterfall.
<taxonomicName id="6C8B93DBFFD5C637E5CBF7162A1CFAC9" authority="Theischinger" authorityName="Theischinger" box="[151,623,1305,1329]" class="Insecta" family="Thaumaleidae" genus="Austrothaumalea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="595" phylum="Arthropoda" rank="species" species="denticulata">
<emphasis id="99FF344AFFD5C637E5CBF71629A6FAC9" box="[151,469,1305,1329]" italics="true" pageId="1" pageNumber="595">Austrothaumalea denticulata</emphasis>
Theischinger
</taxonomicName>
is the only other species of thaumaleid known from the Grampians (
<bibRefCitation id="CF1A95A9FFD5C637E5C2F7322918FAAC" author="Theischinger" box="[158,363,1341,1364]" pageId="1" pageNumber="595" refString="Theischinger, G. (1986) Australian Thaumaleidae. Records of the Australian Museum, 38, 291 - 317. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.38.1986.184" type="journal article" year="1986">Theischinger 1986</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E391BBD3FFD5C637E59BF7502A55FA0E" pageId="1" pageNumber="595" type="etymology">
<paragraph id="AB34E858FFD5C637E59BF7502AC3FA61" blockId="1.[151,1436,151,1433]" pageId="1" pageNumber="595">
<emphasis id="99FF344AFFD5C637E59BF7502933FA8E" bold="true" box="[199,320,1375,1398]" pageId="1" pageNumber="595">Etymology</emphasis>
.
<taxonomicName id="6C8B93DBFFD5C637E413F7512A03FA8E" box="[335,624,1374,1398]" class="Insecta" family="Thaumaleidae" genus="Austrothaumalea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="595" phylum="Arthropoda" rank="species" species="spiculata">
<emphasis id="99FF344AFFD5C637E413F7512A03FA8E" box="[335,624,1374,1398]" italics="true" pageId="1" pageNumber="595">Austrothaumalea spiculata</emphasis>
</taxonomicName>
is so named from the Latin,
<emphasis id="99FF344AFFD5C637E6F3F76F2B94FA8F" box="[943,999,1376,1399]" italics="true" pageId="1" pageNumber="595">spica</emphasis>
(point), in reference to the shape of the paramere, which resembles that of a Roman spear.
</paragraph>
<paragraph id="AB34E858FFD5C637E5CBF7D02A55FA0E" blockId="1.[151,1244,1503,1561]" box="[151,550,1503,1526]" pageId="1" pageNumber="595">
<emphasis id="99FF344AFFD5C637E5CBF7D029B8FA0E" bold="true" box="[151,459,1503,1526]" pageId="1" pageNumber="595">
Modified key from
<bibRefCitation id="CF1A95A9FFD5C637E42EF7D029B8FA0E" author="Sinclair" box="[370,459,1503,1526]" pageId="1" pageNumber="595" refString="Sinclair, B. J. (2008 b) New species of Austrothaumalea Tonnoir from Australia (Diptera: Thaumaleidae). Records of the Australian Museum, 60, 1, 1 - 12. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.60.2008.1490" type="journal article" year="2008" yearSuffix="b">Sinclair</bibRefCitation>
</emphasis>
(2008b)
</paragraph>
</subSubSection>
<subSubSection id="E391BBD3FFD5C637E5CBF40E2DE8F94C" pageId="1" pageNumber="595" type="description">
<paragraph id="AB34E858FFD5C637E5CBF40E2CAFF9E1" blockId="1.[151,1244,1503,1561]" box="[151,1244,1537,1561]" pageId="1" pageNumber="595">
<taxonomicName id="6C8B93DBFFD5C637E5CBF40E29C5F9E1" box="[151,438,1537,1561]" class="Insecta" family="Thaumaleidae" genus="Austrothaumalea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="595" phylum="Arthropoda" rank="species" species="spiculata">
<emphasis id="99FF344AFFD5C637E5CBF40E29C5F9E1" box="[151,438,1537,1561]" italics="true" pageId="1" pageNumber="595">Austrothaumalea spiculata</emphasis>
</taxonomicName>
keys to couplet
<quantity id="6C7345BDFFD5C637E738F40D2AE8F9E1" box="[612,667,1538,1561]" metricMagnitude="-1" metricUnit="m" metricValue="5.08" pageId="1" pageNumber="595" unit="in" value="20.0">20 in</quantity>
<bibRefCitation id="CF1A95A9FFD5C637E7FDF40D2B21F9E1" author="Sinclair" box="[673,850,1538,1561]" pageId="1" pageNumber="595" refString="Sinclair, B. J. (2008 b) New species of Austrothaumalea Tonnoir from Australia (Diptera: Thaumaleidae). Records of the Australian Museum, 60, 1, 1 - 12. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.60.2008.1490" type="journal article" year="2008" yearSuffix="b">Sinclair (2008b)</bibRefCitation>
. This couplet is modified as follows:
</paragraph>
<paragraph id="AB34E858FFD5C637E5CBF4482DE8F94C" blockId="1.[151,1436,1607,1716]" pageId="1" pageNumber="595">
20 Paramere with pair of slender coiled processes............................................................................................
<taxonomicName id="6C8B93DBFFD5C637E1E9F4482DEFF9A4" authority="Theischinger" authorityName="Theischinger" box="[1205,1436,1607,1628]" class="Insecta" family="Thaumaleidae" genus="Austrothaumalea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="595" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="99FF344AFFD5C637E1E9F4482D6AF9A4" box="[1205,1305,1607,1628]" italics="true" pageId="1" pageNumber="595">A. simplex</emphasis>
Theischinger
</taxonomicName>
- Paramere without pair of coiled processes ................................................................................................................................... 20a 20a Paramere with subapical, lateral expansions, spearhead-like in appearance ....................................................
<taxonomicName id="6C8B93DBFFD5C637E18BF48D2D3AF96F" box="[1239,1353,1666,1687]" class="Insecta" family="Thaumaleidae" genus="Austrothaumalea" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="595" phylum="Arthropoda" rank="species" species="spiculata" status="sp. nov.">
<emphasis id="99FF344AFFD5C637E18BF48D2D3AF96F" box="[1239,1353,1666,1687]" italics="true" pageId="1" pageNumber="595">A. spiculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="82CC8931FFD5C637E013F48D2DEFF960" box="[1359,1436,1666,1688]" pageId="1" pageNumber="595" rank="species">
<emphasis id="99FF344AFFD5C637E013F48D2DEFF960" bold="true" box="[1359,1436,1666,1688]" pageId="1" pageNumber="595">sp. nov.</emphasis>
</taxonomicNameLabel>
- Paramere simple, straight, lacking expansions or processes .......................................................................................................... 21
</paragraph>
</subSubSection>
</treatment>
</document>